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Why Mushrooms Have Evolved to Be So Promiscuous: Insights from Evolutionary and Ecological Patterns
fungal biology reviews 29 (2015) 167e178 journal homepage: www.elsevier.com/locate/fbr Review Why mushrooms have evolved to be so promiscuous: Insights from evolutionary and ecological patterns Timothy Y. JAMES* Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, USA article info abstract Article history: Agaricomycetes, the mushrooms, are considered to have a promiscuous mating system, Received 27 May 2015 because most populations have a large number of mating types. This diversity of mating Received in revised form types ensures a high outcrossing efficiency, the probability of encountering a compatible 17 October 2015 mate when mating at random, because nearly every homokaryotic genotype is compatible Accepted 23 October 2015 with every other. Here I summarize the data from mating type surveys and genetic analysis of mating type loci and ask what evolutionary and ecological factors have promoted pro- Keywords: miscuity. Outcrossing efficiency is equally high in both bipolar and tetrapolar species Genomic conflict with a median value of 0.967 in Agaricomycetes. The sessile nature of the homokaryotic Homeodomain mycelium coupled with frequent long distance dispersal could account for selection favor- Outbreeding potential ing a high outcrossing efficiency as opportunities for choosing mates may be minimal. Pheromone receptor Consistent with a role of mating type in mediating cytoplasmic-nuclear genomic conflict, Agaricomycetes have evolved away from a haploid yeast phase towards hyphal fusions that display reciprocal nuclear migration after mating rather than cytoplasmic fusion. Importantly, the evolution of this mating behavior is precisely timed with the onset of diversification of mating type alleles at the pheromone/receptor mating type loci that are known to control reciprocal nuclear migration during mating. -
Ottawa Fury Fc @ Tampa Bay Rowdies
TAMPA BAY ROWDIES OTTAWA FURY FC - VS – USL REGULAR SEASON April 8, 2017 7:30PM ET 2017 Record (W-D-L) & Conference Ranking: Al Lang Stadium, th 2017 Record (W-D-L) & Conference Ranking: 0-0-1 | 0 Point | 12 Place | 2GF 3GA st St. Petersburg, FL 2-0-0 | 6 Points | 1 Place | 5GF 0GA HOW TO WATCH AND LISTEN: TV: Rogers TV (Local) Channels: 22 (SD) & 182 (HD) | RADIO (EN): TSN 1200 | TSN1200.ca STREAM: USL Match Centre | SOCIAL MEDIA: LIVE updates on Twitter @OttawaFuryFC MATCH NOTES: Weekly Storylines: Fury FC dropped a hard-fought 3-2 decision in their USL opener last Saturday in St. Louis. Despite overturning a two-goal deficit, a late Saint Louis FC goal thwarted Ottawa’s comeback Saturday’s match marks the first time that Ottawa and Tampa Bay meet in USL action. The two sides have squared off nine times in NASL action. Fury FC is 2W-5D-2L lifetime versus the Rowdies Ottawa-native Eddie Edward and Onua Obasi notched their first-ever goals in Fury colours in Saturday’s loss. Edward’s goal was Ottawa’s first-ever goal in USL play Ottawa will be without the services of Shane McEleney who received a red card late in Saturday’s match The Rowdies sit atop the Eastern Conference with six points having outscored the opposition 5-0 Striker Georgi Hristov is among the league-leaders with two goals this season. Former Fury FC defender Kyle Porter is now a member of the Tampa Bay Rowdies Scouting the Rowdies: Tampa Bay finished ninth in the NASL last year with a combined record of 9W-12D-11L. -
Female Fellows of the Royal Society
