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€8I ?llE4snv I€Jlu.J ur snu{uoJcrtrt pue ououlold,{rgJ ,uoorC ,ld .I 184 NuYtsiaVol. 3 (2) 1980

It must be emphasised that the key to genera given here is designed to apply only to the inland ALrstialian region; complications present elsewherehave been deliberately ignored. Some indication of the variation outside the present area is given in the generic descriptions. Specialised terminology-The relating to the androecium and gynoecium follows Green (l979iand Creen(1q80). termforal rubeis prelerred rotaly\-tubc. used by Bentharn it8tli fot lhe rea\on\stated by Parlin (1955)and Douglas{1957) The r5eof ttomium for a resion of dehiscencein ihe anther is explained by Esau (1965) and illustrated by Fahn (1974). The rnorphology of the gland on the connective requires some explanation, as it does not appear to have been previously used as a diagnostic character in this group: it may vary'from a near-globular, | featureless structrue, as in M. helmsii (Fig. l12) throueh a serieshaving an increasingly prominent neck, with an apparently porate, terminal orificel In T. patviflira (Fig. 17)and T. nuviculqta (Fig. 30) the gland usually-protrudes onlv shortlv between the microsporangia and is termed trunc(tte or compressed-urceolate; wh6n it js i contracted below the apex, as in T. v,ittv'eri (Fig. 37) and T. hexantlra (Fig. 63). it is termed urceolqte, while at the extreme, when the neck is long and curved, as in T. elliotrii tFie. 54), it is termed urceolate-falcata. The shape of the gland may also be clavqte, as \n-Tl. mqisonneuvcitFigs. 6-7) ot conpound, with smaller lateral bulges, as in M..finbrisepala (Fig. 100). The flower stalk is here interpreted as a peduncle as it appears homologous with the structure bearing two or three flowers with pedicels in related genera Onlv for unusually variable parameters are dimensions qualified as approximate. Characters omitted from sone descriptions can be assumed to be unknown.

Key to the genera,basetl on inland Australian species -i- 1. Stamens less than 10, usually opposite the sepals; ovules 2 or 2 2 super- gland prominent, often equalling 'orDosed. ascending or lateral; anther connective exceedins thi anther, clavate or + urceolate . Thryptomene Endl. (p.184) 1*. Stamens10, or 5 oppositethe petals;ovules 2 or 6 10,collateral, apical; anther connectivegland lessthan half as large as the anther,t'iltl:"O,ill,1i", ;"rth. (p. 195)

THRYPTOMENE Endl. Glabrousslrubs, slenderor spreading,mostly 0 5 2 m high (one speciesoutside the areaarborescent); bark usuallysmooth, dark greyIo bro\.\n. occasionally6brou'. Lenves small. entire. opposite,mostly + flat above and convex below, with severalto many i.-.is.a ghnds'especiallyvisible on the lower surface. F/ouers sessile,subterded by a pair of iomplicatL braCteoles,at least in the bud stage,inflorescence mostly solitary wiih a very short to long peduncle,borre singlyin the-upper leaf axils;some specie-s with inflordscencespairedll|the axrls. Floral tubemostly less than 3 mm long, cylindrical, tulbinate. hemisphirjcator rarely laterally flattened,smooth, ribbed or rugose,adnate to the ovary, sornetimesproduced beyond it making-the. disc- surrounding the style + concave. Sezdlsand petils 5 or rarely 6, borne on the rim of the tube; sepalsas long ur ifr" tot " tb very shbrt, scarious.petaloid or petaloidwith scariousmargins, entire or denticulate;petaliaboul as longas the tubeor shorter.rnostly rounded. entire. moslly (antesepalous)or 10 (in a single 'whorl,pink, white or the two suffused. ,Slarrersusually either 5 not aluays regularlyopposite the perianth parts).1rarely 6 (antesepalous,f. /rex- an,Jra),ya.iable between 6 and 9, or between15 and 30 (in one speciesoutside the area); filurodotfilifotm, .rsuallyat leasttwice as long asthe anthel at maturity; connectivebearing a Drominentgland which may be globular, truncate,urceolate or falcatewith a porate orifice. and sometimesprotruding betweenthe two microsporangia;anther tetraspor- 'squ .puq3cs '8uol lnoq€ s/rdas trnoqlr,r lo eso8ru-l?FpuelS dlolnuru 'eJ?urqJn]JEurpnlfuol ,snor.rBcs ruur 9. L lnoqE f,lpeorq aqnt ,paddrl-ftlsau €eJ€ quprru eql 'JlnJE '8uoT ,el"Joecu€l\oto1g .luelsrsJed,Z .slrxB surbrBrrrrqt utu I lnoq" ,{lp?olq sapapp.tg laddll Jql ur ItElrTos 'elrssJss./ ).|o/!r .[Bla^esspu"l8 ,,r\oleq p3urolloq-}?U ,pEoJq 'e^oq" Io lee{ '8uol daep € qtr,\\ xe^uoc l?q,reulos '{crql 'l?lnarqJo to ,{lpeolq u[u 'ellsses 's,{\oJ 'ole3rJqurr 'elessncep le3rldrllo 'dtededZ I t ur ,{l.leolc sa$aI u,rolq-qsrppoJ 'pEorq IJEq Jo snorqg ^luos qlr^\.sualsJ ru t.Z ol dn pur,;q8rq uI S.I t.0 qn.rF tx A '€uulqeplno_ (€Il0/ lgtr{ :oloq) ..8uno^ le sue)torI :sns:re^eBu€U s,e^Er8snl{ s.luoru uolr rou to qeplnol seuorletsedotd,, :adtrJ .lZ:0I .urierl '.[anl.ll|.d otolD!.trDeuauntd, qJ '(Zll0l'IaI,,{:oloq).,l1€rus pcefi t srlErluoce€rlEr}snv elurC uelrlng py,, . adlq .{rr^nauuosr€W,, '(t98I) 'llenI,,\tr'C 9-t9 :t ru3"rl ra^nouroslsruouauold,{rql'I

