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Вопросы Ихтиологии Том 59 № 4 2019 Июль–Август Основан В 1953 Г

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Вопросы Ихтиологии Том 59 № 4 2019 Июль–Август Основан В 1953 Г Российская академия наук ВОПРОСЫ ИХТИОЛОГИИ Том 59 № 4 2019 Июль–Август Основан в 1953 г. Выходит 6 раз в год ISSN: 0042-8752 Журнал издается под руководством Отделения биологических наук РАН Редакционная коллегия: Главный редактор Д.С. Павлов А.М. Орлов (ответственный секретарь), С.А. Евсеенко (заместитель главного редактора), М.В. Мина (заместитель главного редактора), М.И. Шатуновский (заместитель главного редактора), О.Н. Маслова (научный редактор) Редакционный совет: П.-А. Амундсен (Норвегия), Д.А. Астахов, А.В. Балушкин, А.Е. Бобырев, Й. Вайценбок (Австрия), Ю.Ю. Дгебуадзе, А.В. Долгов, М. Докер (Канада), А.О. Касумян, Б. Коллетт (США), А.Н. Котляр, К.В. Кузищин, Е.В. Микодина, В.Н. Михеев, П. Моллер (США), А.Д. Мочек, Д.А. Павлов, Ю.С. Решетников, А.М. Токранов, В.П. Шунтов Зав. редакцией М.С. Чечёта E-mail: [email protected] Адрес редакции: 119071 Москва, Ленинский проспект, д. 33 Телефон: 495-958-12-60 Журнал “Вопросы ихтиологии” реферируется в Реферативном журнале ВИНИТИ, Russian Science Citation Index (Clarivate Analytics) Москва ООО «ИКЦ «АКАДЕМКНИГА» Оригинал-макет подготовлен ООО «ИКЦ «АКАДЕМКНИГА» © Российская академия наук, 2019 © Редколлегия журнала “Вопросы ихтиологии” (составитель), 2019 1 Подписано к печати 28.12.2018 г. Дата выхода в свет 15.03.2019 г. Формат 60 × 88 /8 Усл. печ. л. 15.5 Тираж 24 экз. Зак.2047 Бесплатно Учредитель: Российская академия наук Свидетельство о регистрации средства массовой информации ПИ № ФС77-66712 от 28 июля 2016 г., выдано Федеральной службой по надзору в сфере связи, информационных технологий и массовых коммуникаций (Роскомнадзор) Издатель: Российская академия наук, 119991 Москва, Ленинский пр., 14 Исполнитель по госконтракту № 4У-ЭА-197-18 ООО «ИКЦ «АКАДЕМКНИГА», 109028 Москва, Подкопаевский пер., 5, мезонин 1, к. 2 16+ Отпечатано в типографии «Book Jet» (ИП Коняхин А.В.), 390005, г. Рязань, ул. Пушкина, 18, тел. (4912) 466-151 СОДЕРЖАНИЕ Том 59, Номер 4, 2019 Материалы к ревизии рода Photonectes (Melanostomiidae) А. М. Прокофьев 375 Видовой состав и распределение донных рыб в элиторали северо-западной части Охотского моря в летний период П. А. Савельев, Е. А. Метелёв, А. С. Сергеев, В. С. Данилов 405 Видовой состав, распределение и некоторые аспекты биологии рыб рода Lycodes (Zoarcidae) в море Лаптевых в 2014 году Е. В. Смирнова, О. В. Карамушко, Н. В. Чернова 416 Современная структура ихтиоцена внешнего края шельфа и верхней части склона северных Курильских островов и Юго-Восточной Камчатки А. Б. Савин, А. А. Баланов, В. В. Панченко 428 К вопросу о становлении жизненных стратегий у тихоокеанских лососей Oncorhynchus (Salmonidae) М. К. Глубоковский, С. Л. Марченко 429 Репродуктивные стратегии и происхождение парапатрических и симпатрических форм арктического гольца Salvelinus alpinus (Salmonidae) в системе озёр Большое и Малое Леприндо (Северное Забайкалье) С. С. Алексеев, М. Ю. Пичугин, Н. В. Гордеева, В. П. Самусенок, А. Л. Юрьев, В. С. Хлыстов, А. Н. Матвеев 430 Репродуктивная биология карликового тетрадона Carinotetraodon travancoricus (Tetraodontidae) К. М. Анупама, Х. С. Хари Санкар, М. Ритин Радж, М. Харикришнан 448 Популяционно-генетическая организация тихоокеанской трески Gadus macrocephalus в Северной Пацифике на основе микросателлитного анализа М. А. Смирнова, С. Ю. Орлова, А. М. Орлов 449 Первая массовая поимка неалота Nealotus tripes (Gempylidae) М. В. Назаркин, О. А. Мазникова, А. М. Орлов 450 Возраст и рост обыкновенной султанки Mullus barbatus в северной части Эгейского моря С. Тюзюн, К. Далян, Л. Эрийльмаз 451 О некоторых вопросах динамики численности темпоральных форм горбуши Oncorhynchus gorbuscha на Сахалине и южных Курильских островах А. М. Каев 452 Оценка обилия донных рыб на шельфе и материковом склоне Северного Приморья Д. Г. Кравченко, Д. В. Измятинский 461 Первое обнаружение и биология моллинезии Poecilia sphenops (Poeciliidae) из Северной Сахары Алжира К. Гази, А. Си Бахир, Т. Иддер 472 Вкусовая привлекательность желчных веществ для рыб А. О. Касумян, М. И. Виноградская 473 КРАТКИЕ СООБЩЕНИЯ Первая находка редкошипого круглопёра Eumicrotremus gyrinops (Cyclopteridae, Cottoidei) от тихоокеанского побережья острова Парамушир О. С. Воскобойникова 483 О новых случаях регистрации гренландской полярной акулы Somniosus microcephalus в Карском море М. В. Бородавкина, Н. В. Чернова, Н. А. Чекменева 487 Микропластик в пище балтийской песчанки Ammodytes tobianus литорали Куршской косы Балтийского моря А. В. Гущин, Я. В. Веремейчик 492 ВОПРОСЫ ИХТИОЛОГИИ, 2019, том 59, № 4, с. 375–404 УДК 597.5 МАТЕРИАЛЫ К РЕВИЗИИ РОДА PHOTONECTES (MELANOSTOMIIDAE) © 2019 г. А. М. Прокофьев1, 2, * 1Институт проблем экологии и эволюции им. А.Н. Северцова РАН – ИПЭЭ, Москва, Россия 2Институт океанологии им. П.П. Ширшова РАН – ИО РАН, Москва, Россия *E-mail: [email protected] Поступила в редакцию 06.10.2018 г. После доработки 07.02.2019 г. Принята к публикации 07.02.2019 г. Критически проанализированы признаки внешней морфологии меланостомовых рыб рода Pho- tonectes. Показано, что признаки, ранее применявшиеся для разграничения подродов, непригодны для этих целей. Подтверждено существование в составе рода только двух чётко диагностируемых подродов, Trachinostomias и Photonectes s. str., для которых даны ревизованные диагнозы на основа- нии нового набора признаков. Вид P. uncinatus перемещен из подрода Trachinostomias в номинатив- ный подрод. Кратко обсуждены основания для выделения групп видов в составе номинативного подрода и филогенетические отношения между этими группами. Описано три новых вида: P. disti- chodon, P. filipendulus и P. xenopogon spp. n. Указаны отличительные признаки ещё одного нового вида, из-за недостатка данных остающегося пока не описанным. Дана таблица для определения всех из- вестных видов рода. Ключевые слова: меланостомовые, Photonectes, внешняя морфология, классификация, подродовой состав, новые виды. DOI: 10.1134/S0042875219040167 Род Photonectes Günther, 1887 является наиме- зывалось наличие удлинённого заднего отростка нее разработанной в таксономическом отноше- anguloarticulare, удлинённое hypohyale и наличие нии группой меланостомовых рыб, видовое раз- плотного соединительнотканного хрящевого об- нообразие которой в настоящее время остаётся в разования между коракоскапулярной пластин- значительной мере недооценённым, а принципы кой и cleithrum (Fink, 