Funghi E Natura Gruppo Di Padova
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Histological Studies of Mycorrhized Roots and Mycorrhizal-Like-Structures in Pine Roots
Benchmark Histological Studies of Mycorrhized Roots and Mycorrhizal-Like-Structures in Pine Roots Carla Ragonezi 1,* and Maria Amely Zavattieri 1,2,3 ID 1 Banco de Germoplasma ISOPlexis, Campus da Penteada, Universidade da Madeira, 9020-105 Funchal, Portugal; [email protected] 2 Departamento de Biologia, Pólo da Mitra Apartado 94, 7002-554 Évora, Portugal 3 Instituto de Ciências da Terra (ICT), Colégio Luís António Verney, Rua Romão Ramalho 59, 7000-671 Évora, Portugal * Correspondence: [email protected]; Tel.: +351-925193860 or +351-291705000 (ext. 5408) Received: 6 July 2018; Accepted: 29 August 2018; Published: 5 September 2018 Abstract: Several studies have shown the potential of using Ectomycorrhizal (ECM) fungi in conifer micropropagation to overcome the cessation of adventitious root development. In vitro inoculation promotes the re-growth of the root system induced previously by auxin treatments, facilitating acclimation and diminishing the losses of plants because of a weak root system that is incapable of water and nutrient absorption. During a series of mycorrhization experiments, cryostat and ultrafine cuts were used to study the morpho-histological transformation of the symbiotic roots. To obtain cryostat cuts from pine roots a method frequently used for animal tissue was adopted. Molecular methods allowed fungi identification in all the mycorrhization phases and in the acclimation of derived plants. Mycorrhizal-like-structures derived from in vitro culture and axenic liquid cultures of roots were microscopically analyzed and compare with mycorrhizal roots. Keywords: ectomycorrhiza; mycorrhiza-like structures; stone pine; adventitious roots; Hartig net 1. Introduction Ectomycorrhizal fungi (ECM fungi) are phylogenetically very diverse and more than 2000 species of ECM fungi worldwide have been identified, primarily from Basidiomycota and Ascomycota. -
Mantar Dergisi
11 6845 - Volume: 20 Issue:1 JOURNAL - E ISSN:2147 - April 20 e TURKEY - KONYA - FUNGUS Research Center JOURNAL OF OF JOURNAL Selçuk Selçuk University Mushroom Application and Selçuk Üniversitesi Mantarcılık Uygulama ve Araştırma Merkezi KONYA-TÜRKİYE MANTAR DERGİSİ E-DERGİ/ e-ISSN:2147-6845 Nisan 2020 Cilt:11 Sayı:1 e-ISSN 2147-6845 Nisan 2020 / Cilt:11/ Sayı:1 April 2020 / Volume:11 / Issue:1 SELÇUK ÜNİVERSİTESİ MANTARCILIK UYGULAMA VE ARAŞTIRMA MERKEZİ MÜDÜRLÜĞÜ ADINA SAHİBİ PROF.DR. GIYASETTİN KAŞIK YAZI İŞLERİ MÜDÜRÜ DR. ÖĞR. ÜYESİ SİNAN ALKAN Haberleşme/Correspondence S.Ü. Mantarcılık Uygulama ve Araştırma Merkezi Müdürlüğü Alaaddin Keykubat Yerleşkesi, Fen Fakültesi B Blok, Zemin Kat-42079/Selçuklu-KONYA Tel:(+90)0 332 2233998/ Fax: (+90)0 332 241 24 99 Web: http://mantarcilik.selcuk.edu.tr http://dergipark.gov.tr/mantar E-Posta:[email protected] Yayın Tarihi/Publication Date 27/04/2020 i e-ISSN 2147-6845 Nisan 2020 / Cilt:11/ Sayı:1 / / April 2020 Volume:11 Issue:1 EDİTÖRLER KURULU / EDITORIAL BOARD Prof.Dr. Abdullah KAYA (Karamanoğlu Mehmetbey Üniv.-Karaman) Prof.Dr. Abdulnasır YILDIZ (Dicle Üniv.-Diyarbakır) Prof.Dr. Abdurrahman Usame TAMER (Celal Bayar Üniv.-Manisa) Prof.Dr. Ahmet ASAN (Trakya Üniv.-Edirne) Prof.Dr. Ali ARSLAN (Yüzüncü Yıl Üniv.-Van) Prof.Dr. Aysun PEKŞEN (19 Mayıs Üniv.-Samsun) Prof.Dr. A.Dilek AZAZ (Balıkesir Üniv.-Balıkesir) Prof.Dr. Ayşen ÖZDEMİR TÜRK (Anadolu Üniv.- Eskişehir) Prof.Dr. Beyza ENER (Uludağ Üniv.Bursa) Prof.Dr. Cvetomir M. DENCHEV (Bulgarian Academy of Sciences, Bulgaristan) Prof.Dr. Celaleddin ÖZTÜRK (Selçuk Üniv.-Konya) Prof.Dr. Ertuğrul SESLİ (Trabzon Üniv.-Trabzon) Prof.Dr. -
