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SYSTEMATICA OF ARNICA, SUBGENUS AUSTROMONTANA AND A NEW SUBGENUS, CALARNICA (ASTERACEAE:SENECIONEAE) by GERALD BANE STRALEY B.Sc, Virginia Polytechnic Institute, 1968 M.Sc, Ohio University, 1974 A THESIS SUBMITTED IN PARTIAL FULFILMENT OF THE REQUIREMENTS OF THE DEGREE OF DOCTOR OF PHILOSOPHY in THE FACULTY OF GRADUATE STUDIES (Department of Botany) We accept this thesis as conforming to the required standard THE UNIVERSITY OF BRITISH COLUMBIA March 1980 © Gerald Bane Straley, 1980 In presenting this thesis in partial fulfilment of the requirements for an advanced degree at the University of British Columbia, I agree that the Library shall make it freely available for reference and study. I further agree that permission for extensive copying of this thesis for scholarly purposes may be granted by the Head of my Department or by his representatives. It is understood that copying or publication of this thesis for financial gain shall not be allowed without my written permission. Department nf Botany The University of British Columbia 2075 Wesbrook Place Vancouver, Canada V6T 1W5 26 March 1980 ABSTRACT Seven species are recognized in Arnica subgenus Austromontana and two species in a new subgenus Calarnica based on a critical review and conserva• tive revision of the species. Chromosome numbers are given for 91 populations representing all species, including the first reports for Arnica nevadensis. Results of apomixis, vegetative reproduction, breeding studies, and artifi• cial hybridizations are given. Interrelationships of insect pollinators, leaf miners, achene feeders, and floret feeders are presented. Arnica cordifolia, the ancestral species consists largely of tetraploid populations, which are either autonomous or pseudogamous apomicts, and to a lesser degree diploid, triploid, pentaploid, and hexaploid populations. -
Chloroplast DNA Evidence for a North American Origin of the Hawaiian
Proc, Natl. Acad. Sci. USA Vol. 88, pp. 1840-1843, March 1991 Evolution Chloroplast DNA evidence for a North American origin of the Hawaiian silversword alliance (Asteraceae) (insular evolution/adaptive radiation/biogeography/long-distance dispersal/phylogenetics) BRUCE G. BALDWIN*t, DONALD W. KYHOS*, JAN DVORAKO, AND GERALD D. CARR§ Departments of *Botany and tAgronomy and Range Science, University of California, Davis, CA 95616; and §Department of Botany, University of Hawaii, Honolulu, HI 96822 Communicated by Peter H. Raven, November 30, 1990 (received for review August 1, 1990) ABSTRACT Chloroplast DNA restriction-site compari- sity, structural (4, 5), biosystematic (6), allozymic (7), and sons were made among 24 species of the Hawaiian silversword chloroplast DNA (cpDNA) (8) data indicate the silversword alliance (Argyroxiphium, Dubautia, and Wilkesia) and 7 species alliance originated from a single colonizing species. of North American perennial tarweeds in Adenothamnus, Ma- Carlquist (1) presented convincing anatomical evidence dia, Raillardella, and Railkrdiopsis (Asteraceae-Madiinae). indicating taxonomic alignment of the Hawaiian silversword These data and results from intergeneric hybridization indi- alliance with the almost exclusively herbaceous American cated surprisingly close genetic affinity of the monophyletic Madiinae or tarweeds. Gray (9) earlier suggested such affinity Hawaiian group to two diploid species of montane perennial for Argyroxiphium, which was disputed by Keck (10) based herbs in California, Madia bolanderi and Raillardiopsis muiri. on presumed morphological dissimilarities and the magnitude Of 117 restriction-site mutations shared among a subset of two of the oceanic barrier to migration. Herein, we compare or more accessions, more than one-fifth (25 mutations) sepa- cpDNA restriction sites between the silversword alliance and rated the silversword alliance, M. -
List of Plants for Great Sand Dunes National Park and Preserve
