Lonicera Xylosteum

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Lonicera xylosteum - Fly Honeysuckle (Caprifoliaceae) -------------------------------------------------------------------------------------------- Lonicera xylosteum is a tough, utilitarian shrub known USAGE for emerging and mature gray-green foliage, compact Function rounded habit of some cultivars, and urban and salt -informal or formal hedge spray tolerance. -border, entranceway, low screen, or specimen shrub -parking lot or street island small shrub FEATURES -can be a facer shrub if kept pruned Form Texture -medium texture in foliage and when bare -thick density in foliage and when bare Assets -densely rounded habit -foliage emerging gray-green to silvery-green -urban tolerant, including exceptional salt spray and drought tolerance -spring flowers -low maintenance plant -medium shrub Liabilities -to 8' tall x 8' wide but frequently pruned much smaller -poor autumn color (but holds green foliage late) -radiating mound -some of the "compact" cultivars can eventually get -slow to medium rate quite spreading and tall if left unpruned Habitat Culture -Zone 4 -full sun to partial shade -Native to Europe -urban tolerant and very adaptable, especially to poor soils, soil pHs, restricted root zones, drought, salt SELECTIONS spray, and heavy pruning, but is not tolerant of wet Alternates sites or poorly drained sites -shrubs for urban stress situations (species of Berberis, -propagated primarily by rooted stem cuttings Forsythia, Lonicera, Viburnum, Weigela, etc.) -Honeysuckle Family, with virtually no disease or pest -shrubs with a tight growth habit that grow slowly problems (Berberis 'Crimson Pygmy', Buxus hybrids, Fothergilla -low availability, in containers or B&B form gardenii, Picea abies [shrub forms], Viburnum plicatum Foliage var. tomentosum 'Newport' [Nanum Newport'], etc.) -gray-green -winter salt-spray-tolerant companion plants -deciduous (Hemerocallis, Myrica pensylvanica, etc.) -opposite Cultivars - Variants - Related species -elliptical -Lonicera xylosteum 'Compacta' ('Emerald Mound') - -short blue-green mature foliage, mounding to 3' as a low petiole informal hedge with low maintenance -one of the -Lonicera x xylosteoides 'Clavey's Dwarf' - earliest hybrid of Fly and Tatarian (Lonicera tatarica) shrubs to Honeysuckles; up to 6' tall, used as a medium-sized leaf out informal hedge that needs little maintenance; this -buds and cultivar is often incorrectly treated as Lonicera young xylosteum 'Clavey's Dwarf' foliage emerging with silvery hairs and remaining pubescent -Lonicera japonica - Japanese Honeysuckle - -green tinged with purple vigorous vine or twisting groundcover, adorned with Flowers fragrant white and light yellow flowers all summer -white to cream long; good choice for a quick vine or an erosion control -in May and effective for 2 weeks groundcover -flowers borne upright with 4 per node -Lonicera maackii - Amur Honeysuckle - probably -not as showy as other Honeysuckles but still attractive the most invasive shrub in northern climates, being Fruits readily seeded near any perching site by birds; habit is -virtually no fruit set an arching vase (15' x 15') that flowers prolifically (and Twigs somewhat fragrantly) in May-June, and sets many red -tan to white-brown fruits in Sept., maturing and devoured by late Nov. by -winter buds extend out at a 45 degree angle birds (Do Not Use! see plantNote on this species) -rather stout, zigzag, and slightly rough Trunk -not applicable.