AR TICLE Diversity of Chroogomphus (Gomphidiaceae, Boletales) In
doi:10.5598/imafungus.2018.09.02.04 IMA FUNGUS · Diversity of ( , ) in Europe, and Chroogomphus Gomphidiaceae Boletales ARTICLE [C. rutilus Ross Scambler1,6, Tuula Niskanen1, Boris Assyov2, A. Martyn Ainsworth1, Jean-Michel Bellanger3, Michael Loizides4 , Pierre- Arthur Moreau5, Paul M. Kirk1, and Kare Liimatainen1 1Jodrell Laboratory, Royal Botanic Gardens, Kew, Surrey TW9 3AB, UK; corresponding author e-mail: [email protected] 2!"#$"%'*+'///<'" 3UMR5175, CNRS, Université de Montpellier, Université Paul-Valéry Montpellier, EPHE, INSERM, 1919, route de Mende, F-34293 Montpellier Cedex 5, France 4P.O. box 58499, 3734 Limassol, Cyprus 5Université de Lille, Fac. Pharma. Lille, EA 4483 IMPECS, F – 59000 Lille, France 6 Present address :Department of Applied Sciences, University of the West of England, Frenchay Campus, Coldharbour Lane, Bristol, BS16 1QY, UK In this study, eight species of Chroogomphus are recognized from Europe: C. britannicus, C. aff. [ 1, C. fulmineus, C. cf. helveticus, C. mediterraneus, C. cf. purpurascens, C. rutilus, and C. subfulmineus. DNA barcode Different candidates for the application of the name C. rutilus[ ITS =>Chroogomphus fulmineus and C. mediterraneus are molecular systematics [C. subfulmineus?[ new taxa a new subgenus Siccigomphus and three new sections, Confusi, Filiformes, and Fulminei are introduced. The taxonomy former sections Chroogomphus and Floccigomphus are elevated to subgeneric level. Comparison of the ITS X[%!?'/\]'!?'*[ of 1.5 %, with the exception of the two species belonging to sect. Fulminei which differ by a minimum of 0.9 %. Ecological specimen data indicate that species of Chroogomphus form basidiomes under members of Pinaceae, with a general preference for species of Pinus. Five European species have been recorded under Picea, while Abies and Larix have also been recorded as tree associates, although the detailed nutritional relationships of the Submitted: 27 November 2017; Accepted: 27 August 2018; Published: 5 September 2018. -
Bulk Isolation of Basidiospores from Wild Mushrooms by Electrostatic Attraction with Low Risk of Microbial Contaminations Kiran Lakkireddy1,2 and Ursula Kües1,2*
Lakkireddy and Kües AMB Expr (2017) 7:28 DOI 10.1186/s13568-017-0326-0 ORIGINAL ARTICLE Open Access Bulk isolation of basidiospores from wild mushrooms by electrostatic attraction with low risk of microbial contaminations Kiran Lakkireddy1,2 and Ursula Kües1,2* Abstract The basidiospores of most Agaricomycetes are ballistospores. They are propelled off from their basidia at maturity when Buller’s drop develops at high humidity at the hilar spore appendix and fuses with a liquid film formed on the adaxial side of the spore. Spores are catapulted into the free air space between hymenia and fall then out of the mushroom’s cap by gravity. Here we show for 66 different species that ballistospores from mushrooms can be attracted against gravity to electrostatic charged plastic surfaces. Charges on basidiospores can influence this effect. We used this feature to selectively collect basidiospores in sterile plastic Petri-dish lids from mushrooms which were positioned upside-down onto wet paper tissues for spore release into the air. Bulks of 104 to >107 spores were obtained overnight in the plastic lids above the reversed fruiting bodies, between 104 and 106 spores already after 2–4 h incubation. In plating tests on agar medium, we rarely observed in the harvested spore solutions contamina- tions by other fungi (mostly none to up to in 10% of samples in different test series) and infrequently by bacteria (in between 0 and 22% of samples of test series) which could mostly be suppressed by bactericides. We thus show that it is possible to obtain clean basidiospore samples from wild mushrooms. -
