Annotated Checklist of Vascular Flora, Black Canyon of The

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Annotated Checklist of Vascular Flora Black Canyon of the Gunnison National Park

Natural Resource Technical Report NPS/NCPN/NRTR—2009/227 ON THE COVER Fendler’s mountain lilac (Ceanothus fendleri). Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Black Canyon of the Gunnison National Park

Natural Resource Technical Report NPS/NCPN/NRTR—2009/227

Authors Tim Hogan Nan Lederer Dina Clark University of Colorado Museum Herbarium 218 UCB Boulder, CO 80309

Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741

Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network National Park Service P.O. Box 848 Moab, UT 84532

July 2009

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Please cite this publication as:

Hogan, T., N. Lederer, D. Clark, and W. Fertig. 2009. Annotated checklist of vascular ﬂ ora: Black Canyon of the Gunnison National Park. Natural Resource Technical Report NPS/NCPN/NRTR— 2009/227. National Park Service, Fort Collins, Colorado.

NPS 144/100146, July 2009 ii Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Contents

Figures ...... v Tables ...... v Abstract ...... vii Acknowledgements ...... ix

1 Introduction ...... 1 1.1 Background and justiﬁ cation ...... 1 1.2 Objectives ...... 1 1.3 Study area ...... 1 1.3.1 History and setting ...... 1 1.3.2 Geology ...... 2 1.3.3 Vegetation ...... 2 1.3.4 Previous botanical studies ...... 4

2 Methods ...... 7

3 Results ...... 9 3.1 Summary of the ﬂ ora of Black Canyon of the Gunnison National Park ...... 9 3.2 Life form and biogeographic diversity ...... 10 3.3 Plants of special concern ...... 10 3.4 Non-native species and noxious weeds ...... 10 4 Discussion ...... 13

5 Literature Cited ...... 15

Appendices ...... 17 Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes...... 19 Appendix B. Potential (unconfi rmed) vascular plant taxa of Black Canyon of the Gunnison National Park...... 71 Appendix C. Rejected (falsely reported or questionable) vascular plant taxa of Black Canyon of the Gunnison National Park...... 89 Appendix D. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by life form with ecological notes...... 91

Contents iii

Figures

Figure 1.3.1. Location of Black Canyon of the Gunnison National Park, Montrose County, Colorado...... 3 Figure 1.3.4. Increase in the number of plant taxa conﬁ rmed as present in Black Canyon of the Gunnison National Park by decade, 1950–2007...... 6

Tables

Table 1.3.4. Chronology of plant-collecting efforts in Black Canyon of the Gunnison National Park...... 5 Table 3.1. Statistical summary of the ﬂ ora of Black Canyon of the Gunnison National Park...... 9 Table 3.3. Plant species of conservation concern from Black Canyon of the Gunnison National Park...... 11 Table 3.4. Noxious weed species of Black Canyon of the Gunnison National Park...... 12

Contents v

Abstract

In 2005–2006, the National Park Service’s Northern Colorado Plateau Network received funding to develop a revised checklist of the vascular plant fl ora of Black Canyon of the Gunnison National Park. This entailed a thorough review of existing literature, re-examination of specimens from the Black Canyon of the Gunnison National Park herbarium and other state and national museum collections, and additional fi eld work to corroborate unverifi ed species reports and locate new species. Based on this study, 532 vascular plant taxa have been confi rmed or reliably reported as occurring in the park (470 are known from validated herbarium specimens and 62 from unvouchered literature reports). Another 20 species formerly considered part of the park fl ora have now been shown to be falsely reported. The current fl ora of Black Canyon of the Gunnison National Park represents about 17% of the known fl ora of Colorado, and 28% of the fl ora of the Colorado Plateau ecoregion within Colorado. Although no plant taxa are listed as threatened or endangered, at least 18 are considered species of concern by the state natural heritage program. Introduced plants make up 13.7% of the park fl ora (73 species), of which 19 species are considered noxious weeds by the State of Colorado. Further surveys in Black Canyon of the Gunnison National Park are likely to fi nd additional new species (at least 290 taxa are known from similar habitats outside the park boundary), especially among late-fl owering, weedy, or wetland taxa.

Contents vii

Acknowledgements

We would like to thank the following individuals for their assistance with this project: Dr. Angela Evenden, botanist and former leader of the Northern Colorado Plateau Network (NCPN) program in Moab, Utah, was an active proponent of this project and helped secure funding for ﬁ eld work and herbarium studies in 2006. Libby Nance (former NCPN database specialist), and Margaret Beer (former NCPN database manager) assisted with development and quality control of the park’s NPSpecies database. Libby Nance, Janet Coles, and Helen Thomas also provided data from recent vegetation mapping and wetland inventories on the park. Thanks to Black Canyon of the Gunnison National Park staﬀ , especially Danguole Bockus and Paul Zaenger, for providing logistical help with the herbarium- review phase of the project and comments on the draft manuscript. To any others whom we may have inadvertently forgotten, please accept our apologies (and thanks). Funding for this project was provided by the National Park Service.

Contents ix

vertebrates and vascular plants for each park unit 1 Introduction in the network (Evenden et al. 2002). This eff ort has entailed reviewing existing species checklists, In 2006, the National Park Service (NPS) estab- corroborating the identifi cation of specimens in lished a cooperative agreement with the Univer- park museum and herbarium collections, locat- sity of Wyoming, through the Rocky Mountains ing additional specimen records from other mu- Cooperative Ecosystem Studies Unit, to produce seums, mining relevant literature and datasets, a series of technical reports summarizing the vas- and conducting fi eld surveys to locate new taxa cular plant fl ora of each of the 16 national park or confi rm unsubstantiated reports.† All of these units of the Northern Colorado Plateau Network data have been synthesized in NPSpecies, the (NCPN). These reports are based on existing in- park service’s standardized database for docu- formation in the park service’s NPSpecies data- menting park status (present, probably present, base system and augmented by further data min- unconfi rmed, falsely reported), abundance, resi- ing, reviews of state and regional herbaria, and dency, nativity, and rarity. With the production of new fi eld surveys to resolve the status of many a technical series of reports on the fl ora of each reported and unconfi rmed species. The objective park, the NPSpecies data will now be available to of these reports is to summarize the current state a broader audience. of knowledge of the fl ora of each park for the benefi t of NPS biologists and interpretive staff , outside researchers, and the general public. The 1.2 Objectives following report addresses the vascular fl ora of The objectives of this project were to: Black Canyon of the Gunnison National Park, in west-central Colorado. Other NCPN park units 1. Document the current status (present, are addressed in separate reports. reported, potential, or falsely reported) of all vascular plant species that have been cited as occurring in Black Canyon of the Gunnison 1.1 Background and justifi cation National Park, based on a review of herbari- One of the primary missions of the NPS is “to um collections and past literature. conserve unimpaired the natural and cultural resources and values of the national park sys- 2. Develop an annotated checklist of the fl ora tem for the enjoyment of this and future genera- of Black Canyon of the Gunnison National tions” (NPS 2000). Due to their high degree of Park, with supplemental information on protection and distribution across a wide variety taxonomic synonyms, distribution, within- of habitat types, the nation’s national park units park abundance, fl owering period, growth make a signifi cant contribution to the protection form, nativity, habitat, and sources for each of biological diversity (Noss and Cooperrider record. 1994). Unfortunately, the full contribution of the National Park System is not known, as many park 1.3 Study area units have traditionally lacked comprehensive biological inventories or monitoring programs to 1.3.1 History and setting identify the composition and condition of their The Black Canyon of the Gunnison River is one of biota. Knowing the status of park fl ora and fauna the steepest and narrowest canyons in the world. is critical for eff ective park management and for At its deepest point, the river is nearly 2,300 feet systematically identifying and fi lling gaps in the below the canyon rim; at its narrowest, it is only nation’s protected area network (Margules and 40 feet wide. Due to its remote location and rug- Pressey 2000). ged depths, the Black Canyon remained poorly explored until the middle of the nineteenth cen- To rectify this shortcoming, Congress passed tury. Captain John Gunnison, who explored the the National Parks Omnibus Management Act Gunnison River drainage as a potential route in 1998, to provide guidance and funding for for a transcontinental railroad in 1853, reached an inventory and monitoring program across the upper end of the canyon (in the area that is the entire National Park System. Since 2000, the now Curecanti National Recreation Area) but NCPN Inventory and Monitoring Program has did not descend into its depths. The Denver and been developing and verifying species lists of

†Taxa refer to all named species, subspecies, and varieties.

Chapter 1: Introduction 1 Rio Grande Railroad constructed a narrow-gauge sists of 1.3–1.7 billion year-old Precambrian rail line to the upper end of the canyon in 1882, rocks. The oldest outcrops in the park are derived and considered extending the route through the from sedimentary and igneous rocks that were re- entire Black Canyon. Byron Bryant led a team of crystallized by heat and pressure to form gneisses surveyors into the canyon in the winter of 1883, and schists interlaced by pinkish pegmatite. Over and, after four months of diffi cult work, con- time, these rock layers were faulted, folded, and vinced the railroad to fi nd an easier route else- compressed into nearly vertical, accordion-like where. An expedition in 1900 sought to explore bands. Later in the Precambrian, volcanic mag- the canyon bottom by wooden boat in search of ma intruded into these rocks, forming a plug of a location to divert Gunnison River water to agri- granite at the mid-reach of the canyon (Chronic cultural communities in the Uncompahgre Valley. 1988). This expedition failed, but in the following year, Will Torrence and Abraham Lincoln Fellows suc- Late in the Precambrian, the ancestral Gunnison ceeded in running the river on boats constructed Plateau had eroded to a level peneplain. During of infl atable rubber mattresses (Vandenbusche most of the Paleozoic and Mesozoic, the area was 1980). Construction of the six-mile Gunnison near or below sea level and subjected to cycles of Tunnel, to divert part of the river’s fl ow, com- deposition and erosion of marine shales, lime- menced at the upper end of the canyon in 1905; it stones, and sandstones. Only the later Jurassic and was completed four years later. Cretaceous sedimentary layers are still present (earlier layers eroded away eons ago). Mesozoic In later years, Torrence became an advocate for outcrops in Black Canyon of the Gunnison NP protecting the Black Canyon for public enjoy- include the Dakota Sandstone (a beach deposit) ment. In March 1933, President Herbert Hoover and the Mancos Shale (marine clays that weather used the Antiquities Act to create Black Canyon into adobe-like badlands), present on mesa tops of the Gunnison National Monument (NM) for and slopes along the park’s western boundary “the preservation of the spectacular gorges and (Chronic 1988). additional features of scenic, scientifi c, and edu- cational interest” (Evenden et al. 2002). Since About 60 million years ago, in the late Cretaceous, 1935, the boundaries of the monument have tectonic forces during the Laramide Orogeny ele- been expanded on six occasions—most recently vated the Gunnison Plateau and adjacent areas of in 1999, when 4,400 acres of adjacent wilderness the Colorado Plateau and southern Rocky Moun- were added and the unit was converted to nation- tains. Starting about 35 million years ago, volcanic al park status (Figure 1.3.1). Presently, the park activity in the West Elk and San Juan mountains covers 30,750 acres (12,444 ha), of which nearly buried much of the Black Canyon area under a half is offi cially designated wilderness (Evenden thick mantle of volcanic ash and mudfl ows. Fol- et al. 2002). lowing the last major volcanic eruptions about 10–15 million years ago, the Gunnison River be- Black Canyon of the Gunnison National Park came entrenched within its current drainage and (NP) is located about 15 miles east of Montrose, began slicing through the volcanic overburden in eastern Montrose County, Colorado. The park and underlying Mesozoic sediments at the same borders Curecanti National Recreation Area time the region experienced a second period of (NRA) along its southeastern border. The recre- uplift. Ultimately, the river reached Precambrian ation area contains three dams that regulate the basement rock about 2 million years ago, and be- fl ow of the Gunnison River into Black Canyon. gan carving the Black Canyon. A steep gradient, Elevation within the park ranges from 5,400 feet abundant runoff from the Rocky Mountains, and (1,645 m) at the bottom of Black Canyon to 8,775 time have allowed the river to downcut at a rate feet (2,675 m) on the summit of Signal Hill. Aver- of 33 cm (1 foot) every thousand years (Chronic age annual precipitation is 16–20 inches (400–508 1988). mm), with about one-quarter coming as snow. Temperature ranges from an average of 15°F 1.3.3 Vegetation (–9°C) in winter to 85°F (29°C) in mid-summer (Evenden et al. 2002). Six general vegetation types have been documented within Black Canyon of the Gunnison NP (see NCPN in ed. for a more detailed classifi cation). 1.3.2 Geology Extensive areas along the summit of the plateau Most of the Black Canyon of the Gunnison con- above the canyon (especially on the south rim) are

2 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Black Canyon of the GunnisonBlackCapito CanyonZion lNational Reef National of National th Park e ParkGunnison Park Colorado Nation al P ark Colorado

Visitor Center

347

012 Map location Kilometers 012 Miles

Established March 2, 1933 Legend 1,645–2,675 m elevation Park boundary (NPS) (5,400–8,775 ft elevation) Park easement 12,444 ha (30,750 ac) Paved roads Park inholdings Unpaved roads Curecanti NRA (NPS) Intermittent streams U.S. Bureau of Land Management Rivers Private

NCPN boundary

MAP CREATED BY ANETH WIGHT, NCPN (DERIVED FROM EVENDEN ET AL. 2002) Figure 1.3.1. Location of Black Canyon of the Gunnison National Park, Montrose County, Colorado.

Chapter 1: Introduction 3 covered by thickets of Gambel oak (Quercus gam- extent of their explorations within the modern belii) and Saskatoon serviceberry (Amelanchier boundaries of Black Canyon of the Gunnison NP alnifolia). Openings within oak scrubland may be will likely be forever unknown. dominated by mountain big sagebrush (Seriphidi- um vaseyanum) grasslands with an understory of The earliest botanist known to have entered needle-and-thread grass (Hesperostipa comata), Black Canyon was Carl Albert Purpus, who col- muttongrass (Poa fendleriana), and an assortment lected the type specimen of Purpus’ sullivantia of perennial forbs. Rocky areas along the north (Sullivantia hapemannii var. purpusii) within the rim of the canyon and on south-facing canyon canyon in 1892 (Table 1.3.4, Figure 1.3.4). In the slopes tend to be occupied by two-needle pinyon ensuing 60 years, botanical exploration within the (Pinus edulis) and Utah juniper (Sabina osteosper- Black Canyon area was limited. Edwin Payson, of ma) interspersed with Gambel oak, serviceberry, the University of Wyoming, collected the type of and alder-leaf mountain mahogany (Cercocar- mountain pepperweed (Lepidium montanum var. pus montanus). Adobe badlands of semi-barren tenellum) in Black Canyon in 1917. William Pen- Mancos Shale occur sporadically along the west- land documented nine new species for the area in ern park boundary and support desert shrubland 1942. Margaret Douglass added matrimony-vine vegetation of scattered Utah juniper, shadscale (Lycium barbarum) to the Black Canyon fl ora in (Atriplex confertifolia), and mat saltbush (Atriplex 1954. corrugata). These badlands contain a large number of desert species that do not occur elsewhere The fi rst systematic eff ort to document the com- in the park. plete fl ora of Black Canyon of the Gunnison NM began in 1959. Over the next four years, the fl ora North-facing slopes within the inner canyon of the monument increased from 12 to 152 spe- support forests of Douglas-fi r (Pseudotsuga men- cies (an increase of 92%) (Figure 1.3.4). Monu- ziesii), blue spruce (Picea pungens), and quaking ment naturalists W. D. Wetlaufer and Pat and aspen (Populus tremuloides). These woodlands Carol Miller added 48 new species in the process may be replaced by Gambel oak on drier sites. of developing the monument’s herbarium. Other Moist, shaded cliff -faces provide a micro-habitat collectors of note included University of Colora- for several rare species, such as Black Canyon do professor William A. Weber (59 new species), gilia (Gilia penstemonoides) and Purpus’ sullivan- J. M. Rominger (23 new grass taxa), A. S. Hyde, tia (Sullivantia hapemanii var. purpusii), or plants Richard Beidelman, and B. Vogel (10 new species usually found at much higher elevations, such as combined) (Hogan 2006). spotted saxifrage (Ciliaria austromontana). The canyon bottom is lined by a narrow band of ri- The next signifi cant eff ort to document the mon- parian forest dominated by box-elder (Negundo ument’s fl ora took place 20 years later, when Wil- aceroides), narrowleaf cottonwood (Populus an- liam Weber resumed his research on the fl ora of gustifolia), and western chokecherry (Padus vir- Black Canyon in collaboration with Hans Beck, giniana), and occasional ponderosa pine (Pinus of the University of Colorado herbarium; Claudia ponderosa) (Evenden et al. 2002; Hogan et al. Rector, of Colorado National Monument; and 2005). Don Kennedy, of Black Canyon of the Gunnison NM. Weber (1983) documented 178 new species for the monument and developed a list of addi- 1.3.4 Previous botanical studies tional species to be expected within the area with Because of the ready access provided by the additional fi eld work. Weber’s work brought the Denver and Rio Grande Railroad, the Gunnison total number of species confi rmed for the monu- River drainage attracted a large number of fi eld ment to 347. botanists in the late nineteenth and early twen- tieth centuries, including Marcus E. Jones, E. L. A new wave of botanical exploration began in the Greene, Herbert Wheeler, Charles Crandall, and early years of the twenty-fi rst century. In 2002, Charles Baker. Unfortunately, the locality data the NCPN (in ed.) vegetation mapping team con- from the specimens procured by these early visi- ducted sampling across Black Canyon of the Gun- tors is usually vague and often limited only to the nison NP, and reported 86 new species for the name of one of the railroad towns (Sapinero or fl ora. Steve Dewey and Kim Andersen, of Utah Cimarron) now located largely within adjacent State University, reported three additional spe- Curecanti NRA (Hogan et al. 2009). Thus the full cies during a survey of noxious weeds in the park

4 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park in 2003 (Dewey and Andersen 2005). Tim Hogan, the Black Canyon of the Gunnison NP herbari- Nan Lederer, Dina Clark, and Luke Tembrock, of um (doubling it in size). Hogan and others (2005) the University of Colorado Museum Herbarium, discovered 93 new species and conﬁ rmed (with conducted a general ﬂ oristic survey of the park in vouchers) 28 of the taxa reported previously by 2004, and collected more than 450 specimens for the NCPN veg mapping crew.

Table 1.3.4. Chronology of plant-collecting efforts in Black Canyon of the Gunnison National Park. # of new Year Collector(s) Comments taxa 1892 C. A. Purpus 1 Carl Albert Purpus collected the holotype of Purpus’ sullivantia (Sullivantia hapemanii var. purpusii) in Black Canyon in 1892. The full extent of Purpus’s collecting within the future national park is not known. 1917 E. B. Payson 1 Edwin Payson collected the type of mountain pepperweed (Lepidium montanum var. tenellum) in Black Canyon. The full extent of Payson’s collecting within the future national park is not known. 1942 C. W. T. Penland 9 William Penland collected at least 9 new species for Black Canyon of the Gunnison NM (deposited at the University of Colorado). The full extent of Penland’s collecting in the monument is not known. 1954 M. Douglass 1 Margaret Douglass collected matrimony-vine (Lycium barbarum). 1959– W. D. Wetlaufer, 140 Park naturalist W. D. Wetlaufer collected 33 new species for Black Canyon of 1963 W. A. Weber, P. the Gunnison National Monument from 1959 to 1961. William Weber, of the Miller, C. Miller, University of Colorado herbarium, discovered 59 new species for the monument R. S. Beidelman, in 1960 and 1962. Pat and Carol Miller, naturalists for Black Canyon and Colorado A. S. Hyde, J. M. national monuments, added 15 new species in 1961. J. M. Rominger collected 23 Rominger, and B. new grass species for the monument in 1962. A. S. Hyde, Richard Beidelman, and Vogel B. Vogel contributed 10 new species from 1961 to 1963. 1974 M. Cottrell and C. 15 Cottrell and Gregory collected 28 specimens for the BLCA herbarium and added Gregory 15 new species to the monument fl ora. 1980 K. Koepsel and W. 2 Kirk Koepsel collected elegant columbine (Aquilegia elegantula) and William Grey Grey discovered Black Canyon gilia (Gilia penstemonoides) during his research on rare plants of Black Canyon. 1983 W. A. Weber, H. 178 William Weber and Hans Beck, of the University of Colorado; Don Kennedy, Beck, D. Kennedy, from Black Canyon of the Gunnison NM; and Claudia Rector, from Colorado NM, and C. Rector collected 239 specimens for the BLCA herbarium and 178 new vascular plant taxa for the monument fl ora. 1984 E. Elder 1 Ellen Elder collected leopard-lily (Fritillaria atropurpurea). 2002 NCPN vegetation 86 Vegetation mappers with the NCPN (in ed.) reported (without vouchers) 140 new mapping team reported species for the park. At least 28 of these were later corroborated with vouchers by Hogan and others (2005), but 58 remain unverifi ed. 2003– T. Hogan, N. 121 + 3 Tim Hogan, Nan Lederer, Dina Clark, and Luke Tembrock, of the University of 2004 Lederer, D. Clark, reported Colorado Museum Herbarium, collected 460 specimens throughout the park L. Tembrock, S. as part of a general fl oristic survey. Hogan and others (2005) discovered 93 Dewey, and K. new species for the Black Canyon NP fl ora and confi rmed 28 additional species Andersen previously reported by the NCPN vegetation mapping crew. Dewey and Andersen (2005) reported 3 additional species during a survey of noxious weeds in 2003.

Chapter 1: Introduction 5 500

450

400

350

300

250

200

150 Number of vascular taxa 100

0 1950 1960 1970 1980 1990 2000 2007 Year

Figure 1.3.4. Increase in the number of plant taxa conﬁ rmed as present in Black Canyon of the Gunnison National Park by decade, 1950–2007.

6 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Each list was annotated with supplemental infor- 2 Methods mation on life form, geographic range (within the context of Colorado), population size in the park In 2005, Tim Hogan and Nan Lederer, of the (derived from Hogan et al. 2005, NCPN in ed., and University of Colorado Museum Herbarium, fi eld observations), fl owering period, distribu- reviewed all mounted specimens in the Black tion across general vegetation types, nativity, data Canyon of the Gunnison National Park her- source (collector and collection number from barium (BLCA) to determine whether each was a sample specimen or literature citation), year correctly identifi ed to species, subspecies, or va- of earliest collection (from herbarium records), riety following the nomenclature of Weber and and relevant synonymy and taxonomic problems Wittmann (2001). Hogan and others (2005) also (based on review of Barkworth et al. 2003, 2007; reviewed relevant herbarium collections from the Barneby 1989; Cronquist 1994; Cronquist et al. University of Colorado (COLO). As necessary, 1972, 1977, 1984, 1997; Dorn 2001; Flora of North they made corrections and annotations to each America Editorial Committee 1993, 1997, 2000, specimen and to the voucher records in the park’s 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c; Automated National Cataloging System (ANCS+) Holmgren et al. 2005; Kartesz 2003; Welsh et al. and NPSpecies database. 2003; and the Integrated Taxonomic Information System [ITIS] website, http://www.itis.gov/). The Based on this herbarium work and a review of annotations to these lists and label data from the park references (Dewey and Andersen 2005; BLCA herbarium review were then incorporated NCPN in ed.; Weber 1983), we developed four into the NPSpecies database for the park. lists:

1. A revised checklist of species that were present in the park (confi rmed by an authenticated voucher; included in Appendices A and D); 2. A list of species that were reported in the literature but not corroborated with a voucher (equivalent to “probably present” in NPSpe- cies terminology; included in Appendices A and D); 3. A list of potential species (“unconfi rmed” in NPSpecies) known from the vicinity of Black Canyon of the Gunnison National Park based on Weber and Wittmann (1992) and data from the University of Colorado Museum Herbarium digital specimen database (www.cumuseum. colorado.edu/ Research/Botany/ Databases/search.php) (included in Appendix B); and 4. A list of species that are now known to be misidentifi ed or highly questionable (Ap- pendix C).

Chapter 2: Methods 7

specimen (Table 3.1, Appendix A). Of these, 59 3 Results were reported by the NCPN (in ed.) and 3 from Dewey and Andersen (2005). Taken together, the 3.1 Summary of the fl ora of Black number of confi rmed and reported vascular plant taxa in Black Canyon of the Gunnison NP is 532 Canyon of the Gunnison National (Table 3.1, Appendices A, D). Park Based on a review of specimens in the BLCA and Following our review of literature and herbarium COLO herbaria (correcting for misidentifi cations collections from neighboring Curecanti National and collections from outside the park) and results Recreation Area, we recognized 290 species (Ap- of the 2003–2004 University of Colorado fl oristic pendix B) that could potentially be found within survey (Hogan et al. 2005), the number of vas- Black Canyon of the Gunnison NP (Weber 1983; cular plant taxa confi rmed as occurring in Black Hogan and Lederer NPSpecies database 2006). Canyon of the Gunnison NP with an authenti- Most of these species are known from compa- cated voucher stands at 470 (Table 3.1, Appendi- rable habitats elsewhere in Montrose County, or ces A, D). This number is slightly lower than the have been collected from within a 5–7-mile radius 485 reported by Hogan and others (2005), due to of the park. the elimination of some duplicate names. An ad- Of the pool of species documented or reported ditional 62 taxa have been reported for the park for Black Canyon of the Gunnison NP, we found but have not been corroborated with a voucher

Table 3.1. Statistical summary of the ﬂ ora of Black Canyon of the Gunnison National Park. Number Number Category conﬁ rmed additionally Total in park reported for park Taxonomic diversity Total taxa 470 62 532 (including varieties and subspecies) Full species 465 61 526 (excluding varieties and subspecies) Families 87 3 90 Life form diversity Tree taxa 13 1 14 Shrub taxa 53 9 62 Perennial forb taxa 242 35 277 Annual forb taxa 73 10 83 Perennial graminoid taxa 73 6 79 Annual graminoid taxa 4 1 5 Fern taxa 12 0 12 Biogeographic diversity Introduced taxa 63 10 73 Native taxa Locally endemic taxa 12 1 13 Regionally endemic taxa 29 6 35 Disjunct taxa 1 0 1 Peripheral taxa 7 4 11 Sparse taxa 2 0 2 Widespread taxa 356 41 397 Total native taxa 407 52 459 The number of taxa and families is based on taxonomic concepts of Weber and Wittmann (2001). See appendices for an ex- planation of the various categories.

Chapter 3: Results 9 that 11 taxa were falsely reported based on mis- 3.3 Plants of special concern identifi ed voucher specimens in the BLCA and Black Canyon of the Gunnison NP has no plants COLO herbaria. We consider an additional 9 spe- listed as threatened or endangered under the cies cited (but not vouchered) by the NCPN veg- U.S. Endangered Species Act. Only three spe- etation mapping team (NCPN in ed.) to be ques- cies, Rocky Mountain spring-parsley (Loma- tionable based on the absence of suitable habitat tium concinnum), Rocky Mountain thistle (Cir- or the distance of Black Canyon of the Gunnison sium perplexans), and Uncompahgre bladderpod NP to the known or expected range of each spe- (Lesquerella vicina), are listed as sensitive by the cies (Appendix C). U.S. Bureau of Land Management’s Colorado of- The confi rmed and reported fl ora of Black Can- fi ce or the U.S. Forest Service (Table 3.3) (Spack- yon of the Gunnison NP represents 16.9% of the man et al. 1997). The Colorado Natural Heritage 3,145 native and naturalized taxa documented Program (2007) tracks 17 taxa known from Black for the state of Colorado by Weber & Wittmann Canyon of the Gunnison NP as species of con- (1992), and 56.3% of the state’s 160 plant families cern, and one other on its watch list of potential (Table 3.1). The park contains 66.7% of the 797 species of concern (Table 3.3). Hogan and others species reported for Montrose County (Kartesz (2005) discovered the fi rst populations of Rocky 2003). At the ecoregional scale, Black Canyon of Mountain thistle and Uncompahgre bladderpod the Gunnison NP contains 28.0% of the 1,898 within the park in 2004, and also mapped occur- species reported for the Colorado portion of the rences of two other rare species: juniper tumbl- Colorado Plateau (Kartesz 2003). emustard (Thelypodiopsis juniperorum) and Pur- pus’ sullivantia. 3.2 Life form and biogeographic 3.4 Non-native species and noxious diversity weeds Although ecologically dominant, tree and shrub species make up only 14.3% of the confi rmed and Introduced plant species (not historically native reported plant taxa of Black Canyon of the Gun- to Colorado or North America) make up 13.7% nison NP (see Table 3.1). Perennial forbs are the of the confi rmed and reported fl ora of Black Can- most species-rich life form in the park fl ora, with yon of the Gunnison NP (see Table 3.1). This fi g- 277 known and reported taxa (52.1% of the total ure is lower than the statewide average of 15.6% fl ora). Annual forbs and grass-like plants com- for non-native species (Rejmanek and Randall prise less than 17% of the fl ora. Like trees and 1994). Of the park’s 73 introduced plant species, shrubs, perennial graminoids are ecologically im- 19 are listed as noxious by the State of Colorado portant but relatively species-poor, representing (Table 3.4) (Dewey and Andersen 2005; CDA just 14.8% of the total fl ora. The diversity of ferns 2007). Plants on the Colorado state noxious weed and fern-allies in Black Canyon of the Gunnison list fall into three categories with diff ering man- NP is especially low (2.2%) (see Table 3.1). agement objectives and requirements (Colorado Noxious Weed Act of 1996 Title 35 Article 5.5). Of the park’s 459 native taxa, 397 (86.5%) are Weeds on List A are designated for full eradica- widespread across Colorado and western North tion wherever they occur. Containment of in- America. Only 13 species (2.8% of the native festations is the goal for species on List B, and fl ora) have extremely limited global distributions voluntary control, education, and research is the (local endemics found within an area of less than objective for species on List C. Currently, 14 spe- 1° of latitude × 2° of longitude; see appendices for cies from Black Canyon of the Gunnison NP are complete defi nitions of biogeographic catego- on List B, and 5 are on List C; none is on List A ries). Regional endemics (largely restricted to the (Table 3.4). Dewey and Andersen (2005) and Ho- Colorado Plateau) comprise an additional 7.6% gan and others (2005) mapped infestations of 19 of the native park fl ora (see Table 3.1). Species at noxious weeds and invasive non-native species the extreme edge of their global range in Colora- within the park. do (peripherals), and those found sparsely over a wide range, make up 2.4% and 0.4% of the park’s native fl ora, respectively (see Table 3.1). Shelton’s violet (Viola biternata) is the only disjunct species presently known from the park.

10 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Table 3.3. Plant species of conservation concern from Black Canyon of the Gunnison National Park.

TNC CNHP Family Species Common name global Legal status status rank Apiaceae Aletes anisatus Rocky Mountain Indian G3 Watch, S3 (Umbelliferae) parsley Apiaceae Cymopterus planosus Rocky Mountain spring- G3 SOC, S3S4 (Umbelliferae) parsley Apiaceae Lomatium concinnum adobe desert-parsley G2G3 CO BLM: Sensitive SOC, S2S3 (Umbelliferae) Asteraceae Cirsium perplexans Rocky Mountain thistle G2 CO BLM: Sensitive, SOC, S2S3 (Compositae) USFS R2: Sensitive Brassicaceae Boechera gunnisoniana Gunnison’s rockcress G3 SOC, S3 (Cruciferae) Brassicaceae Boechera oxylobula Glenwood Springs G3 SOC, S3 (Cruciferae) rockcress Brassicaceae Lesquerella vicina Uncompaghre G2 CO BLM: Sensitive SOC, S2 (Cruciferae) bladderpod Brassicaceae Stanleya albescens white prince’s-plume G3 SOC, S3 (Cruciferae) Brassicaceae Thelypodiopsis juniper tumblemustard G2 SOC, S2 (Cruciferae) juniperorum Capparaceae Cleomella palmeriana Palmer’s stinkweed G3G4 SOC, S3 (Cleomaceae) Crossosomataceae Forsellesia meionandra spiny greasebush G5T3 SOC, S3 Hydrophyllaceae Phacelia splendens Eastwood’s phacelia G3 SOC, S3 Lamiaceae Monardella odoratissima stinking horsemint G4G5 SOC, S2 (Labiatae) T3T5 Loasaceae Nuttallia pterosperma wing-seed stickleaf G3G5 SOC, S3? Polemoniaceae Gilia penstemonoides Black Canyon gilia G3 SOC, S3 Ranunculaceae Myosurus cupulatus horseshoe mousetail G4 SOC, S1? Saxifragaceae Sullivantia hapemanii Purpus’ sullivantia G3T3 SOC, S3 var. purpusii Scrophulariaceae Penstemon retrorsus Adobe Hills G3 SOC, S3 beardtongue This list was derived from the Colorado Natural Heritage Program (CNHP 2007). All species are conﬁ rmed as present in the park.

