1 Multilocus phylogeny of the avian family Alaudidae (larks) 2 reveals complex morphological evolution, non- 3 monophyletic genera and hidden species diversity 4 5 Per Alströma,b,c*, Keith N. Barnesc, Urban Olssond, F. Keith Barkere, Paulette Bloomerf, 6 Aleem Ahmed Khang, Masood Ahmed Qureshig, Alban Guillaumeth, Pierre-André Crocheti, 7 Peter G. Ryanc 8 9 a Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese 10 Academy of Sciences, Chaoyang District, Beijing, 100101, P. R. China 11 b Swedish Species Information Centre, Swedish University of Agricultural Sciences, Box 7007, 12 SE-750 07 Uppsala, Sweden 13 c Percy FitzPatrick Institute of African Ornithology, DST/NRF Centre of Excellence, 14 University of Cape Town, Rondebosch 7700, South Africa 15 d Systematics and Biodiversity, Gothenburg University, Department of Zoology, Box 463, SE- 16 405 30 Göteborg, Sweden 17 e Bell Museum of Natural History and Department of Ecology, Evolution and Behavior, 18 University of Minnesota, 1987 Upper Buford Circle, St. Paul, MN 55108, USA 19 f Percy FitzPatrick Institute Centre of Excellence, Department of Genetics, University of 20 Pretoria, Hatfield, 0083, South Africa 21 g Institute of Pure & Applied Biology, Bahauddin Zakariya University, 60800, Multan, 22 Pakistan 23 h Department of Biology, Trent University, DNA Building, Peterborough, ON K9J 7B8, 24 Canada 25 i CEFE/CNRS Campus du CNRS 1919, route de Mende, 34293 Montpellier, France 26 27 * Corresponding author: Key Laboratory of Zoological Systematics and Evolution, Institute of 28 Zoology, Chinese Academy of Sciences, Chaoyang District, Beijing, 100101, P. R. China; E- 29 mail:
[email protected] 30 1 31 ABSTRACT 32 The Alaudidae (larks) is a large family of songbirds in the superfamily Sylvioidea.