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Quaternary of the Netherlands Contr. Tert. Quatern. Geol. 27(4) 107-112 2 pis. Leiden, December 1990 Two new freshwater molluscan species from the Early Quaternary of The Netherlands T. Meijer RIJKS GEOLOGISCHE DIENST HAARLEM, THE NETHERLANDS Meijer, T. Two new freshwater molluscan species from the Early Quaternary of The Netherlands. — Contr. Tert. Quatern. Geol. 27(4): 107-112, 2 pis. Leiden, December 1990. Two new freshwater molluscan species are described from Early Quaternary deposits in The Netherlands. Valvata (Valvata) salebrosa be related the Recent Valvata sibirica whereas subtile n. does seem n. sp. may to (Middendorff, 1851), Sphaerium (Sphaerium) sp. not have with Recent to any affinity species. Key words — Freshwater Mollusca, Quaternary, The Netherlands, new taxa. T. Meijer, Rijks Geologische Dienst, P.O. Box 157, 2000 AD Haarlem, The Netherlands. Contents LHB — L.W. Hordijk collection (Brielle, The Netherlands) Introduction 107 p. RGD — Rijks Geologische Dienst (Haarlem, Abbreviations in the 107 text p. The Netherlands) collections Acknowledgements p. 107 RGM — Nationaal Natuurhistorisch Museum 108 Descriptions p. (formerly Rijksmuseum van Geologie en subtile 108 Sphaerium n. sp p. Mineralogie) collections (Leiden, The Valvata salebrosa n. sp p. 110 Netherlands) References p. 112 TMA — T. Meijer collection (Alkmaar, The Netherlands) Introduction 45B/7 — RGD borehole file number Earlier this I reviewed the — below surface year, Quaternary m metres freshwater Mollusca of The Netherlands (Meijer, H — height of shell (in front view) HB — of 1990). Several new species were described. On ac- height bodywhorl of the of that for HA — of ceptance manuscript paper height aperture publication two additional (apparently new) spe- W — width of shell (gastropods); in bivalves their could W denotes the semidiameter cies were found. Although occurrence — of still be noted in that paper, there was not enough WA width aperture which L — shell time to include detailed descriptions, are length of in this now given paper. All dimensions are in mm. ABBREVIATIONS IN THE TEXT ACKNOWLEDGEMENTS The abbreviations used: following are I to the for their am grateful following persons sup- ITZ — Instituut Taxonomische Mr voor Zoologie port: R.G. Moolenbeek (ITZ, Amsterdam) collections (Amsterdam, The Nether- for the opportunity to study material of Valvata lands). frigida and Sphaerium transversum housed in the ITZ 108 for rior than the anterior collections; Mr J.W.M. Jagt (Venlo) correc- leg bifid, slightly longer leg, of the tion of the English, and Mr G.A. Peeters both legs almost reaching the edge hinge (Schiedam, The Netherlands) for preparation of plate. the illustrations. In other specimens the width of the ligament pit with the reach 3/4of the widthof the Both The results of this study are published may hinge plate. of the of C3 the of the consent of the Director Rijks Geologische legs may overhang edge hinge Dienst (Haarlem). plate. Left valve: hinge generally corresponding to that of the right valve. All and PII well developed, DESCRIPTIONS short, PII slightly longer than AIL C2 small, in- the verted V-shaped; both legs may overhang edge subtile Sphaerium n. sp. of the hinge plate. C4 thin, semiparallel to the PI. 1, Figs la-c, 2 C4 lower edge of the ligament pit; apex of to c. 3/4 of the width of the 1990 thin — 166. reaching up hinge Sphaerium sp. [a very Sphaerium]. Meijer, p. plate. Holotype — Pl. 1, Fig. la-c, kept in the RGD Dimensions — H 7.6, L 10.5, W 2.4 (holotype). molluscan collections. Material — RGD: pit Maalbeek, Belfeld (province Locus typicus — Pit Maalbeek, Belfeld, province of of Limburg), Tegelen Formation (Tiglian): il- lustrated the Limburg. Dutch coordinates (mapsheet 58E): are holotype (Pl. 1, Fig. la-c) and a additional 29 208.058/369.505. paratype (Pl. 1, Fig. 2), an paratypes — 12 defective adult Stratum typicum Tegelen Formation (Tiglian, (right valves: 3 adult specimens, Early Pleistocene). specimens, 2 juvenile specimens; left valves: 3 — = 7 adult 2 Derivatio nominis L. subtilis, -is, -e (adj.) thin, adult specimens, defective specimens, fragile, alluding to the thickness of the shell. juvenile specimens). of Lim- Diagnosis — A thin-shelled Sphaerium (Sphaerium) RGM: pit Maalbeek, Belfeld (province its 229 796: valve and RGM 229 species, differing from Sphaerium corneum in burg) (RGM 1 right 797: left All the oblong shape. 