Female Fellows of the Royal Society Professor Jan Anderson FRS [1996] Professor Ruth Lynden-Bell FRS [2006] Professor Judith Armitage FRS [2013] Dr Mary Lyon FRS [1973] Professor Frances Ashcroft FMedSci FRS [1999] Professor Georgina Mace CBE FRS [2002] Professor Gillian Bates FMedSci FRS [2007] Professor Trudy Mackay FRS [2006] Professor Jean Beggs CBE FRS [1998] Professor Enid MacRobbie FRS [1991] Dame Jocelyn Bell Burnell DBE FRS [2003] Dr Philippa Marrack FMedSci FRS [1997] Dame Valerie Beral DBE FMedSci FRS [2006] Professor Dusa McDuff FRS [1994] Dr Mariann Bienz FMedSci FRS [2003] Professor Angela McLean FRS [2009] Professor Elizabeth Blackburn AC FRS [1992] Professor Anne Mills FMedSci FRS [2013] Professor Andrea Brand FMedSci FRS [2010] Professor Brenda Milner CC FRS [1979] Professor Eleanor Burbidge FRS [1964] Dr Anne O'Garra FMedSci FRS [2008] Professor Eleanor Campbell FRS [2010] Dame Bridget Ogilvie AC DBE FMedSci FRS [2003] Professor Doreen Cantrell FMedSci FRS [2011] Baroness Onora O'Neill * CBE FBA FMedSci FRS [2007] Professor Lorna Casselton CBE FRS [1999] Dame Linda Partridge DBE FMedSci FRS [1996] Professor Deborah Charlesworth FRS [2005] Dr Barbara Pearse FRS [1988] Professor Jennifer Clack FRS [2009] Professor Fiona Powrie FRS [2011] Professor Nicola Clayton FRS [2010] Professor Susan Rees FRS [2002] Professor Suzanne Cory AC FRS [1992] Professor Daniela Rhodes FRS [2007] Dame Kay Davies DBE FMedSci FRS [2003] Professor Elizabeth Robertson FRS [2003] Professor Caroline Dean OBE FRS [2004] Dame Carol Robinson DBE FMedSci -
Inside Science
SPRING 2009 NEWS FROM THE ROYAL SOCIETY INSIDE SCIENCE YOUNG EXPLORERS TOUCHDOWN IN NEW ZEALAND International Expedition Prize is a ‘once in a lifetime experience’ SCIENCE TAKES TO THE STAGE The Royal Shakespeare Company premiers a new play on the emergence of modern science UPDATE FROM THE ROYAL SOCIETY This third issue of Inside Science contains early information DID YOU KNOW? about exciting plans for the Royal Society’s 350th Anniversary in 2010. The Anniversary is a marvellous STEADY FOOTING, opportunity to increase the profile of science, explore its SHAKY BRIDGE benefits and address the challenges it presents for society On its opening day, crowds of but perhaps most important of all to inspire young minds pedestrians experienced unexpected with the excitement of scientific discovery. swaying as they walked across London’s Our policy work continues to address major scientific issues Millennium Bridge. Whilst pedestrians affecting the UK. In December we cautioned the Government on fondly nicknamed it the ‘wobbly bridge’, the levels of separated plutonium stockpiled in the UK – currently physicists were busy exploring the the highest in the world. With support from our Plutonium Working Group, the Society has reasons for the phenomenon. submitted detailed comment to the Nuclear Decommissioning Authority (NDA) for a report to The view was widely held that the Government on management options for the stockpile. ‘wobble’ was due to crowd loading and Late last year we ran an extremely successful MP-Scientist pairing scheme, helping to build pedestrians synchronising their footsteps bridges between parliamentarians and some of the best young scientists in the UK. -
Smutty Alchemy
University of Calgary PRISM: University of Calgary's Digital Repository Graduate Studies The Vault: Electronic Theses and Dissertations 2021-01-18 Smutty Alchemy Smith, Mallory E. Land Smith, M. E. L. (2021). Smutty Alchemy (Unpublished doctoral thesis). University of Calgary, Calgary, AB. http://hdl.handle.net/1880/113019 doctoral thesis University of Calgary graduate students retain copyright ownership and moral rights for their thesis. You may use this material in any way that is permitted by the Copyright Act or through licensing that has been assigned to the document. For uses that are not allowable under copyright legislation or licensing, you are required to seek permission. Downloaded from PRISM: https://prism.ucalgary.ca UNIVERSITY OF CALGARY Smutty Alchemy by Mallory E. Land Smith