The Science of Spring-Flowering Bulbs
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Roozengaarde's Bulb Planting Guide
15867 Beaver Marsh Road Mount Vernon, WA 98273 Frequently 360-424-8531 or 866-4TULIPS Asked www.Tulips.com / [email protected] Questions WILL BULBS GROW ANYWHERE? Yes, bulbs will grow in many different climates. Warmer climates are more challenging and may require pre-cooling each fall prior to plant- ing. The year we ship your bulbs, we do any necessary pre-cooling here on our farm prior to shipment—your bulbs will be ready to plant when you receive them! DO BULBS NEED TO BE WATERED? Yes, if Mother Nature does not take care of it for you, then water during dry spells. Be sure not to oversoak the planting area. Water just enough so that it absorbs quickly. Water after planting, during growth, and even after topping if the soil is dry. WHAT IS THE BEST TYPE OF SOIL AND FERTILIZER? Soil needs to provide adequate drainage and oxygen for the bulbs. We recommend using a slow release fertilizer specifically designed for flower bulbs - usually a N - P - K formula. DO BULBS NEED TO BE DUG IN THE SUMMER? We recommend digging tulip bulbs each year after the foliage has died down naturally. They are more prone to disease and rot if left in the ground through the summer months. Other bulbs like daffodils can typically be dug on a 3-5 year schedule. If you experience diminished results in the spring, you should dig your bulbs that summer. WHAT IS THE BEST WAY TO STORE MY BULBS IF I DIG THEM? After digging, make sure to dry bulbs thoroughly. -
Bulb Dormancy in Vitro—Fritillaria Meleagris: Initiation, Release and Physiological Parameters
plants Review Bulb Dormancy In Vitro—Fritillaria meleagris: Initiation, Release and Physiological Parameters Marija Markovi´c*, Milana Trifunovi´cMomˇcilov , Branka Uzelac , Sladana¯ Jevremovi´c and Angelina Suboti´c Department of Plant Physiology, Institute for Biological Research “Siniša Stankovi´c“—NationalInstitute of the Republic of Serbia, University of Belgrade, Bulevar Despota Stefana 142, 11060 Belgrade, Serbia; [email protected] (M.T.M.); [email protected] (B.U.); [email protected] (S.J.); [email protected] (A.S.) * Correspondence: [email protected] Abstract: In ornamental geophytes, conventional vegetative propagation is not economically feasible due to very slow development and ineffective methods. It can take several years until a new plant is formed and commercial profitability is achieved. Therefore, micropropagation techniques have been developed to increase the multiplication rate and thus shorten the multiplication and regeneration period. The majority of these techniques rely on the formation of new bulbs and their sprouting. Dormancy is one of the main limiting factors to speed up multiplication in vitro. Bulbous species have a period of bulb dormancy which enables them to survive unfavorable natural conditions. Bulbs grown in vitro also exhibit dormancy, which has to be overcome in order to allow sprouting of bulbs in the next vegetation period. During the period of dormancy, numerous physiological processes occur, many of which have not been elucidated yet. Understanding the process of dormancy will allow us to speed up and improve breeding of geophytes and thereby achieve economic profitability, which is very important for horticulture. This review focuses on recent findings in the area of Citation: Markovi´c,M.; Momˇcilov, bulb dormancy initiation and release in fritillaries, with particular emphasis on the effect of plant M.T.; Uzelac, B.; Jevremovi´c,S.; growth regulators and low-temperature pretreatment on dormancy release in relation to induction of Suboti´c,A. -
