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PRESIDENT OF THE Stephen C. Kowalczykowski Economic Sciences Immunology and Plant Biology ACADEMY Jim Manley Anne C. Case Inflammation David Baulcombe Marcia K. McNutt Kiyoshi Mizuuchi Matthew O. Jackson Robert L. Coffman Philip N. Benfey Brenda A. Schulman Jose A. Scheinkman Peter Cresswell Dominique Bergmann James A. Wells K. Christopher Garcia Gloria M. Coruzzi EDITOR-IN-CHIEF Engineering Sciences Lewis L. Lanier Joseph R. Ecker May R. Berenbaum Biophysics and Alexis T. Bell Tak Wah Mak Stephen P. Long June B. Nasrallah Computational James J. Collins Carl F. Nathan Pablo G. Debenedetti Krishna K. Niyogi ASSOCIATE EDITORS Biology Lawrence Steinman Evelyn L. Hu Tadatsugu Taniguchi Julian I. Schroeder Huda Akil David Baker Douglas E. Soltis Angela M. Gronenborn Yonggang Huang Arthur Weiss Adriaan Bax Rakesh K. Jain Philippa Marrack Axel T. Brunger Plant, Soil, and Microbial Catherine J. Murphy Klavs F. Jensen Mathematics William F. DeGrado Sang Yup Lee Sciences Dolores R. 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