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Lonicera Spp

Species: Lonicera spp. http://www.fs.fed.us/database/feis/plants/shrub/lonspp/all.html SPECIES: Lonicera spp. Choose from the following categories of information. Introductory Distribution and occurrence Botanical and ecological characteristics Fire ecology Fire effects Fire case studies Management considerations References INTRODUCTORY SPECIES: Lonicera spp. AUTHORSHIP AND CITATION FEIS ABBREVIATION SYNONYMS NRCS PLANT CODE COMMON NAMES TAXONOMY LIFE FORM FEDERAL LEGAL STATUS OTHER STATUS AUTHORSHIP AND CITATION: Munger, Gregory T. 2005. Lonicera spp. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/ [2007, September 24]. FEIS ABBREVIATIONS: LONSPP LONFRA LONMAA LONMOR LONTAT LONXYL LONBEL SYNONYMS: None NRCS PLANT CODES [172]: LOFR LOMA6 LOMO2 LOTA LOXY 1 of 67 9/24/2007 4:44 PM Species: Lonicera spp. http://www.fs.fed.us/database/feis/plants/shrub/lonspp/all.html LOBE COMMON NAMES: winter honeysuckle Amur honeysuckle Morrow's honeysuckle Tatarian honeysuckle European fly honeysuckle Bell's honeysuckle TAXONOMY: The currently accepted genus name for honeysuckle is Lonicera L. (Caprifoliaceae) [18,36,54,59,82,83,93,133,161,189,190,191,197]. This report summarizes information on 5 species and 1 hybrid of Lonicera: Lonicera fragrantissima Lindl. & Paxt. [36,82,83,133,191] winter honeysuckle Lonicera maackii Maxim. [18,27,36,54,59,82,83,131,137,186] Amur honeysuckle Lonicera morrowii A. Gray [18,39,54,60,83,161,186,189,190,197] Morrow's honeysuckle Lonicera tatarica L. [18,38,39,54,59,60,82,83,92,93,157,161,186,190,191] Tatarian honeysuckle Lonicera xylosteum L.
Japanese Honeysuckle Wildland (Lonicera Japonica Thunb.) Gary N

Japanese Honeysuckle Wildland (Lonicera japonica Thunb.) Gary N. Ervin, Ph.D., Associate Professor, Mississippi State University John D. Madsen, Ph.D., Extension/Research Professor, Mississippi State University Ryan M. Wersal, Research Associate, Mississippi State University Fig. 1. Japanese honeysuckle vine climbing up a tree. Fig. 2. Flowers of Japanese honeysuckle. Introduction Problems Created Japanese honeysuckle was introduced from Japan in the early 1800s and now is one of the most commonly encoun- tered exotic weeds in the Mid-South. This species frequently overtops and displaces native plants and forestry species in any habitat, but particularly where natural or human activities creates edges. Japanese honeysuckle also is somewhat shade tolerant and can be found in relatively densely canopied forest. This species perenniates with the aid of well de- veloped root and rhizome systems, by which it also is capable of spreading vegetatively, in addition to rooting at nodes along aboveground stems. Both features contribute substantially to its rapid dominance over native vegetation. Regulations Japanese honeysuckle is listed as a noxious weed in CT, MA, NH, and VT. In the southeast, Japanese honeysuckle is considered a severe invasion threat in KY, SC, and TN. It is considered one of the top ten invasive plants in GA, and is listed as a category one invasive in Florida. It is not presently listed in any noxious weed legislation in southern states. Description Vegetative Growth Japanese honeysuckle exhibits a semi-evergreen to evergreen life cycle and is readily identified during winter by its per- sistent green foliage. Its vines may climb and/or spread along the ground to lengths of 80’.
Phylogeny and Phylogenetic Taxonomy of Dipsacales, with Special Reference to Sinadoxa and Tetradoxa (Adoxaceae)

