An update of the revision of Cyathea subgen. Alsophila sect. Gymnosphaera (Cyatheaceae) in Madagascar and the Comoros including a discussion of putative hybridization events Thomas JANSSEN Muséum national d’Histoire naturelle, Département Systématique et Évolution, UMR-CNRS 5202, USM 0602, case postale 39, 57 rue Cuvier, F-75231 Paris cedex 05 (France) and Georg-August-Universität Göttingen, Albrecht-von-Haller-Institut für Pflanzenwissenschaften, Abt. Spezielle Botanik, Untere Karspüle 2, D-37073 Göttingen (Germany)
[email protected] France RAKOTONDRAINIBE École pratique des hautes études and Muséum national d’Histoire naturelle, Département Systématique et Évolution, UMR-CNRS 5202, USM 0602, case postale 39, 57 rue Cuvier, F-75231 Paris cedex 05 (France)
[email protected] Janssen T. & Rakotondrainibe F. 2007. — An update of the revision of Cyathea subgen. Alsophila sect. Gymnosphaera (Cyatheaceae) in Madagascar and the Comoros including a discussion of putative hybridization events. Adansonia, sér. 3, 29 (2) : 195-213. ABSTRACT A new revision of Cyathea sect. Gymnosphaera in Madagascar and the Comoros is proposed. Two new species, Cyathea impolita Rakotondr. & Janssen and C. rouhaniana Rakotondr. & Janssen, as well as a new variety, C. impolita var. michelii Rakotondr. & Janssen are described. As opposed to all other members of the section in the region, C. rouhaniana does not possess bullate scales nor a dense indument on the adaxial face of the costae and costulae; the fertile basal pinnae of each leaf are more rapidly caducous than the sterile pinnae and sori include abundant paraphyses longer than the sporangia. C. impolita and its va- riety michelii have dull, medium brown scales at their petiole base, isomorphic fertile and sterile pinnae and sori usually distributed over the entire abaxial surface in fertile leaves.