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Molecular Phylogeny of Tribolium (Danthonioideae: Poaceae) and Its Taxonomic Implications
BOLUS LIBRARY C24 0007 9273 MOLECULAR PHYLOGENY OF TRIBOLIUM (DANTHONIOIDEAE: POACEAE) AND ITS TAXONOMIC IMPLICATIONS REFILOE NTSOHI NTSREF002 SYSTEMATICS HONOURS PROJECT, 2003 SUPERVISOR: TONY VERBOOM University of Cape Town The copyright of this thesis vests in the author. No quotation from it or information derived from it is to be published without full acknowledgement of the source. The thesis is to be used for private study or non- commercial research purposes only. Published by the University of Cape Town (UCT) in terms of the non-exclusive license granted to UCT by the author. University of Cape Town ABSTRACT Molecular sequence data from two noncoding regions of D A (ITS AND TRNL-F) Were used to produce a phylogeny for the genus Tribolium and its African and Australasian allies. Topological comparisons of the combined molecular data with th e appended existing morphological tree were made. A significant incongruence was I revealed. Molecular data indicate that Tribolium is paraphyletic. The formally defined sections: Uniolae, Acutiflorae and Tribolium have been retrieved by the molecular data. The analysis retrieved Karroochloa as polyphyletic. Monophyly of Schismus is strongly supported. The Australasian species form a monophyletic clade. Data support early divergence of the Merxmuellera species and Pseudopentameris macrantha. I I I I 2 INTRODUCTION Tribolium is a genus of annual and perennial temperate, C3 grasses, currently comprising a total of ten species (Linder and Davidse, 1997). The genus is restricted to southern Africa, where the species are endemic to the winter rainfall region of the western Cape, which includes the Cape Floristic and Namaqualand regions. Here it forms part of the fynbos and succulent Karoo floras, with marginal populations of the most widespread species occurring in the neighbouring biomes. -
Mitteilungen Der Botanischen Staatssammlung München
© Biodiversity Heritage Library, http://www.biodiversitylibrary.org/; www.biologiezentrum.at Mitt. Bot. Staatssamml. München © Biodiversity Heritage Library, http://www.biodiversitylibrary.org/; www.biologiezentrum.at well within their lowermost leaf-sheaths. Besides these characters another can be observed in South American species, namely tufts of hairs in a trans- verse line across the back of the lemma just beneath the Insertion of the cen- tral awn. Species, which exhibit this character, are, however, not confined to South America but occur also in the Easter-Island, in New Zealand, Austra- lia, Tasmania, Indonesia and in the Himalayas. ZoTOV has based his genus Notodanthonia on species from New Zealand. It remains to be decided whether or not this genus should be retained. Only the study of the South American species can elucidate this problem. None of the groups mentioned so far occurs in southern Africa. None- theless, not fewer than 121 Danthonia species have been described from the- re. It goes without saying that many of these names are synonyms. More- over, over the years, quite a number of these species have been separated into different genera which have been recognized by most students of African Gramineae. These include well-known generic names such as Pentameris, Pentaschistis, Strehlochaete, Chaetohromus, Alloeochaete as well as Phacnan- thoecium. But, even so, 36 species remain in Danthonia as can be seen in C. E. Hubbard's treatment of this group in the Flora of Tropical Africa and in Miss Chippindall's cursory guide to South African grasses. In the following account these remaining species of Danthonia will be considered: The first 4 taxa described as Danthonia can be distinguished from typical Danthonia at a first glance by having a long pungent callus at the base of each floret. -
1 Supplementary Information for Invasive Grasses Increase