Great Sand Dunes National Park and Preserve Plant Checklist DRAFT as of 29 November 2005 FERNS AND FERN ALLIES Equisetaceae (Horsetail Family) Vascular Plant Equisetales Equisetaceae Equisetum arvense Present in Park Rare Native Field horsetail Vascular Plant Equisetales Equisetaceae Equisetum laevigatum Present in Park Unknown Native Scouring-rush Polypodiaceae (Fern Family) Vascular Plant Polypodiales Dryopteridaceae Cystopteris fragilis Present in Park Uncommon Native Brittle bladderfern Vascular Plant Polypodiales Dryopteridaceae Woodsia oregana Present in Park Uncommon Native Oregon woodsia Pteridaceae (Maidenhair Fern Family) Vascular Plant Polypodiales Pteridaceae Argyrochosma fendleri Present in Park Unknown Native Zigzag fern Vascular Plant Polypodiales Pteridaceae Cheilanthes feei Present in Park Uncommon Native Slender lip fern Vascular Plant Polypodiales Pteridaceae Cryptogramma acrostichoides Present in Park Unknown Native American rockbrake Selaginellaceae (Spikemoss Family) Vascular Plant Selaginellales Selaginellaceae Selaginella densa Present in Park Rare Native Lesser spikemoss Vascular Plant Selaginellales Selaginellaceae Selaginella weatherbiana Present in Park Unknown Native Weatherby's clubmoss CONIFERS Cupressaceae (Cypress family) Vascular Plant Pinales Cupressaceae Juniperus scopulorum Present in Park Unknown Native Rocky Mountain juniper Pinaceae (Pine Family) Vascular Plant Pinales Pinaceae Abies concolor var. concolor Present in Park Rare Native White fir Vascular Plant Pinales Pinaceae Abies lasiocarpa Present -
Genetic Diversity and Evolution in Lactuca L. (Asteraceae)
Genetic diversity and evolution in Lactuca L. (Asteraceae) from phylogeny to molecular breeding Zhen Wei Thesis committee Promotor Prof. Dr M.E. Schranz Professor of Biosystematics Wageningen University Other members Prof. Dr P.C. Struik, Wageningen University Dr N. Kilian, Free University of Berlin, Germany Dr R. van Treuren, Wageningen University Dr M.J.W. Jeuken, Wageningen University This research was conducted under the auspices of the Graduate School of Experimental Plant Sciences. Genetic diversity and evolution in Lactuca L. (Asteraceae) from phylogeny to molecular breeding Zhen Wei Thesis submitted in fulfilment of the requirements for the degree of doctor at Wageningen University by the authority of the Rector Magnificus Prof. Dr A.P.J. Mol, in the presence of the Thesis Committee appointed by the Academic Board to be defended in public on Monday 25 January 2016 at 1.30 p.m. in the Aula. Zhen Wei Genetic diversity and evolution in Lactuca L. (Asteraceae) - from phylogeny to molecular breeding, 210 pages. PhD thesis, Wageningen University, Wageningen, NL (2016) With references, with summary in Dutch and English ISBN 978-94-6257-614-8 Contents Chapter 1 General introduction 7 Chapter 2 Phylogenetic relationships within Lactuca L. (Asteraceae), including African species, based on chloroplast DNA sequence comparisons* 31 Chapter 3 Phylogenetic analysis of Lactuca L. and closely related genera (Asteraceae), using complete chloroplast genomes and nuclear rDNA sequences 99 Chapter 4 A mixed model QTL analysis for salt tolerance in -
Phylogenies and Secondary Chemistry in Arnica (Asteraceae)
Digital Comprehensive Summaries of Uppsala Dissertations from the Faculty of Science and Technology 392 Phylogenies and Secondary Chemistry in Arnica (Asteraceae) CATARINA EKENÄS ACTA UNIVERSITATIS UPSALIENSIS ISSN 1651-6214 UPPSALA ISBN 978-91-554-7092-0 2008 urn:nbn:se:uu:diva-8459 ! " # $ %&& &'&& ( ( ( ) * + , * - * %&& * ) . /!0* ! * 12%* 34 * * 5. 24 62633$64&2%6&* + /!0 , ( 7 /.+0 , ( , ! 7 * ( ( / ! " 0 / 0 ,6 ! ( ( 8! 55 /#$%&0 , 4 %1* ) , ,6 ' - * 9 : ( , ( (( ! * ( ( ( * .7 ( , ! ( ( 11 ( ; * .+ ! 7 ( ( /** ( 0 , ( * . ( ( ( ( ( , ( * " ( .+ ( # ! * ! 6 ( ( ( ( ( ( "6< ! , ( ( % * ( ( , (( * = , .+ * )* + ! ! 5+. +. 8)% 7 )! "6< ,-.' , ' / ' 0 ( 1 ' ' ,234&5 ' * > - %&& 5.. ;36;%$ 5. 24 62633$64&2%6& ' ''' 6 $32 / '99 *-*9 ? @ ' ''' 6 $320 List of Papers This thesis is based on the following papers, which are referred to in the text by their Roman numerals: I Ekenäs, C., B. G. Baldwin, and K. Andreasen. 2007. A molecular phylogenetic study of Arnica (Asteraceae): Low chloroplast DNA variation and problematic subgeneric classification. Sys- tematic Botany -
Annotated Checklist of Vascular Flora, Cedar Breaks National
