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Begun in 1895
OLUME 106, NUMBER 344 APRIL 5, 1994
Neogene Paleontology in the Northern
Dominican Republic 15.
The Genera Cohimbella, Eurypyrene,
Parametaria, Conella, Nitidella, and
Metulella (Gastropoda: Columbellidae)
by
Peter Jung
Paleontological Research Institution 1259 Irumansburg Road Ithaca, New York, 14850 U.S.A. J PALEONTOLOGICAL RESEARCH INSTITUTION
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(Permanence of Paper). @ This paper meets the requirements of ANSI/NISO Z39.48-1992 'BufOti^ of
Begun in 1895
OLUME 106, NUMBER 344 APRIL 5, 1994
Neogene Paleontology in the Northern
Dominican Republic 15.
The Genera Columbella, Eurypyrene,
Parametaria, Conella, Nitidella, and
Metulella (Gastropoda: Columbellidae)
by
Peter Jung
Paleontolofjical Research Institution 1259 Trumansburg Road Ithaca, New \ork. 14850 U.S.A. ISSN 0007-5779 ISBN 0-87710-432-8 Lihraiy of Congress Catalog Card Number: 94-65486
Printed in the United States of Amenca Allen Press, Inc. Lawrence, KS 66044 U.S.A. CONTENTS
Page Abstract 5 Resumcn 5 Introduction 5 Acknowledgements 5 Biostratigraphy and Paleobiogeography 6 Abbreviations of Repository Institutions 8 Systematic Paleontology Introduction 8
Genus Columhella Lamarck, 1 799 11 C. suhnu'irali^nu Olsson, 1922 11 C aff. nislianik's Heilpnn, 1887 12 C. lopczana. new species 13 Genus Eurypyrcnc Woodrmg, 1928 14 E. injlala (Gabb, 1873b) 14 E. eurynoluin (Woodrmg. 1928) 15 Genus Parametaria Dall, 1916 16 Subgenus Dominilana new subgenus 16 P. (D.) islahispaniolac (Maury, 1917a) 17 P. (D.) lopezana. new species 19
Genus Conclla Swainson, 1 840 20
C. perplcxabilis (Maury, 1 9 1 7a) 20
Genus Nilidclla Swainson. 1 840 21
N. cibaoica (Maury. 1 9 1 7a) 22
N. aff, cihaoica (Maury, 1917a) 24 N. caimilana. new species 24 Genus Melulclla Gabb, 1873a 25
M. fusiformis Gabb, 1 873a 25
M. venusta (G. B, Sowerby II, 1850) 27 References Cited 30 Plates 32 Index 43 LIST OF ILLUSTRATIONS
Text-figure Page
1. Index map showing location of investigated areas in the Cibao Valley. Dominican Republic 6 2. Columnar section of Rio Gurabo showing (discontinuous) "ranges" of species discussed herein 7 3. Columnar section of Rio Cana showing (discontinuous) "'ranges" of species discussed herein 8 4. Columnar section of cliff exposures on Rio '>aque del Norte near Lope/, north of Baitoa. showing "ranges" of species discussed herein and stratigraphic positions of NMB localities 8 5. Schematic column for Rio \aque del Norte showing "ranges" of species discussed herein and relative stratigraphic positions of NMB localities 9
6. Section exposed in Maury's Bluff 2 on Rio Mao showing "range" of Paramelaria Dominitana) islahispamolae (Maury, 1 9 1 7a) and stratigraphic positions of NMB localities 9 7. Section exposed at the downstream (eastern) end of Maury's Bluff 3 on Rio Mao showing "ranges" of species discussed herein and stratigraphic positions of NMB localities 10 8. Section exposed at mouth of Arroyo Bajon on Rio Mao showing (discontinuous) "ranges" of species discussed herein and stratigraphic positions of NMB localities 10
9. Schematic colums for the central ponion of Rio Amina showing "range" of Melu/ella venusla (G. B. Sowerby II, 1850) and relative stratigraphic positions of NMB and TU localities. II 10. Paramelaria (Dominitana) islahispamolae (Maury. 1917a). NMB H 17412. NMB locality 16836: Rio Cana section, Dominican Republic; lowest part of Gurabo Formation (late Miocene) 17
11. Paramelaria (Dominilarial islahispamolae (Maury . 1917a). NMB H 17413. NMB locality 16828: Rio Cana section, Dominican Republic; upper part of Cercado Formation (late Miocene) 18
1 2. (Restored) height/width diagram of Paramelaria (Dominilaria) islahispamolae (Maury, 1 9 1 7a) 19
1 3. (Restored) height/width diagram of Conella perplexahilis (Maury , 1 9 1 7a) 21 14. Nilidella cibaoica (Maury, 1917a). NMB H 17421. NMB locality 16934: Rio Gurabo section, Dominican Republic; lower part of Gurabo Formation (late Miocene). 22 15. Nitidella nitida (Lamarck. 1822). NMB H 17422. NMB locality 17704: Isla de Aves, Venezuela: Recent 23
1 6. (Restored) height/width diagram of SiliJella cibaoica (Maury. 1 9 1 7a) 24
1 7. Melulella fusiformis Gabb. 1 873a. NMB H 1 7429. NMB locality 1 7266; La Barranca of Rio Yaque del Norte, Dominican Republic; upper part of Globorolalia margarilae zone (late early Pliocene) 26 18. (Restored) height/width diagram of Melulella fusiformis Gabb. 1873a 27
1 19. Melulella vemisla (G. B. Sowerby 11. 1 850). NMB H 743 1 . NMB locality 1 5903: Rio Gurabo section, Dominican Republic; Cercado Formation (late Miocene) 28
20. (Restored) height/width diagram of Melulella venusta (G. B. Sowerby II, 1850) 29
LIST OF TABLES
Table Page
1. Numbers of lots and specimens of each of the 1 3 species of columbellid gastropods available for this study 11 2. Measurements of Columbclla submercalona Olsson. 1922 12 3. Measurements of Columbella aff ruslicoides Heilprin, 1887 13 4. Measurements of Columbella lopezana. new species 15
5. Measurements of Eurypyrene inflala (Gabb, 1 873b) 15 6. Measurements of Eurypyrene eurynolum (Woodring, 1928) 16 7. Measurements of Paramelaria (Dominilaria) lopezana, new species 20 8. Measurements of Nilidella caimilana. new species 25 NEOCENE PALEONTOLOGY IN THE NORTHERN DOMINICAN REPUBLIC 15. The Genera Columhella, Eitrypyrene, Parametaria. Conella, Nitidella, and Metulella (Gastropoda: Columbellidae)
Peter Jung Naturhistorisches Museum Augustinergasse 2 CH-4051 Basel Switzerland
ABSTRACT
Thirteen species of columbellid gastropods belonging to six genera are descnbed and figured. Two forms have been identified by means of
"open nomenclature" thus leaving a total of I 1 positively identified species. Three species, Colunihella lopczana and Paramelana iDominilana) topezana. both from the late early to early Middle Miocene Baitoa Formation of the Lopez section on Rio Yaque del Norte, and Nilidella caimitana from the Late Miocene Cercado Formation of the Rio Cana section, and one subgenus, Paramelana (Dommitanal. are descnbed as new.
Although the available material is not rich, the stratigraphic occurrences of these species are discussed. Only two species are well represented; three are moderately well represented, and the remaining species are represented by five or fewer specimens and thus must be considered rare,
RESUMEN
Se han descrito y representado trece especies de gasteropodos in la familia Columbellidae pertenecientes a seis generos. Dos formas han sido identificadas por medio de nomenclatura abierta. dcjando asi un total de one especies positi vamente idcntificadas. Tres especies, Coluinbella lopezana y Paramelana IDominilana) lopezana del Mioceno temprano tardio a medio temprano, Formacion Baitoa de la seccion Lopez in Rio Yaque del Norte, Nilidella caimitana de finales de Mioceno, Formacion Cercado de la seccion del Rio Cana, y un subgenero, Parameiaria (Dominitaria). han sido descritos como nueves.
A pesar de no disponer de amplio matenal, la ocurencia estratigrafica de estas especies es discutida. Solo hay dos especies bien representadas; tres moderadamente representadas, y las expecies restantes son representadas por cinco o menos especimenes y por lo tanto tienen que ser consideradas como raras.
INTRODUCTION 1979, and 1980. The field work was made possible by a grant from the Swiss National Science Foundation This paper is a further contribution to the series of (Grant 2.646-0.76). The financial help and the assis- taxonomic studies dealing with Neogene fossils from tance in the field provided by Institut Francais du Pe- sections situated in the Cibao Valley of the northern trole are gratefully acknowledged. Dominican Republic (Text-fig. 1), The project and the I am indebted to the following persons for the loan framework within which these studies are being carried of specimens under their care: Messrs. Frederick out have been outlined by Saunders et a/. (1982) and J. Collier and Warren Blow, National Museum of Nat- Saunders et al. ( 1 986). Jung ( 1 986, p, 5) listed the most important early collections of molluscs from this area. ural History, Washington, DC, U.S.A.; Dr. Peter Hoo- As is the case for all the contributions to this series, ver, Paleontological Research Institution, Ithaca, NY, the material studied has been collected from measured U.S.A.; Dr. Emily Yokes, Tulane University. New Or- sections. The geographic location of the investigated leans, LA, U.S.A.; Dr. Yves Finet, Natural History areas is shown in Text-figure 1. For detailed infor- Museum, Geneva, Switzerland; Mr, C. P. Nuttall, The mation as to geographic locations and stratigraphic Natural History Museum. London. England; Dr. Ken- position of all the collecting stations, as well as to the neth J, Boss, Museum of Comparative Zoology, Cam- general biostratigraphic framework and the ages, the bridge, MA, U,S.A.; Ms. Elana Benamy. Academy of reader is referred to the paper by Saunders et al. (1986). Natural Sciences. Philadelphia, PA, LI.S,A,
Formational names have been used with care, because In addition I am grateful to Mr. Wolfgang Suter, correlations from section to section are not certain. photographer (now retired) at the Naturhistorisches Museum Basel as well as to Dr. Richard Guggenheim ACKNOWLEDGEMENTS and Mr. Marcel Diiggelin, both of the Scanning Elec-
The material on which this paper is based was col- tron Microscope Laboratory, University of Basel, lected during field work carried out in the years 1978, Switzerland. Bulletin 344
Text-figure 1. — Index map showing location of investigated areas in the Cibao Valley, Dominican Republic (after Jung, 1986, text-fig. 1).
BIOSTRATIGRAPHY AND Eurypyrene inflata (Gabb, 1873b) PALEOBIOGEOGRAPHY Parametaria lopezana. n. sp.
A total of 13 columbellid species belonging to six La Barranca on Rio Yaque del Norte (Text-fig. 5): genera is discussed in this paper. Two forms have been Eurypyrene eurynot um (Woodring, 1928) identified by means of open nomenclature (Richter, Metulella fusiformis Gabb, 1873a 1943, 1948; Matthews, 1973; Komicker, 1979; Bengs- Metulella venusta (G. B. Sowerby II, 1850) ton, 1988) which leaves a total of 1 1 positively iden- Rio Mao section (Text-figs. 6-8): tified species. Parametaria islahispaniolae (Maury, 1 9 1 7a) The stratigraphic "ranges" of these species— or rath- Conella perplexabilis (Maury, 1 9 1 7a) er their stratigraphic occurrences are plotted in Text- — Metulella venusta (G. B. Sowerby II, 1850) figures 2 to 9. As pointed out by Jung and Petit (1990, Arroyo Zalaya section (Saunders et al., 1986, text-fig. p. 88) the species are not continuously present through 36): a given sequence of sediments, rather their occurrences Metulella fusiformis Gabb. 1873a are spotty. Several species occur only in one of the sections considered herein; others have been found in Rio Amina section (Text-fig. 9): more than one section. The distribution of species in Metulella venusta (G. B. Sowerby II. 1850) the various sections is as follows: As found by Jung and Petit (1990, p. 88) for can- Rio Gurabo section (Text-fig. 2): cellariids, the distributional patterns of the columbellid Columbella submercatoria Olssen, 1 922 species discussed herein reflect a considerable degree Nitidella cibaoica (Maury, 1917a) of stratigraphic restriction. Seven of the 1 1 positively Metulella venusla (G. B. Sowerby II, 1850) identified species, i.e.. more than 60% of the discussed Rio Cana section (Text-fig. 3): fauna, have been found in single sections only. This is Parametaria islahispaniolae (Maury, 1917a) a large percentage considering that the sediments of all Nitidella cibaoica (Maury, 1 9 1 7a) the sections in which these species occur are of similar Nitidella caimitana, n. sp. age (late Miocene to early Pliocene) with the exception Metulella venusta (G. B, Sowerby II, 1850) of the Lopez section which is of late early to early Lopez section of Rio Yaque del Norte (Text-fig. 4): middle Miocene age (Saunders et al.. 1986, p. 30). Columbella lopezana, n. sp. Of the seven species restricted to single sections, Dominican Columbellid Gastropods: Jung
three are known from only one locality within the Do- 3 minican Republic: Eurypyrene inflata (Gabb, 1873b) c > is recorded only from NMB locality 17290: Baitoa ffl Formation (late early to early middle Miocene) of the Lopez section on Rio Yaque del Norte (Text-tig. 4); Eurypyrene eurynotum (Woodring, 1928) from TU lo- cality 1404: Arroyo Babosico near La Barranca on Rio
Yaque del Norte (Text-fig. 5), upper part of Globoro- talia margaritae zone (late early Pliocene); and Niti- della cainiilana, n. sp. from TU locality 1230: Cercado ^ '' ~ Formation (late Miocene) ofthe Rio Cana section (Text- 3 fig. 3). The remaining four species which are restricted to single sections are distributed as follows: — Columbella submercatoria Olsson, 1922 has been found at two localities (NMB 1 59 1 0, 1 59 1 2) in the Rio .< -v.-^ Gurabo section: upper part of Cercado Formation (late Miocene). See Text-figure 2. — Columbella lopezana, n. sp. is recorded from four localities (NMB 16935, 17288, 17289, 17290) in the Lopez section on Rio Yaque del Norte: Baitoa For- mation (late early to early middle Miocene).