Supplementary Information For Invasive grasses increase fire occurrence and frequency across U.S. ecoregions Emily J. Fusco1*, John T. Finn2, Jennifer K. Balch3,4, R. Chelsea Nagy3, Bethany A. Bradley1,2 Affiliations: 1 Graduate Program in Organismic and Evolutionary Biology, University of Massachusetts- Amherst, Amherst, Massachusetts, 01003, USA 2 Department of Environmental Conservation, University of Massachusetts- Amherst, Amherst, Massachusetts, 01003, USA 3 Earth Lab, University of Colorado- Boulder, Boulder, Colorado, 80309, USA 4 Department of Geography, University of Colorado-Boulder, Boulder, Colorado, 80309, USA Correspondence to: [email protected] This PDF file includes: Figure S1 Tables S1 to S4 SI References 1 www.pnas.org/cgi/doi/10.1073/pnas.1908253116 Supplemental Table S1: A list of 176 non-native invasive grass and other graminoid species as listed by the Invasive Plant Atlas of the United States (1). For each species, we conducted a Web of Science (WOS) search and recorded whether there was literature suggesting the species altered fire regimes (Yes/No). For each fire promoting species in WOS, we supplemented our determination of whether that species was a fire promoter using the Fire Effects Information System (FEIS; 2). For each species designated as a fire promoter, we searched for available spatial data, and kept only species that were both fire-promoting with spatial data for our analysis. Final species used are highlighted in yellow. WOS FEIS Fire Data Keep for Scientific Name Common Name(s) Search Database Promoter Available Analysis Achnatherum punagrass No - No - No brachychaetum Godr. Barkworth Aegilops cylindrica Host jointed goatgrass No - No - No Aegilops ovate goatgrass No - No - No geniculata Roth Aegilops triuncialis L. -
Recovery Plan for the Sonoran Pronghorn (Antilocapra Americana Sonoriensis) Second Revision
U.S. Fish & Wildlife Service Recovery Plan for the Sonoran Pronghorn (Antilocapra americana sonoriensis) Second Revision Sonoran pronghorn. Photograph by Jim Atkinson, U.S. Fish and Wildlife Service. November 2016 DISCLAIMER Recovery plans delineate such reasonable actions as may be necessary, based upon the best scientific and commercial data available, for the conservation and survival of listed species. Plans are published by the U.S. Fish and Wildlife Service (FWS) and sometimes prepared with the assistance of recovery teams, contractors, state agencies and others. Recovery plans do not necessarily represent the views, official positions or approval of any individuals or agencies involved in the plan formulation, other than FWS. They represent the official position of FWS only after they have been signed by the Regional Director. Recovery plans are guidance and planning documents only; identification of an action to be implemented by any public or private party does not create a legal obligation beyond existing legal requirements. Nothing in this plan should be construed as a commitment or requirement that any federal agency obligate or pay funds in any one fiscal year in excess of appropriations made by Congress for that fiscal year in contravention of the Anti-Deficiency Act, 31 U.S.C. 1341, or any other law or regulation. Approved recovery plans are subject to modification as dictated by new findings, changes in species status, and the completion of recovery actions. LITERATURE CITATION SHOULD READ AS FOLLOWS: U. S. Fish and Wildlife Service. 2016. Recovery Plan for the Sonoran pronghorn (Antilocapra americana sonoriensis), Second Revision. U.S. Fish and Wildlife Service, Southwest Region, Albuquerque, New Mexico, USA. -
Distribution of the Native Grasses of California
HILGARDIA A Journal of Agricultural Science Published by the California Agricultural Experiment Station VOLUME 17 APRIL, 1947 NUMBER 9 CONTENTS DISTRIBUTION OF THE NATIVE GRASSES OF CALIFORNIA ALAN A. BEETLE UNIVERSITY OF CALIFORNIA • BERKELEY, CALIFORNIA HILGARDIA A Journal of Agricultural Science Published by the California Agricultural Experiment Station VOL. 17 APRIL, 1947 NO. 9 DISTRIBUTION OF THE NATIVE GRASSES OF CALIFORNIA1 ALAN A. BEETLE2 THE grasses, supplemented by certain legumes, form the principal basis for range wealth. The natural forage value of the Gramineae as a whole makes an intensive study of their characteristics important, for the broader the knowledge concerning them the more readily may any problem be met. The following paper presents a picture of the current distributions of grasses in California, together with evidences of their floral origins by migration from other regions. Vegetation has many characteristics which are not always apparent at first glance. For instance, certain elements of the vegetation are native in their location, some are native elsewhere and have only recently been introduced. Some are old species often representative of a primitive condition in their genus, still others appear to be recently evolved. Some of the migrants arrived in California from the north during glacial periods, some crossed the ocean, and others came from the south during interglacial periods. Some plants are distributionally restricted for a number of reasons, including: (1) specialization as to habitat or environmental repression, as the species of vernal pools; (2) recent origin (plants sometimes referred to as neoendemics or initiates), as the endemic varieties of Distichlis spicata; (3) ancient origin (paleoendemics or relics); and (4) genotypic specialization (genetic endemics). -
