The Flora of the Ledges Region of Boone County, Iowa

Proceedings of the Iowa Academy of Science

Volume 22 Annual Issue Article 13

1915

William W. Diehl Iowa State College

Let us know how access to this document benefits ouy

Recommended Citation Diehl, William W. (1915) "The Flora of the Ledges Region of Boone County, Iowa," Proceedings of the Iowa Academy of Science, 22(1), 77-104. Available at: https://scholarworks.uni.edu/pias/vol22/iss1/13

This Research is brought to you for free and open access by the Iowa Academy of Science at UNI ScholarWorks. It has been accepted for inclusion in Proceedings of the Iowa Academy of Science by an authorized editor of UNI ScholarWorks. For more information, please contact [email protected]. Diehl: The Flora of the Ledges Region of Boone County, Iowa

FLORA OF THE LEDGE'S 77

'l'HE FLORA OF THE LEDGES REGION OF BOONE' GOUNTY, IOWA.

WILLIAM W. DIEHL. INTRODUCTION. . In the south-central part of Boone county is an area that possesses characteristic plants as well as peculiar geological and top.ographic features. This is located four miles directly south of Boone and extends for about two miles south toward and along the Des Moines river. Its name "The Ledges" is appro priate because of striking sandstone cliffs from ten to over forty feet in height, extending beyond the perpendicular in some cases as much as twelve feet. This ledge rock here comprises an island of younger massive sandstone surrounded uncon formably by somewhat older Carboniferous layers., which are of different structure.1 The conditions peculiar to this restricted locality give the flora a distinctive character. The region has been of sp.ecial interest at various times to students of plant life. Plants from "The Ledges" have been col lected and studied chiefly by students from Ames. Thus the herbarium of the Department of Botany of Iowa State Col lege possesses much material from that source. Hitchcock prob-· ably made us,e of some material from ''The Ledges'' in his Anth0- phyta ·and Pteridophyta of Ames, Iowa. L. H. Pammel in "The Character and Distribution of Forest Trees and Shrubs of Boone County, Iowa '•i gives a list which was hased largely upon .collections and observations made at ''The Ledges.'' Bruce Fink collected lichens at The Ledges in 19033 • Collections pre served in the herbarium of the Department of ·Botany have been made at various times by L. H. Pammel, R. E. Buchanan, C., M. King, Robert Combs, C. R. Ball, J. P. Anderson, J. V. Ellis, and others. THE SCOPE OF 'XHE WORK.

I The scope of this pap.er is to list the Pteridophytes and Sper matophytes of "The Ledges" with an account of the plant .ch~r acters of the re~on as revealed by a detailed study of a strip across a characteri"stic location. . . ' .

Published by UNI ScholarWorks, 1915 1 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

F-L( 78 IOWA ACADEMY OF SCIENCE seventh edition,7 and t The area from which this collection was taken includes prac-. used throughout. Thii tically all that region, originally part of the forest belt adja t~ the limitations of th cent to the Des Moines river, which is drained by the ravines that 'Converge among the ledge outcrops. The more restricted THE OIL area, represented on the map (fig. 1) by the shad1ed part, is It is evident from tl sesses a flora typical c recently covered with of this area was woo limit of the unshade1 mately the northern e: belt. Along the roads: nants of such prairie which }iave not been . the forest once extend of tree stumps over m growing near a clum11 rings. Residents of tl this land was once we confined to the rough slopes of the ravines 1 The catalogue indi, of the vegetation of t and Viburnum Lenta~ tract to the westwar< N Ledges.'' Aesculus while plentiful at Fr about three miles to · what similar, has not · thalictroides L. is abu [-->.I Mile + in Story county, but i FIG, 1.-Map of the "Ledges" region. The shaded part indicates the area characterized. by the sandstone outcrops. · Oaltha palustris L. co Moines river to the '1 characterized by outcropping p.ative sandstone rock. All of the been unable to find it plants listed as collected by the writer were obtained within ''The Ledges'' may this area at various times from September, 1914, until JunB, of Oladonia sylvatica, 1915. Those specimens which are listed as collections by others phyllus, Polypodium may have been obtained within the boundaries of the unshaded Mi'tella diphylla, Jun part of the map'. All the plants in the catalogue are repre ·rus och.roleucus, Phy sented by specimens in the h!')rbarium of the Department of ica and Rhamnus lari Botany of the Iowa State College ·at Ames. The arrangement ' followed is that of Engler and Prantl as used in Gray's Manual,

https://scholarworks.uni.edu/pias/vol22/iss1/13 2 Diehl: The Flora of the Ledges Region of Boone County, Iowa FLORA OF THE LEDGES 79 SCIENCE seventh edition,7 and the nomenclature of this manual has been was taken includes prac-. used throughout. This list· is incomplete in some respects, due t of the forest belt adja to the limitations of the season when this study was undertaken. is drained by the ravines ops. The more restricted THE CHARAOTER OF TBE FLORA. ) by the shadied part, is It. is evident from the appended catalogue that this. area pos sesses a flora typical of a forested region or of a region until recently covered with woodland. Originally . all or nearly all of this area was wooded. It would seem that the northern limit of the unshaded part of the map represents approxi mately the northern extension of the former encroaching forest belt. Along the roadside just north of this line are some rem nants of such prairie· grasses as Spartina Michauxii Hitch, which }J.ave not been found within the area in question. That the forest once ext.ended to this line is shown by the ·presence of tree stumps over much of the area. The stump of a bur oak growing near a clump. in this vicinity showed sixty-one annual rings. Residents of the district also attest that practically all this land was once wooded. Of course the wooded area is now con.fined to the rough land near the river and along the steep slopes of the ravines adjacent. The catalogue indicates the somewhat anomalous character of the vegetation of the region. For. instance, Morus rubra L. · and Viburnum Lentago L., which are plentiful just outside the tract to the westward, have not yet been found within ''The Ledges." Aesculus glabra var. arguta (Buckley) Robinson, while plentiful at Frazer near Boone, and along Honey creek about three miles to the northwest where conditions are some what similar, has not been found at ''The Ledges.'' Anemonella thalictroides L. is abundant on the bluffs along the Skunk river in Story county, but it has not been collected at "Tl)e Ledges." haded part indicates the area ne outcrops. Caltha palustris L. covers the marshy southern bank of the Des Moines river to the west of ''The Ledges,''· but the writer has ndstone rock. All of the been unable to find it within the area in question. er were obtained within ''The Ledges'' may be said to be characterized by the presence · ember, 1914, until June, of Cladonia sylvatica, Po1lytrichum commune, Camptosorus rhizo d as collecti~ns by others phyllus, Polypodium vulgare, Woodsia obtusa, Trillium nivale, ndaries of the unshaded Mi'tella ·diphylla, Juniperus virginiana, Dirca palustris, Lathy the catalogue are repre rus ochroleucus, Physocarpus opuUfoUus, Prunus pennsylvan of the Department of ida, and Rhamnus lanceolata. Ames. The arrangement s used in Gray's Manual,

Published by UNI ScholarWorks, 1915 3 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

80 IOWA ACADEMY OF SCIENCE· FLORA OE

THE BELT TRANSECT;. Plat 6 is the twelve foot c .An idea of _the character of the region is best indicated per plants. haps by a detailed study of a belt transect or strip across some Plat 7 lies from the top. oJ part of the area rep.resentative of the diverse conditions which hogback, a distance of thirty

give ''The Ledges'' their peculiarity. The belt chosen extends to .At this point, where the 1 ward the southeast from the bank of Pease creek, up the north vegetation is immediately o:f facing hillside, over the ridge or hogback and down the south proaching a prairie type. Bt facing slope to Pease creek. The belt is represented on the map established in places, giving (fig. 1) as a black block. plants more commonly found

P.tl

1· Plat 8 extends ten yards d1 Bred chiefly by A.n.tennaria pla Plat 9 is a tract twenty-six growth. Plat 10, sixteen yards long, posed rock, with scant plant-c

013, Plat 11 is a twenty-eight : FIG. 2.-Diagram showing the relative location of the various plots in the · strip survey. dense growth of shrubs and y From this point there is a The strip is six feet in width. The north and south slopes are down a bare sandstone cliff, pl designated ".A" and "B" respectively. Each has been subdi and alluvial bank nine feet ac vided into separate plats because of their somewhat individual characters. stream. PART A. The appended tables give tl centage basis with the numbe1 Plat l is two yards long. It is the area directly along the yard, rated at 100 per cent. stream bank, and is made up of plants growing a part of the time partly submerged. · THE PERCENTAGE DISTRIBUTION O] Plat 2, three yards in length, extends from the edge of plat

1 to the base of the hill ; it has many ch.aracters in common Svecies with 1. Plat 3 extends up the talus slope fo;r six yards to the first Adiantum pedatum ...... ; ___ _ vertical cliff or rock exposure. Here, as on much of the hill *Camptosorus rhizophyllus Cystopteris fragilis ...... side above, the extreme mesophytic conditions are reflected in Woodsia obtusa ...... , the type of plants found. Equisetum arvense ...... Equisetb.m hyemale val'. Plat 4 is the part of the ·belt which extends up the cliff at robustu1n ...... this point. In one respect this location is not typical of the Muhlenbergia Sp...... Cinna arundinacea ...... ledges in that here is found the Moosewood, Dirca palustris, Poa pratensis ...... which is nowhere else in this region. Glyceria aquatica . . . • . . . . + Plat 5 extends for twenty-three yards up. the slope to the Eleocharis palustris...... + *NOTE-For such plants not easily base of the next cliff, and goes up most of the way in a rock cated by + or - ; these are not taken strewn ravine. 6

https://scholarworks.uni.edu/pias/vol22/iss1/13 4 Diehl: The Flora of the Ledges Region of Boone County, Iowa 'SCIENCE· FLORA OF THE LEDGE'S 81

NSECT,. Plat 6 is the twelve foot cliff, which is bare of any vascular

~gion is best indicated per plants. •ansect or strip across some Plat 7 lies from the top. of the cliff to the watershed of the 1e diverse conditions which hogback, a distance of thirty-eight yards. The belt chosen extends to· At this point, where the sun's rays strike the hilltop, the 'Pease creek, up the north~ vegetation is immediately of a different character, even ap g-back and down the south proaching a prairie type. But the woody plants are becoming ; is represented on the map established in places, giving shade sufficient to p;rotect those plants more commonly found on the north slope. PART B. Plat 8 extends ten yards down the hillside. The plat is cov

<. ered chiefly by Antennaria plantaginifolia (L.) Richards. Plat 9 is a tract twenty-six yards long, covered by a shrubby growth. II. Plat 10, sixteen yards long, is rather open clay and dry exposed rock, with scant plant-cover. Plat 11 is a twenty-eight yard strip covered with a fairly lion of the va.rious plots in the . dense growth of shrubs and young trees. From this point there is a sheer drop of twenty-four feet north and south slopes are down a bare sandstone cliff, plat 12, to plat 13, which is a. talus 'ly. Each· has been subdi and alluvial bank nine feet across, reaching to the edge of the their somewhat individual stream. ... The appended tables give the data upon a comparative percentage basis with the number of plants in the average square he area directly along the yard, rated at 100 per cent. nts growing a part of the THE PERCENTAGE DISTRIBUTION OF PLANTS ON nrn NORTH SLOPE (A).