Mycology Praha
f I VO LUM E 52 I / I [ 1— 1 DECEMBER 1999 M y c o l o g y l CZECH SCIENTIFIC SOCIETY FOR MYCOLOGY PRAHA J\AYCn nI .O §r%u v J -< M ^/\YC/-\ ISSN 0009-°476 n | .O r%o v J -< Vol. 52, No. 1, December 1999 CZECH MYCOLOGY ! formerly Česká mykologie published quarterly by the Czech Scientific Society for Mycology EDITORIAL BOARD Editor-in-Cliief ; ZDENĚK POUZAR (Praha) ; Managing editor JAROSLAV KLÁN (Praha) j VLADIMÍR ANTONÍN (Brno) JIŘÍ KUNERT (Olomouc) ! OLGA FASSATIOVÁ (Praha) LUDMILA MARVANOVÁ (Brno) | ROSTISLAV FELLNER (Praha) PETR PIKÁLEK (Praha) ; ALEŠ LEBEDA (Olomouc) MIRKO SVRČEK (Praha) i Czech Mycology is an international scientific journal publishing papers in all aspects of 1 mycology. Publication in the journal is open to members of the Czech Scientific Society i for Mycology and non-members. | Contributions to: Czech Mycology, National Museum, Department of Mycology, Václavské 1 nám. 68, 115 79 Praha 1, Czech Republic. Phone: 02/24497259 or 96151284 j SUBSCRIPTION. Annual subscription is Kč 350,- (including postage). The annual sub scription for abroad is US $86,- or DM 136,- (including postage). The annual member ship fee of the Czech Scientific Society for Mycology (Kč 270,- or US $60,- for foreigners) includes the journal without any other additional payment. For subscriptions, address changes, payment and further information please contact The Czech Scientific Society for ! Mycology, P.O.Box 106, 11121 Praha 1, Czech Republic. This journal is indexed or abstracted in: i Biological Abstracts, Abstracts of Mycology, Chemical Abstracts, Excerpta Medica, Bib liography of Systematic Mycology, Index of Fungi, Review of Plant Pathology, Veterinary Bulletin, CAB Abstracts, Rewicw of Medical and Veterinary Mycology. -
Moeszia9-10.Pdf
Tartalom Tanulmányok • Original papers .............................................................................................. 3 Contents Pál-Fám Ferenc, Benedek Lajos: Kucsmagombák és papsapkagombák Székelyföldön. Előfordulás, fajleírások, makroszkópikus határozókulcs, élőhelyi jellemzés .................................... 3 Ferenc Pál-Fám, Lajos Benedek: Morels and Elfin Saddles in Székelyland, Transylvania. Occurrence, Species Description, Macroscopic Key, Habitat Characterisation ........................... 13 Pál-Fám Ferenc, Benedek Lajos: A Kárpát-medence kucsmagombái és papsapkagombái képekben .................................................................................................................................... 18 Ferenc Pál-Fám, Lajos Benedek: Pictures of Morels and Elfin Saddles from the Carpathian Basin ....................................................................................................................... 18 Szász Balázs: Újabb adatok Olthévíz és környéke nagygombáinak ismeretéhez .......................... 28 Balázs Szász: New Data on Macrofungi of Hoghiz Region (Transylvania, Romania) ................. 42 Pál-Fám Ferenc, Szász Balázs, Szilvásy Edit, Benedek Lajos: Adatok a Baróti- és Bodoki-hegység nagygombáinak ismeretéhez ............................................................................ 44 Ferenc Pál-Fám, Balázs Szász, Edit Szilvásy, Lajos Benedek: Contribution to the Knowledge of Macrofungi of Baróti- and Bodoki Mts., Székelyland, Transylvania ..................... 53 Pál-Fám -
Melanophyllum Eyrei
