Araneae) Described by P

Ber. nat.-med. Verein Innsbruck Band 90 S. 135 - 150 Innsbruck, Nov. 2003

A critical review of the spider species (Araneae) described by P. Drensky in the period 1915-1942 from the Balkans

Christo DELTSHEV*)

Synopsis: Acritical review of the spider species (Araneae) described by P. DRENSKY in the period 1910-1942 is presented. All published and unpublished results of revisions of material in DRENSKY's collection have been used in the present study to clarify the difficult taxonomical situation, caused by insufficient descriptions and loss of materials. From two new genera and 48 new species and subspecies described by DRENSKY only 18 taxa are currently regarded as valid, 6 others stand as species inquirendae. Three new synonyms are proposed: Echemus babunaensis DRENSKY, 1929 = Zelotes tenuis (L. KOCH, 1866), Theridion perestirensis DRENSKY, 1929 = Th. nigrovariegatum SIMON, 1873; Zora affinis DRENSKY, 1929 = Clubiona comta C. L. KOCH, 1839.

1. Introduction: DRENSKY started his studies about the Bulgarian spider fauna in 1910. Altogether he described two new genera and 48 new species and subspecies (DRENSKY 1910, 1915, 1921, 1929, 1931, 1935, 1938, 1940, 1942). The spiders were collected from the territory of the Balkan Peninsula (Bulgaria, Greece, Macedonia, Turkey). The descriptions of most taxa are not well done, accompanied by insufficient and not precise drawings without scale. In many cases types were single specimens only. Some types were lost during the war. Until now types of the following species could not be found and are probably lost: Cyclosa strandjae (1915), Comaroma ressenensis (1929), Gonatium strugaense (1929), Porrhom- ma fons-frigidus (1929), Echemus babunaensis (1929), Troglohyphantes kratochvili (1935), Macedoniella karamani (1935), Mesioteles cyprius scopensis (1935), Coelotes pastor pirinicus (1942). All results of revisions of the material in DRENSKY's collection, published and unpublished, have been used in the present study to clarify the complicated taxonomical situation. Nomenclature follows the internet catalogue of PLATNICK (2002). Species are discussed in chronological order.

*) Anschrift des Verfassers: Prof. Dr. Christo Deltshev, Institute of Zoology, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd., 1000 Sofia, Bulgaria; e-mail [email protected].

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2. Results and discussion: Junior synonyms Cyclosa strandjae DRENSKY, 1915 = C. conica (PALLAS, 1772) (Araneidae) DIMITROV (1999: 23, f. 1), DELTSHEV & BLAGOEV (2001: 110, 111). Type material was not found. The figures given by DRENSKY (1915) suggest that this species has been described from a subadult specimen (DIMITROV 1999).

Amaurobius Jurinitschi var. flavum DRENSKY, 1915 = Coelotes jurinitschi (DRENSKY, 1915) (Amaurobiidae) DELTSHEV (1990: 33), DELTSHEV & BLAGOEV (2001: 110, 111). Type material was examined (5 females).

Amaurobius Jurinitschi var. niger DRENSKY, 1915 = Coelotes jurinitschi (DRENSKY, 1915) (Amaurobiidae) DELTSHEV (1990: 33), DELTSHEV & BLAGOEV (2001: 110, 111). Type material was examined (1 female).

Micaria turcica DRENSKY, 1915 = M. albovittata (LUCAS, 1846) (Gnaphosidae) MIKHAILOV (1991: 78, sub M. romana, DELTSHEV & BLAGOEV (2001: 110, 111, sub M. romana). Type material was examined (1 female).

Theridium cornutum JURINITCH & DRENSKY, 1917 = Theridion impressum L. KOCH, 1881 (Theridiidae) DELTSHEV (1992: 17, 22), DELTSHEV & BLAGOEV (2001:115). Type material was examined (1 female). DRENSKY (1939) wrote incorrectly Th. cornutum DRENSKY, 1917.

Heliophanus flavipes var. nigripes JURINITCH & DRENSKY, 1917 = H. flavipes (HAHN, 1832) (Salticidae) DELTSHEV & BLAGOEV (2001: 110, 111), PLATNICK (2002 ). Type material was examined (1 immature individual).

Theridion pirini DRENSKY, 1921 = Th. petraeum L. KOCH, 1872 (Theridiidae) DELTSHEV (1992: 20), DELTSHEV & BLAGOEV, (2001: 110, 111). Type material was examined (4 females, 3 juv.).