ds from the edge of plat Percentage to Each Plat ny characters in common Svecies l 2 \ 3 I 4 \ 5 I 6 \ 7· --2:6-gy--·--- for six yards to the first Adiantum pedatum ...... *Camptosorus rhizophyllus , as on much of the hill Cystopteris fragilis ...... 172.8 onditions are reflected in Woodsia obtusa ...... 4 0.9 Equisetum arvense ...... + Equiset\lm hyemale va1'. ch extends up the cliff at r

Published by UNI ScholarWorks, 1915 5 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

82 IOWA ACADEMY OF SCIENCE .FLORA OJ PERCENTAGE DISTRIBUTION, NORTH SLOPE (A)-Continued PERCENTAGE DIST.RIBUTI

Percentage to Each Plat Species 6 'l Species Carex pennsylvanica ...... Carex sp...... Gali um circaezans ...... Carex sp ...... Galium boreale ...... Arisaema triphyllum ...... 8.7 2.2 Galium trifidum ...... Polygonatum commutatum. 0~8 Lonicera dioica ...... Salix longifolia ...... 19.4 3.5 2.2 Viburnum pubescens ...... Populus grandidentata ... . 0.1 Vernonia altissima ...... Corylus americana ...... 0.5 Solidago latifolia ...... Ostrya virginiana ...... 0.4 1.8 3.4 Solidago cariadense ...... Carpinus caroliniana ...... 9.6 0.8 1.8 Aster sagittifolius ...... Quercus alba ...... 0.2 E'rigeron philadelphicus .. Quercus rubra ...... I 1.1 Ambrosia trifida ...... Ulmus fulva ...... I 0.8 0.2 Heliopsis scabra ...... Asarum canadense ...... · I 2.6 0.9 Rudbeckia faciniata ...... Thalictrum dioicum ...... 15.7 6.6 9.2 8.7 Prenanthes aiba ...... Hepatica acutiloba ...... 4.3 3.5 10.6 12.9 Anemone virginiana ...... 1..7 Anemone quinquefolia .... . 7.8 0.2 0,6, THE FERC'ENTAGE DIS1'RIRUTION ISopyrum biternatum .... . 1.7 0.6 Aquilegia canadensis ...... 9.6 0.1 Sanguinaria canadensis .. . 1.7 Draba caroliniana ...... 0.2 Species Mitella diphylla ...... 1 13.2 6.4 1.8 Ribes cynosbati...... 1 0.8 0.4 0.2 Amelanchier canadensis .. . 1.6 .Adiantum pedatum ...... Rubus occidentalis ...... J 0.8 0.4 0.2 Equisetum arvense ...... Rosa pratincola ...... 0.2 Equisetum hyemale var. robusta Prunus virginiana ...... 1.7 Panicum lanuginosum ...... Astragalus canadensis .... . 0.4 Muhlenbergia tenuiflora ...... , Desmodium grandiflorum .. 8.7 1.1 Elyrnus robustus ...... Rhus Toxicodendron ...... 1.7 o.~ 0.2 Carex sp ...... '...... Evonymus atropurpureus .. 2.6 Hypoxis hirsuta ...... Celastris scandens ...... 0.8 1.7 Populus grandidentata ...... Acer saccharum ...... 0.8 0.2 Corylus americana ...... Impatiens pallida ...... 1,7 Ostrya virginiana ...... Psedera quinquefolia ...... 0.4 0.6. Carpinus caroliniana ...... Vitis vulpina ...... 1.7. 0.6. Quercus alba ...... Tilia americana ...... 0.8 0.1 Quercus rubra ...... Dirca palustris ...... 0.8 Ulmus fulva ...... Aralia racemosa ...... '0.8 0.8 0.4 ' Ulmus racemosa ...... Aralia nudicaulis ...... 2.0 Thalictrum dioicurn ...... Osmorhiza Claytoni...... 1.7 0.4 Hepatica acutiloba ...... Cornus circinata ...... 0.2 .Anemone virginiana ...... Cornus asperifolia ...... 2.5. .Anemone quinquefolia ...... Cornus alternifolia ...... 0.4 Menispermum canadense ...... Fraxinus americana ...... --·-- 0.4 Ainelanchier canadensis ...... Fraxinus nigra ...... 0.2 Fragaria virginiana var. illinoiS· Hydrophyllum appendicula- ensis ...... tum ...... 1.3 Potentilla argentea ...... Teucrium canadense ...... 0.8 Rubus occidentalis ...... Prunella vulgaris ...... 5;9 Rosa pratincola ...... , ... . Monarda fistulosa ...... 10.6 13.0 2.6 Prunus virginiana ...... Pedicularis canadensis ... . 10.6 13.0 2.6 Melilotus alba ...... Galium aparine .. : ...... Amorpha canescens ...... + + Astragalus canadensis ......

https://scholarworks.uni.edu/pias/vol22/iss1/13 6 Diehl: The Flora of the Ledges Region of Boone County, Iowa . OF SCIENCE FLORA OF THE LEDGES 83 ORTH SLOPE (Al-Continued PERCENTAGE DISTRIBUTION, NORTH SLOPE (Al-Continued

Percentage to Each Plat Percentage to Each Plat 'l. Species j2[sj4/516 7 Galium circaezans .....•. + Galium boreale ...... + + 8.7. 2.2 Galium trifiduin ...... + + Lonicera dioica ...... 0.8 1.3 0.2 0.8 0~8 Viburnum pubescens ...... 2.2 .5 2.2 0.1 Vernonia altissima ..•.... 0.8 2.2 0.9 Solidago latifolia ...... 17.5 33.4 3.6 I- 0.5 3.4 Solidago cana,dense ...... 122.6 31.4 0.4 1.8 Aster sagittifolius ...... 108.1 20.3 9.6 0.8 1.8 0.8 0.2 Erigeron philadelphicus .. 1.7 10.7 1.1 Ambrosia trifida ....•.•. ;. 0.4 Heliopsis seabra ...... 7.0 1.9 0.8 0.2 Rudbeckia laciniata ....•. 17.6 14.8 1.3 0.1 2.6 0.9 Prenanthes alba ...... : 15.7 6.6 9.2 8.7 1.3 4.3 3.5 10.6 12.9 1.7 7.8 0.2 M THE l>E:RC'ENTA:GE DISTRIBUTION OF PJ,A:NTS ON THE SOUTH SLOPE (B). 1.7 0.6 9.6 0.1 Percentage of Plants to Each Plat 1.7 0.2 Species 13.2 6.4 1.8 0.8 0.4 0.2 1.6 Adiantum pedatum .•...... 0.1 0.8 0.4 0.2 Equisetum arvense ....••...... + 0.2 Equisetum hyemale var. robusta + 1.7 Panicum lanugino>:um ...... 0.4 Muhlenbergia tenuiflora ...... + 1.1 Elymus robustus ...... 8.7 Carex sp ...... + 1.7 o.~ 0.2. + 2.6 Hypoxis hirsuta ...... 0.3 3.6 0.8 1.7 Populus grandidentata ...... 0.1 0.8 0.2 Corylus americana ..•...... 2.3 1.7 Ostrya virginiana ...... 2.3 2.9 1.2 0.4 0.6. Carpin us caroliniana ...... 0.9 1.7 0.6 Quercus alba ...... 0.1 0.8 0.1 Quercus rubra ..... ·...... 0.9 0.7 0.2 0.8 tnmus fulva ...... •...... 0.3 1.1 1.0 Ulmus racemosa ...... '0.8 0.8 0.4 ·3.5 2.0 Thalictrum dioicum ...... 31.5 3.5 0.2 1.7 0.4 Hepatica acutiloba ...... 1.1 6.2 0.2 Anemone virginiana ...... 0.4 0.3 2.5. Anemone quinquefolia ...... 1.9 0.4 Menispermum canadense ...... 8.4 0.4 Amelanchier canadensis ...... 1.3 _0.1 0.2 Fragaria virginiana var. illinoisensis ...... 1.3 2.0 1.3 Potentilla argentea ...... 0.1 - 0.8 Rubus occidentalis ...... 0.5 0.4 ;9 Rosa pratincola ...••..•... , ... . 0.6 .6 13.0 2.6 Prunus virginiana ...•..•...... 1.0 .6 13.0 2.6 Melilotus alba ...•..•...•...... 5.5 Amorpha canescens ...•...... 1.4 3.9 + + Astragalus canadensis ...... 1.2 1.G

Published by UNI ScholarWorks, 1915 7 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

84 IOWA ACADEMY OF SCIENCE FLORA OF~

THE PERCENTAGE DISTRIBUTION OF PLANTS ON THE SouTH SLOPE (B).

l'erG€'Iltage of Plants to Each Plat CATALOGUE OF THE PL Species 12 18 PTERID 8 l 9 I 10 l 11 l I Lespedeza capitata ...... 1.3 FILI! Vicia americana ...... 0.4 Polyp1 Lathirus ochroleucus ...... 0.6 0.5 Zanthoxylum amerieanum ...... ---- 0.2 Polypodium vulgare L. L. H. ] Oxalis corniculata ..•...... I ---- 0.5 4.2 M. K., 03; 1, W. W. D., Polygala Senega ....•...... 1.3 ---- posures. Rhus glabra ..•...... 1 ---- 3.5 Rhus Toxicodendron ..•...... ---- 0.1 0.7 Adiantum pedatu.m L. R. E. Vitis vulpina ...... 0.3 Tilia americana ...... 0.1 C. M. K., 03; 2, W.W. D. AraIi a nudicaulis .... ·, ...... 0.9 0.4 0.1 0.9 Asplenium Fiiix-femina (L.) ] Zizia aurea .•...... 0.4 0.7 Cornus asperifolia ...... 3.4 1.3 6.9 C. M. K., 03. (No collec1 Fraxinus americana ...... 1.0 made.) Apocynum androsaemifolium .... 0.1 0.1 Phlox pilosa ...... ; ...... 3.0 0.5 Camptosorus rhizophyUus (L.: Phlox divaricata ..•...... 0.1 0.1 C. M. K., 03; 3, W.W. D. Teucrium canadense ...... 3.1 Monarda fistulosa .•...... 2.5 0.1 15.8 Cystopteris fragilis (L.) Bern Pediculari$ canadensis ...... 03 R. E. B., C. M. K., 03; L Galium circaezans ...... ·.. + + W.W. D. Common on w Gali um boreale ..••.••...... + Galium sp...... + Woodsia obtusa (Spreng.) Tor Lonicera dioica ..•...... 0.6 0.4 97; 3903, L. H. P., R. E. Viburnum pubescens ...... 5.2 1.2 Solidago latifolia (?) ...... 0.3 7.6 On shaded, rocky banks. Solidago ulmifolia (?) ...... 2.9 Solidago canadensis (?) ...... 0.3 0.6 OphiOj Aster sagittifolia ...... 0.9 6.1 Aster sp...... 4.3 2.0 3.9 Botrychium virgitnianum (L.) Erigeron philadelphicus .... ~ .... 40.5 7.3 B., 02; 32, J. V. E., 14; 6 Heliopsis scabra ...... 13.0 0.7 6.3 ---- Prenanthes alba ...... •.. 0.3 0.3. woods. ---- I ---- EQUl EquJ Equisetum.arvense L. E. Bis W. W. D., 15. Common Equisetum hyemale L. var. r R. E. B., 02; 3979, L. E J. V. E,, 14; 8, W.W. D SPERM GYMN CON'

~ Juniperus virginiana L. L. 03; 9, W. W. D., 14. On

https://scholarworks.uni.edu/pias/vol22/iss1/13 8 Diehl: The Flora of the Ledges Region of Boone County, Iowa

[)F SCIENCE FLORA OF THE LEDGES 85 lNTS ON THE SOUTH SLOPE ( B) . ereentage of Plants to Each Plat CATALOGUE OF THE PLANTS OF "THE LE'DGES."