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE LILIACEAE de Jussieu 1789 (Lily Family) (also see AGAVACEAE, ALLIACEAE, ALSTROEMERIACEAE, AMARYLLIDACEAE, ASPARAGACEAE, COLCHICACEAE, HEMEROCALLIDACEAE, HOSTACEAE, HYACINTHACEAE, HYPOXIDACEAE, MELANTHIACEAE, NARTHECIACEAE, RUSCACEAE, SMILACACEAE, THEMIDACEAE, TOFIELDIACEAE) As here interpreted narrowly, the Liliaceae constitutes about 11 genera and 550 species, of the Northern Hemisphere. There has been much recent investigation and re-interpretation of evidence regarding the upper-level taxonomy of the Liliales, with strong suggestions that the broad Liliaceae recognized by Cronquist (1981) is artificial and polyphyletic. Cronquist (1993) himself concurs, at least to a degree: "we still await a comprehensive reorganization of the lilies into several families more comparable to other recognized families of angiosperms." Dahlgren & Clifford (1982) and Dahlgren, Clifford, & Yeo (1985) synthesized an early phase in the modern revolution of monocot taxonomy. Since then, additional research, especially molecular (Duvall et al. 1993, Chase et al. 1993, Bogler & Simpson 1995, and many others), has strongly validated the general lines (and many details) of Dahlgren's arrangement. The most recent synthesis (Kubitzki 1998a) is followed as the basis for familial and generic taxonomy of the lilies and their relatives (see summary below). References: Angiosperm Phylogeny Group (1998, 2003); Tamura in Kubitzki (1998a). Our “liliaceous” genera (members of orders placed in the Lilianae) are therefore divided as shown below, largely following Kubitzki (1998a) and some more recent molecular analyses. ALISMATALES TOFIELDIACEAE: Pleea, Tofieldia. LILIALES ALSTROEMERIACEAE: Alstroemeria COLCHICACEAE: Colchicum, Uvularia. LILIACEAE: Clintonia, Erythronium, Lilium, Medeola, Prosartes, Streptopus, Tricyrtis, Tulipa. MELANTHIACEAE: Amianthium, Anticlea, Chamaelirium, Helonias, Melanthium, Schoenocaulon, Stenanthium, Veratrum, Toxicoscordion, Trillium, Xerophyllum, Zigadenus. -
Nematode, Ditylenchus, Stem and Bulb, Meloidogyne, Root Knot
BIOLOGY AND CONTROL OF STEM AND ROOT KNOT NEMATODES r Becky B. Westerdahl1 Abstract: Plant parasitic nematodes are nonsegmented-microscopic roundworms which are frequently present in alfalfa fields. Although more than 10 different genera have been found in alfalfa fields in California, two (stem and bulb, and root knot) are most commonly associated with damage. A management plan to fit a particular growing situation should be developed using a combination of techniques including: planting site selection, certified seed, clean equipment, weed and irrigation management, resistant varieties, crop rotation, fallow, organic amendments and chemical nematicides. Ke~words nematode, Ditylenchus, stem and bulb, Meloidogyne, root knot, INTRODUCTION Plant parasitic nematodes are nonsegmented-microscopic roundworms which are frequently present in alfalfa fields. Whether or not alfalfa is to be planted in a nematode infested area, a grower should be knowledgeable about nematodes. If nematodes are present, both pre and postplant management strategies should be developed for pathogenic species. If an alfalfa field or a potential planting site is not infested, a grower should be aware of techniques available to prevent the introduction of harmful species. For growers to carry on a nematode pest management program they need to be familiar with (1) nematode biology; (2) symptoms and signs of nematode f damage; (3) how nematodes injure plants; (4) how to sample for nematodes; and (5) the principles underlying various management techniques including: planting site selection, the use of certified seed, the importance of using clean equipment and irrigation water, weed management, the use of resistant varieties, crop rotation, fallow, organic amendments, and chemical nematicides. -
Thermonastic TROPISM
THERMOnastic TROPISM Eleni Katrini | Ruchie Kothari | Mugdha Mokashi Bio_Logic Lab School of Architecture, Carnegie Mellon University ABSTRACT Motivation: To develop dynamic elements which respond to changes in temperature Approach: The thermonastic movements of the Tulip flower were looked at for inspiration to develop conceptual systems based on movement and heat. The tulip flower opens and closes its petals based on the external temperature. The petals open up when the temperature is high and close when the temperature is low. This movement of the petals is facilitated by movement of water through the plant. Experiments were conducted using paper which is a fibrous material exhibiting properties similar to that of a flower petal. Water at different temperatures was studied as a conducting medium. Findings: Based on analysis, different forms of flowers