Germination, Survivorship and Growth of an Invasive Desert Grass
Iowa State University Capstones, Theses and Graduate Theses and Dissertations Dissertations 2018 Germination, survivorship and growth of an invasive desert grass, Schismus arabicus, under varied shade, moisture and soil-nitrogen regimes Sarah Elizabeth Emeterio Iowa State University Follow this and additional works at: https://lib.dr.iastate.edu/etd Part of the Ecology and Evolutionary Biology Commons Recommended Citation Emeterio, Sarah Elizabeth, "Germination, survivorship and growth of an invasive desert grass, Schismus arabicus, under varied shade, moisture and soil-nitrogen regimes" (2018). Graduate Theses and Dissertations. 16347. https://lib.dr.iastate.edu/etd/16347 This Thesis is brought to you for free and open access by the Iowa State University Capstones, Theses and Dissertations at Iowa State University Digital Repository. It has been accepted for inclusion in Graduate Theses and Dissertations by an authorized administrator of Iowa State University Digital Repository. For more information, please contact [email protected]. Germination, survivorship and growth of an invasive desert grass, Schismus arabicus , under varied shade, moisture and soil-nitrogen regimes by Sarah E. Emeterio A thesis submitted to the graduate faculty in partial fulfillment of the requirements for the degree of MASTER OF SCIENCE Major: Ecology and Evolutionary Biology Program of Study Committee: Kirk A. Moloney, Major Professor W. Stanley Harpole Lee Burras The student author, whose presentation of the scholarship herein was approved by the program of study committee, is solely responsible for the content of this thesis. The Graduate College will ensure this thesis is globally accessible and will not permit alterations after a degree is conferred. Iowa State University Ames, Iowa 2018 Copyright © Sarah E. -
Annotated Checklist of the Vascular Plant Flora of Grand Canyon-Parashant National Monument Phase II Report
Annotated Checklist of the Vascular Plant Flora of Grand Canyon-Parashant National Monument Phase II Report By Dr. Terri Hildebrand Southern Utah University, Cedar City, UT and Dr. Walter Fertig Moenave Botanical Consulting, Kanab, UT Colorado Plateau Cooperative Ecosystems Studies Unit Agreement # H1200-09-0005 1 May 2012 Prepared for Grand Canyon-Parashant National Monument Southern Utah University National Park Service Mojave Network TABLE OF CONTENTS Page # Introduction . 4 Study Area . 6 History and Setting . 6 Geology and Associated Ecoregions . 6 Soils and Climate . 7 Vegetation . 10 Previous Botanical Studies . 11 Methods . 17 Results . 21 Discussion . 28 Conclusions . 32 Acknowledgments . 33 Literature Cited . 34 Figures Figure 1. Location of Grand Canyon-Parashant National Monument in northern Arizona . 5 Figure 2. Ecoregions and 2010-2011 collection sites in Grand Canyon-Parashant National Monument in northern Arizona . 8 Figure 3. Soil types and 2010-2011 collection sites in Grand Canyon-Parashant National Monument in northern Arizona . 9 Figure 4. Increase in the number of plant taxa confirmed as present in Grand Canyon- Parashant National Monument by decade, 1900-2011 . 13 Figure 5. Southern Utah University students enrolled in the 2010 Plant Anatomy and Diversity course that collected during the 30 August 2010 experiential learning event . 18 Figure 6. 2010-2011 collection sites and transportation routes in Grand Canyon-Parashant National Monument in northern Arizona . 22 2 TABLE OF CONTENTS Page # Tables Table 1. Chronology of plant-collecting efforts at Grand Canyon-Parashant National Monument . 14 Table 2. Data fields in the annotated checklist of the flora of Grand Canyon-Parashant National Monument (Appendices A, B, C, and D) . -
The Naturalized Vascular Plants of Western Australia 1