Lepthyphantes pirini DRENSKY, 1929 = L. quadrimaculatus KULCZYNSKI, 1897 (Linyphiidae) DELTSHEV (1988a: 52), DELTSHEV & BLAGOEV (2001: 110, 111). Type material was examined (1 female).

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Linyphia phrigiana var. pirini DRENSKY, 1921 = Pityohyphantes phrygianus (C.L. KOCH, 1836) (Linyphiidae) (Figs. 1-4) DELTSHEV & BLAGOEV (2001: 110, 111). Type material was examined (2 males, 26 females) and new figures are presented here (Figs. 3, 4).

Drassus pirini DRENSKY, 1921 = Drassodes lapidosus (WALCKENAER, 1802) (Gnaphosidae) (Figs. 5-7) DELTSHEV & BLAGOEV (1997: 281), DELTSHEV & BLAGOEV (2001: 110, 111). Type material was examined (1 female, 2 juv.) and new figures are presented here (Figs. 6- 7).

Echemus babunaensis DRENSKY, 1929 = Zelotes tenuis (L. KOCH, 1866), syn. nov. (Gnaphosidae) (Figs. 8-10) DRENSKY (1929) incorrectly used the spelling E. babuni in his explanation to plate 1, figs. 3-4. Type material was not found. The original description and figure (fig. 3) correspond well with Zelotes tenuis. New figures of Bulgarian material of Z. tenuis are given (Figs. 9, 10).

Dipoena regia DRENSKY, 1929 = Neottiura suaveolens (SIMON, 1879) (Theridiidae) WUNDERLICH (1977: 289), KNOFLACH (1999: 357). In the description of this species DRENSKY (1929) used various spellings: regia (p. 33), regina (p. 69), regalis (p. 76). Type material was not found.

Theridium peristerensis DRENSKY, 1929 = Theridion nigrovariegatum SIMON, syn. nov. (Theridiidae) (Fig. 11-12) DRENSKY (1929: 76) used another spelling: Th. Peristeri. The type (1 female) was examined and a new figure is presented (Fig. 12).

Comaroma ressenensis DRENSKY, 1929 = Spermophora senoculata (DUGES, 1836) (Pholcidae) (Fig. 13-16) DELTSHEV & BLAGOEV (2001: 110, 111). DRENSKY (1929: 76) also wrote C. ressnen- sis). Type material was not found. The description and figures fit well to S. senoculata (see Figs 13- 16).

Hogna (Tarentula) macedonica DRENSKY, 1929 = H. radiata (LATREILLE, 1817) (Lycosidae) (Fig. 17-18) DELTSHEV & BLAGOEV (2001). Type material was examined (2 subadult females) and a new illustration is presented. The

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1 3

4 5

6 7

Figs. 1-7: 1-4 Pityohyphantes phrygianus (C.L. KOCH, 1836): 1 tibial apophysis; 2, 4 epigyne; 3 male palp, external view (Figs. 1-2 from DRENSKY 1921, Linyphia phrigiana var. pirini; Figs. 3-4 drawn from types). - 5-7: Drassodes lapidosus (WALCKENAER, 1802): 5 epigyne (from DRENSKY 1921, Drassus pirini), 6-7 vulva, ventral and dorsal view (type). Scale: 0.3 mm (3,4,6,7).

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8 9

11 10

Figs. 8-12: 8-10 Zelotes tenuis (L. KOCH, 1866): 8 epigyne (from DRENSKY 1929, Echemus babunaensis); 9-10 epigyne/vulva, ventral and dorsal view (specimen from Bulgaria). - Theridion nigrovariegatum SIMON, 1873: 11 epigyne (from DRENSKY 1929, Th. peristerensis), 12 vulva (type). Scales: 0.1 (12); 0.2 mm (9, 10)

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16 15

Figs. 13-16: Spermophora senoculata (DUGES, 1836): epigyne/vulva, 13 (from DRENSKY 1929, Comaroma ressenensis), 14 (specimen from Bulgaria), 15 (from MILLER & ZITNANSKA 1976), 16 (from DRESCO & HUBERT 1989). Scale: 0.2 mm (14).