12 13 PTERIDOPHYTA. 9 10 11 l ! l I l FILICALES. 1.3 0.4 Polypodiacere. 0.6 0.5 96; 296, R. 0.2 Poiypodium vulgare L. L. H.P., L. H. P., E. B., C. 0.5 4.2 M. K., 03; 1, W. W. D., 14. Common on sandstone ex- 1.3 3.5 :t posures. 0.1 0.7 I Adiantum pedatum L. R. E. B., 02; 3938, L. H. P., R. E. B., 0.3 i C. M. K., 03; 2, W. W. D. Common in rich woods. 0.1 I 0.4 0.1 0.9 I Asplenium Filix-femina (L.) Bernh. 3926, L. H. P., R. E. B., 0.4 . 0.7 ----- C. M. K., 03. (No collection of this plant has since been 3.4 1.3 6.9 1.0 made.) 0.1 0.1 Camptosorus rhizophyiius (L.) Link. 3974, L. H. P., R. E. B., 3.0 0.5 0.1 0.1 C. M. K., 03; 3, W. W. D. On damp sandstone exposures. 3.1 Cystopteris fragilis (L.) Bernh. L. H. P., 98; 3892, L. H. P., 2.5 0.1 15.8 .03 R. E. B., C. M. K., 03; L. H. P., 13; 3, J. V. Ellis, 14; 4, + + W.W. D. Common on woody slopes. + + Woodsia obtusa (Spreng.) Torr. L. H.P., 96; L. H.P. & R. C., 0.4 97; 3903, L. H. P., R. E. B., C. M. K.,03; 5, W. W. D., 15. 5.2 1.2 0.3 7.6 On shaded, rocky banks. 2.9 Ophioglossacere. 0.6 0.9 6.1 Botrychium virginianum (L.) Sw. L. H. P. & R. C., 97; R. E. 2.0 3.9 - 40.5 7.3 B., 02; 32, J. V. E., 14; 6, W. W. D., 15. Common in rich 0 0.7 woods. 3 0.3 -~~~ I ==== EQUISETALES, Equisetacere. Equisetum arvense L. E. Bissell, L. H. P., 10; 36, V. E. 14; 7, W.W. D., 15. Common in sandy alluvium. Equisetum kyemale L. var. robustum (A. Br.) A. A. Eaton. R. E. B., 02; 3979, L. H. P~, R. E. R, C. M. K., 03; 33, J. V. E., 14; 8, W. W. D., 14. SPERMATOPHYTA. / GYMNOSPERM.lE. CONIFERALES. Pinacere. Juniperus virginiana L. L. H. P., 98, 99, i3; 3935, C. M. K., 03; 9, W. W. D., 14. On exposed hillsides, rare.

Published by UNI ScholarWorks, 1915 9 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

FLORA OF 86 IOWA ACADEMY OF SCIENCE

A.NGIOSPERM.1E. M uhlenbergia racemosa (Mich W. D., 15. Moist banks o MONOCOTYLEDONlE. Muhlenbergia Mexicana (L.) ' PANDANALES. Phleum pratense L. 28, J. V. Typhacere. .Agrostis alba L. 3940, L. H. Typha latifolia L. 62, J. V. E.; 10, W. W. D., 15. In artificial P., 13; 68, J. V. E., 14. swainp on upland. .Agrostis hyemalis (Walt.) B NAJADALES. C'. M. K., 03. Alisruacere. .Agrostis perennans (Walt.) Ti 0. M. K., 03. Sagittaria lafifolia Willd. 11, W. W. D., 15, On flood plain of the Des Moines river. Oinna arundinacea L. 3930, ] 23, W. W. D., 15. Moist Alisma Plantago~aqiiatica L. L. II. P., 14. Sphenopholis obtusata (Mich:ii GRAMINALES. sides. Grallllinea· Danthonia spicata (L.) Beauv. Andropogon f'urcatus Muhl. R. E. B., 02; 12, W .. W. D., 15. k., 03; 24, W. W. D., 15. Roadsides. Eragrostis hypnoides (Lam.) I Sorghastrum nutans (L.) Nash. R. E. B., 02. 96~ Digitaria humifusa Pers. ·L. H.P., 91. . Eragrostis capillaris (L.) Nees. 96. Digitaria sanguinalis (L.) Scop. L. H. P., 91; 13, W. W. D., 15. Upland fields. Eragrostis Frankii (Fisch. Ma~ P., 96. Panicum capillare L. L. H. P., 96; R.. E. B., 02 ~ 14, W. W. D., 14. Common in fields. Eragrostis pilosa (L.) Beauv. Panicum virgatum L. 287, W. W. D., 1{). Eragrostis Megastachrya; (Koele Panicum lanuginosum Ell. 15, W.W. D., 15. Dry exposed hill Poa pratensis L. 109, W. W side. common. Panicum latifolium L. L. H. P., R. E. B., C. M. K., 03; W. W. Glyceria nervata (Willd.) Trin D, 14. Upland woods. 15. Banks of Pease creek l 3908, Panicum villosissimum Nash. 3879, L. H. P., R. E. B., C. M. Bromus ptUrgans L. L. H L. · 3924,' L. K., 03. . Elymus virgin'icus E. robustus Scribn. & J. G. Sm. Echinochloa crusgalli (L.) Beauv. 17, W. W. D., 14. Upland fields. K., 02; 111, W. W. D., 14. Moench. L. H. Setaria glauca (L.) Beauv. 18, W.W. D., 14. Up.land fields. H ystrix patula 0. M. K., 03; L. H.P., V Setaria viridis (L.) Beauv. 65, J .. V_. E., 14; 19, W.W. D., 14. J. Upland fields. Cypera Oenchrus carolinianus Walt. 20, W. W. D., 15. Sandy allu- Cyperus esculentus L. 291, W. vium. Eleocharis palustris (L.) R. & Leersia virginica Michx. L. H. P., 96. submerged, along Pease ere Leersia oryzoides (L.) Sw. L. H. P., 15. Scirpus validus Vahl. 67, J. V. Muhlenbergia tenuiflora (Willd.) R S. P., 21, W. W. D., 15. Carex rosea Schkuhr; ~ 197, W. , Exposed, rocky hillsides.

https://scholarworks.uni.edu/pias/vol22/iss1/13 10 Diehl: The Flora of the Ledges Region of Boone County, Iowa FLORA OF THE LEDGES 87 r SCIENCE Muhlenbergia racemosa (Michx.) B. S. P., L. H. P., 96; 22, W. W. D., 15. Moist banks of Pease creek. ·

~NlE. Muhlenbergia Mexicana (L.) Trin. L. R. P., 15.

~. Phleum pratense L. 28, J. V. E., 14. Agrostis alba L. 3940, L. H. P., R. E. B., C. M. K., 03; L. H. P., 13; 68, J. V. E., 14. W. W. D., 15. In artificial Agrostis hyemalis (Walt.) B. S. P. 3942, L. H. P., R. E. B., C'. M. K., 03. Agrostis perennans (Walt.) Tuckerm. 3688, L. H.P.; R. E. B., C. M. K., 03. W. D., 15. On flood plain Cinna arundinacea L. 3930, L. H. P;, R. E. B., · C. M. K.,. 03; 23, W. W. D., 15. Moist shaded banks of Pease creek. P., 14. Spihenopholis obtusata (Michx.) Scribn. L. H. P., 15. Hill sides. s. Danthoma spicata (L.) Beauv. 3951, R. E. B., L. R. ·p., C. M. k., 03; 24, W.W. D., 15.. Exposed hillsides . . B., 02; 12, W. W. D., 15. Eragrostis hypnoides (Lam.) B. S. P. 108, L. H. P., C. R. B., 96~ E. B., 02. Eragrostis capillaris (L.) Nees. L. H.P., 96; C.R. B., L. H.P., 91. 96. R. P., 91; 13, W. W. D., 15. Eragrostis Frankii (Fisch. May. & Lall.) Steud. C.R. B., L. R. P., 96. R.' E. B., 02; 14, W. W. D., Eragrostis pilosa (L.) Beauv. L. H. P., 15. Eragrostis Megastachya (Koeler.) Link. 183, W. W. D.

D., 15. l: Poa pratensis L. 109, W. W. D., 15. Woods and pastures, '. 'TV. D., 15. Dry exposed hill- common. GZyceria nervata (Willd.) Trin. L. H. P., 96; 110, W. W. D., E. B., C. M. K., 03; W. W. 15. Banks of Pease creek and.of Des Moines river. Bromus p'llrgans L. 3908, L. H. P., R. E. B., C. M. K., 03. , L. R. P., R. E. B., C. M. Elymus virgin'icus L. · . 3924, L. H. P., R. E. B., C. M. K., 03. E. robustus Scribn. & J. G. Sm. 3925, L. R. P., R. E. B., C'. M. 17, W. W. D., 14. Upland K., 02; 111, W. W. D., 14, Hystrix patula Moench. L. H. P., 91; 3902, L. H. P., R. E. B., W. D., 14. Upland fields. C. M, K.; 03; L. H. P., J. V. E., 13. V: E., 14; 19, W. W. D., 14. Cyperacere. Oyperus escufontus L. 291, W. W. D., 15. Roadside. 'TV. W. D., 15. Sandy allu- Eleocharis palustris (L.) R. & S. 112, W. W. D., 15. Partly submerged, along Pease creek. 96. Scirpus validus Vahl. 67, J. V. E., 14. '., 15. Carex rosea Schkuhr,t 197, W.W. D., 15. Rich shaded hillside. R S .. P., 21, W. W. D., 15.

