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Teil 12A / Part 12A I , Pachypleurosauria, Nothosauroidea,

by

O. RlEPPEL

With 80 Figures

Verlag Dr. Friedrich Pfeil • Munchen 2000 ISBN 3-931516-78-4 Contents

Foreword (P. WELLNHOFER) V Acknowledgements VI Institutional Acronyms VII Figure Abbreviations VIII Introduction: History of the Concept of Sauropterygia 1 Phylogenetic Relationships of Stem-Group Sauropterygia 4 Stratigraphic and Geographic Distribution of Stem-Group Sauropterygia 6 General Skeletal Anatomy of Stem-Group Sauropterygia 10

Systematic Review 16 Superorder Sauropterygia OWEN, 1860 16 Order Placodontia COPE, 1871 16 Suborder Placodontoidea COPE, 1871 16

Family Paraplacodontidae PEYER & KUHN-SCHNYDER, 1955 17 Anatomy of Paraplacodontidae 17 PEYER, 1931 18 Family Placodontidae COPE, 1871 19 Anatomy of Placodontidae 19 Genus AGASSIZ, 1933 21 Suborder NOPCSA, 1923 23 Anatomy of Cyamodontoidea 23 Interrelationships of Cyamodontoidea 25 Superfamily Cyamodontida NOPCSA, 1923 25 Family Henodontidae F. v. HUENE, 1948 26 Genus F. v. HUENE, 1936 26 Family Cyamodontidae NOPCSA, 1923 27 Genus Cyamodus MEYER, 1863 27 Superfamily Placochelyida ROMER, 1956 32 Family Macroplacidae nov. fam 32 Genus Macroplacus SCHUBERT-KLEMPNAUER, 1975 32 Family Protenodontosauridae nov. fam 33 Genus Protenodontosaurus PINNA, 1990 33 Family ROMER, 1956 34 Genus JAEKEL, 1902 34 Genus MEYER, 1858 36 Genus Psephosaurus E. FRAAS, 1896 38 Cyamodontoidea indet 39

IX Order Eosauropterygia RIEPPEL, 1994 41

Suborder Pachypleurosauria NOPCSA, 1928 41 Family Pachypleurosauridae NOPCSA, 1928 41 Anatomy of Pachypleurosauridae 42 Interrelationships of Pachypleurosauridae 44 Genus DAMES, 1890 45 Genus Dactylosaurus GURICH, 1884 46 Genus Hanosaurus YOUNG, 1972 49 Genus YOUNG, 1958 50 Genus SEELEY, 1882 51 Genus RIEPPEL, 1989 56 Pachypleurosauria incertae sedis 57 Suborder Eusauropterygia TSCHANZ, 1989 58 Infraorder Nothosauroidea BAUR, 1889 58 Superfamily Simosauria nov. superfam 58 Family Simosauridae F. v. HUENE, 1948 58 Anatomy of Simosauridae 58 Genus MEYER, 1842 61 Superfamily Nothosauria BAUR, 1889 64 Family Germanosauridae nov. fam 64 Genus Germanosaurus NOPCSA, 1928 64 Family BAUR, 1889 66 Genus MUNSTER, 1834 66 Genus CURIONI, 1847 86 Nothosauria incertae sedis 96 Infraorder Pistosauroidea BAUR, 1887-90 97 Superfamily Cymatosauria E.v. HUENE, 1944 97 Family Cymatosauridae E.v. HUENE, 1944 97 Anatomy of Cymatosauridae 97 Genus CASE, 1936 98 Genus FRITSCH, 1894 98 Superfamily Pistosauria BAUR, 1887-90 104 Family Pistosauridae BAUR 1887-90 105 Genus SANDER, RIEPPEL & BUCHER, 1997 105 Genus MEYER, 1839 105 Pistosauroidea incertae sedis 106 Genus Chinchenia YOUNG, 1965 107 Genus Kwangsisaurus YOUNG, 1959 107 Problematic taxa of unknown affinities 108 Taxa which have been placed within the Sauropterygia, but must be excluded 109 A list of names currently recognized as nomina dubia 110

Functional Anatomy of Stem-Group Sauropterygia Ill Life History of Triassic Stem-Group Sauropterygia 114 Evolution of Triassic Stem-Group Sauropterygia 114 Historical Biogeography of Triassic Stem-Group Sauropterygia 116 Avenues for Future Research 119 Literature Cited 120 Index of Genera and Higher Taxa 133