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j 'ere,,,oor\\ I r!!H'g 'scBuBU:r Jo ,nnN,(Lur08t) letlu 00€eJ :vI-rydJSnv Hmos (;$I]r;:f ;:3;Jrf,E 7 prere^apue eFu€U e^efsnr\t urr\\trg :(IN .AVt pinil5p :g .yrifi SS6l{i"i,1111 11Sil optpuv]o N ur)tZlresec uosdlurs:(IN,-IAhl) t 16lfEI I Z,otz1Lrpy 9 s.reIol,rqJo ,^^NN (I'u{toj selrlu0t:,\dolTdugJ Nd3HIUoN 1HIt8d) l96I Bny 9Z'L88Z)8roaD S' r'(uorssrtrt),{rreq^\eN o-tu:oJ srlrtu :(HIUfd) 'L6OtL 'dnv Jo 3N 58t'(J t161..{lnf0t a$od Z 6r'uor}e}SuolC "urol :(CV) 9€6I oz .ltl\tltI B^A qzl ,s az.qz)rloH \iJoderoo3eqqpS :(Hlu ,{e11g1 .69!; ;.iray tdr Ltot .s o y roorcJtsuolnnrfJoNllulc)\rtrlut:rHlutd)ot6l {c,\ s.ozqEt )Stort s-vI t0t szl s|,tz) rrr\r(J \pu€s rf?rrD alrl u.qol :bnbu tsnv NU tJS t,\\ :puttttD\J\u.,ut.,)dt Jo tnltrlps 'ql61 3fU ees-1608, u7,:I clehl :II-t,"rnarg .r"q-"no'[qf,(9nl,{ 'frtn.rqeg 11e,no61g) ZZ: pepJocal SurJe,roll uaes .pe8relue jrtq .Xre,lci jd -etuereld tou 2aag lou eql esic] eql r€ou l u uto:g s:red pesodJednso,rt ur Sursue ,tr sa1n,t6 .Biol 'qlqt luJclul wut 9.0 lnoqE a/f/S "lloroJ eqt spl",ro] SurpnJlord,{ltueuluo-rd 'e}"^"[J e^ttcauuo3 ,3uol oql uo pu?13:?rLuots l?urpnlr8uol o,rt ,{q Surcsrqap urrr 2.0 lnoqe dreqtuz i8uol ruur g .0 lnoq P slu.1LuETIJ :slurLUr-"LU orll alquosel qctq,\\"rpourut€ls fu"ru serllr,r Burl?ulell" 'snoJEdJsJtue'9 AJIeuorstJJo snLuDjS .poJ Jo doop ,pellrd ,,no11eqsrsrg .1urd 'rolau"rp 'teln.rqJo lurd .dll Jo etrrl,r ruur Z-9.1 lnoq" splal esnlqo ,,{qseg,1crq1 e 'rrlnBurrrl ,sur8l"ru lrir aJluoJ prol"lod ,{1pro.rqe el"lnor.rne,eurF,{q qrr,^,ri8u

(2) 188 NuYtsiaVol 3 1980

This speciesi5 recordeda5 occrlrringon a varielyofsoil typesinclrrding Iuterite' red with 'hrubby rood- trna, "t'riii sand.sandy "containingc6y and hard-iettin-gred earth asociated iunJ'oi ior"ii country species of Eucalyptus, Callirtis, Grevillea, Dodonuect, Acacia, Eremophila ot Melqleuca. A pood deal of confusion over localities seems to have originated in a rrix-ltp- in S""ti,ufi;, (1861) notes tnder Thryptomtne oligandra. His de.scriptions-correctly dis- linsirirh b.ti"".nihe arborescent f.'oligandra, rvhich has broad lelves and large.flowers, u"E lfr. tftt.UUv T. part'iflora (treated by Bentham as a.rariety of T', oligctntlra), whtch tu, nurro" lear-e.lrjd smallflouers. brrt he hascorrfrr.ed the geognrphicalrrngrs of the ruo speciesuhich are quite distincl.T. olisandraoccrtrring on Cape.York. Penlnsula Be- iit"T.- iirrifluro. is restricted to an area in-cantral nnd south-eastern Queensland .;;; B";ih"; apparentlydrew his publishedcitation from the.incorrectly-labelled MEL iOii-S, *iij.h ii "iultt"a ui seenby hinr, this specimen is assumedto be the holotype rather than the duplicate at K. The poptrlationat Palm \ alle). \.T . preriotr'ly relerrcdto as T' pattiflora b5 .Chip- pendrle(l971). ir inclrrdedhere in Thryprom,tE\itt"cti l ntt\ l:Patr.tllotars co^nsroerecr (enrrill io bc restrictedto Orteenslandltnd occursout'idc the ilreacovered by the I lora oT Australia. T. parviflora is very closely related to a Western Australian species,]d tl(lviculatq, ot *ni"fr'tie onfv known iocalitiei,both near Lake Disappointment,are withir the latitudinal range of T. pani;flora.

3. Thryptomene naviculata, J. W. Green, sp. nov.

Ftutex rctnndalus;fol/d 4-seriata,clliptico-obo\.ltx, + l. o.Inrn longa. Sulra concava, infra rotundata j*iei11ibui 'upeio 'e*iles.soli.lar i. a\illsres:r//b//' ,"r "o,l',rii". in drrridio rninuredenticulari.: /ul: -o'eo i;";J;;-i;;,i;;5lco.rarrsl repdl,rungu.iculata. l 8 mm lorga percloideaalba \cl rinera:r"4/.' ."onf- u* ilnriflua"a uiiquanturn majoia; Va r'lia5, xnte selala positr ; connectivum glandem plominentem po\rla' con Drc.so-,r'ceolalarr'leren.: u'rrla 2 in plcccn a stlbba\all Zlpe . Near Karara Well, Canning Stock Route NE of Lake Disappointmcnt,-W A, )iids's. tzz"zz'v,A. S. Mit(hetf914,2 Mav 1979(holo: PERTH; iso: CANB, K, PERTH).

Rounded shrub lp to 1 m high and broad. Iccves closely-irubricate, decussaie, 4- rowed. broadly elliptical or obovati, 1 3 I 8 mm long, corca\e above, rounded or keeled telo*. rnargin'minutelvdenticulate in the upperhalt Lcol bnresdecurrenl' uhite' corer- ins rhc,rrrfice ol'rhe leafy branche.. F/or'r'rs'es'ile. about 2 mm longrnd 4 6 mm broad. soTitarvin the upper axili of nost leafy branches,forming nunerous condensed,capitate oi "pirie-tit

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'uoojC 'ld 68I ell€Jlsn\/ IeJluoJ ur snU,{woJcrIAIpue eueuotdtuqJ I NuYtsiaVol 3 (2)1980 190 associated with a At the lype locality thc species formed a low, open shrubland' Thttntomene navi(:ulatLt n.urr.^p"ritop!'f, i udia ip. or Pkitrqchne schinzii, on.rid^sand', *Hflr;*;rHm,i+$#;#T#[:;:'#l'$T,:'.t::iir"1,1':H]iili].]X?l# tube.

4. Thryptomene wittrveri J. W. Grcen, sp' nov' ..rn*ii'.1i;i?iili,.It,iliix1?,\iikfi'i1JilliliilS'"?it J;i,li?.,''"|,%.Tl,',1:;;':l'J:",..H:i;; {:ii{ ils\s,: *:lli11i$::';f*l:ii::'J""*;;",trJl"ff't'ggti"1;'-?liii"fafifi;il"l''Tl,,l?"rftiK:::,',?: subbasaliposita. Aug 1973(holo: frrc. Upper[parts of] Mount Augustus,W A', E' lfittt'er 1109,20 pEnrUi iso:PERTH). ' -small i'l,niill'fiJili;ll:l::?i;:","tffiTirfi;l1l , ,"i3i::lil,i.",l,i:;"i."*'Jl'f"}i..' oil glands iFlotrers solitz ffirouridcd'betorv,;;:,-'r.,i"+:fi,'l#ir,:.;:$ff ivith numerous leaving severaLshort, hairlike ;f":#.ifri[}*].|.;;:iii#*-JJ""ll;1'"|.-.i''f'?#: iHn:ru:'m',.;.i:l'"*;:tl'lT#oi#il;'*l?f stanens ;;i:iii; ;"ffi.-';"rals sintilar,sodewhat exceeding the sepals Disc conca\s' base;anthers i. irii,*ph.iir, o.i mmlong;'filaments short, dilaled and flartenedat the l:irm"|,".lffi'lr**"i;ktil'"lni:*$x:tr'nriil::!:L,tl"::**'i fjij,.-'s":li#i''i:r,,?ji4**:,'L:il: Tfl"lljfil!?:s'".:[h';lxffi:i"i','il?:: 32-39, Map 2. l':t'tr;x;,,;;i,\::',{.,y1:Jlll;^,f il$.^i,lit",li.i"ir)S:'[i'i,t].i:lli?]'),,,'s,ll il'fil; i.;sh'1ffo.'\'i,, p""r' i;j6, ;. a. Beatrytrhotr27508. 24 rul\ lqb8'AD, cANBr' occ,.,ningin.a st:lt"i:",1 Recordedas |;3;li:JJJfil,ti1ii1t'i!j'.'rt"""#ffit;IJi Ernsrwittwer, superintende't' or'ii"gi i"* .."9 ;il't-*;^r'J': t;'"li;t'|"iil]iJi:;;f The speciesis i;fii'" c;t;;;, *i.to Jir.ou"t"a the western AustraLian'i:'Ipopulation' fi*y-rYl,f.*"li,l{i:l:r,,;":;{^'#:i"I:;x'i",ilYil;"i""'l*Ti';il.Yi:llil hexirmerottsspccies L hexqndra.