Codes: TNC global rank assesses abundance and conservation priority on a scale of 1–5 (1 being extremely vulnerable and 5 being secure) for full species (G) and varieties or subspecies (T) across their entire range. A “?” indicates uncertainty in the rank, Q = taxonomic questions, and NR = not ranked. A comparable scoring system is used to measure state abundance (S) in Colorado under the column CNHP status. TNC = The Nature Conservancy. Under legal status, USFWS R2= U.S. Fish and Wildlife Service, Region 2. CO BLM = U.S. Bureau of Land Management, Colorado ofﬁ ce. Under CNHP status, SOC denotes plants on the state “Species of Concern” list and Watch indicates species on the watch list that may ultimately warrant listing as species of concern (CNHP 2007).

Chapter 3: Results 11 Table 3.4. Noxious weed species of Black Canyon of the Gunnison National Park.

CO noxious Family Species Common name Park status weed list Asteraceae (Compositae) Acosta diffusa diffuse knapweed Present List B (Centaurea diffusa) Asteraceae (Compositae) Acosta maculosa spotted knapweed Present List B (Centaurea maculosa) Asteraceae (Compositae) Acroptilon repens Russian knapweed Present List B (Centaurea repens) Asteraceae (Compositae) Arctium minus burdock Present List C Asteraceae (Compositae) Breea arvensis Canada thistle Present List B (Cirsium arvense) Asteraceae (Compositae) Breea incana Canada thistle Present List B (Cirsium arvense) Asteraceae Carduus nutans var. macrolepis musk thistle Present List B (Compositae) Asteraceae (Compositae) Cirsium vulgare bull thistle Present List B Boraginaceae Cynoglossum offi cinale common hound’s-tongue Present List B Brassicaceae (Cruciferae) Cardaria draba whitetop Present List B Brassicaceae (Cruciferae) Hesperis matronalis Dame’s rocket Reported List B Chenopodiaceae Halogeton glomeratus halogeton Present List C Convolvulaceae Convolvulus arvensis fi eld bindweed Present List C Geraniaceae Erodium cicutarium stork’s-bill Reported List B Poaceae (Gramineae) Anisantha tectorum cheatgrass Present List C (Bromus tectorum) Poaceae (Gramineae) Elytrigia repens quackgrass Present List B Scrophulariaceae Linaria vulgaris butter-and-eggs Reported List B Scrophulariaceae Verbascum thapsus woolly mullein Present List C Tamaricaceae Tamarix ramosissima fi ve-stamen tamarisk Present List B This list was derived from the Colorado Department of Agriculture (2007).

12 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Many of the “missing” potential or reported taxa 4 Discussion consist of cryptic species that fl ower late in the growing season, or occur in undersampled wet- Since 1983, the confi rmed and reported fl ora of land or weedy habitats. Confi rmation of reported Black Canyon of the Gunnison National Park has species should remain a high priority, as some of increased by 35%, from 348 to 532 taxa due pri- these taxa may ultimately prove to be erroneous. marily to the fl oristic research of Weber (1983), Additional studies would also be useful to better Hogan et al. (2005), and others. Another 290 spe- refi ne the abundance and distribution of species have been documented from the vicinity (Ap- cies of high management interest (especially rare pendix A, B). Should all of these potential species plants and invasive non-natives). be found, the fl ora of Black Canyon of the Gunni- son National Park would increase by nearly 35%.

Chapter 4: Discussion 13

——. 1984. Volume 4, Subclass Asteridae (except 5 Literature Cited Asteraceae). Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Barkworth, M. E., K. M. Capels, S. Long, and M. Bronx, N.Y.: New York Botanical Garden. B. Piep, eds. 2003. Magnoliophyta: Com- 573 pp. melinidae (in part): Poaceae, part 2. Flora of North America north of Mexico, Volume 25. Cronquist, A., N. H. Holmgren, and P. K. Hol- New York: Oxford University Press. 783 pp. mgren, eds. 1997. Volume 3, Part A, Subclass Rosidae (except Fabales). Intermountain Barkworth, M. E., K. M. Capels, S. Long, L. K. fl ora: Vascular plants of the Intermountain Anderton, and M. B. Piep, eds. 2007. Mag- West, U.S.A. Bronx, N.Y.: New York Botani- noliophyta: Commelinidae (in part): Poace- cal Garden. 446 pp. ae, part 1. Flora of North America north of Mexico, Volume 24. New York: Oxford Dewey, S., and K. Andersen. 2005. An inventory University Press. 911 pp. of invasive non-native plants in Black Can- yon of the Gunnison National Park (2003): Barneby, R. C. 1989. Volume 3, Part B. Fabales. Final report. Utah State University. Weed In Cronquist, A., A. H. Holmgren, N. H. Science Research Project #SD0511A. Holmgren, J. L. Reveal, and P. K. Holmgren, eds. Intermountain fl ora: Vascular plants of Dorn, R. D. 2001. Vascular plants of Wyoming. the Intermountain West, U.S.A. Bronx, N.Y.: Third edition. Cheyenne, Wy.: Mountain New York Botanical Garden. 279 pp. West Publishing. 412 pp. Chronic, H. 1988. Pages of stone: Geology of Evenden, A., M. Miller, M. Beer, E. Nance, S. western national parks and monuments: Daw, A. Wight, M. Estenson, and L. Cudlip. Grand Canyon and the plateau country. 2002. Northern Colorado Plateau Vital Seattle, Wa.: The Mountaineers. 158 pp. Signs Network and Prototype Cluster plan for Natural Resources Monitoring: Phase I Colorado Department of Agriculture (CDA). report. October 1. Two volumes. National 2007. Colorado noxious weed list. www. Park Service, Northern Colorado Plateau cepep.colostate.edu/noxious.htm#CONWA. Network, Moab, Ut. 138 pp + app. Colorado Natural Heritage Program (CNHP). Flora of North America Editorial Committee. 2007. Tracked vascular plant species 1993. Pteridophytes and gymnosperms. (8/16/2007 version). www.cnhp.colostate. Flora of North America north of Mexico, edu/tracking/vascular.html. Volume 2. New York: Oxford University Cronquist, A. 1994. Volume 5, Asterales. In Press. 475 pp. Cronquist, A., A. H. Holmgren, N. H. Hol- ——. 1997. Magnoliophyta: Magnoliidae and mgren, J. L. Reveal, and P. K. Holmgren, eds. Hamamelidae. Flora of North America Intermountain fl ora: Vascular plants of the north of Mexico, Volume 3. New York: Ox- Intermountain West, U.S.A. Bronx, N.Y.: ford University Press. 590 pp. New York Botanical Garden. 496 pp. ——. 2000. Magnoliophyta: Alismatidae, Are- Cronquist, A., A. H. Holmgren, N. H. Holmgren, cidae, Commelinidae (in part), and Zin- and J. L. Reveal, eds. 1972. Volume 1, Geo- giberidae. Flora of North America north logical and botanical history of the region, of Mexico, Volume 22. New York: Oxford its plant geography and a glossary. The University Press. 352 pp. vascular cryptogams and the gymnosperms. Intermountain fl ora: Vascular plants of the ——. 2002a. Magnoliophyta: Commelinidae (in Intermountain West, U.S.A. Bronx, N.Y.: part): Cyperaceae. Flora of North America New York Botanical Garden. 270 pp. north of Mexico, Volume 23. New York: Oxford University Press. 608 pp. Cronquist, A., A. H. Holmgren, N. H. Hol- mgren, J. L. Reveal, and P. K. Holmgren, ——. 2002b. Magnoliophyta: Liliidae: Liliales and eds. 1977. Volume 6, The monocotyledons. Orchidales. Flora of North America north Intermountain fl ora: Vascular plants of the of Mexico, Volume 26. New York: Oxford Intermountain West, U.S.A. Bronx, N.Y.: University Press. 723 pp. New York Botanical Garden. 584 pp.

Chapter 5: Literature Cited 15 ——. 2003. Magnoliophyta: Caryophyllidae, part National Park Service (NPS). 2000. Manage- 1. Flora of North America north of Mexico. ment policies. U.S. Department of Interior, Volume 4. New York: Oxford University National Park Service, Washington, D.C. Press. 559 pp. Publication #NPS D1416. 137 pp. ——. 2005. Magnoliophyta: Caryophyllidae, part Northern Colorado Plateau Network (NCPN). 2. Flora of North America north of Mexico. in ed. Black Canyon of the Gunnison Volume 5. New York: Oxford University National Park vegetation classifi cation and Press. 656 pp. mapping project. Natural Resource Techni- cal Report. National Park Service, Denver, ——. 2006a. Magnoliophyta: Asteridae, part 6: Co. Asteraceae, part 1. Flora of North America north of Mexico. Volume 19. New York: Noss, R. F., and A. Y. Cooperrider. 1994. Saving Oxford University Press. 579 pp. nature’s legacy: Protecting and restoring biodiversity. Washington, D.C.: Island Press. ——. 2006b. Magnoliophyta: Asteridae, part 7: 416 pp. Asteraceae, part 2. Flora of North America north of Mexico. Volume 20. New York: Rejmanek, M., and J. M. Randall. 1994. Invasive Oxford University Press. 666 pp. alien plants in California: 1993 summary and comparison with other areas in North ——. 2006c. Magnoliophyta: Asteridae, part 8: America. Madrono 41(3):161–177. Asteraceae, part 3. Flora of North America north of Mexico. Volume 21. New York: Spackman, S., B. Jennings, J. Coles, C. Dawson, Oxford University Press. 616 pp. M. Minton, A. Kratz, and C. Spurrier. 1997. Colorado rare plant fi eld guide. Prepared Hogan, T. 2006. History of botanical surveys in for the Bureau of Land Management, U.S. Black Canyon of the Gunnison National Forest Service, and U.S. Fish and Wildlife Park and Curecanti National Recreation Service by the Colorado Natural Heritage Area. Report prepared for the Northern Program, Ft. Collins, Co. Colorado Plateau Network, National Park Service. 4 pp. Vandenbusche, D. 1980. The Gunnison country. Gunnison, Co.: B & B Printers. Hogan, T., N. Lederer, and D. Clark. 2005. Flo- ristic inventory of Black Canyon of the Gun- Weber, W. A. 1983. Checklist of plants of Black nison National Park and Curecanti National Canyon National Monument. Boulder, Co.: Recreation Area. Report prepared for the University of Colorado Museum. 14 pp. Northern Colorado Plateau Network by the Weber, W. A., and R. C. Wittmann. 1992. Catalog University of Colorado Museum Herbarium, of the Colorado fl ora: A biodiversity base- Boulder, Co. line. Niwot, Co.: University Press of Colo- Hogan, T., N. Lederer, D. Clark, and W. Fertig. rado (revised electronic version, January 2009. Annotated checklist of the vascular 2000). http://cumuseum.colorado.edu/Re- fl ora of Curecanti National Recreation Area. search/Botany/Databases/vascular_plants. National Park Service, Denver, Co. pdf. Holmgren, N. H., P. K. Holmgren, and A. Cron- ——. 2001. Colorado fl ora: Western Slope. Third quist. 2005. Volume 2, Part B Subclass Dille- edition. Boulder: University Press of Colo- niidae. Intermountain fl ora: Vascular plants rado. 488 pp. of the Intermountain West, U.S.A. Bronx, Welsh, S. L., N. D. Atwood, S. Goodrich, and L. N.Y.: New York Botanical Garden. 488 pp. C. Higgins. 2003. A Utah fl ora. Third edition, Kartesz, J. T. 2003. Synthesis of the North Ameri- revised. Provo, Ut: Brigham Young Univer- can fl ora, Version 2.0. sity. 912 pp. Margules, C. R., and R. L. Pressey. 2000. Sys- tematic conservation planning. Nature 405:243–253.

16 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park mgren et al. 2005; Welsh et al. 2003). No eff ort has been Appendices made to identify additional synonyms from monographs and fl oras preceding the early 1970s. The following appendices contain annotated checklists of the vascular fl ora of Black Canyon of the Gunnison Common name (Appendices A, B, C, and D) National Park. Appendix A lists all species that have been Common names are derived primarily from Welsh and confi rmed for the fl ora with an authenticated voucher others (2003) and the Intermountain Flora (Barneby 1989; specimen (including extant and historical records) or that Cronquist 1994; Cronquist et al. 1972, 1977, 1984, 1997; have been reliably reported for the park in the literature Holmgren et al. 2005). (Dewey and Andersen 2005; NCPN in ed.; Weber 1983). Vouchers were examined from the Black Canyon of the Life form (Appendices A, B, C, and D) Gunnison NP and University of Colorado herbaria (BLCA Life form is based on the stature and growth habit of typi- and COLO, respectively). cal members of a species at reproductive maturity. Trees Species entries in Appendix A are organized alphabetically are woody perennials with a single stem or trunk and are by family and genus. typically over 3.5 m tall. Shrubs are woody perennials with one to many trunks arising from near the base of the Appendix B includes species that may potentially occur plant and are usually less than 3.5 m tall. Perennial Forbs within Black Canyon of the Gunnison NP based on the (PerF) are broad-leaved dicots or monocots with non- proximity of the park to other populations or the presence woody stems (at least above ground level) that live for mul- of suitable habitat. tiple years. Annual Forbs (AnnF) are broad-leaved dicots or monocots with non-woody stems that complete their Appendix C lists species that have been previously re- life cycle (mature, fl ower, and die) in one year. Perennial ported for Black Canyon of the Gunnison NP but are now Graminoids (PerG) are linear-leaved, grass-like monocots considered erroneous because they are based on misiden- that live for multiple years. Annual Graminoids (AnnG) tifi ed specimens or questionable literature records. are linear-leaved, grass-like monocots that complete their life cycle (mature, fl ower, and die) in one year. Ferns and Appendix D contains the same confi rmed, historical, and fern-allies (Fern) are non-fl owering vascular plants that reported taxa as in Appendix A, but organized by life form reproduce by spores. rather than by family in order to better meet the needs of NPS interpretive specialists. Range (Appendices A, B, and C)

The codes and references for each fi eld in the appropriate Range represents the distribution of a species within appendices are summarized below. Colorado and the state’s contribution to the plant’s over- all global range. Disjunct (Disj) taxa have their Colorado Family (Appendices A, B, C, and D) distribution separated from the main, contiguous por- Family concepts and nomenclature follow Colorado Flora: tion of their range by a gap of more than 800 km (ca 500 Western Slope, third edition (Weber and Wittmann 2001). miles). Introduced (Intro) taxa are those that are not na- Alternative family names and treatments from other re- tive to Colorado or North America but which have be- gional fl oras (Dorn 2001; Flora of North America Editorial come naturalized (breeding on their own without human Committee 1993, 2002b; Holmgren et al. 2005; Welsh et al. assistance). Local Endemics (LocEn) are taxa whose en- 2003) are listed in parentheses. tire global range is restricted to an area of less than 16,500 km2 (ca 6,370 mi2, or 1° of latitude × 2° of longitude). Pe- Species name (Appendices A, B, C, and D) ripheral (Periph) taxa have a widespread global distribu- Nomenclature for scientifi c names and taxonomic con- tion but occur at the margin of their contiguous range cepts are derived from Weber and Wittmann (2001). Sci- in Colorado and occupy less than 5% of the state’s area entifi c names are listed in alphabetical rather than phylo- (usually only within a few miles of the state border). Re- genetic order. gional Endemics (RegEn) are taxa with a global range of 16,500–250,000 km2 (an area about the size of the state of Synonyms/Taxonomic notes (Appendices A, B, and C) Wyoming). Sparse taxa occur widely across Colorado or North America but their range within Colorado is small Pertinent synonyms are included for species with alterna- and patchy, with populations restricted to specialized or tive scientifi c names in other recent state and regional fl or- uncommon habitats. Widespread (Wide) taxa have global as (Barkworth et al. 2003, 2007; Barneby 1989; Cronquist ranges exceeding 250,000 km2 and occur over at least 10% 1994; Cronquist et al. 1972, 1977, 1984, 1997; Dorn 2001; of the state. Flora of North America Editorial Committee 1993, 1997, 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c; Hol-

Chapter 6: Appendices 17 Park status (Appendices A, C, and D) Year doc. (Appendix A) Park status identifi es whether a species is Present (Pres), Year documented indicates the fi rst year in which a spe- Historical (Hist), or Reported (Rep) for Black Canyon cies was confi rmed or reported to occur in Black Canyon of the Gunnison NP. Species are considered Present if a of the Gunnison NP. This year does not necessarily cor- confi rmed voucher or documented observation has been respond with the date of the voucher specimen cited un- made within park boundaries since 1970. Historical spe- der Source. “NA” indicates that the fi rst year is not known cies have been confi rmed with a voucher from the park (used only for “reported” taxa). but have not been relocated since before 1970. Reported species are listed for the park in the literature (Dewey and Comments (Appendices A, B, C, and D) Andersen 2005; NCPN in ed.; Weber 1983), but have not This column includes additional comments on the status been corroborated with a voucher specimen. Species that of a particular species in the park, the place of origin for may potentially occur in Black Canyon of the Gunnison introduced species, legal status, and other data. NP are listed in Appendix B. Rejected taxa (listed in Ap- pendix C) have either been falsely reported (FalsRep) in Nativity (Appendix D) the literature based on misidentifi ed herbarium specimens Nativity indicates whether a species is native to Colorado or are considered Questionable (Ques?) because park or Introduced (Intro) based on Weber and Wittmann reports are from well outside their known or expected (2001) and the Flora of North America Editorial Commit- range. tee (1993, 1997, 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c). Pop. size (Appendix A) Population size refers to the abundance and frequency Flower time (Appendix D) with which a species occurs within Black Canyon of the Flower time refers to the months in which a species is most Gunnison NP. Abundance data are derived from Hogan likely to be in fl ower. NA indicates a species that is a non- and others (2005), NCPN (in ed.), and personal obser- fl owering fern or gymnosperm. vations. Abundant (Abun) taxa have large populations, a broad ecological amplitude (often being dominant in one Habitat (Appendix D) to several vegetation types), or occur extensively across the The distribution of species across generalized habitats in park. Common (Com) taxa have large local populations Black Canyon of the Gunnison National Park is indicat- but may be restricted to a single vegetation type or do not ed by an “X” in the columns Sage/Grs through Wet and occur across the entire park. Uncommon (Unc) taxa have is derived from Hogan and others (2005) and NCPN (in low to medium-sized populations, typically have a narrow ed.). Sage/Grs = communities dominated by big sage- ecological amplitude (or at least contribute relatively little brush, needle-and-thread, and muttongrass. PinJun = to total vegetative cover), or have a small range across the two-needle pinyon–Utah juniper woodlands. Oak/MtBr park. Rare taxa usually have low population numbers and = woodlands dominated by Gambel oak and other mon- are restricted to a single location within the park. A “?” in- tane shrubs. Dg-fi r = Douglas-fi r–quaking aspen forests. dicates that population size is unknown. Wet = Wetlands (including seeps and springs). Additional Source (Appendix A) comments are included for species that are restricted to a particular soil/geologic type or are found mostly in dis- Sources are the evidence for the assigned park status. In turbed sites. the case of Present and Historical taxa, sources are herbarium records (with collector, collection number, and repository) that were corroborated during this study. For reported taxa, the source is a published or unpublished reference citing the species within the park. Specimen re- positories include the Black Canyon of the Gunnison NP (BLCA) and University of Colorado (COLO) herbaria.

18 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park A. ed Comments Palisades, Mesa Co., CO (Crandall s.n. RM) be base on misidentiﬁ macropetalum Year Year doc. 1962 1962 1983 1959 1983 2004 locality is Type 1959 2004 Native to Eurasia Miller 11375 (BLCA) (BLCA) Source (COLO) (BLCA) & Rector, Kennedy 16750 (BLCA) (BLCA) Hogan 2021 (BLCA) Tembrock Tembrock 4339 (BLCA) Pop. size Park status Range Life Form Shrub Wide Pres Com & Weber PerF Wide Rep Unc NCPN (in ed.) 2002 PerF Wide Pres Com Clark & PerF Intro Pres Unc Hogan & PerF Wide Rep Unc NCPN (in ed.) 2002 Common name Rocky Mountain maple box elder Tree Wide Pres Com Hogan 4110 Harriman’s yuccaHarriman’s Shrub Wide oniontaper-tip Pres PerF onionGeyer’s Wide Unc PerF Pres 17012 Weber WideSan Juan onion Unc Pres PerF Hogan 4130 Unc Wide Weber, Pres Unc Hogan 4230 Fendler’s Fendler’s sandwort textile onion PerF Wide Repball-head Raresandwort NCPN (in ed.) 2002 Report may mouse-ear chickweed Hooker’s Hooker’s sandwort .

var

var. var.

var. var. ssp. & var. hookeri, fendleri congesta, Acer negundo, A. negundo interius var. E. hookeri Includes vars. neomexicanum, douglasii, tripartitum Synonyms/ notes Taxonomic harrimaniae geyeri var. var. E. congesta congesta vulgare hookeri fontanum interius Negundo aceroides ssp. harrimaniaeYucca harrimaniae Y. Acer glabrum Eremogone hookeri Arenaria hookeri Allium acuminatum Allium geyeri A. geyeri Allium macropetalum Allium textile Eremogone fendleri Arenaria fendleri Eremogone congesta Arenaria congesta Cerastium fontanum C. vulgatum, vascular plant taxa of Black Canyon the Gunnison National Park, organized by family and rmed, historical, and reported c name with taxonomic and biogeographic notes. Aceraceae Agavaceae FamilyAceraceae Species name Alsinaceae (Caryophyllaceae) Alliaceae (Liliaceae) Alliaceae (Liliaceae) Alliaceae (Liliaceae) Alliaceae (Liliaceae) Alsinaceae (Caryophyllaceae) Alsinaceae (Caryophyllaceae) Alsinaceae (Caryophyllaceae) Appendix A. Conﬁ scientiﬁ

Chapter 6: Appendices 19 20

Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientiﬁ c name with Annotated Checklist ofVascular Flora: BlackCanyonoftheGunnison NationalPark taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Alsinaceae Pseudostellaria Stellaria jamesiana James’ chickweed PerF Wide Pres Com Weber 11363 1962 (Caryophyllaceae) jamesiana (COLO) Amaranthaceae Amaranthus albus tumble pigweed AnnF Intro Pres Unc Weber & 1983 Native to tropical Beck 16987 America (BLCA) Anacardiaceae Rhus aromatica ssp. R. trilobata var. squawbush Shrub Wide Pres Com Beck & 1983 trilobata trilobata Kennedy 17026 (BLCA) Anacardiaceae Rhus glabra smooth sumac Shrub Wide Pres ? Weber, 1983 Rector, & Kennedy 16844 (BLCA) Anacardiaceae Toxicodendron Rhus radicans, T. western poison Shrub Wide Pres Com Wetlaufer 1961 rydbergii radicans ivy s.n. (BLCA) Apiaceae Aletes anisatus Pteryxia Rocky Mountain PerF Wide Pres Unc Weber 11159 1960 (Umbelliferae) hendersonii Indian parsley (COLO) Apiaceae Cymopterus bulbosus onion spring- PerF Wide Pres Unc Lederer, 2004 (Umbelliferae) parsley Clark, Hogan, & Tembrock 04- 43 (BLCA) Apiaceae Cymopterus planosus Rocky Mountain PerF LocEn Pres Unc Weber & 1962 Type locality (Umbelliferae) spring-parsley Miller 11388 is Minturn, (BLCA) Eagle Co., CO (Osterhout 2572) Apiaceae Cymopterus C. purpureus var. Colorado Plateau PerF RegEn Pres Unc Hogan & 2003 (Umbelliferae) purpureus purpureus spring-parsley Clark 4056 (BLCA) Apiaceae Ligusticum porteri southern PerF Wide Pres Unc Hogan 4139 1959 (Umbelliferae) ligusticum (BLCA) Apiaceae Lomatium adobe desert- PerF LocEn Pres Rare Hogan 4231 2004 Type locality is (Umbelliferae) concinnum parsley (BLCA) Paonia, Delta Co., CO (Osterhout 4515 RM). CO BLM Sensitive Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Apiaceae Lomatium dissectum Includes vars. fern-leaved PerF Wide Pres Unc Lederer & 1962 BLCA material is (Umbelliferae) eatonii & desert-parsley Clark 03-41 var. multiﬁ dum multiﬁ dum (BLCA) Apiaceae Lomatium grayi L. grayi var. grayi milfoil lomatium PerF Wide Pres Unc Penland 1805 1942 (Umbelliferae) (COLO) Apiaceae Lomatium L. simplex var. ternate desert- PerF Wide Pres Unc Lederer & 1961 (Umbelliferae) triternatum var. simplex parsley Clark 03-03 platycarpum (COLO) Apiaceae Osmorhiza chilensis O. berteroi Chilean sweet- PerF Wide Rep Rare NCPN (in ed.) 2002 (Umbelliferae) cicely Apiaceae Osmorhiza blunt sweet-cicely PerF Wide Pres Com Weber, 1983 (Umbelliferae) depauperata Rector, & Kennedy 16830 (BLCA) Apiaceae Osmorhiza western sweet- PerF Wide Pres Unc Weber, 1983 (Umbelliferae) occidentalis cicely Rector, & Kennedy 16828 (BLCA) Apiaceae Pseudocymopterus Cymopterus Lemmon’s spring- PerF Wide Pres Unc Weber, 1983 (Umbelliferae) montanus lemmonii parsley Rector, & Kennedy 16815 (BLCA) Apocynaceae Apocynum Includes vars. common PerF Wide Pres Unc Hogan 4104 1963 cannabinum cannabinum, dogbane (BLCA) glaberrimum, & angustifolium Asclepiadaceae Asclepias cryptoceras A. cryptoceras var. pallid milkweed PerF Wide Pres Unc Lederer, 2004 cryptoceras Clark, Hogan, & Chapter 6:Appendices Tembrock 04- 51 (BLCA) Asclepiadaceae Asclepias speciosa showy milkweed PerF Wide Pres Unc Weber & 1983 Beck 17004 (BLCA) Asclepiadaceae Asclepias A. galioides whorled PerF Wide Pres Unc Clark 2110 1983 subverticillata milkweed (BLCA) 21 22

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Asparagaceae Asparagus offi cinalis asparagus PerF Intro Pres Unc Hogan 4168 2003 Native to Eurasia (Liliaceae) (BLCA) Aspidiaceae Dryopteris fi lix-mas male fern fern Wide Pres Unc Hogan 4162 1983 (Polypodiaceae, (BLCA) Aspleniaceae) Aspleniaceae Cystopteris fragilis Some authors brittle bladder fern Wide Pres Unc Weber & 1962 (Polypodiaceae) include C. fern Miller 11378 utahensis, C. (BLCA) reevesiana, & C. tenuis Aspleniaceae Cystopteris Included in southwestern fern Wide Pres Unc Montgomery 1983 (Polypodiaceae) reevesiana Cystopteris fragilis bladder fern 86-282 by some authors (COLO) Asteraceae Achillea lanulosa Achillea common yarrow PerF Wide Pres Com Wetlaufer 1959 (Compositae) millefolium var. s.n. (BLCA) lanulosa, vars. not recognized in Flora of North America (2006) Asteraceae Acosta maculosa Centaurea stoebe spotted PerF Intro Pres Unc Hogan & 2004 CO state noxious (Compositae) ssp. micranthos, knapweed Lederer 4388 weed (List B). C. maculosa, C. (BLCA) Native to Europe. biebersteinii Asteraceae Acroptilon repens Centaurea repens Russian PerF Intro Pres Unc Hogan & 2004 CO state noxious (Compositae) knapweed Tembrock weed (List B). 4287 (BLCA) Native to Eurasia. Asteraceae Agoseris aurantiaca orange agoseris PerF Periph Pres Unc Weber, 1983 (Compositae) var. aurantiaca Rector, & Kennedy 16825 (BLCA) Asteraceae Agoseris glauca var. short agoseris PerF Periph Pres Com Weber, 1983 (Compositae) dasycephala Rector, & Kennedy 16824 (BLCA) Asteraceae Agoseris glauca var. (Compositae) A. parvifl ora cutleaf agoseris PerF Wide Pres Com Weber, 1983 laciniata Rector, & Kennedy 16745 (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Asteraceae Almutaster Aster pauciﬂ orus alkali aster PerF Wide Rep Unc Hogan et al. 2002 (Compositae) pauciﬂ orus (2005), NCPN (in ed.) Asteraceae Antennaria low pussytoes PerF Wide Pres Com Weber & 1962 (Compositae) dimorpha Miller 11318 (BLCA) Asteraceae Antennaria parvifolia common PerF Wide Pres Com Lederer & 1962 (Compositae) pussytoes Clark 03-12 (BLCA) Asteraceae Antennaria A. anaphaloides tall pussytoes PerF Wide Pres Unc Beck 17054 1983 (Compositae) pulcherrima ssp. (COLO) anaphaloides Asteraceae Antennaria rosea Included in A. rosy pussytoes PerF Wide Pres Unc Beidleman 1961 (Compositae) microphylla by s.n. (BLCA) some authors Asteraceae Arctium minus burdock PerF Intro Pres Unc Weber & 1983 CO state noxious (Compositae) Beck 16940 weed (List C). (BLCA) Native to Eurasia. Asteraceae Arnica cordifolia A. cordifolia var. heartleaf arnica PerF Wide Pres Unc Hogan & 1962 (Compositae) cordifolia; vars. not Tembrock recognized in Flora 4274 (BLCA) of North America (2006). Asteraceae Artemisia carruthii Carruth’s PerF Wide Pres Unc Clark & 2003 (Compositae) wormwood Hogan 1999 (BLCA) Asteraceae Artemisia frigida fringed sagebrush Shrub Wide Pres Com Weber & 1962 (Compositae) Miller 11381 (BLCA) Chapter 6:Appendices Asteraceae Artemisia ludoviciana Louisiana PerF Wide Pres Com Hogan 4107 1983 (Compositae) ssp. incompta wormwood (BLCA) Asteraceae Artemisia ludoviciana Louisiana PerF Wide Pres Com Weber 16916 (Compositae) ssp. ludoviciana wormwood 1983 (COLO) 23 24

