A List of the Herpetological Type Specimens in the Zoologisches Forschungsmuseum Alexander Koenig, Bonn
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Bonn zoological Bulletin Volume 59 pp. 79–108 Bonn, December 2010 A list of the herpetological type specimens in the Zoologisches Forschungsmuseum Alexander Koenig, Bonn Wolfgang Böhme Zoologisches Forschungsmuseum Alexander Koenig, Herpetology Section, Adenauerallee 160, D-53113 Bonn, Germany; E-mail: [email protected]. Abstract. In the herpetological collection of ZFMK 528 scientific species group names are represented by type materi- al. Of these, 304 names are documented by primary type specimens (onomatophores) while for 224 further names sec- ondary type specimens (typoids) are available, ranging chronologically from 1801 to 2010. The list is a shortened pred- ecessor of a comprehensive type catalogue in progress. It lists name bearing types with their catalogue numbers includ- ing information on further type series members also in other institutions, while secondary types are listed only by pres- ence, both in ZFMK and other collections including holotype repositories. Geographic origin and currently valid names are also provided. Key words. Amphibians and reptiles, type list, ZFMK Bonn. INTRODUCTION A first ZFMK herpetological type catalogue was published (currently section) in 1951, for many decades. Nonethe- (Böhme 1974) three years after I had entered Museum less, the present list does comprise some historical “pre- Koenig as a herpetological curator. It contained only 34 ZFMK” material which has been obtained after 1971 from reptilian names documented by type material, 22 of which smaller university museums, first of all from the Zoolog- were name-bearing type specimens (onomatophores), and ical Museum of the University of Göttingen (1977). Sin- 12 further names were documented by paratypes only. gle taxa documented by type specimens were also received Amphibians were not represented by type material at that by the Zoological Museum of the University of Kiel time. A second type catalogue was published ten years lat- (1973), and recently by the Biogeography Department of er, on the occasion of the 50th anniversary of Museum the University of Saarbrücken (via University of Trier, in Koenig as a public zoological museum (Böhme & 2009). Bischoff 1984). It counted 147 names of amphibians (51) and reptiles (96) of which 99 were onomatophores while The following catalogue comprises, with one exception 48 additional names were documented by secondary type (the chameleon Kinyongia vanheygeni), only the extant material (paratypes, paralectotypes). In addition, 22 types while those types which went lost in Göttingen be- names representing primary and secondary type specimens fore the collection transfer to Bonn, and which have been of historical material which had belonged before to the Zo- already mentioned in Böhme & Bischoff (1984), will be ological Museum in Göttingen (transferred to ZFMK in carefully listed and commented on in the forthcoming 1977) were listed as missing types because they were al- monograph mentioned above. ready lost before they ever reached ZFMK. The delimitation and arrangement of families follows Frost Within the present special BzB issue on type lists of the et al. (2006) for the amphibians, and Böhme & Sander vertebrate and invertebrate collections of ZFMK, the pres- (2010) for the reptiles, plus the familial concept of Zaher ent herpetological list is a much shortened predecessor of et al. (2009) for colubroid snakes. For dendrobatid frogs, a comprehensive, critically evaluated type catalogue of I follow the monograph by Grant et al. (2006, see also Löt- ZFMK amphibians and reptiles which will be published ters et al. 2007); for Madagascar, I follow Glaw & Vences in the framework of a monograph dealing with the histo- (2007). Within their families, taxa are arranged in alpha- ry of herpetological research in Bonn since the founda- betical order of genera, and within genera, arrangement tion of the Rheinische Friedrich-Wilhelms-Universität in is again alphabetically. Names of authors are accompanied 1818 which predates the foundation of ZFMK, particular- by an initial letter of their forenames only when a fami- ly the creation of its own herpetological department ly name occurs more than twice. Bonn zoological Bulletin 59: 79–108 ©ZFMK 80 Wolfgang Böhme Catalogue numbers are given here for primary types on- GNHM = Göteborgs Naturhistoriska Museet (see also ly, i.e. for holotypes, lectotypes and neotypes; syntypes NHMG) are also counted under this category because they are the HLMD = Hessisches Landesmuseum, Darmstadt first candidates for lectotype selection and are thus like- IEBR = Institute of Ecology and Biological Resources, ly to contain future lectotypes; for secondary types, i.e. Hanoi paratypes