5. Thryptomene nealensisJ. W. Grecn, sp nov' pedunculati; lrneaa, crassa,3 5 mm longa;-/fo''er solitarii' axillarcs' breviter gl'*lifun:m:tli*:r:***i:'sl,'.';""llfi.":1lg';'?::li:i::l:,Lar,lTt'.:?l:1;Ig:'tliiFrrl.-r humilis;Jblia posta. Fig

(22 km) E of Neale Junction, Great..Victoria Desert' W A ' approx tubl I1'pe; 14 miles 49- ii-:ii's. rz-i;il'i, A. s. Georgigaze. lt o.t. 1966(holo: PERTH; iso: K' cANB' PERTH). Flo 53, 'r8ueU ,tS e-^Er8snr{aqt-puE duulqEplnO u!o^\tog Bro{ /9-9S :096I 'Anv 'unopleg "u s uftl uro{ 'ZS-8} ^J.\J)JoOSU'{J?AO,lAoleq pue qnno.tql .Jqnl { 's.I'S'It9 s'8' SJ leJol{ -/9-99 seln^o pu€ ^Je^o,{Je^o bur,r(\oqs8ur,r(\oqs re.,ilolc-gsle.,ie.,i oJC-olC-gs uoursls -t9 eqnl IeroE paue €g 8ut.{oqs ^{a 're,{old-€9 Surpuelqnsls pu? rEcs eloo}r?rqeloolr?rq 'roaolg-zg'ro1r\olg-29 .SJ .lBrxBqB'lBrxBqB Frxepe'llq"H-8t :Jeol 'l9-8t SJ le.reltlle]ele TEe-I I9-6t :llttolp auauoittlqJ l9-8t sorn8rC 9Zt8 otuoo9 urorJ IIV eluereld qSnorql eqnl selnroseF o puspue,{rE\o xrB^o Buhoqs8ur^\oqs 'S.I Je\1olJ-9tre.{{ol{-9t sururEts-gt-ttsueurE}S-9t-tt .Jeel Surpuelqns pu? IeJolClpJolJ SI-/tSl-tt 'SI 'l€lx€q€ 'Jee'f-'Zr-WS-I recs eloepeJq 're.,i\olC-gt llqeH-ot : slsaaloau auawofiItqtr ./t-{t sem8rd

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,ueero .r,\A.I I6I ellErlsnv IBIUeJ ur snu,{rnoJcrl I puB eueuoldturlJ 192 NuYtsiaVol. 3 (2) 1980

Shrub 0.3 m high. Learcs decussate,closely imbricate, not obviously in 4 rows, linear, thick, 2-5 mm lorg. 0 6 nm broad, :f flat above, rounded or keeled below, + sulcate when dry, covered with numerous oil glands, obtuse or with a minute, acute tip. F/ol ers solitary, axillary, 3 4 mm long, scattered along the upper leafy bratches' Peduncle 0.3 0.7 mm long, somewhat flattened. Bru(Jeolesdeciduous, leaving several short, hair- like strands subtending the flower. Florql tube broadly obcorical or turbinate, flaring above a + obvious constriction above the ovary. Sepclsorbicular, I mm long, + entire, petaloid, pink. Pelals similar, somewhat larger. Drsc concave, shallow. Stantens 5, irntesepalous,or occasionally6, about 0 6 mm long; fllamentsvery short, flattened and ditated at the base; anthers versatile, rounded, 0 8 mm broad, pink, dehiscing by two longitr.rdinal stomia; connective gland urceolate, the neck short. Sl.l,/e short, about 0 5mmlong. Ovall cavity in the upper half of the tube,the placentat basal. Ovules2, elliptical or somewhat angular. Fruit and ^reednot seen. Flowering recorded October. Figures40-47; Map 2. This speciesis knorvn only frorr the type collection,found on a lateritic breakaway. No further details as to habitat or associatedspecies are recorded. It differs fron fri1?- tonlene witt$eri chiefly in having smaller, narrower leaves and a shorter peduncle.

6. Thryptomeneelliottii F. Muell., Fragm. 9:62 (1875). ?/p?. "In eremointer' Youldeh el Beitanam(Beltana); E. Giles." (neo: "BetweenYouldeh and CharlotteWaters, E. Ciles"MEL 7068,1;isoneo: AD 97137326),neo. nov. "On Thrlptomener,fttuae J. M. Black,Trans. & Proc.Roy. Soc.S. Austral.4l: 384 5 (1917).Tlpe: theEast-West Railway, 60 miles (96 km) NW ofPort Augusta.",S.A. white, Jan. 1917 (holoi AD 97534332a iso:MEL 70815. NSW 136171). ShrubO.3 I5nihigh. Zearesdecussate, scarcely imbricate, L 4-ranked,shortly petiolate, obovate-clavate,thick, 2-3 nm long, 1 mm broad, flat or somewhat convex above,rounded, -t keeled (or sometimes sulcate on drying) below, obtuse or very shortly acute-tipped. Flowers solrtary or occasionally in pairs, axillary, 4-5 mm long, 1 mm broad, crowded irrrong the upper leaves. Peduncle I mm long. Bracteoles deciduous. Floral tube oblong, 3 4 mm lorg, somewhat irregularly l0-ribbed often saccateat the base,tangentially to the axis flattened at the level of the ovary cavity, on one side or both, sometimesminutely papillose on the abaxial surface. Sepals znd petttls orbicular, I mm long, petaloid, pink or white. Dl,rc concave, shallow. Stamens5, antesepalous,nearly I nrm long, filaments short, about 0 3 nrm long, flattened and dilated towards the base; anthers rounded, 0 5 mm broad, versatile,dehiscing by two longitudinalstomia; connective gland very prominent, almost as large as the anther, urceolate-falcate. Sf,/e short, about 0'6 mm long. Orarl in the upper half of the lube. Plocentcr btLsal. Ovules 2. Fruit a\d J"ed not seen, Flowering recordedApril-October. Figures48 57; Map 3.

