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Columnals (PDF) 2248 22482 2 4 V. INDEX OF COLUMNALS 8 Remarks: In this section the stratigraphic range given under the genus is the compiled range of all named species based solely on columnals assigned to the genus. It should be noted that this range may and often differs considerably from the range given under the same genus in Section I, because that range is based on species identified on cups or crowns. All other abbreviations and format follow that of Section I. Generic names followed by the type species are based on columnals. Genera, not followed by the type species, are based on cups and crowns as given in Section I. There are a number of unlisted columnal taxa from the literature that are indexed as genera recognized on cups and crowns. Bassler and Moodey (1943) did not index columnal taxa that were not new names or identified genera with the species unnamed. I have included some of the omissions of Bassler and Moodey, but have not made a search of the extensive literature specifically for the omitted citations because of time constraints. Many of these unlisted taxa are illustrated in the early state surveys of the eastern and central United States. Many of the columnal species assigned to genera based on cups or crowns are incorrect assignments. An uncertain, but significant, number of the columnal genera are synonyms of other columnal genera as they are based on different parts of the stem of a single taxon. Also a number of the columnal genera are synonyms of genera based on cups and crowns as they come from more distal parts of the stem not currently known to be associated with the cup or crown. Some of the columnal genera will ultimately prove to have priority over genera named or to be named on the basis of cups and crowns. Some of the columnal taxa may belong to cystoids or other non- crinoid pelmatozoans. Many corrections in this revision of the earlier bibliographies and indices, but by no means most of the taxa, have been made on the stratigraphic age of the species. These are based on new series terms used for the different geologic systems being defined for international terms. Most of these are for the Ordovician and Mississippian. Some corrections are based on more recent publications that incorporate new fossil data. Other corrections will be made for future revisions as ratification of the international terms become available and as time allows for making the corrections. Also the geographic occurrences of the taxa are being revised to take into account the breakup of the U.S.S.R. This has been done for most of the Ordovician listings, and will be made for others as time permits in the future. The corrections are designed to aid in future paleogeographic and evolutionary studies. I consider the columnal classification of Stukalina (1986, 1988) to contain separate families erected for different parts of the stem of a single crinoid genus. Most columnal familes are based on a single genus. Therefore her classification is not used, nor is any columnal classification applied at this time. Hopefully as columnal studies continue the classification applied will be based on the whole animal, however, I do not