Kansas University Quarterly
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Diptera Chamaemyiidae), an Overlooked Family of Biological Control Agents
Bulletin of Insectology 68 (2): 173-180, 2015 ISSN 1721-8861 Seasonal habits of predation and prey range in aphidophagous silver flies (Diptera Chamaemyiidae), an overlooked family of biological control agents 1 2 3 1 4 2 Serdar SATAR , Alfio RASPI , Işıl ÖZDEMIR , Adnan TUSUN , Mehmet KARACAOĞLU , Giovanni BENELLI 1Department of Plant Protection, Faculty of Agriculture, University of Çukurova, Balcali, Adana, Turkey 2Department of Agriculture, Food and Environment, Insect Behaviour Group, University of Pisa, Italy 3Plant Health Central Research Institute, Yenimahalle, Ankara, Turkey 4Biological Control Research Station, Yüreğir, Adana, Turkey Abstract Aphids are among the most widespread and serious groups of pests in agro-ecosystems, and predaceous arthropods have been proposed as biological control agents against them, including parasitic Hymenoptera, lacewings, ladybugs, hoverflies and silver flies (Diptera Chamaemyiidae). Chamaemyiidae is a small family of predaceous flies, including aphidophagous and coccido- phagous species. Little is known about their ecology, and partial failures of Chamaemyiidae-based biological control programs against aphids may be due to poor synchronization of predator-prey seasonal habits. In this study, we investigated seasonality of predation and prey range of aphidophagous Chamaemyiidae. A field survey was conducted on crops and indigenous flora in seven agricultural sites in southern Turkey. Seventeen host plant families were surveyed for Chamaemyiidae presence and 371 silver flies were studied. All Chamaemyiidae larvae were observed to prey on adult and young instar aphids in the field. Silver fly total larval abundance reached a maximum in autumn and early winter. Six species were identified: Leucopis annulipes, L. formosana, L. glyphinivora, L. revisenda, L. rufithorax and L. -
Tylosaurus and Pteranodon
Tylosaurus and Pteranodon Estimate size and measure to check estimate. OBJECTIVES Students will: 1. identify the Tylosaurus and Pteranodon as the two state fossils of Kansas, and 2. estimate and check the wingspan of the Pteranodon or the length of the Tylosaurus. MATERIALS FROM THE TRUNK Tylosaurus model Pteranodon model Fossil sample OTHER MATERIALS Ruler or tape measure Masking tape, post-its or something similar to mark measurements on floor TEACHER PREPARATION Decide which of the two fossils you will measure: Pteranodon had a 25-foot wingspan or the Tylosaurus was 49 feet long. Identify a location with 50 linear feet of space that can be used to measure the wingspan of the Pteranodon or the length of the Tylosaurus. Consider using a hallway, playground, or gym. Once a location is identified, use a tape measure to mark the beginning and end of a 25-foot linear space and a 49-foot linear space. This is where the students will measure the two state fossils. HISTORICAL BACKGROUND In 2014 the Kansas Legislature passed a bill making the Tylosaurus and the Pteranodon the state fossils of Kansas. Both of these reptiles lived at the time of dinosaurs, but neither are dinosaurs. Mike Everhart, adjunct curator of paleontology at the Sternberg Museum, geologist Alan Deitrich, and Steven Fisher, a 4-H geology project member, testified in support of the bill. Fossil hunters and natural history museums initiated the adoption of these state fossils. Kansas 4-H geology project members supported the bill. Pteranodon (teh-RAN-oh-don) – “Pteranodon, a great, winged pterosaur with a wingspread of more than 24 feet, which flew the skies of Kansas during the cretaceous period of the mesozoic era, is hereby designated as the official flying fossil of the state Kansas Symbols Traveling Resource Trunk KANSAS HISTORICAL SOCIETY www.kshs.org ©2014 61 of Kansas.” (House Bill 2595) The first Pteranodon specimens discovered in North America were found in western Kansas in 1870 by Othniel Charles Marsh. -
Diptera: Syrphidae)