Selectionof specimensexaminecl: WESTERN AUSTRALIA: Eeldoun(Iltoon), N of Loongana,l. rR. Main, l7 Arg. 1960(PERTH); SOUTH AUSTRALIA: "Betweenthe Elizabethand Youlden,Ciles Exped.1875, young" (AD, MEL); BetweenOuldabinna and the MusgraveRange (MEL); ca 100km S of VokesHill, N of Cook,T. R. N. Lothialt5691, 20 July 1972(AD); N of Fowler'sBay, Gi1ff (MEL); ca 22 km N of Watsonon Maralingaroad, T. R. N. Lothian5521, 14 July 1972(AD); Balton,E. H. Ising 1312,17 Sep.1920 (AD, PERTH);Wynbring, Herb. J. M. Blaclc,22Sep. 1920 (AD, MEL).

This speciesis widespreadin the vicinity of the Trans-Australianrailway, between Loonganain WesternAustralia and Wynbring ir SouthAustralia, and northwardstowards the Musgrave Ranges. Recorded only from sandy soils, usually on red sand dunes or ridges. The few available label details indicate that mallee (Eucalyptus spp.), spinifex (Triodia spp.) and Acqciq linophylla nray be associatedspecies.

Af thor-rghclosely related to Thtyptotllenenealensis, T. elliottii is d\strlguished by having almost always sone flattening of the floral tube, as well as a very prominent connective granq. '€ere clqq?Jd -oa8 1e:oua8eluEs aqJ ur ,{Jotupxeqpodola^ep e^Erl pllroqs snue8 qaeeJo setcedse18uts e ,{rrurproerlxe sureeslI ,rau1t-Yaqsul{tuorJttl! setcedsctrledul,{s ,{llullr"d pal?lal leqt 'Je}JErtqc -un oql ul s"r{ 1r s€ uotlelndod elqElspuc turoJrunr-' se oJpuDxaq7 uI pe^lo^e 'lIxE e,req o1 snedde ,{reuexe11 eq1 uI lo,llog euo uuql eJout Sut,leq dleuotsecco JEel 'ntpuDx)Ll Jo relf,?l"qJ Iensnun eql llqtqxe oslc oserll Jo eruos 7 e1t1 1de15 ( 1put1) nu1ttr1n lpu1' J'\ooB DLltuD.nlu!7'llsnll I DaD)!DJ ol pel"loJ osl€ sI pue'slsualDaa'J pvr lJa ulr auatuoltlI.ttll ol pel"loJ ,(lesoycst 1l pu€13 e^Illeuuoc el€loecJn pue eqnl l€.rou peqqu oql e.leqsqalq.^\ selceds snoJolul?luad Jo dnoJ8 e o1 s8uoleq setcedsstql

'..qnll: 13ltrru,, \r PePloJer uorl€la8e^ pal€rcoss" ,41uoeq1 drecs e;o e8po 3ql uo ellJolcl 8ul,{lte^o IIos,\\ollsqs pue rueoy ,{purs 'se8prJ pu? sopISIIIq^uols opl'Ilcur selceds slql JoJ pepJooeJ sl€lrq"H

'Dpp!.l7 '€ruoiurrg'elll^soloorg 'a)irnog L96I :(/ SN) 8961^oN'.talotlttv l:1 , Jo N (*n O;SilY (tdg) des'uostawwg rtoN :(.]AW'ruS',{[OV 'CV) Ot :SafV,t HInOS ,4{AN lclB^Epv996I t'eldlrno tz6l 8nV 6Z'SS6tD.t^tllt5)ol4l /fi rt6N :(NV'ISNagnO :p.)uluDx)suaut)2.Is lo llotpalas

t d"tN :99-89 sern8rj 'sqluorll paprocer Suuo^\olg oJnl"rrr elrrb tou uoes :]ln{ 1nq 7 ,{lluaredde spaa5 'pe8J"lueII" 'Z l"q^\eruos oqn] l"rou SuIlInJC BluoJEld lzstqqns " uo ,{Il"Jel"llo3 euJoq saf,ro 'avoI rulu a7,175 e1,{1soql 8ulJ",JeJod eql'el"loeJJn ^llJoqs e^rlceu 9.0'Japuels 'olll?sJe^ -uoc eql uo puolt Erwrols o,\\l fq SuIJStqep'ssoJ3? luol t?c leurpnlt8uol 'suoru?lst.0 s.leqlue:8uol rulu lnoq"'ruJoJITUslueluElg !3uo1 rtrru /.0 lnoq? sftol?led i.0 'snol"desetu" 'g -01u" :F Jo uorlpp" oql fq 8 ol dn Jo suaul9ls J"lnpu€I8 ,{lsnol^qo lou 'e1rq,r--,{ur€er3ro eltq,n 'pro1e1ed'.Ielelu"rp ulor 'l€lnclqJo 'JelItiIIS'9 s1o|adpue slodag 8 0 'sledes ',rloll"qs 'e,\ecuoors./d. ,{tr.rnleur1" Suol urtu g.Z lnoq€ eq} ,no1eqSurqcuerq 'el"lnu€druec-elelo squ eruos Jo 'peqqrJ-O1 ,{lluourruord ',{1"^o eql e^oq" polcetluoJ 'pzoJq 'BLroI -ealn aqu pJolf ruur Z. I o] cln pue 3uo1 utot t lnoq€ sJaaolf ulul g . T I ',(1r;n1eur 'snou?.Iqluour 'leoull 'lclel"l 'Z l" sllolrptc€p sap)pp.tg 3uo1 ruru 1.0 ol dn sapunpal qcu"Jq rlcee puo 0r{1Jo ulc ulq:}L{\se^B3l eql sllx? eql uI € u3^e Jo z Jo 'spuep z I Jo 'oJJnul ,{lFuors"Jao lo ,{JBtrToss.ra,rolg ,{ueur ()1 qllu pallop 1to lyzrus I?Je^es'e^oq€ elnurur ? qlr^{ Jo esnlqo ',{\oleq pelea{ tr1lq8tls to pepunoJ e^€cuoc ,'(l,\\ollEqs 'peoJq 'ole^oqo-,\\o.u1?u 'pe,{oJ-t' ruor y. T o1 dn pue Suol ruur 9 I [re,l ro reeutl ,{lsnor^qo 'clessucep 'elrsoddo 'Peqcu?Jq-qonlrl 'asnJrd lou sa,raaT q31q ui 7 T gr.qs Sutpeerds

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'.{1e,tr1cedser ed,{1oau -osr pue ed,{1oeupeleu8rsop oleq eJ€ ,{eqt 'Je^e,$oq 'Joord sJuosq?eql ur :3d.'{loloq eI{} 'uotltpodxo Jo epnlcur plnoc,'(er{l l?rl1 olqlssod st 11 pllql erll uo ep"tu uollcelloc elSuIS" 01 Suoloq 01 reedd"'sollD fq pelcelloc 's.leqlo egJ tloll"cllqlrd JoJ erull uI Jollelll peqJ"eJ el"rl ol 1rJoJ ol?l oo1"aru 3ur1ce11ocolqlssod erll q8norql Sutsstd'9181^"lN uI q ed JoJ €lsnSnv l.rod Uel qcrq,r uorlrp:dxc qunoJ eql uo pelcelloc ueeq e,\"q ls11Luu3lul33ds erll eJoJ -eJeql :uonrpedxe pJrql oql uo lou s€,\\ oLl,{{Suno^ ,{q polcolloc s?,\\put uotltpedxe prtr11 eql lou tnq qunoJ eql Jo elnoJ eql uo s€,r LIJII{,\\Jo^Iu qloqezlla eql ol s.IeJeJIeq"T euo 'en8o1o1ordeql ur esoql ruorJ Suuogrp suollducsul -{lll"col Jeeq pe}Dcol uoeq e^"q q5tq,{\ 'uoDlpodxo lollentrAlFq ueos suelurceds,{luo eq1 ree,( l€ql pu" qcl?t\ ueo,{\l3q Jo IUdV'gl8l p.lq] ,solrD uo pel.clloc ueeq e^pq lsnu od^l eql aunf uI uoltdlrrsep eql paqs( -qr.Ldoq sy "li14 u\ lluoup 7 jo ed,{1o1oqs.rollon1tl eterol 01 elq"un uoeq e^"r[ I 'ueerC E6l erlerlsnv lurtuo3 ul snlr,{rllorarlI puc euaruotd.,{rql l f NuytsiaVol. 3 (2) 1980

