Rettili Marini Mesozoici

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Rettili Marini Mesozoici 01/06/2011 Rettili Marini Mesozoici Se la pressione selettiva è favorevole, i Rettili ritornano all’acqua. BASSO tasso metabolico Adattamento TOLLERANZA all’anossia favorito da CAPACITA’ di sfruttare il METABOLISMO ANAEROBICO I gruppi di Rettili marini Mesozoici più importanti sono due: Ichthyosauria Sauropterygia Placodontia “Nothosauria” Plesiosauria 1 01/06/2011 ANAPSIDE SINAPSIDE DIAPSIDE “Parapside” EURYAPSIDE ICHTHYOSAURIA Dal Triassico Inferiore al Cretacico Medio-Superiore Derivazione diapside? Cranio simile a quello euryapside, evolutosi indipendentemente ? Finestra temporale superiore Ittiosauro primitivo? Diapside primitivo? Difficile stabilire l’entità delle convergenze Nanchangosaurus (Hupehsucus) Triassico, Cina 2 01/06/2011 Utatsusaurus Triassico inferiore, Giappone Archi neurali ancora connessi con i centri no processi trasversi “mano simile a pinna ma struttura da tetrapode terrestre. Grippia Triassico inferiore, Spitzbergen Contatto prefrontale-postfrontale Dentatura eterodonte Chaohusaurus Triassico inferiore, Cina Denti su due file http://www.ucmp.berkeley.edu/people/motani/ichthyo/ Ichthyosauria Ittiosauri post -Triassico Ittiosauri triassici 3 01/06/2011 Robusto anello sclerotico Pinna caudale e dorsale senza supp. scheletrico Natatoia, iperdattilia e iperfalangia Centri vertebrali discoidali anficeli •Accorciamento porzioni prossimali dell’arto •Perdita di identità elementi carpo metacarpo •Iperfalangia ed iperdattilia •Perdita mobilità articolare fra le diverse sezioni dell’arto 4 01/06/2011 5 01/06/2011 6 01/06/2011 Cymbospondylus 7 01/06/2011 SAUROPTERYGIA: Nothosauria (Triassico) Nothosauroidea Pachypleurosauroidea La scapola si infila ventralmente alla clavicola Il cinto forma un anello ventrale Pachypleurosauroidea Nothosauroidea 8 01/06/2011 Ceresiosaurus 2m circa I Nothosauroidea presentano un trend evolutivo che va verso un progressivo irrigidimento del tronco ed un passaggio del nuoto da assiale ondulatorio verso un nuoto parassiale simmetrico che si riscontrerà nei Plesiosauria. Numerosi caratteri anatomici ne farebbero gli antenati diretti dei Plesiosauria, ma un carattere morfologico, costituisce un ostacolo. I Nothosauria presentano nel palato un carattere derivato assente nei Plesiosauria: le finestre interpterigoidee sono chiuse, mentre nei Plesiosauria rimane la condizione plesiomorfica, ossia le finestre interpterigoidee sono aperte. 9 01/06/2011 Pistosaurus Questo problema è stato risolto con il ritrovamento di Pistosaurus, un Sauropterigio che mostra uno scheletro postcraniale da Nothosauroidea con un cranio da plesiosauro, con finestre interpterigoidee aperte. Pachypleurosauroidea Nothosauroidea Pistosaurus Plesiosauria PLESIOSAURIA (?Triassico Sup. -Cretacico) Notevole adattamento al nuoto Arti anteriori e posteriori simili fra loro Coda non molto lunga Plesiosauri Tronco completamente rigido Cinti “bassi” ma molto espansi ventralmente Dimensioni generalmente superiori ai notosauri 10 01/06/2011 Arti trasformati in natatoie Cinto pettorale robusto ed espanso ventralmente Coda breve non modificata per il nuoto Gastralia che contribuiscono ad irrigidire il tronco ELASMOSAUROIDEA Estremo allungamento del collo, oltre 70 vertebre cervicali 11 01/06/2011 Pliosauroidea 12 01/06/2011 Cranio molto grande con fauci allungate ed armate di robusti denti conici, situati all’estremità anteriori delle mascelle. Lunghezza da due a oltre venti metri. Probabilmente veloci (e voraci) predatori di mare aperto. Neusticosaurus “Notosauro” Plesiosauro Plesiosauro Giurassico Inferiore Giurassico Superiore 13 01/06/2011 Vertebre: •Incremento dimensioni processi trasversi •Avvicinamento e riduzione zigapofisi Pistosaurus Neusticosaurus Cryptocleidus Tartaruga Plesiosauro Otaria Tartaruga Plesiosauro Pinguino 14 01/06/2011 PLACODONTIA Triassico Medio-Superiore Placodontoidea Coda lunga ma Placodontia non compressa Cyamodontoidea lateralmente Arti non modificati per il nuoto Cinto pettorale, vista ventrale Sezione frontale tronco Cinto pelvico vista ventrale •Ampia finestra temporale •Robusta barra sq-j-po (muscolatura mandibolare) •Denti trituranti 15 01/06/2011 Cyamodontoidea Psephoderma, Triassico Superiore Psephoderma Nei ciamodontoidi evoluti si ha scomparsa dei denti premascellari durante l’ontogenesi 16 01/06/2011 Henodus, Triassico Superiore 17.
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