— Parametaria lopezana, n. sp. is recorded from three localities (NMB 16936, TU 1363, TU 1364) in the Lopez section on Rio Yaque del Norte: Baitoa For- mation (late early to early middle Miocene). See Text- figure 4.
— Conella perplexabilis (Maury, 1 9 1 7a) is known from a number of localities in the Rio Mao section: Cercado Formation (late Miocene). See Text-figures 7 and 8. In addition to the seven species with restricted dis- tribution as pointed out above, there are four species which occur in more than one section. The most wide- ^"zr spread species is Metulella venusta (G. B. Sowerby H,
1 850) which has been found in beds cropping out along Rio Cana, Rio Gurabo, Rio Mao, Rio Amina, and Rio
Yaque del Norte (Text-figs. 2, 3, 5, 9). It is followed by Parametaria islahispaniolae (Maury, 191 7a) which is known from a number of localities in the sections of Rio Cana and Rio Mao (Text-figs. 3, 6, 7, 8).
Out of the 1 1 positively identified species eight are known only from the Dominican Republic. This fact points to a remarkable endemism. The remaining three species are also known from other countries. They are:
— Columbella submercatoria Olsson, 1922 is recorded from the late Miocene Cercado Formation of the Do- minican Republic, but is also known from the early Pliocene Bowden Formation ofJamaica and from beds of late Miocene or early Pliocene age in Costa Rica.
Text-figure 2. — Columnar section of Rio Gurabo showing (dis- continuous) "ranges" of species discussed herein (after Saunders et ai. 1986, text-fig.6). Numbers in second column from left refer to thickness in m. Bulletin 344
m o 'c a: (0 o D- U. 400 in
E 10 (0 C < >CD a: JO =) 19 3 CD Dominican Columbellid Gastropods: Jltng
jANriAt.O
Silii .iti.l I Hjv* E
I f:-^=ei7 U31b — I
1 7J'J1 .^ .— I :
Ili'l17 1 7r90 -".LP *>)
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! ? J'Jb
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UJ
W.'HH ' I /l.'-I
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Text-figure 4— Columnar section of cliff exposures on Rio Yaque del Norte near Lopez, north of Baitoa. showing "ranges" of species discussed herein and stratigraphic positions of NMB localities (after
Saunders el al., 1986, text-fig. 25). 10 Bulletin 344
Young ytavel terrace
lell lieiJ
(D
1G912 • IS fallen tubhie thought to be tfoin this beil
(D JO o c (D WQ. !c m 10
S E CO (0 Q.
Text-figure 6. — Section exposed in Maurv's Bluff 2 on Rio Mao showing "range" of Parametana (Dominilaria) islahispanwlae (Maur>', 1917a) and stratigraphic positions of NMB localities: black squares represent localities collected for microfossils and lithologic analyses; black circles represent localities collected for macrofossils (after Saunders el ai. 1986. text-fig. 31). herein are assigned to six genera. The numbers of lots and specimens of each of these species are hsted in
Table 1. It is obvious from this table that the available
material is not rich. Only two species are well repre-
sented (Table 1): Parametaria (Dominitaria) islahis- paniolae (Maury, 1917a) and Metidella vemista (G. B. Sowerby II, 1850). Three species are moderately well 1 I 1
Dominican Columbellid Gastropods: Jung 1
1 3 species Table 1. — Numbers of lots and spetimcns of each of the (0 of columbellid gastropods available for this study. c >CD
1G957 se Silty cljy -• CD TU 124U
)6807« 1 6956 l-^;::^ CldV^V sill TU l?19
'TU 1220 Sdndy, tidyey bill 16805,TU 1217« 169b2 I
CANADA LIE ME RA
15. -=. 1 6955 I
1G954I Sdruly siti
16806 • 169631 IM i^^V
POTRERO DAM ^l-^t-*- 20< 16949B TU 16b04« 16950B HI' Sdruly r>il 1218
1 695 0'
Text-figure 9. — Schematic column for the central portion of Rio Amina showing "range" of MenilcUa vcnusla (G. B. Sowerby 11, positions of NMB and TU localities: 1 850) and relative stratigraphic black squares represent NMB localities collected for microfossils and lithologic analyses; black circles represent NMB localities collected for macrofossils (after Saunders et a!.. 1986, text-fig. 35).
represented: Cone/la pcrplcxahilis (Maury. 191 7a), AV-
1 MctiilcUa fmifor- tidella cihaoica {Maur> , 9 1 7a), and inis Gabb 1873a. The remaining eight species are rep- resented by five or fewer specimens and thus must be considered rare.
Genus COLUMBELLA Lamarck, 1799
ColumbcUa Lamarck, 1799, p. 70.
Type species (by nwnotypyl.— Voliita mercatoria Linnc, 1758, p. 730. Recent. West Indies. Diagnosis. — ShsW of small to medium size (10-30 mm in height), stout, and solid. Protoconch consists of about one and a half smooth volutions. Number of teleoconch whorls usually six to seven. Profile of spire whorls almost straight, slightly convex, or somewhat shouldered. Body whorl large, evenly rounded or shouldered. Sculpture consists of more or less accen-
tuated spiral cords, or it is lacking entirely. Axial sculp- tural element subordinate or missing. Aperture sig- moid and narrow. Outer lip usually with prominent denticles on its inner surface. Columella with denticles. Anterior canal short, posterior canal inconspicuous.
Remarks. — A list of synonyms of the genus Col- iiinbella has been given by Radwin (1977a. p. 411). The generic name Pyrene Roding (1798. p. 134) has 12 Bulletin 344
Table 2. — Measurements of Columbelta submercatoria Olsson. 1922. Figures are in mm.
restored height width
PRI 2 1 069 (lectotype) 17.8 PR! 21070 (paralectotype) USNM 369448: specimen figured by
Woodnng(1928, PI. 16. fig. 9)
NMB H 17394 (PI. 1. figs. 7-9) NMB H 17395 (PI. 1. figs. 10-12) Dominican Columbellid Gastropods: Jung 13
at hand. Three of them are so badly worn that their Table 3. — Measurements lor Columhctia alt. rusticoides Heilprin, spiral sculpture is hardly recognizable. The orthocline 1887. Figures are in mm. riblets on the early teleoconch whorls are visible only
on one of the figured specimens (PI. 3, figs. 7-9). Comparisons. — As pointed out under C. suhmer-
catoha, C. rusticoides Heilprin ( 1 887. p. 8 1 . fig. 9) was originally described from the Caloosahatchee Forma-
tion of Florida, but is still living in the Recent fauna.
C. aff. rusticoides is similar in degree of slenderness to
C. rusticoides. but its later whorls are evenly convex and not shouldered. Its spiral sculpture is more prom- inent that that of C rusticoides. Four recent specimens of C. rusticoides from Key Biscayne, Florida are at
hand for comparison. Although all of them are some- what worn, one specimen shows inconspicuous, or- thocline riblets on the early teleoconch whorls and in the respect is similar to the figured specimens of C. aff. rusticoides (PI. 3, figs. 7-9). C. aff. rusticoides is some- what smaller than C. platynema Woodring (1928, p. 271, pi. 16, fig. 10) from the early Pliocene Bowden Formation of Jamaica. The description of C. platy- nema was based on the holotype only, a specimen with strongly worn spire whorls. It is refigured here (PI. 3, figs. 1-3). It is therefore not known whether C. pla- tynema has any axial sculpture on the early spire whorls as does C. aff. rusticoides. The worn spire whorls of C platynema give the (probably wrong) impression that its spire is lower than that of C. aff. rusticoides. The Dominican species mainly differs from C. platynema by its considerably finer and less prominent spiral sculpture.
Material. — One lot containing five specimens as list- ed below:
1. 5 spec, TU locality 1230: Rio Cana, east bank, just above the ford at Caimito on Los Quemados-Sa- baneta road (= USGS 8534; Maury's Zone H). Upper part of Cercado Formation (late Miocene).
Measurements. — See Table 3. Occurrence. — Unov^n only from TU locality 1230: Rio Cana at Caimito; upper part ofCercado Formation (late Miocene) (Saunders £•/«/., 1986, text-figs. 15, 16). Distribution. — Not known from outside the Domin- ican Republic.
Columbella lopezana, new species
Plate 3, figures 10-15
Description. — SheW of small size (10-15 mm in height), stout. Protoconch consists of about two and a quarter smooth volutions. Number of teleoconch whorls up to five. Profile of the first teleoconch whorls straight, of later whorls slightly convex. Body whorl moderately inflated. Surface of whorls smooth. The inconspicuous growth lines are orthocline to slightly prosocline. Aperture sigmoid and narrow. Outer lip 14 Bulletin 344
17289, 17290; Baitoa Formation (late early to early and body whorl shouldered. Spiral sculpture consists middle Miocene) (Saunders et al., 1986, p. 30. text- of threads on the lower part of the body whorl. Axial figs. 21. 25). sculpture missing except for growth lines, which may Distribution. — Not known from outside the Domin- be somewhat accentuated on the shoulder of the body ican Republic. whorl. Outer lip moderately thickened, with long lir- ations on its inner surface. The outer lip has a shallow sinus near its adapical end. Posterior canal wide and Genus EURYPYRENE Woodring, 1928 prominent with a few short denticles on the callus of Eun'pyrene Woodhng. 1928. p. 272. the parietal wall. Inner lip with denticles. Anterior ca- nal moderately long, hardly recurved. Type species (by original designation).— Pyrene (Eii- Holotype ofE. inflata.- ANSP 3287. Plate 4, figures rypyrene) eurynotum Woodring, 1928. Bowden, Ja- 1-3. maica; Bowden Formation (early Pliocene). Dimensions holotype inflata. Height 25.9 Diagnosis. SheW of medium size (14-26 mm in of ofE. — mm; width 13.8 height), stout, and solid. Protoconch consists of about mm. Type locality inflata. Santo (= one and a half smooth volutions. Shape of its outer lip ofE. — Domingo Do- minican Republic): this is the only information given not known. Number of teleoconch whorls five to six.
by (1873b) and Pilsbry ( 1 922). type locality Profile of early spire whorls straight to slightly convex. Gabb The is here restricted Later spire whorls and body whorl somewhat shoul- to NMB locality 17290: Rio Yaque This locality is in dered. Spiral sculpture consists of threads on the lower del Norte. situated the Lopez section, which is part part of the body whorl. Axial sculptural element sub- of the Baitoa Formation (Saunders et al., ordinate. Growth lines may be somewhat accentuated 1986, p. 30, text-figs. 21, 25), of late early to early middle Miocene age. on shoulder of body whorl or even on last spire whorl. Holotype E. inflata brassica. 1059. Plate Outer lip moderately thickened, with a shallow sinus of — PRI 4, figures 7-9. near its adapical end; its inner surface with long den- Dimensions holotype inflata brassica. — Height ticles. Anterior canal moderately long, only slightly of ofE. 16.6 width 8.9 recurved. Posterior canal more or less prominent. mm; mm. Type locality inflata brassica. — District, Remarks. — The information concerning the proto- ofE. Brasso Central Range, Trinidad. The exact locality of the ho- conch in the above diagnosis is taken from Woodring lotype is not known. Brasso Formation. The Brasso ( 1 928, p. 272), because the protoconch is not preserved includes the foraminiferal zones of in any of the available specimens. Eurypyrene has been Formation Glo- borotalia its described as a subgenus of Pyrene. Pyrene, however, fohsi and subspecies, which are of early Miocene age according to Bolli and Saunders (1985, is limited to the Indo- Pacific region according to Rad- p. 159, fig. 3). win (1978, p. 333) and its early spire whorls are ter- Remarks.— Exctpi for the holotype oi E. inflata the raced, which is not the case in Eurypyrene. Neogene of the Dominican Republic has yielded only As pointed out by Woodring (1928, p. 272) there are no living representatives of the genus. Without one additional specimen of this species. Even Maury's expedition failed to find a specimen in the Dominican citing a species Radwin (1978, p. 333) wrote that the Republic (Maury, 1917a, 1917b, 1925, p. 211). genus is "last encountered in the Pleistocene of Flor- E. inflata brassica was also based on the holotype ida". According to Radwin (1977a, p. 405, fig. 21) only. It is refigured here (PI. 4, figs. 7-9) and is thought Eurypyrene first appeared in the late Miocene. As shown to be a somewhat immature specimen. The collections under E. inflata. however, the genus is already record- of the Naturhistorisches Basel contain three ed from late early to early middle Miocene times. Museum specimens of E. inflata brassica, all from the early middle Miocene upper Brasso Formation of the Cen- Eurypyrene inflata (Gabb, 1873b) tral Range, Trinidad. One of them from NMB locality Plate 4, figures 1-9 10301 (Caparo River) is somewhat immature and has
Slrombina inflata Gabb, 1873b, p. 221. dimensions similar to those of the holotype of is. in- Columhella inflata (Gabb). Pilsbry. 1922, p. 350 pi. 18, fig. 14. Jlata brassica (height 16.3 mm; width 9.1 mm). A sec- Columbella inflata var. brassica Maury, 1925. p. 211, pi. 36. fig. 8. ond specimen from NMB locality 10365 (Guaracara Description. — SheW of medium size (22-26 mm in River) is of similar size but lacks its body whorl. The height), stout, and solid. Protoconch not known. Num- third specimen is also from NMB locality 10365. It ber of teleoconch whorls up to six. Profile of early spire consists of the body whorl and the last spire whorl. Its whorls straight to slightly convex. Later spire whorls width measures 13.2 mm, a dimension comparable to Dominican Columbellid Gastropods: Jung 15
Table 4. — Measurements of Columhclla lopezana. new species. Tablc 5. — Measurements of Eurypyrenc intlatci (Gabb, 1873b). Figures are in mm. Figures are m mm.
rcslorcd restored height width height width
NMB H 17400 (PI. 3. figs. 10-12) (holotype) 15.0 9.3 ANSP 3287: holotype (PI, 4, figs. 1-3) 26.0 13.8 NMB H 17401 (PI. 3, figs. 13-15) (paratype) 14.3 8.5 NMB H 17405 (PI. 4, figs. 4-6) 22.5 12.3 NMB H 17402; NMB locality 16935 12.7 8.1 NMB H 17403; NMB locality 17289 (paratypes) 10.4 7.1 NMB H 17404; NMB localitv 17290 13.4 8.5 tion (late early to early middle Miocene), northern Do- minican Republic.