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Asteraceae Aster foliaceus A. foliaceus Canby’s aster PerF Wide Rep Unc NCPN (in ed.) 2002 (Compositae) var. canbyi, Symphyotrichum foliaceum var. canbyi Asteraceae Aster laevis var. A. laevis, smooth aster PerF Wide Rep Unc NCPN (in ed.) 2002 (Compositae) geyeri Symphyotrichum laeve var. geyeri Asteraceae Balsamorhiza arrowleaf PerF Wide Pres Com Lederer & 1942 (Compositae) sagittata balsamroot Clark 03-15 (BLCA) Asteraceae Breea arvensis Cirsium arvense; Canada thistle PerF Intro Pres Unc Hogan 4101 1983 CO state noxious (Compositae) includes vars. (BLCA) weed (List B). horridum & Native to Eurasia. integrifolium Asteraceae Breea incana Included in Cirsium Canada thistle PerF Intro Pres ? Weber 16890 1983 CO state noxious (Compositae) arvense in Flora (COLO) weed (List B). of North America Native to Eurasia. (2006) Duplicate specimen at BLCA is determined as B. arvensis. Asteraceae Brickellia californica California Shrub Wide Pres Unc Hogan 4172 1983 (Compositae) brickellbush (BLCA) Asteraceae Brickellia grandifl ora tassel-fl ower PerF Wide Pres Unc Beck & 1983 (Compositae) brickellbush Kennedy 17029 (BLCA) Asteraceae Brickellia longifolia B. longifolia var. longleaf Shrub Periph Pres Unc Lederer & 2004 (Compositae) longifolia brickellbush Clark 04-27 (BLCA) Asteraceae Brickellia B. oblongifolia var. Mohave Shrub Wide Pres Unc Hogan & 2004 (Compositae) oblongifolia linifolia brickellbush Tembrock 4383 (BLCA) Asteraceae Carduus nutans (Compositae) var. Includes ssp. musk thistle PerF Intro Pres Unc Weber & 1983 CO state noxious macrolepis leiophyllus Beck 16950 weed (List B). (BLCA) Native to Eurasia. Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Asteraceae Chaenactis douglasii Includes vars. hoary dusty- PerF Wide Pres Unc Hogan & 1962 (Compositae) douglasii & maiden Clark 4059 montana (BLCA) Asteraceae Chrysothamnus dwarf Shrub Wide Pres Com Hogan & 2002 (Compositae) depressus rabbitbrush Lederer 4387 (BLCA) Asteraceae Chrysothamnus Ericameria glabrate Shrub Wide Pres Com Weber & 2003 (Compositae) nauseosus ssp. nauseosa var. rabbitbrush Beck 16989 graveolens graveolens, E. (BLCA) nauseosa var. glabrata Asteraceae Chrysothamnus Ericameria parryi Parry’s Shrub Wide Pres Com Weber & 1983 (Compositae) parryi ssp. parryi var. parryi rabbitbrush Beck 16930 (BLCA) Asteraceae Chrysothamnus green rabbitbrush Shrub Wide Pres Com Weber & 1983 (Compositae) viscidiﬂ orus ssp. Beck 16908 viscidiﬂ orus (BLCA) Asteraceae Cirsium centaureae C. clavatum var. fringed thistle PerF RegEn Pres Unc Weber & 1983 (Compositae) americanum, C. Beck 16891 remotifolium (BLCA) ssp. oregonense; included in C. callilepis by some authors Asteraceae Cirsium eatonii C. tweedyi, C. Eaton’s thistle PerF RegEn Pres Unc Weber & 1983 (Compositae) polyphyllum, Beck 16945 includes vars. (BLCA) eatonii & harrisonii Asteraceae Cirsium C. neomexicanum

Chapter 6:Appendices (Compositae) neomexicanum var. New Mexico PerF Wide Pres Unc Hogan 4154 2003 neomexicanum, thistle (BLCA) vars. not recognized in Flora

of North America (2006) 25 26

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Asteraceae Cirsium perplexans Rocky Mountain PerF LocEn Pres Unc Lederer, 2004 Type locality (Compositae) thistle Clark, is Cimarron, Hogan, & Montrose Co., Tembrock 04- CO (Baker 62 (BLCA) 286 NY). CO BLM Sensitive, USFS Region 2 Sensitive Asteraceae Cirsium tracyi C. undulatum var. Tracy’s thistle PerF RegEn Pres Unc Weber, 1983 Type locality (Compositae) tracyi Rector, & is Mancos, CO Kennedy (Baker, Earle, & 16852 (BLCA) Tracy 90) Asteraceae Cirsium vulgare bull thistle PerF Intro Pres Unc Weber, 1983 CO state noxious (Compositae) Rector, & weed (List B). Kennedy Native to Europe. 16841 (BLCA) Asteraceae Conyza canadensis C. canadense var. Canadian AnnF Wide Pres Unc Hogan 4171 1983 (Compositae) glabrata, vars. not horseweed (BLCA) recognized in Flora of North America (2006) Asteraceae Cyclachaena Iva xanthifolia marsh-elder AnnF Wide Pres Unc Hogan & 1983 (Compositae) xanthifolia Lederer 4399 (BLCA) Asteraceae Erigeron concinnus E. pumilus var. Navajo fl eabane PerF Wide Pres Com Weber, 1983 (Compositae) concinnus, E. Rector, & concinnus var. Kennedy concinnus 16856 (BLCA) Asteraceae Erigeron divergens E. divergens var. spreading daisy PerF Wide Pres Unc Hogan & 1961 (Compositae) divergens Tembrock 4354 (BLCA) Asteraceae Erigeron eatonii E. eatonii var. Eaton’s daisy PerF Wide Pres Com Penland 1802 1942 (Compositae) eatonii (COLO) Asteraceae Erigeron Engelmann’s PerF Wide Rep ? NCPN (in ed.) 2002 (Compositae) engelmannii daisy Asteraceae Erigeron fl agellaris trailing daisy PerF Wide Pres Unc Hogan 4124 1959 (Compositae) (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Asteraceae Erigeron peregrinus E. peregrinus wandering PerF Wide Rep Rare NCPN (in ed.) 2002 (Compositae) var. scaposus, fl eabane E. glacialis var. glacialis Asteraceae Erigeron speciosus E. speciosus var. aspen daisy PerF Wide Pres Com Hogan 4133 1959 (Compositae) macranthus (BLCA) Asteraceae Euthamia Solidago western PerF Wide Pres Unc Hogan 4169 2003 (Compositae) occidentalis occidentalis goldenrod (BLCA) Asteraceae Gnaphalium G. exilifolium slender cudweed AnnF Wide Pres Unc Weber & 1983 (Compositae) uliginosum Beck 16929 (BLCA) Asteraceae Grindelia squarrosa Vars. not curly-cup PerF Wide Pres Unc Clark 2107 2003 May be (Compositae) var. squarrosa recognized in Flora gumweed (BLCA) introduced from of North America C USA (2006) Asteraceae Gutierrezia thread Shrub Periph Rep Rare NCPN (in ed.) 2002 Report may be (Compositae) microcephala snakeweed based on G. sarothrae Asteraceae Gutierrezia sarothrae broom Shrub Wide Pres Unc Weber & 1983 (Compositae) snakeweed Beck 16931 (BLCA) Asteraceae Helianthella fi ve-nerve PerF Wide Pres Unc Hogan 4141 1983 (Compositae) quinquenervis sunfl ower (BLCA) Asteraceae Helianthus annuus H. annuus ssp. common AnnF Wide Pres ? Hogan & (Compositae) lenticularis. Ssp. sunfl ower 2004 not recognized Tembrock in Flora of North 4288 (BLCA) America (2006).

Chapter 6:Appendices Asteraceae Heliomeris multifl ora Viguiera multifl ora showy goldeneye PerF Wide Pres Com Weber & 1961 (Compositae) var. multifl ora var. multifl ora Beck 16918 (BLCA) Asteraceae Heterotheca villosa Chrysopsis villosa hairy golden- PerF Wide Pres Com Hogan 4096 1959 (Compositae) var. villosa aster (BLCA) 27 28

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Asteraceae Hymenopappus Wyoming PerF RegEn Pres Unc Lederer, 2004 (Compositae) fi lifolius var. luteus hyalineherb Clark, Hogan, & Tembrock 04- 60 (BLCA) Asteraceae Lactuca serriola prickly lettuce AnnF Intro Pres Unc Hogan & 2004 Native to Europe (Compositae) Lederer 4393 (COLO) Asteraceae Leucantha cyanus Centaurea cyanus bachelor’s button AnnF Intro Rep Rare NCPN (in ed.) 2002 Native to Europe (Compositae) Asteraceae Leucelene ericoides Aster arenosus, rose-heath PerF Wide Pres Unc Hogan & 2004 (Compositae) Chaetopappa Tembrock ericoides 4379 (BLCA) Asteraceae Ligularia pudica Senecio pudicus bashful groundsel PerF RegEn Pres Unc Beck 17045 1983 (Compositae) (COLO) Asteraceae Machaeranthera M. canescens var. hoary aster PerF Wide Pres Com Wetlaufer 1959 (Compositae) canescens canescens, Aster s.n. (BLCA) canescens, Dieteria canescens var. canescens Asteraceae Machaeranthera Haplopappus slender AnnF Periph Rep Unc NCPN (in ed.) 2002 (Compositae) gracilis gracilis, Xanthisma goldenweed gracile Asteraceae Machaeranthera M. grindelioides gumweed aster PerF Wide Rep Unc NCPN (in ed.) 2002 (Compositae) grindelioides var. grindelioides, Haplopappus nuttallii, Xanthisma grindelioides var. grindelioides Asteraceae Machaeranthera M. pinnatifi da lacy tansy-aster PerF Wide Pres Unc Hogan & 2004 (Compositae) pinnatifi da var. pinnatifi da, Tembrock Haplopappus 4372 (BLCA) spinulosus var. spinulosus Asteraceae Madia glomerata mountain AnnF Wide Pres Unc Weber & 1983 (Compositae) tarweeed Beck 16928 (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Asteraceae Microseris nutans Ptilocalais nutans nodding PerF Wide Pres Unc Weber, 1983 (Compositae) microseris Rector, & Kennedy 16768 (BLCA) Asteraceae Oligosporus O. dracunculus, tarragon PerF Wide Pres Unc Hogan & 1983 (Compositae) dracunculus ssp. Artemisia Lederer 4386 glaucus dracunculus, vars. (BLCA) not recognized Flora of North America (2006) Asteraceae Packera multilobata Senecio basin groundsel PerF Wide Pres Com Weber 11353 1962 (Compositae) multilobatus (COLO) Asteraceae Packera neomexicana Senecio New Mexico PerF Wide Pres Com Weber & 1961 (Compositae) neomexicanus groundsel Miller 11366 var. mutabilis, P. (BLCA) neomexicana var. mutabilis Asteraceae Petradoria pumila P. pumila var. rock-goldenrod PerF Wide Pres Com Hogan 4151 1959 (Compositae) pumila, Solidago (BLCA) petradoria Asteraceae Psilochenia Crepis acuminata mountain PerF Wide Pres Unc Wetlaufer 1959 (Compositae) acuminata hawksbeard s.n. (BLCA) Asteraceae Psilochenia atribarba Crepis atribarba slender PerF Wide Pres Unc Hogan & 1983 (Compositae) hawksbeard Tembrock 4281 (BLCA) Asteraceae Psilochenia Crepis modocensis Modoc PerF Wide Pres Unc Weber, 1983 (Compositae) modocensis hawksbeard Rector, & Kennedy 16795 (BLCA)

Chapter 6:Appendices Asteraceae Psilochenia Crepis occidentalis western PerF Wide Pres Unc Lederer, 2004 (Compositae) occidentalis var. occidentalis hawksbeard Clark, Hogan, & Tembrock 04- 52 (BLCA) Asteraceae Rudbeckia ampla R. laciniata var. cutleaf (Compositae) ampla coneﬂ PerF Wide Pres Unc Hyde s.n. 1961 ower (BLCA) 29 30

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Asteraceae Senecio integerrimus S. integerrimus var. western PerF Wide Pres Com Lederer & 1960 (Compositae) exaltatus groundsel Clark 03-06 (BLCA) Asteraceae Senecio wootonii Wooton’s ragwort PerF Wide Hist ? Penland 1799 1942 (Compositae) (COLO) Asteraceae Seriphidium novum Artemisia nova, A. black sagebrush Shrub Wide Rep Unc NCPN (in ed.) 2002 (Compositae) nova var. nova, A. arbuscula var. nova Asteraceae Seriphidium Artemisia Wyoming big Shrub Wide Rep Unc NCPN (in ed.) 2002 (Compositae) tridentatum ssp. tridentata var. sagebrush wyomingense wyomingensis Asteraceae Seriphidium Artemisia mountain big Shrub Wide Pres Abun Hogan 4178 1961 (Compositae) vaseyanum tridentata var. sagebrush (BLCA) vaseyana, includes A. tridentata var. paucifl ora Asteraceae Solidago velutina S. sparsifl ora, three-nerve PerF Wide Pres Com Weber & 1983 (Compositae) S. velutina var. goldenrod Beck 16979 sparsifl ora (BLCA) Asteraceae Stenotus acaulis Haplopappus stemless PerF Wide Rep Unc NCPN (in ed.) 2002 Report may (Compositae) acaulis var. acaulis, goldenweed be based on S. S. acaulis var. armerioides acaulis Asteraceae Stenotus armerioides Haplopappus (Compositae) armerioides var. thrift PerF Wide Pres Unc Weber & 1962 armerioides, S. goldenweed Miller 11324 armerioides var. (BLCA)

armerioides Asteraceae Stephanomeria brown-plume PerF Wide Pres Unc Weber & 1983 (Compositae) paucifl ora wire-lettuce Beck 16996 (BLCA) Asteraceae Taraxacum offi cinale common PerF Intro Pres Unc Weber, 1983 Native to Eurasia (Compositae) dandelion Rector, & Kennedy 16764 (BLCA) Asteraceae Tetradymia canescens spineless Shrub Wide Pres Unc Clark 2101 2002 (Compositae) horsebrush (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Asteraceae Tetradymia spinosa T. spinosa var. shortspine Shrub Wide Pres Unc Hogan 4226 2002 (Compositae) spinosa horsebrush (BLCA) Asteraceae Tetraneuris acaulis Hymenoxys acaulis stemless PerF Wide Rep ? NCPN (in ed.) 2002 (Compositae) var. acaulis, T. hymenoxys acaulis var. acaulis Asteraceae Tetraneuris ivesiana Hymenoxys Canyonlands PerF Wide Pres Unc Clark & 1959 (Compositae) argentea var. hymenoxys Hogan 1992 ivesiana, H. acaulis (BLCA) var. ivesiana Asteraceae Tetraneuris torreyana Hymenoxys Torrey’s PerF RegEn Pres Unc Lederer & 2003 (Compositae) torreyana hymenoxys Clark 03-24 (BLCA) Asteraceae Townsendia incana hoary townsendia PerF Wide Pres Unc Lederer, 2002 (Compositae) Clark, Hogan, & Tembrock 04- 58 (BLCA) Asteraceae Tragopogon dubius T. dubius yellow salsify PerF Intro Pres Unc Clark & 1983 Native to Europe (Compositae) var. major Hogan 1996 (BLCA) Asteraceae Virgulaster Aster ascendens, Paciﬁ c aster PerF Wide Pres Unc Hogan & 1983 (Compositae) ascendens A. chilensis ssp. Lederer 4385 adscendens, (BLCA) Symphyotrichum ascendens Asteraceae Wyethia mule’s-ears PerF Wide Pres Unc Clark & 2002 (Compositae) amplexicaulis Lederer 2187a (BLCA) Asteraceae Wyethia arizonica Arizona PerF RegEn Pres Unc Hogan 4156 2003 Chapter 6:Appendices (Compositae) mulesears (BLCA) Asteraceae Wyethia x magna Hybrid of W. hybrid wyethia PerF RegEn Pres Com Wetlaufer 1959 (Compositae) amplexicaulus x W. s.n. (BLCA) arizonica

Asteraceae Xylorhiza venusta Machaeranthera Cisco woodyaster PerF RegEn Pres Unc Hogan 4205 2002 Type locality is (Compositae) venusta (BLCA) Cisco, Grand Co., UT (Jones s.n. POM) 31 32

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Berberidaceae Mahonia fremontii Berberis fremontii Fremont’s Shrub Wide Rep Unc NCPN (in ed.) 2002 mahonia Berberidaceae Mahonia repens Berberis repens Oregon-grape Shrub Wide Pres Unc Hogan & 1961 Lederer 4398 (BLCA) Betulaceae Alnus incana ssp. A. tenuifolia, thinleaf alder Shrub Wide Pres Com Weber & 1983 tenuifolia A. incana var. Beck 16997 occidentalis (BLCA) Boraginaceae Cryptantha fendleri sand dune AnnF Wide Pres Unc Weber & 1983 cryptanth Beck 16985 (BLCA) Boraginaceae Cryptantha gracilis slender cryptanth AnnF Wide Pres Unc Weber & 1962 Miller 11351 (BLCA) Boraginaceae Cryptantha C. pterocarya var. wing-nut AnnF Wide Pres Unc Weber & 1962 pterocarya pterocarya cryptanth Miller 11356 (BLCA) Boraginaceae Cryptantha watsonii Watson’s AnnF Wide Pres Unc Hogan & 2003 cryptanth Clark 4057 (BLCA) Boraginaceae Cynoglossum common hound’s- PerF Intro Pres Unc Weber, 1983 CO state noxious offi cinale tongue Rector, & weed (List B). Kennedy Native to Eurasia. 16779 (BLCA) Boraginaceae Hackelia fl oribunda showy stickseed PerF Wide Pres Unc Beck & 1983 Kennedy 17053 (BLCA) Boraginaceae Lappula marginata L. texana, L. cupseed stickseed AnnF Wide Pres Unc Hogan & 2004 redowskii var. Tembrock cupulata, L. 4374 (BLCA) occidentalis var. cupulata Boraginaceae Lappula redowskii L. redowskii var. western stickseed AnnF Wide Pres Com Weber 11314 1962 redowskii, L. (COLO) occidentalis var. occidentalis Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Boraginaceae Lappula squarrosa L. echinata, L. European AnnF Intro Rep Rare NCPN (in ed.) 2002 Native to Eurasia. squarrosa var. stickseed Native race (L. squarrosa erecta) from the Rocky Mtns. is probably extralimital Boraginaceae Lithospermum western PerF Wide Pres Unc Weber & 1962 ruderale stoneseed Miller 11313 (BLCA) Boraginaceae Mertensia arizonica M. arizonica aspen bluebell PerF Periph Pres ? Cottrell & 1974 var. grahamii Gregory 255 (BLCA) Boraginaceae Mertensia brevistyla Weber & Wittmann short-styled PerF Wide Pres Com Hogan 3992 1960 (2001) include M. bluebell (BLCA) fusiformis Boraginaceae Mertensia ciliata streamside PerF Wide Pres Unc Beck & 1959 bluebell Kennedy 17046 (BLCA) Boraginaceae Oreocarya Cryptantha yellow-eye PerF Wide Pres Com Weber & 1962 fl avoculata fl avoculata cryptanth Miller 11328 (BLCA) Boraginaceae Plagiobothrys P. scouleri var. meadow AnnF Wide Pres Unc Clark & 2003 scouleri ssp. hispidulus popcorn-fl ower Hogan 2014 penicillatus (BLCA) Brassicaceae Alyssum alyssoides pale madwort AnnF Intro Rep Rare NCPN (in ed.) 2002 Native to Europe (Cruciferae) Brassicaceae Alyssum desertorum desert madwort AnnF Intro Pres Com Lederer & 2003 Native to Europe (Cruciferae) Clark 03-47 (BLCA) Chapter 6:Appendices Brassicaceae Alyssum parvifl orum A. parvifl orum var. European AnnF Intro Pres Com Weber, 1983 Native to Europe (Cruciferae) micranthum, A. madwort Rector, & simplex, A. minus Kennedy var. micranthum 16742 (BLCA) 33 34

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Brassicaceae Arabis hirsuta Includes vars. hairy rockcress PerF Wide Rep Rare NCPN (in ed.) 2002 Earlier reports (Cruciferae) glabrata & were based on pycnocarpa misidentifi ed specimens of Turritis glabra Brassicaceae Barbarea vulgaris European PerF Intro Pres Unc Hogan 3963 2003 Native to Eurasia (Cruciferae) wintercress (BLCA) Brassicaceae Boechera crandallii Arabis crandallii Crandall’s PerF RegEn Rep Com NCPN (in ed.) 2002 Type locality (Cruciferae) rockcress is Cimarron, Montrose Co., CO (Crandall 6 GH). CO BLM Sensitive Brassicaceae Boechera divaricarpa Arabis confi nis, spreadingpod PerF Wide Pres Com Hogan 3971 1983 Often considered (Cruciferae) A. holboellii var. rockcress (BLCA) to represent a brachycarpa, A. hybrid swarm x divaricarpa, B. involving Arabis brachycarpa drummondii with A. holboellii, A. sparsifl ora, A. pendulina, & A. perennans. Brassicaceae Boechera B. stricta, B. Drummond’s PerF Wide Pres Unc Weber, 1983 (Cruciferae) drummondii angustifolia, Arabis rockcress Rector, & drummondii Kennedy 16748 (BLCA) Brassicaceae Boechera fendleri Arabis fendleri, Fendler’s PerF Wide Rep Rare NCPN (in ed.) 2002 (Cruciferae) A. holboellii var. rockcress fendleri, B. fendleri var. spatifolia, vars. not recognized in Holmgren et al. (2005) Brassicaceae Boechera Arabis Gunnison’s PerF LocEn Pres Unc Weber 1962 Type locality is (Cruciferae) gunnisoniana gunnisoniana rockcress 11340a Saguache Creek, (COLO) Gunnison Co., CO (Rollins 2090 GH) Brassicaceae Boechera lignifera Arabis lignifera Sweetwater PerF Wide Pres Unc Penland 1807 1942 (Cruciferae) rockcress (COLO) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Brassicaceae Boechera oxylobula Arabis oxylobula Glenwood PerF LocEn Pres Unc Weber 11340 1962 Type locality (Cruciferae) Springs rockcress (COLO) is Glenwood Springs, Garﬁ eld Co., CO (Osterhout s.n.) Brassicaceae Boechera pallidifolia Arabis pallidifolia Gunnison County PerF LocEn Pres Unc Weber 16785 1962 Type locality (Cruciferae) rockcress (COLO) is hill above Sapinero, Gunnison Co., CO (Rollins & Rollins 8376) Brassicaceae Boechera perennans Arabis perennans perennial PerF Wide Pres Unc Hogan 4233 2004 (Cruciferae) var. perennans rockcress (BLCA) Brassicaceae Boechera retrofracta B. holboellii var. Holboell’s PerF Wide Pres Unc Lederer & 1983 (Cruciferae) secunda, Arabis rockcress Clark 03-35 holboellii var. (BLCA) secunda, A. holboellii var. retrofracta Brassicaceae Boechera selbyi B. selbyi var. selbyi, Selby’s rockcress PerF Wide Pres Unc Weber 11333 1962 (Cruciferae) Arabis selbyi (COLO) Brassicaceae Camelina microcarpa little-pod false AnnF Intro Pres Unc Clark & 2003 Native to Asia (Cruciferae) ﬂ ax Hogan 2000 (BLCA) Brassicaceae Capsella bursa- shepherd’s-purse AnnF Intro Pres Unc Hogan 4236 1983 Native to Europe (Cruciferae) pastoris (BLCA) Brassicaceae Cardaria chalepensis C. draba ssp. orbicular PerF Intro Pres Unc Clark & 2003 Native to Europe (Cruciferae) chalepensis, C. whitetop Hogan 2009 draba var. repens (BLCA)

Chapter 6:Appendices Brassicaceae Chorispora tenella blue mustard AnnF Intro Pres Unc Clark & 2003 Native to Asia (Cruciferae) Hogan 2008 (BLCA) Brassicaceae Descurainia pinnata D. pinnata var. (Cruciferae) ssp. ﬁ lipes ﬁ western tansy- AnnF Wide Pres Unc Weber & 1962 lipes mustard Miller 11354 (BLCA) 35 36

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Brassicaceae Descurainia pinnata D. richardsonii var. mountain tansy- AnnF Wide Pres Unc Weber, 1983 (Cruciferae) ssp. incisa sonnei, D. incana mustard Rector, & var. incana, D. Kennedy incana var. sonnei 16786 (BLCA) Brassicaceae Descurainia sophia fl ixweed AnnF Intro Pres Unc Hogan 4142 1983 Native to Europe (Cruciferae) (BLCA) Brassicaceae Draba cuneifolia D. cuneifolia var. wedgeleaf draba AnnF Wide Rep Unc NCPN (in ed.) 2002 (Cruciferae) cuneifolia Brassicaceae Draba nemorosa woodland AnnF Intro Pres Unc Hogan 3969 2003 Native to Eurasia (Cruciferae) whitlow-grass (BLCA) Brassicaceae Draba reptans dwarf draba AnnF Wide Pres Unc Lederer & 2003 (Cruciferae) Clark 03-17 (BLCA) Brassicaceae Erysimum capitatum E. capitatum western PerF Wide Pres Unc Weber & 1962 (Cruciferae) var. capitatum, wallfl ower Miller 11392 E. asperum var. (BLCA) arkansanum, E. asperum var. elatum Brassicaceae Erysimum E. cheiranthoides worm-seed PerF Wide Rep Rare NCPN (in ed.) 2002 Sometimes (Cruciferae) cheiranthoides ssp. wallfl ower considered altum adventive from Europe (at least in part). Known from CURE. Brassicaceae Erysimum shy wallfl ower PerF Wide Rep Rare NCPN (in ed.) 2002 (Cruciferae) inconspicuum Brassicaceae Erysimum repandum spreading AnnF Intro Pres Unc Lederer, 2004 Native to Europe (Cruciferae) wallfl ower Clark, Hogan, & Tembrock 04- 44 (BLCA) Brassicaceae Hesperis matronalis dame’s rocket PerF Intro Pres Unk Danguole ? Native to Europe (Cruciferae) Bockus, pers. Comm.. Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Brassicaceae Lepidium L. densifl orum var. prairie AnnF Wide Pres Unc Weber, 1962 (Cruciferae) densifl orum densifl orum pepperwort Rector, & Kennedy 16758 (BLCA) Brassicaceae Lepidium montanum Jones’ PerF Wide Pres Unc Hogan 4223 2004 (Cruciferae) var. jonesii pepperwort (BLCA) Brassicaceae Lepidium montanum mountain PerF Wide Rep Rare NCPN (in ed.) 2002 (Cruciferae) var. montanum pepperwort Brassicaceae Lepidium montanum mountain PerF LocEn Pres Unc Lederer & 1917 Type locality is (Cruciferae) var. tenellum pepperweed Clark 04-26 Black Canyon, (BLCA) Montrose Co., CO (Payson 1033 RM) Brassicaceae Lepidium clasping AnnF Intro Pres Unc Weber & 1962 Native to Europe (Cruciferae) perfoliatum pepperwort Miller 11349 (BLCA) Brassicaceae Lepidium L. bourgeauanum branched AnnF Periph Pres Unc Weber & 1983 (Cruciferae) ramosissimum var. pepperwort Beck 16910 bourgeauanum (BLCA) Brassicaceae Lesquerella rectipes Physaria rectipes Colorado Plateau PerF RegEn Rep Unc NCPN (in ed.) 2002 (Cruciferae) bladderpod Brassicaceae Lesquerella vicina Uncompaghre PerF LocEn Pres Rare Lederer, 2004 Type locality is (Cruciferae) bladderpod Clark, mesa overlooking Hogan, & Uncompahgre Tembrock 04- River, Montrose 56 (BLCA) Co., CO (Reveal 7492 GH). CO BLM Sensitive Brassicaceae Noccaea montana N. montana var. alpine pennycress PerF Wide Pres Unc Weber & 1962 (Cruciferae) montana, Thlaspi Miller 11391 Chapter 6:Appendices montanum var. (BLCA) montanum Brassicaceae Physaria acutifolia P. acutifolia var. sharp-leaf (Cruciferae) acutifolia, P. PerF Wide Pres Unc Weber & 1983 australis twinpod Beck 17062 (BLCA) 37 38

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Brassicaceae Physaria fl oribunda mesa twinpod PerF LocEn Pres Unc Hogan & 2003 Type locality is (Cruciferae) Clark 4051 Sangre de Cristo (BLCA) Creek, Costilla Co., CO (Rydberg & Vreeland 6135 NY) Brassicaceae Physaria rollinsii Rollins’ twinpod PerF LocEn Rep Unc NCPN (in ed.) 2002 Type locality (Cruciferae) is 4 miles NW of Doyleville, Gunnison Co., CO (Mulligan & Crampton 2880) Brassicaceae Schoenocrambe Sisymbrium fl ax-leaved PerF Wide Pres Com Lederer & 1961 (Cruciferae) linifolia linifolium plainsmustard Clark 03-44 (BLCA) Brassicaceae Sisymbrium tumble mustard AnnF Intro Pres Unc Hogan 4097 1961 Native to Europe (Cruciferae) altissimum (BLCA) Brassicaceae Stanleya albescens white prince’s- PerF RegEn Pres Unc Hogan 4207 2002 (Cruciferae) plume (BLCA) Brassicaceae Stanleya pinnata var. S. pinnata prince’s-plume PerF Wide Pres Unc Hogan 4221 2004 (Cruciferae) pinnata (BLCA) Brassicaceae Streptanthus S. cordatus var. heart-leaf PerF Wide Pres Unc Lederer & 2002 (Cruciferae) cordatus cordatus jewelfl ower Clark 03-40 (BLCA) Brassicaceae Thelypodiopsis elegant PerF RegEn Pres Unc Hogan 4220 2004 Type locality (Cruciferae) elegans thelypody (COLO) is Westwater, Grand Co., UT (Jones s.n. POM) Brassicaceae Thelypodiopsis Sisymbrium juniper AnnF LocEn Pres Unc Hogan & 1961 Type locality is (Cruciferae) juniperorum juniperorum tumblemustard Tembrock western Colorado 4381 (BLCA) (Payson) Brassicaceae Turritis glabra Arabis glabra var. tower-mustard PerF Wide Pres Unc Beck & 1983 (Cruciferae) glabra, vars. not Kennedy recognized in 17034 (BLCA) Holmgren et al. (2005) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Cactaceae Echinocereus E. triglochidiatus claretcup PerF Wide Pres Unc Weber & 1962 triglochidiatus var. Miller 11384 melanacanthus, (BLCA) includes var. inermis. Vars. not recognized in Flora of North America (2003). Cactaceae Opuntia fragilis var. O. fragilis brittle prickly- PerF Wide Pres Unc Clark & 1983 brachyartha pear Lederer 2190 (BLCA) Cactaceae Opuntia O. phaeacantha berry prickly-pear PerF Wide Pres Unc Hogan 4113 2003 phaeacantha var. phaeacantha; (BLCA) vars. not recognized in Flora of North America (2003) Cactaceae Opuntia polyacantha O. polyacantha var. plains prickly- PerF Wide Pres Com Hogan 4136 2003 polyacantha. Flora pear (BLCA) of North America (2003) includes vars. ruﬁ spina & trichophora in var. polyacantha. Cactaceae Pediocactus Includes var. Simpson’s PerF Wide Pres Unc Weber, 1983 simpsonii var. minor simpsonii hedgehog cactus Rector, & Kennedy 16784 (BLCA) Calochortaceae Calochortus C. gunnisonii var. Gunnison’s PerF Wide Pres Unc Clark & 1959 (Liliaceae) gunnisonii gunnisonii mariposa Hogan 2031 Chapter 6:Appendices (BLCA) Calochortaceae Calochortus nuttallii sego-lily PerF Wide Pres Unc Lederer, 2002 (Liliaceae) Clark, Hogan, & Tembrock 04- 49 (BLCA) Campanulaceae Campanula harebell PerF Wide Pres Unc Wetlaufer 1959 rotundifolia s.n. (BLCA) 39 40

