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Land Between the Lakes Golden Pond, Kentucky HELD AT BRANDONSPRJNG GROUPCAMP LANDBETWEEN THE Lms MARCH 6, 1993 Sponsored by: The Center for Field Biology Austin Peay State University, Clarksville, Tennessee and Mumy State University Center for Reservoir Research and Tennessee Valley Authority - Land Between The Lakes Golden Pond, Kentucky EDITED BY: Steven W. Hamilton, Edward W. Chester, and A. Floyd Scott The Center for Field Biology Austin Peay State University Published by and available from: The Center for Field Biology, Austin Peay State University, Clarksville, Tennessee 37044 Price: $5.00 . SUGGESTED CITATION Hamilton, S.W., E.W. Chester and A.F. Scott. 1993. Proceedings of the fifth annual symposium on the natural history of lower Tennessee and Cumberland River Valleys. Center for Field Biology, Austin Peay State University, Clarksville, Tennessee. Published July 1993 PREFACE On March 5' and 6", 1993 over 120 students of regional natural history and field biology gathered at Brandon Spring Group Camp in TVA's Land Between The Lakes to participate in the Fifth Symposium on the Natural History of Lower Tennessee and Cumberland River Valleys. Sponsors of this symposium were The Center for Field Biology at Austin Peay State University, the Center for Reservoir Research at Murray State University, and Land Between The Lakes. On Friday afternoon the symposium attendees were welcomed by Dr. Ben Stone, Director of The Center for Field Biology at APSU and Dr. Gary Boggess, Dean of the College of Sciences at Murray State University. Representing Land Between The Lakes was Mr. John Mechler, manager of Land Management. In his welcoming presentation Mr. Mechler described LBL's three-pronged environmental mission of resource management, research, and education. Stream ecology was the theme of Friday afternoon's invited papers presented by Dr. Art Benke of the University of Alabama and Dr. Ike Schlosser of the University of North Dakota. Dr. Benke presented information about what is probably the most poorly studied lotic environment, large rivers. Dr. Benke's studies of coastal plain blackwater rivers in southeastern Georgia have provided critical insights into the function of large rivers and the importance of floodplain inundation. Dr. Schlosser, applying landscape ecology perspectives to stream fish ecology, discussed the importance of large scale habitat heterogeneity, the effects of ecotonal barriers, and the interactions of adjacent habitats on stream fish distributions and life histories. The full text of Dr. Benke's and Dr. Schlosser's papers are presented in these proceedings. Following the evening meal, Dr. Ray Norris, coordiiator of Tennessee Save Our Streams, Izaak Walton League of America, gave an informative and enjoyable slide presentation on the Adopt-A-River program in Tennessee. This program educates the public about water quality problems, sponsors river "clean-up" projects, trains volunteer groups to monitor water quality via sampling of aquatic macroinvertebrates, and even initiates litigation against water quality violators. An informal social followed the presentation by Dr. Norris. The Saturday morning program was filled with two concurrent sessions, each comprised 15 contributed papers. Session I, moderated by Dr. Cindy Taylor of Austin Peay State University, consisted of papers on general zoology and aquatic biology. Dr. William Ellis, Dean of the College of Graduate and Professional Programs and Professor of Biology at Austin Peay, served as moderator for Session 11, which included papers on botany and microbiology. The contributors were invited to publish an abstract or a complete paper in these proceedings. The 15 contributors to Session I opted to publish abstracts only. Five of the contributors to Session 11 chose to publish complete papers, while the remaining ten elected to publish only abstracts. The style and format of these proceedings follow, for the most part, the style established by earlier proceedings. Dr. Floyd Scott served as the organizer and editor for Contributed Papers Session I and Dr. Wayne Chester was the organizer and editor for Session II. I organized and edited the Invited Papers session and acted as the managing editor for these proceedings. Steve Hamilton July 1993 ACKNOWLEDGMENTS The editors would like to thank Ms. Cathy Hoback and Ms. Dorothy Reed for assistance in organizing and coordinating the activities before and during the symposium. We also must recognize the participation of the many APSU undergraduate and graduate research assistants for their help during this event. Manuscripts were typed and corrected by Ms. Reed and Ms. Marilyn Griffy, and Ms. Griffy helped with the final format. Their assistance is greatly appreciated. All complete papers were reviewed by at least one anonymous