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j9- Figur€s 58-66. Thrypto.mene.hexandra,; 58 Habit. Leaf pair. ieaf TS. leaf tips. 60 loflorescence showing flowers io pairs. 61.62-Flower. exrernal longitudioal and oblique from above. 6j_64_ Slamensliom above and inside. 65-66-Fruit. showing rwin, single seededfruirs. 58_61,6j_65 from Althofer s.n., 40 miles (64 km) N of Bourke, Nov. t968: 62 from Lidden s.n.. Brookesvilie.Enneonii- f967. Figures 67 76, Micromyrtus hexamera:67-}labil. 68-phyllotaxy. 69-Flouer and ped-uncle. 7012-Flo\7;e\ extemal and from above. ?3Jtamens. 74-Floral tu6e, LS showing ovaty. ZS- O\,'ulesand placenta., 76-Ovule._and stylar-vein_ 75 from E. Berche,Warrego R. disfict, Sc;p.1900; all othen from Silcock s.n., 30 miles (48 km) SW of Charlevi e, Aus. i969, :6tttZ 'I88OZ'6180r'IAI des 'eqileg 'L{!ra^rUotsele^ 'aipug s,proC ", "rr."J'$'.n:tOSH I) 9881 a .des ,lleqdluel .S se palrJl ro^ru o8orrer1,\pur SullrBC uea^\tag:(I06€tl ^ SN) t68I lA,un8urrreg o] 3)rnog LUorJpDod :sad,ltD.todoeaT(It€Il.'otela.IEtr',,I:osr:006€tI .des.eq:rto{ 'i 'l,Je^IX 'ollequrooC IASN :oloq) 006I oierel ol e)rnog uro{ peol{,, ss pelrrl lJu}srp F^rl{ oSorrelA :(oteq peteu -?lsep) a(lltopaT (1061)ZB :92 sole/ S N JoS uur.I cord'oqctog to pr€l,^lr./a&/Dxaq auauoJdt ii '(916t.) 'eqcleg ,91 stu?ld sol€y'itsN snsueJ ti pr?l\ (eqclegte pl"hl) srausxaq srq uorrrl,{ .I

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's:rrd .01 pesodrednsur 1ou seF^o :lBlelel Burpuelro '1e:jdz 'auauroldtr.tllJ Z l?Jrdaqns o1uaco14 ur ueql lueururold ssel pue reldrurs ,{1lunsn '.(olDdas!.rqLu{ 'W) pu€13 e^rlceuuoJ 1,! pue Dluauouaurtq saneds o,\\l ur ellrJol luorusls '(sg"d ,1:oq,r :(otawoxaq t!) ZI,{ler"r qlu"rred eq1alrsoddo .(pep8a: e18urse ur ,{11uewdde .eltq(,(rueerc Jo snouoruelsoldrpqo) 91 ro (snol4ods1ue) g Jeqlre ,{[?nsn saawDjs .ro ',ro11e,{ 'E]JE ,(sltx" ,ro11e,4elzd sercedsle.re^es ul slDtad eql eprqno sarcodslele,\as ur erll ur ,{J€lrlos 'elcunpad e13u1se uo ' e r) palo^\og-t o1 dt atuacsatogli,t1.luecseroqrz 'sgi?r4s euou s.relJeJ"qcSur,rolloJ oql ur (peuuap aJeqse) auawofiIl{J ruo{ sJeJrO 'qlu3gsoJ,u,{htou)u{tr .l s6l erleJlsnv IErtueJ ur snu,{Lalorrrt{pue ouoruotd,{rg_Ltoa.lC I 196 NuyrsiaVot.3 (2) 1980

Shrub O'6 2.5 m high; branchlcts slender; bark loose, fibrous, pirle yellow. leaves decussate,+ inbricate, sonewhat spreading,l-3 mm long, obovate-oblong,thick, con- cave above,keeled or somewhatflattened near the base,minutely ciliolateon the margins or._entire,_very _shortly petiolate, with several large oil glands below. F/orers solitary, axillary, often clustered near the ends of the upper branches, about 4 mn iong. peluir& 0'7 1.2 mm long, shorterthan or I as long as the subtendingleaf. BracteoTesscarious, about,1 mm long, deciduousin rnost specimens,leaving short, hair-like tracessubtending most flowers. Florc[l tube narrow-obconical or turbilate, 2 rnm long, angled or fluted with 6 ridges, branching into several more below the sepals,the interstiies with numerous oil glands. Sepals6, semi-orbicular.0.5 nrm long, membranoLrs.pctals 6, much larger than the sepals,broadly elliptical, up to 1.8 mnr long, white, turning pink or on some individuals either pink or purple, minutely denticulata. Disc I fla1, often somewhat oblique. S/orrers 12, 1 mnr long; filaments terete,0.9 mm long; anthersrounded, 0-2 mm broad, versatile,dark brown, dehiscingby two longitudinal stomia; connective gland smaller than the anther halvcs, yellow. Sly/e slender, almost 1 mm 1ong. Ovary in the upper quarter of the floral lube. Plecenta aptcal. Ovules 8 ( l0? nde Maiden & Betche). Frirll scarcely enlarged. Mature sceds not seen. Flowering re- corded January, AprifNovember. Figures 66-76; Map 4.

S.ekctioof .sp(dnc \ examieLl: QUEENSLAND: Near Alpha Station, a. S. Snith6411,16June 1955 rBRI)iCalabrh._55 rnilei (88 knrJ \W or Charlc\iilc.A. t4ut,a).t8 Sep.tq67 rBRtl; Bei$eerr Beechal Creekand.30 miles (48 kln) SW of Charle\ilJe on Quitpieroad, -R . G. Sttcock,67 Aug.-1969(BRI, CANB, MEL, NSW):Boatman Station, ,S. L. Ercrist3098, I8 July 1947(BRt, CANB, NSW);48 tm W of Cunnal tqL1, -r: L:-,Sltit,q0ro4, 8 Nov. 1954(B R t); St. Ceorge-Bollorrroi,l, C. ttl. Atthofet18, Juty 1949 (B R I). NEWSOUTH WALES: Kerribree-Laurrdale, J.L. B,nrman,Noy. t 912 (NSW); 6j miles(l0i) kmfr;m (N 9f) Pou_r!9^or'Jn-1e_gniaroad, G.. W. Althufcr 204, Oct. 197:(BRll; l0 mrles(48 kn, N oiBourke,R. Roe, 8 July1940 (CANB); 30 miles(,18 knr) W of Bourke,H. C. Dotman,Aug. I967.(NSW, pERfH).