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Capparaceae Cleomella C. palmeriana var. Palmer’s AnnF RegEn Pres Unc Lederer, 2004 (Cleomaceae) palmeriana palmeriana stinkweed Clark, Hogan, & Tembrock 04- 47 (BLCA) Caprifoliaceae Sambucus S. racemosa var. red elderberry Shrub Wide Pres Unc Hogan & 1983 (Adoxaceae) microbotrys var. microbotrys; Tembrock microbotrys included in S. 4300 (BLCA) racemosa var. racemosa by some authors. Caprifoliaceae Symphoricarpos S. oreophilus mountain Shrub Wide Pres Com Weber, 1983 rotundifolius var. utahensis; snowberry Rector, & includes vars. Kennedy utahensis, parishii 16859 (BLCA) & oreophilus Caryophyllaceae Gastrolychnis Silene drummondii Drummond’s PerF Wide Pres Unc Hogan 4137 1983 drummondii ssp. drummondii catchfl y (BLCA) & ssp. striata, includes var. kruckebergii, Lychnis drummondii Celastraceae Paxistima myrsinites Pachistima mountain-lover Shrub Wide Pres Com Cottrell & 1974 myrsinites, Gregory 282 Pachystima (BLCA) myrsinites Chenopodiaceae Atriplex canescens A. canescens var. fourwing saltbush Shrub Wide Pres Unc Hogan & 2002 occidentalis, A. Tembrock canescens var. 4384 (BLCA) canescens Chenopodiaceae Atriplex confertifolia shadscale Shrub Wide Pres Unc Hogan 4213a 2002 (BLCA) Chenopodiaceae Atriplex corrugata mat saltbush Shrub RegEn Pres Com Lederer, 2002 Clark, Hogan, & Tembrock 04- 41 (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Chenopodiaceae Atriplex gardneri A. gardneri var. Gardner’s Shrub Wide Pres Com Lederer, 2002 gardneri, A. saltbush Clark, nuttallii Hogan, & Tembrock 04- 48 (BLCA) Chenopodiaceae Atriplex powellii A. powellii var. Powell’s orach AnnF Wide Pres Unc Hogan & 2004 powellii Tembrock 4289 (BLCA) Chenopodiaceae Chenopodium album C. album var. lambsquarter AnnF Intro Rep Unc NCPN (in ed.) 2002 Native to Eurasia album Chenopodiaceae Chenopodium C. fremontii var. mountain AnnF Wide Pres Unc Weber & 1983 atrovirens atrovirens goosefoot Beck 16884 (BLCA) Chenopodiaceae Chenopodium C. capitatum var. smallhead AnnF Wide Rep Unc NCPN (in ed.) 2002 foliosum parvicapitatum, C. goosefoot overi Chenopodiaceae Chenopodium C. fremontii var. Fremont’s AnnF Wide Pres Unc Clark 2099 1983 fremontii fremontii goosefoot (BLCA) Chenopodiaceae Halogeton halogeton AnnF Intro Pres Unc Lederer, 2004 CO state noxious glomeratus Clark, weed (List C). Hogan, & Native to Eurasia. Tembrock 04- 37 (BLCA) Chenopodiaceae Krascheninnikovia K. lanata var. winterfat Shrub Wide Pres Unc Hogan 4214 2002 lanata lanata, Ceratoides (BLCA) lanata, Eurotia lanata. Vars. not recognized in Flora of North America

Chapter 6:Appendices (2003). Chenopodiaceae Monolepis Nuttall’s poverty- AnnF Wide Pres Unc Weber, 1983 nuttalliana weed Rector, & Kennedy 16759 (BLCA) Chenopodiaceae Salsola australis S. kali, S. iberica, S. Russian thistle AnnF Intro Pres Unc Clark 2108 2003 Native to Asia pestifer, S. tragus (BLCA) 41 42

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Chenopodiaceae Sarcobatus greasewood Shrub Wide Pres Unc Lederer, 2002 (Sarcobataceae) vermiculatus Clark, Hogan, & Tembrock 04- 36 (BLCA) Convallariaceae Maianthemum M. racemosum false Solomon’s- PerF Wide Pres Com Cottrell & 1961 (Liliaceae, amplexicaule var. amplexicaule, seal Gregory 251 Ruscaceae) Smilacina racemosa (BLCA) Convallariaceae Maianthemum Smilacina stellata starry false PerF Wide Pres Unc Weber, 1983 (Liliaceae, stellatum Solomon’s-seal Rector, & Ruscaceae) Kennedy 16831 (BLCA) Convolvulaceae Convolvulus arvensis fi eld bindweed PerF Intro Pres Unc Weber, 1983 CO state noxious Rector, & weed (List C). Kennedy Native to Eurasia. 16820 (BLCA) Cornaceae Swida sericea Cornus sericea red-osier Shrub Wide Pres Unc Weber & 1962 var. sericea, C. dogwood Miller 11377 stolonifera, C. alba (BLCA) var. alba Crassulaceae Amerosedum Sedum lance-leaved PerF Wide Pres Com Clark & 1959 lanceolatum lanceolatum; stonecrop Hogan 2029 includes ssp. (BLCA) lanceolatum & subalpinum Crossosomataceae Forsellesia Glossopetalon spiny greasebush Shrub RegEn Pres Unc Lederer, 2002 (Celastraceae) meionandra spinescens var. Clark, meionandrum Hogan, & Tembrock 04- 45 (BLCA) Cupressaceae Sabina osteosperma Juniperus Utah juniper Tree Wide Pres Abun Cottrell & 1974 osteosperma Gregory 283 (BLCA) Cupressaceae Sabina scopulorum Juniperus Rocky Mountain Tree Wide Pres Com Beck & 1959 scopulorum juniper Kennedy 17019 (BLCA) Cyperaceae Carex egglestonii Eggleston’s sedge PerG RegEn Rep Rare NCPN (in ed.) 2002 Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Cyperaceae Carex geophila White Mountain PerG Wide Pres Unc Weber & 1962 sedge Miller 11329 (BLCA) Cyperaceae Carex geyeri elk sedge PerG Wide Pres Com Weber & 1962 Miller 11312 (BLCA) Cyperaceae Carex lanuginosa C. pellita woolly sedge PerG Wide Pres Unc Lederer & 2004 Clark 04-20 (BLCA) Cyperaceae Carex microptera C. microptera small-wing sedge PerG Wide Pres Unc Weber, 1983 var. microptera, Rector, & C. festivella, C. Kennedy macloviana var. 16855 (BLCA) microptera Cyperaceae Carex occidentalis western sedge PerG Wide Pres Com Clark & 1962 Hogan 2004 (COLO) Cyperaceae Carex petasata Liddon sedge PerG Wide Pres Unc Weber, 1983 Rector, & Kennedy 16809 (BLCA) Cyperaceae Carex praegracilis blackcreeper PerG Wide Pres Unc Weber & 1983 sedge Beck 16913 (BLCA) Cyperaceae Carex rossii C. deﬂ exa var. Ross’ sedge PerG Wide Pres Unc Clark & 2002 boottii, C. brevipes, Lederer 2186 C. pityophila (BLCA) Cyperaceae Carex vallicola valley sedge PerG Wide Pres Unc Weber, 1983 Rector, & Chapter 6:Appendices Kennedy 16754 (BLCA) Cyperaceae Carex xerantica C. tahoensis dryland sedge PerG Wide Pres Unc Hogan 4127 2003 (BLCA) Cyperaceae Eleocharis bolanderi E. montevidensis Bolander’s PerG Wide Pres ? Weber, 1983 spikerush Rector, & Kennedy 16857 (BLCA) 43 44

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Cyperaceae Eleocharis Includes E. common PerG Wide Pres Unc Weber & 1983 macrostachya palustris, E. spikerush Beck 16905 erythropoda, & E. (BLCA) uniglumis Cyperaceae Schoenoplectus Schoenoplectus hardstem bulrush PerG Wide Rep ? Hogan et al. NA lacustris ssp. acutus acutus var. (2005), NPS occidentalis, vegetation Schoenoplectus plots acutus var. acutus, Scirpus acutus Ephedraceae Ephedra cutleri E. viridis var. viscida Shrub RegEn Pres ? Cottrell & 1974 Gregory 269 (BLCA) Ephedraceae Ephedra viridis E. viridis var. viridis green ephedra Shrub Wide Pres Unc Hogan & 2002 Tembrock 4280 (BLCA) Equisetaceae Equisetum arvense fi eld horsetail fern Wide Pres Unc Lederer & 1974 Clark 04-21 (BLCA) Equisetaceae Hippochaete Equisetum hyemale tall scouring-rush fern Wide Pres Unc Cottrell & 1962 hyemalis var. affi nis var. affi ne, E. Gregory 259 praealtum (BLCA) Equisetaceae Hippochaete Equisetum smooth scouring- fern Wide Pres Unc Cottrell & 1974 laevigata laevigatum, E. rush Gregory 260 kansanum (BLCA) Equisetaceae Hippochaete x Equisetum x Ferriss’ horsetail fern Sparse Pres ? Beck 17018 1983 ferrissii ferrissii, hybrid of (COLO) E. hyemale x E. laevigatum Ericaceae Arctostaphylos uva- A. uva-ursi var. bearberry Shrub Wide Rep Rare NCPN (in ed.) 2002 ursi ssp. coactilis uva-ursi, vars. not recognized in Holmgren et al. (2005) Euphorbiaceae Chamaesyce fendleri Euphorbia fendleri Fendler’s spurge PerF Wide Rep Unc NCPN (in ed.) 2002 Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Euphorbiaceae Chamaesyce Euphorbia ridge-seeded AnnF Wide Pres Unc Weber & 1983 glyptosperma glyptosperma spurge Beck 16933 (BLCA) Euphorbiaceae Tithymalus Euphorbia robusta, shorthorn spurge PerF Wide Pres Unc Lederer, 2004 brachyceras E. brachycera, E. Clark, brachycera var. Hogan, & robusta Tembrock 04- 64 (BLCA) Euphorbiaceae Tragia ramosa noseburn PerF Wide Rep Rare NCPN (in ed.) 2002 Fabaceae Astragalus milkweed PerF RegEn Pres Unc Weber & 1962 Type locality is (Leguminosae) asclepiadoides milkvetch Miller 11320 Cisco, Grand (BLCA) Co., UT (Jones s.n. POM). Weber & Miller 11320 (BLCA) is probably from just outside the park. Fabaceae Astragalus A. diversifolius lesser rushy PerF Wide Pres Com Lederer, 2002 (Leguminosae) convallarius var. campestris, A. milkvetch Clark, convallarius var. Hogan, & convallarius Tembrock 04- 55 (BLCA) Fabaceae Astragalus falcatus Russian sickle PerF Intro Rep Unc NCPN (in ed.) 2002 Native to Asia (Leguminosae) Fabaceae Astragalus fl avus A. fl avus var. yellow milkvetch PerF Wide Pres Com Hogan 4212 2004 (Leguminosae) candicans, A. fl avus (BLCA) var. fl avus Fabaceae Astragalus straggling PerF RegEn Pres Unc Clark & 2003 Type locality

Chapter 6:Appendices (Leguminosae) lentiginosus var. milkvetch Lederer 2185 is Montezuma palans (BLCA) Canyon, San Juan Co., UT (Eastwood s.n. POM) 45 46

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Fabaceae Astragalus Includes great bladdery PerF Periph Pres Unc Lederer, 2004 (Leguminosae) megacarpus vars. parryi & milkvetch Clark, megacarpus. Vars. Hogan, & not recognized in Tembrock 04- Barneby (1989) 59 (BLCA) Fabaceae Astragalus miser var. Rydberg’s weedy PerF Wide Pres Com Clark & 1983 (Leguminosae) oblongifolius milkvetch Lederer 2195 (BLCA) Fabaceae Astragalus small-fl owered AnnF RegEn Rep Unc NCPN (in ed.) 2002 (Leguminosae) nuttallianus var. milkvetch micranthemiformis Fabaceae Astragalus oophorus egg milkvetch PerF Wide Pres Com Weber & 1960 (Leguminosae) var. caulescens Miller 11345 (BLCA) Fabaceae Astragalus Rocky Mountain PerF Wide Pres Unc Hogan 4125 2003 (Leguminosae) scopulorum milkvetch (BLCA) Fabaceae Astragalus tenellus pulse milkvetch PerF Wide Rep Unc NCPN (in ed.) 2002 (Leguminosae) Fabaceae Astragalus Fort Wingate PerF RegEn Pres Unc Clark & 2004 (Leguminosae) wingatanus milkvetch Lederer 2194 (BLCA) Fabaceae Glycyrrhiza lepidota G. lepidota var. licorice-root PerF Wide Pres Unc Beck & 1983 (Leguminosae) lepidota Kennedy 17024 (BLCA) Fabaceae Lathyrus leucanthus L. lanszwertii var. whitefl ower PerF RegEn Pres Com Miller & 1961 Type locality is (Leguminosae) leucanthus sweetpea Miller s.n. Ojo, Huerfano (BLCA) Co., CO (Rydberg & Vreeland 6020 NY). Fabaceae Lupinus argenteus L. argenteus var. silvery lupine PerF Wide Rep Unc NCPN (in ed.) 2002 (Leguminosae) argenteus Fabaceae Lupinus caudatus var. (Leguminosae) L. argenteus Utah lupine PerF Wide Pres Com Hogan & 1959 utahensis var. utahensis, Clark 4058 L. caudatus ssp. (BLCA) caudatus Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Fabaceae Lupinus polyphyllus L. prunophilus meadow lupine PerF Wide Pres Unc Weber, 1983 (Leguminosae) var. prunophilus Rector, & Kennedy 16826 (BLCA) Fabaceae Lupinus sericeus L. sericeus var. silky lupine PerF Wide Rep Unc NCPN (in ed.) 2002 (Leguminosae) sericeus, L. bakeri. Vars not recognized in Barneby (1989) Fabaceae Medicago lupulina black medick AnnF Intro Pres Unc Weber & 1983 Native to Europe (Leguminosae) Beck 16943 (BLCA) Fabaceae Medicago sativa M. sativa var. sativa alfalfa PerF Intro Pres Unc Hogan 4098 2003 Native to Europe (Leguminosae) (BLCA) Fabaceae Melilotus albus M. alba; included white sweet- AnnF Intro Pres Unc Weber & 1983 Native to Europe (Leguminosae) in M. offi cinalis by clover Beck 16974 some authors (BLCA) Fabaceae Melilotus offi cinale M. offi cinalis yellow sweet- PerF Intro Pres Unc Weber, 1983 Native to Europe (Leguminosae) clover Rector, & Kennedy 16853 (BLCA) Fabaceae Trifolium T. gymnocarpon dwarf clover PerF Wide Pres Unc Weber & 1962 (Leguminosae) gymnocarpon var. gymnocarpon Miller 11316 (BLCA) Fabaceae Trifolium pratense red clover PerF Intro Rep Rare NCPN (in ed.) 2002 Native to Europe (Leguminosae) Fabaceae Trifolium repens white clover PerF Intro Pres Unc Weber & 1983 Native to Europe (Leguminosae) Beck 16948 (BLCA) Chapter 6:Appendices Fabaceae Vicia americana var. American vetch PerF Wide Pres Unc Clark & 1961 (Leguminosae) americana Hogan 1994 (BLCA) Fagaceae Quercus gambelii Q. gambelii var. Gambel oak Tree Wide Pres Abun Miller & 1961 gambelii Miller s.n. (BLCA) 47 48

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Fumariaceae Corydalis aurea C. aurea var. aurea golden smoke PerF Wide Pres Unc Cottrell & 1961 Gregory 279 (BLCA) Fumariaceae Corydalis curvisiliqua C. aurea var. golden corydalis PerF Wide Pres Unc Weber & 1962 ssp. occidentalis occidentalis Miller 11383 (BLCA) Gentianaceae Frasera speciosa Swertia radiata elkweed PerF Wide Pres Unc Weber, 1983 Rector, & Kennedy 16819 (BLCA) Geraniaceae Erodium cicutarium stork’s-bill AnnF Intro Rep Unc NCPN (in ed.) 2002 Native to Europe. Weber & Beck 17066 (BLCA) is from just outside the park. Geraniaceae Geranium Richardson’s PerF Wide Pres Unc Weber, 1983 richardsonii crane’s-bill Rector, & Kennedy 16835 (BLCA) Grossulariaceae Ribes cereum R. cereum var. wax currant Shrub Wide Pres Unc Weber & 1962 (Saxifragaceae) pedicellare, Miller 11334 R. cereum var. (BLCA) inebrians Grossulariaceae Ribes leptanthum trumpet Shrub Wide Pres Unc Lederer & 1962 (Saxifragaceae) gooseberry Clark 03-46 (BLCA) Grossulariaceae Ribes wolfi i wolf currant Shrub Wide Pres Unc Cottrell & 1974 (Saxifragaceae) Gregory 275 (BLCA) Helleboraceae Actaea rubra ssp. A. rubra red baneberry PerF Wide Pres Unc Hogan 4160 1983 (Ranunculaceae) aArguta (BLCA) Helleboraceae Aquilegia elegantula elegant PerF Wide Pres Unc Koepsel s.n. 1980 (Ranunculaceae) columbine (BLCA) Helleboraceae Aquilegia micrantha alcove columbine PerF RegEn Pres Unc Hogan & 2002 (Ranunculaceae) Tembrock 4264 (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Helleboraceae Delphinium D. nelsonii Nuttall’s larkspur PerF Wide Pres Unc Clark & 1961 Type locality is (Ranunculaceae) nuttallianum Hogan 2017 Bluff, San Juan (BLCA) Co., UT (Wetherill s.n. CAS) Helleboraceae Delphinium mountain PerF RegEn Rep Unc NCPN (in ed.) 2002 (Ranunculaceae) ramosum larkspur Hydrangeaceae Fendlera falcata F. rupicola var. Fendlerbush Shrub Wide Pres Unc Hogan 3965 1983 (Saxifragaceae) rupicola (BLCA) Hydrangeaceae Fendlera wrightii F. rupicola var. Wright’s Shrub Wide Pres Unc Weber & 1962 (Saxifragaceae) wrightii fendlerbush Miller 11380 (BLCA) Hydrangeaceae Fendlerella utahensis Utah fendlerella Shrub Periph Rep Rare NCPN (in ed.) 2002 (Saxifragaceae) Hydrangeaceae Philadelphus P. microphyllus littleleaf mock- Shrub Wide Pres Unc Hogan 4117 1959 (Saxifragaceae) microphyllus var. occidentalis, orange (BLCA) P. occidentalis var. occidentalis Hydrophyllaceae Hydrophyllum H. capitatum var. capitate PerF Wide Pres Unc Penland 1808 1942 capitatum capitatum waterleaf (COLO) Hydrophyllaceae Phacelia splendens Eastwood’s AnnF RegEn Pres Rare Hogan 4210 2004 Type locality is phacelia (BLCA) Grand Junction, Mesa Co., CO (Eastwood s.n. CAS) Hypericaceae Hypericum H. scouleri, H. western St. PerF Wide Pres Unc Weber & 1983 (Guttiferae, formosum formosum var. John’s-wort Beck 16909 Clusiaceae) scouleri (BLCA) Juncaceae Juncus arcticus ssp. J. balticus var. ater montanus, J. Baltic rush PerG Wide Pres Com Weber, 1983 Chapter 6:Appendices arcticus var. Rector, & balticus; Kennedy includes 16747 (BLCA) var. mexicanus. Juncaceae Juncus bufonius Includes vars. toad rush AnnG Wide Pres Unc Weber & 1983 bufonius, Beck 16901 occidentalis, (BLCA) halophilus, & ambiguus 49 50

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Juncaceae Juncus confusus perplexing rush PerG Wide Pres Unc Weber, 1983 Rector, & Kennedy 16752 (BLCA) Juncaceae Juncus dudleyi J. tenuis var. slender rush PerG Wide Pres Unc Lederer & 2004 dudleyi Clark 04-25 (BLCA) Juncaceae Juncus longistylis longstyle rush PerG Wide Pres Com Weber & 1983 Beck 16914 (BLCA) Juncaceae Juncus saximontanus J. ensifolius mountain rush PerG Wide Pres Unc Weber & 1983 var. montanus; Beck 16907 includes J. tracyi, J. (BLCA) xiphoides Juncaceae Juncus torreyi Torrey’s rush PerG Wide Rep Hogan et al. NA (2005), NPS vegetation plots Lamiaceae Agastache urticifolia nettle-leaf PerF Wide Pres Unc Weber & 1983 (Labiatae) horsemint Beck 16960 (BLCA) Lamiaceae Dracocephalum Moldavica smallfl ower PerF Wide Pres Unc Hogan & 1983 (Labiatae) parvifl orum parvifl ora dragonhead Tembrock 4303 (BLCA) Lamiaceae Marrubium vulgare common PerF Intro Pres Unc Hogan & 2003 Native to Eurasia (Labiatae) horehound Tembrock 4378 (BLCA) Lamiaceae Mentha arvensis M. arvensis var. fi eld mint PerF Wide Pres Unc Lederer & 2004 (Labiatae) canadensis, M. Clark 04-23 arvensis var. (BLCA) glabrata Lamiaceae Monardella M. odoratissima stinking PerF Wide Pres Unc Hogan & 2004 (Labiatae) odoratissima var. glauca, M. horsemint Tembrock glauca 4342 (BLCA) Lemnaceae Lemna turionifera L. minor lesser duckweed PerF Wide Pres Unc Weber & 1983 Beck 16946 (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Liliaceae Fritillaria leopard-lily PerF Wide Pres Unc Lederer & 1984 atropurpurea Clark 03-07 (BLCA) Linaceae Adenolinum lewisii L. lewisii, L. lewisii blue ﬂ ax PerF Wide Pres Unc Clark & 2002 var. lewisii, L. Lederer 2184 perenne ssp. lewisii (BLCA) Loasaceae Acrolasia albicaulis Mentzelia white-stem AnnF Wide Pres Unc Hogan 4224 2004 albicaulis, M. blazingstar (BLCA) obscura. Most authors include A. gracilis Loasaceae Acrolasia dispersa Mentzelia dispersa; Nevada stickleaf AnnF Wide Pres Unc Weber & 1983 includes vars. Beck 16981 compacta, dispersa, (BLCA) & latifolia Loasaceae Acrolasia humilis Nuttallia humilis, Thompson’s AnnF RegEn Pres Unc Hogan 4229 2004 Type locality Mentzelia humilis, stickleaf (BLCA) is 7 miles E M. thompsonii of Montrose, Montrose Co., CO (Weber 7470) Loasaceae Nuttallia Mentzelia wing-seed PerF Wide Pres Rare Lederer, 2004 Type locality is pterosperma pterosperma stickleaf Clark, Willow Creek, Hogan, & San Juan Co., UT Tembrock 04- (Eastwood 31 63 (BLCA) CAS) Malvaceae Malva neglecta common mallow AnnF Intro Pres Unc Clark & 2004 Native to Eurasia Lederer 2191 (BLCA) Malvaceae Sphaeralcea coccinea Includes ssp. scarlet globe- PerF Wide Pres Unc Hogan 4206 2002

Chapter 6:Appendices ssp. dissecta coccinea, elata, & mallow (BLCA) dissecta Melanthiaceae Toxicoscordion Zigadenus foothills death- PerF Wide Pres ? Miller s.n. 1961 (Liliaceae) paniculatum paniculatus camas (BLCA) Melanthiaceae Toxicoscordion Zigadenus meadow death PerF Wide Pres Unc Penland 1806 (Liliaceae) venenosum venenosus var. camas (COLO) 1942 venenosus 51 52

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Nyctaginaceae Abronia argillosa clay sand-verbena PerF RegEn Pres Unc Hogan & 2004 Type locality is 6 Tembrock miles S of Cisco, 4295 (BLCA) Grand Co., UT (Welsh et al. 16689 BRY) Nyctaginaceae Oxybaphus linearis O. decumbens, narrow-leaf four- PerF Wide Pres Unc Weber & 1983 Mirabilis linearis o’clock Beck 16988 var. linearis; some (BLCA) authors include M. glabra Oleaceae Fraxinus anomala singleleaf ash Shrub Wide Pres Com Weber & 1974 Beck 17001 (BLCA) Onagraceae Camissonia C. scapoidea Paiute suncup AnnF Wide Pres Unc Hogan 4225 2004 scapoidea var. scapoidea, (BLCA) Oenothera scapoidea. Includes ssp. brachycarpa Onagraceae Epilobium E. paniculatum var. autumn willow- AnnF Wide Pres Unc Weber & 1983 brachycarpum paniculatum herb Beck 16906 (BLCA) Onagraceae Epilobium ciliatum E. ciliatum northern willow- PerF Wide Pres Com Hogan & 1983 var. ciliatum, herb Tembrock E. watsonii, E. 4337 (BLCA) adenocaulon Onagraceae Epilobium halleanum Hall’s willow-herb PerF Wide Pres ? Weber 16912 1983 Duplicate (COLO) collection at BLCA is identifi ed as E. ciliatum. Onagraceae Gayophytum G. diffusum var. diffuse AnnF Wide Pres Unc Beck & 1983 diffusum ssp. strictipes, G. groundsmoke Kennedy parvifl orum lasiospermum 17061 (BLCA) Onagraceae Gayophytum branching AnnF Wide Pres Unc Weber, 1983 ramosissimum groundsmoke Rector, & Kennedy 16842 (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Onagraceae Oenothera long-tube PerF Wide Pres Com Weber & 1962 caespitosa ssp. evening-primrose Miller 11385 marginata (BLCA) Onagraceae Oenothera hairy-throat PerF Wide Pres Unc Clark & 1961 coronopifolia evening-primrose Hogan 1998 (BLCA) Orchidaceae Corallorhiza Includes vars. spotted coral-root PerF Wide Rep Rare NCPN (in ed.) 2002 maculata maculata & occidentalis Orchidaceae Epipactis gigantea giant helleborine PerF Wide Rep Unk Michael Scott ? veg plot Orobanchaceae Aphyllon unifl orum Orobanche unifl ora naked broomrape PerF Wide Pres Unc Clark & 2004 var. occidentalis Lederer 2189 (BLCA) Pinaceae Picea pungens blue spruce Tree Wide Pres Unc Weber & 1962 State tree Miller 11367 of Utah and (BLCA) Colorado Pinaceae Pinus edulis two-needle Tree Wide Pres Abun Cottrell & 1974 pinyon Gregory 253 (BLCA) Pinaceae Pinus ponderosa var. ponderosa pine Tree Wide Pres Unc Weber & 1983 scopulorum Beck 16703 (BLCA) Pinaceae Pseudotsuga P. menziesii var. Douglas-fi r Tree Wide Pres Com Cottrell & 1974 menziesii glauca Gregory 276 (BLCA) Plantaginaceae Plantago major Includes var. common plantain PerF Intro Pres Unc Weber & 1983 pachyphylla Beck 16902 (BLCA) Chapter 6:Appendices Plantaginaceae Plantago patagonica P. patagonica var. woolly plantain AnnF Wide Rep Rare NCPN (in ed.) 2002 Mostly native to patagonica Europe Poaceae Achnatherum Oryzopsis Indian ricegrass PerG Wide Pres Com Hogan & 1962 (Gramineae) hymenoides hymenoides, Stipa Clark 4077 hymenoides (BLCA) Poaceae Achnatherum (Gramineae) lettermanii Stipa lettermanii Letterman’s PerG Wide Pres Unc Rominger 1962 needlegrass s.n. (BLCA) 53 54

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Poaceae Achnatherum A. nelsonii var. Nelson’s PerG Wide Pres Unc Hogan 4146 1962 (Gramineae) nelsonii nelsonii, Stipa needlegrass (BLCA) nelsonii var. nelsonii Poaceae Achnatherum Stipa pinetorum pine needlegrass PerG Wide Pres Unc Hogan & 2002 (Gramineae) pinetorum Clark 4088 (BLCA) Poaceae Achnatherum x Stiporyzopsis x Bloomer’s PerG Wide Pres ? Rominger 1962 (Gramineae) bloomeri bloomeri, Stipa x ricegrass s.n. (BLCA) bloomeri, hybrids between Stipa hymenoides and other Stipa species except S. comata Poaceae Agropyron A. cristatum var. clustered PerG Intro Pres Unc Hogan & 1983 Native to Eurasia (Gramineae) desertorum desertorum; wheatgrass Tembrock included within A. 4347 (BLCA) cristatum (without var.) in Flora of North America (2007) Poaceae Agropyron A. cristatum var. crested PerG Intro Pres Unc Weber & 1962 Native to Eurasia (Gramineae) mongolicum cristatum, A. wheatgrass Beck 16919 cristatum ssp. (BLCA) pectinatum, A. pectinatum, A. sibiricum, A. fragile, A. mongolicum Poaceae Agrostis gigantea Included in A. redtop PerG Intro Pres Com Beck & 1962 Native to Eurasia (Gramineae) stolonifera by Kennedy some authors 17059 (BLCA) Poaceae Agrostis scabra A. hiemalis, A. (Gramineae) ticklegrass PerG Wide Pres Unc Weber & 1983 geminata Beck 16977 (BLCA) Poaceae Alopecurus aequalis A. aequalis var. shortawn foxtail PerG Wide Pres Unc Weber & 1983 (Gramineae) aequalis Beck 16944 (BLCA) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Poaceae Anisantha tectorum Bromus tectorum cheatgrass AnnG Intro Pres Com Weber, 1983 CO state noxious (Gramineae) Rector, & weed (List C). Kennedy Native to Eurasia. 16878 (BLCA) Poaceae Aristida purpurea A. glauca, A. purple three-awn PerG Wide Pres Com Weber & 1983 (Gramineae) var. longiseta purpurea var. Beck 16994 fendleriana, A. (BLCA) fendleriana Poaceae Blepharoneuron pine dropseed PerG Wide Pres Unc Beck 14 1983 (Gramineae) tricholepis (COLO) Poaceae Bromopsis Bromus ciliatus; fringed brome PerG Wide Pres Unc Weber 16968 1983 (Gramineae) canadensis includes vars. (COLO) ciliatus & richardsonii Poaceae Bromopsis inermis B. inermis var. smooth brome PerG Intro Pres Com Hogan & 1962 Native to Eurasia (Gramineae) inermis Tembrock 4294 (BLCA) Poaceae Bromopsis lanatipes Bromus lanatipes woolly brome PerG Wide Pres Unc Weber & 1983 (Gramineae) Beck 16980 (BLCA) Poaceae Bromopsis porteri Bromus frondosus, nodding brome PerG Wide Pres Unc Hogan & 1983 (Gramineae) Bromus anomalus. Lederer 4394 Material ascribed (BLCA) to Bromus anomalus in regional ﬂ oras is mostly B. porteri or B. lanatipes. Poaceae Calamagrostis Jones’ reedgrass PerG Wide Rep Rare NCPN (in ed.) 2002

Chapter 6:Appendices (Gramineae) scopulorum Poaceae Calamagrostis stricta C. neglecta, C. slimstem PerG Wide Pres Unc Rominger 1962 (Gramineae) inexpansa reedgrass s.n. (BLCA) Poaceae Ceratochloa carinata Bromus mountain brome PerG Wide Pres Unc Weber, 1983 (Gramineae) marginatus, B. Rector, & polyanthus, B. Kennedy carinatus 16823 (BLCA) 55 56

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Poaceae Chondrosum gracile Bouteloua gracilis blue grama PerG Wide Pres Com Weber & 1983 (Gramineae) Beck 16938 (BLCA) Poaceae Critesion jubatum Hordeum jubatum foxtail barley PerG Wide Pres Unc Rominger 1962 (Gramineae) s.n. (BLCA) Poaceae Dactylis glomerata orchard grass PerG Intro Pres Unc Hogan 4126 1983 Native to Eurasia (Gramineae) (BLCA) and Africa Poaceae Elymus canadensis E. canadensis var. Canada wildrye PerG Wide Pres Unc Beck & 1983 (Gramineae) canadensis Kennedy 17023 (BLCA) Poaceae Elymus elymoides E. elymoides var. squirreltail PerG Wide Pres Com Hogan & 2002 (Gramineae) elymoides, Sitanion Clark 4076 hystrix (BLCA) Poaceae Elymus glaucus E. glaucus var. blue wildrye PerG Wide Rep Rare NCPN (in ed.) 2002 (Gramineae) glaucus Poaceae Elymus lanceolatus E. lanceolatus thickspike PerG Wide Pres Unc Weber & 1962 (Gramineae) var. lanceolatus, wheatgrass Beck 16978 Agropyron (BLCA) dasystachyum var. dasystachyum, A. albicans var. albicans, A. albicans var. griffi thsii. Includes var. riparium Poaceae Elymus longifolius Sitanion hystrix, bottlebrush PerG Wide Pres Unc Rominger 1962 (Gramineae) E. elymoides var. squirreltail s.n. (BLCA) brevifolius Poaceae Elymus trachycaulus Agropyron slender PerG Wide Pres Unc Hogan & 1962 (Gramineae) caninum, A. wheatgrass Tembrock subsecundum, 4349 (BLCA) A. trachycaulum, A. latiglume, Elymus alaskanus ssp. latiglumis, E. trachycaulus var. trachycaulus Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Poaceae Elytrigia repens Agropyron repens, quackgrass PerG Intro Pres Unc Weber, 1962 CO state noxious (Gramineae) Elymus repens Rector, & weed (List B). Kennedy Native to Eurasia. 16806 (BLCA) Poaceae Eremopyrum Agropyron annual AnnG Intro Pres Unc Lederer, 2004 Native to Asia (Gramineae) triticeum triticeum wheatgrass Clark, Hogan, & Tembrock 04- 38 (BLCA) Poaceae Festuca arizonica F. ovina var. Arizona fescue PerG Wide Rep Rare NCPN (in ed.) 2002 (Gramineae) arizonica Poaceae Festuca arundinacea Schedonorus tall fescue PerG Intro Pres Unc Weber & 1983 Native to Europe (Gramineae) phoenix, S. Beck 16955 arundinaceus, (BLCA) Lolium arundinaceum Poaceae Festuca idahoensis F. ovina var. ingrata Idaho fescue PerG Wide Pres Unc Hogan 4128 2003 (Gramineae) (BLCA) Poaceae Festuca saximontana F. ovina var. Rocky Mountain PerG Wide Pres Unc Weber 11338 1962 (Gramineae) rydbergii, F. ovina fescue (COLO) var. saximontana Poaceae Festuca thurberi Thurber’s fescue PerG Wide Pres Unc Weber, 1983 (Gramineae) Rector, & Kennedy 16817 (BLCA) Poaceae Hesperostipa comata H. comata var. needle-and- PerG Wide Pres Com Rominger 1962 (Gramineae) comata, Stipa thread s.n. (BLCA) comata var. comata Poaceae Hilaria jamesii Pleuraphis jamesii galleta PerG Wide Pres Unc Lederer, 2002 Chapter 6:Appendices (Gramineae) Clark, Hogan, & Tembrock 04- 40 (BLCA) Poaceae Koeleria macrantha K. cristata, K. Junegrass PerG Wide Pres Com Hogan & 1962 (Gramineae) pyramidata, K. Clark 4091 nitida (BLCA) 57 58