A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF DOCTOR OF PHILOSOPHY GRADUATE PROGRAM IN ENGLISH CALGARY, ALBERTA JANUARY, 2021 © Mallory E. Land Smith 2021 MELS ii Abstract Sina Queyras, in the essay “Lyric Conceptualism: A Manifesto in Progress,” describes the Lyric Conceptualist as a poet capable of recognizing the effects of disparate movements and employing a variety of lyric, conceptual, and language poetry techniques to continue to innovate in poetry without dismissing the work of other schools of poetic thought. Queyras sees the lyric conceptualist as an artistic curator who collects, modifies, selects, synthesizes, and adapts, to create verse that is both conceptual and accessible, using relevant materials and techniques from the past and present. This dissertation responds to Queyras’s idea with a collection of original poems in the lyric conceptualist mode, supported by a critical exegesis of that work. -
The ST Cross College Magazine 2015 Ad Quattuor Cardines Mundi
CROSSWORD THE ST CROSS COLLEGE MAGAZINE 2015 AD QUAttUOR CARDINES MUNDI Contents ST Cross COLLEGE West Quad Campaign UNIVERSITY OF OXFORD 04 An update on the progress towards achieving this landmark project COVER STORY – The 161st Boat Race 05 St Cross students making history on the Tideway The Body in the Garden 06 Recent investigations undertaken by Oxford Archaeology ahead of the construction of the West Quad revealed a body in the garden The St Cross 50th Anniversary Lecture Series 08 This series of termly lectures brought three eminent speakers to Oxford to celebrate the th Crossword – Issue 23 College’s 50 Anniversary Editor: Susan Berrington St Cross Merchandise 09 A selection of gifts, books and momentos Managing Editor: Ella Bedrock Design: Broccoli Creative Design AI Risk 10 Stuart Armstrong looks at the risks associated with Contact details: Artificial Intelligence The Development & Alumni Relations Office St Cross College Students’ News 61 St Giles 12 Oxford ‘Four Corners’ - The St Cross International Poetry OX1 3LZ 13 Competition 2015 Tel: +44 (0)1865 278480 Kate Venables talks of the success of the ‘Four Corners’ Email: [email protected] International Poetry Competition www.stx.ox.ac.uk St Cross College Photography Competition 2015 Cover Image: 14 Students Jamie Cook (MSc Engineering Science) and Shelley Pearson (MSc Child Development Sports News and Education), who were in the winning Dark 16 Blue boats in this year’s Oxford and Cambridge Members’ News Boat Race, with Olympic gold medallist rower 18 Tim Foster (Dip Social Studies, 1996). Matriculation and College Photographs Photo credit: Phil Sills 20 The 2015 Telethon This edition of Crossword is printed using an 22 A conversation from the call room environmentally friendly, waterless printing process, on Forest Stewardship Council (FSC) certified paper and to Eco Management Audit The Four Corners of the World Scheme (EMAS) standards. -
UEFA EURO 2016 FINAL TOURNAMENT DRAW PRESS KIT Paris, France Saturday 12 December 2015 18.00CET Last Updated 14/12/2015 08:04CET
UEFA EURO 2016 FINAL TOURNAMENT DRAW PRESS KIT Paris, France Saturday 12 December 2015 18.00CET Last updated 14/12/2015 08:04CET UEFA EURO 2016 OFFICIAL SPONSORS Team profiles: Pot 4 2 Legend 80 1 UEFA EURO 2016 - FINAL TOURNAMENT DRAW Saturday 12 December 2015 - 18.00CET (18.00 local time) Press kit Paris, France Team profiles: Pot 4 Turkey Republic of Ireland Iceland Wales Albania Northern Ireland 2 UEFA EURO 2016 - FINAL TOURNAMENT DRAW Saturday 12 December 2015 - 18.00CET (18.00 local time) Press kit Paris, France Turkey Turkey - Team profile Best result: Semi-finals (2008) Coach: Fatih Terim Leading scorers: all-time – Hakan Şükür (51); current – Burak Yılmaz (19) Most appearances: all-time – Rüştü Reçber (120); current – Emre Belözoğlu (93) Association formed: 1923 