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Systematics of the Antillean Blind Snakes of the Genus Typhlops (Serpentes: Typhlopidae)

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Systematics of the Antillean Blind Snakes of the Genus Typhlops (Serpentes: Typhlopidae) Louisiana State University LSU Digital Commons LSU Historical Dissertations and Theses Graduate School 1976 Systematics of the Antillean Blind Snakes of the Genus Typhlops (Serpentes: Typhlopidae). John Paul richard Thomas Louisiana State University and Agricultural & Mechanical College Follow this and additional works at: https://digitalcommons.lsu.edu/gradschool_disstheses Recommended Citation Thomas, John Paul richard, "Systematics of the Antillean Blind Snakes of the Genus Typhlops (Serpentes: Typhlopidae)." (1976). LSU Historical Dissertations and Theses. 2993. https://digitalcommons.lsu.edu/gradschool_disstheses/2993 This Dissertation is brought to you for free and open access by the Graduate School at LSU Digital Commons. It has been accepted for inclusion in LSU Historical Dissertations and Theses by an authorized administrator of LSU Digital Commons. For more information, please contact [email protected]. INFORMATION TO USERS This material was produced from a microfilm copy of the original document. While the most advanced technological means to photograph and reproduce this document have been used, the quality is heavily dependent upon the quality of the original submitted. The following explanation of techniques is provided to help you understand markings or patterns which may appear on this reproduction. 1. The sign or "target" for pages apparently lacking from the document photographed is "Missing Page(s)". If it was possible to obtain the missing page(s) or section, they are spliced into the film along with adjacent pages. This may have necessitated cutting thru an image and duplicating adjacent pages to insure you complete continuity. 2. When an im ^e on the film is obliterated with a large round black mark, it is an indication that die photographer suspected that the copy may have moved during exposure and thus cause a blurred image. You will find a good image of the page in the adjacent frame. 3. When a map, drawing or chart, etc., was part of the material being photographed the photographer followed a definite method in "sectioning" the material. It is customary to begin photoing at the upper left hand corner of a large sheet and to continue photoing from left to right in equal sections with a small overlap. If necessary, sectioning is continued again — beginning below the first row and continuing on until complete. 4. The majority of users indicate that the textual content is of greatest value, however, a somewhat higher quality reproduction could be made from "photographs" if essential to the understanding of the dissertation. Silver prints of "photographs" may be ordered at additional charge by writing the Order Department, giving the catalog number, title, author and specific pages you wish reproduced. 5. PLEASE NOTE: Some pages may have indistinct print. Filmed as received. Xerox University MIScrofaims Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. 7 6 - 2 8 ,8 3 4 THOMAS, J o h n P a u l R i c h a r d , 1 9 3 8 - SYSTEMATICS OF THE ANTILLEAN BLIND SNAKES OF THE GENUS TYPHLOPS (SERPENTES: TYPKLOPIDAE). The Louisiana State University and A gricultural and Mechanical College, Ph.D., 1976 Z o o lo g y Xerox University Microfilms, Ann Arbor, Michigan 48106 Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. SYSTîMTICS OF THE ANTILLE&N BUND SNAKES OF THE o m is TYPHLOPS (S5CPENTES: TYPHLOPIDAE) A Dissertation Submitted to the Graduate Faculty of the Louisiana State University and Agricultural and Mechanical College in partial fulfillment of the requirements for the degree of Doctor of Philosophy The Department of Zoology and Physiology by- John Paul R ichard Thomas B. A., University of South Florida, 1969 August, 1976 Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. ACKNOlflEDGMMTS AND ABERE7IATI0NS Without the support, encouragement, and prodigious field work of Albert Schwartz, this study would never have had a reason for being, and it is to him that I owe a prime debt of gratitude. Douglas A. Rossman, as both major professor and friend, has provided help, guidance, encouragement, perceptive criticism , patience, and paronomasia —often beyond the requirements of duty. George H. Lowery, Jr., has in many ways encouraged and aided me during th e course o f t h is study. Ernest E. Williams has endured and responded to numerous loan requests —and re-requests—and has helpfully answered various inquiries about specimens and obscure l i t e r a t u r e . W alter J . Harman has a t various times facilitated the aquisition of funds and equipment