Specialized Roots and Stems Text Pages: 561 – 587

Total Page:16

File Type:pdf, Size:1020Kb

Specialized Roots and Stems Text Pages: 561 – 587 57 Specialized Roots and Stems Text Pages: 561 – 587. Objectives: 1. Be able to describe the various types of specialized roots or stems on some species of plants. 2. Be able to describe and explain propagation procedures used to multiply plants by specialized roots or stems. 3. Be able to describe and explain limitations of propagating plants by specialized roots or stems. 4. Be able to predict how physical manipulations or treatments affect propagation of specialized roots or stems. I. SPECIALIZED STEMS AND ROOTS A. Introduction – specialized structures B. Tubers 1. Tuber - is a swollen, modified stem that functions a. A tuber has all the parts of a stem, and i. a tuber has buds, leaf scars, and ii. eyes - are the buds on iii. a terminal bud is at iv. tubers exhibit apical dominance b. The tuber is borne on c. Examples: 58 2. The growth pattern is that the tuber forms the first year, a. The tuber is used as a food source and b. Certain environmental conditions favor 3. Propagate tubers by 4. Tubercles - are small tubers C. Tuberous Roots and Stems - these structures are 1. Tuberous root - is an enlarged a. It is a root b. Buds that are formed are c. Example: d. Growth is as a biennial i. tuberous root forms one year ii. then in spring, new shoots grow and produce iii. the swollen root provides 2. Tuberous stems - include swelling of the hypocotyl, lower epicotyl, and upper 59 a. Note: this structure is vertically oriented b. More then one bud can be produced c. Examples: d. A tuberous stem is perennial 3. Propagation of these specialized roots and stems is by D. Rhizome - is another type of 1. Rhizome is defined as a specialized stem that is the main axis of the plant, and a. A rhizome has nodes and internodes that can be b. Two basic types of rhizomes: i. pachymorph - is a rhizome that is aa. it has a compressed, squatty appearance bb. examples: ii. leptomorph - has very long, slender appearance and aa. examples: 2. Growth pattern of pachymorph rhizomes: 60 a. This type of rhizome appears as a b. It has determinate growth meaning each clump ends c. Older sections of rhizomes d. Roots usually grow on e. Since vegetative growth, meaning lateral shoots and rhizome production, as well as flower buds form after flowering, all foliage 3. Growth pattern of leptomorph rhizomes: a. The rhizome is symmetrical and b. Most lateral buds on the "stem" remain dormant 4. Propagation of rhizomes is a. Plants are divided before growth starts in b. Each rhizome piece should contain a E. Bulbs 1. Bulb - is a specialized underground organ made of a short, vertical stem a. Vertical stem axis i. it is fleshy or 61 ii. it bears a growing point b. Bulbs scales are fleshy, i. outer scales store food ii. inner scales are 2. Bulblets - are new bulbs that form a. Consider these meristems as new growing b. Bulblets must be grown to a certain size before 3. Two types of bulbs are 4. Tunicate Bulb a. This bulb has its outer scales covered by b. Scales are continuous in c. Examples: d. Growth pattern includes vegetative growth first, i. in the vegetative stage, the bulb must grow to ii. the developing bulb gains most of its size 62 iii. bulbs go dormant in 5. Nontunicate (scaly) bulbs: a. This bulb consists of separate scales b. These bulbs also lack a i. these bulbs are less tolerant of ii. nontunicate bulbs must be kept c. Many species of these bulbs have i. contractile roots are ii. these roots shorten and pull the bulb down d. Example: e. On some lily species, new bulbs form in the leaf axis on the stem i. bulblets form ii. bulbils form on f. Growth pattern 6. Propagation of Bulbs 63 a. Offsets - sufficiently rapid method for b. Bulblet formation on stems c. Bulblet formation on scales i. scaling ii. adventitious bulblets form d. Basal cuttage e. Bulb cuttings F. Corms 1. Corm is a swollen base of a stem and is enclosed a. Note: corms lack b. Since a corm is a stem, it has c. Corms have two types of roots: i. fibrous roots develop from ii. contractile roots - are thick or fleshy roots 2. Growth pattern a. As the stem growth emerges from the corm, the base of i. this swelling eventually forms 64 ii. this year's corm deteriorates b. The new corm puts out stolon-like structures that i. these small corms are called ii. cormels develop between 3. Propagation of corms includes: a. New corms – b. Cormels – c. Corm cuttings – G. Summary 65 c:\plsc300\12notes-6 .