Eurypyrene eurynotum (Woodring, 1928) that of the specimens from the Dominican Republic. Plate 5, figures 7-12 For this reason E. inflata brassica is taken in the syn- Pyrene onymy of E. injlata. (Eurypyrene) euryrwtum'Woodnng, 1928, p. 272, pi. 16, figs. 11, 12. Comparisons.— E. eurynotutu (Woodring) (1928, p. 272, pi. 16, figs. 11, 12), the type species of the genus Description. — SheW of small to medium size (14-17 from the early Pliocene Bowden Formation ofJamaica, mm in height), solid. Protoconch consists of one and is considerably smaller than E. inflata; it is more slen- a half volutions. Teleoconch whorls six. Profile of spire der, i.e., its body whorl is less bulging. whorls practically straight. Late spire whorls and body E. veneziielana (Weisbord) (1962, p. 332, pi. 29, figs. whorl shouldered. Body whorl and late spire whorls 19, 20) was based on a single specimen, the holotype, sculptured by weak spiral threads. In the zone of the which is refigured here (PI. 5, figs. 1-3). It had been greatest inflation of the body whorl these spiral threads collected from the Mare Formation of the Cabo Blanco are obsolete. The shoulder of the body whorl and late area, Venezuela, the age of which is middle to late spire whorls is accentuated by a more prominent spiral
Pliocene (age assignment discussed by Jung, 1989, p. thread. Except for growth lines there is no axial sculp-
20). Eleven specimens from the type locality of the ture. Outer lip moderately thickened, with long lira- Mare Formation are at hand (Gibson-Smith collec- tions on its inner surface. Posterior canal moderately tion). They are somewhat smaller than E. injlata, and prominent, with a few inconspicuous denticles on the due to their larger apical angle they have a stouter callus of the parietal wall. Inner lip with denticles. appearance. Anterior canal moderately long, only slightly recurved. E. occidentalis (Weishord) (1962, p. 334, pi. 29, figs. Holotype. -VSNM 135512. See Plate 5, figures 7-9. 21, 22), also from the middle to late Pliocene Mare Dimensions of holotype. — Height 14.0 mm; width Formation of Venezuela, was based on the holotype 7.8 mm. only, which is refigured here (PI. 5, figs. 4-6). The Type locality. — Bo-wden, Jamaica. Bowden Forma- holotype is not quite adult, having a thin (and broken) tion (early Pliocene). outer lip without lirations on its inner surface and no Remarks. — E. eurynotum was based on the holotype columellar callus developed. This very feature was used and a single paratype, and the Dominican Neogene has by Weisbord as a base for his comparative remarks. It yielded only one specimen, which is somewhat worn. is possible that E. occidentalis represents immature E. It is larger than the Jamaican specimens, and its body veneziielana. whorl is slightly more inflated. Its protoconch is not Material. — One lot consisting of a single specimen preserved and its description given above is taken from as listed below: Woodring (1928, p. 272). 1. 1 spec, NMB H 17405; NMB locaHty 17290: Comparisons. — For comparison with E. injlata see Lopez section on Rio Yaquc del Norte; Baitoa For- under that species. mation (late early to early middle Miocene). Material. — One lot consisting of a single specimen Measurements. Set Table 5. as listed below:
Occurrence. — \n the Dominican Republic this 1 spe- . 1 spec, NMB H 1 7406. TU locality 1 404: Arroyo cies is known only from NMB locality 17290: Lopez Babosico, which enters Rio Yaque del Norte from the section on Rio Yaque del Norte; Baitoa Formation, of West at La Barranca, approximately 8 km upstream late early to early middle Miocene age (see Saunders from Santiago de los Caballeros, north side at second et ai, 1986, p. 30, text-figs. 21, 25). bluff below bridge on Las Charcas-La Barranca road; Distribution. — Brasso Formation (early middle Mio- upper part of Globorotalia margaritae zone (late early cene) of the Central Range, Trinidad. Baitoa Forma- Pliocene) according to Saunders et al. (1986, p. 30). 16 Bulletin 344
Table 6. Measurements of Eunpyrene eurynotum (Woodring. — (1989, p. 43) stated that "it is not very diagnostic, 1928). Figures are in mm. whether the inner surface of the outer lip is smooth or
restored sculptured by denticles or lirae". height width Farametaria is a paciphile genus (Woodring, 1966,
USNM 135512: holotype (PI. 5, figs. 7-9) 14.0 7.8 p. 428). In the western Atlantic part of the Tertiary
USNM 369450: paratype 15.7 8.1 Caribbean Faunal Province it is represented by the NMB H 17406 (PI. 5, figs. 10-12) 17.4 9.6 following two species: F. prototypus (Guppy) (1867, p. 171) and its synonym P. schideri (Rutsch) (1942, p. 148, pi. 5, figs. 5a, 5b) from the Savaneta Glauconitic Measurements. Sice Table 6. — Sandstone Member and the Melajo Clay Member, both Occurrence. — \x\ the Dominican Republic this spe- of the Springvale Formation of Trinidad, now assigned cies is known only from TU locality 1404: Arroyo to the early Pliocene; the second species is F. rutschi Babosico (for more details see above); upper part of Jung (1969, p. 500, pi. 52, figs. 3-6) from the Matura Globorotalia margaritae zone (late early Pliocene) shell bed of the Talparo Formation of Matura. Trini- (Saunders et ai, 1986, p. 30). dad, which is now thought to be of Pleistocene age. Distribution. — Bo-wdsn Formation (early Pliocene) The subgenus Farametaria (see below) is therefore of Bowden, Jamaica. Beds of late early Pliocene age. short-lived and restricted in geographic distribution Dominican Republic. northern being known only from the early Pliocene and the Pleistocene of Trinidad. In the Recent fauna of the Genus PARAMETARIA Dall. 1916 Panamic Province it is represented by F. dupontii. Farametaria Dall, 1916, p. 25. Substitute name tor Meta Reeve (1858-1859, pi. 32, under remarks lor Columlu'tia picata. 1859). Subgenus DOMINITARIA, new subgenus
Type species of Meta Reeve (by subsequent desig- Derivation of name. — Dominitaria is an artificial nation. Reeve, 1859. pi. 1, under remarks for Meta combination of syllables referring to the word "Do- o\u\o\dQ%J. — Comisdupontii¥AenQT. 1834-1880(1846, minican" and using the ending oi Farametaria. pi. 61; 1849, p. 273) (for dates see Sherbom and Wood- Type species (herein designated). — Columbella (Meta) ward. 1901, p. 218). Recent, Gulf of California to Tres islahispaniolae Maury (1917a, p. 93, pi. 15, fig. 3). Marias Islands, western Mexico. Cercado Formation (late Miocene) of the northern Do- Diagnosis. S\\t\\ of small to medium size (11-28 minican Republic. mm in height), stout to moderately slender, coniform. Diagnosis. S\\e\\ of small size (8-16 mm in height), Protoconch consists of one to two volutions. Teleo- coniform, moderately slender. Protoconch consists of conch whorls usually six to seven and one half Profile a little more than one and a half volutions. Teleoconch of spire whorls ranging from slightly concave to straight whorls five to seven. Apical angle variable. Profile of to shouldered. Spire whorls smooth (except for growth spire whorls more or less concave to slightly convex. lines) or shouldered. Spiral sculpture on body whorl The first two to three and a half teleoconch whorls usually restricted to its abapical part. Aperture straight sculptured by knobs or axially elongated knobs situated to slightly sigmoid. Outer lip somewhat extended adap- near the abapical suture. There are nine to ten knobs ically. its inner surface smooth or with denticles. Outer per whorl. Later spire whorls sculptured by slightly lip moderately thickened, but with a thin edge. Colu- opisthocyrt growth lines only. Sculptured early teleo- mellar callus inconspicuous, but usually absent. Pos- conch whorls more or less shouldered. Late spire whorls terior canal inconspicuous to moderately prominent. shouldered or not shouldered. Body whorl shouldered, Anterior canal straight. smooth except for growth lines and spiral threads near Remarks. — The genus Farametaria has never been its base. Aperture straight to slightly sigmoid. Outer studied in detail. Certain generic characteristics are lip somewhat extended adapically. its inner surface contradictory depending on the author. Thus the inner with numerous denticles. Outer lip moderately thick- surface of the outer lip is smooth according to Keen ened, but with a thin edge. Columellar callus absent.
(1971, p. 598), has weak denticles (Coomans, 1 967. p. Posterior canal moderately prominent to well devel- 76), or carries denticles or lirations (Grant and Gale, oped. Anterior canal straight. 1931, p. 680). Discussing the variability of P. dupontii Remarks. — The subgenus Dominitaria is distin-
(Kiener), the type species of the genus, Jung (1969, p. guished from Farametaria s.s. mainly by the sculp- 500) pointed out that the denticles on the inner surface tured early teleoconch whorls and the higher spire. of the outer lip may be strongly developed, weak, or Jung (1989. p. 38) pointed out that "the presence or absent. Stressing the relative importance of certain di- absence of sculpture on early teleoconch whorls seems agnostic features of other columbellid genera, Jung to be a fairly important diagnostic feature" on the sub- Dominican Columbellid Gastropods: Jung 17
X 30
i5Jit>rrF-; „" x90 X 45 5 X 45 6
Text-figure 10. — Paramelaria (Dominitanal islahispaniolae (Maury, 1917a). NMB H 17412. NMV locality 16836: Rio Cana section.
x x x 1 Dominican Republic; upper part of Cercado Formation (late Miocene). 1 . apical view, 25; 2. apical view, 45; 3. spire, 0; 4. protoconch,
X 30; 5. protoconch, x45; 6. transition protoconch-teleoconch. x90.
generic level. In addition the species of Dominitaria There are nine to ten knobs on the first teleoconch are smaller than those of the subgenus Parametaria. whorl. Later spire whorls sculptured only by slightly So far only two species are assigned to Dominitaria. opisthocyrt growth lines. Late spire whorls shouldered. In geographic distribution they are restricted to the Height of spire variable, i.e.. apical angle not constant. northern Dominican Republic, and stratigraphically Body whorl shouldered and practically not inflated; they occur in the Baitoa Formation (late early to early sculptured by growth lines and spiral threads near its middle Miocene) on one hand, and on the other hand base. Aperture straight to slightly sigmoid. Outer lip in the Cercado and Gurabo Formations (late Miocene somewhat extended adapically; its inner surface with to early Pliocene). numerous denticles. Outer lip moderately thickened, but with a thin edge. Columellar callus absent. Pos- Parametaria (Dominitaria) islahispaniolae terior canal moderately prominent. Anterior canal (Maury, 1917a) straight. figures 1-3. Plate 6, figures 1-15; Plate 7, figures 1-3 Holotype. -PR\ 28727. Plate 6, Text-figures 10-12 Dimensions of holotype. — Height 11.2 mm; width 5.8 mm. Columbella (Afeta) islahispaniolae Maury. 191 7a, p. 93. pi. 15. fig. 3. Type locality.— Cercado de Mao, Bluff 3 of Maury.
Description. — SheW of small size (8-16 mm in height), Cercado Formation (late Miocene). See Saunders et al. , coniform, moderately slender. Protoconch consists of 1986, text-fig. 29. a little more that one and a half volutions; its outer lip Remarks.— This species is represented by numerous opisthocline to slightly opisthocyrt. Teleoconch whorls specimens. They show that the height of the spire is five to seven. Profile of spire whorls more or less con- variable. Low-spired specimens (PI. 6, figs. 4-6) look cave. The first two to three teleoconch whorls are sculp- rather unlike high-spired specimens (PI. 6, figs. 7-9). tured by knobs or axially elongated knobs situated near The holotype is an intermediate specimen as to height the abapical suture and sometimes overhanging it. of the spire. An unusually large, low-spired specimen Bulletin 344
"•v^^ ^-
t- f'^%.-l-/-': '
'" -^ «'^
^iP^tiSia^,/
x20
4 X 70
Text-figure W. — Parametaria (Dominilarial tslahispanwiae (Maury, 1917a). NMB H 17413. NMB locality 16828: Rio Cana section,
Dominican Republic; lowest part of Gurabo Formation (late Miocene). 1. apical view, x 30; 2. apical view, x 70; 3. early whorls, x 20; 4. protoconch, x 70.
