DOCSLIB.ORG

T-Ïfrsükt UNIVERSITEIT GENT

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
T-Ïfrsükt UNIVERSITEIT GENT t-ïfrsüKT UNIVERSITEIT GENT Fa c u l t y o f Sc ie n c e s * \o lo 9/0 ■».■'•Si Vakgroep Biologie Krijgslaan 281-S8. B-9000 Gent. BELGIUM E c o l o g y o f macrobenthos a s A BASELINE FOR AN ECOLOGICAL ADJUSTMENT OF BEACH NOURISHMENT E c o l o g ie v a n macrobenthos a l s e e n b a s is v o o r e e n ecologische b ij s t u r in g v a n strandsuppleties J e r o e n Sp e y b r o e c k Thesis submitted in partial fulfillment of the requirements for the degree of Doctor in Sciences: Biology Promotor: Prof. dr. Magda Vincx Co-promotor: dr. Steven Degraer Academic year 2006-2007 M e m b e r s o f t h e r e a d in g c o m m it t e e : Prof. dr. Magda Vincx, promotor (UGent, B) dr. Steven Degraer, co-promotor (UGent, B) dr. Gerard Janssen (RIKZ, NL) dr. ir. Jan van de Graaft (TU-Delft, NL) M e m b e r s o f t h e examination c o m m it t e e : Prof. dr. Wim Vyverman, chair (UGent, B) Prof. dr. Magda Vincx, promotor (UGent, B) dr. Steven Degraer, co-promotor (UGent, B) dr. Gerard Janssen (RIKZ, NL) dr. ir. Jan van de Graaff (TU-Delft, NL) Prof. dr. Jean-Pierre Maelfait (UGent and IN BO, B) dr. Jan Seys (VLIZ, B) dr. Maaike Steyaert (UGent, B) dr. Eric Stienen (INBO, B) Table of contents Dankw oord Summary - Samenvatting ........ 3 Summary .......... 3 Samenvatting .......... 7 Ou t l in e. .......... 13 Rationale ......... 13 Thesis contents and structure 16 Candidate's contribution to the different chapters 18 I. I ntroductory c h a p t e r s Chapter 1. The Belgian sandy beach ecosystem -a review. .... 21 Abstract.. 23 Introduction .......... 24 The physical environment ......... 24 The Belgian coast ........ 24 General features of beaches sampled within this thesis research (Chapters 3 and 4) 28 The biological environment. ........ 32 Vascular plants ......... 32 Terrestrial arthropods ........ 33 Microphytobenthos ........ 36 Zoobenthos ......... 37 Avifauna. ......... 39 Discussion .......... 43 Acknowledgements ......... 44 Chapter 2. Beach nourishment: an ecologically sound coastal defence alternative? A review. 45 Abstract.. 47 Introduction .......... 48 Ecological effects of beach nourishment ....... 50 Differential ecological effects of beach nourishment related aspects . 51 Activities during construction ...... 51 Quality characteristics of the nourishment sediment .... 52 Sediment composition ...... 52 Shells and other coarse material .... 52 Fines ....... 53 Grain size distribution ..... 54 Beach morphology ...... 55 Presence of toxic substances and/or dead organisms . 55 Sediment colour ....... 55 Quantity characteristics of the nourishment sediment .... 56 Factors determining scale of impact. ...... 57 Timing, place and size of beach nourishment. .... 57 Nourishment strategies and techniques ..... 57 Nourishment strategies ...... 57 Nourishment techniques ...... 59 Indirect and cumulative effects ....... 59 Conclusions .......... 60 Guidelines for an ecologically good practice of beach nourishment . 61 Sediment ........ 61 Timing ......... 61 Size ......... 61 Nourishment strategy ....... 62 Shortcomings of former research and options for future research . 62 Acknowledgements ......... 63 II. F ie ld r e s e a r c h - p a t t e r n s : l if e c y c l e , p o p u l a t io n d y n a m ic s a n d z o n a t io n Chapter 3. Understanding the life of a sandy beach polychaete of functional importance S- colelepis squam ata (Polychaeta: Spionidae) on Belgian sandy beaches (northeastern Atlantic, North Sea) 65 Abstract.. 67 Introduction .......... 68 Material & methods ......... 69 Sampling ......... 69 Lab procedures ......... 69 Data analysis ......... 70 Results ........... 71 Physical environment ........ 71 Life cycle, demography, reproduction and recruitment. 71 Abundance and biomass patterns ....... 74 Growth and mortality ........ 75 Mean annual biomass, secondary production and P/B ratio .... 75 Zonation. ......... 75 Discussion .......... 77 Life cycle, demography, reproduction and recruitment. 77 Abundance and biomass ........ 78 Growth and mortality ........ 79 Mean annual biomass, secondary production and P/B ratio .... 