© Demetrio Merino Alcántara [email protected] Condiciones de uso Melanophyllum eyrei (Massee) Singer, Lilloa 22: 436 (1951) [1949] 10 mm Agaricaceae, Agaricales, Agaricomycetidae, Agaricomycetes, Agaricomycotina, Basidiomycota, Fungi Sinónimos homotípicos: Schulzeria eyrei Massee, Grevillea 22(no. 102): 38 (1893) Chlorospora eyrei (Massee) Massee, Trans. Br. mycol. Soc. 2(5): 170 (1907) [1906] Glaucospora eyrei (Massee) Rea, Brit. basidiomyc. (Cambridge): 62 (1922) Lepiota eyrei (Massee) J.E. Lange, Fl. Agaric. Danic. 1: 27 (1935) Cystoderma eyrei (Massee) Singer, Annls mycol. 41(1/3): 170 (1943) Material estudiado: Francia, Aquitania, Pirineos Atlánticos, Osse en Aspe, L'Aidy, 30TXN8762, 1.122 m, en suelo bajo Fagus sylvatica, 1-X-2012, leg. Dianora Estrada y Demetrio Merino, JA-CUSSTA: 9411. Descripción macroscópica: Píleo de 17-29 mm de diám., cónico convexo a plano con la edad, con umbón obtuso, margen agudo, festoneado. Cutícula pruino- sa, mate, de color crema rojizo, con el disco ocráceo. Láminas libres, separadas, de color crema que se va manchando de azul turquesa a medida que van madurando las esporas, arista entera, concolora. Estípite de 36-44 x 2-4 mm, cilíndrico, curvado, con la base bulbillosa provista de cordones miceliares crema blanquecinos, sin anillo, con abundante pruina de color crema sobre fondo marrón ocráceo. Olor agradable. Descripción microscópica: Basidios claviformes a fusiformes, tetraspóricos, con fíbula basal, de (15,1-)15,9-19,7(-22,0) × (5,2-)5,6-7,4(-7,9) µm; N = 40; Me = 17,8 × 6,5 µm. Basidiosporas elipsoidales a subcilíndricas, lisas, de color verdoso, apiculadas, gutuladas, de (3,7-)4,4-5,2(-5,6) × (2,4-)2,8-3,3(-3,5) µm; Q = (1,3-)1,4-1,7(-1,9); N = 129; V = (13-)17-28(-36) µm3; Me = 4,8 × 3,0 µm; Qe = 1,6; Ve = 23 µm3. -
Looking for Lepiota Psalion Huijser & Vellinga (Agaricales, Agaricaceae)
A peer-reviewed open-access journal MycoKeys 52: 45–69 (2019) Looking for Lepiota psalion Huijser & Vellinga 45 doi: 10.3897/mycokeys.52.34021 RESEARCH ARTICLE MycoKeys http://mycokeys.pensoft.net Launched to accelerate biodiversity research Looking for Lepiota psalion Huijser & Vellinga (Agaricales, Agaricaceae) Alfredo Vizzini1,2, Alessia Tatti3, Henk A. Huijser4, Jun F. Liang5, Enrico Ercole1 1 Department of Life Sciences and Systems Biology, University of Torino, Viale P.A. Mattioli 25, I-10125, Torino, Italy 2 Institute for Sustainable Plant Protection (IPSP)-CNR, Viale P.A. Mattioli 25, I-10125, Torino, Italy 3 Department of Environmental and Life Science, Section Botany, University of Cagliari, Viale S. Ignazio 1, I-09123, Cagliari, Italy 4 Frederikstraat 6, 5671 XH Nuenen, The Netherlands5 Research Institute of Tropical Forestry, Chinese Academy of Forestry, Guangzhou, 510520, China Corresponding author: Alfredo Vizzini ([email protected]) Academic editor: T. Lumbsch | Received 21 February 2019 | Accepted 11 April 2019 | Published 9 May 2019 Citation: Vizzini A, Tatti A, Huijser HA, Liang JF, Ercole E (2019) Looking for Lepiota psalion Huijser & Vellinga (Agaricales, Agaricaceae). MycoKeys 52: 45–69. https://doi.org/10.3897/mycokeys.52.34021 Abstract Lepiota psalion is fully described based on a recent collection from Sardinia (Italy) and the holotype. NrITS- and nrLSU-based phylogeny demonstrates that sequences deposited in GenBank as “L. psalion” and generated from two Dutch and one Chinese collections are not conspecific with the holotype and represent two distinct, undescribed species. These species are here proposed asLepiota recondita sp. nov. and Lepiota sinorecondita ad int. Keywords Agaricomycetes, Basidiomycota, cryptic species, hymeniform pileus covering, taxonomy Introduction Recent molecular analyses have indicated that the genus Lepiota (Pers.) Gray is a para- phyletic assemblage that is monophyletic only if it is considered together with spe- cies of Cystolepiota Singer, Echinoderma (Locq. -
Boletín Micológico De FAMCAL Una Contribución De FAMCAL a La Difusión De Los Conocimientos Micológicos En Castilla Y León Una Contribución De FAMCAL