MEMOIRS of THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Number 9 THE FLOWER FLIES OF THE WEST INDIES (DIPTERA: SYRPHIDAE) by F. CHRISTIAN THOMPSON Agricultural Research Service Agricultural Research, Sci. and Educ. Admin. U.S. Department of Agriculture, Washington, D.C. Published by THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Washington, D.C. 1981 PUBLICATIONS COMMITTEE of THE ENTOMOLOGICAL SOCIETY OF WASHINGTON 1981 E. Eric Grissell John M. Kingsolver Wayne N. Mathis George C. Steyskal Thomas E. Wallenmaier David R. Smith, Editor Printed by Allen Press, Inc. Lawrence, Kansas 66044 Date issued: 2 September 1981 TABLE OF CONTENTS Abstract ...................................................................... 4 Acknowledgments .......................... " ................ ,................. 5 Introduction .................. , ........................... ,.................... 7 Economic Importance ........ , ........................................ ,........ 7 Distribution .................................................... ,.............. 9 Taxonomy .............................................................. ,..... 13 Key to Genera of West Indian Syrphidae ......................................... 17 Syrphus Fabricius .............................................................. 20 Allograpta Osten Sacken .............................................. ,........ 23 Pseudodoros Becker .................................. , . 33 Ocyptamus Macquart ........................................................... 34 Salpingogaster Schiner ..................................... -
Diptera Chamaemyiidae) Described by Camillo Rondani: a Revision of Eight Silver Fly Species
Bulletin of Insectology 69 (2): 199-219, 2016 ISSN 1721-8861 On the identity of Leucopis (Diptera Chamaemyiidae) described by Camillo Rondani: a revision of eight silver fly species Alfio RASPI, Giovanni BENELLI Department of Agriculture, Food and Environment, University of Pisa, Italy Abstract The dipterologist Camillo Rondani described a wide number of new Diptera species, including 12 species of silver flies (Diptera Chamaemyiidae). The first Rondani description of Chamaemyiidae was carried out in 1847 in his research “Osservazioni sopra parecchie specie di esapodi afidicidi e sui loro nemici”. Among them, Ochtiphila obscuripes Rondani 1875 was considered by Czerny (1936) synonymous of Chamaemyia geniculata (Zetterstedt 1838). Later on, Raspi shed light on the identity of Leucopis palumbii Rondani 1872 (Raspi, 1983b; 1988); Leucopis bursaria Rondani 1848 (Raspi, 2003) and Ochtiphila frontella Rondani 1875 (Raspi, 2006). In this research, the identity of the other Leucopis species described by Rondani is estabilished. The valid species are Leucopis (Leucopis) aphidivora Rondani 1847, of which Leucopis fiorii Raspi 1986 is a new synonym; Leucopis (Leucopis) aphidiperda Rondani 1848; Leucopis (Leucopis) misaphida Rondani 1848; Leucopis (Leucopis) minuscula Rondani 1875; Leucopis (Leucopis) talaria Rondani 1875; Leucopomyia palliditarsis (Rondani 1875), of which Leucopis alticeps Czerny 1936 is a new synonym. Leucopis (Leucopis) armillata Rondani 1875 is a new synonym of Leucopis palumbii Rondani 1872, and Leucopis ballestrerii Rondani 1875 is a new synonym of Leucopomyia silesiaca (Egger 1862). Overall, concerning the twelve species of Chamaemyiidae described by Camillo Rondani, nine of them are still considered valid species. This study adds basic knowledge to silver fly systematics and has helpful implications concerning Chamaemyiidae-based biological control programs. -
Diptera: Syrphidae) Including the Description of a New Species
BLOTCH etal.: 13-36 Studia dipterologica 10 (2003) Heft 1 • ISSN 0945-3954 A review of the Mesoamerican flower flies of the genus Meromacrus (Diptera: Syrphidae) including the description of a new species [Eine Ubersicht der mittelamerikanischen Schwebfliegen der Gattung Meromacrus (Diptera: Syrphidae) nebst der Beschreibung einer neuen Art] by Sydella A. BLATCH, F. Christian THOMPSON and Manuel A. ZUMBADO College Park (USA) Washington (USA) Santo Domingo (Costa Rica) Abstract The Mesoamerican species of the genus Meromacrus are revised. A key to the nine species, along with synonymies, descriptions, distributions, and figures for all species are given. Meromacrus melansoni BLATCH spec, nov., is described and one new synonym is proposed (Meromacrus panamensis CURRAN, 1930 = Milesia laconica WALKER, 1852). Key words Flower flies, Syrphidae, Meromacrus, new species, Neotropis, key Zusammenfassung Die mittelamerikanischen Arten der Gattung Meromacrus werden revidiert. Ein Bestimmungs- schliissel zu den 9 Arten nebst der Synonymien, eine Auflistung der wichtigsten Merkmale und Abbildungen zu jeder Spezies sowie Angaben zu ihrer Verbreitung werden gegeben. Meromacrus