79 Zonation ......... 80 Towards global patterns of intraspecific variation in life history characteristics 81 Conclusions .......... 84 Acknowledgements ......... 84 Chapter 4. In s itu study of the autecology of two closely related, co-occurring sandy beach amphipods 85 Abstract.. 87 Introduction .......... 88 Methods .......... 89 Sampling ......... 89 Laboratory procedures ........ 90 Data analysis ......... 90 Results ........... 91 Physical environment ........ 91 Abundance and biomass ........ 92 Life cycle, demography, reproduction and recruitment. 93 Mean annual biomass, secondary production and P/B ratio .... 97 Cross-shore zonation ........ 98 Median grain size related to peak abundance ..... 100 Discussion .......... 100 Abundance and biomass . 100 Life cycle, demography, reproduction and recruitment. 102 Growth and mortality ........ 104 Secondary production 105 Habitat(1)-cross-shorezonation . 105 Habitat (2) - width of the occupied vertical range 106 Habitat (3) - median grain size at peak abundance 107 Acknowledgements .... 107 III. E xperimental r e s e a r c h - p r o c e s s e s : h a b it a t preferences and tol e r a n c e s Chapter 5. The ecological impact of beach nourishment and the restrained habitat of a key species - experimental insights and field data ....... 109 Abstract. .......... 111 Introduction 112 Materials and methods 113 Experimental data 113 Collection of experimentally used specimens. 113 Survival and feeding frequency related to median grain size 113 Burrowing time related to median grain size . 114 Sediment preference - grain size selection . 114 Field data 115 Results . 115 Experimental data 115 Survival and feeding frequency related to median grain size 115 Burrowing time related to median grain size . 116 Experimental sediment preference - grain size selection 117 Field data 118 Individual responses 118 Relatedness of environmental variables 119 Modelling density. 120 Discussion .......... 121 Conclusions-guidelines for beach nourishment ...... 123 Acknowledgements 123 Chapter 6. Competition and sediment-related responses explaining segregated zonation of two closely related, co-occurring key species on sandy beaches? 125 Abstract. .......... 127 Introduction .......... 128 Material & methods ......... 129 Sediment preference 130 Sediment tolerance 131 Results ........... 131 Sediment preference 131 Grain size ........ 131 Fines ......... 133 Sediment tolerance 135 Discussion .......... 136 Both species exhibit the same sediment preferences ..... 136 Occupied grain size range is not influenced by avoidance .... 137 Bathyporeia pilosa is more tolerant towards grain size extremes than B. sarsi . 138 Conclusions 138 Acknowledgements 139 IV. G e n e r a l d is c u s s io n Chapter 7. Towards a methodological framework for the prediction and mitigation of the ecological impact of beach nourishment - a case study on intertidal benthic macrofauna 141 Abstract. .......... 143 Introduction .......... 144 Data requirements, methodology and cost-benefit considerations .... 146 Physical data collection ........ 146 Biological data collection ........ 147 Prioritisation among data requirements-options for economising . 149 An example from intertidal macrobenthos 150 CITED LITERATURE . 155 A p p e n d ix - P u b lic a t io n list J er o en Sp e y b r o e c k (a s on 10A//2007) 185 Dankwoord Met groot risico op een melige epiloog, aan het einde van de rit nog even dit. Men vertelt mij dat dit onderdeel het enige is dat echt gelezen wordt. Daarom dacht ik misschien te moeten beginnen met een of ander smakelijk grapje of een opmerkelijke anekdote. Gezien mijn dissidente aard, doe ik dit echter niet en verwijs ik de lezer door naar de overvloedige prozaïsche praat die ongetwijfeld zal te horen zijn tijdens mijn doctoraatsreceptie. Bij het afronden van een doctoraat past grote dankbaarheid. In mijn geval geldt dit zeker: zonder de steun en praktische hulp van heel veel mensen, had ik het niet gehaald. Uiteraard dank ik zij die mij (grotendeels) betaalden, zijnde MD&K (the goverment agency formerly known as AWZ). Op dagelijkse, meer concrete basis, heb ik de steun en het