(PI. 7, figs. 1-3) caiTies a few spiral ribs on the lower 4. 1 spec, NMB locality 16820: Rio Cana section; half of its outer lip. Low-spired and high-spired spec- lowest part of Gurabo Formation (latest Miocene). imens may be found in the same lot. Thus the variable 5. 23 spec. NMB locality 16828: Rio Cana section; height of the spire is considered as true intraspecific lowest part of Gurabo Formation (latest Miocene). variability. 6. 6 spec, NMB locality 16835: Rio Cana section; Co«;par/,?o«5. — Comparative remarks are given un- upper part of Cercado Foramtion (late Miocene). der P. lopezarw, n. sp. 7. 7 spec, NMB locality 16836: Rio Cana section; Ma/ma/. — Twelve lots with a total of 89 specimens upper part of Cercado Formation (late Miocene). as listed below: 8. 13 spec, NMB locality 16844: Rio Cana section; 1. 1 spec, PRI 28727: holotype; Bluff 3, Cercado upper part of Cercado Formation (late Miocene). de Mao; Cercado Formation (late Miocene). 9. 5 spec, NMB locality 16879: Rio Cana section; 2. 8 spec, NMB locality 16817: Rio Cana section; lower part of Gurabo Formation (early Pliocene). lower part of Gurabo Formation (early Pliocene). 10. 2 spec, NMB locality 16913: Rio Mao section
3. 8 spec, NMB locality 16818: Rio Cana section; (same as Maury's Bluff 3); Cercado Formation (late lower part of Gurabo Formation (early Pliocene). Miocene). I
Dominican Columbellid Gastropods: Jung 19
locality Bajon 11. 13 spec, NMB 16928: Arroyo on 16 -I Rio Mao; Cercado Formation (late Miocene). 12. 2 spec, NMB locality 16929: Rio Mao, about 15 50 m upstream from Maury's Bluff 2; Cercado For- mation (late Miocene). Measiirc?ncnts. — Ploned in Text-figure 12. 14H Occurrence. — Rio Mao section: Cercado Formation (late Miocene): localities 16913, 16928, 16929. NMB 13 Rio Cana section: in this section the species occurs in the Cercado and the Gurabo Formations. Cercado For- 12- mation (late Miocene): NMB localities 16835, 16836, 16844; Gurabo Formation (latest Miocene part of the Formation): NMB localities 16820, 16828; Gurabo 11 - Formation (early Pliocene part of the formation): NMB localities 16817, 16818, 16879. See Saunders et al., 10- (1986, text-figs. 15, 16, 29, 30). Distribution. — Noi known from outside the Domin- ican Republic. 9
Parametaria (Dominitaria) lopezana, new species 8- Plate 7, figures 4-9 Derivation of name. — Named after the Lopez sec- 7- tion, the type locality. Description. — SheW of small size (10-15 mm in
1 1 height), coniform, fairly slender. Protoconch probably — 1 T T consists of about one and a half volutions. Teleoconch 3 4 5 6 7 8 whorls five to six and a half Profile of teleoconch whorls Text-figure 12. — (Restored) height/width diagram oi Parameiana
. 1917a). slightly concave to moderately convex. The first three (Dominitaria) islahispaniolae (Maury to three and a half teleoconch whorls are sculptured by knobs situated near the abapical suture and some- lip seems to be opisthocline. The figured paratype (PI.
times overhanging it. There are 1 1 to 1 2 knobs on the 7, figs. 7-9) is the largest specimen. The sculpture on
first teleoconch whorl. Later spire whorls sculptured its early teleoconch whorls is weak, and the suture of only by slightly opisthocyrt growth lines. Late spire Its late spire whorls is somewhat channelled. whorls slightly shouldered and moderately convex in Comparisons.— Ihe general shape of P. lopezana is profile. Body whorl slightly shouldered, very little in- less coniform than that of P. islahispaniolae. P. lope-
flated, sculptured by growth lines and a few spiral zana is more slender, i.e.. its apical angle is smaller. threads near its base. Aperture slightly sigmoid. Outer The sculpture on the early teleoconch whorls is less lip somewhat extended adapically; its inner surface prominent in P. lopezana. The shoulder of the spire with numerous denticles. Outer lip not much thick- and body whorls is more pronounced in P. islahispa- ened, with a thin edge. Columellar callus absent. Pos- niolae. The unsculptured spire whorls of P. lopezana terior canal well developed. Anterior canal straight. are convex in profile, whereas the profile of the cor- Holotype. -NMB H 17414. Plate 7, figures 4-6. responding whorls of P. islahispaniolae is concave to Dimensions of holotype. — HeighX 11.9 mm; width almost straight. The aperture of P. lopezana is more 5.7 mm. sigmoid, and the posterior canal is more pronounced Type locality. — NMB locality 16936: Lopez section than that of P. islahispaniolae. on Rio Yaque del Norte; Baitoa Formation (late early Material.— Three lots with a total of only five spec- to early middle Miocene) (Saunders et al.. 1986, p. 30, imens as listed below (all from the late early to early text-figs. 21, 25). middle Miocene Baitoa Formation):
Remarks.— This new species is represented by five 1. 3 spec, NMB locality 16936: Lopez section on
specimens only. In the above description it is stated Rio Yaque del Norte.
that the protoconch consists of about one and a half 2. 1 spec, TU locality 1363: Arroyo Hondo, Lopez
volutions. This is an assumption, however, because no section on Rio Yaque del Norte.
complete protoconch is preserved. Although the state 3. 1 spec, TU locality 1364: mouth of Arroyo Hon-
of preservation is unsatisfactory, the shape of its outer do, Lopez section of Rio Yaque del Norte. 20 Bulletin 344
12.0 Dominican Columbellid Gastropods: Jung 21
Columella with a thin callus. Anterior canal short and 17 - straight. Posterior canal moderately prominent. Holotype. -PRl 28728. See Plate 8, figures 7-9. 16 - Dimensions of holotype. — VleighX 12.7 mm; width 6.2 mm. Type locality. — B\uff 3 of Maury, Cercado de Mao; 15 - Cercado Formation (late Miocene). For location see Saunders et ai, 1986, text-fig. 29. 14 - Remarks. — A.\\ho\i%h this species is not particularly rare, there is no protoconch preserved well enough to - be suitable for scanning electron micrography. The ho- 13 lotype is a rather small specimen; it has only five and a half teleoconch whorls and therefore has a somewhat 12- more slender appearance than other specimens. Comparisons. — The only species with which C per- 11 - ple.xabilis can be compared is the living Caribbean C ovulata (Lamarck) (1882, p. 295). Both species have a similar degree of stoutness. C. perplexabilis differs from L 1 1 r- C. ovulata by the lack of spiral sculpture on the spire 6 7 8 9 3. per- whorls and on the adapical part of the body whorl. C. Text-figure 1 —(Restored) height/width diagram of Conella
plexabilis (Maury, 1 9 1 7a). ovuloides (C. B. Adams) (1850, p. 53) is considerably more slender than C. perplexabilis. Its lectotype (se- sists of about one and a half volutions; its outer lip lected by Clench and Turner, 1950, p. 321, pi. 41, fig. slightly opisthocyrt. Teleocnch whorls five to six in 4) is refigured here for comparison (PI. 8, figs. 1-6). number. Profile of early teleoconch whorls straight, of Material.— Vive lots with a total of 26 specimens as later spire whorls somewhat convex. Body whorl little listed below. All specimens have been collected from inflated, not shouldered. Besides orthocline to slightly the late Miocene Cercado Formation. opisthocline growth lines all the whorls are smooth 1. 3 spec. NMB locality 16913: Rio Mao; corre- with the exception of the early teleoconch whorls, which sponds to Bluff 3 of Maury. may be sculptured by more or less conspicuous, prac- 2. 3 spec, NMB locality 16927: Rio Mao, at mouth tically orthocline ribs. Aperture moderately narrow. of Arroyo Bajon. Outer lip not greatly thickened, somewhat extended 3. 6 spec, NMB locality 16928: Rio Mao, near adapically, its inner surface with more or less well de- mouth of Arroyo Bajon. veloped denticles or smooth. Columella smooth, with 4. 9 spec, TU locality 1294: Rio Mao; corresponds two weak plications, or with denticles. Anterior canal to BlufT3 of Maury. straight, short, and wide. Posterior canal inconspicu- 5. 5 spec, TU locality 1 379: Rio Mao, around mouth ous. of Arroyo Bajon. Remarks. — In an attempt to see and refigure the type Measurements. — YXoWtd in Text-figure 13. specimen of Columbella nitida Lamarck, 1822 I have
Occurrence . — 1h\% species is recorded only from the received the following information from Dr. Yves Fi- late Miocene Cercado Formation of the Rio Mao sec- net of the Natural History Museum in Geneva (written tion: localities 16913, 16927, 16928, and TU NMB communication, February 25, 1988): localities 1294, 1379. For location see Saunders et al. (1986, text-fig. 29). "In our original volumes of Anim. s. vert., La- Distribution. — Not known from outside the Domin- marck's daughter, Rosalie de Lamarck, made man- ican Republic. ual annotations that mention the number of speci- mens originally present in her father's collection. For C. nitida. she wrote "21 specimens" (= syntypes). Genus NITIDELLA Swainson, 1 840 There is no C. nitida in our Lamarck collection, Nitidella Swainson, 1840. p. 313. but 2 lots of 15 specimens of this species in our Type species (by monotypy). — Nitidella marmorata general collection, from the Delesscrt collection (let-
Swainson ( 1 840, p. 1 5 1 , fig. 1 7c.) (= Columbella nitida ter D left and above on the cardboard plate). In the Lamarck, 1822, p. 295). Recent, West Indies. index file that Mermod made for the species de- Diagnosis. — ShtW of small to medium size (10-19 scribed by Lamarck and eventually present in our mm in height), moderately slender. Protoconch con- Lamarck collection, he mentioned that one of these 77 Bulletin 344
x30 4 x65 5 x65
Text-figure 14. — Nitidella cibaoica (Maury, 1917a). NMB H 17421. NMB locality 16934: Rio Gurabo section, Dominican Republic; lower
part of Gurabo Formation (late Miocene). 1. apical view, x30; 2. apical view, x65; 3. early whorls, x30; 4. protoconch, x65; 5. protoconch and transition to teleoconch, x65.
30 specimens is probably the one figured by Kiener ied the history of the case in more detail than has been
(pi. 15, fig. 1) and possibh could be the type. But possible here.
this is not certain. I think that a loctotype has to be A', nitida is widespread in the living fauna of the designated, eventually the specimen pointed out by Caribbean Province. Contrary to the statement by
Mermod." Radwin (1977a. p. 414) it does not occur in the Pleis-
tocene of Venezuela. Although Weisbord ( 1962. p. 320) After comparison of the figure given by Kiener ( 1 834- recorded N. nitida from the Distrito Federal, Vene-
1 880. pi. 1 5, fig. 1 . 1 84 ) with the specimen mentioned 1 zuela, he figured only Recent specimens and stated that above, of which Mermod thought it might correspond .". "not a single fossil specimen was observed . . The to Kiener's figure, it is obvious that this is not so. The only fossil species of Nitidella I am aware of is A^. protoconch of the specimen is strongly eroded, whereas cibaoica (Maury) from the late Miocene of the Do- the figure shows a pointed apex. In addition the color minican Republic, which is described below. The pattern does not match. There is no evidence that any stratigraphic range of Nitidella is therefore late Mio- of the specimens from the Delessert collection were cene to Recent. ever part of the original Lamarck collection. For this reason it is not advisable to select a lectotype. On the Nitidella cibaoica (Maury. 1917a) contrary: it would be preferable to designate a speci- Plate 9. figures 1-12 men with good locality information as the neotype of Text-figures 14, 16 Cohimbella nitida Lamarck. I hesitate to designate a neotype of the species, however, without having stud- Coluinbella (Nitidella) cibaoica Maury, 1 9 1 7a, p. 99, pi. 2 1 . figs. 3, 4. Dominican Columbellid Gastropods: Jung 23
'SS'-
^40^f
x25 x25
>^-^
7v'-
x65 4 x80
Text-figure \5.-Nilidella nitida (Lamarck, 1822). NMB H 17422. NMB locality 17704: Isla de Aves, Venezuela; recent. 1. apical view. x25; 2. apical view, x65; 3. early whorls, x 25; 4. protoconch x 80.
Description . — ShtW of small to medium size (10-19 Lectotype (herein selected). — ?K\ 28794. See Plate mm in height), moderately slender. Protoconch con- 9, figures 1-3. sists of about one and a half volutions: its outer lip Dimensions of lectotype. — HeighX 15.0 mm: width slightly opisthocyrt. Teleoconch whorls usually six in 8.0 mm. number. The profile of the first two teleoconch whorls Type locality. — Rio Cana at Caimito; Cercado For- is straight; they are sculptured by inconspicuous, or- mation (late Miocene). For location see Saunders et al. thocline ribs, which are more prominent abapically. (1986, text-fig. 15).