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Poaceae Leymus cinereus Elymus cinereus Great Basin PerG Wide Pres Unc Weber, 1962 (Gramineae) wildrye Rector, & Kennedy 16827 (BLCA) Poaceae Leymus salina Elymus salina, Salina wildrye PerG Wide Pres Unc Lederer, 2004 (Gramineae) E. salinus, E. Clark, ambiguus, Hogan, & L. salinus, L. Tembrock 04- ambiguus 61 (BLCA) Poaceae Melica porteri Porter’s melic PerG Wide Pres Unc Hogan 4161 1983 (Gramineae) (BLCA) Poaceae Monroa squarrosa Munroa squarrosa false buffalograss AnnG Wide Pres ? Weber & 1983 (Gramineae) Beck 16937 (BLCA) Poaceae Muhlenbergia scratchgrass PerG Wide Pres Unc Rominger 1962 (Gramineae) asperifolia s.n. (BLCA) Poaceae Muhlenbergia green muhly PerG Wide Pres Unc Hogan 4173 1962 (Gramineae) racemosa (BLCA) Poaceae Pascopyrum smithii Agropyron smithii, western PerG Wide Pres Com Hogan & 1962 (Gramineae) Elymus smithii; wheatgrass Tembrock includes vars. 4346 (BLCA) smithii & palmeri Poaceae Phalaroides Phalaris reed canary grass PerG Wide Pres Com Hogan 4100 2002 (Gramineae) arundinacea arundinacea (BLCA) Poaceae Phleum pratense timothy PerG Intro Pres Unc Weber & 1983 Native to Eurasia (Gramineae) Beck 16923 (BLCA) Poaceae Piptatherum Oryzopsis littleseed PerG Wide Pres Unc Hogan 4114 1962 (Gramineae) micranthum micrantha ricegrass (BLCA) Poaceae Poa agassizensis P. pratensis ssp. Agassiz’s PerG Wide Pres Unc Rominger 1962 (Gramineae) agassizensis; bluegrass s.n. (BLCA) included in P. pratensis by some authors Poaceae Poa bigelovii Bigelow’s AnnG Periph Rep Rare NCPN (in ed.) 2002 (Gramineae) bluegrass Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Poaceae Poa bulbosa bulbous bluegrass PerG Intro Pres Unc Hogan 3975 2003 Native to Eurasia (Gramineae) (BLCA) and Africa Poaceae Poa compressa Canada bluegrass PerG Intro Pres Unc Hogan & 1983 Native to Eurasia (Gramineae) Lederer 4396 (BLCA) Poaceae Poa fendleriana P. longiligula, P. muttongrass PerG Wide Pres Com Lederer & 1962 (Gramineae) fendleriana ssp. Clark 03-34 longiligula (BLCA) Poaceae Poa glaucifolia Included in P. arida pale-leaf PerG Wide Pres Unc Weber, 1983 (Gramineae) by some authors bluegrass Rector, & Kennedy 16751 (BLCA) Poaceae Poa nemoralis ssp. P. glauca var. interior bluegrass PerG Wide Pres Unc Weber, 1983 (Gramineae) interior glauca, P. interior Rector, & Kennedy 16848 (BLCA) Poaceae Poa pratensis Most authors Kentucky PerG Intro Pres Unc Weber & 1983 Native to Europe (Gramineae) include P. bluegrass Beck 16885 agassizensis (BLCA) Poaceae Poa secunda Includes P. Sandberg’s PerG Wide Pres Com Clark & 1962 (Gramineae) gracillima, P. bluegrass Lederer 2196 juncifolia, P. (BLCA) sandbergii, P. canbyi, P. scabrella, P. nevadensis, P. ampla, & all vars. of P. secunda Poaceae Pseudoroegneria Agropyron bluebunch PerG Wide Pres Unc Hogan & 2003 (Gramineae) spicata spicatum, Elymus wheatgrass Clark 4079

Chapter 6:Appendices spicatus; includes (BLCA) ssp. inermis & spicata Poaceae Sporobolus sand dropseed PerG Wide Pres Unc Weber & 1983 (Gramineae) cryptandrus Beck 16936 (BLCA) 59 60

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Poaceae Thinopyrum Agropyron intermediate PerG Intro Pres ? Weber & 1983 Native to Eurasia (Gramineae) intermedium ssp. intermedium, wheatgrass Beck 16954 barbulatum Elytrigia (BLCA) intermedia, T. intermedium, Elymus hispidus var. ruthenicus Poaceae Thinopyrum Agropyron intermediate PerG Intro Pres ? Weber & 1983 Native to Eurasia (Gramineae) intermedium ssp. intermedium, wheatgrass Beck 16952 intermedium Elytrigia (BLCA) intermedia, Elymus hispidus var. hispidus Polemoniaceae Collomia linearis small collomia AnnF Wide Pres Com Clark & 1983 Hogan 2013 (BLCA) Polemoniaceae Gilia ophthalmoides Included in G. eyelash gilia AnnF Wide Pres Unc Lederer & 1962 Includes reports inconspicua by Clark 03-45 of G. inconspicua some authors (BLCA) from NCPN (in ed.) Polemoniaceae Gilia penstemonoides Aliciella Black Canyon PerF LocEn Pres Rare Grey 714 1980 Type locality penstemonoides gilia (BLCA) is Cimarron, Montrose Co., CO (Jones s.n.) Polemoniaceae Gilia pinnatifi da Aliciella pinnatifi da sticky gilia PerF Wide Rep Unc NCPN (in ed.) 2002 Polemoniaceae Ipomopsis aggregata Gilia aggregata var. scarlet gilia PerF Wide Pres Unc Hogan & 1959 ssp. aggregata aggregata Clark 4075 (BLCA) Polemoniaceae Linanthus harknessii Leptosiphon northern AnnF Wide Pres Unc Weber, 1983 var. septentrionalis septentrionalis, linanthus Rector, & Linanthus Kennedy septentrionalis 16775 (BLCA) Polemoniaceae Microsteris gracilis M. gracilis var. slender phlox AnnF Wide Pres Unc Weber, 1962 humilior, Phlox Rector, & gracilis ssp. humilis Kennedy 16760 (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Polemoniaceae Navarretia breweri yellow navarretia AnnF Wide Pres Unc Weber, 1983 Rector, & Kennedy 16771 (BLCA) Polemoniaceae Phlox austromontana P. austromontana desert phlox PerF Wide Pres Unc Lederer & 2004 var. Clark 04-34 austromontana; (BLCA) var. not recognized in Cronquist et al. (1984) Polemoniaceae Phlox hoodii P. hoodii ssp. Hood’s phlox PerF Wide Pres Unc Penland 1804 1942 canescens; includes (COLO) ssp. glabrata, hoodii, & viscidula Polemoniaceae Phlox longifolia Includes vars. long-leaf phlox PerF Wide Pres Unc Lederer & 1942 longifolia & Clark 03-08 stansburyi (BLCA) Polygonaceae Acetosella vulgaris Rumex acetosella sheep sorrel PerF Intro Pres Unc Weber & 1983 Native to Eurasia Beck 16957 (BLCA) Polygonaceae Eriogonum ﬂ avum yellow wild- PerF Wide Pres Unc Weber & 1959 ssp. ﬂ avum buckwheat Beck 17015 (BLCA) Polygonaceae Eriogonum gordonii Gordon’s wild AnnF Wide Pres Unc Hogan 4227 2002 buckwheat (BLCA) Polygonaceae Eriogonum E. leptocladon var. sand wild Shrub RegEn Rep Rare NCPN (in ed.) 2002 leptocladon leptocladon buckwheat Polygonaceae Eriogonum E. lonchophyllum longleaf wild Shrub RegEn Pres Com Weber, 1983 lonchophyllum var. lonchophyllum, buckwheat Rector, & Chapter 6:Appendices E. intermontanum, Kennedy E. corymbosum var. 16843 (BLCA) humivagans. Vars. not recognized in Flora of North America (2005) Polygonaceae Eriogonum E. racemosum var. redroot wild PerF Wide Pres Com Hogan & racemosum racemosum buckwheat Tembrock 1959

61 4355 (BLCA) 62

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Polygonaceae Eriogonum sulphur-fl ower PerF RegEn Pres Com Weber 16925 1959 umbellatum var. wild-buckwheat (COLO) aureum Polygonaceae Polygonum P. aviculare, P. yard knotweed AnnF Intro Pres Unc Hogan & 1983 Native to Eurasia arenastrum rurivagum Lederer 4389 (BLCA) Polygonaceae Polygonum douglasii P. douglasii var. Douglas’ AnnF Wide Pres Unc Weber & 1983 douglasii knotweed Beck 16895 (BLCA) Polygonaceae Polygonum P. kelloggii var. Kellogg’s AnnF Wide Pres Unc Weber, 1983 polygaloides ssp. kelloggii knotweed Rector, & kelloggii Kennedy 16798 (BLCA) Polygonaceae Pterogonum alatum Eriogonum alatum winged wild PerF Wide Pres Unc Hogan & 2002 buckwheat Clark 4064 (BLCA) Polygonaceae Rumex crispus curly dock PerF Intro Pres Unc Weber & 1983 Native to Eurasia Beck 16999 (BLCA) Polygonaceae Rumex triangulivalvis R. salicifolius var. willow dock PerF Wide Pres Unc Weber & 1983 triangulivalvis, Beck 17006 R. mexicanus, (BLCA) R. salicifolius var. mexicanus, R. utahensis, R. triangulivalvis var. mexicanus, R. triangulivalvis var. montigenitus. Portulacaceae Claytonia lanceolata C. lanceolata var. lanceleaf PerF Wide Pres Unc Weber & 1962 lanceolata. Some springbeauty Miller 11359 authors include C. (BLCA) rosea Portulacaceae Oreobroma pygmaea Lewisia pygmaea alpine bitter-root PerF Wide Pres ? Weber, 1983 Rector, & Kennedy 16753 (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Primulaceae Androsace western rock- AnnF Wide Pres Unc Lederer & 1983 occidentalis jasmine Clark 03-28 (BLCA) Primulaceae Androsace Includes ssp. pygmy-fl ower AnnF Wide Pres Com Hogan 3977 1983 septentrionalis puberulenta, rock-jasmine (BLCA) subumbellata, & var. subilifera Ranunculaceae Atragene Clematis Columbian PerF Wide Rep Rare NCPN (in ed.) 2002 columbiana columbiana var. virgin’s-bower columbiana, C. pseudoalpina Ranunculaceae Ceratocephala Ranunculus bur buttercup AnnF Intro Pres Unc Hogan 3979 1983 Native to Eurasia orthoceras testiculatus, C. (BLCA) testiculata Ranunculaceae Clematis ligusticifolia white virgin’s- PerF Wide Pres Unc Beck & 1983 bower Kennedy 17021 (BLCA) Ranunculaceae Corifl ora hirsutissima Clematis sugar-bowls PerF Wide Pres Unc Clark & 1961 hirsutissima Hogan 2016 (BLCA) Ranunculaceae Halerpestes Ranunculus marsh buttercup PerF Wide Pres Unc Weber & 1983 cymbalaria ssp. cymbalaria var. Beck 16947 saximontana cymbalaria. Vars. (BLCA) not recognized in Flora of North America (1997) Ranunculaceae Hecatonia scelerata Ranunculus blister buttercup AnnF Wide Pres Unc Weber & 1983 sceleratus var. Beck 16942 multifi dus (BLCA)

Chapter 6:Appendices Ranunculaceae Myosurus cupulatus horseshoe AnnF Periph Pres Unc Hogan 3982 2003 mousetail (BLCA) Ranunculaceae Pulsatilla ludoviciana Anemone nuttalliana, pasquefl ower PerF Wide Pres Unc Weber & 1962 A. patens var. Miller 11390 multifi da, P. patens (BLCA) ssp. multifi da 63 64

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Ranunculaceae Ranunculus Includes R. sagebrush PerF Wide Pres Com Lederer & 1960 glaberrimus var. oreogenes buttercup Clark 03-09 ellipticus (BLCA) Rhamnaceae Ceanothus fendleri Fendler’s Shrub Wide Pres Unc Weber & 1983 mountain-lilac Beck 16973 (BLCA) Rhamnaceae Ceanothus velutinus C. velutinus var. deer-brush Shrub Wide Pres Unc Weber, 1962 velutinus Rector, & Kennedy 16854 (BLCA) Rosaceae Amelanchier alnifolia A. alnifolia var. Saskatoon Shrub Wide Pres Abun Cottrell & 1974 alnifolia serviceberry Gregory 272 (BLCA) Rosaceae Amelanchier Utah serviceberry Shrub Wide Pres Unc Lederer & 1961 utahensis Clark 03-18 (BLCA) Rosaceae Argentina anserina Potentilla anserina silverweed PerF Wide Pres Unc Cottrell & 1974 Gregory 257 (BLCA) Rosaceae Cercocarpus alder-leaf Shrub Wide Pres Com Miller & 1961 montanus mountain Miller s.n. mahogany (BLCA) Rosaceae Drymocallis arguta Potentilla arguta tall cinquefoil PerF Wide Pres Unc Weber, 1983 var. convallaria, Rector, & includes ssp. arguta Kennedy 16793 (BLCA) Rosaceae Erythrocoma trifl ora Geum trifl orum prairie-smoke PerF Wide Pres Unc Lederer & 1983 var. trifl orum Clark 03-22 (BLCA) Rosaceae Fragaria vesca ssp. F. vesca woodland PerF Wide Pres Unc Beck & 1983 bracteata strawberry Kennedy 17044 (BLCA) Rosaceae Fragaria virginiana F. virginiana Virginia PerF Wide Pres Unc Cottrell & 1974 ssp. glauca strawberry Gregory 274 (BLCA) Rosaceae Holodiscus discolor H. dumosus bush oceanspray Shrub Wide Pres Com Weber 17011 1959 (COLO) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Rosaceae Padus virginiana ssp. Prunus virginiana western Shrub Wide Pres Com Weber, 1983 melanocarpa var. melanocarpa chokecherry Rector, & Kennedy 16816 (BLCA) Rosaceae Peraphyllum squaw-apple Shrub Wide Pres ? Miller & 1959 ramosissimum Miller s.n. (BLCA) Rosaceae Petrophyton Spiraea caespitosa rock spiraea PerF Wide Pres Unc Hogan & 2004 caespitosum Tembrock 4266 (BLCA) Rosaceae Potentilla norvegica P. norvegica ssp. Norwegian AnnF Intro Pres Unc Beck & 1983 Native to Europe monspeliensis cinquefoil Kennedy and possibly E 17028 (BLCA) North America Rosaceae Potentilla P. gracilis var. beautiful PerF Wide Pres Unc Weber, 1983 hybrids with P. pulcherrima pulcherrima cinquefoil Rector, & hippiana also Kennedy occur in BLCA 16762 (BLCA) (Clark & Hogan 2012) Rosaceae Purshia tridentata antelope Shrub Wide Rep Unc NCPN (in ed.) 2002 bitterbrush Rosaceae Rosa woodsii R. woodsii var. Woods’ rose Shrub Wide Pres Unc Weber, 1983 ultramontana; Rector, & some authors Kennedy include R. manca 16845 (BLCA) Rosaceae Rubus armeniacus Rubus discolor Himalayan Shrub Intro Rep Unc Dewey & 2003 Native to Eurasia blackberry Andersen (2005) Rosaceae Rubus idaeus ssp. R. idaeus var. common red Shrub Wide Pres Unc Beck & 1983 melanolasius strigosus, R. idaeus raspberry Kennedy Chapter 6:Appendices var. aculeatissimus 17039 (BLCA) Rubiaceae Galium bifolium twin-leaf AnnF Wide Pres Unc Weber, 1983 bedstraw Rector, & Kennedy 16774 (BLCA) 65 66

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Rubiaceae Galium coloradoense G. multifl orum var. Colorado PerF RegEn Pres Com Hogan & 1962 Type locality coloradoense bedstraw Clark 4053 is Mancos, (BLCA) Montezuma Co., CO (Eastwood s.n.) Rubiaceae Galium G. boreale northern PerF Wide Pres Com Weber & 1983 septentrionale bedstraw Beck 16961 (BLCA) Rubiaceae Galium spurium G. aparine var. cleavers AnnF Wide Pres Unc Weber, 1962 echinospermon Rector, & Kennedy 16807 (BLCA) Salicaceae Populus angustifolia narrow-leaved Tree Wide Pres Unc Cottrell & 1974 cottonwood Gregory 287 (BLCA) Salicaceae Populus deltoides P. fremontii var. Rio Grande Tree Wide Pres Unc Hogan & 2004 ssp. wislizenii wislizenii cottonwood Tembrock 4375 (BLCA) Salicaceae Populus tremuloides quaking aspen Tree Wide Pres Unc Weber 11374 1962 (COLO) Salicaceae Salix exigua S. exigua var. coyote willow Shrub Wide Pres Com Hogan & 1983 exigua, S. exigua Tembrock var. stenophylla 4377 (BLCA) Santalaceae Comandra umbellata C. umbellata var. bastard toadfl ax PerF Wide Pres Com Hogan 4150 1962 pallida (BLCA) Saxifragaceae Ciliaria Saxifraga spotted saxifrage PerF Wide Pres Unc Hogan & 1962 austromontana bronchialis var. Tembrock austromontana 4275 (BLCA) Saxifragaceae Heuchera parvifolia Includes vars. littleleaf PerF Wide Pres Unc Clark & 1959 BLCA material is major, parvifolia, & alumroot Hogan 2022 var. parvifolia. utahensis (BLCA) Saxifragaceae Lithophragma L. bulbifera, L. glabrum fringecup PerF Wide Pres Unc Lederer & 2003 glabra, L. glabrum woodlandstar Clark 03-25b var. ramulosum (BLCA) Saxifragaceae Lithophragma L. tenella slender PerF Wide Pres Unc Cottrell & 1960 tenellum woodlandstar Gregory 265 (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Saxifragaceae Sullivantia hapemanii S. hapemanii Purpus’ sullivantia PerF RegEn Pres Unc Hogan & 1892 Type locality is var. purpusii Tembrock Black Canyon, 4334 (BLCA) Montrose Co., CO (Purpus 512 UC) Scrophulariaceae Castilleja chromosa C. angustifolia desert paintbrush PerF Wide Pres Com Cottrell & 1961 var. dubia, C. Gregory 263 applegatei ssp. (BLCA) martinii Scrophulariaceae Castilleja linariifolia Wyoming PerF Wide Pres Unc Lederer & 1959 paintbrush Clark 04-35 (BLCA) Scrophulariaceae Castilleja miniata Includes C. scarlet paintbrush PerF Wide Rep Unc NCPN (in ed.) 2002 gracillima Scrophulariaceae Collinsia parvifl ora blue-eyed Mary AnnF Wide Pres Com Lederer & 1961 Clark 03-02 (BLCA) Scrophulariaceae Limosella aquatica mudwort PerF Wide Pres ? Weber & 1983 Beck 16900 (BLCA) Scrophulariaceae Linaria vulgaris butter-and-eggs PerF Intro Rep Unc Dewey & 2003 CO state noxious Andersen weed (List B). (2005) Native to Eurasia. Scrophulariaceae Mimulus fl oribundus seep AnnF Wide Pres Unc Kennedy s.n. 1983 monkeyfl ower (BLCA) Scrophulariaceae Mimulus rubellus reddish AnnF Wide Pres Unc Hogan 3981 2003 monkeyfl ower (BLCA) Scrophulariaceae Orthocarpus luteus yellow owl-clover AnnF Wide Pres Com Weber & 1983 Beck 16897 (BLCA) Chapter 6:Appendices Scrophulariaceae Pedicularis dwarf lousewort PerF Wide Pres Unc Hogan & 2003 centranthera Clark 4068 (BLCA) Scrophulariaceae Pedicularis procera Gray’s lousewort Perf Wide Rep Rare NCPN (in ed.) 2002 Scrophulariaceae Penstemon barbatus P. barbatus var. beardlip PerF Wide Pres Unc Lederer & 1983 var. torreyi barbatus penstemon Clark 04-30 (BLCA) 67 68

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Scrophulariaceae Penstemon P. caespitosus mat penstemon PerF RegEn Pres Com Beidleman 1961 caespitosus var. caespitosus; s.n. (BLCA) includes P. teucrioides Scrophulariaceae Penstemon retrorsus Included in P. Adobe Hills PerF LocEn Pres Unc Hogan 4219 2004 Type locality is caespitosus by beardtongue (BLCA) near Montrose, some authors Montrose Co., CO (Payson 673 US) Scrophulariaceae Penstemon strictus Rocky Mountain PerF Wide Pres Com Hogan & 1959 penstemon Clark 4090 (BLCA) Scrophulariaceae Penstemon watsonii Watson’s PerF Wide Pres Unc Weber, 1983 penstemon Rector, & Kennedy 16778 (BLCA) Scrophulariaceae Pocilla biloba Veronica biloba two-lobe AnnF Intro Pres Unc Clark & 2003 Native to Asia speedwell Hogan 2026 (BLCA) Scrophulariaceae Scrophularia lance-leaf fi gwort PerF Wide Pres Unc Hogan & 1983 lanceolata Tembrock 4270 (BLCA) Scrophulariaceae Verbascum thapsus woolly mullein PerF Intro Pres Unc Weber & 1983 CO state noxious Beck 16956 weed (List C). (BLCA) Native to Eurasia. Scrophulariaceae Veronica americana American PerF Wide Pres Unc Weber & 1983 brooklime Beck 16941 (BLCA) Scrophulariaceae Veronica catenata Included in V. chain speedwell PerF Wide Pres Unc Hogan 4111 2003 anagallis-aquatica (BLCA) by some authors Scrophulariaceae Veronica peregrina purslane AnnF Wide Pres Unc Weber, 1983 var. xalapensis speedwell Rector, & Kennedy 16797 (BLCA) Selaginellaceae Selaginella densa Includes vars. dense spike-moss fern Wide Pres Unc Weber & 1962 standleyi, densa, & Miller 11348 scopulorum (BLCA) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Black Canyon of the Gunnison National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Selaginellaceae Selaginella mutica S. mutica var. blunt-leaf spike- fern Wide Pres Unc Hogan & 1962 mutica moss Tembrock 4268 (BLCA) Simaroubaceae Ailanthus altissima tree-of-heaven Tree Intro Rep Rare Dewey & 2003 Native to China Andersen (2005) Sinopteridaceae Argyrochosma Notholaena Fendler’s cloak- fern Sparse Pres Unc Hogan & 2004 (Polypodiaceae, fendleri fendleri fern Tembrock Adiantaceae) 4269 (BLCA) Sinopteridaceae Cheilanthes feei slender lip-fern fern Wide Pres Unc Lederer & 2004 (Polypodiaceae, Clark 04-33 Adiantaceae) (BLCA) Solanaceae Lycium barbarum L. halmifolium matrimony-vine Shrub Intro Pres Rare Douglass 54- 1954 Native to Eurasia 300 (COLO) Tamaricaceae Tamarix ramosissima T. pentandra, T. ﬁ ve-stamen Shrub Intro Pres Unc Hogan & 2003 CO state noxious chinensis tamarisk Tembrock weed (List B). 4376 (BLCA) Native to Eurasia. Thalictraceae Thalictrum fendleri Fendler’s PerF Wide Pres Unc Hogan & 2004 (Ranunculaceae) meadow-rue Tembrock 4353 (BLCA) Typhaceae Typha latifolia broad-leaved PerG Wide Pres Unc Weber & 1983 cattail Beck 16949 (BLCA) Ulmaceae Celtis reticulata C. laevigata var. netleaf hackberry Tree Wide Pres Unc Hogan 4109 2003 (Celtidaceae) reticulata (BLCA) Ulmaceae Ulmus pumila Siberian elm Tree Intro Pres Rare Weber, 1983 Native to Asia Rector, & Kennedy 16787 (BLCA) Chapter 6:Appendices Urticaceae Parietaria Pennsylvania AnnF Wide Pres Unc Hogan & 2004 pensylvanica pellitory Tembrock 4335 (BLCA) Urticaceae Urtica gracilis ssp. U. dioica var. stinging nettle PerF Wide Pres Unc Weber, 1983 gracilis procera Rector, & Kennedy 16836 (BLCA) 69 70

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes Form status size doc. Uvulariaceae Prosartes trachycarpa Disporum rough-fruited PerF Wide Pres Unc Hogan 3990 1974 (Liliaceae, trachycarpum fairy-bells (BLCA) Calochortaceae) Valerianaceae Valeriana capitata V. acutiloba var. mountain PerF Wide Pres Unc Weber & 1983 Type locality var. acutiloba acutiloba; includes valerian Beck 16818 of V. acutiloba var. pubicarpa (BLCA) is Sangre de Cristo Range, CO (Rydberg & Vreeland 5576 NY). Verbenaceae Verbena bracteata prostrate vervain PerF Wide Pres Unc Weber & 1983 Beck 17000 (BLCA) Violaceae Viola adunca hook-spur violet PerF Wide Pres Unc Hogan 3983 1983 (BLCA) Violaceae Viola biternata V. sheltonii Shelton’s violet PerF Disj Pres ? Weber & 1962 Miller 11389 (BLCA) Violaceae Viola scopulorum V. canadensis Canada violet PerF Wide Pres Unc Weber 11364 1962 var. scopulorum; (COLO) included in V. canadensis by some authors Violaceae Viola vallicola V. nuttallii var. valley violet PerF Wide Pres Unc Clark & 2004 vallicola, V. Lederer 2175 vallicola var. (BLCA) vallicola Viscaceae Phoradendron juniper mistletoe PerF Wide Pres Unc Hogan & 2004 Typically parasitic juniperinum Tembrock on Juniperus 4344 (BLCA) osteosperma Woodsiaceae Woodsia oregana Oregon woodsia fern Wide Pres Unc Hogan 4174 1983 (Polypodiaceae, ssp. cathcartiana (BLCA) Aspleniaceae) Zannichelliaceae Zannichellia palustris horned PerF Wide Pres Unc Clark & 2003 pondweed Hogan 2015 (BLCA) Native to Eurasia. Known from CURE. PerF Wide Known from CURE Common name Life form Range Comments nodding onionpink funnel-lily PerFcommon chickweed PerFcreeping nailwort AnnF Wide Known from CURE long-leaved starwort Wide Intro PerF PerFprostrate pigweed Native to Eurasia redroot pigweed Widegiant angelica Wide AnnF Known from CURE Known from CURE caraway AnnF Wide PerF Present in CURE Intro Native to Central America RegEn Known from CURE PerF Intro CO state noxious weed (List B). water-hemlockrock lovage PerF spring- Constance’s parsley Wide Known from CURE PerF Wide Known from CURE cow parsnip cowbaneFendler’s PerFwater parsnip PerFspreading dogbane Wide Wide Known from CURE clasping leaved dogbane PerF PerF PerF Known from CURE milkweedHall’s Wide Wide Sparse Known from CURE Known from CURE PerF Sparse Known from CURE

var.

var. cernuum var. androsaemifolium obtusum. Synonyms/ notes Taxonomic Includes vars. & C. maculata angustifolia A. androsaemifolium var. hypericifolium H. sphondylium lanatum, H. maximum, H. lanatum

ssp. orum exus ora rmed) vascular plant taxa of Black Canyon the Gunnison National Park. Allium cernuum Androstephium brevifl Alsine mediaParonychia sessilifl Stellaria longifolia Stellaria media Amaranthus blitoidesAmaranthus retrofl Angelica ampla A. graecizans Carum carvi Conioselinum scopulorum Cymopterus constancei Cicuta douglasii C. occidentalis, Oxypolis fendleri Sium suave Apocynum androsaemifolium Apocynum sibiricumAsclepias hallii A. cannabinum montanum Heracleum sphondylium FamilyAlliaceae (Liliaceae) Species name Alliaceae (Liliaceae, Themidaceae) Alsinaceae (Caryophyllaceae) Alsinaceae (Caryophyllaceae) Alsinaceae (Caryophyllaceae) Amaranthaceae Amaranthaceae Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apocynaceae Apocynaceae Asclepiadaceae Apiaceae (Umbelliferae) Appendix B. Potential (unconfi

Chapter 6: Appendices 71 B). Native to Mediterranean. Known from CURE. Common name Life form Range Comments Siskiyou asterwestern aster PerFcutleaf Wide PerF Known from CURE nodding bur-marigold PerF Widestevia dusty-maiden Known from CURE Widechicory AnnF PerF Known from CURE Wide Wide Known from CURE Perf thistleParry’s Intro Native to Eurasia PerF Wide Known from CURE Caineville thistle PerF RegEn diffuse knapweeddiffuse PerF Intro CO state noxious weed (List Chamisso’s arnicaChamisso’s hairy arnica PerFbiennial wormwood Wide PerF PerF Known from CURE Intro Wide Known from CURE Known from CURE skeleton-leaf bursage PerFpearly-everlastingsmall-leaf pussyoes Wide PerF Known from CURE PerF Wide Wide

var.

var. 2006) biennis var. var. Flora of ( included var. var. A. ovata C. arizonicum A. hesperius, Symphyotrichum lanceolatum hesperium Symphyotrichum spathulatum calcareum; in bipinnatum North America Synonyms/ notes Taxonomic A. chamissonis andina Includes Some authors include A. rosea

ssp. ssp. rmed) taxa of Black Canyon the Gunnison National Park, cont. hesperius Aster spathulatusBahia dissecta A. occidentalis, Bidens cernua Chaenactis stevioides Amauriopsis dissecta Cichorium intybus Cirsium calcareum C. calcareum Cirsium parryi Acosta diffusa Centaurea diffusa foliosa Arnica mollis Artemisia biennisAster lanceolatus A. biennis Anaphalis margaritacea Antennaria microphylla Arnica chamissonis Ambrosia tomentosa Franseria tomentosa Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Appendix B. Potential (unconﬁ FamilyAsteraceae (Compositae) Species name Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae)

72 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Native to Eurasia. Known from CURE. collected just outside the park Wide Native to C North America PerF Intro Known from CURE Forb ower PerF Wide Known from CURE ower AnnF Wide Known from CURE ower owered neleaf woollywhite PerF RegEn Known from CURE Common name Life form Range Comments Nuttall’s sunfl Nuttall’s elk thistleorange sneezeweedpappose glandweed PerF PerF annual Wide Wide Known from CURE large-fl blanketfl pineapple-weedox-eye daisy AnnF Wide PerF Intro CO state noxious weed (List B). prairie sunfl hoary daisysmooth daisysmooth daisy PerFblueleaf aster PerF Wide PerF Wide PerF Known from CURE Wide Known from CURE Wide Known from CURE fi poverty-weedblue lettuce PerF Wide & Beck # 17063 (CURE) Weber PerF Wide Known from CURE ;

var.

fallax axillaris

nuttallii var. ssp. var. ssp. var. var. G. aristata Hymenoxys hoopesii scariosum G. pulchella Synonyms/ notes Taxonomic leucanthemum matricarioides, M. discoidea glabellus Eurybia glauca, Herrickia glauca glauca Hybrid of vars. not recognized in Flora of North America (2006). L. pulchella, oblongifolia, Mulgedium pulchellum

var.

ora lifolius var. grandifl x rmed) taxa of Black Canyon the Gunnison National Park, cont. Helianthus nuttallii H. nuttallii Dugaldia hoopesiiDyssodia papposa Helenium hoopesii, Leucanthemum vulgare Chrysanthemum Lepidotheca suaveolens Matricaria Helianthus petiolaris H. petiolaris Erigeron canus Erigeron glabellusErigeron leiomerus Eucephalus glaucus E. glabellus Gaillardia Aster glaucodes, Cirsium scariosum C. scariosum Iva axillaris Iva axillaris parvulus pulchella Hymenopappus fi Lactuca tatarica Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Appendix B. Potential (unconfi FamilyAsteraceae (Compositae) Species name Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae)

Chapter 6: Appendices 73 Native to Europe. Known from CURE. Known from CURE. AnnF Wide ower PerF RegEn Known from CURE ower PerF Wide Known from CURE Common name Life form Range Comments Canadian goldenrod PerF Wide Known from CURE Bigelow’s groundselBigelow’s desert- Torrey’s PerFdandelion RegEn Known from CURE streambank groundsel PerFsticky cudweed Wide Known from CURE prairie coneﬂ AnnF Wide scentless chamomile PerF Intro CO state noxious weed (List B). black-eyed Susan PerF Wide Native to E North America. montane coneﬂ saw-toothed groundsel PerF RegEn Known from CURE arrowleaf groundselbasin big sagebrush PerF Shrub Wide Known from CURE Wide cut-leaved groundsel PerFbroom groundsel Wide Known from CURE PerF Wide Known from CURE

var. var.

var.