vertrouwen van mijn promotor, Magda, en mentor, Steven, zeer sterk gewaardeerd. Ik hoop dan ook dat ze tevreden zijn met het eindresultaat van ongeveer vier jaar strandonderzoek. Ik hoop dat hierop verder kan gebouwd worden, zodat onze stranden nog lang geteisterd worden door enthousiaste biologen. In het bijzonder vond ik doorheen mijn jaren op de mariene de open sfeer (en open deur) van beide beschermengelen enorm aangenaam en constructief. Talloze uren ‘open debat' hebben mij heel veel geleerd en mijn inzicht en motivatie aangesterkt. Mijn onderzoek werd ondersteund door de bijdrage van heel veel mensen. Mijn drie lieftallige thesisstudentes - Lotte, Annelies en Joke - leverden prima werk, waarmee ik uitstekend geholpen was (zie hoofdstukken 3, 4 en 6). Een deel van mijn onderzoek vereiste intensieve staalnames, die werden uitgevoerd door een massa lieve helpende handen, waaronder o.a. zowat alle collega's van toen - An, Ann, Annelies DB, Annelies G, Annick VK, Annick V, Annick VH, Bart, Chrissie, Danielle, Danny, Eveline, Gert, Guy, Hannelore, Hendrik, Ilse, Isolde, Jan VAB, Jan W, Jurgen, Lotte, Maaike, Maarten, Magda, Marijn, Marnix, Matthew, Nancy, Pieter, Sabine, Sandra, Saskia, Sofie DR, de andere Sofie DR, Sofie VDD, Stefanie, Steven, Thomas de student, Thomas de andere strandjanet, Tim, Tom Gh en Véronique. In het bijzonder ook dank voor de praktische hulp van Danny, Bart, Annick VK, Danielle en Steven. Ondanks uitgebreide bolwassing en smalend hoongelach in ons aller koffiekot naar aanleiding van het megalomane dispersie-experiment, wil ik zeker zij die daarbij hielpen bedanken: studenten biologie '07 en Annelies, Annick, Bart, Bea, Cari, Cristiana, Danielle, Dirk, Frauke, Hugo, Karen, Karl, Saskia, Sharon, Sofie DR en Virginie. Bij het al-doende-leren, waaruit dit werk ontsproot, werd ik ook bijgestaan door o.m. Tom Gh, Marijn, Frederik H, Annick VK, Tim en mijn beider bureaugenoten, Jan Wittoeck en Cari Van Colen.
Load more

Recommended publications
  • Green-Tree Retention and Controlled Burning in Restoration and Conservation of Beetle Diversity in Boreal Forests
    Dissertationes Forestales 21 Green-tree retention and controlled burning in restoration and conservation of beetle diversity in boreal forests Esko Hyvärinen Faculty of Forestry University of Joensuu Academic dissertation To be presented, with the permission of the Faculty of Forestry of the University of Joensuu, for public criticism in auditorium C2 of the University of Joensuu, Yliopistonkatu 4, Joensuu, on 9th June 2006, at 12 o’clock noon. 2 Title: Green-tree retention and controlled burning in restoration and conservation of beetle diversity in boreal forests Author: Esko Hyvärinen Dissertationes Forestales 21 Supervisors: Prof. Jari Kouki, Faculty of Forestry, University of Joensuu, Finland Docent Petri Martikainen, Faculty of Forestry, University of Joensuu, Finland Pre-examiners: Docent Jyrki Muona, Finnish Museum of Natural History, Zoological Museum, University of Helsinki, Helsinki, Finland Docent Tomas Roslin, Department of Biological and Environmental Sciences, Division of Population Biology, University of Helsinki, Helsinki, Finland Opponent: Prof. Bengt Gunnar Jonsson, Department of Natural Sciences, Mid Sweden University, Sundsvall, Sweden ISSN 1795-7389 ISBN-13: 978-951-651-130-9 (PDF) ISBN-10: 951-651-130-9 (PDF) Paper copy printed: Joensuun yliopistopaino, 2006 Publishers: The Finnish Society of Forest Science Finnish Forest Research Institute Faculty of Agriculture and Forestry of the University of Helsinki Faculty of Forestry of the University of Joensuu Editorial Office: The Finnish Society of Forest Science Unioninkatu 40A, 00170 Helsinki, Finland http://www.metla.fi/dissertationes 3 Hyvärinen, Esko 2006. Green-tree retention and controlled burning in restoration and conservation of beetle diversity in boreal forests. University of Joensuu, Faculty of Forestry. ABSTRACT The main aim of this thesis was to demonstrate the effects of green-tree retention and controlled burning on beetles (Coleoptera) in order to provide information applicable to the restoration and conservation of beetle species diversity in boreal forests.