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17 18

19 20

21 22

Figs. 17-22: Hogna radiata (LATREILLE, 1817): 17-18 epigyne/vulva (subadult specimen), 17 (from DRENSKY 1929, H. (Tarentula) macedonica), 18 (type). - Callilepis nocturna (LINNAEUS, 1758): 19- 22, male palp, expanded, 19-20 (from DRENSKY 1929, Poecilochroa ochridana), 21-22 (types). Scales: 0.1 (18); 0.3 mm (21-22).

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epigyne/vulva is not yet fully developed (Fig. 18). Shape and pattern of cephalothorax and abdomen closely resemble H. radiata.

Tarentula Konstantinovi DRENSKY, 1929 = Alopecosa sulzeri (PAVESI, 1873) (Lycosidae) LUGETTI & TONGIORGI (1969: 67), DELTSHEV & BLAGOEV (2001: 110, 111). Type material (1 male, 3 females, 1 juv) was examined and corresponds well with A. sulzeri.

Agroeca exortus DRENSKY, 1929 = Robertus frivaldskyi (CHYZER, 1894) (Theridiidae) KNOFLACH & THALER (2000: 433). DRENSKY (1936) changed the species name to A. exorta , according to gender of the genus name. Type material was examined (1 male).

Poecilochroa ochridana DRENSKY, 1929 = Callilepis nocturna (LINNAEUS, 1758), syn. nov. (Gnaphosidae) (Figs. 19-22) Types (2 males with expanded palps) were examined. The new synonomy is obvious (Figs. 21-22).

Prosthesima bitolensis DRENSKY, 1929 = Zelotes gracilis (CANESTRINI, 1869) (Gna- phosidae) (Figs. 23-24) DELTSHEV & BLAGOEV (2001: 110, 111). The type (1 female) was examined and a new illustration is presented (Fig. 24).

Zora affinis DRENSKY, 1936 = Clubiona comta C. L. KOCH, 1839, syn. nov. (Clubionidae) (Figs. 25-27) This species was first tentativelly assigned by DRENSKY (1929: 20, 66, Zora sp. ?) to this genus or to Z. manicata respectively (1929: 76, tab. 2, fig. 3. Later he renamed it as a full species (1936: 131, Zora affinis). 1936 therefore must stand as the year of its formal introduction to science. Type material was not found. The original description and figures fit well to C. comta, see Figs. 25-27.

Lepthyphantes slivnensis DRENSKY, 1931 = L. istrianus (KULCZYNSKI, 1914) (Liny- phiidae) DEELEMAN-REINHOLD (1985: figs: 10-15), DELTSHEV & BLAGOEV (2001: 118). Type material was examined (1 male).

Troglohyphantes Bureschi DRENSKY, 1931 = Lepthyphantes centromeroides KULC- ZYNSKI, 1914 (Linyphiidae) DELTSHEV (1972: 143), DELTSHEV & BLAGOEV (2001: 110, 111). Type material was examined (2 males, 6 females).

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Euxinella strandi DRENSKY, 1938 = Nurscia albosignata SIMON, 1874 (Titanoecidae) LEHTINEN (1967: 253), DELTSHEV & BLAGOEV (2001:127). Type material was examined (1 female).

Nomen nudum Araneus riloensis DRENSKY, 1934 (Araneidae) DELTSHEV & BLAGOEV (2002: 110, 111).

Species inquirendae Lepthyphantes melanotus DRENSKY, 1921 = L. nebulosus var. melanotus (DRENSKY, 1921) (Linyphiidae) WUNDERLICH (1977: 59, figs. 7-8 redrawn from original illustrations, elevated from subspecies). Type material was not found.

Porrhomma fons-frigidus DRENSKY, 1929 (Linyphiidae) DRENSKY (1935), THALER (1968: 384). Type material was not found.

Gnaphosa expilator DRENSKY, 1929 (Gnaphosidae) (Figs. 28-29) DRENSKY (1955: 362, G. explicatrix, misspelling). Type material (1 subadult female) was examined and new figures are presented. As this is a subadult specimen, its epigyne/vulva is not yet developed. Therefore the species must be reinvestigated.

Pterotricha exitiabilis DRENSKY, 1929 (Gnaphosidae) In the explication to tab. 1, fig. 8-9, DRENSKY (1929) wrote P. extitiabilis, which must be regarded as a misspelling. Type material is a female cephalothorax only. Therefore the species awaits rediscovery and redescription.