were created. This includes flowers with varying surface areas, flowers made using composite materials, laminated and chiseled flowers, flowers with varying edge exposure etc. The study was focused on exploring the effects of capillary action on fibrous materials. The observations were focused on- - Range of movement of petals - Duration for capillary action - Effect of hot and cold temperature It was seen that the range of movement was the same for most of the prototypes. All the flowers other than the flowers for which the range of movement was restricted opened up to 180o. The time taken for the water to rise depended on the density and the surface area of the paper in contact with the water. Increase in density of the paper in contact with water, increases the time taken for the water to rise up. -
Tyler Schmidt, Plant Science Major, Department of Horticultural Science
Interspecific Breeding for Warm-Winter Tolerance in Tulipa gesneriana L. Tyler Schmidt, Plant Science Major, Department oF Horticultural Science 19 December 2015 EXECUTIVE SUMMARY Focus on breeding of Tulipa gesneriana has largely concentrated on appearance. Through interspecific breeding with more warm-tolerant species, tolerance of warm winters could be introduced into the species, decreasing dormancy requirements and expanding the range of tulips southward. Additionally, long-lasting foliage can be favored in breeding to allow plants to store more energy for daughter bulbs. Continued virus and fungal resistance breeding will decrease infection. Primary benefits are for gardeners and landscapers who, under the current planting schedule, are planting tulip bulbs annually, wasting money. Producers benefit from this by reducing cooling times, saving energy, greenhouse space, and tulip bulbs lost to diseases in coolers. UNIVERSITY OF MINNESOTA AQUAPONICS: REPORT TITLE 1 I. INTRODUCTION A. Study species Tulips (Tulip gesneriana L.) are one of the most historically significant and well-known horticultural crops in the world. Since entering Europe via Constantinople in the mid-sixteenth century, the Dutch tulip market became one of the first “economic bubbles” of modern civilization, creating and destroying fortunes in four brief years (Lesnaw and Ghabrial, 2000). Since this time, tulips have remained extremely popular as more improved cultivars are released. However, a problem remains: even though viral resistance and long-lasting cultivars are introduced, few are capable of surviving in a climate with truly mild winters and only select cultivars are able to store enough energy for another year of flowering, even in climates with colder winters. Current planting schemes suggest planting annually, wasting tulip bulbs (Dickey, 1954). -
Bulbs: Culture and Maintenance by Diane Relf and Elizabeth Ball, Revised by Joyce Latimer, Virginia Cooperative Extension
GARDENING FACT SHEET Harris County Cooperative Extension 3033 Bear Creek Drive, Houston, Texas 77084 281.855.5600 • http://harris-tx.tamu.edu/hort Bulbs Culture and Maintenance Originally published in the Arizona Master Gardener Manual, produced by the Cooperative Extension, College of Agriculture, The University of Arizona, 1998. Edited and reformatted by Texas Cooperative Extension, Harris County, September 2007 he term bulb is loosely used to include corms, tubers, tuberous roots, and rhizomes, as well as true bulbs. This publication will T refer to all of the above as bulbs. leaves A true bulb is a complete or nearly complete miniature of a plant stem encased in fleshy modified leaves called scales which contain reserves of food. Corms are the base of a stem that becomes swollen and solid with nutrients. It has no fleshy scales. The lateral bud tuber, which is an underground stem that stores food, differs from the true bulb internode Bulb or corm in that it has no covering of dry node leaves and no basal plant from which the old corm roots grow. Usually short, fat and rounded, it has a knobby surface with growth buds, or eyes, from which the shoots of the new plant emerge. Tuberous roots are the only ones Corm from this group that are real roots; their food supply is kept in root tissue, not in stem or leaf tissue as in other bulbs. Rhizomes, which are sometimes called rootstocks, are thickened stems that grow horizontally, weaving their way Tuber along or below the surface of the soil and at intervals sending stems above ground. -