melansoni BLATCH spec, nov., wird beschrieben und eine neue Synonymic wird vorgeschlagen (Meromacrus panamensis CURRAN, 1930 = Milesia laconica WALKER, 1852). Stichworter Schwebfliegen, Syrphidae, Meromacrus, neue Art, Neotropis, Bestimmungsschliissel Introduction Meromacrus flower flies are common pollinators as adults and filter-feeders as maggots in rot- holes in trees. As such they contribute to diverse fauna and are indicators of rich habitats. As part of the effort to document the biodiversity of Costa Rica, we here describe a new species of Meromacrus and provide additional details on the eight other Mesoamerican species. The genus Meromacrus is endemic to the New World the included species ranging from the southern United States to northern Argentina. -
Baccha (Ocyptamus) Medina, B
The Syrphidae of Puerto Rico1'2 H. S. Telford3-* One cannot state with certainty when the first syrphid was collected from Puerto Rico and adjacent islands. Fabricius described a number of 1 Manuscript submitted to Editorial Board October 30, 1972. 2 Scientific paper number 3914. College of Agriculture Research Center, Washing ton State University, Pullman, Washington. Work was conducted under Project No. 0046. 3 Professor and Entomologist, Department of Entomology, Washington State University; Visiting Scientist, Department of Entomology, Agricultural Experiment Station, Mayagiiez Campus, Uío Piedras, Puerto Rico, September 1968-March 1969. This study was made possible by financial support from the Department of En tomology, Agricultural Experiment Station, Mayagiiez Campus, University of Puerto Rico, Río Piedras. I wish to thank Dr. L. P. R. F. Martorell, formerly Chairman, Department of Entomology, Agricultural Experiment Station, especially for his support and aid in all aspects of the project. Mr. Silverio Medina Gaud, Associate Entomologist, Agricultural Experiment Station, was of considerable help. He ac companied me on almost all field trips, assisted in sorting and preparing the material and made valuable field trips on his own. Dr. J. R. Vockeroth, Entomology Research Institute, Ottawa, Canada, verified many determinations and offered advice on nomenclatural problems. Others who materially aided in the loan of specimens, verified determinations or in other ways were: Dr. George Drury, U.S. Atomic Energy Commission, El Verde-Caribbean National Forest, Puerto Rico; Dr. Y. S. Sedman, Western Illinois University; Dr. L. V. Knutson, Systematic Entomology Laboratory, Agricultural Research Service, United States Department of Agriculture; Dr. P. W. Wygodzinsky, American Museum of Natural History; Dr. -
Diptera): a Life History, Molecular, Morphological
The evolutionary biotogy of Conopidae (Diptera): A life history, molecular, morphological, systematic, and taxonomic approach Joel Francis Gibson B.ScHon., University of Guelph, 1999 M.Sc, Iowa State University, 2002 B.Ed., Ontario Institute for Studies in Education/University of Toronto, 2003 A thesis submitted to the Faculty of Graduate and Postdoctoral Affairs in partial fulfillment of the requirements for the degree of Doctor of Philosophy in Biology Carleton University Ottawa, Ontario © 2011 Joel Francis Gibson Library and Archives Bibliotheque et 1*1 Canada Archives Canada Published Heritage Direction du Branch Patrimoine de Pedition 395 Wellington Street 395, rue Wellington Ottawa ON K1A 0N4 Ottawa ON K1A 0N4 Canada Canada Your Tile Votre r&ference ISBN: 978-0-494-83217-2 Our file Notre reference ISBN: 978-0-494-83217-2 NOTICE: AVIS: The author has granted a non L'auteur a accorde une licence non exclusive exclusive license allowing Library and permettant a la Bibliotheque et Archives Archives Canada to reproduce, Canada de reproduire, publier, archiver, publish, archive, preserve, conserve, sauvegarder, conserver, transmettre au public communicate to the public by par telecommunication ou par I'lnternet, preter, telecommunication or on the Internet, distribuer et vendre des theses partout dans le loan, distribute and sell theses monde, a des fins commerciales ou autres, sur worldwide, for commercial or non support microforme, papier, electronique et/ou commercial purposes, in microform, autres formats. paper, electronic and/or any other formats. The author retains copyright L'auteur conserve la propriete du droit d'auteur ownership and moral rights in this et des droits moraux qui protege cette these. -