12 Plant Protection Quarterly Vol.19(1) 2004 Distribution in IBRA Regions Western Australia is divided into 26 The naturalized vascular plants of Western Australia natural regions (Figure 1) that are used for 1: Checklist, environmental weeds and distribution in bioregional planning. Weeds are unevenly distributed in these regions, generally IBRA regions those with the greatest amount of land disturbance and population have the high- Greg Keighery and Vanda Longman, Department of Conservation and Land est number of weeds (Table 4). For exam- Management, WA Wildlife Research Centre, PO Box 51, Wanneroo, Western ple in the tropical Kimberley, VB, which Australia 6946, Australia. contains the Ord irrigation area, the major cropping area, has the greatest number of weeds. However, the ‘weediest regions’ are the Swan Coastal Plain (801) and the Abstract naturalized, but are no longer considered adjacent Jarrah Forest (705) which contain There are 1233 naturalized vascular plant naturalized and those taxa recorded as the capital Perth, several other large towns taxa recorded for Western Australia, com- garden escapes. and most of the intensive horticulture of posed of 12 Ferns, 15 Gymnosperms, 345 A second paper will rank the impor- the State. Monocotyledons and 861 Dicotyledons. tance of environmental weeds in each Most of the desert has low numbers of Of these, 677 taxa (55%) are environmen- IBRA region. weeds, ranging from five recorded for the tal weeds, recorded from natural bush- Gibson Desert to 135 for the Carnarvon land areas. Another 94 taxa are listed as Results (containing the horticultural centre of semi-naturalized garden escapes. Most Total naturalized flora Carnarvon). -
Schismus Barbatus Global Invasive
FULL ACCOUNT FOR: Schismus barbatus Schismus barbatus System: Terrestrial Kingdom Phylum Class Order Family Plantae Magnoliophyta Liliopsida Cyperales Poaceae Common name Mediterranean grass (English), common Mediterranean grass (English) Synonym Festuca barbata , Loefl. ex L. Similar species Schismus arabicus Summary Schismus barbatus is invasive in the Mojave and Sonoran Deserts of the southwestern United States. However, this species also provides food for desert turtles and rodents. It is most common in disturbed areas of the deserts and becomes most abundant during years of favourable winter rains. view this species on IUCN Red List Species Description Schismus barbatus is a tufted annual up to 40cm tall. The inflorescence is a narrow, erect, greenish-purple panicle (branched, cluster), produced in spring [in western United States].” Devender et al. (1997) describes S. barbatus as a cool-season annual. Wilken and Hannah (1998) describe S. barbatus as being a slow growing grass with smooth stems that ascend to somewhat grow on the ground. The leaves are alternate with irregular jagged edges. The stem is smooth for the most part with scattered soft hairs. The blades are narrowly linear, 5-10cm long, 0.5-2mm wide, and curled inward. The grass has dense infloresence 1-5cm long and ovate to elliptic in shape. Spikelets are 4-8mm long, becoming reddish, and are composed of 3-8 florets. Brooks (2000a) indicates that S. barbatus is so similar to S. arabicus physiognomically and in terms of the habitats it invades that the two species may have very similar ecological effects. Notes According to Devender et al. (1997), S. -
On the Flora of Australia
L'IBRARY'OF THE GRAY HERBARIUM HARVARD UNIVERSITY. BOUGHT. THE FLORA OF AUSTRALIA, ITS ORIGIN, AFFINITIES, AND DISTRIBUTION; BEING AN TO THE FLORA OF TASMANIA. BY JOSEPH DALTON HOOKER, M.D., F.R.S., L.S., & G.S.; LATE BOTANIST TO THE ANTARCTIC EXPEDITION. LONDON : LOVELL REEVE, HENRIETTA STREET, COVENT GARDEN. r^/f'ORElGN&ENGLISH' <^ . 1859. i^\BOOKSELLERS^.- PR 2G 1.912 Gray Herbarium Harvard University ON THE FLORA OF AUSTRALIA ITS ORIGIN, AFFINITIES, AND DISTRIBUTION. I I / ON THE FLORA OF AUSTRALIA, ITS ORIGIN, AFFINITIES, AND DISTRIBUTION; BEIKG AN TO THE FLORA OF TASMANIA. BY JOSEPH DALTON HOOKER, M.D., F.R.S., L.S., & G.S.; LATE BOTANIST TO THE ANTARCTIC EXPEDITION. Reprinted from the JJotany of the Antarctic Expedition, Part III., Flora of Tasmania, Vol. I. LONDON : LOVELL REEVE, HENRIETTA STREET, COVENT GARDEN. 