and paralectotypes, only their presence in IFAN = Institut Fondamental d’Afrique Noire, Dakar ZFMK and/or other collection is reported: due to the ne- INHS = Illinois Natural History Survey [descended in part cessity to avoid redundancies with the future big cata- from Illinois State Laboratory of Natural History], Uni- logue. All names are given in their original combinations; versity of Illinois, Champaign, Illinois if taxonomic assignment or rank has changed, the current INPA = Instituto Nacional de Pesquisas da Amazônia, Ma- name believed to be valid today is given. This shortened naus preliminary type list thus offers the following informa- IRSNB = Institut Royal des Sciences Naturelles de Bel- tions: gique, Bruxelles / (= KBIN) Koninklijk Belgisch Instituut voor Natuurwetenschappen, Brussel – All names which are represented at ZFMK by type ma- IZUG = Institute of Zoology, University of Genova terial, in their original taxonomic combinations. LACM = Natural History Museum of Los Angeles – Primary, name-bearing types and their catalogue num- County, Los Angeles bers; also the repositories of additional secondary type LBUM = Laboratoire de Biogéographie, Université de material, if at all existent. Montpellier – If only secondary types are involved at ZFMK, the col- LSUZM = Louisiana State University Zoological Museum lection housing the respective onomatophore is given, MCC = Museo Civico di Storia Naturale, Carmagnola together with all collections holding additional second- MCNSC = Museo Insular de Ciencias Naturales de San- ary type material. ta Cruz de Tenerife – The rough geographic origin of the respective taxa. MCZ = Museum of Comparative Zoology, Cambridge, – The current name combination, if the original combi- Massachusetts nation has changed. MGL = Musée Guimet d’Histoire Naturelle, Lyon MHNCP = Museo de Historia Natural, Universidad Na- For more details including full bibliographic references cional de San Antonio Abad del Cusco, Cusco of each name, for full locality and collector’s data, as well MHNG = Muséum d’Histoire Naturelle, Genève as for taxonomic and/or historical comments the reader MHNSM = Museo de Historia Natural San Marcos, Li- is referred to the comprehensive monograph in progress. ma MM = Museum Magdeburg MNCN = Museo Nacional de Ciencias Naturales, Madrid Abbreviations for museum collections used: MNHN = Muséum National d’Histoire Naturelle, Paris MPEG = Museu Paraense “Emilio Goeldi”, Zoologia, Be- AMNH = American Museum of Natural History, New lém, Pará York MRAC = Musée Royal de l’Afrique Centrale, Tervuren ANSP = Academy of Natural Sciences, Philadelphia MRSN = Museo Regionale di Scienze Naturali, Torino BEV = Laboratoire de Biogéographie et Ecologie des Ver- MTKD (or MTD respectively) = Museum für Tierkunde tébrés, École Pratique des Hautes Études, Montpellier Dresden BIUB = Mongolian Academy of Sciences, Ulan Bataar MUSM = Museo de Historia Natural San Marcos BM(NH) = British Museum (of Natural History), The Nat- MZB = “Museum Zoologicum Bogoriense” = Zoological ural History Museum, London Museum, Bogor CAS = California Academy of Sciences, San Francisco MZUF = Museo Zoologico, Universitá di Firenze, Flo- CBF = Colección Boliviana de Fauna, La Paz rence CET = Centro de Estudios Tropicales, Sevilla, Spain MZUN = Museo di Zoologia dell’Università di Napoli, CM = Carnegie Museum, Pittsburgh Naples CNHM = Chicago Natural History Museum MZUSP = Museu de Zoologia da Universidade de São DZCHU = Department of Zoology, Charles University, Paulo, São Paulo (also as MZSP and MSP) Prague NHCY = Natural History Collection Yemen, Sana’a DZUL = Departamento de Zoología, Universidad de La NHMG = Naturhistoriska Riksmuseet, Göteborg (see al- Laguna, Tenerife so GNHM) EBD = Estación Biologica de Doñana, Sevilla NKA = Museo de Historia Natural “Noël Kempff Mer- EPN = Escuela Politécnica Nacional, Quito cado”, Santa Cruz de la Sierra FMNH = Field Museum of Natural History, Chicago NMB = Naturhistorisches Museum Basel Bonn zoological Bulletin 59: 79–108 ©ZFMK Type specimens in ZFMK: Section of Herpetology 81 NMBE = Naturhistorisches Museum Bern ences, Tbilisi NMK = National Museums of Kenya, Nairobi ZLU = Departamento de Zoología de la Universidad de NMNW = National Museum of Namibia, Windhoek La Laguna, Tenerife NMP = National Museum Prague ZMA = Zoölogisch Museum, Universiteit van Amsterdam NMSR = National Museum of South Rhodesia (= now ZMB = Zoologisches Museum = Museum für Naturkun- Zimbabwe), Harare de, Berlin NMW = Naturhistorisches Museum Wien, Vienna ZMH = Zoologisches Museum der Universität Hamburg NUOL = National University of Laos, Vientiane ZMMU = Zoological Museum, Moscow State Universi- OMNH = Osaka Museum of Natural History, Laborato- ty ry of Zoology, Osaka ZMUC = Zoological Museum, University of Copenhagen PEM = Port Elizabeth Museum ZRC = Zoological