expect that to occur in the near future. Index: 2249 22492 2 4 9 Abraded columnal Pabian and Holterhoff, 1988, p. 83, Fig. 1, no. B. Webster, 1993, p. 127. Pennsylvanian, Virgilian, Calhoun Shale. United States, Kansas ACANTHOCRINUS Roemer, 1850. Haude, 1981, p. 188, Figs. 1, no. H; 2, middle. Seilacher and MacClintock, 2005, p. 229, Fig. 6, no. F. Acanthocrinus (sensu lato) Stukalina, 1986, p. 83. Webster, 1993, p. 127. Recognized number of named columnal species: 4 Devonian species, 1 subspecies; 3 Emsian (+1 subspecies), 1 early Devonian, series uncertain Devonian, Emsian. Europe; Kazakhstan Acanthocrinus (sensu lato) Stukalina, 1986 = ACANTHOCRINUS Acanthocrinus faouensis Le Menn, 1985. Le Menn, 1987b, p. 817, Pl. 2, figs. 7-9. Gluchowski, 1993b, p. 164, Pl. 1, figs. 1-2. Webster, 1993, p. 127. Devonian, late Emsian, Verveur Formation; Bukowa Gora Shales Formation. France; Poland Acanthocrinus gracilior vermicularis Schmidt, 1941, p. 211, Fig. 58, no. a. Webster, 1973, p. 277. Devonian, Emsian, upper part of Coblenzian Beds. Germany Acanthocrinus kersivienensis Le Menn, 1985, p. 42, Pl. 5, figs. 1-9. Le Menn, 1987b, p. 817, Pl. 2, figs. 10, 15. Webster, 1988, p. 169. Webster, 1993, p. 127. Devonian, late Emsian, Bolast Formation. France, Armorican Massif Acanthocrinus? monstruosus Yeltyschewa and Stukalina in Stukalina, 1971, p. 197, Pl. 2, fig. 20. Stukalina, 1977c, p. 151, Pl. 31, figs. 1-3. Webster, 1977, p. 184. Stukalina, 1978a, p. 56, Pl. 13, fig. 15. Gluchowski, 1981b, p. 32, Pl. 2, fig. 5. Stukalina, 1986, p. 83, Pl. 20, figs. 18-14. Webster, 1986, p. 325. Stukalina, 1988c, p. 64, Pl. 13, figs. 1-2. Webster, 1988, p. 169. Stukalina, 1991, p. 154, Pl. 42, figs. 9-10; Fig. 11, no. a. Webster, 1993, p. 127. Devonian, Emsian, Sardzhal Horizon. Kazakhstan, Pribalkhash. Poland Acanthocrinus rex Jaekel, 1895. Ubaghs in Moore and Teichert, 1978, p. T93, Fig. 69, no. 1. Webster, 1986, p. 325. Südkamp, 1995, p. 114, unnumbered fig. p. 114. Early Devonian, series uncertain. Germany Acanthocrinus sp. Breuer et al., 1992, p. 66, Pl. 12, fig. 4. Early Devonian, series uncertain. Germany ACANTHODOCRINUS Yeltsychewa, 1979. *Acanthodocrinus rakverensis Yeltyschewa, 1979. Yeltyschewa, 1979, p. 102. Webster, 1986, p. 325. Recognized number of named columnal species: 2 Ordovician, Katian species Ordovician, Katian. Estonia 2250 22502 2 5 0 Acanthodocrinus magnispinosus Yeltyschewa, 1979, p. 102, Pl. 1, fig. 12. Webster, 1986, p. 325. Ordovician, Katian, Vormsi Horizon. Estonia *Acanthodocrinus rakverensis Yeltsychewa, 1979, p. 102, Pl. 1, figs. 6-7. Webster, 1986, p. 325. Ordovician, Katian, Vezenberg Horizon. Estonia ACBASTAUCRINUS Sisova, 1979 = FLORICYCLUS ACBASTAURCRINUS Sisova, 1979 = FLORICYCLUS *Acbastaucrinus acbastauensis Sisova, 1979 = Floricyclus acbastauensis Acbastaucrinus affectatus Sisova, 1988 = Floricyclus affectatus Acbastaucrinus? martynovae Sisova, 1983 = Floricyclus martynovae Acbastaucrinus quinquangularis Sisova, 1983 = Floricyclus martynovae Acbastaucrinus undosus Sisova, 1979 = Floricyclus undosus ACTINOCRINITES Miller, 1821. Recognized number of named columnal species: 9 species. 