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Araneae) Described by P © Naturwiss.-med. Ver. Innsbruck; download unter www.biologiezentrum.at Ber. nat.-med. Verein Innsbruck Band 90 S. 135 - 150 Innsbruck, Nov. 2003 A critical review of the spider species (Araneae) described by P. Drensky in the period 1915-1942 from the Balkans by Christo DELTSHEV*) Synopsis: Acritical review of the spider species (Araneae) described by P. DRENSKY in the period 1910-1942 is presented. All published and unpublished results of revisions of material in DRENSKY's collection have been used in the present study to clarify the difficult taxonomical situati- on, caused by insufficient descriptions and loss of materials. From two new genera and 48 new spe- cies and subspecies described by DRENSKY only 18 taxa are currently regarded as valid, 6 others stand as species inquirendae. Three new synonyms are proposed: Echemus babunaensis DRENSKY, 1929 = Zelotes tenuis (L. KOCH, 1866), Theridion perestirensis DRENSKY, 1929 = Th. nigrovariegatum SIMON, 1873; Zora affinis DRENSKY, 1929 = Clubiona comta C. L. KOCH, 1839. 1. Introduction: DRENSKY started his studies about the Bulgarian spider fauna in 1910. Altogether he described two new genera and 48 new species and subspecies (DRENSKY 1910, 1915, 1921, 1929, 1931, 1935, 1938, 1940, 1942). The spiders were collected from the territory of the Balkan Peninsula (Bulgaria, Greece, Macedonia, Turkey). The descriptions of most taxa are not well done, accompanied by insufficient and not precise drawings without scale. In many cases types were single specimens only. Some types were lost during the war. Until now types of the following species could not be found and are probably lost: Cyclosa strandjae (1915), Comaroma ressenensis (1929), Gonatium strugaense (1929), Porrhom- ma fons-frigidus (1929), Echemus babunaensis (1929), Troglohyphantes kratochvili (1935), Macedoniella karamani (1935), Mesioteles cyprius scopensis (1935), Coelotes pastor pirinicus (1942). All results of revisions of the material in DRENSKY's collection, published and unpublished, have been used in the present study to clarify the complicated taxonomical situation. Nomenclature follows the internet catalogue of PLATNICK (2002). Species are discussed in chronological order. *) Anschrift des Verfassers: Prof. Dr. Christo Deltshev, Institute of Zoology, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd., 1000 Sofia, Bulgaria; e-mail [email protected]. 135 © Naturwiss.-med. Ver. Innsbruck; download unter www.biologiezentrum.at 2. Results and discussion: Junior synonyms Cyclosa strandjae DRENSKY, 1915 = C. conica (PALLAS, 1772) (Araneidae) DIMITROV (1999: 23, f. 1), DELTSHEV & BLAGOEV (2001: 110, 111). Type material was not found. The figures given by DRENSKY (1915) suggest that this spe- cies has been described from a subadult specimen (DIMITROV 1999). Amaurobius Jurinitschi var. flavum DRENSKY, 1915 = Coelotes jurinitschi (DRENSKY, 1915) (Amaurobiidae) DELTSHEV (1990: 33), DELTSHEV & BLAGOEV (2001: 110, 111). Type material was examined (5 females). Amaurobius Jurinitschi var. niger DRENSKY, 1915 = Coelotes jurinitschi (DRENSKY, 1915) (Amaurobiidae) DELTSHEV (1990: 33), DELTSHEV & BLAGOEV (2001: 