    [Show full text]
  • Diptera, Empidoidea) 263 Doi: 10.3897/Zookeys.365.6070 Research Article Launched to Accelerate Biodiversity Research
    A peer-reviewed open-access journal ZooKeys 365: 263–278 (2013) DNA barcoding of Hybotidae (Diptera, Empidoidea) 263 doi: 10.3897/zookeys.365.6070 RESEARCH ARTICLE www.zookeys.org Launched to accelerate biodiversity research Using DNA barcodes for assessing diversity in the family Hybotidae (Diptera, Empidoidea) Zoltán T. Nagy1, Gontran Sonet1, Jonas Mortelmans2, Camille Vandewynkel3, Patrick Grootaert2 1 Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny (JEMU), Rue Vautierstraat 29, 1000 Brussels, Belgium 2 Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny (Ento- mology), Rue Vautierstraat 29, 1000 Brussels, Belgium 3 Laboratoire des Sciences de l’eau et environnement, Faculté des Sciences et Techniques, Avenue Albert Thomas, 23, 87060 Limoges, France Corresponding author: Zoltán T. Nagy ([email protected]) Academic editor: K. Jordaens | Received 7 August 2013 | Accepted 27 November 2013 | Published 30 December 2013 Citation: Nagy ZT, Sonet G, Mortelmans J, Vandewynkel C, Grootaert P (2013) Using DNA barcodes for assessing diversity in the family Hybotidae (Diptera, Empidoidea). In: Nagy ZT, Backeljau T, De Meyer M, Jordaens K (Eds) DNA barcoding: a practical tool for fundamental and applied biodiversity research. ZooKeys 365: 263–278. doi: 10.3897/zookeys.365.6070 Abstract Empidoidea is one of the largest extant lineages of flies, but phylogenetic relationships among species of this group are poorly investigated and global diversity remains scarcely assessed. In this context, one of the most enigmatic empidoid families is Hybotidae. Within the framework of a pilot study, we barcoded 339 specimens of Old World hybotids belonging to 164 species and 22 genera (plus two Empis as outgroups) and attempted to evaluate whether patterns of intra- and interspecific divergences match the current tax- onomy.
    [Show full text]
  • Local and Regional Influences on Arthropod Community
    LOCAL AND REGIONAL INFLUENCES ON ARTHROPOD COMMUNITY STRUCTURE AND SPECIES COMPOSITION ON METROSIDEROS POLYMORPHA IN THE HAWAIIAN ISLANDS A DISSERTATION SUBMITTED TO THE GRADUATE DIVISION OF THE UNIVERSITY OF HAWAI'I IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF DOCTOR OF PHILOSOPHY IN ZOOLOGY (ECOLOGY, EVOLUTION AND CONSERVATION BIOLOGy) AUGUST 2004 By Daniel S. Gruner Dissertation Committee: Andrew D. Taylor, Chairperson John J. Ewel David Foote Leonard H. Freed Robert A. Kinzie Daniel Blaine © Copyright 2004 by Daniel Stephen Gruner All Rights Reserved. 111 DEDICATION This dissertation is dedicated to all the Hawaiian arthropods who gave their lives for the advancement ofscience and conservation. IV ACKNOWLEDGEMENTS Fellowship support was provided through the Science to Achieve Results program of the U.S. Environmental Protection Agency, and training grants from the John D. and Catherine T. MacArthur Foundation and the National Science Foundation (DGE-9355055 & DUE-9979656) to the Ecology, Evolution and Conservation Biology (EECB) Program of the University of Hawai'i at Manoa. I was also supported by research assistantships through the U.S. Department of Agriculture (A.D. Taylor) and the Water Resources Research Center (RA. Kay). I am grateful for scholarships from the Watson T. Yoshimoto Foundation and the ARCS Foundation, and research grants from the EECB Program, Sigma Xi, the Hawai'i Audubon Society, the David and Lucille Packard Foundation (through the Secretariat for Conservation Biology), and the NSF Doctoral Dissertation Improvement Grant program (DEB-0073055). The Environmental Leadership Program provided important training, funds, and community, and I am fortunate to be involved with this network.