PHYLOGENY AND PHYLOGENETIC TAXONOMY OF DIPSACALES, WITH SPECIAL REFERENCE TO SINADOXA AND TETRADOXA (ADOXACEAE) MICHAEL J. DONOGHUE,1 TORSTEN ERIKSSON,2 PATRICK A. REEVES,3 AND RICHARD G. OLMSTEAD 3 Abstract. To further clarify phylogenetic relationships within Dipsacales,we analyzed new and previously pub- lished rbcL sequences, alone and in combination with morphological data. We also examined relationships within Adoxaceae using rbcL and nuclear ribosomal internal transcribed spacer (ITS) sequences. We conclude from these analyses that Dipsacales comprise two major lineages:Adoxaceae and Caprifoliaceae (sensu Judd et al.,1994), which both contain elements of traditional Caprifoliaceae.Within Adoxaceae, the following relation- ships are strongly supported: (Viburnum (Sambucus (Sinadoxa (Tetradoxa, Adoxa)))). Combined analyses of C ap ri foliaceae yield the fo l l ow i n g : ( C ap ri folieae (Diervilleae (Linnaeeae (Morinaceae (Dipsacaceae (Triplostegia,Valerianaceae)))))). On the basis of these results we provide phylogenetic definitions for the names of several major clades. Within Adoxaceae, Adoxina refers to the clade including Sinadoxa, Tetradoxa, and Adoxa.This lineage is marked by herbaceous habit, reduction in the number of perianth parts,nectaries of mul- ticellular hairs on the perianth,and bifid stamens. The clade including Morinaceae,Valerianaceae, Triplostegia, and Dipsacaceae is here named Valerina. Probable synapomorphies include herbaceousness,presence of an epi- calyx (lost or modified in Valerianaceae), reduced endosperm,and distinctive chemistry, including production of monoterpenoids. The clade containing Valerina plus Linnaeeae we name Linnina. This lineage is distinguished by reduction to four (or fewer) stamens, by abortion of two of the three carpels,and possibly by supernumerary inflorescences bracts. Keywords: Adoxaceae, Caprifoliaceae, Dipsacales, ITS, morphological characters, phylogeny, phylogenetic taxonomy, phylogenetic nomenclature, rbcL, Sinadoxa, Tetradoxa.
Japanese Honeysuckle)

No. 09 March 2010 Lonicera japonica (Japanese Honeysuckle) Initial Introduction and Expansion in Range Introduced to the United States in the early to mid-1800s as an ornamental plant, Lonicera japonica is native to East Asia, including Japan and Korea. It is still promoted by some landscapes architects for its rapid growth and fragrant flowers that linger on the vine throughout most of the summer. Wildlife managers have promoted the plant as winter forage, particularly for deer. Still others are nostalgic about this plant, remembering the sweet nectar they enjoyed as children. Lonicera japonica is now found across the southern United States from California to New England and the Great Lakes Region. Lonicera japonica spreads locally by long aboveground runners and underground rhizomes. The runners develop roots at the nodes (stem and leaf junctions) so this plant often forms dense mats. Under high light conditions, the plants are able to flower and produce fruits that can be dispersed long distances primarily by birds. Description and Biology • Perennial trailing or twining vine. In North Carolina, it is considered semi-evergreen to evergreen. • Young stems are slender, while older stems are hollow and up to 2 inches in diameter with brownish bark that peels in long strips. • Oblong to oval shaped leaves are 1 to 2 and a half inches long arranged in opposite pairs along the stem. Mature leaves have smooth edges and young leaves are often lobed. • White and pale yellow, trumpet-shaped flowers occur in pairs from between the leaves and bloom from late April to August. • Small, nearly spherical, black fruits mature in autumn.
Viburnum Opulus Var. Americanum

Viburnum opulus L. var. americanum (Mill.) Ait. (American cranberrybush): A Technical Conservation Assessment Prepared for the USDA Forest Service, Rocky Mountain Region, Species Conservation Project May 8, 2006 James E. Nellessen Taschek Environmental Consulting 8901 Adams St. NE Ste D Albuquerque, NM 87113-2701 Peer Review Administered by Society for Conservation Biology Nellessen, J.E. (2006, May 8). Viburnum opulus L. var. americanum (Mill.) Ait. (American cranberrybush): a technical conservation assessment. [Online]. USDA Forest Service, Rocky Mountain Region. Available: http://www.fs.fed.us/r2/projects/scp/assessments/viburnumopulusvaramericanum.pdf [date of access]. ACKNOWLEDGMENTS Production of this assessment would not have been possible without the help of others. I wish to thank David Wunker for his help conducting Internet searches for information on Viburnum opulus var. americanum. I wish to thank Dr. Ron Hartman for supplying photocopies of herbarium specimen labels from the University of Wyoming Rocky Mountain Herbarium. Numerous other specimen labels were obtained through searches of on-line databases, so thanks go to those universities, botanic gardens, and agencies (cited in this document) for having such convenient systems established. I would like to thank local Region 2 botanists Bonnie Heidel of the Wyoming Natural Heritage Program, and Katherine Zacharkevics and Beth Burkhart of the Black Hills National Forest for supplying information. Thanks go to Paula Nellessen for proofing the draft of this document. Thanks go to Teresa Hurt and John Taschek of Taschek Environmental Consulting for supplying tips on style and presentation for this document. Thanks are extended to employees of the USDA Forest Service Region 2, Kathy Roche and Richard Vacirca, for reviewing, supplying guidance, and making suggestions for assembling this assessment.
Mistaken Identity? Invasive Plants and Their Native Look-Alikes: an Identification Guide for the Mid-Atlantic