Recommended publications
  • Aeroponic and Hydroponic Systems for Medicinal Herb, Rhizome, and Root Crops Anita L
    Aeroponic and Hydroponic Systems for Medicinal Herb, Rhizome, and Root Crops Anita L. Hayden1 Native American Botanics Corporation, P.O. Box 44287, Tucson, AZ 85733 Additional index words. Arctium, Urtica, Anemopsis, Zingiber, Scutellaria, greenhouse Summary. Hydroponic and aeroponic production of medicinal crops in controlled environments provides opportunities for improving quality, purity, consistency, bioactivity, and biomass production on a commercial scale. Ideally, the goal is to optimize the environment and systems to maximize all five characteristics. Examples of crop production systems using perlite hydropon- ics, nutrient film technique (NFT), ebb and flow, and aeroponics were studied for various root, rhizome, and herb leaf crops. Biomass data comparing aeroponic vs. soilless culture or field grown production of burdock root (Arctium lappa), stinging nettles herb and rhizome (Urtica dioica), and yerba mansa root and rhizome (Anemopsis californica) are presented, as well as smaller scale projects observing ginger rhizome (Zingiber officinale) and skullcap herb (Scutellaria lateriflora). Phytochemical concentration of marker compounds for burdock and yerba mansa in different growing systems are presented. Production of medicinal herb and root crops the plants hydrated. NFT is a gutter (channel) of the crop are suspended in a spray chamber in controlled environments (CE) provides op- system without any aggregate medium, and where they are fully accessible for monitoring portunities for improving the quality, purity, where the fertilizer
    [Show full text]
  • Nematode, Ditylenchus, Stem and Bulb, Meloidogyne, Root Knot
    BIOLOGY AND CONTROL OF STEM AND ROOT KNOT NEMATODES r Becky B. Westerdahl1 Abstract: Plant parasitic nematodes are nonsegmented-microscopic roundworms which are frequently present in alfalfa fields. Although more than 10 different genera have been found in alfalfa fields in California, two (stem and bulb, and root knot) are most commonly associated with damage. A management plan to fit a particular growing situation should be developed using a combination of techniques including: planting site selection, certified seed, clean equipment, weed and irrigation management, resistant varieties, crop rotation, fallow, organic amendments and chemical nematicides. Ke~words nematode, Ditylenchus, stem and bulb, Meloidogyne, root knot, INTRODUCTION Plant parasitic nematodes are nonsegmented-microscopic roundworms which are frequently present in alfalfa fields. Whether or not alfalfa is to be planted in a nematode infested area, a grower should be knowledgeable about nematodes. If nematodes are present, both pre and postplant management strategies should be developed for pathogenic species. If an alfalfa field or a potential planting site is not infested, a grower should be aware of techniques available to prevent the introduction of harmful species. For growers to carry on a nematode pest management program they need to be familiar with (1) nematode biology; (2) symptoms and signs of nematode f damage; (3) how nematodes injure plants; (4) how to sample for nematodes; and (5) the principles underlying various management techniques including: planting site selection, the use of certified seed, the importance of using clean equipment and irrigation water, weed management, the use of resistant varieties, crop rotation, fallow, organic amendments, and chemical nematicides.