Published by UNI ScholarWorks, 1915 11 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

88 IOWA ACADEMY OF SCIENCE FLORA OF~ Carex gromida Bailey. 462, L. H. P., R. H. C., 97; L. H. P., 98. Carex pennsylvanrica Lam. 113, W. W. D., 15. Common on hill- Amaryl sides. Hypoxis hirsuta (L.) Coville. Carex panicea L. var. Meadi"i (Dewey) Bailey. 193, W. W. D. on hilltop. Carex eburnea Boot.? 305, W. \V. D., 15. ORCHl Carex laxiflora Lam. L. H. P., R. H. C., 97. Orchl Cafcex laxiflora Lam. var. latifolia Boot. 114, W. W. D., 15. Cypripedium candidum. Muhl. Rich wooded hillsides, frequent. Cypripedium hirsutum Mill. l Carex oligocarpa Schkuhr. L. H. P., C. R. B., 97; L. H. P., 13. Orchis spectabilis L. L. H. P., Carex grisea Wahlenb.? Too young. 194, W.W. D., 15. Habenaria bracteata (Willd.) 1

Carex Crawei Dewey.? Too young. 195, W. W. D., 15. DICOTY Carex Assin'iboinensis W. Boot.¥ 196, W. W. D., 15. Archicl

ABALES. SALI1 Aracere Salii Arisaema triphyllum (L.) ·Schott. L. H. P., R. H. C., 97; L. Salix nigra Marsh. L. H. P., H. P., 14; 115, W. W. D. Woods, common. Pease creek and Des Moim Arisaema Dracon.tium (L.) Schott. · L. H. P., 97, 13; L. H. P., Salix amygdaloides Anders. 3, R. H. C., 97; L. H. P., C. R. B., 98. 01; 3876, L. H.P., R. E. l r,I XYRIDALES. 15. Along Pease. creek an Salix longifoZia Muhl. 3870, I CommeUnacere H. S. Kellogg, 06; 125, l Tradescantia refiexa Raf. 315, W. W. D., 15. creek and Des Moines rive

LILIALES. Salix discolor Muhl. L. H. P., Liliacere Salix humilis Marsh. 3873, L. Populus tremuloides Michx. 1: Uvularia grandiflora Sm. 116, W.W. D., 15. Woods, common. tops and level upland. Erythronium albidurn Nutt. 117, W. W. D., 15. Woods, not Populus grandidentata Michx. uncommon. E. B., C. M. K., 03; L. H. l R. Allium canadense L. L. H. P., H. C., 97. Populus deltoides Marsh. L. 1 liilium philadelphicum L. L. H.P., 15. D. Valley of Pease creek Smilacina racemosa (L.) Desf. R. E. B., 02. JUGL.A Polygonatum commutatum (R. & S.) Dietr. 118, W.W. D., 15. Jugla Woods, common. Juglans cinerea L. 129, W. W. Trillium nivale Riddell. E. Bissell, L. H. P., 10; 119, W. W. D., Juglans nigra L. 130, W. W. · 15. Common. Carya ovata (Mill.) K. 'Koch. Smilax herbacea L. 120, W. W. D., 15. Edge of upland wood. Carya cordiform'is (Wang.) K. Smilax ecirrhata (Engelm.) Wats. 121, W.W. D., 15. sides. Smilax hispida Muhl. L. H. P., 98; 45, J. V. E., 14; 122, W. FA~ W. D., 15. BefaJ Corylus americana Walt. 13:'. abundant.

https://scholarworks.uni.edu/pias/vol22/iss1/13 12 Diehl: The Flora of the Ledges Region of Boone County, Iowa

Y OF SCIENCE FLORA OF THE LEDGE'S 81} L P., R. H. 0., 97; L. H. P., 98. W. W. D., 15. Common on hill- Amaryllidacere Hypoxis hirsuta (L.) Coville. 123, W. W. D., 15. With moss )ewey) Bailey. 193, W. W. D. on hilltop. W. D., 15. ORCHIDALES. R.H. C., 97. Orchidacere 1lia Boot. 114, W. · W. D., 15. Cypripedium candidum. Muhl. Edith C'airns, 97. ent. Cypripedium hirsutum Mill. Edith Cairns, 03. L P., C. R. B., 97; L. H. P., 13. Orchis spectabilis L. L. H. P., 15. mng. 194, W. W. D., 15. Habenaria bracteata (Willd.) R. Br. mg. 195, W. W. D., 15. DICOTYLEDONJE. 196, W. W. D., 15. A rchiclamydeae. ,ES. SALICALES. Salicacere tt. L. H. P., R. H. C., 97; L. SaUx nigra Marsh. L. H. P., 13; 124, W. W. D., 15. Along Woods, common. Pease creek and Des Moines river. 1tt. L. H. P., 97, 13; L. H. P., Salix amygdaloides Anders. 3473, W. D. Fitzgerald, R. F. M., , B., 98. 01; 3876, L. H. P., R. E. B., C. M. K., 03; 125, W. W. D.,

~LES. 15. Along Pease. creek and in wet places on upland. Salix lonrfifolia Muhl. 3870, L. H. P., R. E. B., C. M. K., 03; inacere H. S. Kellogg, 06; 125, W. W. D., 15. Valley of Pease W.W. D., 15. creek and Des Moines river, abundant.

LES. Salix discolor Muhl. L. H. P., 98. cere Salix humilis Marsh. 3873, L. H. P., R. E. B., C. M. K., 03. Populus trem.uloides Michx. 126, W.W. D., 15. · Exposed hill- ~. W_. D., 15. Woods, common. tops and level upland. .7, W. W. D., 15. Woods, not Pop1tlus grandidentata Michx. L. H. P., 02; 3898, L. H: P., R. E. B., C. M. K., 03; L. H. P., 13; 127, W. W. D. Dry slopes. ft. H. C., 97. Populus deltoides Marsh. L. H. P., A. F. M., 12; 128, W. W. , P., 15. D. Valley of Pease creek and of Des Moines river. R. E. B., 02. JUGLANDALES, 1 S.) Dietr. 118, W.W. D., 15. Juglandacere Juglans cvn.erea L. 129, W.W. D., 14. Wooded slop.es, common. all, L. H. P., 10; 119, W. W. D., Juglans nigra L. 130, W. W. D., 14. Common. Carya ovata (Mill.) K. "Koch. 131, W. W. D., 15. Frequent. D., 15. Edge of upland wood. Carya cordiform'is (Wang.) K. Koch. 132, W.W. D., 14. Hill- ;s. 121, W. W. D., 15. sides. 98; 45, J. V. E., 14; 122, W. FAGALAS. Betulacere. Corylus americana Walt. 133, W. W. D., 14. Dry hillsides, abundant.

Published by UNI ScholarWorks, 1915 13 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

FLORA OF 90 IOWA ACADEMY OF SCIENCE ARISTOI Ostrya virginiana (Mill.) K. Koch. R. E. B., 02; L. H. P., 13; AristoJ J. P. A., L. H. P., 13; 134, W. W. D., 15. Hillside, abun Asarum canadense L. 3936, ] . dant. L. H. P., 10; 149, W. W. Carpinus caroliniana Walt. L. H. P., 98; R. E. B., 02; 135, W. common. W. D., 14. Shaded hillsides, common. POLY1 Fagacere. Poly1

Quercus alba L. 3897, L. H. P., R. E. B., C. M~ K., 03; 136, W. Rumex crispus L. 150, W. W W. D., 15. Hillside and upland, common. Rumex altissimus Wood. 151 Quercus macrocarpa Michx. R. E. B., 02; 154, J. V. E., 98; of Des Moines river. 137~ W.W. D.,14. Common. Rumex acetosella L. L. H.P., Quercus Muhlenbergii Engelm. L. H. P., C. R. B., 98; 3906, L. Polygonum aviculare L. 152, H. P., R. E. B., C. M. K., 03; 138, W. W. D., 15. Upland Polygonum convolvulus L. 3( and dry hillsides,' not uncommon. Polygonum virginianum L. I Quercus rubra L. 139, W.W. D., 15. Common. Fagopyrum esculentum Moern

URTICALES. CHEN ' Urticacere. Chen< I· Ulmus fulva Michx. L. H. P., C. R. B., 98; L. H.P., 98; 3943, Ohenopodium album L. 153, i L. H. P., R. E. B., C. M. K., 03; 18, J. V. E., 14; 140, W. Salsola; Kali L. var. tenuifolii W. D., 15. Common. 15. Weed, rare. Ulm.us americana L. 141, W. W. D., 15. Ravines and valleys. A:mai Ulmus racemosa Thomas. 142, .W. W. D., 15. Dry, rocky hill Amaranthus retroflexus L. 1 sides, and along Pease creek, rare. Amaranthu~ graecizans L. 11 Celtis accidentalis L. 143, W.W. D., 15. Valley of Des Moines CARY(] river. Caryc Oannab.is sativa L. 144, W. W. D., 15.. Pasture, east end of "Ledges". Silene stellata (L.) Ait. 386: Humulus lupulus L. 145, W.W. D., 15. Valley of Pease creek. 51, J. V. E., 14; 158, W. Morus rubra L. L. H. P., 98. (Probably just outside the Silen.e nivea (Nutt.) Otth. mapped area.) · . K., 03. Laportea canadensis (L.) Gaud. · 3927, L. H. P., R. E. B., C. M. Saponaria officinalis L. 159, K., 03; 42, J. V. E., 14; 146, W.W. D., 15. Woods, common. Glaytonia virginica L. 160, · Pilea pumila (L.) Gray. 147, W. W. D., 15. Low woods, com RANl mon. Por1 SANTALALES. SantaJ.acere. Ranun:culus aquatalis L. var. Ranunculus rhomboideus Go: Comandra umbellata (L.) Nutt. L. H. P., 92, 02, 12, 13; 3897, Ranun:culus abortivus L. 16 L. H.P., R .. E. B., C. M. K, 03; 148, W.W. D. Dry ex common. posed south slope.

https://scholarworks.uni.edu/pias/vol22/iss1/13 14 Diehl: The Flora of the Ledges Region of Boone County, Iowa

FLORA OF THE LEDGES 91 :>F SCIENCE ARISTOLOCHIALES • . R. E. B., 02; L. H.P., 13; Aristolochiacere. r. W. D., 15. Hillside, abun- Asarum canadense L. 3936, L. H. P., R. E. B., C. M. K., 03; L. H. P., 10; 149, W. W. D., 15. Damp shaded locations, P., 98; R. E. B., 02; 135, W. common. common. POLYGON ALES.

1• E. B., C. M~ K., 03; 136, W. Rumex crispus L. 150, W.W. D., 15. Fields. 1,nd, common .. Rumex altissimus Wood. 151, W. W. D., 15. Fields. in valley liJ. B., 02; 154, J. V. E., 98; of Des Moines river. Rumex acetoseUa L.' L. H. P., 15. ~ H. P., C. R. B., 98; 3906, L. Polygonum (J!l)iculare L. 152, W. W. D., 15. Common weed. I; 138, W. W. D., 15. Upland Polygon:um COlft.Volvitlus L. 300, w. vV. D., 15. Field, common. tnon. Polygonum virginianum L. L. H. P., R. F. M., 12. 15. Common. Fagopyrum esculentum Moench. L. H. P., 02.

,ES. . CHENOPODIALES. ere. Chenopodiacere. R. B., 98; L. H.P., 98; 3943, Chenopodium album L. 153, W.W. D., 15. Common wee¢!. 03; 18, J. V. E., 14; 140, W. Salsola; Kali L. var. tenuifolia G. F. W. Mey. 154, W. W. D.,' 15. Weed, rare. D., 15. Ravines and valleys. AmarantJ;iacere. r. W. D., 15. Dry, rocky hill Amaranthus retroflexus L. 155, W. W. D., 15. Common weed. rare. Amaranth"'!-~ graecizans L. 156, W. W. D., 15. D., 15. Valley of Des Moines CABYOPHYLLALES. D., 15. Pasture, east end of Caryophyllacere. Silene stellata (L.) Ait. 3863, L. H. P., R. E. B., C. M. K., 03; i D., 15. Valley of Pease creek. 51, J. V. E., 14; 158, W. W. D., 15. I (Probably just outside the Silene nivea (Nutt.) Otth. 3954, L. H. P., R. E. B., C. M. . K., 03. l Saponaria officinalis L. 159, W. W. D., 15. Roadside. I 3927, L. H. P., R. E. B., C. M. ii W.W. D., 15. Woods, common. Claytonia virginica L. 160, W. W. D., 15. Common in woo_ds. I ·. W. D., 15. Low woods, com- RANUNCUJ,ALES. II Portulacacere. ~LES. Ranun'Culus aquatalis L. var. capillaceous DC. L. H. P., 9. ~cere. Ranitnculus rhomboideus Goldie. L. H. P., 98. L. H. P., 92, 02, 12, 13; 3897, Ranunculus abortivus L. 161, W. W. D., 15. Damp hillsides, ., 03; 148, W. W. D. Dry ex- common.