Mistaken Identity ? Invasive Plants and their Native Look-alikes an Identification Guide for the Mid-Atlantic Matthew Sarver Amanda Treher Lenny Wilson Robert Naczi Faith B. Kuehn www.nrcs.usda.gov http://dda.delaware.gov www.dsu.edu www.dehort.org www.delawareinvasives.net Published by: Delaware Department Agriculture • November 2008 In collaboration with: Claude E. Phillips Herbarium at Delaware State University • Delaware Center for Horticulture Funded by: U.S. Department of Agriculture Natural Resources Conservation Service Cover Photos: Front: Aralia elata leaf (Inset, l-r: Aralia elata habit; Aralia spinosa infloresence, Aralia elata stem) Back: Aralia spinosa habit TABLE OF CONTENTS About this Guide ............................1 Introduction What Exactly is an Invasive Plant? ..................................................................................................................2 What Impacts do Invasives Have? ..................................................................................................................2 The Mid-Atlantic Invasive Flora......................................................................................................................3 Identification of Invasives ..............................................................................................................................4 You Can Make a Difference..............................................................................................................................5 Plant Profiles Trees Norway Maple vs. Sugar
Elderberry, Or Have Respect for Your Elders!

Enchanted Elderberry, or Have Respect for Your Elders! Elderberries have been blooming along roadsides. Their blossoms spread across the bushes like white lace doilies. Do you remember infamous poison-laced elderberry wine that was used to put lonely gentlemen out of their misery by little old ladies in the comedy, "Arsenic and Old Lace"? Did "old lace" refer to the ladies, the "laced" wine, or the appearance of elderberry blossoms? Who knows? Image 1: A nice colony blooming several years ago off Gilgal Road in Abbeville County. Many tiny white flowers form a large more or less flat-topped blossom known botanically as a cyme. Each flower is "complete" with five sepals, five petals, five stamens that produce pollen, and one pistil that produces eggs and seed. And as complete flowers they are also "perfect" because both male and female parts are present. Incomplete flowers lacking sepals or petals can still be perfect as long as both male and female structures are present! Image 2: One cluster of flowers, a cyme. Image 3: Individual flowers. Can you count petals and stamens? By summer’s end bushes will be in fruit, and, botanically speaking each fruit is a drupe! Other familiar examples of drupes are cherries, peaches, and olives, fruits that have a single seed surrounded by the fleshy part that developed from the flower's ovary. Image 4: Beautiful juicy ripe drupes ready for pie, jelly or wine. Elderberry is in the honeysuckle family, Caprifoliaceae, whose Greek root words mean "goat leaves." That makes sense for most honeysuckles, with undivided (simple) leaves in twos arranged opposite one another on the stem, and shaped sort of like goat’s ears! Leaves of elderberry also have opposite arrangement, but they are compound leaves, with the green blade divided into smaller leaflets not at all resembling goat’s ears! Their leaves actually resemble somewhat those of an ash tree.
FINAL Phyton-Lonicera Maackii