Nickname: Ay-Yıldızlılar (The Crescent-Stars) Where they play: Various The pinnacle of Turkey's achievement came at the 2002 FIFA World Cup, where Şenol Güneş's side upset the odds to finish third. It was only their second World Cup appearance and they have not been back since. It took Turkey ten attempts to qualify for their first UEFA European Championship final tournament, and after losing all three matches at EURO '96 they reached the quarter-finals at UEFA EURO 2000 under Mustafa Denizli. They went one step further eight years later in Austria/ Switzerland, a dramatic penalty shoot-out win over Croatia putting Fatih Terim's team into the semi-finals, where they lost 3-2 to Germany. After missing out in Poland/Ukraine, they returned to the finals at UEFA EURO 2016 by qualifying as the best third-placed side over the nine preliminary sections. -
Evolution of Sexual Reproduction: a View from the Fungal Kingdom Supports an Evolutionary Epoch with Sex Before Sexes
fungal biology reviews xxx (2015) 1e10 journal homepage: www.elsevier.com/locate/fbr Review Evolution of sexual reproduction: A view from the fungal kingdom supports an evolutionary epoch with sex before sexes Joseph HEITMAN* Department of Molecular Genetics and Microbiology, Duke University Medical Center, Durham, NC 27710, USA article info abstract Article history: Sexual reproduction is conserved throughout each supergroup within the eukaryotic tree Received 16 February 2015 of life, and therefore thought to have evolved once and to have been present in the last eu- Received in revised form karyotic common ancestor (LECA). Given the antiquity of sex, there are features of sexual 10 June 2015 reproduction that are ancient and ancestral, and thus shared in diverse extant organisms. Accepted 17 August 2015 On the other hand, the vast evolutionary distance that separates any given extant species from the LECA necessarily implies that other features of sex will be derived. While most Keywords: types of sex we are familiar with involve two opposite sexes or mating types, recent studies Evolution in the fungal kingdom have revealed novel and unusual patterns of sexual reproduction, Heterothallic including unisexual reproduction. In this mode of reproduction a single mating type can Homothallic on its own undergo self-fertile/homothallic reproduction, either with itself or with other Inbreeding members of the population of the same mating type. Unisexual reproduction has arisen MAT independently as a derived feature in several different lineages. That a myriad of different Mating type types of sex determination and sex determinants abound in animals, plants, protists, and Meiosis fungi suggests that sex specification itself may not be ancestral and instead may be a Outcrossing derived trait. -
Chemistry a Level
World class thinking. World class achieving. St Mary’s College CHEMISTRY (AQA) Pre-course reading and guidance A Level Chemistry WHY SHOULD I CHOOSE CHEMISTRY? › Chemistry will give you the opportunity to work in many areas including dentistry, veterinary science and medicine. The course is delivered by members of staff that have a great deal of expertise in delivering the subject. Group sizes are normally a maximum of twenty five pupils. › Pick up a can of soft drink and you’ll find chemistry everywhere, from the metal can you’re holding, to the paint used to cover it and the liquid inside. Just breathe in and out and you’re performing a chemical reaction. Chemistry is sometimes known as the "central science" because it helps to connect physical sciences, like maths and physics, with applied sciences, like biology, medicine and engineering. WHAT IS THE COURSE STRUCTURE LIKE? › The course is a two year course. It consists of three exams. Papers 1 and 2 are worth 35% of the qualification with Paper 3 being