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Fokienia Hodginsii (Dunn) A Fokienia A. Henry & H. H. Thomas, Gard. Chron., ser. 3, 49: 67. 1911. Type: Fokienia hodginsii (Dunn) A. Henry & H. H. Thomas [Cupressus hodginsii Dunn] (Cupressaceae). Fokien is an earlier European spelling for Fujian, a on flattened ultimate branchlets but increasingly less province of China. so on older (whip) shoots, becoming long decur- rent to 15 mm, with facials usually smaller and/or Description narrower than the laterals, green or glaucous green. Facial leaves appressed, obtuse, variable in size from 384 See the species description. 2–7 × 1–2 mm on ultimate branchlets; stomata in two bands separated by a midrib. Lateral leaves lon- Distribution ger than facials or nearly equal in length, bilaterally flattened, variable, 4–10 × 2–3(–4) mm on ultimate As for the species. branchlets; apex incurved, free to appressed, acute to obtuse; stomata in a single whitish band, eglandular. Pollen cones terminal, 4–5 × 1.5–2 mm, initially sub- Fokienia hodginsii (Dunn) A. Henry & globose but elongating when maturing, with 10–12 H. H. Thomas, Gard. Chron., ser. 3, 49: 66–68, decussate, broadly ovate-acute microsporophylls f. 32–35. 1911. Cupressus hodginsii Dunn, J. Linn. each bearing 3 small, abaxial pollen sacs. Seed cones Soc., Bot. 38: 367. 1908; Chamaecyparis hodginsii terminal, maturing in the second year, caducous, to (Dunn) Rushforth, J. Biol. (Viet Nam) 29 (3): 38. 15–25 × 12–22 mm, subglobose, brown. Bract-scale 2007. Type: China: Fujian, Min Jiang, Nanping, complexes (10–)12–16, decussate, spreading at matu- [“Yenping”], A. E. N. Hodgins HK 3505 (holotype rity, to 10 × 16 mm (uppermost 2–4 much smaller and K). Pl. 14, Fig. 127 sterile), cuneate-peltate, flattened, woody, with irreg- ular margins, distally with a central depression and a Etymology small mucronate umbo (bract tip), dull brown, with striated anterior part, orange- to reddish brown with The species epithet is after A. E. N. Hodgins, who whitish seed marks. Seeds 20–30 per cone, 4–5 mm collected it in 1905. long, ovoid and 3–4 ridged, with acute apex and trun- cate base (hilum), brown, wings 2, very unequal in Vernacular names size and shape; larger wing ca. 6 × 5 mm, smaller one ca. 1.5 mm, or a mere strip near the seed apex. Fujian Cypress; fu jian bai (Chinese) Taxonomic notes Description Some authors have placed this species either in Trees to 30 m tall, evergreen, monoecious; trunk Cupressus or (recently) in Chamaecyparis based on monopodial, to 1 m d.b.h. Bark on trunk nearly comparisons of (some) gross-morphological char- smooth, becoming irregularly fissured and flak- acters. Detailed analyses of the ontogeny of the seed ing, purplish brown. Branches long, spreading or cones (Jagel, 2001; Farjon, 2005a) and phylogenetic down-curved, self-pruning lower down the trunk, analyses based on morphology (Farjon, 2005a) as forming a broadly pyramidal, in old trees rounded well as DNA sequence data (Gadek & Quinn, 1993) and dense crown. Foliage branches spreading in flat- have demonstrated that Fokienia hodginsii represents tened sprays (plagiotropic), ultimate branchlets pin- an evolutionary lineage separate from these two gen- nately arranged, forming frond-like sprays, but in era and therefore merits recognition at the rank of older trees of irregular and shorter length forming genus. Similarities observed, especially in the foliage flat, rounded sprays, deciduous. Leaves decussate, and in later (mature) phases of cone development, imbricate, decurrent, scale-like, strongly dimorphic are likely to be the result of convergent evolution. 385 plate 14. Fokienia hodginsii. 1. Habit of tree. 2. Branch with foliage. 3. Branchlet with juvenile leaves (lower side). 4. Branchlet with adult leaves. 5. Pollen cones. 6. Microsporophyll with pollen sacs. 7. Seed cones. 8, 9. Seeds..
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