Liliaceae Lily Family
Liliaceae lily family While there is much compelling evidence available to divide this polyphyletic family into as many as 25 families, the older classification sensu Cronquist is retained here. Page | 1222 Many are familiar as garden ornamentals and food plants such as onion, garlic, tulip and lily. The flowers are showy and mostly regular, three-merous and with a superior ovary. Key to genera A. Leaves mostly basal. B B. Flowers orange; 8–11cm long. Hemerocallis bb. Flowers not orange, much smaller. C C. Flowers solitary. Erythronium cc. Flowers several to many. D D. Leaves linear, or, absent at flowering time. E E. Flowers in an umbel, terminal, numerous; leaves Allium absent. ee. Flowers in an open cluster, or dense raceme. F F. Leaves with white stripe on midrib; flowers Ornithogalum white, 2–8 on long peduncles. ff. Leaves green; flowers greenish, in dense Triantha racemes on very short peduncles. dd. Leaves oval to elliptic, present at flowering. G G. Flowers in an umbel, 3–6, yellow. Clintonia gg. Flowers in a one-sided raceme, white. Convallaria aa. Leaves mostly cauline. H H. Leaves in one or more whorls. I I. Leaves in numerous whorls; flowers >4cm in diameter. Lilium ii. Leaves in 1–2 whorls; flowers much smaller. J J. Leaves 3 in a single whorl; flowers white or purple. Trillium jj. Leaves in 2 whorls, or 5–9 leaves; flowers yellow, small. Medeola hh. Leaves alternate. K K. Flowers numerous in a terminal inflorescence. L L. Plants delicate, glabrous; leaves 1–2 petiolate. Maianthemum ll. Plant coarse, robust; stems pubescent; leaves many, clasping Veratrum stem. -
Seed, Tuber, Bulb
Garden Education from the Salmon Center Seed, Tuber, and Bulb Exploration Activity Ages 9+ (can be adapted for younger age group if focus is primarily on observation) Overview: Most students know that plants grow from seeds, but did they know that they also grow from bulbs and tubers? The purpose of this activity is to investigate the differences and similarities between seeds, bulbs, and tubers through the use of observational skills. Students will also learn about the anatomy and function of seeds, bulbs, and tubers. Essential Questions: What do seeds, tubers, and bulbs have in common? What are their differences? Why does a seed, tuber, or bulb grow when planted, but if a leaf or stem is planted, it decomposes? Definitions: Tuber: A swollen, fleshy, usually underground part of a plant that provides food and bears buds from which a new plant arises (Examples include potatoes, artichokes, Jicama, and yams) Bulb: A short underground stem surrounded by fleshy leaves, which contain stored food for the embryo inside (Examples include garlic, tulips, daffodils, and lilies) Bud: Compact growth on a tuber and inside a bulb that develops into a leaf, flower, or shoot Seed: An embryonic plant enclosed in a protective outer layer Seed coat: The outer layer that protects the seed/embryo Embryo: The baby plant inside a seed. It has only two tiny leaves and the beginnings of a root Cotyledon: The part of the plant that provides food for the embryo Materials: ● Seeds of different shapes and sizes (If using beans, consider soaking beforehand to allow for easier dissection) ● A tuber (a potato is an easy one!) ● A bulb (try garlic or a flower bulb) ● Magnifying glass ● Dissection tools (tweezers, knife, fork, etc.) ● Seed, Tuber, and Bulb Anatomy Guide (included) Start the Activity: 1. -
Specialized Roots and Stems Text Pages: 561 – 587