Año Año 2011 2011 Nº6 Nº 6 Boletín Micológico de FAMCAL Una contribución de FAMCAL a la difusión de los conocimientos micológicos en Castilla y León Una contribución de FAMCAL Con la colaboración de Boletín Micológico de FAMCAL. Boletín Micológico de FAMCAL. Una contribución de FAMCAL a la difusión de los conocimientos micológicos en Castilla y León PORTADA INTERIOR Boletín Micológico de FAMCAL Una contribución de FAMCAL a la difusión de los conocimientos micológicos en Castilla y León COORDINADOR DEL BOLETÍN Luis Alberto Parra Sánchez COMITÉ EDITORIAL Rafael Aramendi Sánchez Agustín Caballero Moreno Rafael López Revuelta Jesús Martínez de la Hera Luis Alberto Parra Sánchez Juan Manuel Velasco Santos COMITÉ CIENTÍFICO ASESOR Luis Alberto Parra Sánchez Juan Manuel Velasco Santos Reservados todos los derechos. No está permitida la reproducción total o parcial de este libro, ni su tratamiento informático, ni la transmisión de ninguna forma o por cualquier medio, ya sea electrónico, mecánico, por fotocopia, por registro u otros métodos, sin el permiso previo y por escrito del titular del copyright. La Federación de Asociaciones Micológicas de Castilla y León no se responsabiliza de las opiniones expresadas en los artículos firmados. © Federación de Asociaciones Micológicas de Castilla y León (FAMCAL) Edita: Federación de Asociaciones Micológicas de Castilla y León (FAMCAL) http://www.famcal.es Colabora: Junta de Castilla y León. Consejería de Medio Ambiente Producción Editorial: NC Comunicación. Avda. Padre Isla, 70, 1ºB. 24002 León Tel. 902 910 002 E-mail: [email protected] http://www.nuevacomunicacion.com D.L.: Le-1011-06 ISSN: 1886-5984 Índice Índice Presentación ....................................................................................................................................................................................11 Favolaschia calocera, una especie de origen tropical recolectada en el País Vasco, por ARRILLAGA, P. -
Contribución Al Conocimiento Del Género Boletus L., Sensu Lato, En La Provincia De Albacete
Número Páginas Año 13 11-36 2019 CONTRIBUCIÓN AL CONOCIMIENTO DEL GÉNERO BOLETUS L., SENSU LATO, EN LA PROVINCIA DE ALBACETE Por César A. RODRÍGUEZ RODRÍGUEZ 1 José FAJARDO RODRÍGUEZ 1, 2, 3, 4, Domingo BLANCO SIDERA 1, 4, * Alonso VERDE LÓPEZ 1, 2, 4 * [email protected] Recibido: 22 de septiembre de 2018 Aprobado: 19 de abril de 2019 1 Sociedad Micológica de Albacete 2 Grupo de Investigación en Botánica, Etnobiología y Educación. Laboratorio de Sistemática y Etnobotánica. Instituto Botánico (UCLM). Jardín Botánico de Casti- lla-La Mancha. Campus de Albacete. 3 Universidad Popular de Albacete 4 Instituto de Estudios Albacetenses —11— CÉSAR A. RODRÍGUEZ RODRÍGUEZ • JOSÉ FAJARDO RODRÍGUEZ • DOMINGO BLANCO SIDERA • ALONSO VERDE LÓPEZ RESUMEN Con este trabajo tratamos de clarificar la segregación que se ha he- cho actualmente, atendiendo a nuevos criterios genéticos y moleculares, del género Boletus L. sensu lato, derivando en una nueva clasificación y nombres aceptados desde el punto de vista de la taxonomía micológica. Se actualiza el catálogo de especies presentes en la provincia de Albacete, indicando las citas respaldadas por muestras de herbario, de- positadas en el herbario ALBA-SMA, así como la presencia probable de ciertas especies, debido a sus requerimientos ecológicos. Se aportan tres nuevas citas de especies para la micobiota albace- tense. Palabras clave: Boletus, micobiota, Albacete. ABSTRACT With this work we try to clarify the segregation that has been done, taking into account new genetic and molecular criteria, of the genus Bo- letus L. sensu lato, resulting in a new classification and names accepted from the point of view of mycological taxonomy. -