110, 111). Type material was examined (1 female). Micaria turcica DRENSKY, 1915 = M. albovittata (LUCAS, 1846) (Gnaphosidae) MIKHAILOV (1991: 78, sub M. romana, DELTSHEV & BLAGOEV (2001: 110, 111, sub M. romana). Type material was examined (1 female). Theridium cornutum JURINITCH & DRENSKY, 1917 = Theridion impressum L. KOCH, 1881 (Theridiidae) DELTSHEV (1992: 17, 22), DELTSHEV & BLAGOEV (2001:115). Type material was examined (1 female). DRENSKY (1939) wrote incorrectly Th. cornutum DRENSKY, 1917. Heliophanus flavipes var. nigripes JURINITCH & DRENSKY, 1917 = H. flavipes (HAHN, 1832) (Salticidae) DELTSHEV & BLAGOEV (2001: 110, 111), PLATNICK (2002 ). Type material was examined (1 immature individual). Theridion pirini DRENSKY, 1921 = Th. petraeum L. KOCH, 1872 (Theridiidae) DELTSHEV (1992: 20), DELTSHEV & BLAGOEV, (2001: 110, 111). Type material was examined (4 females, 3 juv.). Lepthyphantes pirini DRENSKY, 1929 = L. quadrimaculatus KULCZYNSKI, 1897 (Linyphiidae) DELTSHEV (1988a: 52), DELTSHEV & BLAGOEV (2001: 110, 111). Type material was examined (1 female). 136 © Naturwiss.-med. Ver. Innsbruck; download unter www.biologiezentrum.at Linyphia phrigiana var. pirini DRENSKY, 1921 = Pityohyphantes phrygianus (C.L. KOCH, 1836) (Linyphiidae) (Figs. 1-4) DELTSHEV & BLAGOEV (2001: 110, 111). Type material was examined (2 males, 26 females) and new figures are presented here (Figs. 3, 4). Drassus pirini DRENSKY, 1921 = Drassodes lapidosus (WALCKENAER, 1802) (Gnaphosidae) (Figs. 5-7) DELTSHEV & BLAGOEV (1997: 281), DELTSHEV & BLAGOEV (2001: 110, 111). Type material was examined (1 female, 2 juv.) and new figures are presented here (Figs. 6- 7). Echemus babunaensis DRENSKY, 1929 = Zelotes tenuis (L. KOCH, 1866), syn. nov. (Gnaphosidae) (Figs. 8-10) DRENSKY (1929) incorrectly used the spelling E. babuni in his explanation to plate 1, figs. 3-4. Type material was not found. The original description and figure (fig. 3) correspond well with Zelotes tenuis. New figures of Bulgarian material of Z. tenuis are given (Figs. 9, 10). Dipoena regia DRENSKY, 1929 = Neottiura suaveolens (SIMON, 1879) (Theridiidae) WUNDERLICH (1977: 289), KNOFLACH (1999: 357). In the description of this species DRENSKY (1929) used various spellings: regia (p. 33), regina (p. 69), regalis (p. 76). Type material was not found. Theridium peristerensis DRENSKY, 1929 = Theridion nigrovariegatum SIMON, syn. nov. (Theridiidae) (Fig. 11-12) DRENSKY (1929: 76) used another spelling: Th. Peristeri. The type (1 female) was examined and a new figure is presented (Fig. 12). Comaroma ressenensis DRENSKY, 1929 = Spermophora senoculata (DUGES, 1836) (Pholcidae) (Fig. 13-16) DELTSHEV & BLAGOEV (2001: 110, 111). DRENSKY (1929: 76) also wrote C. ressnen- sis). Type material was not found. The description and figures fit well to S. senoculata (see Figs 13- 16). Hogna (Tarentula) macedonica DRENSKY, 1929 = H. radiata (LATREILLE, 1817) (Lycosidae) (Fig. 17-18) DELTSHEV & BLAGOEV (2001). Type material was examined (2 subadult females) and a new illustration is presented. The 137 © Naturwiss.