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The Malacological Collection from Riparo Dalmeri (Trento)

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The Malacological Collection from Riparo Dalmeri (Trento) The malacological collection from Riparo Dalmeri (Trento) GIAMPAOLO DALMERI & CHIARA FIOCCHI - ABS1RACT The collection is composed of twenty-five shells and has been studied from a paleontological and palethnological point of view. A total of five species have been determined which are all present in the modern fauna in the Mediterranean area and which are probably contemporary with the age of the site. An ornamental use is very probable because there are a lot of perforations which have also been studied using both a stereornicroscope and a scanning electron microscope. Comparisons were made with other malacological Keycollections words: from Epigravettian sites on the southern slopes of the Eastern Alps. Parole chiave: Riparo Dalmeri, Veneto, Final Epigravettian, Malacology, Ornaments Riparo Dalmeri, Veneto, Epigravettiano recente, Malacologia, Ornamenti Giampaolo Dalmeri - Chiara Fiocchi - Museo Tridentino di Scienze Naturali, Via Calepina 14, 1-38100 Trento Dip. Scienze Geologiche e Paleontologiche, Universita di Ferrara, Corso Ercole I d'Este 32, 1-44100 Ferrara. E-mail: fcc @dns.unife.it 1. THE FAUNA AND THE STATE OF PRESER­ cklist of the Italian fauna species has been used for the VATION general systematic outline. Class: Gastropods Subclass: Prosobranchia The malacological collection found in the final Superorder: Caenogastropods Epigravettian levels at Dalmeri rockshelter, at the end Order: Neogastropds of the excavation campaign which took place in the Family: Buccinidae year 1997, is composed of twenty-five mollusca shells Genus: Cyclope Risso, 1826 belonging to the Class of Gastropods. Five species were determined; two of these are in open nomenclature: Cyclope sp. Cyclope sp., Cyclope neritea (LlNNAEus, 1758), Cyclo­ Material: 1 shell. pe pellucida Risso, 1826, Columbella rustica (LINNA­ Observations: this is a fragment of only part of the EUS, 1758) and Mitra sp. (Fig.1). The shells are preser­ columella callus and of the columella, ved quite well and only a low percentage is fragmented so it was impossible to determine the (Fig.2). The Columbella rustica specimens are slightly exact species that it belongs to. decorticated, while the Cyclope shells present more evident alterations of the surface. Traces of the origi­ Cyclope neritea (LINNAEUS, 1758) nal colour are present in one of the specimens of Cyclope Material: 5 shells, four of which were almost neritea which features a flamey brown colour. completely intact. Habitat: this species lives on sandy mobile su­ bstrata on the intercoastal area. It is an euryhaline species diffused in estua­ 2. OBSERVATION ON THE SPECIES ries and brackish lagoon enviroments, belonging to the Bio-complexes of the A systematic general outline of the identified Euryhaline and Eurytherm Lagoons species, the number of samples found, the habitat and (PEREs & PICARD, 1964). the geographic distribution hereby follows. The classi­ Geographic fication presented by BoooN et al. (1995) in the Che- distribution: diffusedin allof the Mediterraneanarea. 186 I I Cyclope pellucida Risso, 1826 Apart from just the one fragment belonging to Material: 2 shells, one of which is almost com­ the Mitra sp., whose form vaguely reminds us of the iI I pletely intact. Columbella rustica, only the Cyclope and Columbella Habitat: lives in the mobile substrata on the top species were present. Due to the fact that only two spe­ level of the intercoastal area, associa­ cies were found, it would seem that they were purpo­ ted with the bio-complexes of the san­ sely chosen by man and were probably gathered along ds which are quite protected from the the beaches. The two species have, in fact, very distin­ breaking of the waves (PER.Es & Pr­ ct forms: low orbicular (genus Cyclope) or spiral (ge­ CARD, 1964).This species is also pre­ nus Columbella and Mitra.) forms. In natural condi­ sent in the bio-complexes of fine su­ tions the shells are shiny with a porcelain-coloured perficial layers of sands at a depth of smooth surface and their colour is characterised by 0 to 2.5cm (PER.Es & PrcARD, 1964; blotches and small flames. Neither of the species show SABELLI & SPADA, 1977). In waters with evident signs of ornaments or sculpture. Both the spe­ a low content of salt it is substituted cies have a smooth surface. by C. neritea (LINNAEus, 1758). Geographic distribution: species common in all of the Mediter­ ranean area. 4. THE PERFORATIONS Family COLUMBELLIDAE Each of the fourteen specimens have one perfo­ Type Columbella Lamarck, 1799 ration: Cyclope neritea (one perforation), Cyclope pel­ lucida (two), and Columbella rustica (eleven). These Columbella rustica (LINNAEUS, 1758) perforations are always located near the border