57 Specialized Roots and Stems Text Pages: 561 – 587. Objectives: 1. Be able to describe the various types of specialized roots or stems on some species of plants. 2. Be able to describe and explain propagation procedures used to multiply plants by specialized roots or stems. 3. Be able to describe and explain limitations of propagating plants by specialized roots or stems. 4. Be able to predict how physical manipulations or treatments affect propagation of specialized roots or stems. I. SPECIALIZED STEMS AND ROOTS A. Introduction – specialized structures B. Tubers 1. Tuber - is a swollen, modified stem that functions a. A tuber has all the parts of a stem, and i. a tuber has buds, leaf scars, and ii. eyes - are the buds on iii. a terminal bud is at iv. tubers exhibit apical dominance b. The tuber is borne on c. Examples: 58 2. The growth pattern is that the tuber forms the first year, a. The tuber is used as a food source and b. Certain environmental conditions favor 3. Propagate tubers by 4. Tubercles - are small tubers C. Tuberous Roots and Stems - these structures are 1. Tuberous root - is an enlarged a. It is a root b. Buds that are formed are c. Example: d. Growth is as a biennial i. tuberous root forms one year ii. then in spring, new shoots grow and produce iii. the swollen root provides 2. Tuberous stems - include swelling of the hypocotyl, lower epicotyl, and upper 59 a. Note: this structure is vertically oriented b. More then one bud can be produced c. -
Vegetative Vs. Reproductive Morphology
Today’s lecture: plant morphology Vegetative vs. reproductive morphology Vegetative morphology Growth, development, photosynthesis, support Not involved in sexual reproduction Reproductive morphology Sexual reproduction Vegetative morphology: seeds Seed = a dormant young plant in which development is arrested. Cotyledon (seed leaf) = leaf developed at the first node of the embryonic stem; present in the seed prior to germination. Vegetative morphology: roots Water and mineral uptake radicle primary roots stem secondary roots taproot fibrous roots adventitious roots Vegetative morphology: roots Modified roots Symbiosis/parasitism Food storage stem secondary roots Increase nutrient Allow dormancy adventitious roots availability Facilitate vegetative spread Vegetative morphology: stems plumule primary shoot Support, vertical elongation apical bud node internode leaf lateral (axillary) bud lateral shoot stipule Vegetative morphology: stems Vascular tissue = specialized cells transporting water and nutrients Secondary growth = vascular cell division, resulting in increased girth Vegetative morphology: stems Secondary growth = vascular cell division, resulting in increased girth Vegetative morphology: stems Modified stems Asexual (vegetative) reproduction Stolon: above ground Rhizome: below ground Stems elongating laterally, producing adventitious roots and lateral shoots Vegetative morphology: stems Modified stems Food storage Bulb: leaves are storage organs Corm: stem is storage organ Stems not elongating, packed with carbohydrates Vegetative -
General Introduction
01hfdst.02-055 09-07-2002 12:31 Pagina 9 Chapter 1 General introduction Flower bulbs are a major export product of the Netherlands. In 2001, the Netherlands produced an estimated number of 10 billion flower bulbs, which is 65% of the total world production.Tulip, narcissus and lilies are the most important crops. More than half of the flower bulbs is used for the forcing market for cut flowers and pot plants and the rest for the dry sales market for parks and home gardens. More than 75% of the total Dutch production is exported to the United States, Japan and to countries within the European Union. (Sources: Internationaal Bloembollen Centrum, Productschap Tuinbouw) 9 01hfdst.02-055 09-07-2002 12:31 Pagina 10 Chapter 1 Flower bulbs Periodicity of flower bulbs Under natural circumstances, flower bulbs are exposed to seasonal changes in temperature, rainfall and light (Le Nard and De Hertogh, 1993a).The growth cycle of the bulbs is adapted to that periodicity. Depending on the season of flowering, growth activity of the bulbs takes place during spring or autumn. Spring flowering bulbs have a rest period during the summer and resume growth during autumn (Le Nard and De Hertogh, 1993a). They require a warm-cold-warm temperature sequence for growth and development.Typical examples are the tulip, freesia, narcissus and hyacinth. Summer flowering bulbs rest in winter and resume growth in spring. Typical examples of summer flowering bulbs are lily, allium and gladiolus. The time of flowering is not related to the time of flower initiation. Flower initiation can take place during different periods of the year and at different stages of bulb development.