Published by UNI ScholarWorks, 1915 15 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

92 IOWA ACADEMY OF SCIENCE FLORA OFT

Raniinculus septentrionalis Poir. 162, W. W. D., 15. Wooded Sisymbrium o1ficinale (L.) Scop hillsides, common. Erysimum cheiranthoides L. 3 Thalictrum diowum L. I,. H. P., 98, 13; R. E. B., 03; 163, W. K., 03. W. D., 15. Hillsides:, common. Radicula palustris (L.) Moench. Hepatica acutiloba DC. 164, W. ·w. D., 15. Shaded hillside, K., 03. common. Dentaria laciniata Muhl. 177, -V Anemone virginiana L. R. E. B., 02; 3884, L. H. P., R. E. B., Oardamine bulbosa (Schreb.) l C. M. K., 03; L. H. P., 13; 165, W. W. D., 15. Hillsides, fiths, R. H. C., 97; 1:57, W. frequent. Oardamine h~rsuta L. 3928, 3971 Anemone quirnquefolia L. 294, W. W. D., 15. Woods, common. Arabis laevigata (Muhl) .Poir. Clematis vir[jiniana L. 30, J. V. E., 14. K., 03. Isowrum biternatum (Raf.) L. & G. 166, W. W. D., 15. Hill Arabis oanadensis L. R. E. B., sides, common. ROSJ Aquilegia canadensis L. L. H. P., 98; 167, W. W. D., 15. Hill sides, common. Saxifri Actaea rubra (Ait.) Willd. 3917, L. H.P., R. E. B., C. M. K., H euchera villosa Michx. L. H. 03; 168, W. W. D., 14. Shaded hillsides. Mitella diphylla L. L. H. P., ( Menispermacere. B., C. M. K., 03; 180, W. V'i Menispermum canadense L. 6, J. V. E., 14; 169, W. W. D., 14. Ribes Oynosbati L. L. H. P., . Common. W. W. D., 15. Common . Berberidacere. Rib es fioridum L 'Her. L. H. I Podophyllum. p'Bltatum L. 170, W. W. D., 15. Common in Ribes gracile Michx. 182, W. · woods. Ros PAPA VERALES. Physocarpus opulifolius (L.) J\I Papa.veracere. D., 15. Wooded slop.es. Sanguinaria canadensis L. 171, W. W. D., 15. Open woods, Pyrus·ioensis (Wood.) Bailey. frequent. Fuilllllriacere. . 06; 185, W. W. D., 15. F Amelanchier canadensis (L.) ] Dicentra cucullaria (L.) Bernh. 172, W.W. D., 15. W.D., 15. Oruciferre. Ordtaegus punctata J acq. L. E Draba verna L. 173, W. W. D., 15. Roadsides, fields. C. M. K., 03; 187, W. W. Draba caroliniana Walt. 174, W.W. D., 15. Woods and fields. Orataegus tomen.tosa L. R. E. Lepidium virginicum L. 175, W. W. D., 15. Fields and road- Ora'taegus mollis (T. & G.) Sc sides, common. D., 15. Common. Lepidium apetalum Willd. L. H.P., 15. Fragaria virginiana Duchesne Capsella bursa-pastoris (L.) Medic. 176, W. W. D., 15. Fields 189, W. W. D., 15. ComIJ and roadsides, common. Frag~ri.a vesca L. var. americ BriMsica nigra (L.) Koch. R. E. B., 02. L. H. P., R. E. B., C. M. Sisymbrium canescens Nutt. L. H. P., 98; L. H. P., C. R. B., rocky hillside. 98. ~ Potentilla arguta Pursh. R. 1

https://scholarworks.uni.edu/pias/vol22/iss1/13 16 Diehl: The Flora of the Ledges Region of Boone County, Iowa I' ~ '

OF SCIENCE FLORA OF THE LEDGE'S 93

162, W. W. D., 15. Wooded Sisymbrium o1ficinale (L.) Scop. 301, W.W. D., 15. Fields. Erysimum cheiranthoides L. 3869, L. H. P., R. E. B., C. M. , 98, 13; R. E. B., 03; 163, W. K., 03. n . Radicula palustri,s (L.) Moench. 3954, L. H. P., R. E. B., C. M. . W. D., 15. Shaded hillside, K., 03. Dentaria laciniata Muhl. 177, W.W. D., 15. Hillside, common. ,, 02 '· 3884 ' L. H. P~, R. E. B., Oardamine bulbosa (Schreb.) B. S. P. L. H. P., C. J. Grif 165, W. W. D., 15. Hillsides, fiths, R. H. C., 97; 1'57, W. W. D., 15. Roadside ditch. Oard,amine h~rsuta L .. 3928, 3975, L. H. P., R. E. B., C. M. K., 03. ~. W. D., 15. Woods, common. Arabi,s laevigata (Muhl) .Poir. 3870, L. H. P., R. E. B., C. M. E., 14. . K., 03. ~ G. 166, W. W. D., 15. Hill- Arabi,s canadensi,s L. R. E. B., 03.

ROSALES. '., 98; 167, W. W. D., 15. Hill- Saxifragacere. 7, L. H. P., R. E. B., C. M. K., H euchera villosa Michx. L. H. P ., 02. ~ded hillsides. Mitella diphylla L. L. H. P., C. R. B., 98; 162, L. H. P., R. E. macere. B., C. M. K., 03; 180, W. W. D., 15. Damp shaded hillsides. · r. V. E., 14; 169, W. W. D., 14. Ribes Cynosbati L. L. H. P., 96, 02, 13; 8, J. V. E., 14; 181, W. W. D., 15. Common. [ifwere. Ribes fioridum L'Her. L. H.P., 15. Ribes gracile Michx. 182, W.W. D., 15. Common. , W. W. D., 15. Common m Rosacere. ~ALES. Physocarpus opulifolius (L.) Maxim. L. H. P., 13; 184, W. W racere. D., 15. Wooded slop.es. , W. W. D., 15. Open woods, Pyrudoensi,s (Wood.) Bailey. L. H.P., R.H. G., 97; L. H.P., iacere. · 06; 185, W. W. D., 15. Frequent. ' Amelanchier canadensis (L.) Medic. L. H. P., 02, 13; 186, W. 172, W.W. D., 15. W.D., 15. ferre. - ' Crr.ttaegus punctata Jacq. L. H.P., 98; 3427, L. H. P., R. E. B., , 15. Roadsides, fields. C. M. K., 03; 187, W. W. D., 15. Frequent. V. W. D., 15. Woods and fields. Crataegus tomentosa L. R. E. B., 02. r!. W. D., 15. Fields and road- Cra'taegus moll-is (T. & G.) Scheele. L. H. P., 98; 188, W. W. D., 15. Common. [. P., 15. Fragaria virginia.na Duche~ne var. illinoisensis (Prince) Gray. die. 176, W. W. D., 15. Fields 1~9, W. W. D., 15. Common. Fragaria vesca L. var. americana Porter. L. H. P., 98; 3964, !}. B., 02. L. H. P., R. E. B., C. M. K., 03; 190, W. W. D., 15. On I. H. P., 98; L. H. P., 0. R. B., rocky hillside. Potentilla arguta Pursh. R. E. B., 02.

Published by UNI ScholarWorks, 1915 17 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

94 IOWA ACADEMY OF SCIENCE FLORA OF Potentilla monspeliensis L. 292, W. W. D., 15. Fields. Desmodium canad.ense (L.) D Potentilla canadensis 'L. L. H. P., 02. Lespedeza oapitata Michx. ] Rubus idaeus var. aculeatissimus (C. A. Mey.) Regel & Tiling. Dry exposed slopes. hH.~~ - .Vioia oaroliniana Walt. L. I · Rubus occidentalis L. 191, W. W. D., 1'5. Frequent.. Vioia americana Muhl. L. I Rubus villosus Ait. 192, W. W. D., 15. Frequent. woods, common. Agrimonia striata Michx. 3956, L. H. P., R. E. B., C. M. K., 03. Lathyrus oohroleuous Hook. : Rosa pratirncola Greene. L. H. P., 98; 193, W. W. D., 15. Fre~ ' i L. H. ·P., R. M., 12; . quent. F. 2~ Strophostyles helvola (L.) Br Rosa blanda Ait. L. H. P., 98. .Amphioarpa monoica (L.) El Prunus serotirna Ehrh. L. H. P., G. R. B., 98; 194, W. W. D., 15. Not uncommon. GER Prunus virginiana L. 3923, L. H. P., R. E. B., C. M. K., 03; On 195, W. W ..D., 15. Common. Oxalis oornioulata L. R. E. 1 Pru,nus piennsylvanica L. L. H. P., 14. Rare. M. K., 03. Common. Prunus americana Marsh. L. H. P., 97; 3881, L. H. P., R., E. B., Oxalis vio,lacea L. L. H. P., 15 C. M. K.; 03; 196, W. W. D., 15. Common. Ger: LegUminosre. Geranium maoulatum L. 208 . Gymnocladus diolica (L.) Koch. 197, W. W .D., 15. Rare. .R1 Gleditsia triacanthos L. L. H. P., 00; 197, W. W. D., 15. Fre Zanthoxylum amerioanum Mi~ quent. Ba;ptisia bracteata (Muhl.) Ell. 135, J. V. E., 14. Pol3' Trifol~um pratense L. 198, W. W. D., 14. Escaped to fence Polygala senega L, L. H. P. rows. W. W. D., 15. Dry slopes Melilotus ojficinalis (L.) Lam. ·199, W. W. D., 14. Roadsides. Polygala sangu~nea L. R. E. Meluotus alba Desr. 200, W. W. D., 15. Roadsides and old C. M. K., 03. fields. Polygala vertioillata L. 3855, Medicago sativa L. 201, W. W. D., 15. Old field. Euph Amorpha canescens Pursh. 3991, L. H. P., R. E. B., C. M. K., Acalypha virginioa L. 211, "V 03; 202, W. W. D., 15. Euphorbia Preslii Guss. R. E. Petalostemum pu.rpu.reu.m (Vent.) Rydh.. R. E. B., 02 .. Euphorbia oorollata L. 3668, Petalostemum candidum Michx .. 3865, L. H. P., R. ,E. B., C. M. K., 03. SAP: Robinia pseudo-acacia L. 3901, L. H. P., R. E. B., C. M. K., 03. Anac Astragalus caryocarpus Ker. L. H. P., 00. Rhus glabra L. R. E. B., 02; .Astragalus canadensis L. R. E. B.,' 02; 3918, L. H. P., R. E. B., dry slopes. I C. M. K., 03; 203, W. W. D., 15. Exposed hillside. Rhus toxicodendron L. 22, J Desmodium grandiflorum (Michx.) DO. R. E. B., 02; ~04, W. Frequent. Cela 111,1 W. D., 15. Rich woods. II Desmodium Dillenii Darl. R. E. B,, 02. Evonymus atropurpureus J ac1 ,11''ii!1 1 14. Common. 'l.'11

1 Ii' ii.. !. '11 '''I https://scholarworks.uni.edu/pias/vol22/iss1/13 18 Diehl: The Flora of the Ledges Region of Boone County, Iowa

OF SCIENCE FLORA OF THE LEDGES 95

W. W. D., 15. Fields. Desmodium canadense (L.) DC. 52, J. V. E., 14. 02. Lespedeza capitata Michx. R. E. B., 02; 205, W. W. D., 15. (C. A. Mey.) Regel & T_g~g. Dry exposed slopes. Vicia caroZiniana Walt. L. H. P., R. H. C., 97. '. D., 1•5. Frequent. Vicia americana Muhl. L. H. P., 13; 206, W. W. D., 15. In D., 15. Frequent. woods, common. ~-H.P., R. E. B., C. M. K., 03. Lathyrus ochroleucus Hook. L. H.P., C.R. B., 98; L. H.P., 99; ., 98; 193, W. W. D., 15. Fre- L. H. P., R. F. M., 12; 207, W. W. D., 15. Dry ~lopes. Strophostyles helvola (L.) Britton. L. H. P., 15. Amphicarpa monoica (L.) Ell. 56, J. V. E., 14.