There are about 15 ribs per whorl. The subsequent Remarks. — N. cibaoica is not a common species. whorls have a somewhat convex profile and are smooth The denticles on the inner surface of the outer lip are except for faint, practically orthoclinc growth lines. usually well developed, but not so in the lectotype and
Body whorl not greatly inflated. Aperture moderately the figured paralectotype (PI. 9, figs. 4-6). The orthoc- narrow. Outer lip thickened, somewhat extended adap- linc ribs on the early teleoconch whorls arc sometimes ically, its inner surface with numerous more or less poorly developed and hardly recognizable. The figured prominent denticles. Columella smooth or with two specimens from the Rio Gurabo section (PI. 9, figs. 7- hardly recognizable plications. Anterior canal straight, 1 2) are larger than the average size of the available short, and wide. Posterior canal inconspicuous. material. Bulletin 344
3. 8 spec, NMB locality 16856: Rio Cana; Cercado 19- Formation (late Miocene). 4. 4 spec, NMB locality 16934: Rio Gurabo; lower - 18 part of Gurabo Formation (late Miocene). 5. 5 spec, TU locality 1420: Arroyo Bellaco, trib- 17 utary of Rio Cana; most probably Cercado Formation (late Miocene). 16- Distribution. — Not known from outside the Domin- ican Republic.
15- Nitidella aff. cibaoica (Maury, 1 9 1 7a) Plate 8, figures 10-12
14- Remarks. — This form is represented by a single specimen from TU locality 1250 (Rio Verde; age prob- 13- ably early Pliocene) (for location see Saunders et al., 1 986, text-fig. 38). Its protoconch is not preserved. The
general appearance is stouter than that of TV. cibaoica. 12- The shell consists of almost seven teleoconch whorls.
The profile of the early teleoconch whorls is straight,
11 - that of the subsequent whorls moderately convex. The first three and a half teleoconch whorls are sculptured by orthocline to slightly opisthocline ribs, which are 10- considerably more prominent in their abapical part. There are 12 to 15 ribs per whorl. The aperture is 9- narrower than that of N. cibaoica. The inner surface of the outer lip carries prominent denticles. The col- umella is covered by a callus with six denticles. The L 1^ posterior canal is more prominent than that of N. ci- 7 8 10 baoica. This shell obviously represents a new species.
Text-figure 16.— (Restored) height/width diagram of Mtidella ci- I feel, however, that more material is needed to de- baoica (Maury, 1917a). scribe it formally.
Nitidella caimitana, new species Comparisons . — To my knowledge the living Cainb- Plate 8, figures 16-21 bean A^. nitida (Lamarck) (1822, p. 295) is the only other species of Nitidella so far described. A^. cibaoica Derivation of name.— Named after Caimito on Rio (Maury) mainly differs from N. nitida by having sculp- Cana, the type locality. tured early teleoconch whorls. In addition the denticles Description. -Shell of small size (six to eight mm in on the inner surface of the outer lip are usually more height), slender. Protoconch consists of one and a half
Measiireiywnts . — VXonQd in Text-figure 16. to two volutions; the shape of its outer lip is not known. Occurrence.— Ccrcado Formation (late Miocene) of Number of teleoconch whorls five and a half Profile the Rio Cana section: NMB locality 16856 and TU of early teleoconch whorls straight, of later whorls locality 1420. Lower part of Gurabo Formation (late slightly convex. Except for faint, orthocline growth lines
Miocene) of the Rio Gurabo section: NMB localities all the teleoconch whorls are smooth. Body whorl only 15849, 15857, 16934. For location see Saunders, Jung, slightly inflated. Aperture narrow, slightly sigmoid. and Biju-Duval (1986, text-figs. 4, 6, 15, 16). Outer lip thickened, only slightly extended adapically, strongly developed in A', cibaoica. Two recent speci- its inner surface with numerous, well developed den- mens of N. nitida from Isla de Aves, Venezuela, are ticles. Columella with two plications. Anterior canal figured for comparison (PI. 9, figs. 1 3-18; Text-fig. 1 5). straight, short, and moderately narrow. Posterior canal Material. — ¥i\t lots with a total of 19 specimens as inconspicuous. listed below: Holotype. -'NMB H 17425. See Plate 8, figures 16-
1. 1 spec, NMB locality 15849: Ri'o Gurabo: lower 18. part of Gurabo Formation (late Miocene). Dimensions ofholotype. — Height 7.2 mm; width 3.4
2. 1 spec, NMB locality 15857: Rio Gurabo; lower mm. part of Gurabo Formation (late Miocene). Type locality. — T\J 1 230: Rio Cana at Caimito; Cer- Dominican Columbellid Gastropods: Jung 25
cado Formation (late Miocene). For location see Saun- Tabic 8. — Measurements o{ Nitidella caimilana. new species. Fig- ures are in mm. ders f? a/.. (1986, text-fig. 15). single Remarks. — This species is represented by a restored lot containing only three specimens. This is admittedly height width a low number of specimens upon which to base a new NMB H 17425: holotype (PI. 8, figs. 16-18) 7.2 3.4 species. N. cainutana, however, is so similar to N. NMB H 17426: figured paratype 8.0 3.7 nilida(P\. 9. figs. 13-18 for comparison), however, that NMB H 17427: unfigured paratype 5.7 2.7
it may be its ancestor and therefore well worth being described formally. Although the protoconch of the holotype is pre- Remarks.— Metulella columbellata (Dall) (1889, p.
served, its preservation is not good enough to recognize 182), known from the Recent fauna of the Gulf of
the shape of its outer lip; the number of its volutions Mexico, was first figured by Dall (1890, pi. 6, fig. 8). as indicated in the above description is therefore not According to Radwin (1977b, p. 126, fig. 19) it is the fully reliable. The figured paratype is lacking its pro- only living species of the genus, and its habitat seems toconch. The unfigured paratype has its protoconch, to be restricted to deeper water. Unfortunately no ma- but the surface of the whole shell is eroded. terial oi M. columbellata is available for comparison. Comparisons. — N. caimitana is not only smaller than The collection Gibson-Smith, however, contains two
N. cibaoica, but it lacks the axial sculpture on the first apparently undescribed species collected in Venezue-
teleoconch whorls. In this latter respect it is like the lan waters. All these three living species have fewer
living A', nitida. N. nitida and A^. caimitana have about teleoconch whorls than the fossil species described be- the same number of teleoconch whorls, but N. caim- low. The protoconch of M. columbellata as shown in
itana is considerably smaller. The prominence of the the figure given by Dall (1890, pi. 6, fig. 8) is quite denticles on the inner surface of the outer lip is variable similar to those of the two fossil species recorded here- in N. nitida; sometimes they are hardly developed. In in. On the other hand the protoconch of the Venezu- the available material of N. caimitana they are con- elan material mentioned above is quite different: the spicuous. apex is pointed, and there are almost three volutions. Material. — One lot with a total of only three spec- In his discussion of the genus Metulella Woodring imens as listed below: (1928, p. 278) proposed to use the subgeneric name
1. 3 spec, TU locality 1230: Rio Cana at Caimito; Metulella for slender species with a long anterior canal
Cercado Formation (late Miocene). and the subgeneric name Thiarinella Sacco ( 1 890) for Measurements. — See Table 8. stout species with a short anterior canal. As a conse- Occurrence.— Cercado Formation (late Miocene) of quence he assigned M. fusiformis to the subgenus Me- the Rio Cana section: TU locality 1230. tulella and M. venusta, which is described below, to Distrihution. — Nol known from outside the Domin- Thiarinella. Fusus comptus Bronn, 1831, is the type ican Republic. species of Thiarinella. According to the description and figure given by Sacco (1890, p. 56, pi. 2, fig. 74) Genus METULELLA Gabb, 1873a it completely lacks spiral sculpture on the spire whorls.
Metii/ella Gahb. 1873a, p. 270. It therefore seems advisable to consider Metulella as a genus with Caribbean distribution only. Type species (by original designation).— Metulella Metulella (Thiarinella) olssoni Perrilliat ( 1 972, p. 90, fusiformis Gahh, 1873b. Neogene, Dominican Repub- pi. 42, figs. 7-16) from the Agueguexquite Formation lic. (early Pliocene) of Mexico is a species of the subgenus Diagnosis. -Shell of small to medium size (10-30 Lirastrombina Jung (1989, p. 96). Erroneously it has mm in height), slender. Protoconch consists of a little not been discussed by Jung (1989). less than two volutions; its outer lip is slightly opisth- The stratigraphic range of the genus Metulella as ocyrt. Number of teleoconch whorls six to nine. Profile understood here is late Miocene to Recent. of teleoconch whorls straight to moderately convex. incised. Sculpture teleoconch Suture usually strongly of Metulella fusiformis Gabb, 1873a whorls consists of conspicuous axial ribs and less Plate 10, figures 1-17 spiral threads. Body whorl moderately in- prominent Text-figures 17, 18 flated. Aperture narrow. Outer lip moderately thick- Metulella fusiformis Gabb. 1873a. p. 270, pi. 1 1, fig. 3. Gabb, 1873b. ened, with prominent denticles on its inner surface. p. 206. Maury, 1917a, p. 89. pi. 15, fig. 28. Pilsbry. 1922. p. 352. Columella with a callus carrying denticles. Anterior pi. 18, fig. 16. canal narrow, moderately long to long. Posterior canal Metulella williamgabbi Maury, 1917a, p. 90, pi. 15. fig. 29. Pilsbry, inconspicuous. 1922. p. 352. 1
26 Bulletin 344
x30 x25
x50 x50
Text-figure 17 .-.\feliilellafusifomiis Gahh. 1873a. NMB H 17429. NMB locality 17266: La Barranca on Rio Yaque del Norte. Dominican x x x 4. Republic; upper part of Globorotalta margantae zone (late early Pliocene). 1 . apical view, 30; 2. apical view, 50; 3. early whorls, 25; protoconch and transition to teleoconch. x 50.
Metulella dominicensis Pilsbry and Johnson, 1917. p. 169. Pilsbry, Between the lowest spiral thread and the suture there 1922. p. 352. pi. 18. fig. 18. is a narrow but usually steep slope. The lower part of
is spiral threads, but the Description. — S^itW of small to medium size (15-19 the body whorl sculptured by axial ribs fade out toward the base. Outer lip not strongly mm in height), slender. Protoconch consists of a little its inner surface with prominent denticles. less than two volutions; its outer lip is slightly opisth- thickened; denticles, ocyrt. Number of teleoconch whorls six and a half to Inner lip with a callus carrying conspicuous recurved. seven and a half Profile of teleoconch whorls only interior canal long and narrow, somewhat Pilsbry, slightly convex to moderately convex. Suture strongly Lectotype.-A.nSP 3216 (selected by 1922. incised. Sculpture of teleoconch whorls consists of con- p. 352). See Plate 10. figures 1-3. spicuous axial ribs and usually less prominent spiral Dimension oflectotype. — He'i^l 1 8.8 mm; width 6. threads. The axial ribs are orthocline to slightly pro- mm.
socline, and their number per whorl is 12 on early Type locality. — SanXo Domingo. As noted b\ Pilsbry teleoconch whorls and up to 18 on late teleoconch (1922, p. 307) this is the only indication as to locality whorls. On the first one or two teleoconch whorls the that can be found on the original labels of the Gabb spiral threads are poorly developed, but their promi- collection in the Academy of Natural Sciences in Phil- nence gradually increases on successive whorls. Their adelphia. The same can be said of the type localities number is four or five on all the teleoconch whorls. of A/, williamgabbi and M. dominicensis. because these Dominican Columbellid Gastropods: Jung 27
two species are based on material from the Gabb col- 20 lection. The type locality of Af. fusijorniis is here res- triced to NMB locality 1 7266: La Barranca on Rio 19- Yaque del Norte. Its location is given by Saunders et
1 a/. ( 1 986, text-lig. 2 1 ). According to these authors ( 986, p. 30) the silts cropping out at NMB locality 18- 17266 are of late early Pliocene age (upper part of Globorotalia niargaritae Zone). 17- HolorypeofM. williamgabbi.-PRI 28745. See Plate 10, figures 9-11. Dimensions ofholotypeofM. williamgabbi. — Height 16- 16.5 mm; width 5.7 mm. Type locality o/'M. williamgabbi. — Santo Domingo 15- (see above). llo/olypc M. dominicensis. ANSP 3217. See of — 14- Plate 10, figures 4-6. holotype dominicensis. — Dimensions of of M. Height L 16.4 mm; width 5.2 mm. -r 1^ Type locality o/'M. dominicensis. — Santo Domingo 4 6 (see above). Text-figure 18— (Restored) height/width diagram of Metiilellu fu- Remarks. — MosX authors have considered M. fiisi- siformis Gabb. 1873a. fornus to be a rare species. The present description of the species is based on enough material to allow for some remarks on variability. The profile of the teleo- Yaque del Norte; upper part of Globorotalia niargar- conch whorls is variable to some degree: it ranges from itae Zone (late early Pliocene). almost straight to moderately convex. Specimens form 2. 6 spec, NMB locality 1 7267: La Barranca on Rio Arroyo Zalaya (NMB locaHty 17271; PI. 10, figs. 15- Yaque del Norte; upper part of Globorotalia niargar- 17) have a more convex profile than specimens from itae Zone (late early Pliocene). other localities. 3. 2 spec, NMB locality 17271: Arroyo Zalaya; Glo- M. williamgabbi and M. dominicensis are both based borotalia margaritae Zone (early Pliocene). on material from the Gabb collection, and their exact Measurements. — VloXXed in Text-figure 18. type localities are therefore not known. Pilsbry (1922, Occurrence. — Globorotalia margaritae Zone (early p. 352) noted that M. dominicensis is "perhaps not Pliocene) of La Barranca on Rio Yaque del Norte (NMB distinct from M. williamgabbi". Both have somewhat localities 17266, 17267) and Arroyo Zalaya (NMB lo- reduced axial and spiral sculptures, and the profile of cality 17271). For location and age see Saunders et al. their teleoconch whorls is less convex. These features (1986. pp. 30, 34, text-figs. 21, 36). are here considered to fall within the variability of M. Distribution. — Nol known from outside the Domin- fusiformis. ican Republic. Comparisons. — The living Af. columbellata (Dall) Metulella venusta (G. B. Sowerby II, 1850) (1889, p. 182) from the Gulf of Mexico basically is a Plate 10, figures 18-20; Plate 11, figures 1-9 smaller species. It has a larger apical angle, the profile Text-figures 19, 20 of its teleoconch whorls is practically straight, and its
anterior canal is considerably shorter than that of M. ColumbellavenustaG. B. Sowerby II. 1850, p. 46. pi. 9. fig. 6.Guppy. fusiformis. M. elongata (Toula) (1911, p. 478, pi. 29. 1876. p. 526. Metulella venusta Sowerby, Gabb. 1873b. p. 206. Maury, 1917a, p. fig. 8) from the Pliocene (?) of Tehuantepec, Mexico, 89, pi. 15, figs. 26, 27. Pilsbry. 1922. p. 352. is obviously closely related to A/, fusiformis. No top- otypes are available for comparison, however. The Description. -Shell of medium size (20-30 mm in
specimen figured by Toula measures only 1 1.8 mm in height), moderately slender. Protoconch consists of a height and is thus smaller than all the specimens of A/. little less than two volutions; its outer lip is slightly fusiformis at hand. Judging from Toula's figure the opisthocyrt. Number of teleoconch whorls up to nine. axial ribs are more prominent, and there seem to be Profile of teleoconch whorls moderately convex. Su- more spiral threads in AI. elongata. ture incised. Sculpture of teleoconch whorls consists Material. — Three lots with a total of only 15 spec- of orthocline to slightly prosocline axial ribs and spiral
imens as listed below: threads and bands. There are about 1 7 axial ribs per early 27 on late 1 . 7 spec, NMB locality 1 7266: La Barranca on Rio whorl on teleoconch whorls and about 28 Bulletin 344
S^V
^^^•n'a^^v x25
Text-figure \9. — Metu!ella venustaiG. B. Sowerby 11. 1850). NMB H 17431. NMB locality 15903: Rio Gurabo section. Dominican Republic;
Cercado Formation (late Miocene). 1. apical view, x 30; 2. apical view, x 60; 3. early whorls, x 25; 4. protoconch and transition to teleoconch, x50.