Coelotes pastor pirinicus DRENSKY, 1942 (Amaurobiidae) DELTSHEV (1990a: 42). Type material was not found. DRENSKY’s description is very short and does not allow safe recognition. In recent collections from the type region C. kulczynskii (DRENSKY, 1915) only was discovered.

Macedoniella karamani DRENSKY, 1935 (Liocranidae) Type material was not found. Taxonomic status of this new genus and species need confir- mation and must be reinvestigated.

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23 24

25 27

Figs. 23-27: 23-24 Zelotes gracilis (CANESTRINI, 1869): 23 epigyne (from DRENSKY 1929, Prosthesima bitolensis); 24 vulva (type). – 25-27 Clubiona comta C. L. KOCH, 1839: epigyne, 25 (from DRENSKY 1929, Zora affinis); 26 (specimen from Bulgaria); 27 (from WIEHLE 1965). Scales: 0.1 (24); 0.2 mm (26).

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28 29

30 31

Figs. 28-32: 28-29 Gnaphosa expilator DRENSKY, 1929: epigyne (subadult specimen), 28 from DRENSKY (1929); 29 new figure. – 30-32 Zora prespaensis DRENSKY, 1929: 30 epigyne (from DRENSKY 1929); 31-32 vulva, ventral and dorsal view (type). Scale: 0.1 mm (29, 31, 32).

Valid species Theridium adrianopoli DRENSKY, 1915 = Theridion adrianopoli (Theridiidae) DELTSHEV (1992: 117, f. 1-3), KNOFLACH (1997: 133, f. 1-12), DELTSHEV & BLAGOEV (2001: 115). Type material was examined (1 female).

Amaurobius jurinitschi DRENSKY, 1915 = Coelotes jurinitschi (DRENSKY, 1915) (Amaurobiidae) DRENSKY (1915: 156; A. j. var. flavus), DRENSKY (1915: 156; A. j. var. niger), DRENSKY (1942: 42, f 5; Coelotes j.), DELTSHEV (1990: 33, f. 4.1-4;), DELTSHEV & BLAGOEV (2001: 127, C. j.). Type material was examined (4 females).

Amaurobius Kulczynsky DRENSKY, 1915 = Coelotes kulczynskii (DRENSKY, 1915) (Amaurobiidae) DRENSKY (1939: 86), DRENSKY (1942: 41, fig. 5e; Coelotes K.), DELTSHEV (1990: 36, f. 6.1-6; C. kulczynskii), DELTSHEV & BLAGOEV (2001: 127; C. k.). Type material was examined (2 females).

Hahnia pirini DRENSKY, 1921 = Cryphoeca pirini (DRENSKY, 1921) (Hahniidae) DRENSKY (1942: 56, f. 19; Cryphoeca p.), DELTSHEV & BLAGOEV (2001: 126; C. p.). Type material was examined (1 female).

Chiracanthium macedoinica DRENSKY, 1921 = Cheiracanthium macedonicum DRENSKY, 1921 (Clubionidae) DELTSHEV & BLAGOEV (2001: 127). Type material was examined (1 female). This species is closely related to Ch. striolatum SIMON, 1878. Both species should be thoroughly compared.

Zodarium gallicum var. pirini DRENSKY, 1921 = Z. pirini DRENSKY, 1921 (Zodariidae) DRENSKY (1940: 173, f. 3a), DELTSHEV (1987: 20, fig. 2.1-3, 3.1-3, Z. p., elevated from subspecies), DELTSHEV & BLAGOEV (2001: 128). Type material was examined (7 females).

Theridium refugus DRENSKY, 1929 = Theridion refugum DRENSKY, 1929 (Theridiidae) DELTSHEV (1992: 20, fig. 1-3), KNOFLACH (1998: 589, fig. 46, 52, 55d), DELTSHEV & BLAGOEV (2001: 115). Type material was examined (3 females).

Gonatium strugaensis DRENSKY, 1929 = Gonatium (?) strugaense DRENSKY, 1929 (Linyphiidae: Erigoninae)

Type material was not found. The figures given by DRENSKY for the male palp suggest that this species is misplaced in Gonatium (MILLIDGE 1981). The species needs revision.

Zora prespaensis DRENSKY, 1929 (Zoridae) (Figs. 30-32) Type material (1 female) was examined and new figures are presented (Figs. 31-32). The species is related to Zora spinimana (SUNDEVALL, 1833).