reviewer. We acknowledge all reviewers for their contribution to the quality of these papers. While we hope these proceedings are error fie, experience suggests that this is not the case. SYMPOSIUM REGISTRANTS Following, in alphabetical order, is a list of those individuals who registered at the 1993 symposium. Institutional affiliation (when available), city (of the person's institution or home), and state are also given. Melissa Allen, Austin Peay State University, Clarksville, Tennessee; Donna Barnes, Austin Peay State University, Clarksville, Tennessee; Joseph Barnes, Vanderbilt University, Nashville, Tennessee; Randall Barnes, Austin Peay State University, Clarksville, Tennessee; Carol Baskin, University of Kentucky, Lexington, Kentucky; Jerry Baskin, University of Kentucky, Lexington, Kentucky; Arthur Benke, University of Alabama, Tuscaloosa, Alabama; Alison Bennett, Austin Peay State University, Clarksville, Tennessee; James Bittner, Union University, Jackson, Tennessee; Scott Blane, Austin Peay State University, Clarksville, Tennessee; Steve Bloemer, Tennessee Valley AuthorityLand Between the Lakes, Golden Pond, Kentucky; Gary Boggess, Murray State University, Murray, Kentucky; Bob Bosserman, University of Louisville, Louisville, Kentucky; Leana Brown, Austin Peay State University, Clarksville, Tennessee; Kevin Buchanan, Austin Peay State University, Clarksville, Tennessee; Toni Buchanan, Jackson State Community College, Jackson, Tennessee; C. D. Buffington, Tennessee Valley AuthorityILand Between the Lakes, Golden Pond, Kentucky; Fkank Bulow, Tennessee Tech University, Cookeville, Tennessee; Gerald Burnett, Tennessee Valley AuthorityILand Between the Lakes, Golden Pond, Kentucky; Willodean Burton, Austin Peay State University, Clarksville, Tennessee; DyAnn Campbell, Austin Peay State University, Clarksville, Tennessee; Trudy Carr, Tennessee Valley AuthorityLand Between the Lakes, Golden Pond, Kentucky; Edward W. Chester, Austin Peay State- University, Clarksville, Tennessee; Richard Clements, University of Kentucky, Lexington, Kentucky; Mrs. Richard Clements, University of Kentucky, Lexington, Kentucky; Don Dailey; Austin Peay State University, Clarksville, Tennessee; Toni Dalasandro; Murray State University, Murray, Kentucky; Hal DeSelm, University of Tennessee, Knoxville, Tennessee; Jonathan Dey, lllinois Wesleyan, Bloornington, Illinois; Darrin Dodge, Murray State University, Murray, Kentucky; Elgenor Douglas, Austin Peay State University, Clarksville, Tennessee; L. Dubonis-Gray; Murray StateUniversity, Murray, Kentucky; William Ellis, Austin Peay State University, Clarksville, Tennessee; Tim Evans, Tennessee Valley AuthorityILand Between the Lakes, Golden Pond, Kentucky; Susan Eyer, Illinois Wesleyan, Bloomington, Illinois; Mack Finley, Austin Peay State University, Clarksville, Tennessee; Kevin Etch, Austin Peay State University, Clarksville, Tennessee; Paul Florence, University of Louisville, Louisville, Kentucky; James Fralish, Southern Illinois University, Carbondale, Illinois; Scott Franklin, Southern Illinois University, Carbondale, Illinois; Marty Gamble, Austin Peay State University, Clarksville, Tennessee; Deborah LingleGillis, Tennessee Department of Environment and Conservation, Nashville, Tennessee; Hannah Gillis, Clarksville, Tennessee; Patricia Gladu, Lindsey Wilson, Columbia, Kentucky; Don Green, Tennessee Department of Environment and Conservation, Nashville, Tennessee; Patricia Grimes, Austin Peay State University, Clarksville, Tennessee; Alma Hamilton, Northeastern State Community College, Blountville, Tennessee; Steve Hamilton, Austin Peay State University, Clarksville, Tennessee; Brad Harub, Kentucky Wesleyan College, Owensboro, Kentucky; Tera Haslem, Clarksville, Tennessee; Wallob Hebel, Austin Peay State University, Clarksville, Tennessee; Laura Helms, Austin Peay State University, Clarksville, Tennessee; Lori Hicks, Austin Peay State University, Clarksville, Tennessee; Cathy Hoback, Austin Peay State University, Clarksville, Tennessee; Gina Howe, Southern lllinois University, Carbondale, lllinois; Annie Isenhour, Tennessee Department of Environment and Conservation, Nashville, Tennessee; Jeannette Jones, Murray State University, Murray, Kentucky; James Joyner, Austin Peay State University, Clarksville, Tennessee; Steven Karr, Carson-Newman College, Jefferson, Tennessee; Thomas Kind, Murray State University, Murray, Kentucky; R. Kingsolzer, Kentucky-Wesleyan, Owensboro, Kentucky; John Koons, Jackson State University, Jackson, Tennessee; Mina Kobraei, Murray State University, Murray, Kentucky; Jill Kruper, Hood College, Maryland; Matt Kulp, Tennessee Technological University, Cookeville, Tennessee; Jeff Lebkeucher, Austin Peay State University, Clarksville, Tennessee; Bobby Lee, Hancock Biological
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