teleoconch whorls. On the first teleoconch whorls the Dimensions of lectotype. — Hq'\^\ 23.4 mm; width spiral threads are not or only poorly developed, but 9.5 mm. their prominence gradually increased on successive Type locality. — The locality information accompa- whorls. On later teleoconch whorls the number of spi- nying the lectotype simply states "Yaque River", Do- ral threads and bands is five to six. Sometimes the minican Republic. This general indication is unsatis- whorls may be slightly shouldered. The entire height factory. Among the available material there is only one of the body whorl is sculptured by spiral threads and specimen from Rio Yaque del Norte (NMB locality bands, but the axial ribs fade out toward the base. 17267: La Barranca). The bulk of the material, how- Outer lip moderately thickened, with a fairly sharp ever, has been collected from the Rio Gurabo section edge, its inner surface with well developed denticles. and on Rio Mao, areas which are not far from the Inner lip with a strong callus carrying prominent den- "Yaque River" (Rio Yaque del Norte). ticles. Anterior canal short and practically straight. Remarks. — Af. vemista is a fairly common species. Posterior canal inconspicuous. Its morphological characters are more or less constant Lectotype (herein selected). — BMNH GG 20044. See with the exception that sometimes the spire whorls are
Plate 1 1, figures 1-3. This is the specimen figured by somewhat shouldered and not regularly convex.
G. B. Sowerby II (1850, pi. 9, fig. 6). Comparisons.— M. vemista cannot really be com- Dominican Columbellid Gastropods: Jung 29
pared with M. fusiformis. It is not only a considerably 30- larger species, but is stouter, i.e., has a larger apical angle, and has a much shorter anterior canal. In the two latter respects Af. vemista is closer to the living M. columbellata (Dall) (1889, p. 182) from the Gulf of Mexico. M. coluinbellata, however, is much smaller than M. vemista, and the profile of its teleoconch whorls is less convex or even straight. Material. — Ill lots with a total of 129 specimens as listed below:
1. 2 spec, NMB locality 15814: Rio Gurabo; early Pliocene part of Gurabo Formation. 2. 6 spec, NMB locality 15842: Rio Gurabo; late Miocene part of Gurabo Formation. 3. 2 spec, NMB locality 15844: Rio Gurabo; late Miocene part of Gurabo Formation. 4. 3 spec, NMB locality 15863: Rio Gurabo; lower part of Gurabo Formation (late Miocene). 5. 2 spec, NMB locality 15865: Rio Gurabo; lower part of Gurabo Formation (late Miocene).
6. 1 spec, NMB locality 15866: Rio Gurabo; lower part of Gurabo Formation (late Miocene). 7. 2 spec, NMB locality 15867: Rio Gurabo; lower part of Gurabo Formation (late Miocene). 8. 5 spec, NMB locality 15871: Rio Gurabo; lower part of Gurabo Formation (late Miocene). 9. 7 spec, NMB locality 15898: Rio Gurabo; Cer- cado Formation (late Miocene). 10. 5 spec, NMB locality 15899: Rio Gurabo; Cer- cado Formation (late Miocene).
11.1 spec, NMB locality 1 5902: Rio Gurabo; Cer- cado Formation (late Miocene).
12. 1 spec, NMB locality 15903: Rio Gurabo; Cer- cado Formation (late Miocene).
1 3. 1 spec, NMB locality 1 5905: Rio Gurabo; Cer- cado Formation (late Miocene). 14. 5 spec, NMB locality 15906: Rio Gurabo; Cer- cado Formation (late Miocene). 15. 2 spec, NMB locality 15907: Rio Gurabo; Cer- cado Formation (late Miocene).
16. 1 spec, NMB locality 1 6808: Rio Gurabo; lower part of Gurabo Formation (late Miocene).
17. 1 spec, TU locality 1375: Rio Gurabo; Cercado Formation (late Miocene). 18. 2 spec, NMB locality 16807: Rio Amina; beds of probably late Miocene age. 19. 2 spec, NMB locality 16821: Rio Cana; lowest part of Gurabo Formation (late Miocene).
20. 1 spec, NMB locality 1 6838: Rio Cana; Cercado Formation (late Miocene). 21. 2 spec, NMB locality 16857: Rio Cana; Cercado Formation (late Miocene). 22. 74 spec, NMB locality 16910: Rio Mao; beds of late Miocene age. 30 Bulletin 344
calities see Saunders et at., (1986, text-figs. 4, 5, 6, 15, Distribution.— Not known from outside the Domin- 16, 21, 29, 34, 35). ican Republic.
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manding. .\'.\7.V. Report on the Mollusca. Part II. — Gas- Bulletins of Amencan Paleontology, vol. 98. No. 334. pp.
tropoda and Scaphopoda. Bulletin of the Museum of Com- 83-144, pis. 15-29, text-figs. 1-18, table 1. parative Zoology, vol. 18. pp. 1^92. pis. 10-40. Keen, A. M. 1890. Scientific results of exploration by the U.S. Fish Commis- 1971. Sea shells of tropical West .America. Second edition. Stan- '//. sion Steamer .Albatross. I Preliminary report on the ford University Press, Stanford. California. 1064 pp.. il- collection ofMollusca and Brachiopoda obtained in 1887- lustr. 8. Proceedings of the United States National Museum, Keroher, G. C. and others. vol. 12, pp. 219-362, pis. 5-14. 1966. Lexicon of Geologic .Wames ofthe United States for 1936-
1916. Notes on the H 'est .A merican Columbellidae. The Nautilus, 1960. U.S. Geological Survey Bulletin 1200, iv + 4341 vol. 30, No. 3. pp. 25-29. pp. Dall, W. H.. and Bartsch. P. Kiener, L.-C. 1911. New species of shells from Bermuda. Proceedings of the 1834-1880. Species general et iconographie des coquiles vi- 1-1 livr. United States National Museum, vol. 40, No. 1820, pp. vantcs. . . (continue par P. Fischer). Paris, vol. 1, 277-288, pi. 35. 1-165 (for dates see Sherbom, C. D., and Woodward. B. Gabb, W, M. B.. 1901, Proceedings of the Malacological Society of Lon- 1873a. Description of some new genera of .Mollusca. Proceedings don, vol.4, pp. 216-219). of the Academy of Natural Sciences of Philadelphia, vol. Kornicker, L. S. 24. pp. 270-274, pis. 9-11. 1979. The question mark in tu.xonomic literature. Journal of Pa- 1873b. On the topography and geology ofSanto Domingo. Trans- leontology, vol. 53, p. 761. actions of the American Philosophical Society, new series, Lamarck, J.B.P.A. vol. 15, pp. 49-259. 2 maps. 1799. Prodrome d'linc nouvcllc classification des coquillcs. Me- Gibson-Smith, J,, and Gibson-Smith, W. moires de la Societe d'Histoire Naturelle de Paris, \ol. 1. 1979. The genus Arcinella (Mollusca: Bivahia) in Venezuela and pp. 6.3-71. 1- some associated faunas. Geos [Caracas], No. 24, pp. 1 1822. llisloirc naturelle des animau.x sans vertebres. Pans, vol. 32, pi. 1-3, figs. 1-2. 7.711 pp. Dominican Columbellid Gastropods: Jung 31
Linne, C. 1 948. Finfiihrung m die Zoologi.sche Noiiicnk laiiir. 2nd cd. Kra- 1758. Syslcnta naturae per regna tria naturae. Editio decima, mer, Frankfurt, 252 pp. reformata. Stockholm, vol. I. pp. 1-824. Rios, K., and Tostes, L. R.
Lyons, W. G. 1981. I new Brasilian Pyrenc (Prosobranchia: Columbellidae).
1 99 1 . Post- Miocene species of Latirus Monlfort. IS 10 (Mollusca: The Nautilus, vol. 95, No. 4, pp. 178-179, figs. 1-3. Fasciolarudae) of southern Florida, with a review of re- Roding, P. F. gional marine bioslraligraphy. Bulletin of the Florida Mu- 1 798. Museum Boltenianiiin. Pars secuiida. Hamburg, pp. i-viii of Natural seum History, Biological Sciences, vol. 35. No. -I- 1-199. Facsimile reprint. 1906. by C. D. Sherbom and 3, pp. 131-208, figs. 1-97. E. R. Sykes. Matthews, S. C. Rutsch, R. F.
1973. Notes on open nomenclature and upon synonymy lists. 1 942. Die Mollusken der Spring\-ale-Schichten (Obermiocaen) von Palaeontology, vol. 16, pp. 713-719. Trinidad (Britisch-West-lndien). Verhandlungen der Na- Maury, C. J. turforschenden Gesellschaft in Basel, vol. 54. pp. 96-182. 1917a. .Santo Domingo type .sections and fossils. I't. I: Mollusca. pis. 3-9. text-figs. 1-2. Bulletins of American Paleontology, vol. 5, No. 29, pp. Sacco, F. 1-251. 1-39. pis. 1890. / molluschi dei terreni terziani del Piemonte e della Li- 1917b. Santo Domingo type .sections and fo.ssils. Pt. 2: .Stratig- guria. Parte 17. (I'olutidae. Marginellidae. Columbelli- raphy. Bulletins of American Paleontology, vol. 5. No. 30, dae). Memorie della Reale Accademia delle Scienze di
1-43, pis. 1-3. I pp. text-fig.. 1 table. Torino. Serie II. vol. 40. pp. 1-76. pis. 1-2. 1925. .4 further contribution to the paleontology ofTrinidad (.Mio- Saunders, J. B., Jung, P., and Biju-Duval, B. cene horizons). Bulletins of American Paleontology, vol. 1986. Neogene Paleontology in the Northern Dominican Repub- 10, No. 42. pp. 1-250. pis. 1-43. lic. 1. Field surveys, lithotogy. environment, and age. Bul- Olsson, A. A. letins of American Paleontology, vol. 89, No. 323, pp. 1- 1922. The .Miocene of northern Costa Rica. Bulletins of Amer- 79, pis. 1-9, text-figs. 1-39, tables 1-4. ican Paleontology, vol. 9. No. 39. pp. 1-168, pis. 1-32. Saunders, J. B., Jung, P., Geister, J., and Biju-Duval, B. Olsson, A. A., and Harbison, .\. 1982. The neogene of the south flank of the Cibao Valley. Do- 1953. Pliocene .Mollusca of southern Florida, with special refer- minican Republic: a stratigraphic study. Transactions of ence to those from North Saint Petersburg. With special the 9th Caribbean Geological Conference (Santo Domin-
chapters on Turridae by William G. Fargo, and Vilrinel- go, 1980), vol. I, pp. 151-160, figs. 1-4.
lidae and freshwater mollusks by Henry .4. Pilsbry. The Sowerby, G. B. 11
Academy of Natural Sciences of Philadelphia. Mono- 1 844. Thesaurus Conchyliorum or .Monographs ofgenera ofshells. graphs, No. 8. pp. 1-458. pis. 1-65, text-figs. 1-2, 2 maps. I 'ol. 1. part 4: Monograph of the genus Columbella. Lon- Perrilliat, M. C. don, pp. 109-146. pis. 36^0. 1972. .Monografia de los moluscos del Mioceno medio dc Santa 1847. [editor]. Thesaurus Conchyliorum or Monographs ofgen- Rosa. Veracruz. Mexico. Parte L (Gasteropodos: Fissu- era of shells. I ol. I: .Monograph of the genus Columbella. rellidae a Olividaej. Paleontologia Mexicana, No. 32, pp. London, pp. 109-146, pis. 36^0. Facsimile reprint. Rot- 1-130. pis. 1-51, I fig., 1 table. terdam and Lisboa. 1981 (W. Backhuys. Rotterdam, Luis Pilsbry, H. A. Pisani Bumay, Lisboa, publishers).