    [Show full text]
  • Enzymatic Hydrolysis of Lotus Rhizome Starch Using Α-Amylase and Glucoamylase
    Journal of Food and Nutrition Research (ISSN 1336-8672) Vol. 56, 2017, No. 4, pp. 372–380 Enzymatic hydrolysis of lotus rhizome starch using α-amylase and glucoamylase LI GUO Summary To study the susceptibility of lotus root starch to digestive enzymes and its potential impact on glycemic response, enzyme kinetics and in vitro digestibility of the granular, gelatinized and retrograded starches were analysed. The results showed that the digestion rate coefficient values of the granular, gelatinized and retrograded starches were 4.6 × 10-3 min-1, 9.8 × 10-3 min−1 and 2.3 × 10-3 min−1, respectively. Compared to the granular starch, content of rapid digestible starch (RDS) increased by 39.0 %, content of slowly digestible starch (SDS) and resistant starch (RS) decreased by 9.6 % and 15.0 % after gelatinization, respectively. While content of RDS decreased by 21.1 %, content of SDS and RS increased by 2.1 % and 20.8 % after retrogradation, respectively. As for glycemic index (GI) and hydroly- sis index (HI), GI (70.57) and HI (56.21) of the gelatinized starch were higher than GI (66.63) and HI (49.03) of the granular starch, and GI (57.83) and HI (33.01) of the retrograded starch. The results provide an interesting information about exploring novel and slow digestible foods made of lotus root starch for potential health benefits. Keywords lotus root starch; digestibility; α-amylase; glucoamylase Lotus (Nelumbo nucifera) is a well-known and mucosal α-glucosidase in the human gastroin- medicinal plant widely cultivated in Asian coun- testinal tract [5, 6].
    [Show full text]
  • Ongoing Evolution in the Genus Crocus: Diversity of Flowering Strategies on the Way to Hysteranthy
    plants Article Ongoing Evolution in the Genus Crocus: Diversity of Flowering Strategies on the Way to Hysteranthy Teresa Pastor-Férriz 1, Marcelino De-los-Mozos-Pascual 1, Begoña Renau-Morata 2, Sergio G. Nebauer 2 , Enrique Sanchis 2, Matteo Busconi 3 , José-Antonio Fernández 4, Rina Kamenetsky 5 and Rosa V. Molina 2,* 1 Departamento de Gestión y Conservación de Recursos Fitogenéticos, Centro de Investigación Agroforestal de Albadaledejito, 16194 Cuenca, Spain; [email protected] (T.P.-F.); [email protected] (M.D.-l.-M.-P.) 2 Departamento de Producción Vegetal, Universitat Politècnica de València, 46022 Valencia, Spain; [email protected] (B.R.-M.); [email protected] (S.G.N.); [email protected] (E.S.) 3 Department of Sustainable Crop Production, Università Cattolica del Sacro Cuore, 29122 Piacenza, Italy; [email protected] 4 IDR-Biotechnology and Natural Resources, Universidad de Castilla-La Mancha, 02071 Albacete, Spain; [email protected] 5 Department of Ornamental Horticulture and Biotechnology, The Volcani Center, ARO, Rishon LeZion 7505101, Israel; [email protected] * Correspondence: [email protected] Abstract: Species of the genus Crocus are found over a wide range of climatic areas. In natural habitats, these geophytes diverge in the flowering strategies. This variability was assessed by analyzing the flowering traits of the Spanish collection of wild crocuses, preserved in the Bank of Plant Germplasm Citation: Pastor-Férriz, T.; of Cuenca. Plants of the seven Spanish species were analyzed both in their natural environments De-los-Mozos-Pascual, M.; (58 native populations) and in common garden experiments (112 accessions).
    [Show full text]
  • The Iris- Empress of Flowers
    The Iris- Empress of Flowers by Susan Camp If the rose is the queen of flowers, then the regal iris must be the empress. She stands tall, elegantly nodding her head to lesser flowers and mere mortals. The tall bearded iris, in particular, always attracts attention and admiration from gardeners and passersby. One cannot help but pause and appreciate the delicate construction of the blossom and breathe in the sweet fragrance. The colors of the iris range from white through sherbet shades to deeper hues, all the way to purples so deep they are almost black. The colors seem especially vivid this spring. The iris is named for the Greek messenger goddess, symbolized by the rainbow. While it is fun to romanticize the iris and imagine it as a regal representation of the flower world, the iris is a plant with specific cultural needs and several pests and diseases. Iridaceae is a huge genus of 200-300 species. Most species grow from either rhizomes or bulbs. A few grow from fleshy tubers. The species are immensely diverse. The most popular irises grown in the United States are the tall bearded and other bearded varieties. The tall bearded iris, which is rhizomatous, is the focus of this column, but if you enjoy the beauty of the flower, the possibilities for your garden are almost infinite. The tall bearded iris can reach a height of 2 ½ feet. The leaves are vivid green, fleshy, and sword-shaped. The showy flowers consist of three upright inner petals called standards and three outer hanging petal-like sepals, known as falls.