_ Apparently fairly plentiful within its restricted distribr.rtion, Milomyrtus hexamercr has been.recordedmainly on red, brown or grey-brownsand or sandy )oam, sometimes on sandridges or in rocky places. Thc few notes on record list Triocliq, Atqcia "n.rulga aneuro, Eucab)ptusmelanophloia or box" as associatedvegetation.

I have found no evidencefor more than 8 ovulesin any of the specimensexamined, "Ovules including the three syntypes,despite the description 8 to l0'; in the protologue. Ovule number seernsto be remarkably constant in this species,though it could not be determinedwith certaintyin NSW 143901which might have had 6 oi7.

, ,Mi(:rom!-rtLts hexanerq belongs to a group of species,some as yet urdescribed, in- cluding M. cilictto (Sm.) Druce, which occurs widely in south-eastern Australia. It is very like_a Queenslandspecies of this group, M. Ieptocalyx(F. MLrell.)Benth., in floral tube, peduncleand leaf, but is distinguishedby its hexamery,style ard stamens. A note on the occurrenca of hexamery in the same area of distribution is included utder Thrvp- Iomc ( hcxqnlKt.

2. Micromyrtushymenonema (F. Muell.) C. A. Gardn., Enr"Lm.PI. Austral. Occ. 96 (193]).

Thrlptomenchymenonema F. Muell.,Fragm. 10: 26(1876)_Tper VictoriaSprings, young, 30 Sep. l875(holo: MEL 71345).

Shrub 0 7 m high. Zeales closely to loosely imbricate, decussate,linear-obovate, ,4-0 .5 2 4 mm long, 1-l mm broad, I sessile,minutely ciliolate-membranous,concave above, rounded below and keeled where broadest,the apex with a minute hyaline mucro 0.1 mm long, the suface dr-rll or glossy, oil glands several each side of the midrib. I/orl,ers solitary in the upper axils, 3-4 mm long, subtendedby two complicate, petaloid bracteoles I.2 I.8 mm long, slightly fleshyalong the midrib, and with membranousmarsins: ped- uncle flattened,0.7 mm long. .Floral tube turbinate, I 1.5 n.rmlong. deeply-5-ridgea, at least when dry, otherwise without sculpture, bulging slightly below the sepals. ,.!epals '29l0I surellll\ ruo{ '86-26'06-69 'uro,r 'eqnl 001-66'16 :91€8 o8roeD luo{ I0I lel,{}s pu€ saln^o Suuoqs S.I I€rolC-I0J 'eprstno ruorJ ueulets--{ol 'slleut"ls pu" 'leled-96 sledos-r6 .oloel.elg-S6 'SJ 'lEuretxolElad-66 'roa.\olg-26 J6 pu€ leurelxo'oqnt l€lolg-t6 €6 pue prx€q€ .r(xel -oll{qd-06 '10l-69 lelelel.9t89 :er-I-16.ura !lqeH-68 :olDdasuEu![sntrtluoDlw samBrC oAJooCtuo{ IIV rBIXls pu€ d4ropue snouerqruour'seln^O-88 eprsutpue oplslno (uo4'suaru€ls '/g-99 .eldls pue sueul€ls 'eloelr"rg .l€lxe 8ur^\oqs S-I'raa\olC a8 l€led-t8 Iedos-g8 Z8 seloepeJq pue ro^\old-I8

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'oeorD 'lA L6l ellerlsnv Ierlue3 ur snU,{1tIoJ.rII pu€ ououotd,{rql f 198 NuytsiaVol. 3 (2).1980 ovate-oblong,l-l 8 mn long and I 4 rrm broad, petaloidexcept near the ciliate margins, :t auriculate at the base. Petals pink, exceedingthe sepals,entire, + orbicular, 2.7 mm long. Dr,rc flat or slightly convex. Stamens 10, L I mm long, the antepetalous alternating with the somewhat shorter antesepalousin a single whorl; filaments lorate, 0.5 1 mm long,0.4 mm broad;anthersversatile, dehiscing by two longitudinalstomia; gland often 3-lobed,orange-red and red. Sly/e 0.8 mm long, not exceedingthe stamens. Ovules6, collaterally attached to the stylar vein near the summit of the ovary and contained within a fine membrane. Fruit unknown. Flowering recorded June-October. Figures 77-88; Map 5. Specimensexamifie(1: WESTERN AUSTRALIA: RawlinsonRange, herb. C. ,r. Gardner,A,uE-Sep. 1962 (PERTH);70 miles(113 km) W of NealeJunction, A. S. Geotge8411, 10 Oct. 1966 (PERTH); 27 miles (43km) NE of Lavertonon Warburtonroad, A. S. George2841,24 Aug. 1961(PERTH); N of Cundeelee, A. S. Georgc5846.21 Sep. 1963 (PERTH); Victolia Desert Camp 59 (ca 200 km E of Kalgoorlie),R. -l1elras, 20 Sep.1891 (AD, MEL, NSW);Queen Victoria Spring, R. D. Royce5517, 1 Oct.1956 (PERTH). This very distinctive species,inhabits red (and yellow) sand-dunecountry rvith spinifex (Triodiu spp.), over an elongated tract some 750 km long, to the north-east of Kalgoorlie. Only three collectionsare known outside the type area around Queen Victoria Spring. The specieshas only one close relative, M. Jinbrisepala, which also has broad, llattened lilaments.

3, Micromyrtus fimbrisepala J. W. Green, sp. nov. fuutex; folia late ellipticaad orbicularia,l 5 2-2 mm longa,marginibus minute ciliolatis, supra concava,infta propeapicen carinata.; flotes solitarij, axjllares, breviter pedunculati; trlrrr florisprofunde 5-plicatus,a sepalisvalde fimbriatis aLlriculatisfere celatus;petala rosea, orbicularia, ca 2 mn1diam.; stamina10;filamenta late lorata; glansconnectivi composiia, rubra; o,'ula6, collateralia. Type: 21 6 miles (34 8 km) W of Warburton, Gibson Desert, W.A. Shrub 0 7 m; fls. pink. In red sand among spinifex, between dunes. l. S. George8375, 9 Oct. 1966 (holo: PERTH; iso: CANB, K, PERTH).