1 922. Revision W. M. 's Tertiary of Gabb Mollusca ofSanto Dom- 1850. Descriptions of new species of fossil shells found by J. S. ingan. Proceedings the of Academy of Natural Sciences Heniker Esq.. in Moore. J. C. On some Tertiary Beds in of Philadelphia, vol. 73. pp. 305^35, pis. 16-47,text-figs. the Island of San Domingo: from notes by J. S. Heniker. 1^8. Esq.. with remarks on the fossils. Quarterly Journal of the Pilsbry, A., H. and Johnson, C. W. Geological Society of London, vol. 6, pp. 44-53, pis. 9- 1917. New Mollusca ofthe Santo Domingan Oligocene. Proceed- 10. ings of the Academy of Natural Sciences of Philadelphia, Swainson, W. vol. 69. pp. 150-202. 1 840. .4 treatise on malacology or the natural classification of Radnin, G. E. shells and shell-fish. London, 419 pp., 130 figs. \911a. The family Coliimhcllidac m the li'estcrn .illantic. The Toula, F. Veliger. vol. 19. No. 4. pp. 403-417, 2 pis.. 4 text-figs. 1911, Nachlrdge zur jungleriidren (pliocdnen) Fauna von Te- 1977b. The Family Columbcllidac in the Western Atlantic. Part huantepec. Jahrbuch der k. k. geologischen Reichsanstalt. Ila. — ThePyreninae. Veliger, vol. 1 19- The 20. No. 2. pp. vol. 61. No. 3 -I- 4, pp. 473-486. pl. 29. 2 figs. 133, figs. 1-47. Weisbord, N.E. 1978. The family Columbellidae in the Western .Atlantic. Pan 1962. Late Cenozoic gastropods from northern Venezuela. Bul- lib. -The Pyrenmae (Continued). The Veliger, vol. 20, letins of American Paleontology, vol. 42, No. 193, pp. 1- No. 4. pp. 328-344, figs. 1^8. 672, pis. 1-48. Reeve, L. A. Woodring, W.P. 1858-1859. Monograph of the genus Columbella. Conchologia 1928. .Miocene mollusks from Bowden. Jamaica. Part 2. Gastro- Iconica (London), vol. I 1. 37 pis. pods and discussion of results. Carnegie Institution of
1 859. Monograph the genus of Meta. Conchologia Iconica (Lon- Washington, Publication No. 385, pp. 1-564, pis. 1-40, don), vol. II, 1 pl. 3 text-figs. Richter, R. 1966. The Panama land bridge as a sea barrier. Proceedings of 1 943. lunfiilirung in die /.oologische Nomcnklatiir. Scnckcnbcr- the American Philosophical Society, vol. 1 10. No. 6. pp. gische Naturforschendc Gesellschaft, Frankfurt, 1 54 pp. 425-433. text-figs. 1-3. tables 1-6. Bulletins of American Paleontology, Volume 106 Plate 2
allx4 Dominican Columbellid Gastropods: Jung 33
Explanation of Plate 2
All figures x 4
Figure Page 1-6. Columbella mercatoria (Linne, 1758) 12 NMB H 17396. NMB locality 17666: Adicora, Paraguana Peninsula. Falcon, Venezuela; Recent. 1-3: not whitened to show color
pattern; 4-6: whitened. 1. 4. front views; 2, 5. rear views; 3, 6. from right side. Height 20.5 mm; width 1 1.9 mm.
7-9. Columbella williamgabbi Weisbord, 1 962 12 Holotype. PRI 26213. Quebrada Mare Abajo, Cabo Blanco, Venezuela; Mare Formation (middle to late Pliocene). 7. front view;
8. rear view; 9. from right side. Height 21.4 mm; width 12.5 mm. 34 Bulletin 344
Explanation of Plate 3
All figures x4
Figure Page 1-3. Columbella platynema Woodnng, 1928 12
Holotype. USNM 369449. Bowden, Jamaica. Bowden Formalion (early Pliocene). 1. front view. 2. rear view; 3. from right side. Height 17.1 mm; width 9.6 mm.
4-9. Columbella alT. rusticoides Heilprin, 1887 12 4-6. NMB H 17397. Locality TU 1230: Rio Cana at Caimito, Dominican Republic; upper part of Cercado Formalion (late Miocene). 4. front view; 5. rear view; 6. from right side. Height 15.0 mm; width 8.4 mm. 7-9. NMB H 17398. Locality TU 1230; Rio Cana at Caimito. Dominican Republic; upper part of Cercado Formation (late Miocene). 7. front view; 8. rear view; 9. from right side. Height 13.4 mm; width 7.8 mm. Note inconspicuous, orthocline riblcts on early teleoconch whorls. 10-15. Columbella lopezana new species 13 10-12. Holotype. NMB H 17400. NMB locality 16935; Lopez section on Rio Yaque del Norte. Dominican Republic; Baitoa Formation (late early to early middle Miocene). 10. front view; II. rear view; 12. from right side. Height 14.9 mm; width 9.3 mm. 13-15. Paratypc. NMB H 17401. NMB locality 17288; Lopez section on Rio Yaque del Norte. Dominican Republic; Baitoa Formation (late early to early middle Miocene). 13. front view; 14. rear view; 15. from right side. Height 14.2 mm; width 8.5 mm. Bulletins of American Paleontology, Volume 106 Plate 3
all x4 Bulletins of American Paleontology, Volume 106 Plate 4
all x4 Dominican Columbellid Gastropods: Juno 35
Explanation of Plate 4
All figures x4
Figure Page 1-9. Eurypyrene inflata (Gabb. 1873b) 14 1-3. Holotype. ANSP 3287. "Santo Domingo". 1. front view, 2. rear view; 3. from right side. Height 25.9 mm; width 13.8 mm. 4-6. NMB H 17405. NMB locality 17290: Lope/ section on Rio Yaque del Norte. Dommican Republic; Baitoa Formation (late early to early middle Miocene). 4. front view; 5. rear view; 6. from right side. Height 22.2 mm; width 12.3 mm. 7-9. Holotype of Columhclla nitlala brassica Maury, 1925. PRI 1059. Brasso District. Central Range, Trinidad; Brasso Formation
(early middle Miocene). 7. front view; 8. rear view; 9. from right side. Height 16.6 mm; width 8.9 mm. 36 Bulletin 344
Explanation of Plate 5
All figures x4
Figure Page 1-3. Eurypyrene venezuelana (Weisbord, 1962) 15
Holotype. PRI 26242. Quebrada Mare Abajo. Cabo Blanco. Venezuela; Mare Formation (middle to late Pliocene). 1. front view;
2. rear view; 3. from right side. Height 18.6 mm; width 1 1.4 mm. 4—6. Eurypyrene occidentalis (Weisbord. 1962) 15 Holotype. PRI 26243. Quebrada Mare Abajo. Cabo Blanco. Venezuela; Mare Formation (middle to late Pliocene). 4. front view; 5. rear view; 6. from right side. Height 22.1 mm; width 12.4 mm. 7-12. Eurypyrene eurynotum (Woodring, 1928) 15 7-9. Holotype. USNM 135512. Bowden. Jamaica; Bowden Formation (early Pliocene). 7. front view; 8. rear view; 9. from right side. Height 14.0 mm; width 7.8 mm.
10-12. NMB H 1 7406. LocalitN TU 1404; Arroyo Babosico. La Barranca, Rio Yaquc del Norte. Dominican Republic; upper part
of Globnrolalia margantae zone (late early Pliocene). 10. front view; 1 1. rear view; 12. from right side. Height 17.2 mm; width 9.6 mm. Bulletins of American Paleontology, Volume 1 06 Plate 5
all x4 Bulletins of American Paleontology, Volume 106 Plate 6
all x6 Dominican Columbellid Gastropods: Jung 37
Explanation of Plate 6
All figures x6
Figure Page 1-15. Parametaria (Dominitaria) islahispaniolae (Maury, 1917a) 17 1-3. Hololypc. PRl 28727. Cercado dc Mao, Bluff 3 of Maury. Dominican Republic; Cercado Formation (late Miocene). 1.
front view; 2. rear view; 3. from right side. Height 1 1.2 mm; width 5.8 mm. 4-6. NMB H 17407. NMB locality 16844: Rio Cana, Dominican Republic; Cercado Formation (late Miocene). 4. front view;
5. rear view; 6. from right side. Height 1 1.2 mm; width 6.1 mm. Low-spircd specimen. 7-9. NMB H 17408. NMB locality 16818: Rio Cana. Dominican Republic; early Pliocene part of Gurabo Formation. 7. front
view; 8. rear view; 9. from right side. Height 10.3 mm; width 5.1 mm. High-spired specimen. 10-12. NMB H 17409. NMB locality 16828: Rio Cana. Dominican Republic; lowest part of Gurabo Formation (latest Miocene).
10. front view; 1 1. rear view; 12. from right side. Height 8.2 mm; width 4.2 mm. 13-15. NMB H 17410. NMB locality 16928: Arroyo Bajon on Rio Mao. Dominican Republic; Cercado Formation (late Miocene).
13. front view; 14. rear view; 15. from right side. Height 1 1.1 mm; width 5.5 mm. Bulletin 344
Explanation of Plate 7
All figures x 6
Figure Page 1-3. Parametaria (Dominitaria) islahispaniolae (Maun, 1917a) 17
NMB H 1741 1. NMB locality 16844; Rio Cana, Dominican Republic; upper part of Cercado Formation (late Miocene). 1. front view; 2. rear view; 3. from right side. Height 16.0 mm; width 8.0 mm. Exceptionally large, low-spired specimen. 4-9. Parametaria (Dominitaria) lopezana new species 19
4-6. Hololype. NMB H 1 74 14 NMB locality 16936: Lopez section on Rio Vaquedcl Norte. Dominican Republic: Baitoa Formation
(late early to early middle Miocene). 4. front view; 5. rear view; 6. from right side. Height I 1.9 mm; width 5.7 mm. 7-9. Paratype. NMB H 17415. Locality TU 1363: Lope? section on Rio >'aque del Norte. Dominican Republic; Baitoa Formation (late early to early middle Miocene). 7. front view: 8. rear view; 9. from right side. Height 15.2 mm; width 6.2 mm. Large specimen. Bulletins of American Paleontology, Volume 106 Plate 7
allx6 Bulletins of American Paleontology, Volume 1 06 Plate 8
1-15: x4 16-21: x8 Dominican Columbellid Gastropods: Jung 39
Explanation of Plate 8
Figures 1-15: x4 Figures 16-21: x8
Figure Page 1-6. Conella ovuloides (C. B. Adams, 1850) 20 Lectotype. MCZ 177372. Turks Island, Bahamas; recent; 1-3. not whitened to show color pattern; 4-6. whitened. 1, 4. front
views; 2, 5. rear views; 3, 6. from right side. Height 16.2 mm; width 6.4 mm.
7-9. Conella perplexabilis (Maury, 1 9 1 7a) 20 Holotype. PRI 28728. Cercado de Mao, Bluff 3 of Maury, Dominican Republic; Cercado Formation (late Miocene). 7. front
view; 8. rear view; 9. from right side. Height 12.7 mm; width 6.2 mm.
10-12. Nitidella alT. cibaoica (Maury, 1917a) 24 NMB H 17424. Locality TU 1250: Rio Verde, Dominican Republic; age probably early Pliocene. 10. front view; 11. rear view; 12. from right side. Height 12.9 mm; width 6.6 mm. 13-15. Conella perplexabilis (Maury, 1917a) 20 NMB H 17418. NMB locality 16913: Rio Mao, Bluff 3 of Maury, Dominican Republic; Cercado Formation (late Miocene). 13. front view; 14. rear view; 15. from right side. Height 15.7 mm; width 8.3 mm. 24 1 6-2 1 . Nitidella caimitana, new species 16-18. Holotype. NMB H 17425. Locality TU 1230: Rio Cana at Caimito, Dominican Republic; Cercado Formation (late Miocene). 16. front view; 17. rear view; 18. from right side. Height 7.2 mm; width 3.4 mm. 19-21. Paratype. NMB H 17426. Locality TU 1230: Rio Cana at Caimito, Dominican Republic; Cercado Formation (late Miocene). 19. front view; 20. rear view; 21. from right side. Height 7.8 mm; width 3.7 mm. 40 Bulletin 344
Explanation of Plate 9
All figures x 4
Figure Page 1-12. Nitidella cibaoica (Maur>-. 1917a) 22
1-3. Lectotype. PRI 28794. Rio Cana at Caimito, Dominican Republic; Cercado Formation (late Miocene). 1. front view; 2.
rear view; 3. from nght side. Height 15.0 mm; width 8.0 mm. 4-6. Paralectotype. PRI 33071. Rio Cana at Caimito, Dominican Republic; Cercado Formation (late Miocene). 4. front view; 5. rear view; 6. from nght side. Height 15.6 mm; width 7.5 mm. 7-9. NMB H 17419. NMB locality 15849: Rio Gurabo. Dominican Republic; lower part ofGurabo Formation (late Miocene). 7. front view; 8. rear view; 9. from right side. Height 18.9 mm; width 9.2 mm. Large specimen. 10-12. NMB H 17420. NMB locality 16934: Rio Gurabo, Dominican Republic; lower part ofGurabo Formation (late Miocene).