    [Show full text]
  • Introduced Staphylinidae (Coleoptera) in the Maritime Provinces of Canada
    48 Introduced Staphylinidae (Coleoptera) in the Maritime Provinces of Canada Christopher G. Majka1 Nova Scotia Museum, 1747 Summer Street, Halifax, Nova Scotia, Canada B3H 3A6 Jan Klimaszewski Laurentian Forestry Centre, Canadian Forest Service, Natural Resources Canada, 1055 de P.E.P.S., P.O. Box 10380, Stn. Sainte-Foy, Québec, Quebec, Canada G1V 4C7 Abstract—The fauna of introduced rove beetles (Staphylinidae) in the Maritime Provinces of Canada is surveyed. Seventy-nine species have now been recorded. Of these, 73 have been found in Nova Scotia, 29 on Prince Edward Island, and 54 in New Brunswick. Twenty-five species are newly recorded in Nova Scotia, 16 on Prince Edward Island, and 10 in New Brunswick, for a total of 51 new provincial records. Of these, 15 species, Tachinus corticinus Gravenhorst, Mycetoporus lepidus (Gravenhorst), Habrocerus capillaricornis (Gravenhorst), Aleochara (Xenochara) lanuginosa Gravenhorst, Gnypeta caerulea (C.R. Sahlberg), Atheta (Microdota) amicula (Stephens), Cordalia obscura (Gravenhorst), Drusilla canaliculata (Fabricius), Deleaster dichrous (Gravenhorst), Coprophilus striatulus (Fabricius), Carpelimus subtilis (Erichson), Leptacinus intermedius Donisthorpe, Tasgius (Rayacheila) melanarius (Heer), Neobisnius villosulus (Stephens), and Philonthus discoideus (Gravenhorst), are newly recorded in the Maritime Prov- inces. Two of these, Atheta (Microdota) amicula and Carpelimus subtilis, are newly recorded in Canada. Leptacinus intermedius is removed from the faunal list of New Brunswick and Philhygra botanicarum Muona, a Holarctic species previously regarded as introduced in North America, is re- corded for the first time in the Maritime Provinces. An examination of when species were first de- tected in the region reveals that, on average, it was substantially later than comparable dates for other, better known families of Coleoptera — an apparent indication of the comparative lack of at- tention this family has received.
    [Show full text]
  • Page 1 Concessionaria Per La Progettazione, Realizzazione E
    Concessionaria per la progettazione, realizzazione e gestione del collegamento stabile tra la Sicilia e il Continente Organismo di Diritto Pubblico (Legge n° 1158 del 17 dicembre 1971, modificata dal D.Lgs. n°114 del 24 aprile 2003) P O N T E S U L L O S T R E T T O D I M E S S I N A PROGETTO DEFINITIVO ALTERNATIVE AI SITI DI DEPOSITO (Richieste CTVA del 22/12/2011 Prot. CTVA/2011/4534 e del 16/03/2012 Prot. CTVA/2012/1012) EUROLINK S.C.p.A. IMPREGILO S.p.A. SOCIETÀ ITALIANA PER CONDOTTE D’ACQUA S.p.A. COOPERATIVA MURATORI E CEMENTISTI - C.M.C. DI RAVENNA SOC. COOP. A.R.L. SACYR S.A.U. ISHIKAWAJIMA - HARIMA HEAVY INDUSTRIES CO. LTD A.C.I. S.C.P.A. - CONSORZIO STABILE IL CONTRAENTE GENERALE STRETTO DI MESSINA STRETTO DI MESSINA IL PROGETTISTA PROJECT MANAGER Direttore Generale Amministratore Delegato Dott. Ing. D. Spoglianti (Ing. P.P. Marcheselli) Ing. G. Fiammenghi Dott. P.Ciucci Ordine Ing. Milano n° A 20953 Dott. Ing. E. Pagani Ordine Ing. Milano n° 15408 Firmato digitalmente ai sensi dell’ ”Art.21 del D.Lgs. 82/2005” Unità Funzionale GENERALE AMV0258_F0 Tipo di sistema AMBIENTE Raggruppamento di opere/attività STUDIO DI IMPATTO AMBIENTALE Opera - tratto d’opera - parte d’opera QUADRO DI RIFERIMENTO AMBIENTALE - FAUNA Titolo del documento RELAZIONE GENERALE – AMBIENTE TERRESTRE C G 0 7 0 0 P R G V G A M I A Q 3 0 0 0 0 0 0 0 8 F0 CODICE REV DATA DESCRIZIONE REDATTO VERIFICATO APPROVATO F0 31/05/2012 Emissione finale P.MICHELI M.SALOMONE D.SPOGLIANTI NOME DEL FILE: AMV0258_F0.doc revisione interna:F0 Ponte sullo Stretto di Messina PROGETTO DEFINITIVO RELAZIONE GENERALE – AMBIENTE Codice documento Rev Data TERRESTRE AMV0258_F0.doc F0 31/05/2012 INDICE INDICE ..........................................................................................................................................