1859. PRINTED BY JOHN EDWARD TAYLOR, LITTLE QUEEN STREET, LINCOLN'S INN FIELDS. CONTENTS OF THE INTRODUCTORY ESSAY. § i. Preliminary Remarks. PAGE Sources of Information, published and unpublished, materials, collections, etc i Object of arranging them to discuss the Origin, Peculiarities, and Distribution of the Vegetation of Australia, and to regard them in relation to the views of Darwin and others, on the Creation of Species .... iii^ § 2. On the General Phenomena of Variation in the Vegetable Kingdom. All plants more or less variable ; rate, extent, and nature of variability ; differences of amount and degree in different natural groups of plants v Parallelism of features of variability in different groups of individuals (varieties, species, genera, etc.), and in wild and cultivated plants vii Variation a centrifugal force ; the tendency in the progeny of varieties being to depart further from their original types, not to revert to them viii Effects of cross-impregnation and hybridization ultimately favourable to permanence of specific character x Darwin's Theory of Natural Selection ; — its effects on variable organisms under varying conditions is to give a temporary stability to races, species, genera, etc xi § 3. -
SUPPLEMENTARY MATERIAL African Journal of Range & Forage Science, 2021 Biogeog
SUPPLEMENTARY MATERIAL https://doi.org/10.2989/10220119.2021.1918247 African Journal of Range & Forage Science, 2021 Biogeographical patterns of grasses (Poaceae) indigenous to South Africa, Lesotho and Eswatini Marike Trytsman, Francuois L Muller, Craig D Morris and Abraham E van Wyk Supplementary material 4. Grass species endemic/near-endemic to southern Africa (South Africa, Lesotho and Eswatini) (based on Fish et al. 2015) as recorded in each grasschorion. IOCB: Indian Ocean Coastal Belt Central Arid Region Fynbos Grassland IOCB Savanna Succulent Karoo 1 Agrostis eriantha Agrostis bergiana Agrostis barbuligera Anthephora argentea Agrostis barbuligera Agrostis bergiana 2 Anthephora argentea Agrostis eriantha Agrostis bergiana Aristida diffusa Agrostis bergiana Aristida dasydesmis 3 Aristida canescens Agrostis polypogonoides Agrostis eriantha Aristida transvaalensis Agrostis eriantha Aristida diffusa 4 Aristida dasydesmis Agrostis schlechteri Agrostis subulifolia Brachypodium flexum Anthephora argentea Aristida engleri 5 Aristida diffusa Anthoxanthum dregeanum Anthoxanthum dregeanum Bromus firmior Aristida canescens Aristida parvula 6 Aristida engleri Anthoxanthum tongo Aristida canescens Capeochloa cincta Aristida dasydesmis Brachypodium bolusii 7 Aristida parvula Aristida canescens Aristida diffusa Cymbopogon prolixus Aristida diffusa Brachypodium flexum 8 Aristida spectabilis Aristida diffusa Aristida effusa Cynodon bradleyi Aristida effusa Capeochloa arundinacea 9 Brachypodium bolusii Brachypodium flexum Aristida monticola Cynodon -
California Grasslands and Range Forage Grasses
CALIFORNIA GRASSLANDS AND RANGE FORAGE GRASSES ARTHUR W. SAMPSON AGNES CHASE DONALD W. HEDRICK BULLETIN 724 MAY, 1951 CALIFORNIA AGRICULTURAL EXPERIMENT STATION i THE COLLEGE OF AGRICULTURE UNIVERSITY OF CALIFORNIA . CALIFORNIA GRASSLANDS AND RANGE FORAGE GRASSES provides useful and technical information on the uncultivated or wild grass- lands, and the native and naturalized range forage grasses of California. This bulletin will be helpful to you if you are among these readers: 1 Stockmen who have had some botanical training and who will want to use the illustrated keys and descriptions to determine the identity and the relative usefulness of the grasses growing on their range; 2. Range technicians and range appraisers who are chiefly concerned with management, evaluations, and economic considerations of the state's range lands; and 3. Students of range management and related fields whose knowl- edge of ecology, forage value, and taxonomy of the range grasses is an essential part of their training or official work. THE AUTHORS: Arthur W. Sampson is Professor of Forestry and Plant Ecologist in the Experiment Station, Berkeley. Agnes Chase is Research Associate, U. S. National Herbarium, Smithsonian Institu- tion; formerly Senior Agrostologist, U. S. Department of Agriculture. Donald W. Hedrick is Research Assistant in the Department of Forestry; on leave from the Soil Conservation Service, U. S. Department of Agriculture. Manuscript submitted for publication August 22, 1949. CONTENTS PAGE Introduction . 5 Where Grasses Grow 7 Topography, climate, grassland soils, life zones, grasslands in relation to other plant associations Plant Succession and the Climax Cover 17 The Nutrition of Range Grasses 18 Annual vs.