(2006). var. Flora of var. var. var. vars. not ssp. pulcherrima var. Synonyms/ notes Taxonomic salebrosa, S. lepida salebrosa pseudaureus macounii G. macounii, P. hallii pseudoaurea, Senecio pseudaureus Tripleurospermum Tripleurospermum inodorum, perforata, T. maritima T. maritima R. montana tridentata spartioides; recognized in North America

var. hallii Senecio bigelovii var. var. admirabilis var. rmed) taxa of Black Canyon the Gunnison National Park, cont. Malacothrix torreyi Pseudognaphalium viscosum Gnaphalium viscosum, Ratibida columnifera Ligularia bigelovii Matricaria perforata M. maritima, Rudbeckia hirta R. hirta Packera pseudaurea pseudaurea P. Rudbeckia occidentalis montana Senecio spartioides S. spartioides Seriphidium tridentatum Artemisia tridentata Solidago canadensis S. canadensis Senecio eremophilus Senecio serra Senecio triangularis kingii Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Appendix B. Potential (unconﬁ FamilyAsteraceae (Compositae) Species name Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae)

74 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Montezuma Co., CO (Baker et al. 827) Common name Life form Range Comments golden crownbeardwater birch AnnFthick-sepaled cryptanth AnnF Widerecurved cryptanth Known from CURE showy stoneseed Wide Tree cryptanthBaker’s AnnF Known from CURE Wide PerF Periph PerF Known from CURE Wide RegEn Known from CURE locality is Mancos River, Type falcate asterrough cocklebur PerF AnnF Wide Wide stemless townsendia PerF Wide Known from CURE late goldenrodslender wirelettuce PerF PerF Wide Wide Known from CURE northern wintercress PerFhoary alyssumnodding rock cress Wide Known from CURE AnnF PerF Intro LocEn Native to Europe Known from CURE James’ cryptanth PerF Wide

var. (2006). (2003) Flora of Flora of falcatum var. minor, minor, vars. not jamesii vars. not var. var. var. exauriculata falcatus, Synonyms/ notes Taxonomic X. saccharatum, X. strumarium canadense, recognized in North America var. var. Symphyotrichum falcatum S. tenuifolia tenuifolia, recognized in North America cinerea Arabis demissa

var.

var. rmed) taxa of Black Canyon the Gunnison National Park, cont. Betula fontinalisCryptantha crassisepala Cryptantha recurvata Lithospermum incisum B. occidentalis Oreocarya bakeri Cryptantha bakeri elachantha Xanthium strumarium X. pensylvanicum, Ximenesia encelioides encelioides Verbesina Virgulus falcatus Aster falcatus Stephanomeria tenuifolia S. minor exscapa Townsendia Solidago gigantea Berteroa incana Boechera demissa demissa Oreocarya suffruticosaBarbarea orthoceras Cryptantha jamesii, C. Betulaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Appendix B. Potential (unconﬁ FamilyAsteraceae (Compositae) Species name Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Boraginaceae Brassicaceae (Cruciferae)

Chapter 6: Appendices 75 is from just outside the park. CURE. Native to Europe. Known from CURE. CURE. CURE. AnnF Wide Known from CURE PerF RegEn Known from CURE ax AnnF Intro & Rector 17308 (COLO) Weber eld pepperwort AnnF Intro Native to Asia. Known from Common name Life form Range Comments graceful false ﬂ watercress PerF Intro Native to Europe. Known from whitetop PerF Intro CO state noxious weed (List B). heartleaf bittercress PerF Wide Known from CURE ﬁ hairy whitetop PerF Intro Native to Asia. Known from hairy yellowcress PerF Wide Known from CURE slender-leaved tansy- slender-leaved mustard hare’s-ear mustardhare’s-ear AnnF Intro Native to Europe western tansy-mustard AnnFmountain draba Sparse splendid draba AnnFSyrian mustard RegEn PerFovalpurse Known from CURE AnnF tall Eastwood’s RegEnpepperwort Known from CURE Intro AnnF Native to Europe Sparse silver bladderpodAfrican mustard PerF AnnF Wide Intro Native to Africa

viscosa, hispida

var. viscosa, D. oxyloba var. leptophylla var. var. var. draba var. var. var. Synonyms/ notes Taxonomic aquaticum draba cordifolia richardsonii D. incisa D. pinnata osmiarum Includes vars. spectabilis Hutchinsia procumbens L. montanum spathulatum, L. eastwoodiae

viscosa D. incana hispida R. islandica ssp. var. ssp. var. cinalenasturtium- Rorippa rmed) taxa of Black Canyon the Gunnison National Park, cont. Camelina rumelica Nasturtium ofﬁ Cardaria draba Lepidium draba, C. Cardamine cordifolia C. cordifolia Neolepia campestre Lepidium campestre Cardaria pubescens Rorippa palustris Descurainia incisa Conringia orientalis Descurainia pinnata halictorum Draba rectifructa Draba spectabilis Euclidium syriacum Hymenolobus procumbens Hornungia procumbens, Lepidium alyssoides eastwoodiae Malcolmia africana Lesquerella ludoviciana Physaria ludoviciana Appendix B. Potential (unconﬁ FamilyBrassicaceae (Cruciferae) Species name (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae Brassicaceae (Cruciferae)

76 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park authors. Known from CURE CURE. CURE. CURE. Intro PerF Sparse Not included for CO by many AnnF Wide Known from CURE Forb y AnnF Wide eld pennycress AnnF Intro Native to Europe. Known from Common name Life form Range Comments pitseed goosefoot AnnF Wide Known from CURE southern marsh yellowcress spreading yellowcress PerF Wide Known from CURE entire-leaf thelypody PerFslate thelypodyﬁ Wide PerF Wide Parry’s harebellParry’s common hopsyellow beeplant PerFRocky Mountain PerFbeeplant Wide AnnF Known from CURE Wide Wide Known from CURE pincushion cactus PerF Wide sleepy catchﬂ Menzies’ campion PerF Wide Known from CURE black twinberry Shrub Wide Known from CURE common hornworttumbling orach PerF AnnF Wide Intro Native to Eurasia. Known from Russian pigweed annual summer-cypress AnnF Intro Native to Eurasia. Known from

var.

var. var.

var. (2003) Flora of

var. menziesii var. parryi var. var. var. ; varieties not Synonyms/ notes Taxonomic berlandieri integrifolium vivipara recognized in North America H. lupulus neomexicanus menziesii, Silene menziesii scoparia zschackei, C. album

orum ssp. rmed) taxa of Black Canyon the Gunnison National Park, cont. Rorippa teres Rorippa sinuata Thelypodium laxiﬂ Thlaspi arvense Thelypodium integrifolium integrifolium T. Humulus lupulus Cleome lutea Cleome serrulata americanus Coryphantha vivipara Escobaria vivipara Campanula parryi C. parryi Silene antirrhina Anotites menziesii A. menziesii Distegia involucrata Lonicera involucrata Ceratophyllum demersum Atriplex rosea Axyris amaranthoides Bassia sieversiana Kochia scoparia, B. Chenopodium berlandieri C. berlandieri Brassicaceae (Cruciferae) Appendix B. Potential (unconﬁ FamilyBrassicaceae (Cruciferae) Species name Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Cannabaceae Capparaceae (Cleomaceae) Capparaceae (Cleomaceae) Cactaceae Campanulaceae Caryophyllaceae Caryophyllaceae Caprifoliaceae Ceratophyllaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae

Chapter 6: Appendices 77 C. aurea. Wide Forb Common name Life form Range Comments golden sedge sedgeDewey’s PerGDouglas’ sedge PerG sedgeEmory’s Wide sedgeHasse’s Wide Known from CURE PerG Known from CURE PerG Wide PerG Wide Known from CURE Disj CO material may belong in water sedge PerG Wide Known from CURE common juniper Shrub Wide Known from CURE strawberry-blite annual Nebraska sedgePennsylvania sedge PerG PerGmeadow sedgenarrowleaf sedge Wide Wide Known from CURE Known from CURE PerG PerG Wide Wide Known from CURE Known from CURE oakleaf goosefootnarrow-leaf goosefoot AnnFslender Russian thistle AnnFparsley-fern AnnF Wide Wide Known from CURE Intro Known from CURE Native to Russia fern Wide Known from CURE

var. var. salinum, aquatilis var. var. var. by some

var. var. included in (sensu lato) beaked sedge PerG Wide Known from CURE heliophila var. ssp. Synonyms/ notes Taxonomic deweyana; C. bolanderi authors J. communis depressa capitatum C. pensylvanica digyna, C. heliophila, inops C. eleocharis, stenophylla, C. duriuscula C. salinum C. crispa acrostichoides

ssp. ssp. ssp. rmed) taxa of Black Canyon the Gunnison National Park, cont. Carex deweyanaCarex douglasii C. deweyana Carex emoryi Carex hassei Carex aurea C. garberi Carex aquatilis C. aquatilis alpina Chenopodium capitatum C. capitatum heliophila Carex pensylvanica Carex praticola Carex stenophylla Carex utriculata C. rostrata eleocharis Carex nebrascensis Chenopodium glaucumChenopodium leptophyllum C. glaucum Salsola collina Cryptogramma acrostichoides Juniperus communis Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Appendix B. Potential (unconﬁ FamilyChenopodiaceae Species name Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Cryptogrammaceae (Polypodiaceae, Adiantaceae) Cupressaceae

78 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Native to Europe. CURE. CURE. (Parry 32 ISC). Known from CURE. eld milkvetch PerF Wide Known from CURE Common name Life form Range Comments three-square bulrush PerG Wide panicled bulrushRussian-olive PerG Wide Tree Known from CURE Intro CO state noxious weed (List B). silver buffaloberryleafy spurge Shrubﬁ Wide milkvetchSheldon’s PerFGunnison milkvetch PerF Wide PerF RegEn LocEn CO BLM Sensitive. Known from two-grooved milkvetch PerFchickpea milkvetch Wide PerF Intro Native to Europe. Known from Drummond’s milkvetchDrummond’s PerF milkvetchHall’s Wide milkvetchHayden’s Known from CURE PerFgreat rushy milkvetch PerF PerF RegEn Wide Known from CURE Wide Known from CURE locality is ‘southern Utah’, Type skiff milkvetchskiff PerF LocEn CO BLM Sensitive

var.

var. var.

hallii var. var. var. var. polyphyllus, Synonyms/ notes Taxonomic var. Schoenoplectus pungens longispicatus, Schoenoplectus pungens polyphyllus uralensis bisulcatus haydenianus

var. rmed) taxa of Black Canyon the Gunnison National Park, cont. Schoenoplectus pungens Scirpus pungens Scirpus microcarpus Elaeagnus angustifolia Tithymalus uralensisTithymalus Astragalus agrestis Euphorbia esula Astragalus amphioxys A. dasyglottis Astragalus anisus vespertinus Shepherdia argentea Astragalus cicer Astragalus bisulcatus A. bisulcatus Astragalus halliiAstragalus haydenianusAstragalus lonchocarpus A. bisulcatus A. hallii Astragalus drummondii Astragalus microcymbus Appendix B. Potential (unconﬁ FamilyCyperaceae Species name Cyperaceae Elaeagnaceae Euphorbiaceae Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Elaeagnaceae Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae)

Chapter 6: Appendices 79 Canyon of the Grand River, Canyon of the Grand River, Grand Co., CO (Patterson s.n. GH) CURE. CURE. ower lupine PerF Wide Known from CURE Common name Life form Range Comments Brandegee’s corydalisBrandegee’s PerF LocEn locality is in Colorado Type Patterson’s milkvetchPatterson’s PerF RegEn locality is near Grand Type montane golden pea PerFAlsike clover Wide Known from CURE PerF Intro Native to Europe. Known from stinking milkvetch milkvetchWetherill’s PerF PerFSiberian pea-tree Wide RegEn Shrub USFWS: 3C, USFS R2 Sensitive Intro Native to Siberia. Known from King’s lupineKing’s AnnF Wide northern sweet-vetch PerFseemly sweetpea trefoilWright’s Wide PerF Wide PerFstemless lupine Known from CURE RegEnsmallﬂ Known from CURE PerF Wide cloverRusby’s PerF Wide Known from CURE var. var. T.

ovata

var. var. utahensis, var. var. var. kingii var. var. orus ; includes var. orus, L. argenteus pygmaeum parviﬂ longipes rusbyi, T. Synonyms/ notes Taxonomic montana rhombifolia eucosmus L. lepidus L. caespitosus laxiﬂ var. rhombifolia longipes brachypus, T. var. ssp. boreale utahensis

var.

ssp. ssp. orusargenteus L. rmed) taxa of Black Canyon the Gunnison National Park, cont. brandegeei Astragalus pattersonii Trifolium hybridum Trifolium Astragalus wetherillii Caragana arborescens Astragalus praelongus praelongus Lupinus lepidus longipesTrifolium longipes T. caespitosus Lathyrus eucosmusLotus wrightii L. brachycalyx Lupinus kingiiLupinus parviﬂ L. kingii Thermopsis montana pinetorum, T. T. Corydalis caseana Hedysarum boreale Hedysarum boreale Appendix B. Potential (unconﬁ FamilyFabaceae (Leguminosae) Species name Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fumariaceae Fabaceae (Leguminosae)

80 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park PerF Wide Known from CURE Common name Life form Range Comments Engelmann’s gentianEngelmann’s PerF Wide Tracy’s rushTracy’s PerG Wide Known from CURE wand phaceliabracken fern PerFRocky Mountain iris WideIdaho blue-eyed grass fern Known from CURE PerF PerFmountain blue-eyed grass Wide Wide Wide Known from CURE Known from CURE Known from CURE white scorpion-weed AnnF Wide Known from CURE Columbian monkshood PerFshale columbine Widecommon mare’s-tail Known from CURE Canadian waterweed PerFwestern waterweed PerF PerF waterleafFendler’s RegEn PerF Wide USFWS: 3C Sparse PerF Known from CURE Known from CURE Sparse Wide Known from CURE Rocky Mountain gentian PerF Widewhitestem gooseberry Known from CURE Shrubspiny swamp currant Wide ShrubSiberian water-milfoil Known from CURE PerF Wide Known from CURE Wide purple cluster crane’s-bill PerF Periph Known from CURE knotted rush PerG Wide Known from CURE

alba var. var.

fendleri & by some var. var. parryi and var. and var. nis var. var. J. ensifolius var. montanus nis Synonyms/ notes Taxonomic Included in var. heterosepala authors Includes ssp. heterophylla virgata aquilinum P. pubescens ochroleucum afﬁ Includes vars. columbianum Includes var. Includes var.

var. ssp. var. nisaffi Gentiana rmed) taxa of Black Canyon the Gunnison National Park, cont. Gentianella heterosepala G. amarella Pneumonanthe affi Pteridium aquilinum Iris missouriensis Sisyrinchium idahoense Sisyrinchium montanum lanuginosum occidentale Phacelia heterophylla Aquilegia barnebyi Hippuris vulgaris Elodea canadensis Elodea nuttallii Hydrophyllum fendleri Phacelia alba H. fendleri neomexicana P. Geranium caespitosum Ribes lacustre Myriophyllum sibiricum Aconitum columbianum caespitosum Ribes inerme Juncus tracyi Juncus nodosus Appendix B. Potential (unconfi FamilyGentianaceae Species name Gentianaceae Hypolepidaceae (Polypodiaceae, Dennstaedtiaceae) Iridaceae Iridaceae Iridaceae Hydrophyllaceae Helleboraceae (Ranunculaceae) Hippuridaceae Hydrocharitaceae Hydrocharitaceae Hydrophyllaceae Hydrophyllaceae Geraniaceae Grossulariaceae (Saxifragaceae) Haloragaceae Helleboraceae (Ranunculaceae) Grossulariaceae (Saxifragaceae) Juncaceae Juncaceae

Chapter 6: Appendices 81 PerF Wide PerF Wide Known from CURE PerG Wide PerF Wide Known from CURE ower willow- ower owered reweed PerF Wide Known from CURE white-fl herb Common name Life form Range Comments scarlet gaura PerF Wide fi New Mexico checker- mallow white checker-mallow PerF Wide Known from CURE hairy hedge-nettleivy-leaf duckweedcommon bladderwort PerFfalse mermaid PerF PerF stickleafRusby’s Wide Wide Wide Known from CURE AnnF Known from CURE PerF Wide Wide Known from CURE small-fl woodrush western evening- primrose splitlip hempnettleself-heal AnnFannual sagelittle skullcap Intro Known from CURE PerF PerF PerF Wide Wide Known from CURE Wide Known from CURE pale evening-primrose PerF Periph yellow evening-primrose PerF Wide Known from CURE

desert olive desert Shrub Wide

ssp.

crenulata candida var. ava fl var. var. var. orum orum, E. orum, ava canescens, canescens, C. neomexicana lactifl & circumvagum lactifl hornemannii Includes vars. glabrata Includes vars. Synonyms/ notes Taxonomic O. elata angustifolium var. C. angustifolium var. angustifolium lanceolata S. galericulata

var.

pilosa ssp. ssp. ssp. orumalpinum E. ora da avafl O. exa rmed) taxa of Black Canyon the Gunnison National Park, cont. hirsutissima Oenothera elata Gaura coccinea Epilobium lactifl Sidalcea neomexicana Lemna trisulca Utricularia macrorhizaFloerkea proserpinacoides Nuttallia rusbyi U. vulgaris Sidalcea candida Mentzelia rusbyi Forestiera pubescensChamerion danielsii neomexicana F. Epilobium Prunella vulgarisSalvia refl Scutellaria galericulata vulgaris P. Stachys palustris epilobifolia Galeopsis bifi Luzula parvifl Oenothera fl Oenothera pallida pallida Onagraceae Onagraceae Onagraceae Malvaceae Lemnaceae Lentibulariaceae Limnanthaceae Loasaceae Malvaceae Oleaceae Onagraceae Lamiaceae (Labiatae) Lamiaceae (Labiatae) Lamiaceae (Labiatae) Lamiaceae (Labiatae) Lamiaceae (Labiatae) Appendix B. Potential (unconfi FamilyJuncaceae Species name Onagraceae Onagraceae

82 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park CURE. CURE. CURE. PerF Wide Known from CURE Common name Life form Range Comments biennial evening- primrose slender bog orchid PerFclustered broomrapeEngelmann spruce PerF Widewoolly-base plantainEnglish plantain Known from CURE Tree PerF Widecrested wheatgrass Known from CURE Wide Wide PerF Known from CURE PerG Intro Intromeadow foxtail Native to Eurasia. Known from Native to Eurasia PerG Intro Native to Eurasia. Known from stinkgrass AnnG Intro Native to Eurasia. Known from desert saltgrass PerG Wide American sloughgrassbluejoint reedgrass AnnG PerG Widebrookgrass Known from CURE mat grama Widemeadow barley Known from CURE tufted hairgrass PerG PerG AnnG Wide PerG Wide Wide Known from CURE Known from CURE Wide Known from CURE northern mannagrass PerG Wide Known from CURE spike fescue PerG Wide fowl mannagrassAmerican mannagrass PerG PerG Wide Wide Known from CURE Known from CURE

& by

included stricta var. G. striata var. A. cristatum Synonyms/ notes Taxonomic O. strigosa, biennis P. gracilis, Habenaria P. saccata pectiniforme; within (without variety) in Flora of North America (2007) Includes vars. canadensis, imberbis, macouniana, C. scribneri brachyantherum Included in Includes G. elata fowl mannagrass PerG Wide Known from CURE some authors

ssp. rmed) taxa of Black Canyon the Gunnison National Park, cont. Oenothera villosa strigosa Aphyllon fasciculatumPicea engelmannii Plantago eriopoda Orobanche fasciculata Plantago lanceolata Alopecurus pratensis Limnorchis stricta Platanthera stricta, Eragrostis cilianensis Calamagrostis canadensis Catabrosa aquatica Chondrosum prostratumCritesion brachyantherum Bouteloua simplex Hordeum Deschampsia cespitosa Distichlis stricta D. spicata Agropyron pectiniforme A. cristatum Beckmannia syzigachne Glyceria elata Glyceria borealis Leucopoa kingii Hesperochloa kingii Glyceria grandis Glyceria striata Appendix B. Potential (unconﬁ FamilyOnagraceae Species name Orobanchaceae Pinaceae Plantaginaceae Plantaginaceae Poaceae (Gramineae) Orchidaceae Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae)

Chapter 6: Appendices 83 & Beck 17067 (BLCA) was collected just outside the park. AnnF Wide Known from CURE Common name Life form Range Comments tall wheatgrass PerG Intro Native to Eurasia. Weber alkali sacaton PerG Wide mountain muhlyspike muhlygreen needlegrass PerGfowl bluegrass alkaligrassNuttall’s PerG Wide PerG PerG Known from CURE Wide PerG Wide Known from CURE Wide Wide Known from CURE Known from CURE annual muhly AnnG Wide Known from CURE spike trisetumprickly-phlox PerG Wide Shrubblue leafy Jacob’s-ladder Known from CURE PerF Wide Known from CURE Wide Known from CURE cushion wild buckwheat PerFcream wild buckwheat Wide PerF Wide Known from CURE water smartweed PerF Wide Known from CURE Rocky Mountain phlox PerF Widenodding wild buckwheat Known from CURE

var. var. & var. var. (2005) airoides Flora of T. alaskanum depressa, patula

. var. var. & var. vars. not ora, pungens, Linanthus subalpinum; emersum, congdonii, Synonyms/ notes Taxonomic Elymus elongatus ponticus, Agropyron elongatum & montanum var. pungens Includes ssp. majus, E. umbellatum var. includes vars dichrocephalum desereticum var. Polygonum coccineum, Persicaria coccinea Includes ssp. multiﬂ foliosissimum cernuum, recognized in North America

var. ora rmed) taxa of Black Canyon the Gunnison National Park, cont. Thinopyrum ponticum Elymus elongatus, Muhlenbergia wrightii Nassella viridulaPoa palustris Puccinellia airoidesSporobolus airoides Stipa viridula nuttalliana P. S. airoides Muhlenbergia montana Muhlenbergia minutissima Leptodactylon pungens Leptodactylon pungens Phlox multiﬂ Eriogonum cernuum E. cernuum Eriogonum subalpinum E. umbellatum Persicaria amphibia Polygonum amphibium ovalifolium Trisetum spicatum Trisetum Polemonium foliosissimum foliosissimum P. Eriogonum ovalifolium Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Appendix B. Potential (unconﬁ FamilyPoaceae (Gramineae) Species name Polemoniaceae Polemoniaceae Polygonaceae Polygonaceae Polygonaceae Poaceae (Gramineae) Polemoniaceae Polygonaceae

84 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park CURE. CURE. Wide Forb PerF Wide Known from CURE PerF Wide Known from CURE owered Common name Life form Range Comments water smartweedwillow-weed PerF Wide Known from CURE AnnF Intro Native to Eurasia. Known from lady’s-thumb AnnF Intro Native to Eurasia. Known from western dockgolden dock PerF Wide annual Known from CURE fennel-leaf pondweed PerF Wide water-crowfoot PerFlittle buttercup Wide PerF RegEn Known from CURE dark-throat shooting- star green-ﬂ wintergreen thread water-buttercup PerF Wide

var.

. UT

by some

var. Included var Includes var. huberi ssp. & coriaceum, orum.

emersum eradicatus, B. circinatum Synonyms/ notes Taxonomic Polygonum amphibium var. lapathifolium R. occidentalis, R. aquaticus fenestratus R. fueginus “subvars.” from Flora: Coleogeton pectinatus ranunculinus pulchellum, D. pauciﬂ authors. atwoodii, trichophyllus. in diffusus, R. circinatus, diffusus, R. longirostris, R. trichophyllus var. B. longirostre, circinatum subrigidum trichophyllus

var. ssp. rmed) taxa of Black Canyon the Gunnison National Park, cont. Persicaria coccinea Polygonum coccineum, Persicaria lapathifolia Polygonum Persicaria maculosa Polygonum persicaria Rumex aquaticus occidentalis Rumex maritimus fueginus Potamogeton pectinatusDodecatheon pulchellum Stuckenia pectinata, D. pulchellum Cyrtorhyncha ranunculina Ranunculus Pyrola chlorantha virens P. Batrachium circinatumaquatilis Ranunculus Batrachium trichophyllum Ranunculus Appendix B. Potential (unconﬁ FamilyPolygonaceae Species name Polygonaceae Polygonaceae Polygonaceae Polygonaceae Potamogetonaceae Primulaceae Ranunculaceae Pyrolaceae (Ericaceae) Ranunculaceae Ranunculaceae

Chapter 6: Appendices 85 CURE. CURE. Native to Asia. Known from CURE. Common name Life form Range Comments shrubby cinquefoil Shrub Widemountain ninebark Known from CURE biennial cinquefoilred cinquefoil Shrub AnnFgraceful cinquefoil Wide Known from CURE Widewoolly cinquefoil PerF PerFPennsylvania cinquefoil Known from CURE PerFwestern thimbleberry Wide Disj PerFsweet-scented bedstraw Shrub Known from CURE PerF willowBebb’s Wide Wide willowBooth’s Wide Known from CURE Wide willowDrummond’s Known from CURE crack willow Known from CURE Shrub Shrub Shrub Wide Wide Wide Known from CURE Known from CURE Tree Known from CURE Intro Native to Eurasia. Known from Macoun’s buttercupMacoun’s creeping buttercup PerF PerF Wide Intro Known from CURE Native to Eurasia. Known from drab buttercup PerF Wide Known from CURE large-leaved avens PerF Wide Known from CURE Oriental clematis PerF Intro CO state noxious weed (List B). willow hawthornerect avens Shrub RegEn PerF Known from CURE Wide Known from CURE river hawthorn Shrub Wide Known from CURE rivularis hippiana repens

alpeophilus orus var. var. var. var. ssp. oribunda, D. oribunda, oribunda abelliformis Synonyms/ notes Taxonomic fl fl fl Includes var. Includes var. Pentaphylloides fruticosa, Dasiphora fruticosa

var.

ssp. var. oribundafruticosa, Potentilla orumparvifl Rubus abelliformisgracilis P. orum rmed) taxa of Black Canyon the Gunnison National Park, cont. Physocarpus monogynus Potentilla biennis Potentilla concinna Potentilla fl Potentilla hippiana Potentilla pensylvanica Rubacer parvifl Galium trifl hippiana P. Salix bebbiana Salix boothii Salix drummondiana Salix fragilis concinna Ranunculus repens R. repens Ranunculus macounii Ranunculus inamoenus inamoenus Pentaphylloides fl perincisum Viticella orientalis Clematis orientalis Geum aleppicum Geum macrophyllum Crataegus rivularisCrataegus saligna C. douglasii Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rubiaceae Salicaceae Salicaceae Salicaceae Salicaceae Ranunculaceae Ranunculaceae Appendix B. Potential (unconfi FamilyRanunculaceae Species name Rosaceae Ranunculaceae Rosaceae Rosaceae Rosaceae Rosaceae

86 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park may be adventive in CO. Known from CURE. PerF Sparse ower PerF Wide Known from CURE ower PerF Wide Known from CURE recracker PerF Periph ower Common name Life form Range Comments smallfl woodlandstar whiplash willowmountain willow willowScouler’s red alumroot Shrub Shrub Wide Shrub Wide Known from CURE Wide Known from CURE PerF Known from CURE Wide whiplash willow Shrub Wide Known from CURE Geyer’s willowGeyer’s strapleaf willow Shrub Shrub Wide Wide Known from CURE Known from CURE brook saxifragediamondleaf saxifrage coraldropsRitter’s PerF PerF paintbrushEastwood’s sulphur paintbrush PerF Wide PerF Wide Known from CURE PerF RegEn glabrous monkeyfl RegEn Known from CURE Wide yellow monkeyfl Eaton’s fi Eaton’s penstemonRydberg’s PerFthyme-leaved speedwell PerF WideVirginia ground-cherry Wide Known from CURE cut-leaf nightshade PerF Known from CURE AnnF Wide Known from CURE Wide Native to C North America,

caudata scabrida var. var. var. var. var. var. var. var. ora jamesii humifusa Synonyms/ notes Taxonomic parvifl ligulifolia geyeriana sulphurea, C. septentrionalis odontoloma fremontii, M. glabratus var. Veronica serpyllifolia, Veronica serpyllifolia Veronica var. rydbergii orum Lithophragma lasiandra caudata S. lasiandra orum ssp. ssp. rmed) taxa of Black Canyon the Gunnison National Park, cont. humifusum Salix monticola Salix scouleriana Heuchera rubescens Lithophragma parvifl Salix lucida Salix ligulifoliaSalix lucida S. eriocephala Salix geyeriana S. geyeriana Micranthes rhomboideaBesseya ritteriana Castilleja scabrida Saxifraga rhomboidea Castilleja sulphureaMimulus glabratus C. scabrida C. rhexifolia M. glabratus Micranthes odontoloma Saxifraga arguta, S. Mimulus guttatus Penstemon eatonii serpyllifolium Veronicastrum ssp. Physalis virginiana Solanum trifl Penstemon rydbergii rydbergii P. Salicaceae Salicaceae Saxifragaceae Saxifragaceae Salicaceae Salicaceae Salicaceae Appendix B. Potential (unconfi FamilySalicaceae Species name Saxifragaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Saxifragaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Solanaceae Solanaceae Scrophulariaceae

Chapter 6: Appendices 87 rmed Native to Eurasia. Known from CURE. ncluded here. trose County, Colorado, but have not previously been reported or conﬁ been reported Colorado, but have not previously County, trose ina Clark (2005) for Black Canyon of the Gunnison National Park’s NPSpecies ina Clark (2005) for Black Canyon of the Gunnison National Park’s Common name Life form Range Comments clasping twisted-stalk PerF Widetobacco-root Known from CURE Canadian white violet PerF PerF Widepuncture vine Known from CURE Wide Known from CURE AnnF Intro CO state noxious weed (List C).