Lepthyphantes monticola balcanica DRENSKY, 1931 = Antrohyphantes balcanicus (Drensky, 1931) (Linyphiidae) DELTSHEV (1972: 137, fig. 1-8; L. balcanica, elevated to species rank), DELTSHEV (1988b: 296, fig. 10-14, 27; Antrohyphantes balcanica), DELTSHEV & BLAGOEV (2001: 116, A .b.). Types (3 females) were examined.

Troglohyphantes rhodopensis DRENSKY 1931 = Antrohyphantes rhodopensis (DRENSKY, 1931) (Linyphiidae) DUMITRESCU (1971: 168, fig. 1-13; Antrohyphantes rodopicus nov. gen. n. sp.), DELTSHEV (1975: 213, Lepthyphantes (A.) rhodopensis), DELTSHEV (1988b: 297, fig. 15-19, 28; A. rodopensis), DELTSHEV & BLAGOEV (2001: 116, A.r.). Type material (2 females) was examined.

Bolyphantes sophianus DRENSKY, 1931 = Antrohyphantes sophianus (DRENSKY, 1931) (Linyphiidae) DRENSKY (1931) also wrote B. sofianus which must be regarded as misspelling. MILLER (1958: 578, fig. 48-55; L. tranteevi), DELTSHEV (1972: 145, Lepthyphantes sophianus), BRIGNOLI (1983: 287, A. s.), DELTSHEV (1988b: 299, figs. 21-24, 29), DELTSHEV & BLAGOEV (2001: 116). Type material (1 male, 1 female) was examined.

Troglohyphantes bulgarianus DRENSKY, 1931 = Centromerus bulgarianus (DRENSKY, 1931) (Linyphiidae) KRATOCHVÍL & MILLER (1938: 111, fig. 2; C. b.), DELTSHEV (1973: 117, figs. 1-4, C.b.) DELTSHEV & CURCIC (1997: 54, figs. 3, 6, 9, 12; 2002: 170, fig. 5.), DELTSHEV & BLAGOEV (2001: 118). Type material was examined (1 male, 3 females).

Troglohyphantes lakanikensis DRENSKY, 1931 = Centromerus lakatnikensis (DRENSKY, 1931) (Linyphiidae) KRATOCHVÍL & MILLER (1938: 112, C. l), DELTSHEV (1973: 121, figs. 5-10; 1983a: 54, pl. 1, figs. 5-6), DELTSHEV & BLAGOEV (2001: 116). Type material was examined (1 male, 3 females).

Troglohyphantes trnovensis DRENSKY, 1931 = Lepthyphantes trnovensis (DRENSKY, 1931) (Linyphiidae) KRATOCHVÍL & MILLER (1938: 108, L. t.), DELTSHEV (1972: 145, fig. 10, sub L. spela- eorum), MILLER & SVATON (1978: 13, pl. VI, figs. 4-5, L. t.), DELTSHEV (1980: 50, pl. 6), DEELEMAN-REINHOLD (1985: 42, figs. 20-26), DELTSHEV & BLAGOEV (2001: 118). Type material was examined (2 females).

Troglohyphantes kratochvili DRENSKY, 1935 (Linyphiidae) DEELEMAN-REINHOLD (1978: 150, figs. 62a-h, 63d-g, 65i). Type material was not found.

Mesiotelus cyprius scopensis DRENSKY, 1935 (Liocranidae) HADJISSARANTOS (1940: 96, figs. 32a-b). Type material was not found.

Amaurobius claustrarius balcanicus DRENSKY, 1940 = Callobius claustrarius balcanicus (DRENSKY, 1940) (Amaurobiidae) DELTSHEV & BLAGOEV (2001: 126, C. c.b.). Type material was examined (1 male, 7 females).

Acknowledgements: I am especially indebted to my colleagues Gergin Blagoev and Stoyan Lazarov for discussion and helpful assistance and to Boris Andrev for preparing of the illustrations of Drensky's figures.