-med. Ver. Innsbruck; download unter www.biologiezentrum.at 1 3 2 4 5 6 7 Figs. 1-7: 1-4 Pityohyphantes phrygianus (C.L. KOCH, 1836): 1 tibial apophysis; 2, 4 epigyne; 3 male palp, external view (Figs. 1-2 from DRENSKY 1921, Linyphia phrigiana var. pirini; Figs. 3-4 drawn from types). - 5-7: Drassodes lapidosus (WALCKENAER, 1802): 5 epigyne (from DRENSKY 1921, Drassus pirini), 6-7 vulva, ventral and dorsal view (type). Scale: 0.3 mm (3,4,6,7). 138 © Naturwiss.-med. Ver. Innsbruck; download unter www.biologiezentrum.at 8 9 11 10 12 Figs. 8-12: 8-10 Zelotes tenuis (L. KOCH, 1866): 8 epigyne (from DRENSKY 1929, Echemus babuna- ensis); 9-10 epigyne/vulva, ventral and dorsal view (specimen from Bulgaria). - Theridion nigrova- riegatum SIMON, 1873: 11 epigyne (from DRENSKY 1929, Th. peristerensis), 12 vulva (type). Scales: 0.1 (12); 0.2 mm (9, 10) 139 © Naturwiss.-med. Ver. Innsbruck; download unter www.biologiezentrum.at 13 14 16 15 Figs. 13-16: Spermophora senoculata (DUGES, 1836): epigyne/vulva, 13 (from DRENSKY 1929, Comaroma ressenensis), 14 (specimen from Bulgaria), 15 (from MILLER & ZITNANSKA 1976), 16 (from DRESCO & HUBERT 1989). Scale: 0.2 mm (14). 140 © Naturwiss.-med. Ver. Innsbruck; download unter www.biologiezentrum.at 17 18 19 20 21 22 Figs. 17-22: Hogna radiata (LATREILLE, 1817): 17-18 epigyne/vulva (subadult specimen), 17 (from DRENSKY 1929, H. (Tarentula) macedonica), 18 (type). - Callilepis nocturna (LINNAEUS, 1758): 19- 22, male palp, expanded, 19-20 (from DRENSKY 1929, Poecilochroa ochridana), 21-22 (types). Scales: 0.1 (18); 0.3 mm (21-22). 141 © Naturwiss.-med. Ver. Innsbruck; download unter www.biologiezentrum.at epigyne/vulva is not yet fully developed (Fig. 18). Shape and pattern of cephalothorax and abdomen closely resemble H. radiata. Tarentula Konstantinovi DRENSKY, 1929 = Alopecosa sulzeri (PAVESI, 1873) (Lycosidae) LUGETTI & TONGIORGI (1969: 67), DELTSHEV & BLAGOEV (2001: 110, 111). Type material (1 male, 3 females, 1 juv) was examined and corresponds well with A. sul- zeri. Agroeca exortus DRENSKY, 1929 = Robertus frivaldskyi (CHYZER, 1894) (Theridiidae) KNOFLACH & THALER (2000: 433). DRENSKY (1936) changed the species name to A. exorta , according to gender of the genus name. Type material was examined (1 male). Poecilochroa ochridana DRENSKY, 1929 = Callilepis nocturna (LINNAEUS, 1758), syn. nov. (Gnaphosidae) (Figs. 19-22) Types (2 males with expanded palps) were examined. The new synonomy is obvious (Figs. 21-22). Prosthesima bitolensis DRENSKY, 1929 = Zelotes gracilis (CANESTRINI, 1869) (Gna- phosidae) (Figs. 23-24) DELTSHEV & BLAGOEV (2001: 110, 111). The type (1 female) was examined and a new illustration is presented (Fig. 24). Zora affinis DRENSKY, 1936 = Clubiona comta C. L. KOCH, 1839, syn. nov. (Clubionidae) (Figs. 25-27) This species was first tentativelly assigned by DRENSKY (1929: 20, 66, Zora sp. ?) to this genus or to Z. manicata respectively (1929: 76, tab. 2, fig. 3. Later he renamed it as a full species (1936: 131, Zora affinis). 1936 therefore must stand as the year of its formal introduction to science. Type material was not found. The original description and figures fit well to C. comta, see Figs. 25-27. Lepthyphantes slivnensis DRENSKY, 1931 = L. istrianus (KULCZYNSKI, 1914) (Liny- phiidae) DEELEMAN-REINHOLD (1985: figs: 10-15), DELTSHEV
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