by some included in & rugulosa, V. rugulosa, V. Synonyms/ notes Taxonomic Includes vars. amplexifolius, americanus, chalazatus. var. scopulorum; canadensis V. authors rmed) taxa of Black Canyon the Gunnison National Park, cont. Streptopus fassettii S. amplexifolius. Valeriana edulis Valeriana Viola rydbergii terrestris Tribulus canadensis V. Appendix B. Potential (unconﬁ FamilyUvulariaceae (Liliaceae, Calochortaceae) Species name Valerianaceae Violaceae Zygophyllaceae The vascular plant species shown in this table are known from the vicinity of Black Canyon Gunnison National Park in Mon known from The vascular plant species shown in this table are for the park. Species on this list are derived from a checklist of potential species developed by Tim Hogan, Nan Lederer, and D a checklist of potential species developed by Tim derived from Hogan, Nan Lederer, for the park. Species on this list are not i as potentially occurring in the park, but are Appendix C could be considered (1983). Some species from database and Weber

88 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park ed ed ed ed ed . orus. (Weber & Beck (Weber S. linifolius. E. triglochidiatus. C. viscidifl A. ludoviciana. P. splendens. P. C. rotundifolia. E. ciliatum . ed ed ed ed ed E. concinnus C. tracyi. C. planosus. Pseudostellaria jamesiana. Y. harrimaniae Y. ed specimen of ed specimen of outside the known range of this species in & Wittmann 2001). May Colorado (Weber be based on misidentifi outside the known range of this species in & Wittmann 2001). May Colorado (Weber be based on misidentifi specimens of specimens of outside the known range of this species in & Wittmann 2001). May Colorado (Weber be based on misidentifi habitat probably not present in park. May be based on misidentifi specimens of Comments specimens of specimen of 17012). misidentifi misidentifi & is not known from Colorado (Weber Wittmann 2001). outside the known range of this species in & Wittmann 2001). May Colorado (Weber be based on misidentifi Park status Range PerF Wide Ques? Reported by NCPN (in ed.), but BLCA is PerF Wide Ques? Reported by NCPN (in ed.), but BLCA is Shrub Periph Ques? Reported by NCPN (in ed.), but BLCA is well PerF WidePerF FalsRep Wide Reports from BLCA based on misidentifi Ques? Reported by NCPN (in ed.), but suitable Life Form PerF Wide FalsRep (1983) based on a Report by Weber eabane PerF Wide FalsRep Reports from BLCA based on misidentifi eld chickweed PerF Wide FalsRep Reports from BLCA based on misidentifi scarlet claret-cup cactus Arctic harebell PerF Wide FalsRep (1983) based on a Report by Weber shaggy fl slim-leaf plainsmustard wavy-leaved thistle PerF Wide FalsRep Reports from BLCA based on misidentifi Greene’s Greene’s rabbitbrush long-stalk spring- parsley Michaux’s wormwood New Mexico yucca Shrub RegEnfi FalsRep Report from BLCA based on misidentifi Scribner panicgrass PerG Wide Ques? Reported by NCPN (in ed.), but this species Common name Constance’s phacelia Constance’s PerF RegEnRocky Mountain Ques?willow-herb Reported by NCPN (in ed.), but BLCA is

var. var.

var. var. var. var. scribnerianum strictum pumilus Hesperidanthus linearifolius undulatum longipes C. longipes neomexicana ssp. Panicum oligosanthes var. Synonyms/ notes Taxonomic

ora var. Echinocereus coccineus Campanula uniﬂ Schoenocrambe linearifolia Erigeron pumilus E. pumilus Cirsium undulatum C. undulatum Chrysothamnus greenei Artemisia michauxiana Cymopterus longipes Yucca neomexicanaYucca harrimaniae Y. Cerastium strictum C. arvense, arvense Dichanthelium oligosanthes scribnerianum Phacelia constancei Epilobium saximontanum Cactaceae Campanulaceae Brassicaceae (Cruciferae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Apiaceae (Umbelliferae) FamilyAgavaceae Species name Alsinaceae (Caryophyllaceae) Poaceae (Gramineae) Hydrophyllaceae Onagraceae Appendix C. Rejected (falsely reported or questionable) vascular plant taxa of Black Canyon the Gunnison National Park. Appendix C. Rejected (falsely reported

Chapter 6: Appendices 89 e or . . aureum. P. fendleriana. P. O. pygmaea. P. norvegica P. latifolia. T. G. coloradense ed var. ed ed ed specimen of ed specimen of ed specimen of ed specimen of Comments outside the known range of this species in & Wittmann 2001). May Colorado (Weber be based on misidentifi misidentifi outside the known range of this species in & Wittmann 2001). May Colorado (Weber be based on misidentifi outside the known range of this species in & Wittmann 2001). Colorado (Weber misidentifi misidentifi misidentifi Park status and unpublished literature (NCPN in ed., Weber 1983) or from herbarium 1983) or from (NCPN in ed., Weber and unpublished literature Range Life Form PerF Periph Ques? Reported by NCPN (in ed.), but BLCA is arid wild buckwheatwild arid PerF WideNevada bitterroot Ques? PerF Reported by NCPN (in ed.), but BLCA is Wide FalsRep (1983) based on a Report by Weber skyline bluegrass PerG Wide Ques? Reported by NCPN (in ed.), but BLCA is Common name hybrid cattail PerG Wide FalsRep (1983) based on a Report by Weber brook cinquefoil AnnF Sparse FalsRep (1983) based on a Report by Weber rough bedstrawentire-leaved PerFpaintbrush Sparse FalsRep (1983) based on a Report by Weber x cations (False Reports or FalsRep) because the park is well outside known expected range of species (Questionabl by some P. T. latifolia T. Lewisia nevadensis fendleriana authors Included in Synonyms/ notes Taxonomic T. domingensis T. Sterile hybrid between var. var. var. var. glauca x asperrimum Eriogonum umbellatum subaridum Oreobroma nevadensis Poa cusickii epilis Typha Potentilla rivalis Galium mexicanum ssp. Castilleja integra Polygonaceae Portulacaceae FamilyPoaceae (Gramineae) Species name Typhaceae Rosaceae Rubiaceae Scrophulariaceae Appendix C. Rejected (falsely reported or questionable) taxa of Black Canyon the Gunnison National Park, cont. Ques?). The vascular plant species shown in this table have been reported for Black Canyon of the Gunnison National Park from published for Black Canyon of the Gunnison National Park from The vascular plant species shown in this table have been reported because of misidentifi now believed to be erroneous specimens, but are

90 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Rock crevices X X X X X Wet Comments r ﬁ Dg- X XX XXX Oak/ MtBr Jun Pin/ X XXX XXX Grs Sage/ Flower time Park status Native Pres Apr–Jun r Native Pres NA X X longleaf brickellbush Native Pres Aug–Sep California brickellbush Native Pres Aug–Sep Siberian elmRocky Mountain maple Native yuccaHarriman’s Introsquawbush Pres Native Pres Apr–Jun Pres Mar–Apr Native Apr–May X Pres X X X May–Jul X X X X Disturbed areas smooth sumacwestern poison ivyfringed sagebrush Native Native Pres Pres Native May–Jun Pres Jun–Jul Jul–Sep X X X X Rio Grande cottonwood Native Pres Mar–Jun quaking aspentree-of-heavennetleaf hackberry Native Intro Native Pres Pres Rep Mar–Jun Apr–May Jun–Jul X X X Disturbed areas box elder Native Pres Apr–Jun Utah juniperRocky Mountain juniper NativeGambel oak Presblue spruce Nativetwo-needle pinyon NAponderosa pine Pres Native NativeDouglas-ﬁ NA Native Pres Presnarrow-leaved Native Prescottonwood NA Apr–Jun Pres NA X X NA X X X X X X X X X X X X X X X

ssp.

ssp. var. ssp. Brickellia longifolia Brickellia californica Ulmus pumila Acer glabrum harrimaniae Yucca Rhus aromatica trilobata Rhus glabra Artemisia frigida Toxicodendron Toxicodendron rydbergii wislizenii Populus tremuloides Ailanthus altissima Celtis reticulata interius Sabina osteosperma Pseudotsuga menziesii Populus deltoides Sabina scopulorum Quercus gambelii Picea pungens Pinus edulis Pinus ponderosa scopulorum Populus angustifolia Negundo aceroides vascular plant taxa of Black Canyon the Gunnison National Park, organized by life form with rmed, historical, and reported Asteraceae (Compositae) Asteraceae (Compositae) Ulmaceae Shrubs Aceraceae Agavaceae Anacardiaceae Anacardiaceae Asteraceae (Compositae) Anacardiaceae Salicaceae Simaroubaceae Ulmaceae (Celtidaceae) Cupressaceae Pinaceae Salicaceae Cupressaceae Fagaceae Pinaceae Pinaceae Pinaceae Salicaceae FamilyTrees Aceraceae Species name Common name Nativity Appendix D. Conﬁ ecological notes.

Chapter 6: Appendices 91 Adobe Hills, Mancos Shale Rocky slopes s, cont. X Wet Comments r X ﬁ Dg- Oak/ MtBr Jun Pin/ XX XXX X XXX XX XXX XXX XX X XXX XXX X XXXX X XXX XX X X XXX Grs Sage/ Flower time Park status Native Pres Jul–Sep Native Rep Jul–Sep thinleaf alder Native Pres Apr–May Fremont’s mahoniaFremont’s Oregon-grape Native Rep Native Apr–Jul Pres X Apr–Jun X X X X X X X shortspine horsebrush Native Pres Apr–Jun spineless horsebrush Native Pres Jun–Sep mountain big sagebrush Wyoming big sagebrush black sagebrush Native Rep Sep–Oct broom snakeweed Native Pres Jul–Oct thread snakeweed Native Rep Jul–Oct green rabbitbrush Native Pres Jul–Sep Parry’s rabbitbrushParry’s Native Pres Jul–Sep glabrate rabbitbrush Native Pres Aug–Oct dwarf rabbitbrush Native Pres Jul–Sep Mohave brickellbush Native Pres Jun–Aug mountain snowberry Native Pres Jun–Aug red elderberry Native Pres Jun–Jul

ssp. ssp. ssp. ssp. orus orus parryi microbotrys tenuifolia Mahonia fremontii Mahonia repens Alnus incana Tetradymia spinosa Tetradymia Tetradymia canescens Tetradymia Seriphidium vaseyanum wyomingense Seriphidium tridentatum Seriphidium novum Gutierrezia sarothrae Gutierrezia microcephala viscidifl Chrysothamnus viscidifl Chrysothamnus parryi ssp. graveolens Chrysothamnus nauseosus Chrysothamnus depressus Brickellia oblongifolia Symphoricarpos rotundifolius Sambucus microbotrys var. rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Caprifoliaceae (Adoxaceae) Berberidaceae Berberidaceae Betulaceae Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) FamilyShrubs, cont. Asteraceae (Compositae) Species name Common name Nativity Caprifoliaceae Appendix D. Confi

92 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Rocky areas Rocky slopes Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale s, cont. Wet Comments r fi Dg- XX XX Oak/ MtBr XXX XXX X XX X X Jun Pin/ XXX X XXX XX XX X XX Grs Sage/ Flower time Park status littleleaf mock-orange Native Pres May–Sep trumpet gooseberry Native Pres Apr–Jun wolf currantFendlerbush Native PresUtah fendlerella Native May–Aug Pres Native Apr–Jun Rep Apr–Sep Wright’s fendlerbushWright’s Native Pres Apr–Jun Cutler’s ephedraCutler’s green ephedrabearberry Native Pres Native Pres NA Native NA Rep X May–Aug X X X wax currant Native Pres Apr–Aug greasewoodred-osier dogwoodspiny greasebush Native Native Pres Pres Native Pres May–Jul May–Aug Mar–Jun X X winterfat Native Pres May–Jul Gardner’s saltbushGardner’s Native Pres May–Aug mat saltbush Native Pres Apr–Sep mountain-loverfourwing saltbushshadscale Native Native Pres Pres May–Aug May–Sep Native X Pres May–Sep X X X X X Sandy soils singleleaf ash Native Pres Apr–May X X X X i coactilis Philadelphus microphyllus Ribes leptanthum Ribes wolfi Fendlera falcata Fendlerella utahensis Fendlera wrightii Ephedra cutleri Ephedra viridis Arctostaphylos uva-ursi ssp. Ribes cereum Sarcobatus vermiculatus Swida sericea Forsellesia meionandra Krascheninnikovia lanata Atriplex gardneri Atriplex corrugata Paxistima myrsinites Atriplex canescens Atriplex confertifolia Fraxinus anomala rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Hydrangeaceae (Saxifragaceae) Grossulariaceae (Saxifragaceae) Grossulariaceae (Saxifragaceae) Hydrangeaceae (Saxifragaceae) Hydrangeaceae (Saxifragaceae) Hydrangeaceae (Saxifragaceae) Ephedraceae Ephedraceae Ericaceae Grossulariaceae (Saxifragaceae) Chenopodiaceae (Sarcobataceae) Cornaceae Crossosomataceae (Celastraceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Appendix D. Confi FamilyShrubs, cont. Celastraceae Chenopodiaceae Chenopodiaceae Species name Common name Nativity Oleaceae

Chapter 6: Appendices 93 Adobe Hills, Mancos Shale Rocky slopes Adobe Hills, Mancos Shale s, cont. X X Wet Comments r ﬁ Dg- Oak/ MtBr X XXX Jun Pin/ X XXX XXX X XXX X XXX Grs Sage/ Flower time Park status Native Pres May–Jun Native Pres Jun–Oct ve-stamen tamarisk ve-stamen Intro Pres Apr–Sep X textile onionmouse-ear chickweed Introball-head sandwort Native Pres Native Rep Apr–Sep Pres May–Jun May–Aug X X coyote willowmatrimony-vineﬁ Native Intro oniontaper-tip Pres onionGeyer’s PresSan Juan onion Apr–Aug Native Mar–Oct Pres Native Native Pres May–Jul Pres May–Sep May–Jun X X X X X X X X X X Disturbed areas X common red raspberry Native Pres Jun–Aug squaw-apple Native Pres May–Jun antelope bitterbrush roseWoods’ NativeHimalayan blackberry Rep Intro Apr–Jul Native Rep Pres Jun–Sep X May–Jul X X X X X X Disturbed areas western chokecherry Native Pres May–Jul bush oceanspray Native Pres Jun–Aug X X X X Rock crevices Fendler’s mountain-lilacFendler’s Nativedeer-brush PresSaskatoon serviceberry NativeUtah serviceberry Jun–Aug mountain alder-leaf Presmahogany Native Native May–Jul Pres Pres X X Jun–Jul Apr–Jun X X X X X X X X X X X X X sand wild buckwheatlongleaf wild Nativebuckwheat Rep Jun–Oct X

ssp. ssp. Allium textile Eremogone congesta Cerastium fontanum Salix exigua Allium acuminatum Lycium barbarum Lycium ramosissima Tamarix Allium geyeri Allium macropetalum melanolasius Peraphyllum ramosissimum Purshia tridentata Rosa woodsii Rubus armeniacus Rubus idaeus melanocarpa Holodiscus discolor Padus virginiana Ceanothus fendleri Ceanothus velutinus Amelanchier alnifolia Amelanchier utahensis Cercocarpus montanus Eriogonum leptocladon Eriogonum lonchophyllum rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Alliaceae (Liliaceae) Alsinaceae (Caryophyllaceae) Alsinaceae (Caryophyllaceae) Salicaceae Forbs Perennial Alliaceae (Liliaceae) Solanaceae Tamaricaceae Alliaceae (Liliaceae) Alliaceae (Liliaceae) Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rhamnaceae Rhamnaceae Rosaceae Rosaceae Rosaceae FamilyShrubs, cont. Polygonaceae Polygonaceae Species name Common name Nativity Appendix D. Conﬁ

94 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Granite cliffs Disturbed areas s, cont. X Wet Comments r ﬁ Dg- XXX XXX XX X XX XX Oak/ MtBr X X Jun Pin/ XXXX X XXX X XXX XXX X X XXX XX X XXX XX Grs Sage/ Flower time Park status Native Pres Apr–Jun Native Pres Apr–May Native Pres Apr–Jun Native Pres May–Jul Native Pres Jul–Aug Lemmon’s spring- Lemmon’s parsley Chilean sweet-cicely Native Rep May–Jul blunt sweet-cicely Native Pres Jun–Jul western sweet-cicely Native Pres May–Jul ternate desert-parsley Native Pres May–Jul milfoil lomatium Native Pres Apr–May southern ligusticum Nativeadobe desert-parsley Pres Nativefern-leaved desert- parsley Pres Jun–Aug May Rocky Mountain spring- parsley Colorado Plateau spring-parsley onion spring-parsley Native Pres Apr–May Rocky Mountain Indian parsley James’ chickweed Native Pres Jun–Aug Fendler’s sandwortFendler’s sandwortHooker’s Native Rep Native May–Sep Rep Apr–Aug common dogbane Native Pres Jun–Aug platycarpum Pseudocymopterus montanus Osmorhiza chilensis Osmorhiza depauperata Osmorhiza occidentalis Lomatium triternatum var. Lomatium grayi Ligusticum porteri Lomatium concinnum Lomatium dissectum Cymopterus planosus Cymopterus purpureus Cymopterus bulbosus Aletes anisatus Pseudostellaria jamesiana Eremogone hookeri Eremogone fendleri Apocynum cannabinum rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Apiaceae (Umbelliferae) Alsinaceae (Caryophyllaceae) Alsinaceae (Caryophyllaceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Alsinaceae (Caryophyllaceae) Species name Common name Nativity Apocynaceae

Chapter 6: Appendices 95 Disturbed areas Disturbed areas Disturbed areas Disturbed areas Adobe Hills, Mancos Shale s, cont. X X Wet Comments r ﬁ Dg- XX Oak/ MtBr XX X XX X Jun Pin/ XX X XXX X XXX X XXX XXX XXX XX X XXXX XX Grs Sage/ Flower time Park status tall pussytoes Native Pres Jun–Aug rosy pussytoesburdock Nativeheartleaf arnica Pres Native Jun–Aug Intro Pres Pres May–Aug Jul–Sep common pussytoes Native Pres May–Jul low pussytoes Native Pres Apr–Jun alkali aster Native Rep May–Sep cutleaf agoseris Native Pres May–Sep short agoseris Native Pres May–Sep Russian knapweedorange agoseris Intro Pres Native Jun–Sep Pres Jun–Aug spotted knapweed Intro Pres Jul–Sep common yarrow Native Pres May–Sep pallid milkweedshowy milkweedwhorled milkweed Nativeasparagus Pres Native Native Pres Apr–Jun Pres May–Aug May–Sep Intro X Pres X Jun–Jul X X Disturbed areas Disturbed areas wormwoodCarruth’s Native Pres Aug–Sep

orus

var. var. cinalis ssp. aurantiaca anaphaloides Antennaria rosea Arnica cordifolia Arctium minus Antennaria pulcherrima Antennaria parvifolia Antennaria dimorpha Almutaster paucifl Agoseris glauca laciniata Agoseris glauca dasycephala Acroptilon repens Agoseris aurantiaca var. Acosta maculosa Achillea lanulosa Asclepias speciosa Asclepias cryptoceras Asclepias subverticillata Asparagus offi Artemisia carruthii rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Asteraceae Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asclepiadaceae FamilyPerennial Forbs, cont. Asclepiadaceae Species nameAsclepiadaceae Asparagaceae (Liliaceae) Common name Nativity (Compositae) Appendix D. Confi

96 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Disturbed areas Adobe Hills, Mancos Shale Disturbed areas Adobe Hills, Mancos Shale s, cont. X Wet Comments r X XX fi Dg- X Oak/ MtBr X XXX XX X Jun Pin/ XX X XXX XX XX X XXX XXX X XXX X X XXX X XXXX X XXX Grs Sage/ Flower time Park status Native Pres Jul–Oct eabane Native Pres May–Aug ower bull thistle Intro Pres Jul–Aug Tracy’s thistleTracy’s Native Pres Jun–Sep Rocky Mountain thistle Native Pres May–Jul New Mexico thistle Native Pres May–Jun Eaton’s thistleEaton’s Native Pres Jun–Sep fringed thistle Native Pres Jul–Aug hoary dusty-maiden Native Pres May–Sep Canada thistle Intro Pres Jul–Sep tassel-fl brickellbush musk thistle Intro Pres Jun–Sep Canada thistle Intro Pres Jul–Sep Canby’s asterCanby’s Native Rep Jul–Sep smooth asterarrowleaf balsamroot Native Native Pres Rep Apr–Jul Jul–Sep Louisiana wormwood Native Pres Jul–Oct Louisiana wormwood Native Pres Jul–Oct Navajo fl

ora var. geyeri var. incompta ludoviciana Erigeron concinnus Cirsium vulgare Cirsium tracyi Cirsium perplexans Cirsium neomexicanum Cirsium eatonii Cirsium centaureae Chaenactis douglasii Breea incana Brickellia grandiﬂ Carduus nutans macrolepis Breea arvensis Aster foliaceus Aster laevis Balsamorhiza sagittata Artemisia ludoviciana ssp. Artemisia ludoviciana ssp. rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Asteraceae (Compositae) Species name Common name Nativity Asteraceae (Compositae)

Chapter 6: Appendices 97 Adobe Hills, Mancos Shale Disturbed areas s, cont. X Wet Comments r X XX fi Dg- Oak/ MtBr XX X Jun Pin/ X XXX XX X XX XX XX XX XXX XXX X XXX XX X XXX X XXX X XXXX XX Grs Sage/ Flower time Park status ower Native Pres Jun–Aug eabane Native Rep Jul–Aug ve-nerve sunfl ve-nerve gumweed aster Native Rep May–Jul hoary aster Native Pres May–Sep bashful groundsel Native Pres Jul–Oct rose-heath Native Pres May–Sep hairy golden-aster Native Pres May–Sep Wyoming hyalineherb Native Pres May–Oct showy goldeneye Native Pres Jun–Sep fi curly-cup gumweed Native Pres Jul–Sep western goldenrod Native Pres Jul–Oct wandering fl aspen daisy Native Pres Jun–Aug trailing daisy Native Pres May–Aug Engelmann’s daisyEngelmann’s Native Rep May–Jul Eaton’s daisyEaton’s Native Pres May–Jul lacy tansy-aster Native Pres Apr–Sep nodding microseris Native Pres Apr–Aug spreading daisy Native Pres May–Sep

var. ora luteus ora agellaris var. da multifl lifolius Machaeranthera grindelioides Machaeranthera canescens Ligularia pudica Leucelene ericoides Heterotheca villosa Hymenopappus fi var. Heliomeris multifl Helianthella quinquenervis squarrosa Grindelia squarrosa Euthamia occidentalis Erigeron peregrinus Erigeron speciosus Erigeron fl Erigeron engelmannii Erigeron eatonii Machaeranthera pinnatifi Microseris nutans Erigeron divergens rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) FamilyPerennial Forbs, cont. Asteraceae (Compositae) Species name Common name Nativity Appendix D. Confi

98 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Disturbed areas s, cont. X Wet Comments r ﬁ Dg- Oak/ MtBr Jun Pin/ XX X XXXX XX X XXX XXX X XXX X X XXX XX X XXX XXX X XXX X XXX XXX X XXX XX XX Grs Sage/ Flower time Park status Native Pres May–Oct ower Native Pres Jul–Sep stemless hymenoxys Native Rep Apr–Jun common dandelion Intro Pres Apr–Dec brown-plume wirelettuce thrift goldenweed Native Pres May–Jun stemless goldenweed Native Rep May–Jul three-nerve goldenrod Native Pres Aug–Oct Wooton’s ragwortWooton’s Native Hist Apr–Aug western groundsel Native Pres Jun–Aug cutleaf coneﬂ western hawksbeard Native Pres May–Jul Modoc hawksbeard Native Pres May–Jul slender hawksbeard Native Pres May–Jul mountain hawksbeard Native Pres May–Jul rock-goldenrod Native Pres Jul–Sep New Mexico groundsel Native Pres Jun–Aug basin groundsel Native Pres Apr–Aug tarragon Native Pres Jul–Oct

cinale ssp. ora Tetraneuris acaulis Tetraneuris Taraxacum offi Taraxacum Stephanomeria paucifl Stenotus armerioides Stenotus acaulis Solidago velutina Senecio wootonii Senecio integerrimus Rudbeckia ampla Psilochenia occidentalis Psilochenia modocensis Psilochenia atribarba Psilochenia acuminata Petradoria pumila Packera neomexicana Packera multilobata glaucus Oligosporus dracunculus rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Appendix D. Confi FamilyPerennial Forbs, cont. Asteraceae (Compositae) Species name Common name Nativity

Chapter 6: Appendices 99 Adobe Hills, Mancos Shale Disturbed areas Adobe Hills, Mancos Shale s, cont. X Wet Comments r fi Dg- X Oak/ MtBr XXXX X XXX Jun Pin/ XXX X XXXX XXX X XXXX XX XX XXX XXX X XX X X Grs Sage/ Flower time Park status Intro Pres May–Jul Native Pres Apr–Jun c aster c Native Pres Jul–Oct European wintercress Intro Pres May–Aug showy stickseedwestern stoneseedaspen bluebell Native Native Pres Pres Native Jun–Aug Apr–Jun Pres X X Jun–Aug X X X X X X short-styled bluebellstreamside bluebell Nativeyellow-eye cryptanth Native Preshairy rockcress Native Pres Apr–Jun Pres Jun–Aug Native Apr–Jun X Rep X Jun–Aug X X X X X X common hound’s- tongue hybrid wyethia Native Pres May–Jul Cisco woodyaster Native Pres Apr–Jun Arizona mulesears Native Pres Jun–Aug mule’s-ears Native Pres May–Jun Pacifi yellow salsify Intro Pres May–Jul hoary townsendia Native Pres Apr–Jul Torrey’s hymenoxysTorrey’s Native Pres May–Jul spreadingpod rockcress Native Pres Jun–Jul Canyonlands hymenoxys Crandall’s rockcressCrandall’s Native Rep May–Jun cinale avoculata magna oribunda x grahamii major Barbarea vulgaris Boechera crandallii Hackelia fl Lithospermum ruderale Mertensia arizonica var. Mertensia brevistyla Mertensia ciliata Oreocarya fl Arabis hirsuta Cynoglossum offi Xylorhiza venusta Wyethia arizonica Wyethia Wyethia amplexicaulis Virgulaster ascendens Tragopogon dubius Tragopogon var. Townsendia incana Townsendia Tetraneuris torreyana Tetraneuris Boechera divaricarpa Tetraneuris ivesiana Tetraneuris rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Brassicaceae (Cruciferae) Boraginaceae Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Brassicaceae (Cruciferae) FamilyPerennial Forbs, cont. Asteraceae (Compositae) Species name Common name Nativity Appendix D. Confi

100 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Adobe Hills, Mancos Shale Disturbed areas Disturbed areas Disturbed areas Adobe Hills, Mancos Shale s, cont. X X Wet Comments r fi Dg- Oak/ MtBr XX XX Jun Pin/ XX X X XXX X X XXX X XXX X XXX XXX XX XX X XXXX XX XX Grs Sage/ Flower time Park status Native Pres May–Jun Native Pres May–Jun ower Native Rep Jun–Aug ower Native Pres Apr–Aug ower Native Rep Apr–Jul Jones’ pepperwort Native Pres Apr–Sep dame’s rocketdame’s Intro Pres Apr–Jun western wallfl shy wallfl worm-seed wallfl orbicular whitetop Intro Pres May–Aug Selby’s rockcressSelby’s Native Pres May–Aug Holboell’s rockcressHolboell’s Native Pres May–Jul perennial rockcress Native Pres Mar–Jul Gunnison County rockcress Glenwood Springs rockcress Sweetwater rockcress Native Pres May–Jul Gunnison’s rockcressGunnison’s Native Pres May–Jun Fendler’s rockcressFendler’s Native Rep Apr–Aug mountain pepperweed Native Pres Jun–Jul Drummond’s rockcressDrummond’s Native Pres May–Sep mountain pepperwort Native Rep Apr–Sep

ssp. jonesii tenellum montanum Lepidium montanum var. Lepidium montanum var. Hesperis matronalis Erysimum capitatum Erysimum inconspicuum Erysimum cheiranthoides altum Cardaria chalepensis Boechera selbyi Boechera retrofracta Boechera perennans Boechera pallidifolia Boechera oxylobula Boechera lignifera Boechera gunnisoniana Boechera fendleri Lepidium montanum var. Boechera drummondii rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Brassicaceae (Cruciferae) Species name Common name Nativity

Chapter 6: Appendices 101 Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Talus Adobe Hills, Mancos Shale s, cont. Wet Comments r fi Dg- Oak/ MtBr X X X Jun Pin/ X XXXX XXX X XXX XXX XXX X XXX XXX X XX X XXX XXX XXX XX X Grs Sage/ Flower time Park status Native Pres Apr–Jul Native Pres Apr–May Native Rep Apr–Aug Native Pres May–Jul ower Native Pres Apr–Jul ax-leaved berry pricklypear Native Pres Apr–Jul X X brittle prickly-pear Native Pres Jun–Jul claretcup Native Pres Apr–Jun tower-mustard Native Pres May–Sep heart-leaf jewelfl elegant thelypody Native Pres Apr–Jul prince’s-plume Native Pres Apr–Sep white prince’s-plume Native Pres May–Jun fl plainsmustard mesa twinpod Native Pres May–Jul Rollins’ twinpod Native Rep May–Jun sharp-leaf twinpod Native Pres May–Jul alpine pennycress Native Pres Apr–Aug Uncompaghre bladderpod Gunnison’s mariposaGunnison’s Native Pres Jun–Aug Colorado Plateau bladderpod plains prickly-pear hedgehog Simpson’s cactus Native Pres May–Jun X X X X

var. var. oribunda minor Opuntia phaeacantha brachyartha Opuntia fragilis Echinocereus triglochidiatus Turritis glabra Turritis Streptanthus cordatus Thelypodiopsis elegans Stanleya pinnata pinnata Stanleya albescens Schoenocrambe linifolia Physaria ﬂ Physaria rollinsii Physaria acutifolia Noccaea montana Lesquerella vicina Calochortus gunnisonii Lesquerella rectipes Opuntia polyacantha Pediocactus simpsonii var. rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Cactaceae Cactaceae Cactaceae Cactaceae Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Calochortaceae (Liliaceae) FamilyPerennial Forbs, cont. Brassicaceae (Cruciferae) Species name Common name Nativity Cactaceae Appendix D. Conﬁ

102 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale s, cont. Wet Comments r X fi Dg- Oak/ MtBr X X XXX Jun Pin/ X XXX XXX X XXX XXX X XXX X XXXX X XXX XXX X XXX XX Grs Sage/ Flower time Park status Native Pres Apr–Jul Native Pres Apr–Jun Native Pres Jun–Aug y Native Pres May–Sep eld bindweed Intro Pres Jun–Aug X X Disturbed areas great bladdery milkvetch straggling milkvetch Native Pres Apr–Jun Russian sickleyellow milkvetch Intro Native Rep Pres Jun–Aug Apr–Jul Disturbed areas noseburnmilkweed milkvetch Nativelesser rushy milkvetch Native Pres Native Pres May–Jun Rep May–Aug May–Aug X X Fendler’s spurgeFendler’s shorthorn spurge Native Native Rep Pres May–Sep May–Jun X X X fi lance-leaved stonecrop Native Pres Jun–Aug false Solomon’s-seal Native Pres Jun–Jul starry false Solomon’s- seal harebell catchfl Drummond’s Native Pres Jun–Sep egg milkvetch Native Pres May–Jul sego-lily Native Pres Jun–Jul Rydberg’s weedy Rydberg’s milkvetch

var. avus palans caulescens Astragalus miser Astragalus megacarpus Astragalus lentiginosus var. Astragalus falcatus Astragalus ﬂ Tragia ramosa Tragia Astragalus convallarius Astragalus asclepiadoides Chamaesyce fendleri Tithymalus brachyceras Tithymalus Convolvulus arvensis Amerosedum lanceolatum Maianthemum amplexicaule Maianthemum stellatum Campanula rotundifolia Gastrolychnis drummondii Astragalus oophorus var. Calochortus nuttallii oblongifolius rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Euphorbiaceae Fabaceae (Leguminosae) Fabaceae (Leguminosae) Euphorbiaceae Euphorbiaceae Convolvulaceae Crassulaceae Convallariaceae (Liliaceae, Ruscaceae) Convallariaceae (Liliaceae, Ruscaceae) Campanulaceae Caryophyllaceae Fabaceae (Leguminosae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Calochortaceae (Liliaceae) Species name Common name Nativity