Shrub 0 7 m high. leales imbricate, decussate,broadly elliptical to orbicular, I 2.2 mm long, up to 1 .5 mm broad, :1.:sessile, minutely ciliolate-membranous,obtuse, concave above, keeled below near the apex, lustrous, with several proninent oil glands. ,F/on'ers solitary, axillary, clustered near the branch endings, 4-5 mm long, subtended by two complicate bracteoles petaloid in the middle, with the margins membranous and minutely fimbriate, l 8 mm long; peduncle | flattened, 0 5-l mrl lotg. Florul lr./bcturbinate, 2 mm long, deeply 5-ridged,somewhat curved, almost entirely obscuredby the sepals. Sepals - orbicular, auriculate or sometimes peltate, strongly fimbriate, about 2 mm diameter, parchment-like. Petals pink, orbicular, clawed, scarcely exceeding the scpals, about 2 mm diameter. DrJc flat or slightly convex. Stamens 10, :jr I mm long, the antepetalous alternating with the somewhat shofter antesepalous, in a single whorl; filaments lorate,0 4 mm broad,0.5 {.8 mm long; anthersversatile, dehiscing by two longitudinal stomia; connective gland often 3Jobed, orange-red and red. St1/e 0.8 mm long, not exceeding the stanens. Otules 6, collaterally attached to the stylar vein near the summit of the ovary. Otary snall, near the summit of the floral tube, endocarp membranous. Seerls not seen mature. Flowering recorded February, October. Fig- ures89 l0l: Mao 5. Setuctionofspe?imens ?rdmlr?dr WESTERN AUSTRALIA: Type.SOUTH AUSTRALIA: SEof Cheese- man'sPeak, S. Batket31,15 Aug. 1979 (AD, PERTH); Serpentine Lakes, 54 km E of WA border,l' D. tvilians 10702,30Jul. 1979 (AD, PERTH);Vokes Hill roadjunction,228 km N ofCook,L. D.Witlioms 10796,6 Aus. I979(AD, PERTH). Until recentlythis specieswas known only from the sanddunes of the Gibson Desert west of Warburton, W.A.: discoveriesof severaloccLlrrences in the far west of South Australia have now shown its range to extend over a distance of some 500 km. Mr L. D. Williams, who first discoveredthe speciesin South Australia, thought that it appeared to proliferateafter burning (pers.comm.). .re'a .u,.n,"1r1,p,",,1nf6%t"',f,i.,u',iY:,'i^l5Yr 9,1 Lzr_.redos-ezr'lerxeqe i?;'"9,;3H:llH,i'";X"$: puE lerolel .t\qEH-Izl :x.qozoua$ M ttuo.r)tw sornSrg .Is69 J€e.I Ezl-zzt .ledes 8zt_Izl adrooC uro{ llv eprsur uro{ ueulets 0ZI Lelod 6lT glt pros Burdolo^opBuu{oqs 'ro.r^olJ .1er:;e1.lerieqe 'lrqsHS-I /II soloolcerqpu€ re.rou Jo ^\er^ prpold\a-g1L Jer-I 9IT -tIt :DtDlnrtassnrfluo!4w .0zl-tll srrnSrl pue g€ ,u 'eprsurtll ruoLu€tS-_ZII 6€ sdLuEJ]rasrc cuotcr^ s sluleH Luo{ llv ruorj pue a^oqe uro{ .€]ueteld pue sep,r6 urei_Je1,{1s-!irb 'al,{ls 'suelue}s lerelel.'oqnl .pdas .l€}ed 1y1 61y seln^o Sur.roqs S-I Isrol{-6OI .ijcunpoit pue :erb1g -90I 'l€rxeqe .llqeH 80I t6I .711 SI pue lejllel J€o-I 9OI-€OI Z0I :ltsalaq sntrtuotottr4J 761 serntij

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Because M. fimbrisepala is the first lO-stamened species of eilher Thryptomene ot MicroruIrtus to be recorded in South Australia, it will no longer be possible to distinguish between the genera there solely on the character of stamensantesepalous or antepetalous. While relatedlo M. hymenonema,as is evidencedby the broad, strapJike filaments, the new speciesis distinguishedby its most unusual,fimbriate sepals, as rvellas the smaller, rounded leaves.

4. Micromyrtus helmsii (F. Muell. et Tate) J. W. Green, comb. nov. Thryptometrcl,e/tnsli F. Muell.et Tate,Trans. Roy. Soc.S. Austral.16: 356(-1896). Tlpe: yictorla DesertCanps 38 and 39, W.A., R. Helns,2 Sep.I891 (holo: MEL 70695;iso: AD 97448028,AD 97534340, NSw l36l6l, PERTHlex NSw 136242]). Shrub 0 9 l 5 m high, branchlets slender. Ieaycs decussate,imbricate, appressed to the branchlets,obovate-oblong, 1.5-3 5 mn long, I mm broad, concaveabove, keeled near the apex below, obtuse, shortly petiolate, with several conspicuous oil glands below. Flor,ers solitary, axillary, sparse along the upper leafy branchlets, 3 4 mn long, 2 mm broad. Peduncletp to 1 5 mm long. Brqcteolesdeciduous, rarely present on herbarium specimens. Floral tube turbinate to obconical, 2.5 mm long, S-ribbednear the base, the ribs tendingto branch above,interstices rugose. Seprzlssemi-orbicular, 0. 8 mm long and I mm broad, marginsmemblanous. Petalsmuch larger,orbicular, I .5 mm diameter, margins minutely denticulate. Dirc concave,shallow. Starrers 10 in tbe bud, some- times some shed at anthesis("about seven" in the protologue),0 8 mm long; filaments 0 5 mm long, terete.somewhat thickened below; anthers0.5 mm broad, rounded, de- hiscing by two longitudinal stomia; connectivegland small, about 0.1 mm diameter, irregularly globular. ,Sry/eabout 0.5 mm long. Owry in the upper 1,/5 of the floral t\be. Placenta ?Lpicalbut on one side, adjacent to the stylar trein, Ovules7 8, dependent, collaterally arranged about the placenta. Fruit a\d seed not seen. Flowering recorded September. Figures 102-l 12; Map 6.

This spcciesis known only from the type locality which, accordingto Eardley(1950), lies in lat. 27'S, long. 127'E (near Mount Squires). Its habitat is not recordedwith the specimens. The species clearly belongs lo Mioomlrtus because of the character of apical placentation of the ovules which was not noted by Mueller and Tate. It has no very closerelatives within Micromvtus but has the samenumber of stamensand ovules as M, hymenonemaa.nd M. fmbrisepala.

5. Micromyrtus serrulata J. W. Green, sp. nov.

- Ftrtex;Jbliao\ato-oblonga, 1-2 mm longa, marginibus minule ciliolati!, supra concava, infi-a carinala; /d/zs solitarii,axillares, subsessiles, ad apicesramulorunt aggregati; bl'acteoli persistentes; tubus florti angusto-campanulatus,obscure lo-costatus; s4dl4 membranacea,0 6 mm longa:petala sepalis duplo longiora,ern, inea; stamina l0: filamentafiliformia; glans connectivi parva; ovula 2, iollateralia.

Type: 32 mlles(51 km) E of Karonie, Trans-Australiarailway, W.A., A. S. George5951, 9 Nov. 1963(holo: PERTH; iso: CANB, K, PERTH).

Slrrub up to 0.8 rn h\gh. Leaves appressed on the flowering branches, otherwise + spreading, obovate-oblong, l-2 mm long, shallowly concave above, keeled below, nrargins minutely serrulate,apex obtuse,surface dull, coveredwith numerousoil glands. F/oileru solitary, in the upper axils, aggregatedat or near the branch endingsin pseudo-corymbs 3-4 mm long, sessile, each subtended by two conspicuous, persistent, cymbiform-com- plicate bracteoles with a petaloid, glandular centre and membranous ed,ges. Floral tube about 2 mm long, narrow-campanulate,sonewhat indistinctly10-ribbed. Seprils0.6 mm long, I orbicular, broadly clawed, membranous, Petals twice as large ai the sepals, orbicular to broadly elliptical,creamy-white, with a few inconspicuousoil glands. Drc 'psorq '8uol .se^Bol rllur E.I urut S.Zaqry prol! eyt Surpeecxe,Suolullu S. . 'st1o11pnop'8uol 'el"crTdruo, .seq3u"Jq 9.2 l lt€nsn al)unpad ruuJ Z saloaQug oql uo 'poralsnlJ ,^l?llrx? ,,{Julrlos l?urru.relqns SurJ"JddpuJUo'8uoL ulru t,{ll?Jrd,{l srrJ.rolJ-ol€lncrluap Ilatnurul,{llEnsnsrll3]"LU 'spoEl8 qlr,r snocn€la ,p?olq ultrl g.0,8uol 'pJIJJ)l LrolBle^os l"q,\\euos I urLu- 'xJ\uoJ-ouBld 'ele^oqo-Suolqo,el€ssncep ,el?crJqlur satqaT ,]LJEq Z T'uMorq-rlsrpp:J + AJJdEd qlr,r sluJls :q8rq ru g.l-€.0,8utp?oJds to ssool ,lJalo'gn./{S' 'tlt6t) 'loup.rDD 96 tro le.rtsnv ld tunua V J ( llenj I J\ xqD;(tpD.ttsntl[luo.! W