Implementation and Status of Biological Control of the Hemlock Woolly Adelgid
Forest Health Technology Enterprise Team TECHNOLOGY TRANSFER Biological Control IMPLEMENTATION AND STAtuS OF BIOLOGICAL CONTRol OF THE HEMLOCK WOOLLY AdelgId United States Forest Forest Health FHTET-2011-04 Department Service Technology December 2011 of Agriculture Enterprise Team Forest Health Technology Enterprise Team TECHNOLOGY TRANSFER Biological Control IMPLEMENTATION AND STAtuS OF BIOLOGICAL CONTRol OF THE HEMLOCK WOOLLY AdelgId United States Forest Forest Health FHTET-2011-04 Department Service Technology December 2011 of Agriculture Enterprise Team IMPLEMENTATION AND STATUS OF BIOLOGICAL CONTROL OF THE HEMLOCK WOOLLY ADELGID The Forest Health Technology Enterprise Team (FHTET) was created in 1995 by the Deputy Chief for State and Private Forestry, Forest Service, U.S. Department of Agriculture, to develop and deliver technologies to protect and improve the health of American forests. This book was published by FHTET as part of the technology transfer series. http://www.fs.fed.us/foresthealth/technology/ On the cover: Center photo: Hemlock woolly adelgid white woolly masses on a hemlock branch (USDA Forest Service, Karen Felton) Top right: Sasajiscymnus tsugae predatory beetle (USDA Forest Service, Lynn Jones); Middle right: Collecting and checking hemlock branch samples for Laricobius nigrinus predatory beetle larvae (USDA Forest Service, Brad Onken); Bottom center: Collecting Laricobius nigrinus predatory beetles in Idaho (USDA Forest Service, Brad Onken); Middle left: Releasing Laricobius nigrinus predatory beetles (USDA Forest Service, Brad Onken); Top left: Laricobius nigrinus predatory beetle (USDA Forest Service, Lynn Jones) For additional copies of this publication, contact: Brad Onken Richard Reardon U.S. Forest Service U.S. Forest Service 180 Canfi eld Street 180 Canfi eld Street Morgantown, WV 26505 Morgantown, WV 26505 (304) 285-1546 (304) 285-1566 [email protected] [email protected] The entire publication is available online at http://www.fs.fed.us/na/morgantown/fhp/hwa The U.S. -
Allometric Growth in the Skull of Tylosaurus Proriger (Squamata: Mosasauridae) and Its Taxonomic Implications Robert F
Vertebrate Anatomy Morphology Palaeontology 6:75–90 75 ISSN 2292-1389 Allometric growth in the skull of Tylosaurus proriger (Squamata: Mosasauridae) and its taxonomic implications Robert F. Stewart1 and Jordan C. Mallon2,* 1Department of Earth Sciences, Carleton University, Ottawa, Ontario, Canada, K1S 5B6; [email protected] 2Palaeobiology, Canadian Museum of Nature, PO Box 3223, Station D, Ottawa, Ontario, Canada, K1P 6P4; [email protected] Abstract: Ontogeny—the growth and development of an organism—is among the more poorly understood aspects of the life history of mosasaurs, largely owing to a dearth of fossil material from young individuals. We describe the par- tial and nearly complete skulls of two subadult individuals of the mosasaurid Tylosaurus proriger from the upper Smoky Hills Chalk Member of the Niobrara Formation (upper Santonian) in Kansas. We include the more complete of the two specimens in an allometric analysis to better understand proportional changes in the skull through growth. Although our small sample size produces several instances of ‘soft isometry’, we recover the length of the edentulous rostrum as significantly negatively allometric, and quadrate height as significantly positively allometric. In light of our findings, we consider the question of whether T. kansasensis represents an immature ontogimorph of T. nepaeolicus, and find substan- tive evidence to reject this hypothesis. INTRODUCTION Seaway of North America (Williston 1898; Russell 1967; Everhart 2017). These are among the smallest skulls known Mosasauridae is a clade of carnivorous, mostly marine for the species, and they help to elucidate the allometric reptiles known from Upper Cretaceous deposits world- changes undergone by T. proriger through life. -
A New Addition to the Cretaceous Seaway of ND