10. front view; 1 1. rear view; 12. from right side. Height 17,0 mm; width 8.2 mm, 13-18. Nitidella nitida (Umarck, 1822) 24
NMB H 1 7423, NMB locality 17704: Islade Aves. Venezuela; recent; 13-1 5. not whitened to show color pattern; 16-18. whitened. 13, 16, front views; 14, 17, rear views; 15, 18, from right side. Height 12.1 mm; width 5.8 mm. Bulletins of American Paleontology, Volume 1 06 Plate 9
all x4 Bulletins of American Paleontology, Volume 106 Plate 10 Dominican Columbellid Gastropods: Jung 41
Explanation of Plate 10
All figures x4
Figure Page
1-1 7. Metulella fusiformis Gabb, 1873a 25
1-3. Leclolype. ANSP 3216. "Santo Domingo". 1. front view; 2. rear view; 3. from right side. Height 18.8 mm; width 6.1 mm. 4-6. Holotype of Metulella donmucensis Pilsbry and Johnson, 1917. ANSP 3217. "Santo Domingo". 4. front view; 5. rear view; 6. from right side. Height 16.4 mm; width 5.2 mm. 7, 8. Paratype o( Metulella dommicensis Pilsbry and Johnson, 1917. ANSP 70427. "Santo Domingo". 7. front view; 8. rear view. Height 12.4 mm; width 4.4 mm.
9-1 1 . Holotype of Metulella williamgahhi Maury, 1 9 1 7a. PRI 28745. "Santo Domingo". 9. front view; 10. rear view; 1 1 . from right side. Height 16.5 mm; width 5.7 mm.
12-14. NMB H 1 7428. NMB locality 17266: La Barranca on Rio Yaquedel Norte, Dominican Republic; upper part of G/oftoro/a//a Duirganlae /.one (late early Pliocene). 12. front view; 13. rear view; 14. from right side. Height 19.3 mm; width 6.2 mm. 15-17. NMB H 17430. NMB locality 17271: Arroyo Zalaya, Dominican Republic; Globorotalia iiiciriianlae zone (car\y Pliocene). 15. front view; 16. rear view; 17. from right side. Height 16.4 mm; width 5.6 mm. 18-20. Metulella venusta (G. B. Sowerby II, 1850) 27
NMB H 17433. NMB locality 16910: Rio Mao, Bluff 1 of Maury, Dominican Republic; beds of late Miocene age; 18. front view; 19. rear view; 20. from right side. Height 21.4 mm; width 9.2 mm. 1
42 Bulletin 344
Explanation of Plate 1
All figures x 4
Figure Page
1-9. Metulella venusta (G. B. Sowerby II, 1850) 27
1-3. Lectotype. BMNH GG 20044. "Yaque River", Dominican Republic; 1. front view; 2. rear view; 3. from right side. Height 23.4 mm; width 9.5 mm.
4-6. NMB H 17432. NMB locality 16910: Rio Mao, Bluff 1 of Maury. Dominican Republic; beds of late Miocene age; 4. front view; 5. rear view; 6. from right side. Height 26.3 mm; width 10.3 mm. 7-9, NMB H 17434. NMB locality 15842: Rio Gurabo, Dominican Republic; late Miocene part of Gurabo Formation; 7. front view; 8. rear view; 9. from right side. Height 24.6 mm; width 9.9 mm. 1
Bulletins of American Paleontology, Volume 106 Plate 1
all x4
Dominican Columbellid Gastropods: Jung 43
INDEX
Note: Page numbers are in light face; plate numbers are in bold face type, the page numbers on which principal discussions occur are in italics.
Abbott, 1974 20 Coomans, 1967 16,20 Adams, 1850 20,21 Costa Rrca 7.1 1,12 aif. cihaoica. Nitidella 8,24 atT. ruslicoides. ColiimhcUa 3,/2.13 Dall, 1889 25,27,28 Agueguexquite Formation 25 Dall, 1890 25 Arroyo Babosico 7,15,16 Dall, 1916 16 Arroyo Bajon 10,19,21 Dall and Bartsch, 1911 12 Arroyo Bellaco 24 dominicensis, Metulella 25-27 Arroyo Hondo 19 Dominitaria. new subgenus 8,/(^.17 Arroyo Zalaya 27 dupontii Arroyo Zalaya section 6 Conus 16 Parametaria 16 Baitoa Formation 5,7,8,13-15,17,19,20 elongala. Metulella 27 Berm uda 12 eurynotum
5,6-8 .^ Bolli and Saunders, 1985 12,14 Euiypyrene , / Bowden Formation 7,8,12-16 Pyrene (Eurypyrene) 14,15 Brasso Formation 8,14,15 Eurypyrene Woodhng. 1928 S.I4 Bronn, 1831 25 eurvnotum (Woodnr\g. 1928) 5,6-8, /5 Brown and Pilsbry, 1913 12,20 />;//aW (Gabb, 1873b) 4,6-8, /< 15
I n/lata brassica Maury . 1925 4,14,15 cainiiltina, Nitidella S.5-U4.25 occidenialis (Wehbord. 1962) 5,15
Caloosahatchee Formation 12,13 v('?;<;'rz(c/(j/;t; (Wcisbord, 1962) 5,15 Carter, 1984 12 Cercado de Mao 17,18,21 Florida 12-14 Cercado Formation 5,7,12,13,16-21,23-25,28,29 fusifonnis. Metidella 10,6,11,25,26-28 cihaoica Fusus comptus Bronr\, 1831 25 Columbella (Nitidella) 22 9,6,10,22,23-25 Nitidella Gabb, 1873a 25 Nitidella. aff. 8.24 Gabb, 1873b 6-8,11,14,27 Cibao Valley 5,6 Gibson-Smith and Gibson-Smith, 1979 12 1950 20,21 Clench and Turner, Globorotalia fohsi zone 14 8,7/ Coluinhellal.ama.rck. 1799 Globorotalia inargaritae zone 7,15.26,27.29 1887 3,/2.13 afT. n/5r(cw(/c5 Heilprin, Grant and Gale, 1931 16 14 inflata (Gabb) Gulf of California 16 14 injiata vzT. brassica}\\auTy, 1925 Gulf of Mexico 25,27,28 /i lopczana n. sp 3,5-7, Guppy, 1867 16 menatoria(l.\nne. 1758) 2,12 Guppy, 1876 27 ?!iV;(/a Lamarck, 1822 21,22 Gurabo Formation 17-19,22,24,29 onilata Lamarck, 1822 20 picalaReeve. 1859 16 Heilprin, 1887 12,13 plalynema Woodnng. 1928 3,12,13 ruslicoides Heilprin, 1887 12,13 injlata somersiana Dal\ and Barxsch. 1911 12 Eurypyrene 6-8 .14.\5 sid»nercalona 0]sson. 1922 1,6,7,//, 12,13 Stroinbina 14 v«!i«;a G. B. Sowerby 11, 1850 27 inflata brassica. Eurypyrene 4,14,15 vi7///u;);,?(;/)/>; Weisbord, 1962 2,12 var. brassica. Columbella 14 Columbella (Meta) inflata Isla de Aves, Venezuela 23,24 islahispaniolae Maury, 1917a 16,17 islahispaniolae perple.xabilis Maury. 1917a 20 Columbella (Meta) 16,17 Columbella (Nitidella) cibaoica Maury, 1917a 22 Parametaria 6,7 columbellata. Metulella 25,27,28 Parametaria (Dominitaria) 6,7, 10, /7. 18,19 comptus. Fiisus 25 Co/!e//a Swainson. 1840 8,20 ovi/toa (Lamarck, 1822) 20,21 Jamaica 7,8,12-16 fn'i(/,Wo(C.B. Adams, 1850) 8,20,21 Jung, 1969 16
/> 44 Bulletin 344 Keen, 1971 11,13,16 Panama 12 Keroher and others, 1965 12 Parametana VaW. 1916 16 Kiener, 1834-1880 16,22 Parametana duponia (Kiener. 1834-1880) 16 La Barranca 6-8,15,26-29 islahispanwlae (Maury, 1917a) 6,7 lopezana n. sp 6,7 Lamarck, 1799 11 prototypus (Gup-py , Lamarck, 1822 20,21,23 1867) 16 rutschi iung. 1969 16 ledaluciae. Pyrene (Conella) 20 schiden (KuX%ch, 1942) 16 Linne, 1758 11,12 Parametana (Dominilaria) Liraslrombma Jung, 1989 25 5 islahispanwlae (Maury, 1917a) 6,7.10.77,18.19 lopezana lopezana n. sp 7,5,18,/y,20 Columbella 3,5-7, /i Parameiaria (Parametaria) 16,17 Parametana 6,7 perptexabilis Paramelana (Dominilaria) 7,5.1 8, /9,20 Columbella (Mela) 20 Lopez section 5-7,13-15,19,20 Conella 8,6,7,20,21 Lyons, 1991 12 Perrilliat, 1972 25 picata Marc Formation 12,15 Columbella 16 Matura shell bed 16 Conella 20 Maur>. 1917a 6,7,10,11,16-20,22,24,25,27 Pilsbry, 1922 14,25-27 Maury. 1917b 14 Pilsbry and Johnson, 1917 25 Maury, 1925 14 plalvnenia. Columbella 3.12.13 Melajo Clay Member 16 prototypus. Parametaria 16 mercatoria Pyrene (Conella) ledaluciae K\o& and!oiXe%, 1981 20 Columbella 2,12 Pyrene (Eurypyrene) eurynotum Woodring, 1928 14,15 ]'olula 11 Pyrene KoAing. 1798 11,14 A/c/o Reeve, 1859 16 Mela ovuloides 16 Radwm, 1977a 11,12,14,22 Afeiiilella Gabb. 1873a 8,25 Radwm, 1977b 25 McliiMla Radwin, 1978 14,20 cohimbellala {DaW. 1889) 25,27,28 Reeve, 1859 16 r/(in!;/i/(C«s/5 Pilsbry and Johnson. 1917 25-27 Rio Amina section 6,29 clongala (Tou\a. 1911) 27 Rio Cana section 5-7,17-19,25,29 fiisiformis Gabb. 1873a 10,6.11,25,26-28 Rio Gurabo section 6,7,12,22,23,28,29 vc/)i«/a (G. B. Sowerby II, 1850) 10,11,6,7,11.27.28,29 Rio Mao section 6,7,18,19,21,29 nilliamgabhi Maury\ 1917a 25-27 Rios and Tostes, 1981 20 Mclulella (Melulella) 25 Rio Verde 24 Melutella {Thiannella) 25 Roding, 1798 11 (jfa()«; Perrilliat, 1972 25 rusticoides Mexico 25,27 Columbella 12,13 Columbella af[. 3,/2, 13 Rutsch. 1942 16 nitida rutschi. Parametaria 16 Columbella 21,22 NUidella 22-25 Nitidella Svia\n%on. 1840 8,2/. 22,23 Sacco. 1890 25 Nittdella Saunders « a/., 1982 5 air. c-;7)a«aa( Maury, 1917a) 8.24 Saunders « a/., 1986 5.8-10.12-15,17,19-21,23,24,26,27,29 caimitana n. sp 8.5-7.24,25 Savaneta Glauconitic Sandstone Member 16 W/wo/ta (Maury, 1917a) 9,6,10,22.23-25 schiden. Parametana 16 /na/7«ora/a Swainson. 1840 21 Sherbom and Woodward, 1901 16 mnrfa (Lamarck. 1822) 22-25 somersiana. Columbella 12 Sowerby, 1850 6,7,11,27-29 Spnngvale Formation 16 occidcntahs. Eurypyrene 5, 1 Strombina injlala Gab. 1873b 14 Olsson, 1922 6,7,11,12 St. Thomas, Virgin Islands 20 Olsson and Harbison, 1953 12 submercatoria. Columbella 1,6,7,//, 12,13 olssoni. Melulella (Thiannella) 25 Swainson, 1840 20,21 oyulaia Columbella 20 Conella 20,21 Talparo Formation 16 ovuloides Toula, 1911 27 Conella 8,20,21 Tres Marias Islands, Mexico 16 Meta 16 Trinidad 8,14-16 Dominican Columbellid Gastropods: Jung 45 Venezuela 12,15.22-24 Weisbord, 1962 12.15,22 veneziuiiina. Eurvpvrene 5.15 williamgahbi veniista C'oliiiiilHihi 2,12 Columbella 27 Melulella 25-27 Melulella 10,11,6,7,11.27,28.29 Woodring, 1928 6-8,11-15,25 \ 'oltita mercaloria hinne. 1758 11 Woodring, 1966 16 PREPARATION OF MANUSCRIPTS Bulletins of American Paleontology usually comprises two or more sep- arate papers in two volumes each year. The series is a publication outlet for significant, longer paleontological monographs (i.e.. more than approximately 50 printed pages), for which high quality photographic illustrations and the large quarto format are required. Submissions are welcome from any author, regardless of institutional or organizational affiliation. 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