8 Devonian,7 Eifelian, 1 Famennian; 1 Mississippian, Tournaisian Devonian, Eifleian, Famennian. Germany Devonian, Famennian. United Kingdom Mississippian, Tournaisian. Ireland Actinocrinites amphora Gilbertson, 1836. Gilbertson in Portlock, 1843, p. 348, Pl. 16, figs. 1a-1b. Mississippian, Tournaisian. Ireland Actinocrinites cingulatus Goldfuss, 1866 = Pestericrinus cingulatus Actinocrinites cingulatus nobis Goldfuss, 1866 = Pentagonocyclicus cingulatus = Salairocrinus cingulatus = Pestericrinus cingulatus Actinocrinites cingulatus Goldfuss, 1831, p. 195, Pl. 59, figs. 7a-7b. Bassler and Moodey, 1943, p. 268. Melocrinus cingulatus (Goldfuss, 1831). d'Orbigny, 1840, p. 104. Actinocrinus cingulatus (Goldfuss, 1831). Bronn, 1848, p. 14. Dujardin and Hupé, 1862, p.139. Asterocrinus cingulatus (Goldfuss, 1831). d’Orbigny, 1850, p. 104. Devonian, Eifelian. Germany 2251 22512 2 5 Actinocrinites laevis Miller, 1821. (Note: Probably not A. laevis and more than one 1 species. GDW) Goldfuss, 1831, p. 193, Pl. 59, figs. 3b-3e. Early Carboniferous. Belgium Kutorga, 1845-1846, p. 134, Pl. 8, fig. 6. Devonian, Limestone of Wochana. Poland Quenstedt, 1852, p. 622, Pl. 55, figs. 5-6. Quenstedt, 1867, p. 746, Pl. 69, figs. 5-6. Quenstedt, 1885, p. 961, Pl. 78, figs. 4-5. Devonian, Eifelian. Germany Actinocrinites moniliferus Goldfuss, 1831, p. 190, Pl. 59, figs. 10a-10b. Bassler and Moodey, 1943, p. 271. Melocrinus moniliferus (Goldfuss, 1831). d'Orbigny, 1840, p. 104. Bassler and Moodey, 1943, p. 271. Asterocrinus moniliferus (Goldfuss, 1831). d’Orbigny, 1850, p. 104. Devonian, Eifelian. Germany ?Actinocrinites moniliformis Miller, 1821 (non Phillips, 1839) = Cyathocrinites pinnatus Actinocrinites muricatus Goldfuss, 1831, p. 195, Pl. 59, figs. 8a-8c. Roemer, F. A., 1843, p. 9, Pl. 3, fig. 17. Roemer, F. A., 1866b, p. 227. Bassler and Moodey, 1943, p. 272. Actinocrinus muricatus (Goldfuss, 1831). Bronn, 1848, p. 14. Dujardin and Hupé, 1862, p.139. Quenstedt, 1876, p. 649, Pl. 112, fig. 128. Asterocrinus muricatus (Goldfuss, 1831). d’Orbigny, 1850, p. 104. Devonian, Eifelian, Eifel Limestone. Germany Actinocrinites cf. muricatus Goldfuss, 1831. Quenstedt, 1876, p. 649, Pl. 112, fig. 127. Devonian, Eifelian, Eifel Limestone. Germany Actinocrinites nodiger Eichwald, 1856, 1857 = Hexacrinus nodiger Actinocrinites nodulosus Goldfuss, 1831, p. 195, Pl. 59, fig. 9. Roemer, F. A., 1843, p. 9, Pl. 12, fig. 7. Quenstedt, 1852, p. 623, Pl. 54, figs. 33a-33b. Roemer, F. A., 1866b, p. 227. Quenstedt, 1867, p. 746, Pl. 68, figs. 33a-33b. Quenstedt, 1885, p. 962, Pl. 78, fig. 8. Bassler and Moodey, 1943, p. 272. Actinocrinus nodulosus (Goldfuss, 1831). Bronn, 1848, p. 14. Actinocrinus nodulus (Goldfuss, 1831). Dujardin and Hupé, 1862, p. 139 (sic), Asterocrinus nodulosus (Goldfuss, 1831). d’Orbigny, 1850, p. 104. Devonian, Eifelian, Schalke Limestone; Kahleberg Schraubstone. Germany Actinocrinites plumosus (Hall); Shumard, 1868 = Eomyelodactylus ?plumosus Actinocrinites tenuistriatus Phillips, 1841. Roemer, F. A., 1843, p. 9, Pl. 3, fig. 16. Roemer, F. A., 1866b, p. 229. Asterocrinus tenuistriatus (Phillips, 1841). d’Orbigny, 1850, p. 104. Early - Late Devonian, Famennian, Kalkberg Ironstone; Pilton Beds. Germany; United Kingdom 2252 22522 2 5 2
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