    [Show full text]
  • Effects of Urbanisation and Urban Areas on Biodiversity
    From genes to habitats – effects of urbanisation and urban areas on biodiversity Inauguraldissertation zur Erlangung der Würde eines Doktors der Philosophie vorgelegt der Philosophisch-Naturwissenschaflichen Fakultät der Universität Basel von Gwendoline (Wendy) Altherr aus Trogen, Appenzell-Ausserrhoden Basel, 2007 Genehmigt von der Philosophisch–Naturwissenschaftlichen Fakultät auf Antrag von Prof. Dr. Peter Nagel, Prof. Dr. Patricia Holm, Prof. (em.) Dr. Bernhard Klausnitzer Basel, den 18. September 2007 Prof. Dr. Hans-Peter Hauri Dekan TABLE OF CONTENTS Summary 1 General introduction – biodiversity in the city 3 Chapter I – genetic diversity 21 Population genetic structure of the wall lizard (Podarcis muralis) in an urban environment Manuscript Chapter II – species diversity 47 How do small urban forest patches contribute to the biodiversity 47 of the arthropod fauna? Manuscript Leistus fulvibarbis Dejean – Wiederfund einer verschollenen 79 Laufkäferart (Coleoptera, Carabidae) in der Schweiz Veröffentlicht in den Mitteilungen der Entomologischen Gesellschaft Basel 56(4), 2006 Chapter III – habitat diversity 89 How do stakeholders and the legislation influence the allocation of green space on brownfield redevelopment projects? Five case studies from Switzerland, Germany and the UK Published in Business Strategy and the Environment 16, 2007 General discussion and conclusions 109 Acknowledgements 117 Curriculum Vitae 119 SUMMARY Urban areas are landscapes dominated by built-up structures for human use. Nevertheless, nature can still be found within these areas. Urban ecosystems can offer ecological niches, sometimes only found in cities. This biodiversity in the form of genetic diversity, species diversity and habitat diversity provided the structure of this thesis. First, we studied the effects of urbanisation on genetic diversity. We analysed the population structure of the wall lizard with highly variable genetic markers.