, C. R. B., 98; 194, W. W. D., GERANIALES. Oxalidacere. H. P., R. E. B., C. M. K., 03;. Oxalis corniculata L. ·R. E. B., 02; 3871, L. H.P., R. E. B., C. n. c,, M. K., 03. Common. P., 14. Rare. , P., 97; 388i, L. H~ P., R. E. B., Oxalis violapea L .. L. H. P., 15. , 15. Common. Geraniacere. nosre. Geranium maculatum L. 208, W. W. D., 15; Common. 97, W. W .D., 15. Rare. Rutacere • ., 00; 197, W. W. D., 15. Fre- Zanthoxylum americanum Mill. 209, W.W. D., 15. Frequent. I Polygalacere. 135, J. V. E., 14. Polygala senega L. L. H. P., C. R. B., 98; L. H. P., 02; 210, W. D., 14. Escaped to fence W.W. D., 15. Dry slopes. \, Polygala sangu~nea L. R. E. B., 02; 3957, L. H. P., R. E. B., ·199, W.W. D., 14. Roadsides. C. M. K., 03. . W. p., 15. Roadsides and old Polygala verticillata L. 3855, L. H. P., R. E. B., C. M. K., 03. Euphorbiacere. D., 15. Old field. 1, L. H. P., R. E. B., C. M. K., Acalypha virginica L. 211, W.W. D., 15. Fields, common. Euphorbia Preslii Guss. R. E. B., 02; 212, W. W. D., 15. Fields. t.) Rydb. R. E. B., 02. .. Euphorbia coroUata L. 3668, L. R. P., R. E. B., C. M. K., 03 . . 3865, L. H.P., R. E. B., C. M. SAPINDALES. Anacardiaicere. L. H.P., R. E. B., C. M; K., 03. Rhus glabra L. R. E. B., 02 ;. 213, W. W. D., 15. Common on . H.P., 00. dry slopes . . B.,' 02; 3918, L. H. P., R. E. B., Rhus toxicodendron L. 22, J. V. E., 14; 214, W. W. D., 14 . ., 15. Exposed hillside. . Frequent. hx.) DC. R. E. B., 02; ~04, W. Celastracere. Evonymus atropurpureus Jacq. R. E. B., 02; 215, W. W. D., . B., 02. 14. Common.

Published by UNI ScholarWorks, 1915 19 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

FLORA OF' 96 IOWA ACADEMY OF SCIENCE

MYR'. Celastrus scandens L. R. E. B., 02; J. P. A., 13; 216, W. W. D., Thym 14. On wooded slopes. 8,t.a.phylacere. Dirca palustris L. 4020, R. E. damp shaded' ledge; one st1 Staphyfoa trifolia L. 2900, L. H. P., R. E. B., C. M. K., 03; 217, W. W. D., 15: On damp shaded slopes. Onag Aceracere. Oenothera biMinis L. 3868, L. 1 W.W. D., 15. Fields. Acer saccharum Marsh. var. nigrum (Michx. f.) Britton. L. H. P., 98; R. E. B., 02; 218, W. W. D., 14. Common on slopes UMBEi and in upland woods. AJ.·aIJ Acer saccharinum L. 9, J. V. E., 14; 219, W. W. D., 15. Valley Aralia racemosa L. R. E. B., o: of Des Moines river. G. M. K., 03; 312, W. W. I Acer negundo L. 3885, R. E. B., 02; 220, W. W. D., 15. Along Aralia nudicaulis h L. H. P., Pease creek and Des Moines river. common .. Balsaminacere. Pana,x quinqnefoliwn L. 314, V

Impatiens pallida Nutt. 3939, L. H. P., R. E. B., C. M. K., 03; Umbel · 221, W. W. D., 15. In damp shaded locations. Sanicula marilandiica L. L. H. Impatiens bi/fora Walt. 61, J. V. E., 14. Chaerophyllum procumbens (L. RHAMNALES. Osmorhiza Cla.ytoni (Michx.) C Rhamnacere. K., 03; 227, W. W. D., 15. Rhamrnus lanceolata Pursh. L. H. P., 0. R. B., 98. Osmorhiza longistylis (Torr.) D Ceanothus americanus L. R. E. B., 02; 35, J. V. E., 14; 222, W. woods, infrequent. W. D., 15. Cicuta macitlata L. 3944, L. H. Vitacere. W.W. D., 14. Valley of D Psedera quirnquefolia L. 223, W.W. D., 15. Common. Crypto'taenia canadensis (L.) D Vitis vulpina L. L. H.P., 98; R. E. B., 02; 26, J.V. E., 14; R. E. B., C. M. K., 03. 224, W. W. D., 15. . Zizia aitrea (L.) Koch. L. H. J MALVALES. W.D., 15. Tiliacere. Zizia cordata (Walt.) DC. R. Tilia americana L. 225, W. W. D., 15. Frequent in fields. Taenidia integerrima (L.) Drud Abutilm Theophrasti Medic. 226, W. W. D., 15. Fields. P., 98; 3898, L. H.P., R. E Pastinaca sativa L. 231; W.W. VIOLALES. H eracleum lanatum Michx. L. · Cistacere.· • W.W. D., 15. Lechea villosa Ell. 3952, L. H.P., R. E. B., C. M. K., 03. Oorm Vfolacere. Corn.its circinata L 'Her. 3968, l Viola cucullata Ait. 3946, L. H.P., R. E. B., C. M. K., 03; 308, 232, W'l\ W. D., 15. Rocky : W. W. D., 15. Common. Cornus asperifolia Michx. 3886, Viola pubescens Ait. L. H. P., R. H. C., 97; 309, W. W. D., 233, W. W. D., 15. Hillside 15. Common. 7

https://scholarworks.uni.edu/pias/vol22/iss1/13 20 Diehl: The Flora of the Ledges Region of Boone County, Iowa

FLORA OF THE LEDGES '97 'OF SCIENCE MYRTALES. ~2; J. P. A., 13; 216, W. W. D., Thymelll:cere. Dirca palustris L. 4020, R. E. B., 02; 310, W. W. D., 15. On acere. damp shaded ledge; one station only. ' . [. P., R. E. B., C. M. K., 03; Onagraeere. p shaded slopes. · Oenothera biMinis L. 3868, L. H. P., R. E. B., C. M. K., 03; 311, ~ere. W. W. D., 15. Fields. µm (Michx. f.) Britton. L. H. UMBELLALES. , W~ D., 14. Common on slopes Araliacere. , 14; 219, W.W. D., 15. Valley Aralia racemosa L. R. E. B., 02; 2976, 2933, L. H. P., R. E. B., O'. M. K., 03; 312, W. W. D., 15. Damp woods. , 02; 220, W. W. D., 15. Along Ara.lia n1tdicaulis I.I. L. H.P., 13; 313, W.W. D., 15. Woods, river. common .. nacero. Panax quinqicefolium L. 314, W.W. D., 15. Very rare. 1. H. P., R. E. B., C. M. K., 03; Umbelliferre• .p shaded locations. Sanicula marilandica L. L. H. P., C. R. B., 98; L. H. P., 13. ·. E., 14. Chaerophyllum procumbens (L.) Crantz. L. H. P., 98, 00. rALES. Osmorhiza Claytoni (Michx.) Clarke. L. H. P., R. E. B., C. M. iacere. K., 03; 227, W.W. D., 15. In woods, :frequent. H. P., C. R. B., 98. Osmorhiza lo·ngistylis (Torr.) DC. 228, W.W. D., 15. In damp B., 02; 35, J. V. E., 14; 222, W. woods, infrequent. Cicuta macitlata L. 3944, L. H. P., R. E. B., C. :M:. K., 03; 229, cere. W.W. D., 14. Valley of Des Moines river. v. W. D., 15. Common. Crypto'taenia canadensis (L.) DC. L. H. P., 98; 3932, L. H. P., R. E. B., 02; 26, J. V. E., 14; R. E. B., C. M. K., 03. Zizia aitrea (L.) Koch. L. H. P., 98; 23, J. V. E., 14; 230, W.

'ALES. W.D., 15. Zizia cordata (Walt.) DC. R. E. B., 02. Taenidia integerrima (L.) Drude. L. H.P., R. H. C., 97; L. H. D., 15. Frequent in fields. P., 98; 3898, L. H. P., R. E. B., C. M. K., 03. 26, W. W. D., 15. Fields. Pastinaca sa.tiva L. 231, W.W. D., 15. Roadside. ALES. Heracleum lanatum Michx. , L. H. P., 13; 70, J. V. E., 14; :nfi, ~cere • W.W. D., 15. .P., R. E. B., C. M. K., 03. Oornacere. acere. Cornus circinata L 'Her. 3968, L. H. P., R. E. B., C. M. K., 0:1; [. P., R. E. B., C. M. K., 03; 308, 232, ·WI\ W. D., 15. Rocky hillsides, infrequent. Cornus asperifolia Michx. 3886, L. H.P., R. E. B., C. M~ K., oa; ., R. H. C., 97; 309, W. W. D., 233, W. W. D., 15. Hillsides . 7

Published by UNI ScholarWorks, 1915 21 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

98 IOWA ACADEMY OF SCIENCE FLORA OF

Corni/,S amomitm Mill. R. E. B., 02; 3932, L. H. P., R. E. B., C. M. K., 03; 234, W. W. D., 15 .. Corn.us alternifolia L. f. 3961, L. H.P., R. E. B., C. M. K., 03; L. H. P., J. P. A., 13; 235, W. W. D., 15. Frequent. '\

ERIC ALES.

Ericacem. Monotropa itnifiora L. L. H. P., 15.