1 valve). specimens are from Description — Shell compressed, transversely Tegelen Formation (Tiglian). of oblong. Umbones situated distinctly in front of the RGD: boring Rosmalen 45B/6-7 (province in 110.00-120.00 defective vertical midline, resulting a produced posterior Noord-Brabant), m (2 rounded Formation end. Posterior margin more broadly than right valves); Tegelen (Late Tiglian). the anterior margin. Shell thin, comparable with RGD: boring Rosmalen 45B/7 (province of surface 110.80-113.00 Sphaerium lacustre (Muller, 1774). Outer Noord-Brabant), m (1 right valve); lines which Formation with fine, regular are more pronounc- Tegelen (Late Tiglian). Shell of ed at both sides of the umbo. irregularly un- RGD: boring Asperen 38H/178 (province dulate, according the growth lines, resulting in Zuid-Holland), 103.00-104.00 m (1 hinge frag- which the of left Formation pseudoribs are especially prominent on ment a valve); Tegelen (Late of the Inner central part shell. surface smooth; Tiglian). pallial line and adductor impressions shallow but distinct. Hinge with pronounced cardinals. Hinge between the cardinals and PLATE 1 plate narrow, especially the anterior laterals. subtile Sphaerium n. sp. Right valve (holotype): Ligament pit externally Fig. Right valve a: exterior view, x 6; b: visible, distinct, rather long, reaching about 1/2 of 1. (holotype); interior view, x 6; c: hinge, x 12. the the posterior part of hinge plate. The largest Fig. 2. Left valve (paratype), hinge, x 12. width of the ligament pit equals about 2/3 of the Valvata (Valvata) salebrosa n. width of the hinge plate. Cardinal teeth closer to sp. the anterior than to the laterals. Lateral posterior Fig. 3. Holotype, x 12; a: apical view, b: apertural teeth with umbilical view. well developed, short, bluntly view, c: PI and AI of about size. triangular cusps. equal All specimens from Maalbeek pit, Belfeld (pro- than the likewise sized PIII Both are larger equally vince of Limburg, The Netherlands); Tiglian, inverted and AIII. C3 distinct, U-shaped; poste- Tegelen Formation. RGD collection. 109\1 PLATE 1 110 — At the of Dutch Remarks the type locality, preservation Limburg. coordinates (mapsheet 58E): the shells is excellent. In most of the specimens 208.058/369.505. is still — (parts of) the periostracum present. Stratum typicum Tegelen Formation (Tiglian, subtile has overall Sphaerium n. sp. an Early Pleistocene). North American Derivatio nominis — L. = resemblance with the Recent salebrosus, -a, -urn (adj.) The of the the shell Sphaerium transversum (Say, 1829). hinge uneven, rough, alluding to sculpture. differs from the — A Valvata latter species, however, new Diagnosis species of (Valvata) in the the width of the characterised size and axial species following respects: by a large pronounced central hinge plate is much smaller and the shape and longitudinal sculpture. of the cardinals differs considerably. Curiously, Description — Shell planorbid-shaped and with 2 the of S. subtile in whorls which well hinge (the cardinals particular) 3/4 rapidly expanding are shows rather close resemblance with that of rather a rounded and separated by a deep suture. On the side of the shell each whorl is Sphaerium rosmalense Meijer, 1990, from which it upper more than twice wide the The older differs in almost all other respects. Records of the as as preceding one. 37C/554 new species from boring Zuurland (Meij- whorls are slightly depressed. There is a wide um- have A closer in which all whorls visible. The er, 1990) proved to be erroneous. bilicus are greatest look at the material of that boring confirmed the height of the shell lies at the aperture: the upper rivicola original identification as Sphaerium and lower apertural margins project beyond the of of the The (Lamarck, 1818). Indeed, juvenile specimens plane spire. aperture is detached from with the whorl. The is that species show an overall resemblance preceding peristome continuous, subtile rivicola almost and thickened. Sphaerium n. sp. However, Sphaerium circular, not of has and comparable size is much flatter, a more The protoconch comprises about 1/2 whorl ovate shape, a less pronounced umbo and coarser is not sharply separated from the teleoconch. On sculpture. Differences in shape of juvenile speci- the teleoconch, regularly spaced fine growth lines of several shown in mens species of Sphaerium are are present. Text-Fig. 1. The teleoconch shows axial and spiral orna- The recently described Sphaerium icenicum ment. On the last whorl about 30 axial ribs can be 1986 with Holyoak & Preece, has a taller shell a less posterior end. The sculpture is produced PLATE 2 the is broader and less coarser and hinge plate This is similar of straight. species to Sphaerium Juvenile specimens Sphaerium spp. rivicola. From corneum the Sphaerium (Linne, 1758) subtile Maalbeek Fig. 1. Sphaerium n.sp., paratypes. pit, new differs in characters; particu- species hinge Belfeld (province of Limburg, The Netherlands), the cardinals of different More- larly are a shape. Tiglian (RGD). Fig. 2. Sphaerium corneum (Linné, 1758). Zandvoort and over, the shape of the shell is more oblong and Diemen (province of Noord Holland, The of inequilateral than that Sphaerium corneum. Netherlands), Recent (TMA). the subtile At type locality, Sphaerium n. sp. co- Fig. 3. Sphaerium lacustre (Müller, 1774). Amsterdam (pro- with S. rosmalense and S. corneum. The first occurs vince of Noord Holland, The Netherlands), Re- two of these species were also found together at cent (TMA). 4. rivicola Dordrecht Rosmalen. Fig. Sphaerium (Lamarck, 1818). (province of Zuid Holland, The Netherlands), from The new species is so far known only Holocene (RGD).
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