of the Material: 16 shells, ten of which were almost labium and have a subcircular outline and irregular completely intact. margins with very sharp edges (Fig.3). Three hypothe­ Habitat: lives on the intercoastal levels, either ses have been put forward to explain the origin of these under stones or associated with algae perforations: they could be fractures caused by the and Zosteracae (SABELLI & SPADA, impact on hard surfaces due to the waves, or traces of 1986). predators such as Crustacean Decapods (RoBBA & Geographic OsTINELLI, 1975), or otherwise they could be made by distribution: species abundant in all of the Mediter­ man (TABORIN, 1993a,1 993b). Experiments carried out ranean area. show that if the perforations result from anthropic ac­ tivity, pressure and indirect percussion techniques must Family MITRIDAE have been used to produce similar perforations (FRAN­ Type Mitra Lamarck, 1798 crs, 1982; TABoRIN, 1993a). Analyses carried out using both a stereomicroscope and a scanning electron mi­ Mitra sp. croscope have not produced conclusive results in di­ Material: 1 shell. stinguishing how the perforations came about since there Observations: part of the last circle is missing and are no traces (Fig.3). The shells could have been used the columella is exposed showing the for ornamental reasons. Perhaps they were already per­ characteristic columella pleats. forated, or they were perforated later. The stereomi­ croscopic analysis showed that there were traces of a reddish colouring which was probably ochre, along the sutures and inside some of the specimens. 3. THE SELECTION AND THE POSSIBLE PROVENANCE All the species that were determined belong to 5. COMPARISONS the modern fauna present in the Mediterranean area (e.g. BonoN et al., 1955; PoPPE & GoTo, 1991; Cossr­ Parallelisms can be found in the molluscfauna GNANI D'ANGELO G et al. , 1992; & GAR IULLO, 1981). The of Dalmeri rockshelter and the collections recovered absence of fo ssil species and the well-conserved speci­ on three Epigravettian sites on the Southern slopes of mens suggest that the shells are contemporary with the the EasternAlps: Villabruna rockshelter A (Sovramon­ age of the site, although the absence of radiocarbon te-BL), Tagliente rockshelter (Stallavena di Grezzana­ dates of the shells does not allow to confirm this VR), and Biarzo rockshelter (Biarzo-UD) (Fig.4). Four hypothesis. marine shells were found at Riparo Villabruna A (Ar- 187 MAR et al. , 1994) respectively from the subunits 9A, 5, terials. The limited amount of species present indica­ 4A and from the rearranged deposit. Three of these tes that man chose the shells when gathering them, belong to the Class of Gastropods: two specimens of considering that the thanato-complexes of the shell Columbella rustica (LINNAEus, 1758) and one speci­ detritus on the shores or the fossil finds were usually men of Mitra sp. One of the shells has been attributed more varied. The fact that only a few species with to the Class of Scafopods, Dentalium cf. inaequico­ particular morphologies were gathered is obviously statum Dautzemberg, 1891 species. related to their use. The shells were used mainly for 127 specimens were identified at Tagliente their symbolic significance. Each different shape was rockshelter belonging both to the Class of Gastropods probably attributed with a different significance, as and the Class of Bivalves (BENINIAccoRsr, 1974). Two for example the family origin or the social class. The species of the Class of Gastropods were identified, one shells could have also been used as amulets and have of which is in open nomenclature: a specimen of Aphor­ a magic significance. rais sp. and 124 specimens of Cyclope neritea (llNNE­ The specimens were found in sites far from the AUS, 1758). Two specimens belonging to the Class of sea and could have been obtained in two different ways: Bivalves were identified, attributed to the Glycymeris special organized journeys from the site to the gathe­ sp. in open nomenclature. All the shells belonging to ring area or through exchange. Considering that the the Cyclope type feature a perforation near the border present distance from Dalmeri rockshelter to the nea­ of the labium, while in the Glycymeris sp. and in the rest north Adriatic coast is about 80Km, the journey Aporrhais sp. there is evidence of polishing as well was very long. During the Tardiglacial period, it was (BENINIAccoRsr, 1974). During excavation campaigns even further away because the coasts were much lower which took place after 1973, many other shells were and were -60 m from the present sea level as estimated found (Guerreschi, pers. corn.). by many authors, (e.g. LABEYRIE et al. , 1987; Shackle­ Thirteen shells and two fragments belonging to ton, in BARD et al. , 1990; FAVERO , 1984). the Class of Gastropods and to the Class of Bivalves The specimens
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