3. References: DEELEMAN-REINHOLD, C. (1978): Revision of the cave-dwelling and related spiders of the genus Troglohyphantes Joseph (Linyphiidae), with special reference to the Yugoslav species. - Acad. Scient. Artium Slovenica, IV: Historia naturalis 23: 1 - 219. - (1985): Contribution à la connaissance des Lepthyphantes du groupe pallidus (Araneae, Linyphiidae) de Yugoslavie, Grece et Chypre. - Mém. Biospéol. 12: 37 - 50. DELTSHEV, C. (1972): A contribution to the study of spiders (Araneae) from the caves in Bulgaria. II. Genus Lepthyphantes in Bulgarian caves. - Izv. zool. Inst. Sof. 36: 137 - 147. -(1973): Redescription of Centromerus bulgarianus (Drensky, 1931) and Centromerus lakatnikensis (Drensky, 1931) (Araneae, Linyphiidae). - Int. J. Speleol. 5: 117 - 126. - (1975): The genus Lepthyphantes in Bulgarian caves. - Proc. 6th int. arachnol. Congr., Amsterdam: 210 - 213. - (1980): A contribution to the taxonomical study of pallidus group of genus Lepthyphantes Menge (Araneae, Linyphiidae) in Bulgaria. - Acta zool. bulg. 16: 44 - 56. - (1983a): A contribution to the study of sylvaticus group of genus Centromerus F. Dahl (Araneae, Linyphiidae) in Bulgaria. - Acta zool. bulg. 21: 53 - 58. - (1983b): A contribution to the taxonomical and faunistic study of genus Lepthyphantes Menge (Araneae, Linyphiidae) from Pirin Mountain. - Acta zool. bulg. 23: 25 - 32. - (1987): A critical review of genus Zodarion Walckenaer (Araneae, Zodariidae) in Bulgaria. - Acta zool. bulg. 33: 19 - 25.

- (1988a): A contribution to the study of genus Lepthyphantes Menge (Araneae, Linyphiidae) from the Pirin Mountain with a description of a new species (Lepthyphantes rectilamellus sp. n.). - Acta zool. bulg. 36: 52 - 55. - (1988b): The genus Fageiella Kratochvil and the genus Antrohyphantes Dumitresco (Araneae, Linyphiidae, Lepthyphanteae) in the caves of Balkan peninsula. - In J. Haupt (ed.), TUB- Dokument. Kongr. Tagung. Berlin 38: 293 - 299. - (1990): A critical review of genus Coelotes Blackwall in Bulgaria with description of a new species (Coelotes drenskii sp. n., Araneae, Agelenidae). - Acta zool. bulg. 40: 29 - 44. - (1992): A critical review of family Theridiidae (Araneae) in Bulgaria. - Acta zool. bulg. 43: 13 - 22. DELTSHEV, C. & G. BLAGOEV (1997): The spiders of Pirin Mountain (Bulgaria). Taxonomic, faunistic and zoogeographical analysis (Araneae). - Ber. nat.-med. Verein Innsbruck 84: 269 - 286. - (2001): A check list of Bulgarian spider (Araneae). - Bull. Br. arachnol. Soc. 12: 110 - 138. DIMITROV, D. (1999): The spider fauna of the Strandzha Mountain (South-East Bulgaria). I. Faunistic data and taxonomic re-marks (Arachnida: Araneae). - Acta zool. bulg. 52: 15 - 26. DRENSKY, P. (1910): [Die Gattung Tarentula (Sund.) in Bulgarien]. - Trudy russk. ent. Obsch. (= Horae Soc. Ent. Ross.) 39: 411 - 414. - (1915): [Araneides nouveaux ou peu connus de Bulgarie]. - Spis. bulg. Akad. Nauk 12: 141 - 176. - (1921): [Contribution à 1'étude des araignées de la Macédoine orientale et de Pirine planina]. - Spis. bulg. Akad. Nauk 23: 1 - 50. - (1929): [Spinnen (Araneae) aus Mittel und Süd-West Mazedonien]. - Spis. bulg. Akad. Nauk 39: 1 - 76. - (1931): [Höhlen-Spinnen aus Bulgarien]. - Spis. bulg. Akad. Nauk 44: 1 - 50. - (1935): Ueber die von Dr. Stanko Karaman in Jugoslavien und besonders in Mazedonien gesammelten Spinnen (Araneae). - Izv. tsarsk. prirod. Inst. Sof. 8: 97 - 110. - (1936): Katalog der echten Spinnen (Araneae) der Balkanhalbinsel. - Sb. bulg. Akad. Nauk 32: 1 - 223. - (1938): Zur Morphologie und Biologie einer neuen bulgarischen Spinnenart, Euxinella Strandi n. g. n. sp. - Festschrift Strand 4: 569 - 574. Riga. - (1939): [Referate u. Mitteilungen]. Izv. b˘ulg. ent. Druzh. 10: 163 - 168. - (1940): [Die Spinnenfauna Bulgariens. IV. Unterordnung Arachnomorphae, II. Gruppe Trio- nichia, Familien: Zodariidae, Dictynidae und Amaurobiidae]. - Izv. tsarsk. prirod. Inst. Sof. 13: 169 - 194. - (1942). [Die Spinnenfauna Bulgariens. V. Unterordnung Arachnomorphae, II. Gruppe Trio- nichia, Familie: Agelenidae]. - Izv. tsarsk. prirod. Inst. Sof. 15: 33 - 60. - (1966): [Zoogeographical districts in the distribution of terrestrial fauna (in Bulgaria) ]. - In: GERASSIMOV, I. P. & Z. S. GULUBOV (eds.), Geography of Bulgaria 1: 500 - 506. Sofia. DRESCO, E. & M. HUBERT (1989): Araneae Speluncarum Italiae I. - Fragm. entom. 6: 167 - 181. DUMITRESCO, M. (1971): Une araignée nouvelle des grottes de Bulgarie, Antrohyphantes rodopicus n. g., n. sp. (fam. Linyphiidae, sous-fam. Linyphiinae, série de genres Lepthyphanteae). - Trav. Inst. Spéol. "E. Racovitza" 10: 167 - 174. HADJISSARANTOS, H. (1940): [Les araignées de l'Attique]. - Athens, 132 pp. JURINITCH, S. & P. DRENSKY (1917): [Contribution a 1'étude des araignées de Bulgarie]. - Spis. bulg. Akad. Nauk 15: 109 - 136. KNOFLACH, B. (1997): Zur Taxonomie, Verbreitung und Sexualbiologie von Theridion adrianopoli Drensky. - Ber. nat.-med. Verein Innsbruck 84: 133 - 148. - (1998): Mating in Theridion varians HAHN and related species (Araneae: Theridiidae). – J. nat.