Thompson, R.L. and D.B. Poindexter. 2011. Species richness after Lonicera maackii removal from an old cemetery macroplot on Dead Horse Knob, Madison County, Kentucky. Phytoneuron 2011-50: 1–16. Published 10 Oct 2011. ISSN 2153 733X SPECIES RICHNESS AFTER LONICERA MAACKII REMOVAL FROM AN OLD CEMETERY MACROPLOT ON DEAD HORSE KNOB, MADISON COUNTY, KENTUCKY RALPH L. THOMPSON 1, 2 1 Hancock Biological Station Murray State University Murray, Kentucky 42071 2 Berea College Herbarium, Biology Department Berea, Kentucky 40404 [email protected] DERICK B. POINDEXTER I.W. Carpenter, Jr. Herbarium Appalachian State University, Biology Department Boone, North Carolina 28608 [email protected] ABSTRACT The predominance of Lonicera maackii (Rupr.) Herder (Amur Honeysuckle) in central Kentucky has made it a significant invasive for continued community interaction studies. To better understand the dynamics of vegetation succession with respect to this species and overall species richness, quantitative floristics of two macroplots were made at the summit (312 m) of Dead Horse Knob (Rucker’s Knob) near Berea, in Madison County, east-central Kentucky. An old abandoned cemetery was cleared of L. maackii and a macroplot (20 x 12 m) served as a test plot, while a second macroplot was placed within a dense thicket of L. maackii to serve as a reference plot. Thirty quadrats (1 x 1 m) were randomly placed within each macroplot as a means to determine species frequency. A full floristic survey was then conducted of each macroplot. Results from frequency data suggest that native annual and perennial species will quickly recolonize an area after removal of Amur Honeysuckle and become the most important components of the site.
Flora of South Australia 5Th Edition | Edited by Jürgen Kellermann

Flora of South Australia 5th Edition | Edited by Jürgen Kellermann KEY TO FAMILIES1 J.P. Jessop2 The sequence of families used in this Flora follows closely the one adopted by the Australian Plant Census (www.anbg.gov. au/chah/apc), which in turn is based on that of the Angiosperm Phylogeny Group (APG III 2009) and Mabberley’s Plant Book (Mabberley 2008). It differs from previous editions of the Flora, which were mainly based on the classification system of Engler & Gilg (1919). A list of all families recognised in this Flora is printed in the inside cover pages with families already published highlighted in bold. The up-take of this new system by the State Herbarium of South Australia is still in progress and the S.A. Census database (www.flora.sa.gov.au/census.shtml) still uses the old classification of families. The Australian Plant Census web-site presents comparison tables of the old and new systems on family and genus level. A good overview of all families can be found in Heywood et al. (2007) and Stevens (2001–), although these authors accept a slightly different family classification. A number of names with which people using this key may be familiar but are not employed in the system used in this work have been included for convenience and are enclosed on quotation marks. 1. Plants reproducing by spores and not producing flowers (“Ferns and lycopods”) 2. Aerial shoots either dichotomously branched, with scale leaves and 3-lobed sporophores or plants with fronds consisting of a simple or divided sterile blade and a simple or branched spikelike sporophore ..................................................................................
Vascular Plants for High Park and the Surrounding Humber Plains, 2008

Annotated Checklist of the Vascular Plants for High Park and the Surrounding Humber Plains September 2008 Steve Varga Ontario Ministry of Natural Resources Aurora District Acknowledgements The Ministry has undertaken biological inventories of High Park and its environs over the past 32 years. The first botanical survey was carried out by Karen L. McIntosh as part of an ecological survey of High Park that focussed on Grenadier Pond and the surrounding uplands (Wainio et al. 1976). Botanical surveys were also carried out by the author in 1980, 1982 and 1988. In 1989, this survey information was put together into an Area of Natural and Scientific Interest (ANSI) inventory report of the Park that recognized the natural areas of High Park as a provincial life science ANSI. Subsequent botanical surveys were carried out by the author between 1997 and 2008. Botanical records from High Park have also been kindly provided by Dr. Paul. M. Catling, Dr. Paul F. Maycock, Roger Powley, Charles Kinsley, Gavin Miller, Bohdan Kowalyk, Diana Banville and other members of the Toronto Field Naturalists (TFN), City of Toronto Parks, Forestry & Recreation Division including present and former staff such as Terry Fahey, Cara Webster and Richard Ubens among others, the High Park Community Advisory Committee and it Natural Environment Committee chaired by Karen Yukich, and James Kamstra who in 2007 and 2008 is mapping the location and numbers of rare plants in the Park. The Vascular Plant Herbarium (TRT) at the Royal Ontario Museum and the University of Toronto, Erindale College Herbarium (TRTE) were also examined for historical and recent records from the Park and its surrounding areas.
Appendix 6: Invasive Plant Species