    [Show full text]
  • Culture of Iris Anne M
    G1741 Culture of Iris Anne M. Streich, Extension Educator Dale T. Lindgren, Horticulture Specialist Iris culture emphasizes the best in site selection and preparation, planting, culture, and insect and dis- ease control. Irises are among the most popular and beautiful garden flowers for Midwest landscapes (Figure 1). More than 200 species of irises have been found in the wild and from these species, thousands of varieties have been named and made available for public use. Iris plants range in height from just a few inches to over 3 feet and are adapted to a variety of environmental conditions. The standard iris, Japanese iris, Siberian iris, Spuria and yellowflag types are suitable for Nebraska. Iris flowers can be from 1 or 2 inches across up to 8 to Figure 1. Irises 10 inches across and come in almost every color and often in two-color combinations. Irises can be selected to have continuous flowering from early April through June by using Planting an assortment of iris species and cultivars. Irises can be divided into “bearded” and “beardless” Irises grow from an enlarged underground stem called types. The term “bearded” refers to the presence of bushy a rhizome. These rhizomes grow just below the soil surface. “beards” on each of three drooping, petal-like sepals, called They are the source of growth for fans of leaves, flowers falls. The true petals are called standards and are upright. and the roots that anchor the plant. Rhizomes are used to Bearded irises, commonly called standard irises, are the most vegetatively propagate new plants of the same type.
    [Show full text]
  • The Impacts of the Entanglement Concentration on the Hydrodynamic Properties of Kudzu and Lotus Rhizome Starch Aqueous Solutions
    Journal of Food and Nutrition Research, 2016, Vol. 4, No. 11, 750-759 Available online at http://pubs.sciepub.com/jfnr/4/11/8 ©Science and Education Publishing DOI:10.12691/jfnr-4-11-8 The Impacts of the Entanglement Concentration on the Hydrodynamic Properties of Kudzu and Lotus Rhizome Starch Aqueous Solutions Li Guo*, Shuilin Wang, Chenchen Zhu, Jian Hu, Juanjuan Zhang, Xianfeng Du* Department of Food Sciences, Anhui Agricultural University, Hefei, China *Corresponding author: [email protected]; [email protected] Abstract With the rapid development of consumer demands for health, kudzu and lotus rhizome starches have been widely utilized as the nutritiously and naturally medicinal drinks after they are suspended in aqueous solutions. However, it is difficult to control the suitable concentrations to obtain the ideal textures of the kudzu and lotus rhizome starch solutions. In this study, on the basis of starch structure characteristics, the hydrodynamic properties of the kudzu and lotus rhizome starch aqueous solutions around entanglement concentration (the boundary between the semi-dilute regime and the concentrated regime of a polymer solution, ce) were studied. The results indicated that the two starch solutions showed a clear up-turn curve of the ηsp/c versus c curves in dilute solutions. The ce values of the kudzu and lotus rhizome starch aqueous solutions were determined to be 1.56% and 0.6%, respectively. The impact of the ce value on the network formation of the kudzu starch solutions was much more significant compared with the impact on the lotus rhizome starch solutions. Shear thinning behaviour hardly occurs when the concentrations of the kudzu and lotus rhizome starch aqueous solutions were lower than ce, and shear thinning behaviour develops when the concentrations are equal to or greater than ce.