A New Addition to the Cretaceous Seaway of North Dakota Clint A. Boyd In July of 2015, 17-year-old Deborah Shepherd from Green Cove Springs, Florida was visiting the Pembina Gorge State Recreation Area in northeastern North Dakota (Cavalier County) with her family. One member of Deborah’s family had previously attended the Pembina Gorge public fossil dig, and they had brought the family up to the roadside marker near the fossil site to see the area. The group was exploring the area and had dispersed a bit when they heard Deborah excitedly call out. She came running up to the group holding a fist-sized piece of white bone encased in a crust of black shale (fig. 1). Along one side of the bone four large teeth were present. Deborah had found part of the jaw of an ancient sea monster: a mosasaur. Mosasaurs were large aquatic reptiles that lived in the oceans during the Mesozoic while dinosaurs were ruling the land. Though they lived at the same time as the dinosaurs, they are actually more closely related to snakes and monitor lizards (like the Komodo dragon) than they are to dinosaurs. They swam using four large flippers and an extremely long, stiff tail, and had to return to the surface to breathe (fig. 2), just like modern whales and dolphins. They were the top predators of the seas during their time, with some species reaching lengths Figure 2. Reconstruction of a mosasaur. Painting by Becky Barnes. of close to 50 feet and displaying teeth as large as whom, and took temporary possession of the fossil. -
Educator Guide
Educator Guide BACKGROUND INFORMATION Welcome to the world of the late Cretaceous Period, filled with huge carnivorous marine reptiles with double-hinged jaws and teeth in the middle of their palates. Come see gigantic flesh-eating fish big enough to swallow an adult human being whole, flying reptiles with 3-foot skulls, and the biggest sea turtles to have ever lived. Many bizarre and gigantic forms of life populated the prehistoric waters of the late Cretaceous Period. The Midwest was actually underwater at one time. Kansas has only been above sea level for the last 65 million years. Before that, it was home to a variety of sea creatures, including a 45-foot long mosasaur, a sea turtle the size of a small truck, a giant carnivorous fish, and a long-necked plesiosaur. Although these prehistoric marine animals lived during the time of Tyrannosaurus and Triceratops, they are not dinosaurs. Dinosaurs lived on land and did not have wings for flying or fins for swimming. Many Cretaceous marine fossils have been found in Western Kansas. These fossils have been found in thousands of feet of marine sediments made up of shale, chalk, limestone, and sandstone. Common Questions When was the Cretaceous period? The Cretaceous Period extended from 144 to 65 million years ago. What is a mosasaur? A mosasaur is a large marine lizard with a long body and paddle-like limbs. Mosasaurs are not dinosaurs. The chief feature that distinguishes them from dinosaurs is the great flexibility and power of their jaws. Unlike most monstrous reptiles of the past, they still have living relatives, the giant monitor lizards such as the Komodo Dragons. -
The Conopid Flies of California (Diptera)
Pbysocephala bwgessi (Williston). Courtesy E. S. Ross. 4 BULLETIN OF THE CALIFORNIA INSECT SURVEY VOLUME 6, NO. 2 THE CONOPID FLIES OF CALIFORNIA (DIPTERA) BY SIDNEY CAMRAS (Chicago, Illinois) and PAUL D. HURD, JR. (Department of Entomology and Parasitology, University of California, Berkeley) UNIVERSITY OF CALIFORNIA PRESS BERKELEYANDLOSANGELES 1957 BULLETIN OF THE CALIFORNIA INSECT SURVEY Editors: E. G. Linsley, S. B. Freeborn, P. D. Hurd, R. L. Ushger Volume 6, No. 2, pp. 19-50, 4 figures in text, 25 maps, frontis. Submitted by Editors, October 26, 1956 Issued Sept. 3, 1957 Price, 75 cents & UNIVERSITY OF CALIFORNIA PRESS BERKELEY AND LOS ANGELES CALIFORNIA CAMBRIDGE UNIVERSITY PRESS LONDON, ENGLAND PRINTED BY OFFSET IN THE UNITED STATES OF AMERICA THE CONOPID FLIES OF CALIFORNIA (Diptera) BY Sidney Camras and Paul D. Hurd, Jr. INTRODUCTION The conopid flies are characterized by the closed Zo~logy,Harvard College (M.C.Z.); California In- or narrowed first posterior cell of the wing (fig. 2) sect Survey, Department of Entomology and Para- and in all the Nearctic forms by the elongated sitology, University of California (C.I.S.); Mont A. proboscis. These flies, especially the members of Cazier, American Museum of Natural History the subfamily Conopinae which have the base of (A.M.N.H.); H. Dietrich, Department of Entomolcgy, the abdomen narrowed and somewhat thread-waisted, Cornell University (C.U.); H. J. Dybas, Chicago superficially resemble some of the wasps and cer- Museum of Natural History, (C.M.N.H.); G. F. fain asilids, bombyliids, syrphids, and tachinids. Knowlton, Utah State Agricultural College They ate found frequently around flowers.