Mushroom Diversity in Shaki District of Azerbaijan
Plant & Fungal Research (2019) 2(1): 23-31 © The Institute of Botany, ANAS, Baku, AZ1004, Azerbaijan http://dx.doi.org/10.29228/plantfungalres.14 June 2019 Mushroom diversity in Shaki district of Azerbaijan Elgun H. Mustafabayli INTRODUCTION Dilzara N. Aghayeva1 Information of the first mushroom records (Polyporus Institute of Botany, Azerbaijan National Academy of Sciences, officinalis (Vill.) Fr., Tuber album Bull., Tuber Badamdar 40, Baku, AZ1004, Azerbaijan melanosporum Vittad., Morchella esculenta (L.) Pers.) Abstract: Mushroom diversity in Shaki district of in Azerbaijan was mentioned in the medieval manu- Azerbaijan was studied during 2013-2018. Totally 346 scripts [Alakbarli, 2006]. Based on available litera- specimens were collected and identified based on the ture the first fungal collections were made by foreign morphologically main diagnostic features. Phenotypic botanists and mycologists who visited Azerbaijan characters, as well as microscopic features (size, shape, [Georgi, 1800; Kolenati 1858; Voronov 1922-1923]. color of spores) were considered. In total 111 taxa (106 But the recorded and reported information on mush- species, three forms and two variations) belonging to rooms is scanty, only few published works about fungi 67 genera, 32 families and 11 orders of Ascomycota reached us [Aghayeva, 2018]. Significant contribution and Basidiomycta were identified in view of the towards studying mushrooms belongs to A. Sadiqov, latest taxonomic and nomenclature changes. Two taxa who launched first investigations on diversity since 60s Morchella esculenta var. rotunda and Suillus luteus f. of the last century. In the result of this effort a wide albus represent new records for Azerbaijan. Ecological range of mushrooms were investigated in the country groups of mushrooms were reflected. -
Le Specie Più Comuni Del Genere Suillus
MicoPonte n. 3 - 2009: pp. 11-17 Le specie più comuni del genere Suillus SERGIO MATTEUCCI via Per Gignano 151, 55050 Vinchiana (LU) [email protected] INTRODUZ I ONE Quando l’autunno veste i boschi d’arlecchino ed il calore estivo lo ritrovi nel canto del camino, nei nostri boschi di conifere o misti spuntano i primi Suillus comunemente chiamati pinacci. Il genere Suillus appartiene alla famiglia delle Boletaceae e comprende circa 30 specie in Europa. Sono specie terricole ritenute quasi tutte simbionti, con cuticola vischiosa e facilmente asportabile (con eccezione per il S. variegatus e per il S. bovinus), con struttura omogenea, cioè con gambo che non si separa in modo netto dal cappello come ad esempio avviene nel genere Amanita; i tubuli sono separabili dalla carne del cappello (con eccezione per il S. bovinus), mentre i pori negli esemplari adulti sono irregolari e angolosi e secernano, in alcune specie, goccioline lattiginose che seccando lasciano granuli rossastri sul gambo. Suillus bellinii (Inzenga) Kuntze Etimologia: da nome proprio, in onore di Vincenzo Bellini (1801-1835), compositore italiano. Sinonimi: Boletus bellinii Inzenga, Ixocomus bellinii (Inzenga) Gilb., Rostkovites bellinii (Inzenga) Reichert Nomi volgari: pinarolo, pinacchiotto, polpetta Principali caratteri macroscopici Specie di aspetto tozzo, con cappello convesso-spianato con orlo involuto che può raggiungere i 15 cm di diametro; la cuticola è totalmente asportabile, liscia e molto vischiosa, maculata, bianco-grigio o bianco-marrone a volte anche con tonalità giallastre. Tubuli corti, adnati, da bianchi a gialli, giallo-verde. Gambo sempre più corto del diametro del cappello, cilindrico, attenuato alla base, privo di anello, ornato da granulazioni scure su tutta la superficie, alla fine rossastre verso l’alto.