    [Show full text]
  • The Invertebrate Fauna of Dune and Machair Sites In
    INSTITUTE OF TERRESTRIAL ECOLOGY (NATURAL ENVIRONMENT RESEARCH COUNCIL) REPORT TO THE NATURE CONSERVANCY COUNCIL ON THE INVERTEBRATE FAUNA OF DUNE AND MACHAIR SITES IN SCOTLAND Vol I Introduction, Methods and Analysis of Data (63 maps, 21 figures, 15 tables, 10 appendices) NCC/NE RC Contract No. F3/03/62 ITE Project No. 469 Monks Wood Experimental Station Abbots Ripton Huntingdon Cambs September 1979 This report is an official document prepared under contract between the Nature Conservancy Council and the Natural Environment Research Council. It should not be quoted without permission from both the Institute of Terrestrial Ecology and the Nature Conservancy Council. (i) Contents CAPTIONS FOR MAPS, TABLES, FIGURES AND ArPENDICES 1 INTRODUCTION 1 2 OBJECTIVES 2 3 METHODOLOGY 2 3.1 Invertebrate groups studied 3 3.2 Description of traps, siting and operating efficiency 4 3.3 Trapping period and number of collections 6 4 THE STATE OF KNOWL:DGE OF THE SCOTTISH SAND DUNE FAUNA AT THE BEGINNING OF THE SURVEY 7 5 SYNOPSIS OF WEATHER CONDITIONS DURING THE SAMPLING PERIODS 9 5.1 Outer Hebrides (1976) 9 5.2 North Coast (1976) 9 5.3 Moray Firth (1977) 10 5.4 East Coast (1976) 10 6. THE FAUNA AND ITS RANGE OF VARIATION 11 6.1 Introduction and methods of analysis 11 6.2 Ordinations of species/abundance data 11 G. Lepidoptera 12 6.4 Coleoptera:Carabidae 13 6.5 Coleoptera:Hydrophilidae to Scolytidae 14 6.6 Araneae 15 7 THE INDICATOR SPECIES ANALYSIS 17 7.1 Introduction 17 7.2 Lepidoptera 18 7.3 Coleoptera:Carabidae 19 7.4 Coleoptera:Hydrophilidae to Scolytidae
    [Show full text]
  • Zootaxa, Empidoidea (Diptera)
    ZOOTAXA 1180 The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera) BRADLEY J. SINCLAIR & JEFFREY M. CUMMING Magnolia Press Auckland, New Zealand BRADLEY J. SINCLAIR & JEFFREY M. CUMMING The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera) (Zootaxa 1180) 172 pp.; 30 cm. 21 Apr. 2006 ISBN 1-877407-79-8 (paperback) ISBN 1-877407-80-1 (Online edition) FIRST PUBLISHED IN 2006 BY Magnolia Press P.O. Box 41383 Auckland 1030 New Zealand e-mail: [email protected] http://www.mapress.com/zootaxa/ © 2006 Magnolia Press All rights reserved. No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any means, without prior written permission from the publisher, to whom all requests to reproduce copyright material should be directed in writing. This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose other than private research use. ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition) Zootaxa 1180: 1–172 (2006) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA 1180 Copyright © 2006 Magnolia Press ISSN 1175-5334 (online edition) The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera) BRADLEY J. SINCLAIR1 & JEFFREY M. CUMMING2 1 Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, 53113 Bonn, Germany. E-mail: [email protected] 2 Invertebrate Biodiversity, Agriculture and Agri-Food Canada, C.E.F., Ottawa, ON, Canada
    [Show full text]
  • ARTHROPODA Subphylum Hexapoda Protura, Springtails, Diplura, and Insects
    NINE Phylum ARTHROPODA SUBPHYLUM HEXAPODA Protura, springtails, Diplura, and insects ROD P. MACFARLANE, PETER A. MADDISON, IAN G. ANDREW, JOCELYN A. BERRY, PETER M. JOHNS, ROBERT J. B. HOARE, MARIE-CLAUDE LARIVIÈRE, PENELOPE GREENSLADE, ROSA C. HENDERSON, COURTenaY N. SMITHERS, RicarDO L. PALMA, JOHN B. WARD, ROBERT L. C. PILGRIM, DaVID R. TOWNS, IAN McLELLAN, DAVID A. J. TEULON, TERRY R. HITCHINGS, VICTOR F. EASTOP, NICHOLAS A. MARTIN, MURRAY J. FLETCHER, MARLON A. W. STUFKENS, PAMELA J. DALE, Daniel BURCKHARDT, THOMAS R. BUCKLEY, STEVEN A. TREWICK defining feature of the Hexapoda, as the name suggests, is six legs. Also, the body comprises a head, thorax, and abdomen. The number A of abdominal segments varies, however; there are only six in the Collembola (springtails), 9–12 in the Protura, and 10 in the Diplura, whereas in all other hexapods there are strictly 11. Insects are now regarded as comprising only those hexapods with 11 abdominal segments. Whereas crustaceans are the dominant group of arthropods in the sea, hexapods prevail on land, in numbers and biomass. Altogether, the Hexapoda constitutes the most diverse group of animals – the estimated number of described species worldwide is just over 900,000, with the beetles (order Coleoptera) comprising more than a third of these. Today, the Hexapoda is considered to contain four classes – the Insecta, and the Protura, Collembola, and Diplura. The latter three classes were formerly allied with the insect orders Archaeognatha (jumping bristletails) and Thysanura (silverfish) as the insect subclass Apterygota (‘wingless’). The Apterygota is now regarded as an artificial assemblage (Bitsch & Bitsch 2000).