Hist. 32: 545 - 604. - (1999): The comb-footed spider genera Neottiura and Coleosoma in Europe (Araneae, Theridiidae). - Mitt. schweiz. ent. Ges. 72: 341 - 371. KNOFLACH, B. & K. THALER (2000): Notes on Mediterranean Theridiidae (Araneae) - I. - Mem. Soc. entomol. ital. 78: 411-442. KRATOCHVIL, J. & F. MILLER (1938): Sur le probléme des araignées cavernicoles du genre Centro- merus de la Péninsule balkanique. - Izv. tsarsk. prirod. Inst. Sof. 11: 107 - 113. LEHTINEN, P. (1967): Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. - Annls zool. fenn. 4: 199 - 468. LUGETTI, G. & P. TONGIORGI (1969): Ricerche sul genere Alopecosa Simon (Araneae, Lycosidae). - Atti Soc. tosc. Sci. nat. 76: 1 - 100. MIKHAILOV, K. (1991): On the identity and distribution of Micaria romana (Aranei, Gnaphosidae). - Vestnik zoologii 1991: 77 - 79. MILLER, F. (1958): Eine neue Lepthyphantes (L. tranteevi sp. n.) aus den bulgarischen Höhlen. - Prace brn. Zakl. c˘sl. Akad. Ved. 30: 577 - 583. MILLER, F. & O. ZITNANSKA (1976): Ein Beitrag zur Kenntnis der Slowakischen Spinnenfauna. - Biologia (Bratislava) 31: 313 - 318. MILLER, F. & J. SVATON (1978): Einige seltene und bisher unbekannte Spinnenarten aus der Slowakei. - Ann. Zool. Bot. Bratislava 126: 1 - 19. MILLIDGE, A. F. (1981d): A revision of the genus Gonatium (Araneae: Linyphiidae). - Bull. Br. arachnol. Soc. 5: 253 - 277. PLATNICK, N. (2002): The world spider catalog, version 3.0. American Museum of Natural History, online at http://research.amnh.org/entomology/spiders/catalog81-87/index.html. WIEHLE, H. (1965): Die Clubiona-Arten Deutschlands, ihre natürliche Gruppierung und die Einheit- lichkeit im Bau ihrer Vulva (Arach., Araneae). - Senckenbergiana biol. 46: 471 - 505. WUNDERLICH, J. (1977): Zur Kenntnis der Lepthyphantes nebulosus-Gruppe (Arachnida: Araneida: Linyphiidae). - Senckenbergiana biol. 58: 57 - 61.

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