    [Show full text]
  • Tyler Schmidt, Plant Science Major, Department of Horticultural Science
    Interspecific Breeding for Warm-Winter Tolerance in Tulipa gesneriana L. Tyler Schmidt, Plant Science Major, Department oF Horticultural Science 19 December 2015 EXECUTIVE SUMMARY Focus on breeding of Tulipa gesneriana has largely concentrated on appearance. Through interspecific breeding with more warm-tolerant species, tolerance of warm winters could be introduced into the species, decreasing dormancy requirements and expanding the range of tulips southward. Additionally, long-lasting foliage can be favored in breeding to allow plants to store more energy for daughter bulbs. Continued virus and fungal resistance breeding will decrease infection. Primary benefits are for gardeners and landscapers who, under the current planting schedule, are planting tulip bulbs annually, wasting money. Producers benefit from this by reducing cooling times, saving energy, greenhouse space, and tulip bulbs lost to diseases in coolers. UNIVERSITY OF MINNESOTA AQUAPONICS: REPORT TITLE 1 I. INTRODUCTION A. Study species Tulips (Tulip gesneriana L.) are one of the most historically significant and well-known horticultural crops in the world. Since entering Europe via Constantinople in the mid-sixteenth century, the Dutch tulip market became one of the first “economic bubbles” of modern civilization, creating and destroying fortunes in four brief years (Lesnaw and Ghabrial, 2000). Since this time, tulips have remained extremely popular as more improved cultivars are released. However, a problem remains: even though viral resistance and long-lasting cultivars are introduced, few are capable of surviving in a climate with truly mild winters and only select cultivars are able to store enough energy for another year of flowering, even in climates with colder winters. Current planting schemes suggest planting annually, wasting tulip bulbs (Dickey, 1954).
    [Show full text]
  • Bulbs: Culture and Maintenance by Diane Relf and Elizabeth Ball, Revised by Joyce Latimer, Virginia Cooperative Extension
    GARDENING FACT SHEET Harris County Cooperative Extension 3033 Bear Creek Drive, Houston, Texas 77084 281.855.5600 • http://harris-tx.tamu.edu/hort Bulbs Culture and Maintenance Originally published in the Arizona Master Gardener Manual, produced by the Cooperative Extension, College of Agriculture, The University of Arizona, 1998. Edited and reformatted by Texas Cooperative Extension, Harris County, September 2007 he term bulb is loosely used to include corms, tubers, tuberous roots, and rhizomes, as well as true bulbs. This publication will T refer to all of the above as bulbs. leaves A true bulb is a complete or nearly complete miniature of a plant stem encased in fleshy modified leaves called scales which contain reserves of food. Corms are the base of a stem that becomes swollen and solid with nutrients. It has no fleshy scales. The lateral bud tuber, which is an underground stem that stores food, differs from the true bulb internode Bulb or corm in that it has no covering of dry node leaves and no basal plant from which the old corm roots grow. Usually short, fat and rounded, it has a knobby surface with growth buds, or eyes, from which the shoots of the new plant emerge. Tuberous roots are the only ones Corm from this group that are real roots; their food supply is kept in root tissue, not in stem or leaf tissue as in other bulbs. Rhizomes, which are sometimes called rootstocks, are thickened stems that grow horizontally, weaving their way Tuber along or below the surface of the soil and at intervals sending stems above ground.
    [Show full text]
  • Rhizome Elongation and Seagrass Clonal Growth
    MARINE ECOLOGY PROGRESS SERIES Published November 26 Mar Ecol Prog Ser Rhizome elongation and seagrass clonal growth Nuria Marball*, Carlos M. ~uarte* 'Centre for Estuarine and Coastal Ecology, NIOO, Korringaweg 7, 4401 NT Yerseke. The Netherlands 2~entred'Estudis Avanqats de Blanes, CSIC, Cami de Sta. Barbara sln, E-17300 Blanes, Spain ABSTRACT. A compilation of published and original data on rhizome morphometry, horizontal and vertical elongation rates and branching patterns for 27 seagrass species developing in 192 seagrass stands allowed an examination of the variability of seagrass rhizome and clonal growth programmes across and within species. Seagrass horizontal rhizomes extend at rates ranging between 1.2 and 574 cm yr-l, develop a branch, with an angle from 19 to 72", for every 6 to 1800 horizontal internodes, and add a new shoot for every 1.1 to 7.5 cm of rhizome produced. Vertical rhizomes elongate at rates between 0.1 and 34 cm yr-' and the probability that they will branch varies over 3 orders of magnitude. Much (between 40 and 173%) of the variability of seagrass horizontal rhizome and clonal growth pro- grammes is species-specific, largely (21 to 63% of the variance) associated with differences in size among species, although seagrasses also show important intraspecific variability. The broad repertoire of seagrass rhizome and clonal growth programmes explains the different rates and efficiency at which the species occupy space. The implications of specific growth programmes for space occupation were examined by simulating the development of seagrass rhizome networks of 3 seagrass species encom- passing the range of horizontal rhizome growth [Halophila ovalis, Thalassodendron ciliaturn, Posidonia oceanica).