National Park Service U.S. Department of the Interior Natural Resource Program Center Annotated Checklist of Vascular Flora Cedar Breaks National Monument Natural Resource Technical Report NPS/NCPN/NRTR—2009/173 ON THE COVER Peterson’s campion (Silene petersonii), Cedar Breaks National Monument, Utah. Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Cedar Breaks National Monument Natural Resource Technical Report NPS/NCPN/NRTR—2009/173 Author Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741 Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532 February 2009 U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The Natural Resource Publication series addresses natural resource topics that are of interest and applicability to a broad readership in the National Park Service and to others in the management of natural resources, including the scientifi c community, the public, and the NPS conservation and environmental constituencies. Manuscripts are peer-reviewed to ensure that the information is scientifi cally credible, technically accurate, appropriately written for the intended audience, and is designed and published in a professional manner. The Natural Resource Technical Report series is used to disseminate the peer-reviewed results of scientifi c studies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service’s mission. The reports provide contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. Current examples of such reports include the results of research that addresses natural resource management issues; natural resource inventory and monitoring activities; resource assessment reports; scientifi c literature reviews; and peer- reviewed proceedings of technical workshops, conferences, or symposia. -
Rocky Mountain Draba): a Technical Conservation Assessment
Draba graminea Greene (Rocky Mountain draba): A Technical Conservation Assessment Prepared for the USDA Forest Service, Rocky Mountain Region, Species Conservation Project September 30, 2004 Lynn Moore1 and Sandy Friedley2 1Windom Floristics 361 South Camino Del Rio #222 Durango, CO 81303 2Ecosphere Environmental Services 2257 Main Avenue Durango, CO 81301 Peer Review Administered by Society for Conservation Biology Moore, L. and S. Friedley. (2004, September 30). Draba graminea Green (Rocky Mountain draba): a technical conservation assessment. [Online]. USDA Forest Service, Rocky Mountain Region. Available: http:// www.fs.fed.us/r2/projects/scp/assessments/drabagraminea.pdf [date of access]. ACKNOWLEDGEMENTS The authors would like to thank Mike Menefee of the Colorado Natural Heritage Program for providing assistance and occurrence data. We are grateful to the Rocky Mountain Herbarium (RM), Kathryn Kalmbach Herbarium (KHD), San Juan College Herbarium (SJNM), University of Northern Colorado Herbarium (GREE), Fort Lewis College Herbarium (FLD), and University of Colorado Museum (COLO) for providing herbarium label data. Thanks go to Peggy Lyon for providing photographs and information concerning recent surveys in the San Juan Mountains. We would also like to express our appreciation to Ken Heil for his comments and review of the manuscript. Thanks to Barry Johnston and Jeff Redders for providing information concerning the species. AUTHORS’ BIOGRAPHIES Lynn Moore Botanist/Owner Windom Floristics. Lynn graduated from Fort Lewis College with a Bachelor of Science in Biology in 1992 and continued her graduate work at the University of Wyoming, where she finished a Masters of Science degree in Botany in 1998. Lynn has worked in the private sector independently contracting services in the Four Corners area since 1997. -
A Second Annotated Checklist of Vascular Plants in Wells Gray Provincial Park and Vicinity, British Columbia, Canada