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trlsrtsnv lurtual ur snlr,{Luorrrt\lpue euetuo}d,{lql .uoorD .y' .I 201 NuytsiaVol.3 (2)1980 '7 Shrub 0 25-0 m bjgh. Leat'es imbricate, decussate,nauowly obovate-oblong, plano- convex, t keeled below towards the apex, l-3 mm long, 0 7 mm broad, obtuse or with a minute, terminal mucro, somewhatglaucous, with severaloil glands; margins minutely ciliate. Floweff solitary in the upper axils, ofteu several clustered together subterminally, at maturity 7 mm long, enclosedin the bud by two folded bracteoles with ciliate margins. Peduncle 7-1. 5 mm long. Floral tube obconical, densely bearded with yellow, spreading or rellexed hairs up to I 5 mm long. Sepalssemi-elliptical I 5 mm diameter, the margins flmbriate. Petals elliptical, twice as long as the sepals,creany-white or white, the margins entire. Drsc deeply concave. Stantens5,2 mm long; filamentsfiliform, l 6 mm long; anthers versatile,dehiscing by two longitudinal, parallel stomia; gland subglobular, narrowing between the thecae to a tnincate tip, Ovules 2, collaterally attached to the stylar vein in the upper quarter cf the ovary and contained within a fine membrane. Frril not seen. Flowering recordedJuly. Figures l35 143; Map 7.

Aclditioliallpecime exa,r?/iredi wESTERN AUSTRALIA : 28 5 nriles(46 knr) Carnegie to MountEverard, A. R. Faircll2090,28 July 1966 (PERTH).

This very distinctive and apparently rare speciesis known only from the above local- ities,some 340 km apart, in red sandcountry ofthe Gibson Desert. The type was collected near a creekline, on a Triodia plain. The distribution is within the targe of Micromlrtus faviflora, to which M. barbqta is related but from which it differs markedly in the bearded floral tube, fewer ovulesand Ionger stamens.

Acknowledgements

I acknowledgewith pleasurethe advice and assistancegiven by my colleagueMr. P. G. Wilson, particularly in matters of typification and nomenclature. I also thank my other colleaguesand my wife for helping me with discussionson rrrany occasions. The Latin descriptionswere supplied by Mr. A. S. George, while the manuscript was read by Mr. R. J. Henderson,Dr. N. G. Marchant and Mr. P. G. Wilson, all of whom made many helpful suggestionsfor improvement. Mrs Wendy Lee-Frampton is thanked for drawing the maps and for other technical assistance. Miss V. L. Hamley is thanked for typing the nanuscript.

References Austin,M. P. & Nix, H. A. (1978). Regionalclassification of climaleand its relationto Australianranae- land. In "Studiesof the AustralianArid Zone,ltl, Waler in Rangelands"(CSIRO: Melbourne). Bentham,G. (186?)."Flora Australiensis" v.3 (Reeve: London). Chippendale,G. M. (1971). Checklist of NorthernTerritory Plants. Proc.Linn. Soc.N.S. Wales96: 20'7-26'7. Douglas,G. E. (1957).The inferior ovary IJ. Bot.Rev. 23: l-46. Eardley,C..M. (1950). Someearly Alrstralian plant collections. Robert Helms and the ElderExpedition Collections.Australas. Herb. News 7: 10 14. Esau,K. (1965).Plant Anatomy. (Wjley:New York). Fahn,A. (1974). PlantAnatomy. (Pergamon I Oxford). Green,J. W. (1979). Corynanthcrala new gerus of Myrtaceae(subfamily Leptospermoideae, Tribe Chamelaucieae).Nuytsia 2: 168374. Green,J. W. (1980). A revisedterminology lbr the spole-containingparts of anihers. New phvtoi. 84:401406. Parkin,l. (1955).A plealor simplefgynoecium. Phytomorphol. 5,46 47. Rye, B. L..(1979).-Chr_omosonre number variation in the Myrtaceaeand its taxonomicintplications. Austral.J. Bot-2'71 547-573. 'lL\ LSVLD d 'uostr^ :(t) 'Lroslrtd:(€) sLuerrlrlA:(t) o)!q,^^l(/) uouIs :Q) Z t\t'(r) H 'I'q}!usZ9l0t Z/8tt tttz 66nt'Q) E\ZI {.eds :(I) tlt9 t009 S :0) ^ruo^res'(/) tgt1t'Q) 6Lnor:(Z) rl9g oc.{od:(I) / ue8eu 1(I) 8ZtZ Ielped:(I) t98€'(I) 98!g eroor{ :(l) 9tg0l eo)rlr'{ :(I) 8t zsuo e}{ :(r) /Zl rofenl :(Z) 299 ure'^{:(r) '0.) '(t) '(r) '(t) erqrouocet{:(/) '(/) :(t) sepu€ze'I:(t) 898t 9081 €6€l 908'G) 6t9 0rt8 6628 9!I9 288zle'I:0,) vol, f,tteouuex:(l) 8916 6609 v H'uosuqof:(t) t s8ur{coH:(/) zI8 uDrqro'IT IIIH :(/) .r0dreH:(/) ep :(I) uopjoD :(9) :(9 '(Z) '(s) Z '(r) 9tr'0.) tBero 9€'(9) '(9) 9662'(9)ltrpoo5 6LrZr'(D'U) zz16'(r) 6S06 I ltg '(r) EZs8 s6r8 0608'(r)!(Z) 9008 9009 rs69'Q.) 6/8E'(z) '(t) 9r8s 8099'Q\ 69n9'(L)zLtg €E8,'0.) t96Z'Q) IggZ IvSZeEIooD :(r) 6€061 9t€rl €srz raupr€D:(8) 0602'(r) 9€02 Ip eg :(I) tt olglA T lsue^A :(I) 860€lsrre^E :(t) Ztg uqosroqg:(D ZgVn'Q) 86€t reuuoc :(t) t9t dsu] :(t) ZZ urnqlco) :(/) 6tI a)lel) :(r) 816ZetDpuaddrqJ :Q) 629'Q) Itg )IoouurqJ :(Z) OtJ Ilea\sog :(D :(l) oltr^eznegap :(r) euueg:(r) etoqat8neeg1(/) '(t) VZn'Q) n9b'O) 9St ltelcelg Itt'(I) 0I 56892 tt99 S68t preeg :(D L6tZ uqdy :(t) toz 8t reJoq{V :(/) 0Z tq8rrtv :(r) sSVH uosurq3}nHT' rrollrv snl.L{urorrll{l

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