1985). Нужно, однако, отме- видовой диагностики и надвидовой классифика- тить, что Финк (Fink, 1985) изучил только немно- ции – недостаточно разработанными. Между тем гие виды рода и с разной степенью детализации, наряду с Bathophilus Giglioli, 1882 это, вероятно, более того, типовой вид рода (P. albipennis) остал- одна из наиболее диверсифицированных групп ся им неисследованным, в связи с чем монофи- меланостомовых рыб, по-видимому, уступающая лию рода нельзя считать доказанной. Сам Финк по числу видов лишь гипердиверсифицированно- (Fink, 1985) отмечал, что существенные остеоло- му роду Eustomias Vaillant, 1888. С учётом настоя- гические различия были выявлены им даже меж- щей публикации в составе рода содержится 34 но- ду экземплярами, считающимися конспецифич- минальных вида, из которых 24 или 25 являются ными (что не удивляет, так как, судя по списку валидными (табл. 1). Кроме них мне известны материала, речь идет о P. margarita, под именем ещё по меньшей мере четыре неописанных вида. которого ранее смешивалось не менее четырёх Внешне виды рода Photonectes отличаются от разных видов). Несомненно, что полноценный других меланостомовых рыб длинной изогнутой морфологический анализ всех представителей и выступающей вперед нижней челюстью, сим- рода приведёт к значительным изменениям суще- физ которой помещается в широкий промежуток ствующих представлений о морфологической из- между симфизными концами praemaxillaria. По- менчивости и классификации данного рода. добное строение нижней челюсти отмечено еще В настоящее время проведение такого анализа только у рода Tactostoma Bolin, 1939, отличающе- затруднительно, так как ряд видов исключитель- гося от Photonectes намного более удлинённым те- но редки в коллекциях и рассеяны по многим му- лом, многорядными зубами в челюстях и рядом зеям мира, а некоторые из них известны только остеологических особенностей, свидетельствую- по голотипам; ряд видов известен только по юве- щих против их близкого родства (Fink, 1985). В нильным стадиям; кроме того, вследствие осо- качестве апоморфных признаков Photonectes ука- бенностей строения кожи и подкожного соедини- 375 376 ПРОКОФЬЕВ Таблица 1. Список номинальных и валидных видов рода Photonectes Валидное Номинальный вид Примечание название Подрод Trachinostomias Eustomias aguayoi Barbour, 1942 P. margarita P. bifilifer Beebe, 1933 P. parvimanus P. f ilipendulus sp. nova P. filipendulus P. fimbria Regan et Trewavas, 1930 P. parvimanus P. flagellatus Parr, 1927 P. margarita P. (Trachinostomias) gorodinskii P. gorodinskii Отличия от P. parvimanus нуждаются в уточнении на более представительном материале P. intermedius Parr, 1927 P. margarita Под названием “intermedius” описана молодь P. margarita Echiostoma margarita Goode et Bean, 1896 P. margarita Возможно, сборный вид в настоящем понимании P. monodactylus Regan et Trewavas, 1930 P. margarita P. munif icus Gibbs, 1968 P. munificus Родственные отношения с другими видами рода нуждаются в уточнении P. parvimanus Regan et Trewavas, 1930 P. parvimanus Echiostoma richardi Zugmayer, 1913 P. margarita P. xenopogon sp. nova P. xenopogon Подрод Photonectes s. str. P. achirus Regan et Trewavas, 1930 P. achirus Группа “caerulescens”; наибольший известный экземпляр
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