in support of this study. H. B. Boudreaux, George C. Kent, and J. P. Woodring, all members of my comm ittee, and my w ife, Kim, have c r i t i c a l l y read th e manuscript. A number of people—curatorial staff and private persons—have loaned or been instrumental in the lending of specimens: C. M. Bogert, R. G. Zweifel, G. M. Foley (AMNH; see abbreviations beyond); E. V. Malnate (ANSP); A. G. C. G randison, A. F, Stimson (BMH); C. J . McCoy, N. D. Richmond (CM); D. R. Paulson (DRP); F. G. Thompson (PGT, UF/FSM); 0 . G arrido (iZ ); W. E. Duellman (KU); L. D. Ober (LDO); M. L. Jaume (MVZH); E. E. W illiams (MCZ); J . Guibe (MI'IHN); S. J . Inchaustegui (MNHNSD); R. Pinchon (SC); R. M ertens, K. Klemmer (SMF); J . R. Dixon (TCWC); H. A. Dundee (TU); W. Auffenberg (UF/FSM); C. W alker, A. Kluge (UMMZ); J . A. Rivero (UPRM); D. R. Cochran, J . A. P e te rs , G. R. Zug, R. I . Crombie (USNM); C. A. Reed (YPB). Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. Several people have kindly donated specimens of Typhlops collected by them: D. W- Buden, R. McCranie, L. Porras, J. Rindfleish, and C. R. Warren. A. C. l,#ieeler of the British Museum most helpfully provided me with information on Gronovius, Tfdiose collection was the source of Linnaeus’ description of Typhlops lumbricalis. G. Underwood kindly provided information on the holotype of Meditoria nasuta. Large numbers of specimens upon which this study was based were collected during field work sponsored by the National Science Foundation. Grants G —3865 and G-6252 to Albert Schwartz supported field work in Cuba; GB-6944 to Emest E. Williams and G-7977 and B-O23603 to Albert Schwartz supported field work in Hispaniola. The following is a list of abbreviations used in this study for collections from which specimens were examined or cited. AMNH - American Museum of Natural History ANSP — Academy o f N atu ral Sciences o f P h ila d e lp h ia ASFS - A lbert Schwartz F ie ld S e rie s , Miami BMNH - B ritish Museum (Natural History) CM - C arnegie Museum DRP - Dennis R. Paulson private collection (now in Los Angeles County Museum) FGT - Fred G. Thompson f ie l d s e r ie s , F lo rid a S ta te Museum IZ - Institute de Zoologia, La Habana, Cuba JRD - James R. Dixon field series, Texas A & M University KU - University of Kansas Nîuseum of Natural History LDO - Levn_s D. Ober p riv a te c o lle c tio n , Miami Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. LSUMZ - Louisiana State University Museum of Zoology MBZH - Museo y Biblioteca de Zoologia de la Habana (probably no longer extant) MCZ - Museum of Comparative Zoology, Harvard University MNHN - Museum National d'Histoire Naturelle, Paris MNHNSD - Museo N acional de H is to ria N a tu ra l, Santo Domingo, Republica Dominicana RNH - Rijksmuseum van Natuurlijke H istorié, Leiden RT - Richard Thomas private collection. Baton Rouge, Louisiana SC - Séminaire College, Fort de France, Martinique SMF - Natur-Museum und Forschungs-Institut Senckenberg, Frankfurt am Main TCWC - Texas Cooperative W ildlife Collection, Texas A & M University TU - Tulane University UF/FSM - F lo rid a S ta te Museum, U n iv e rsity o f F lo rid a UT#Z - University of Michigan Museum of Zoology UPRM - University of Puerto Rico, Mayagüez USNM - National Museum of Natural History YPB - Yale-Peabody Museum, Y ale U n iv e rsity Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. TABLE OF CONTMTS ACKNOWLEDGMENTS AND ABBREVIATIONS..............................................................................i i LIST OF TABLES......................................................................................................................v i i LIST OF FIGURES................................................................. ix ABSTRACT..................................................................................................................................... INTRODUCTION................................................................................................................................ METHODS AND CHARACTERS.......................................................................................................... SYSTEMATIC ACCOUNTS............................................................................................................. 19 Typhlops .jamaicensis ............................................................................................19 Typhlops sulcata .................................................................................................... 22 Typhlops 1 .................................................................................................................. 25 Typhlops lumbricalis. ........................................................................................
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