    [Show full text]
  • Seed, Tuber, Bulb
    Garden Education from the Salmon Center Seed, Tuber, and Bulb Exploration Activity Ages 9+ (can be adapted for younger age group if focus is primarily on observation) Overview: Most students know that plants grow from seeds, but did they know that they also grow from bulbs and tubers? The purpose of this activity is to investigate the differences and similarities between seeds, bulbs, and tubers through the use of observational skills. Students will also learn about the anatomy and function of seeds, bulbs, and tubers. Essential Questions: What do seeds, tubers, and bulbs have in common? What are their differences? Why does a seed, tuber, or bulb grow when planted, but if a leaf or stem is planted, it decomposes? Definitions: Tuber: A swollen, fleshy, usually underground part of a plant that provides food and bears buds from which a new plant arises (Examples include potatoes, artichokes, Jicama, and yams) Bulb: A short underground stem surrounded by fleshy leaves, which contain stored food for the embryo inside (Examples include garlic, tulips, daffodils, and lilies) Bud: Compact growth on a tuber and inside a bulb that develops into a leaf, flower, or shoot Seed: An embryonic plant enclosed in a protective outer layer Seed coat: The outer layer that protects the seed/embryo Embryo: The baby plant inside a seed. It has only two tiny leaves and the beginnings of a root Cotyledon: The part of the plant that provides food for the embryo Materials: ● Seeds of different shapes and sizes (If using beans, consider soaking beforehand to allow for easier dissection) ● A tuber (a potato is an easy one!) ● A bulb (try garlic or a flower bulb) ● Magnifying glass ● Dissection tools (tweezers, knife, fork, etc.) ● Seed, Tuber, and Bulb Anatomy Guide (included) Start the Activity: 1.
    [Show full text]
  • Tecophilaeaceae 429 Tecophilaeaceae M.G
    Tecophilaeaceae 429 Tecophilaeaceae M.G. SIMPSONand P.J. RUDALL Tecophilaeaceae Leyb., Bonplandia JO: 370 (1862), nom . cons . Cyanastraceae Engler (1900). Erect, perennial, terrestrial herbs. Roots fibrous. Subterranean stem a globose to ellipsoid corm, 1- 4 cm in diameter, in some genera with a membra ­ nous to fibrous tunic consisting of persistent sheathing leaves or fibrovascular bundles . Leaves basal to subbasal, or cauline in Walleria, spiral; base sheathing or non-sheathing, blades narrowly linear to lanceolate -ovate, or more or less petiolate in Cyanastrum and Kabuyea; entire, glabrous, flat, or marginally undulate; venation parallel with a major central vein. Flowers terminal and either Fig. 122A-F. Tecophilaeaceae. Cyanastrum cordifolium . A Flowering plant. B Tepals with sta mens. C Stamens. D Pistil. E solitary (or in small groups) and a panicle or (in Ovary, longitudinal section. F Capsule. (Takh tajan 1982) Walleria) solitary in the axils of cauline leaves. Bracts and bracteoles (prophylls) often present on pedicel. Flowers 1- 3 cm long, pedicellate, bisexual , trimero us. Perianth variable in color, zygomor­ fibrous scale leaves or leaf bases or the reticulate phic or actinomorphic, homochlamydeous, ba­ fibrovascular remains of these scale leaves (Fig. sally syntepalous; perianth lobes 6, imbricate in 2 123). The tunic often continues above the corm, in whorls, the outer median tepal positioned anteri ­ some cases forming an apical tuft. Corms of orly; minute corona appendages present between Walleria, Cyanastrum, and Kabuyea lack a corm adjacent stamens in some taxa. Androecium aris­ tunic (Fig. 122). ing at mouth of perianth tube, opposite the tepals Leaves are bifacial and spirally arranged.
    [Show full text]