A second annotated checklist of vascular plants in Wells Gray Provincial Park and vicinity, British Columbia, Canada Version 1: April, 2011 Curtis R. Björk1 and Trevor Goward2 ENLICHENED CONSULTING LTD. Box 131, Clearwater, BC, V0E 1N0, Canada [email protected], [email protected] Vascular Plants in Wells Gray SUMMARY Wells Gray Provincial Park is a vast wilderness preserve situated in the mountains and highlands of south-central British Columbia. The first major floristic study of the vascular plants of Wells Gray and its vicinity was published in 1965 by Leena Hämet-Ahti, who documented 550 taxa, including a first Canadian record of Carex praeceptorium. The present study contributes nearly 500 additional taxa documented by us between 1976 and 2010 in connection with our personal explorations of the Clearwater Valley. The vascular flora of Wells Gray Park and vicinity now stands at 1046 taxa, including 881 native species and 165 species introduced from Eurasia and other portions of British Columbia. Wells Gray Park is notable both for the presence of numerous taxa (45) at or near the northern limits of their range, as well as for an unexpectedly high number of taxa (43) accorded conservation status by the British Columbia Conservation Data Centre. Antennaria corymbosa has its only known Canadian locality within Wells Gray, while five additional species reported here are known in Canada from fewer than six localities. About a dozen unknown, possibly undescribed taxa have also been detected. Botanical inventory has thus far been confined to the southern portions of Wells Gray. Future studies in northern half of the park will certainly greatly increase our knowledge of the biological diversity safeguarded in this magnificent wilderness preserve. -
White River National Forest Rare Plant Survey 2006
White River National Forest Rare Plant Survey 2006 Prepared for White River National Forest By Peggy Lyon, Janis Huggins and Georgia Doyle Colorado Natural Heritage Program 254 General Services Building Colorado State University Fort Collins CO 80523 Acknowledgments This project would not have been possible without the help of many people in the White River National Forest: Kevin Atchley for first requesting this survey and working to get a contract in place; Beth Brenneman and Keith Giezentanner who took over for Kevin when he left the area; Joe Doerr and Philip Nyland, for sharing their experience and caring about the rare plants on the forest. Also, we depend on the support of the staff of CNHP in Fort Collins, especially Dave Anderson, Jill Handwerk, Susan Spackman, and Amy Lavender. Thanks also to Tim Hogan at the CU Herbarium for assisting with identification of problem plants; Dee Malone for her expertise and transportation to Taylor Pass; and John Chapman, FS Volunteer, for his time and transportation to Warren Lakes. Colorado Natural Areas Program, under the energetic direction of Brian Kurzel, and members of the Colorado Native Plant Society assisted with a survey of the Hoosier Ridge State Natural Area. Finally, thanks to our spouses, George and Rick, for putting up with our absences over the summer, and our faithful companions Molly and Misia, who protected us from Ptarmigan and other scary things. 2 Executive Summary The White River National Forest contracted with Colorado State University in 2005 to have the Colorado Natural Heritage Program (CNHP) conduct a survey of sensitive plants on the forest. -
Great Sand Dunes National Monument and Preserve Vascular Plant Inventory
Great Sand Dunes National Monument and Preserve Vascular Plant Inventory Susan Spackman Panjabi and Karin Decker Colorado Natural Heritage Program, Colorado State University Ft. Collins, Colorado 80523 December 30, 2002 I. Background As part of its biological inventory program, the National Park Service (NPS) contracted the Colorado Natural Heritage Program (CNHP) in 2001-2002 to conduct a field inventory of vascular plants of Great Sand Dunes National Monument and Preserve (GRSA). In 2001 the Rocky Mountain Network of the NPS developed a study plan for biological inventories for parks in the Network. Although detailed botanical research had been conducted in GRSA by H.N. Dixon (1999) and others, additional research was needed, particularly regarding the distribution of high elevation, rare, and non-native plants. The Network in cooperation with CNHP submitted a Biological Inventory Study Plan to the NPS Inventory and Monitoring Program, which was approved. That Plan provides detailed information and guidance for biological inventories in all Rocky Mountain Network parks. The checklist of vascular plants of Great Sand Dunes National Monument (Dixon 1999) was developed over a 30-year period from 1969-1999, and documents about 406 taxa that occur in the Monument. This is one of the most thorough botanical inventories in all of Colorado. Given the thoroughness of this inventory, the checklist is considered to be comprehensive for the Monument. Although, many of the species on the checklist are not represented in the GRSA Herbarium, many voucher specimens are housed at the Alamosa State College Herbarium (pers. comm. Dixon 2002). On November 22, 2000, the boundaries of the Great Sand Dunes National Monument were expanded to include the Great Sand Dunes Preserve. -