Chapter 6: Appendices 103 Disturbed areas Disturbed areas Disturbed areas Disturbed areas x s, cont. X X Wet Comments r ﬁ Dg- XX Oak/ MtBr XX Jun Pin/ X XXX XX X XXX X X XXX X XXX X XXX X XXX XXX X XXXX X XXX XXX Grs Sage/ Flower time Park status Native Pres May–Aug ower sweetpea ower Native Pres May–Aug golden smokegolden corydalis Native Native Pres Pres Apr–Sep Apr–Sep X X X white cloverAmerican vetch Intro Native Pres Pres Apr–Sep May–Aug red clover Intro Rep May–Oct dwarf clover Native Pres Apr–Aug yellow sweet-clover Intro Pres May–Oct alfalfa Intro Pres May–Sep silky lupine Native Rep May–Aug meadow lupine Native Pres Jun–Aug Utah lupine Native Pres May–Aug whiteﬂ licorice-root Native Pres May–Jul silvery lupine Native Rep May–Aug Fort Wingate milkvetch Native Pres Apr–Jul pulse milkvetch Native Rep elkweed crane’s-billRichardson’s Native Pres Jun–Aug Native Pres Jun–Aug X X X X X X Rocky Mountain milkvetch

var. var. cinale occidentalis prunophilus Corydalis aurea Corydalis curvisiliqua ssp. Trifolium repens Trifolium Vicia americana americana Trifolium pratense Trifolium Trifolium gymnocarpon Trifolium Melilotus ofﬁ Medicago sativa Lupinus sericeus Lupinus polyphyllus var. Lupinus caudatus utahensis Lathyrus leucanthus Glycyrrhiza lepidota Lupinus argenteus Astragalus wingatanus Astragalus tenellus Frasera speciosa Geranium richardsonii Astragalus scopulorum rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Fumariaceae Fumariaceae Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Gentianaceae Geraniaceae FamilyPerennial Forbs, cont. Fabaceae (Leguminosae) Species name Common name Nativity Appendix D. Conﬁ

104 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Rocky slopes Disturbed areas Hanging garden s, cont. X X X X Wet Comments r X fi Dg- XXX Oak/ MtBr Jun Pin/ X XXX XX X XXX XX X XXX XXX XXX X XXX X XXX Grs Sage/ Flower time Park status Native Pres Jun–Aug ower ax Native Pres Apr–Aug X X X eld mint eld Native Pres Jun–Sep scarlet globe-mallow Native Pres Apr–Oct lesser duckweedleopard-lilyblue fl Nativewing-seed stickleaf Pres Native Native Jun–Oct Pres Pres Mar–Aug Apr–Jul X X X X X fi stinking horsemint Native Pres May–Aug common horehound Intro Pres May–Oct smallfl dragonhead nettle-leaf horsemint Native Pres Jun–Aug western St. John’s-wort Native Pres May–Sep mountain larkspur Native Rep Jun–Aug capitate waterleaf Native Pres Apr–Jul Nuttall’s larkspurNuttall’s Native Pres Apr–Jul red baneberry Native Pres May–Aug elegant columbinealcove columbine Native Pres Native Jun–Jul Pres May–Jun foothills death-camas Native Pres Mar–Jun

ssp. orum dissecta Sphaeralcea coccinea ssp. Lemna turionifera Nuttallia pterosperma Fritillaria atropurpurea Adenolinum lewisii Mentha arvensis Monardella odoratissima Marrubium vulgare Dracocephalum parviﬂ Agastache urticifolia Hypericum formosum Delphinium ramosum Hydrophyllum capitatum Delphinium nuttallianum arguta Aquilegia elegantula Aquilegia micrantha Actaea rubra Toxicoscordion paniculatum rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Melanthiaceae Malvaceae Lemnaceae Loasaceae Liliaceae Linaceae Lamiaceae (Labiatae) Lamiaceae (Labiatae) Lamiaceae (Labiatae) Lamiaceae (Labiatae) Lamiaceae (Labiatae) Hypericaceae (Guttiferae, Clusiaceae) Helleboraceae (Ranunculaceae) Hydrophyllaceae Helleboraceae (Ranunculaceae) Helleboraceae (Ranunculaceae) Helleboraceae (Ranunculaceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Helleboraceae (Ranunculaceae) Species name Common name Nativity (Liliaceae)

Chapter 6: Appendices 105 Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale s, cont. Wet Comments r ﬁ Dg- Oak/ MtBr X Jun Pin/ X XXX XX X XXXX X XXX X XXX X XXXX X XXXX X XXX Grs Sage/ Flower time Park status Native Pres Jun–Oct Native Pres Jun–Aug Native Pres Apr–Jun Native Pres May–Aug Native Pres Jun–Sep ower wild- ower redroot wild buckwheat desert phlox phloxHood’s long-leaf phloxsheep sorrel Native Native Native Pres Pres Pres Apr–Jun Intro Apr–Jun Apr–Jul Pres X X X Apr–Sep X X X X X X X X Disturbed areas yellow wild-buckwheat Native Pres Jun–Sep spotted coral-rootgiant helleborinenaked broomrape Nativecommon plantain Native RepBlack Canyon gilia Native Repsticky gilia Intro Jun–Jul Presscarlet gilia Native Apr–Aug Pres May–Jul Pres May–Aug Jun–Aug X Native Native Rep X Pres X May–Sep X May–Aug X X X X X Rock crevices hairy-throat evening- primrose northern willow-herb willow-herbHall’s Nativelong-tube evening- Presprimrose Native Jun–Aug Pres Jun–Aug X X narrow-leaf four- o’clock clay sand-verbena Native Pres May–Aug winged wild buckwheat Nativecurly dock Pres Jun–Oct Intro X Pres X May–Sep X Sandy soils X Disturbed areas meadow death camas Native Pres Apr–Jul sulphur-ﬂ buckwheat

ssp.

orum da var. avum aggregata marginata avum aureum Eriogonum umbellatum Eriogonum racemosum Phlox austromontana Phlox hoodii Phlox longifolia Acetosella vulgaris Eriogonum fl fl Corallorhiza maculata Epipactis gigantea Aphyllon unifl Plantago major Gilia penstemonoides Gilia pinnatifi Ipomopsis aggregata ssp. Oenothera coronopifolia Epilobium ciliatum Epilobium halleanum Oenothera caespitosa ssp. Oxybaphus linearis Abronia argillosa Pterogonum alatum Rumex crispus Toxicoscordion Toxicoscordion venenosum rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Polygonaceae Polygonaceae Polemoniaceae Polemoniaceae Polemoniaceae Polygonaceae Polygonaceae Orchidaceae Orchidaceae Orobanchaceae Plantaginaceae Polemoniaceae Polemoniaceae Polemoniaceae Onagraceae Onagraceae Onagraceae Onagraceae Nyctaginaceae Nyctaginaceae Polygonaceae Polygonaceae FamilyPerennial Forbs, cont. Melanthiaceae (Liliaceae) Species name Common name Nativity Appendix D. Confi

106 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Rock crevices Rock crevices s, cont. X X Wet Comments r XX fi Dg- XXX XXX Oak/ MtBr Jun Pin/ X XXXX X XXX X XXX Grs Sage/ Flower time Park status Native Rep May–Sep ax Native Pres Apr–Aug X X X X ower Native Pres Apr–Jun X beautiful cinquefoilColorado bedstraw Nativenorthern bedstraw Nativebastard toadfl Presspotted saxifrage Native Pres Jun–Auglittleleaf alumroot Presfringecup woodlandstar May–Aug Native Native Xslender woodlandstar Jun–Aug Native X Pres Pres Native Pres X Pres Jun–Sep Mar–Aug X May–Sep X X Apr–Aug X X X X X X X X X X X X X Rocky slopes face Cliff rock spiraea Native Pres Aug–Sep Virginia strawberry Native Pres Jun–Jul woodland strawberry Native Pres Jun–Jul silverweedtall cinquefoilprairie-smoke Native Native Native Pres Pres Pres May–Aug Jun–Jul Jun–Aug X X X sagebrush buttercup Native Pres Apr–Jun Purpus’ sullivantia Native Pres Jun–Jul willow docklanceleaf springbeauty Nativealpine bitter-rootColumbian virgin’s- Presbower Native Nativewhite virgin’s-bower Apr–Jul Pressugar-bowls Pres Nativemarsh buttercup May–Sep May–Aug Prespasquefl X X Jul–Aug Native X Native Pres X Pres X May–Sep X May–Aug X X X X X X

ssp.

ora

ssp. var. ora hirsutissima ora purpusii saximontana Sullivantia hapemanii var. Potentilla pulcherrima Galium coloradoense Galium septentrionale Comandra umbellata Ciliaria austromontana Heuchera parvifolia Lithophragma glabrum Lithophragma tenellum Petrophyton caespitosum glauca Fragaria virginiana bracteata Argentina anserina Drymocallis arguta Erythrocoma trifl Fragaria vesca Ranunculus glaberrimus ellipticus Clematis ligusticifolia Pulsatilla ludoviciana Rumex triangulivalvis Claytonia lanceolata Oreobroma pygmaea Atragene columbiana Corifl Halerpestes cymbalaria ssp. rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Saxifragaceae Rosaceae Rubiaceae Rubiaceae Santalaceae Saxifragaceae Saxifragaceae Saxifragaceae Saxifragaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Ranunculaceae Ranunculaceae Ranunculaceae Appendix D. Confi FamilyPerennial Forbs, cont. Polygonaceae Portulacaceae Portulacaceae Species nameRanunculaceae Common nameRanunculaceae Ranunculaceae Nativity

Chapter 6: Appendices 107 Rocky slopes Adobe Hills, Mancos Shale s, cont. Wet Comments r ﬁ Dg- XX X XXX XX Oak/ MtBr XXX X Jun Pin/ XXX X XXX Grs Sage/ Flower time Park status Native PresNative May–Jun Pres Jun–Aug gwort Native Pres Jun–Jul X X X rough-fruited fairy-bells Native Pres Jun–Aug mountain valerian Native Pres Jun–Jul prostrate vervainhook-spur violet violetShelton’s NativeCanada violet Native Pres Native Pres May–Sep Pres Native Mar–Aug X Pres Apr–Jun X Apr–Aug X X X X X X X Disturbed areas Watson’s penstemonWatson’s lance-leaf ﬁ Native Pres Jun–Aug X X X X woolly mulleinAmerican brooklimechain speedwell Native meadow-rueFendler’s Intro Pres Native Native Pres Pres Jul–Aug Pres Jun–Aug Jun–Aug Jun–Aug X X X X X Disturbed areas X stinging nettle Native Pres Jul–Aug desert paintbrushWyoming paintbrushscarlet paintbrush Native Nativemudwort Pres Presbutter-and-eggs Nativedwarf lousewort Jul–Aug Apr–Jun Rep Intro X Jul–Aug X Native Native Rep Pres Pres X X X Jun–Aug Apr–Jun Jun–Aug X X X X X X X X Disturbed areas Gray’s lousewortGray’s beardlip penstemon Native Nativemat penstemonAdobe Hills Pres Repbeardtongue Rocky Mountain Native Jun–Jul Jul–Augpenstemon Pres May–Jul X X X X

var.

ssp. torreyi Prosartes trachycarpa Valeriana capitata Valeriana acutiloba Verbena bracteata Verbena Viola adunca Viola biternata Viola scopulorum Penstemon watsonii Scrophularia lanceolata Verbascum thapsus Verbascum Veronica americana Veronica catenata Veronica Thalictrum fendleri gracilis Urtica gracilis Castilleja chromosa Castilleja linariifolia Castilleja miniata Limosella aquatica Linaria vulgaris Pedicularis centranthera Pedicularis procera Penstemon caespitosus Penstemon strictus Penstemon barbatus var. Penstemon retrorsus rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Uvulariaceae (Liliaceae, Calochortaceae) Valerianaceae Verbenaceae Violaceae Violaceae Violaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Thalictraceae (Ranunculaceae) Urticaceae FamilyPerennial Forbs, cont. Scrophulariaceae Scrophulariaceae Scrophulariaceae Species nameScrophulariaceae Scrophulariaceae Scrophulariaceae Common name Nativity Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Appendix D. Conﬁ

108 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Disturbed areas Disturbed areas Disturbed areas Adobe Hills, Mancos Shale Disturbed areas s, cont. X X X Wet Comments r fi Dg- XX Oak/ MtBr X XX Jun Pin/ XX XXX X X XXX Grs Sage/ Flower time Park status Native Pres May–Aug ower Native Pres Jun–Sep ower sand dune cryptanthslender cryptanth Nativewing-nut cryptanth Pres cryptanthWatson’s Native Nativecupseed stickseed May–Jul Pres Nativewestern stickseed PresEuropean stickseed Pres Native Apr–Jul X Apr–Julmeadow popcorn- Native Presfl Intro May–Jul X X Prespale madwort X Apr–Jun Rep X X Apr–Jun X X X Jun–Aug X Intro X X X X X X Rep X X X May–Jul X X X Disturbed areas Disturbed areas Disturbed areas mountain tarweeed Native Pres Jul–Sep slender goldenweed Native Rep May–Oct bachelor’s buttonbachelor’s Intro Rep Jun–Sep valley violetjuniper mistletoe Native Native Pres Pres Jul–Sep Mar–Jul Xprickly lettuce X Intro Pres Jul–Sep horned pondweedtumble pigweed NativeCanadian horseweed Pres Native Intro Mar–Nov Pres Presslender cudweed Jul–Sep Jun–Octcommon sunfl Native X Pres Jul–Oct X Disturbed areas marsh-elder Native Pres Aug–Oct penicillatus Cryptantha fendleri Alyssum alyssoides Cryptantha gracilis Cryptantha pterocarya Cryptantha watsonii Lappula marginata Lappula redowskii Lappula squarrosa Plagiobothrys scouleri ssp. Madia glomerata Machaeranthera gracilis Leucantha cyanus Lactuca serriola Viola vallicola Phoradendron juniperinum Zannichellia palustris Gnaphalium uliginosum Helianthus annuus Amaranthus albus Conyza canadensis Cyclachaena xanthifolia rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Boraginaceae Brassicaceae (Cruciferae) Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Asteraceae (Compositae) Appendix D. Confi FamilyPerennial Forbs, cont. Violaceae Viscaceae Species name Common name Nativity Zannichelliaceae Asteraceae (Compositae) Asteraceae (Compositae) Annual Forbs Amaranthaceae Asteraceae (Compositae) Asteraceae (Compositae)

Chapter 6: Appendices 109 Disturbed areas Adobe Hills, Mancos Shale Disturbed areas Disturbed areas Disturbed areas Disturbed areas Disturbed areas s, cont. Wet Comments r fi Dg- Oak/ MtBr XXX X Jun Pin/ XXX X XXX XXX XX XXX XX XX XXX XX XXX X XX XX X XX Grs Sage/ Flower time Park status Intro Pres Apr–Jun Native Pres Apr–Jul ax Intro Pres Apr–Jul ower Intro Pres Apr–Jun ixweed Intro Pres Mar–Aug branched pepperwort Native Pres Jun–Aug clasping pepperwort Intro Pres May–Jul prairie pepperwort Native Pres May–Jul spreading wallfl dwarf draba Native Pres Apr–Jul woodland whitlow- grass wedgeleaf draba Native Rep Feb–Jul fl mountain tansy- mustard blue mustardwestern tansy-mustard Native Intro Pres Pres Apr–Jul Apr–Jun Disturbed areas shepherd’s-purse Intro Pres Apr–Sep little-pod false fl juniper tumblemustard Native Pres May–Jun European madwort Intro Pres Apr–Jul tumble mustard Intro Pres May–Aug desert madwort Intro Pres Apr–Jul

ssp. ssp.

orum var. orum lipes Lepidium ramosissimum bourgeauanum Lepidium perfoliatum Lepidium densifl Erysimum repandum Draba reptans Draba nemorosa Draba cuneifolia Descurainia sophia incisa Chorispora tenella Descurainia incisa Descurainia incisa fi Capsella bursa-pastoris Camelina microcarpa Thelypodiopsis juniperorum Alyssum parvifl Sisymbrium altissimum Alyssum desertorum rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) Brassicaceae (Cruciferae) FamilyAnnual Forbs, cont. Brassicaceae (Cruciferae) Species name Common name Nativity Appendix D. Confi

110 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Disturbed areas Disturbed areas Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale s, cont. Wet Comments r fi Dg- Oak/ MtBr X X X X Jun Pin/ XX X XX XXX XX XX X X XXX XXX X XXX XX XX Grs Sage/ Flower time Park status Native Rep Apr–Jun owered common mallow Intro Pres Apr–Oct Disturbed areas Nevada stickleaf stickleafThompson’s Native Native Pres Pres Apr–Jun Apr–Aug X X X X white-stem blazingstar Native Pres Apr–Aug stork’s-bill phaceliaEastwood’s Native Pres Intro May–Jun Rep Feb–Oct X X X X Disturbed areas white sweet-clover Intro Pres May–Oct black medick Intro Pres May–Oct small-fl milkvetch Russian thistleridge-seeded spurge Native Intro Pres Pres Jun–Aug Jun–Sep Disturbed areas Nuttall’s poverty-weedNuttall’s Native Pres Apr–Jul halogeton Intro Pres Jul–Aug smallhead goosefoot goosefootFremont’s Native Native Rep Pres Jun–Sep Aug–Oct X X lambsquartermountain goosefoot Native Intro Pres Rep Jul–Sep Jun–Sep Palmer’s stinkweedPalmer’s orachPowell’s Native Pres Apr–Oct Native Pres Jun–Sep Paiute suncup Native Pres May–Jun micranthemiformis Malva neglecta Camissonia scapoidea Acrolasia dispersa Acrolasia humilis Acrolasia albicaulis Erodium cicutarium Phacelia splendens Melilotus albus Medicago lupulina Astragalus nuttallianus var. Salsola australis Chamaesyce glyptosperma Monolepis nuttalliana Halogeton glomeratus Chenopodium foliosum Chenopodium fremontii Chenopodium album Chenopodium atrovirens Atriplex powellii Cleomella palmeriana rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Malvaceae Onagraceae Loasaceae Loasaceae Loasaceae Geraniaceae Hydrophyllaceae Fabaceae (Leguminosae) Fabaceae (Leguminosae) Fabaceae (Leguminosae) Chenopodiaceae Euphorbiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Appendix D. Confi FamilyAnnual Forbs, cont. Capparaceae (Cleomaceae) Species name Common name Nativity

Chapter 6: Appendices 111 Disturbed areas Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale s, cont. Wet Comments r fi Dg- Oak/ MtBr X XX Jun Pin/ XX X XXX XXX XX X XXX XXX X XXXX Grs Sage/ Flower time Park status Native Pres May–Aug Native Pres May–Oct Native Pres May–Aug ower Native Pres May–Jul Xslopes Rocky ower rock- ower blister buttercuphorseshoe mousetailNorwegian cinquefoil Native Nativetwin-leaf bedstraw Intro Prescleavers Pres Native Pres Apr–May May–Sep Pres Jun–Aug X May–Aug Native X Pres X Apr–Jun X X X X X X X X X bur buttercup Intro Pres Apr–Jun western rock-jasminepygmy-fl Nativejasmine Pres Apr–Jun X X X X Kellogg’s knotweedKellogg’s Native Pres Jun–Sep yard knotweedDouglas’ knotweed Native Intro Pres Pres Jun–Oct May–Oct X X X X X X Disturbed areas slender phlox yellow navarretia wild Gordon’s buckwheat Native Native Pres Pres Mar–Jun May–Jul X X X X X X X X northern linanthus Native Pres May–Aug woolly plantain small collomiaeyelash gilia Native Rep Native Pres Native Mar–Aug Pres May–Aug X Apr–Jul X X X X X X branching groundsmoke diffuse groundsmokediffuse Native Pres May–Aug autumn willow-herb Native Pres Jul–Sep blue-eyed Maryseep monkeyfl Native Pres Apr–Jun X X X X

ora ssp. oribundus orum parvifl septentrionalis Hecatonia scelerata Myosurus cupulatus Potentilla norvegica Galium bifolium Galium spurium Collinsia parvifl Ceratocephala orthoceras Androsace occidentalis Androsace septentrionalis kelloggii Polygonum arenastrum Polygonum douglasii Polygonum polygaloides Microsteris gracilis Navarretia breweri Eriogonum gordonii Linanthus harknessii var. Plantago patagonica Collomia linearis Gilia ophthalmoides Gayophytum ramosissimum Gayophytum diffusum Gayophytum diffusum ssp. Epilobium brachycarpum Mimulus fl rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Ranunculaceae Ranunculaceae Rosaceae Rubiaceae Rubiaceae Scrophulariaceae Ranunculaceae Primulaceae Primulaceae Polygonaceae Polygonaceae Polygonaceae Polemoniaceae Polemoniaceae Polygonaceae Polemoniaceae Plantaginaceae Polemoniaceae Polemoniaceae Onagraceae Onagraceae FamilyAnnual Forbs, cont. Onagraceae Species name Common name Nativity Scrophulariaceae Appendix D. Confi

112 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park s, cont. X X X Wet Comments r ﬁ Dg- Oak/ MtBr Jun Pin/ X XXX Grs Sage/ Flower time Park status ower Native Pres Apr–Jun X X X X reddish monkeyﬂ Pennsylvania pellitory Native sedgeEggleston’s PresWhite Mountain sedge Nativeelk sedge Mar–Aug Nativewoolly sedge Pres Repsmall-wing sedge May–Augwestern sedge Jul–AugLiddon sedge Native Native Nativeblackcreeper sedge X PresRoss’ sedge Pres Pres Native Native Xvalley sedge Jun–Aug Native Pres May–Sep Apr–Juldryland sedge Pres X spikerush PresBolander’s Jun–Augcommon spikerush May–Aug Native Native X May–Jul Native X Native Preshardstem bulrush Pres Native X Pres X Pres May–Aug PresBaltic rush May–Jul X May–Jul Native Jul–Augperplexing rush X X May–Aug X Repslender rush Xlongstyle rush X X X X Jun–Aug X Nativemountain rush Native rushTorrey’s X X X Pres Pres NativeIndian ricegrass Native X X Jun–Aug Native X Pres May–Jul Pres Pres X Native Native Jun–Aug Jun–Aug Rep Jun–Aug Pres X Jun–Aug May–Jul X X X X X X X yellow owl-clovertwo-lobe speedwellpurslane speedwell Native Intro Pres Native Pres Pres Jul–Aug May–Jun Jun–Aug X X X X X X X Disturbed areas

var. ater ssp. acutus ssp. Parietaria pensylvanica Schoenoplectus lacustris Juncus arcticus Mimulus rubellus Carex egglestonii Carex geophila Carex geyeri Carex lanuginosa Carex microptera Carex occidentalis Carex petasata Carex praegracilis Carex rossii Carex vallicola Carex xerantica Eleocharis bolanderi Eleocharis macrostachya Juncus confusus Juncus dudleyi Juncus longistylis Juncus saximontanus Juncus torreyi Achnatherum hymenoides Orthocarpus luteus Pocilla biloba peregrina Veronica xalapensis rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Urticaceae Cyperaceae Juncaceae Appendix D. Conﬁ FamilyAnnual Forbs, cont. Scrophulariaceae Species name Common name Graminoids Perennial Cyperaceae Cyperaceae Nativity Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Poaceae (Gramineae) Scrophulariaceae Scrophulariaceae Scrophulariaceae

Chapter 6: Appendices 113 Disturbed areas Disturbed areas Disturbed areas Disturbed areas s, cont. X X X Wet Comments r X X XX ﬁ Dg- XX XXX Oak/ MtBr XXXX Jun Pin/ X XXX X XXX X XXX XX XXX X XXX XXX X XXX Grs Sage/ Flower time Park status Jones’ reedgrass Native Rep Jul–Aug slimstem reedgrass Native Pres Jul–Aug pine dropseed Native Pres Jul–Sep fringed bromesmooth brome Nativewoolly brome Pres Intronodding brome Jul–Sep Pres Native Native Pres Jun–Aug Pres Jun–Aug Jun–Aug ticklegrassshortawn foxtailpurple three-awn Native Native Pres Pres Native Pres Jul–Aug Jun–Aug Mar–Sep redtop Intro Pres Jun–Sep crested wheatgrass Intro Pres Jun–Aug clustered wheatgrass Intro Pres Jun–Aug Bloomer’s ricegrassBloomer’s Native Pres Jun–Aug pine needlegrass Native Pres Jun–Aug Nelson’s needlegrassNelson’s Native Pres Jun–Sep Letterman’s needlegrassLetterman’s Native Pres Jul–Sep

var. x Calamagrostis scopulorum Calamagrostis stricta Blepharoneuron tricholepis Bromopsis canadensis Bromopsis inermis Bromopsis lanatipes Bromopsis porteri Agrostis scabra Alopecurus aequalis Aristida purpurea longiseta Agrostis gigantea Agropyron mongolicum Agropyron desertorum Achnatherum bloomeri Achnatherum pinetorum Achnatherum nelsonii Achnatherum lettermanii rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) FamilyPerennial Graminoids, cont. Poaceae (Gramineae) Species name Common name Nativity Appendix D. Conﬁ

114 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Disturbed areas Disturbed areas Disturbed areas s, cont. X Wet Comments r X ﬁ Dg- XX XX XX XXX Oak/ MtBr Jun Pin/ X XXX XXX X XXXX XX X XXXX X XXXX XXX X XXXX XX X XX Grs Sage/ Flower time Park status Rocky Mountain fescue Native Pres Jul–Aug Thurber’s fescueThurber’s needle-and-thread Native Native Pres Pres Jul–Aug May–Jul Idaho fescue Native Pres May–Aug Arizona fescuetall fescue Native Rep Intro Jul–Aug Pres May–Jul quackgrass Intro Pres Jun–Aug slender wheatgrass Native Pres Jun–Aug bottlebrush squirreltail Native Pres May–Aug blue wildrye Native Rep Jul–Aug thickspike wheatgrass Native Pres Jun–Aug squirreltail Native Pres May–Aug Canada wildrye Native Pres Jul–Aug orchard grass Intro Pres Jun–Aug foxtail barley Native Pres Jun–Aug blue grama Native Pres Jul–Sep mountain brome Native Pres May–Aug Festuca saximontana Festuca thurberi Hesperostipa comata Festuca idahoensis Festuca arizonica Festuca arundinacea Elytrigia repens Elymus trachycaulus Elymus longifolius Elymus glaucus Elymus lanceolatus Elymus elymoides Elymus canadensis Dactylis glomerata Critesion jubatum Chondrosum gracile Ceratochloa carinata rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Appendix D. Conﬁ FamilyPerennial Graminoids, cont. Poaceae (Gramineae) Species name Common name Nativity

Chapter 6: Appendices 115 Disturbed areas Adobe Hills, Mancos Shale Adobe Hills, Mancos Shale s, cont. X X X Wet Comments r X XX XX X ﬁ Dg- X Oak/ MtBr Jun Pin/ XX X XXX X XXXX X XXX XXX XX X XXXX X XXX XX Grs Sage/ Flower time Park status interior bluegrass Native Pres Jun–Aug pale-leaf bluegrass Native Pres Jun–Aug Canada bluegrassmuttongrass Intro Pres Native Jun–Aug Pres May–Aug Agassiz’s bluegrassAgassiz’s Native Pres May–Aug bulbous bluegrass Intro Pres Apr–Jul reed canary grass Native Pres Jun–Aug timothylittleseed ricegrass Native Intro Pres Pres Jun–Aug Jun–Sep Porter’s melicPorter’s scratchgrassgreen muhly Native Preswestern wheatgrass Native Jul–Oct Native Pres Native Pres Pres Jul–Aug Jun–Aug Jul–Sep Salina wildrye Native Pres Jun–Jul Great Basin wildrye Native Pres Jun–Aug Junegrass Native Pres Jun–Aug galleta Native Pres May–Aug

ssp. Poa nemoralis interior Poa glaucifolia Poa compressa Poa fendleriana Poa agassizensis Poa bulbosa Phalaroides arundinacea Phleum pratense Piptatherum micranthum Melica porteri Muhlenbergia asperifolia Muhlenbergia racemosa Pascopyrum smithii Leymus salina Leymus cinereus Koeleria macrantha Hilaria jamesii rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) FamilyPerennial Graminoids, cont. Poaceae (Gramineae) Species name Common name Nativity Appendix D. Conﬁ

116 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park Rock crevices Rock crevices Rock crevices Adobe Hills, Mancos Shale Disturbed areas Disturbed areas Disturbed areas s, cont. Wet Comments r X ﬁ Dg- XXX XX Oak/ MtBr Jun Pin/ XX XX XX XX X XXX X XX X XXX X XXXX X XXXX Grs Sage/ Flower time Park status Native Pres NA Intro Pres Jun–Aug Intro Pres Jun–Aug eld horsetail eld Native Pres NA X X X X ﬁ southwestern bladder fern brittle bladder fern Native Pres NA male fern Native Pres NA Bigelow’s bluegrassBigelow’s Native Rep Mar–May false buffalograss Native Pres Jun–Sep annual wheatgrass Intro Pres May–Jul broad-leaved cattail Nativetoad rushcheatgrass Pres Jun–Aug Native Intro Pres Pres Jun–Aug May–Jul X X intermediate wheatgrass intermediate wheatgrass sand dropseed Native Pres Jun–Aug bluebunch wheatgrass Native Pres Jun–Aug Sandberg’s bluegrassSandberg’s Native Pres Apr–Sep Kentucky bluegrass Intro Pres May–Aug

ssp. ssp. lix-mas Equisetum arvense Cystopteris reevesiana Cystopteris fragilis Dryopteris ﬁ Poa bigelovii Monroa squarrosa Eremopyrum triticeum Typha latifolia Typha Juncus bufonius Anisantha tectorum intermedium Thinopyrum intermedium barbulatum Thinopyrum intermedium Sporobolus cryptandrus Pseudoroegneria spicata Poa secunda Poa pratensis rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Equisetaceae Aspleniaceae (Polypodiaceae) Aspleniaceae (Polypodiaceae) Ferns and Allies Aspidiaceae (Polypodiaceae, Aspleniaceae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Typhaceae Annual Graminoids Juncaceae Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Poaceae (Gramineae) Appendix D. Conﬁ FamilyPerennial Graminoids, cont. Poaceae (Gramineae) Species name Common name Nativity

Chapter 6: Appendices 117 Rock crevices Rock crevices s, cont. X Wet Comments r ﬁ Dg- Oak/ MtBr Jun Pin/ X XXX X XXX Grs Sage/ Flower time Park status Oregon woodsia Native Pres NA tall scouring-rushsmooth scouring-rush NativeFerriss’ horsetail Nativedense spike-moss Pres Presblunt-leaf spike-moss NA Native cloak-fernFendler’s NA Native Native Pres Pres Native Presslender lip-fern NA Pres NA NA NA Native X Pres X NA X X X X X X Rock crevices Rock crevices

ssp. ferrissii x nis afﬁ cathcartiana Woodsia oregana Woodsia Hippochaete laevigata Hippochaete hyemalis var. Hippochaete Selaginella densa Selaginella mutica Argyrochosma fendleri Cheilanthes feei rmed, historical, and reported taxa of Black Canyon the Gunnison National Park, organized by life form with ecological note Woodsiaceae Woodsiaceae (Polypodiaceae, Aspleniaceae) Equisetaceae FamilyFerns and Allies, cont. Equisetaceae Species nameEquisetaceae Selaginellaceae Selaginellaceae Common nameSinopteridaceae (Polypodiaceae, Adiantaceae) Nativity Sinopteridaceae (Polypodiaceae, Adiantaceae) Appendix D. Conﬁ

118 Annotated Checklist of Vascular Flora: Black Canyon of the Gunnison National Park The Department of the Interior protects and manages the nation’s natural resources and cultural heritage; provides scientiﬁ c and other information about those resources; and honors its special responsibilities to American Indians, Alaska Natives, and afﬁ liated Island Communities.

NPS 144/100146, July 2009 National Park Service U.S. Department of the Interior

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