    [Show full text]
  • Kenai National Wildlife Refuge Species List, Version 2018-07-24
    Kenai National Wildlife Refuge Species List, version 2018-07-24 Kenai National Wildlife Refuge biology staff July 24, 2018 2 Cover image: map of 16,213 georeferenced occurrence records included in the checklist. Contents Contents 3 Introduction 5 Purpose............................................................ 5 About the list......................................................... 5 Acknowledgments....................................................... 5 Native species 7 Vertebrates .......................................................... 7 Invertebrates ......................................................... 55 Vascular Plants........................................................ 91 Bryophytes ..........................................................164 Other Plants .........................................................171 Chromista...........................................................171 Fungi .............................................................173 Protozoans ..........................................................186 Non-native species 187 Vertebrates ..........................................................187 Invertebrates .........................................................187 Vascular Plants........................................................190 Extirpated species 207 Vertebrates ..........................................................207 Vascular Plants........................................................207 Change log 211 References 213 Index 215 3 Introduction Purpose to avoid implying
    [Show full text]
  • Thesis, University of Amsterdam, the Netherlands
    UvA-DARE (Digital Academic Repository) Ecological effects of plant invasions van Hengstum, T. Publication date 2013 Document Version Final published version Link to publication Citation for published version (APA): van Hengstum, T. (2013). Ecological effects of plant invasions. General rights It is not permitted to download or to forward/distribute the text or part of it without the consent of the author(s) and/or copyright holder(s), other than for strictly personal, individual use, unless the work is under an open content license (like Creative Commons). Disclaimer/Complaints regulations If you believe that digital publication of certain material infringes any of your rights or (privacy) interests, please let the Library know, stating your reasons. In case of a legitimate complaint, the Library will make the material inaccessible and/or remove it from the website. Please Ask the Library: https://uba.uva.nl/en/contact, or a letter to: Library of the University of Amsterdam, Secretariat, Singel 425, 1012 WP Amsterdam, The Netherlands. You will be contacted as soon as possible. UvA-DARE is a service provided by the library of the University of Amsterdam (https://dare.uva.nl) Download date:01 Oct 2021 THOMAS VAN HENGSTUM van Hengstum, T. 2013. Ecological effects of plant invasions. PhD thesis, University of Amsterdam, The Netherlands. The research presented in this thesis was funded by the Dutch Organization for Scientific Research (NWO) as part of the ERGO program (838.06.111). ISBN: 978-90-821099-0-0 ECOLOGICAL EFFECTS OF PLANT INVASIONS Academisch proefschrift ter verkrijging van de graad van doctor aan de Universiteit van Amsterdam op gezag van de Rector Magnificus prof.
    [Show full text]
  • 2011 Biodiversity Snapshot. Isle of Man Appendices
    UK Overseas Territories and Crown Dependencies: 2011 Biodiversity snapshot. Isle of Man: Appendices. Author: Elizabeth Charter Principal Biodiversity Officer (Strategy and Advocacy). Department of Environment, Food and Agriculture, Isle of man. More information available at: www.gov.im/defa/ This section includes a series of appendices that provide additional information relating to that provided in the Isle of Man chapter of the publication: UK Overseas Territories and Crown Dependencies: 2011 Biodiversity snapshot. All information relating to the Isle or Man is available at http://jncc.defra.gov.uk/page-5819 The entire publication is available for download at http://jncc.defra.gov.uk/page-5821 1 Table of Contents Appendix 1: Multilateral Environmental Agreements ..................................................................... 3 Appendix 2 National Wildife Legislation ......................................................................................... 5 Appendix 3: Protected Areas .......................................................................................................... 6 Appendix 4: Institutional Arrangements ........................................................................................ 10 Appendix 5: Research priorities .................................................................................................... 13 Appendix 6 Ecosystem/habitats ................................................................................................... 14 Appendix 7: Species ....................................................................................................................
    [Show full text]