Checklist of Montana Vascular Plants
Checklist of Montana Vascular Plants June 1, 2011 By Scott Mincemoyer Montana Natural Heritage Program Helena, MT This checklist of Montana vascular plants is organized by Division, Class and Family. Species are listed alphabetically within this hierarchy. Synonyms, if any, are listed below each species and are slightly indented from the main species list. The list is generally composed of species which have been documented in the state and are vouchered by a specimen collection deposited at a recognized herbaria. Additionally, some species are included on the list based on their presence in the state being reported in published and unpublished botanical literature or through data submitted to MTNHP. The checklist is made possible by the contributions of numerous botanists, natural resource professionals and plant enthusiasts throughout Montana’s history. Recent work by Peter Lesica on a revised Flora of Montana (Lesica 2011) has been invaluable for compiling this checklist as has Lavin and Seibert’s “Grasses of Montana” (2011). Additionally, published volumes of the Flora of North America (FNA 1993+) have also proved very beneficial during this process. The taxonomy and nomenclature used in this checklist relies heavily on these previously mentioned resources, but does not strictly follow anyone of them. The Checklist of Montana Vascular Plants can be viewed or downloaded from the Montana Natural Heritage Program’s website at: http://mtnhp.org/plants/default.asp This publication will be updated periodically with more frequent revisions anticipated initially due to the need for further review of the taxonomy and nomenclature of particular taxonomic groups (e.g. Arabis s.l ., Crataegus , Physaria ) and the need to clarify the presence or absence in the state of some species. -
Senator Beck Basin Long-Term Vegetation Study 2009 Re-Survey Final Report Presented to the Center for Snow and Avalanche Studies, Silverton, Colorado
Senator Beck Basin Long-term Vegetation Study 2009 Re-Survey Final Report Presented to the Center for Snow and Avalanche Studies, Silverton, Colorado January 2010 by Julie A. Crawford, PhD Contract Ecologist 336 North Main Street Mancos, CO 81328 970-533-7646 [email protected] Abstract Climate greatly influences vegetation composition, abundance, and distribution. Predicted climate change in the Rocky Mountains of the western United States may lead to the reduction of snow pack and stream flow, and increase land surface drying. These events, as well as the movement of plant species upslope in response to warming, and the reduction of endemic species at the exterior of their ranges have already been documented. In southwest Colorado, there have been overall increases in mean temperature and decreases in total precipitation over the last century. In addition to the impact of these long-term trends, vegetation also has an immediate response to short-term fluctuations in water availability. Repeated measures of permanently placed vegetation plots can show the impacts of both long- and short-term climate patterns on vegetation. In 2004, a long-term vegetation study was initiated in Senator Beck Basin in the San Juan Mountains of southwestern Colorado. These permanent vegetation transects were read again in 2009 and a comparison of the two readings is presented here. Results show a significant difference in the vegetation of the basin in the five-year period since initial transect set-up. An additional 39 species were recorded from the transects in 2009, while 26 species found in 2004 were not recorded in 2009.