USDA Forest Service Understanding i-Tree – Appendix 6: Invasive Plant Species APPENDIX 6 Invasive Plant Species The following is a list of invasive tree and shrub species by state that are included in i-Tree database (version 6). Each list of invasive species is followed by the reference of the source which were obtained circa 2014. Some of the Web addresses are no longer working; some have been relocated to alternative sites. State-specific invasive species lists will be updated in the future. Alabama Ailanthus altissima Lonicera japonica Poncirus trifoliate Albizia julibrissin Lonicera maackii Pyrus calleryana Ardisia crenata Lonicera morrowii Rosa bracteata Cinnamomum camphora Lonicera x bella Rosa multiflora Elaeagnus pungens Mahonia bealei Triadica sebifera Elaeagnus umbellata Melia azedarach Vernicia fordii Ligustrum japonicum Nandina domestica Wisteria sinensis Ligustrum lucidum Paulownia tomentosa Ligustrum sinense Polygonum cuspidatum Alabama Invasive Plant Council. 2007. 2007 plant list. Athens, GA: Center for Invasive Species and Ecosystem Health, Southeast Exotic Pest Plant Council. http://www.se-eppc.org/ alabama/2007plantlist.pdf Alaska Alnus glutinosa Lonicera tatarica Sorbus aucuparia Caragana arborescens Polygonum cuspidatum Cytisus scoparius Prunus padus Alaska National Heritage Program. 2014. Non-Native plant data. Anchorage, AK: University of Alaska Anchorage. http://aknhp.uaa.alaska.edu/botany/akepic/non-native-plant-species- list/#content Arizona Alhagi maurorum Rhus lancea Tamarix parviflora Elaeagnus angustifolia Tamarix aphylla Tamarix ramosissima Euryops multifidus Tamarix chinensis Ulmus pumila Arizona Wildland Invasive Plant Working Group. 2005. Invasive non-native plants that threaten wildlands in Arizona. Phoenix, AZ: Southwest Vegetation Management Association https:// www.swvma.org/wp-content/uploads/Invasive-Non-Native-Plants-that-Threaten-Wildlands-in- Arizona.pdf (Accessed Sept 3.
CAPRIFOLIACEAE Christine Pang, Darla Chenin, and Amber M

Comparative Seed Manual: CAPRIFOLIACEAE Christine Pang, Darla Chenin, and Amber M. VanDerwarker (Completed, May 10, 2019) This seed manual consists of photos and relevant information on plant species housed in the Integrative Subsistence Laboratory at the Anthropology Department, University of California, Santa Barbara. The impetus for the creation of this manual was to enable UCSB graduate students to have access to comparative materials when making in-field identifications. Most of the plant species included in the manual come from New World locales with an emphasis on Eastern North America, California, Mexico, Central America, and the South American Andes. Published references consulted1: 1998. Moerman, Daniel E. Native American ethnobotany. Vol. 879. Portland, OR: Timber press. 2009. Moerman, Daniel E. Native American medicinal plants: an ethnobotanical dictionary. OR: Timber Press. 2010. Moerman, Daniel E. Native American food plants: an ethnobotanical dictionary. OR: Timber Press. Species included herein: Symphoricarpos spp. 1 Disclaimer: Information on relevant edible and medicinal uses comes from a variety of sources, both published and internet-based; this manual does NOT recommend using any plants as food or medicine without first consulting a medical professional. Symphoricarpos spp. Family: Caprifoliaceae Common Names: Snowberry, Snowberry Bush, Waxberry, Ghostberry, Coralberry Habitat and Growth Habit: The distribution of this plant ranges from North to Central America. Human Uses: Humans have used this plant for ornamental purposes, after flowering the berries change color seasonally. Sources Consulted: http://urbanbutterflygarden.co.uk/snowberry-symphoricarpos-spp, accessed May 10, 2019 https://www.fs.fed.us/database/feis/plants/shrub/symlon/all.html, accessed May 10, 2019 .