PRIMULALES. Pri:m.uJ.acere. · Steironema ciiiatilm (L.) Raf. 3907, L. H. P., R. E. B., C. M. K., 03. Bor1 GENTIAN ALES. Cynoglossitm officinale L. L. Oleacere. Cynoglossum boreale Fernald. Luppula vir[/iniana (L.) Gre1 Fraxinus americana L. 236, W. W. D., 15. Uplands and wooded \ M. K., 03; 246, W. W. D., hillsides, common. Symphy'tum officinµle L. L. Fraxinus pennsyfoanica Marsh. 37, J. V. E., 15; 237, W.W. D., creek. 15. Infrequ()nt. Mertensia virginica (L.) Link Fraxinits pennsylvanica Marsh. var. lanceolata (Borkh.) Sarg. D., 15. Valley of Pease 1 L. H.P., 15. Lithospermum latifolium Mi, F'raxinus nigra Marsh. L. H. P., 98; 3895, L. H. P., R. E. B., M. K., 03. C. M. K., 03; L. H. P., 13; 238, W. W .. D., 15. Frequent in Lithospermum ca;nescens (Mic ravines and in valley of Pease creek. Onosriiodium occidentale Mac Apocynacere. p.osed hillsides. · Verl Apocynum androsaem·ifolium L. 239, W. W. D., 15. On dry wooded hillsides. · Verbena urticaefolia L. 3920 Verbena hastata L. 3872 L. : Apocyn1tm cannabinum L. L. H. P., 14. . . - . ' J. V. E., 14. Asclepiadacere. Verbena stricta Vent. 290 W ' ' ' Asclepias incarnata L. 69, J. V. E., 14. Verbena bracteosa M1chx. 38E Asclepias syriaca L. 240, W. W. D., 15. 317, W.W. D., 15. Asclepias verticillata L. L. H. P., 15. liippia lanceolata Michx. L. ]

POLEMONIALES. L~

Convolvulacere. Teucrium canadense L. 3971 L. H. P., J. P. A.; 39, ,' Convolvitlits sepiitm L. 241, W. W. D., 15. In fiel&; along Des Shaded hillsides. Moines river. Agastache nepetoides (L.) Kt lpomoea hederacea Jacq. L. H.P., 15. in valley of Pease creek.

https://scholarworks.uni.edu/pias/vol22/iss1/13 22

------'------Diehl: The Flora of the Ledges Region of Boone County, Iowa

FLORA OF THE LEDGE'S 99 y OF SCIENCE Polemoniacere. 02 · 3932 L. H. P., R. E. B., ., ' ' Phlox divaricata L. J. V. E., 14; 242, W. W. D., .15. Gommon ., 15.· L . H.P., R. E. B., 0. M. K., 03; . in woods . W. D., 15. Frequent. Phlox pilosa L. L. H. P., 00; 243, W. W. D., 15. On dry slopes, frequent. ALES. Hydrophyllacere.

cem. Hydrrophyllum appendiculatum Michx. 3885, L. H.P., R. E. B., 0. M. K., 03 . ., 15. Hydrophyllum virginianurn L. 244, W. W. D., 15. Common in LES. woods. nJ.acere. · Ellisia nyctelea L. 245, W. W. D., 15. Damp woods, common .. 3907, L. H.P., R. E. B., O. M. Boraginacere. Cynoglossum officinale L. L. H. P., 15. IAN ALES. Cynoglossum bo'f"eale Fernald. L. H. P., 15. eacere. Luppula virrf'im,iana (L.) Greene. 3921, L. H. P., R. E. B., .C. 15 Uplands ~nd wooded '~ <\.-: M. K., 03; 246, W. W. D., 15. Woods, common. ' v.W.D., •· I Symphy'tum officinale L. L. H. P., 15. Near mouth of Pease i. 37, J. V. E., 15; 237, W.W. D., creek. Mertensia virginica (L.) Link. L. H.P., E. Bissel; 247, W.W. ih. var. lanceolata (Borkh.) Sarg. D., 15. Valley of Pease creek. Lithospermum latifolium Michx. 3859, L. H. P., R. E. B., C. · p 98 · 3895 L. H. P., R. E. B., M. K., 03. B; 2s8, W. w:' D., 15. Frequent in Lithospermum canescens (Michx.) Lehm. 19, J. V. E., 14. · Pease creek. Onosniodium occidentale Mackenzie. 299, W. W. D , lh. Ex p.osed hillsides. :>cynacere. Verbenacere. , L. 239, W. W. D., 15. On dry Verbena urticaefolia L. 3920, L. H.P., R. E. B., 0. M. K., D3. Verbena hastcita L. 3872, L. H.P., R. E. B., 0. M. K., 03; 155, L. H.P., 14. J. V. E., 14. lepiadacere. Verbena stricta Vent. 290, W.W. D., 15. Pastures. Verbena bracfeosa 1\1'.ichx. 3882, L. H. P., R. E. B., 0. M. K., 03; . V. E., 14. 317, W.W. D., 15. ·. W.D., 15. Lippia lanceola.ta Michx. L. H. P., 13. H.P.,15. . Labiatre. ~EMONIALES. Teucrium canadense L. 3971, L. H. P., R. E. B., 0. M. K., 03; 1volvulacere. L. H. P., J. P. A.; 39, J. V. E., 14; 248, W. W. D., 15. W. W. D., 15. In field:$ along Des Shaded hillsides. Agastache nepetoides (L.) Ktze. 249, W. W. D., 15. Gommon .:·,. . H.P.,15. in valley of Pease creek .

Published by UNI ScholarWorks, 1915 23 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

100 IOWA ACADEMY OF SCIENCE FLORA 0 Nepeta cataria L. 3893, L. H. P., R. E. B., C. M. K., 03; 250,. Galium circaezans Michx. 2! W.W. D., 15. About abandoned pwellings ...· . . Galium boreale L. 3969, L; : "itepeta hederacea (L.) Trevisan .. L. H. P., 13; 288, W. W. D:, P., 13; 274, W.W. D., 1 15. In pastured ravine_. Gali¥m boreale L; 3969, L. : Prunella vulgaris L. 3862, L. H. P., R. E. B., C.. M. K., 03; 251 · P., 13; 275, W. W. D., 11 W.W. :D., 15. Galium trifidu.m· L; 3916, L. Stachys palustris L, L. H. P., 98; 3931, L. H. P., R. E. B., C. W.W. D., 15. M,K., 03. Oap· Monarda fistulosa L. R. E. B., 02; 252, W. W. D., 14. Common Lonicera aiocica L. L. H.P. along Pease creek. ~iburnum pubescens (Ait.) Hedeoma hispida Pursh. L. H.P., 99, 00. B., 02; 25, J. V. E., 14; ~ Pycnanthemum virginia1Jiitm (L.) Durand & Jackson. L. H.P., Sambucus canadensis L. J. I 15. 15. Mentha spicata L. 253, W.W. D., 15. ·· OAMJ Cuc1 Solanacere. Sicyos angulatus L. 26, L. I Solanum carolinense L .. L. H. P~, C.R. B., 97. 15. Des Moines valley, ci Solanum nigrum L. L. H. P., C. R. :J?., 97; 270, W. W. D., 14. Campanula americana L. 39i Physalis subglabrata (L.) Mackenzie & Bush. R. E. B., 97. Physa.zis pruinosa L. 306, W. W. D., 15. Des Moines river Lol valley. Lobelia spicata Lam. 3880, I Lycium halimifolium J':fill. 254, W. W. D., 15. Escaped fro:m J. V. E., 14. door-yard. · · .. Lobelia siphilitica L. R. E. B Datura stramonium L. R. E. B., 02; 271, W. W. D:, 14. Om Scrophulariacere. Vernonia f ascwulata Michx. l Verbascitm thapsus L. L. H. P., 02; 13, J. V. E., 14. woodS~ Scrophularia leporella. Bicknell. 296, W. W. D., .14. Woods, Eupatorium urticaefolium Re frequent. common. Chelone glabra L. L. H.P., 15. . Eupaforium perfoliatiim L. J Mim'.ulus ringens L. 3875, L. H. P., R. E. B., C. M. K., 03. Eupatoriu.m purpureum L. vai Veronica peregrina L. ?95, W, W, D,, 15. 15. In damp woods. Pedicularis canadensis L. L. H.P., 95;•J. V. E., 14; 293, W.W. Kuhnia eupatoroides L. 258, D., 15. Rocky hillsides, etc. sides. ' Phrymacere. Liatris pycnostachya Michx. : Solidago latifolia L. 28, 29, J, Phryma leptostachya L. L. H.P., R. E. B., 0. M. K., 03. Solidago ulmifolia Muhl. . R .. l . Plantaginacere. C. M. K., 03; 260, W. W. Plantago major L. J. V. E., 14; 255, W. W. D., 15.. Solidago radula Nutt. R. E. E Rubiacere. Solidago canadensis L. 261, V Galium apa:rine L. · 3947, L. H. P., R. E. B., C: M. K., 03; 272, #dlidago altis~ma L. 262,: W. W.W. D., 15. Solidago rigida L .. R. E. B., O

https://scholarworks.uni.edu/pias/vol22/iss1/13 24 Diehl: The Flora of the Ledges Region of Boone County, Iowa

Y OF.SCIENCE FLORA OF THE LEDGES .·101

P., R. E. B., C. M. K., 03; 250,. Galium circaezans Michx. 24, J V E., 14; 273, W. W. D., 15. · loned dwellings. Galium boreale L. 3969, L; H; P., R. E. B., C. M. K., 03; L. H. .. L. :Er. :P., 13; 288, w. w. D.', P., 13; 274, W. W. D., 15. G.ali'f'm boreale L; 3969, L. H. P., R. E. B., C. M. K., 03; L. H. :. P., R. E. B., C. M. K., 03; 251 P., 13; 275, W. W. D., 15. .Galium trifidum·L. 3916, L. H.P., R. E. B., C. M.·K., 03; 276, 98; 3931, L. H. P., R. E. B., C. W.W. D., 15. Oaprifolia.cere. )2; 252, W.W. D., 14. Common Lonicera dJiocica L. L. H. P., R. H. C., 97; 277, W. W. D.; 15. 'V_iburnum pubescens (Ait.) Pursh. L. H. P., 98; 3965, R. E. .P., 99, 00; B., 02; 25, J. V. E., 14; 278, W. W. D., 14. .) Durand & Jackson.' L. H.P., Sambucus canadensis L. J. P. A., L. H. P., 13; 279, W. W;, D., 15. ·· CAMPANULALES. D., 15. Oucurbitacere. iacere. Sicyos angulatus L. 26, L. H. P., 0. R. B., 96; 256, W. W. D., ?~, c. R. B., 97. \ 15. Des Moines valley, common. C. R. ~., 97; 270, W. W. D., 14. Oampanula americana L. 3950, L. H. P., R. E. B., 0. M. K., 03. enzie & Bush. R. E. B., 97. W. D., 15. Des Moines river Lobeliacere. Lobelia spicata Lam. 3880, L. H. P., R. E. B., C. M. K., 03; 21, 1, W. W. D., 15. Escaped :from J. V. E., 14. Lobelia sipkilitica L. R. E. B., 02. :., 02; 211, w. w. D:, 14. Compositre. 1lariacere. Vernonia fasciculata Michx. 257, W.W. D., 15. In open upland •., 02; 13, J. V. E., 14. woods: · 1. 296, W. W. D., 14. Woods, Eupatorium urticaefolium Reichard. L. H. P., 15.. In woods, common.

). Eupatorium perfoliatum L. L. H. P., 15. Along Pease creek. I. P., R. E. B., C. M. K., 03. Eitpatorium purp·ureum L. var. maculatum (L.) Dari. L. H.P., , W, D,, 15. 15. In damp woods. LP., 95;•J. V. E., 14; 293, W.W. Kuknia eupatoroides L. 258, W. W. D., 14. Dry, exposed hill- c. sides. ' ·· · rmacere. Liatris pycnostackya Michx. L. H. P., 15. Dry hillside. Solidago latifolia L. 28, 29, J. V. E., 14; 259, W. W. D., 14. :. P., R. E. B., C. M. K., 03. Solidago ulmifolia Muhl. . R.. E. B., 02; 3894, L. H. P., R; E. B., lginacere. 0. M. K., 03; 260, W. W. D., 14. Dry slopes. 4; 255, W. W. D., 15. Solidago radula Nutt. R. E. K, 02. 11biacere. Solidago canadensis L. 261, W.W. D., 15. Wooded hillsides; I. P., R. E. B., 0: M. K., 03; 272, Bolidago altiSsiina L. 262,: W. w. D., 15. Wooded slopes. Solidago rigida L. R. E. B., 02.

Published by UNI ScholarWorks, 1915 25 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

102 IOWA ACADE"fy[Y OF SCIENCE FLORA OF

Boit<.mia asteroides (L.) L'Her. L. H. P., 15. Matricaria suaveolens (Pursh.) Aster cordifoUus L. 48, L. H. P., C. R. B., 96; 263, W. W. D., dwelling at head of Pease , 14. Wooded slopes. Tanacetum vulgare L. L. H. I Aster sagittifolius Wedemeyer. R. E. B.,.02; 3967, L. It,. P., Oacalia tuberosa Nutt. h H. 1 ,R. E. B., C. M. K., 03; 264, W. W. D., 15. Senecio balsamifera Muhl. L. Erlgeron philadelphicus L. ;L. H. P., 02; 12, J. V. E., 14; 265, spaces. W.W. D., 15. Common. Arctium minor Bernh. 63, J. Erigeron anmuus (L.) Pers. L. H.P., R.H. C., 97; 3864, L. H. Roadsides. . P., R. E. B., c. M. K., 03; 307, w. w. D., 15. Fields and Oirsium discolor (MuhL) "Spre1 . exposed slopes. Cirsium lanceola.tum (L.) Hil Erigeron canadensis L. L. H.P., 02; 12, J. V. E., 14. creek. Antenmaria plantaginifolia (L.) Richards. 266, W. W. D., 15. Oirsium ioense (Pammel) Fe1 Common on hillsides. . p.Iaces. Silphium perfoliatum. L. 3910, L. H. P., R. E. B., C. M. K., 03. Taraxacum officinale Weber. 2 Silphium lacin.iatum L. 298, W. W. D. Dry exposed hillsides. roadsides, common. , Infrequent. Sonchus oleraceus L. 284, W. AmbroS'ia trifida L. 267, W. W. D., 14. River valley, etc., tures. .. cm;nmon. Lactuca canadensis L. 43, J. V Ambrosia artemisiifolia L. 267, W. W. D., 14. Upland fields, Lactuca scariola Ii. L. H. P., J com;m.on. Lactuca fioridana (Lam.) Hite: Ambrosia psilostachya DC. L. H.P., 15. frequent. Xanthium canadense Mill. R. E. B., 02; 268, W. W. D., 15. Prenanthes alba L. R. E. B., ( Fields, common. D., 14. Common on woode1 Heliopsis scabra Dunal. 3913, L. H. P., R. E. B., C. M. K., 03; Key to initials used in the 02 ·· 268, W. W. D., 15. Frequent. J.P. A ...... Rudbeckia hirta L. 3948, L. H.P., R. E. B., C. M. K., 03; 269, C._R. B ...... W. W.D., 15. R. E. B ...... Helianthus grosseseratus Martens. L. H. P., 15. R.H. C...... Helianthus strumosus L. L. H. P., 15. W.W. D .. ~ ...... Helianthus tuberosus L. L. H. P., C. R. B., 96; R. E. B., 02; 49, J.V.E...... J. V. E., 14; 270, W. W. D., 15. C. M. K ...... Ooreopsis palma.ta Nutt.· 3.866, L. H. P., R. E. B., C. M. K., 03; R.F.M...... L. H.P., fa. . L.H.P...... Bidens dJiscoidea (T. & G.) Britton. L. H. P., 15. Acknowledgment is grate.full; Bidens frondosa L. L. H. P., 15. under whos·e direction this stud BidJens cernum L. L. H. P., C. R. B., 98; 280, W. W. D., 15. in the work and for help in d1 Valley o.f Pease creek.. · Thanks are due also to the m Helenium autumna-le L. L. H. P., 15. partment of Botany of Iowa Sta Dyssodia papposa (Vent.) Hitchc. L. H. P., 15. in effecting the work. Acknow Achillea Millefolium L. L. H. P.,.13; 281, W.W." D., 15. Valley Cratty of Armstrong, Iowa, for of Pease creek. of Carex. ·

https://scholarworks.uni.edu/pias/vol22/iss1/13 26 Diehl: The Flora of the Ledges Region of Boone County, Iowa

OF SCIENCE FLORA OF THE LEDGES

J. H.P., 15. Matricaria suaveolens (Pursh.) Buchenau. L. H. P., 15. About C. R. B., 96; 263, W. W. D., dwelling at head of Pease creek. Tanacetum vulgare L. L. H. P.; J. ·P, A., 13; :t. E. B., 02; 3967, L. I!· P., Cacalia tuberosa Nutt. L. H.P., 15. Fietds an,d open spaces~· W. W.D.,15 . Senecio balsamifera Muhl. L. H. P., 15. Hillsides and ope~ . P., 02; 12, J. V. E., 14; 2.65, spaces. Arctium minor Bernh. 63, J. V. E., 14; 282, W. W. ,D., 15. :. P., R. H. C., 97; 3864, L. H. Roadsides. )7, w. W. D., 15. Fields arid Cirsium discolor (Muhl:) 'Spreng. R. E .. B., 02. Cirsium lanceolatum (L.) Hill. L. H. P., 15. Along Pease 02; 12, J. V. E., 14. creek. Richards. 266, W. W. D., 15. Cirsium ioense (Pammel) Fernald. L.' H. P., 15. In open places. . H. P., R. E. .B., C. MK. ., 03 . Taraxacum ojficinale Weber. 283, W.W. D., 15. Pastures and W. D. · Dry exposed hillsides. roadsides, common. Sonchus oleraceus L. 284, W. W. D., 15. Roadsides and pas- 'V. D., 14. River valley, etc., tures. Lactuca canadensis L. 43, J. V. E., 14. W. W. D., 14. Upland fields, Lactuca scariola ]j, L. H. P., 15. Along Pease creek. Lactuca ftoridana (Lam.) Hitch. 297, W. W. D., 15. Woods, :. P., 15. frequent. · !1. B., 02; 268, W. W. D., 15. Prenanthes alba L. R. E. B., 02; 47, J. V. E., 14; 285, W. W. D., 14. Gommon on wooded hillsides. . H. P., R. E. B., C. M. K., 03; Key to initials used in the Catalogue : it. J. P. A ...... J. P. Anderson P., R. E. B., G. M. K., 03; 269, C. R. B...... C. R. Ball R. E. B...... '. R. E. Buchanan s. L. H. P., 15. R. H. C...... R. H. Combs P., 15. W. W. D...... W. W. Diehl •., C. R. B., 96; R. E. B., 02; 49, J. V. E ...... J. V. Ellis 15. C. M. K...... Miss C. M. King :i. H. P., R. E. B., C. M. K., 03; R. F. M...... R. F. Miller L. H~ P ...... L. H. Pammel. con. L. H. P., 15. Acknowledgment is gratefully made to Dr. L. H. Pammel, 5. under whose direction this study was made, for aid and advice . R. B., 98; 280, W. W. D., 15. in the work and for help in determining many of the plants. Thanks are due also to the members of the staff of the De P., 15. partment of Botany of Iowa State College for many suggestions he. L. H. P., 15. in effecting the work. Acknowledgment is made to Mr. R. I. '.,.l3; 281, w. w. D., 15. Valley Cratty of Armstrong, Iowa, for determining the various species of Carex. ·

Published by UNI ScholarWorks, 1915 27 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

104 IOWA ACADE'MY OF. SCIENCE

BIBLIOGRAPHY. 1. Beyer, S. W. Geology of Boone County, Ia. Geol. Surv. V: 187- 190. f. 16, 17. 1895. 2. Hitchcock, A. S. A Catalogue of the Anthophyta and Pterido phyta of Ames, Iowa. St. Louis Acad. Sci. 5: 447-532. f. 1891. (Reprint.) 3. Miller, K. A. The Lichens of the Ledges, Boone County, Iowa. Proc. .Ia. Acad. Sci. II: 139-146. 1904. 4. Pammel, L. H. Character and Distribution of Forest Trees and Shrubs in Boone County, Iowa. Ia. Ge.ol. ·surv. V: 232-239. 1895. 5. Pammel, L. H. A Coµiparative Study of the Vegetation of Swamp, Clay, and Sandstone Are.as in Western ·Wisconsin, Southeastern Minnesota, Northeastern Minnesota, . Central and Southeastern Iowa. Proc. Ia. Acad. Sci. 10: 106-111. f. 5, 33, 34, 42. 1905. 6. Pammel, L. Ii., and King, •O. M. The 'vascular Cryptogams of Iowa and Adjoining Parts of Sovtheastern Minnesota and Western Wisconsin. Proc. Ia. Acad. Sci. 9: 134-150. f. 2-6. pl. 10-31. 1902. 7. Pammel, L. H., Weems, J. B., and Lamson-Scribner, F. The Grasses of Iowa. Ia. Geo!/ Surv, Bull. 1: 1-525. f. 1-220. 1901. 8. Robinson, L. B., :and Fernald, M. L. A Handbook of the Flower ing Plants and Ferns, etc. 1-926. 1908. BOTANICAL LABORATORY, IowA STATE COLLEGE.

North slo1

https://scholarworks.uni.edu/pias/vol22/iss1/13 28 Diehl: The Flora of the Ledges Region of Boone County, Iowa

~ y OF SCIENCE iRAPHY. o.e County, Ia. Geol. Surv. V: 187- e of the Anthophyta and Pterido Louis Acad. Sci. 5: 447-532. f. PLA'£E XVIII. the Ledges, Boone County, Iowa. -146. 1904. l Distribution of Forest Trees and owa. Ia. Geol. ·surv. V: 232-239 .

.Hve Study of the Vegetation of one Are.as in Western ·Wisconsin, Northeastern Minnesota, . Central Proc. Ia. Acad. Sci. 10: 106-111.

M:. The Vascufar Cryptogams of s of 1Sout'heastern Minnesota and Ia. Acad. Sci. 9: 134-150. f. 2-6.

3., ~nd Lamson-Scribner, F. The 11,I Surv, Bull. 1: 1-525. f. 1-220.

M. L. A Handbook of the Flower- 1-926. 1908.

' •• j

Nor th slope of the strip survey,

Published by UNI ScholarWorks, 1915 29 Proceedings of the Iowa Academy of Science, Vol. 22 [1915], No. 1, Art. 13

PLATE XIX.

• South slope of the strip su rvey.

https://scholarworks.uni.edu/pias/vol22/iss1/13 30