worth 30%. There is no coursework. World class thinking. World class achieving. A LEVEL Chemistry Exams (sat in Y13) Paper Topics Weighting (%) Time (hours) • Physical Chemistry 1 35 2 • Inorganic Chemistry • Physical Chemistry 2 35 2 • Organic Chemistry • All content 3 30 2 Exam Syllabus for Chemistry: https://bit.ly/345So5z World class thinking. World class achieving. A LEVEL Chemistry Course Texts (Year 12) You will be loaned the following text book in Y12: ▪ AQA Chemistry: A Level Year 1 and AS by Ted Lister Additionally, you will have the opportunity to purchase the following revision guide: ▪ CGP A-Level Chemistry: AQA Year 1 & AS Complete Revision & Practice with Online Edition World class thinking. -
Smithson Tennant Celebration 23 November Flyer (PDF , 489Kb)
International Year of Chemistry Ir Os Ir Os Programme 16.30 Exhibition Viewing and Refreshments Wednesday 23 November 2011 17.00 Introduction: Professor Robin Perutz (York) National Science Learning Centre, University of York 17.05 Professor Martyn Poliakoff FRS (Nottingham) From test tube to YouTube November 2011 marks the 250th anniversary 17.45 Refreshments and Exhibition of the birth of Smithson Tennant 18.15 Dr Adam Hart-Davis Chemical heroes of Smithson Tennant was born in Selby, North Yorkshire, from modest Yorkshire beginnings he went on to: 19.00 Closing Remarks: David Lewis Discover iridium and osmium Hold the 1702 Chair of Chemistry at Cambridge The exhibition will be available for viewing throughout Be elected a Fellow of the Royal Society the event. Win the Royal Society’s Copley Medal Become a confidante of many of Europe's well-known scientists. Adam Hart-Davis is the presenter of numerous TV programmes about science and history, and often about both at once. He has presented for BBC, Yorkshire Television, ITV and the Os History Channel. Programmes include Ir What the Romans (and others) did for The event is sponsored by: us, How London was built and The Scientific Eye. He has also written many Johnson Matthey Catalysts, BP Plc, The Royal Society of books. The latest, out Oct 2011, is The book of time: everything you need to Chemistry (Dalton Division and Central Yorkshire Sec- know about the biggest idea in the tion) Universe (Mitchell Beazley). He studied for his DPhil in the & The University of York University of York and also holds an honorary doctorate of the University. -
Annual Conference Washington, D.C
Association for Asian Studies ANNUAL CONFERENCE WASHINGTON, D.C. MARCH 22-25 2018 Spatial Data Center & China Data Center UNIVERSITY OF MICHIGAN 330 Packard St, Ann Arbor, MI 48106-1248, USA TEL: (734)647-9610 / FAX: (734)763-0335 / EMAIL: [email protected] Please join the following workshop organized by China Data Center: “Recent Development and New Features of China Data Online” Time: 3:00pm - 5:30pm, Thursday, March 22, 2018 Site: Roosevelt 3, Marriott Wardman Park Hotel, Washington, D.C. New Data and Features of China Data Online The following new databases have been added to China Statistics Database and Census Database: Statistical Datasheets provides about 270,000 statistical tables from all provincial yearbooks and some other sources with full text search function and metadata, including citation information and unique table ID for direct access. Census Maps covers more than 7 million census maps with data by province, city, county or even township, including population census 2000 and 2010, economic census 2004 and 2008, basic unit census 2001, and industrial census 1995. Statistical Charts provides a rich collection of statistical charts for those monthly and yearly statistics at country, province, prefecture city and county levels with full text search function and metadata, including citation information and unique chart ID for direct access. New Features of China Geo-Explorer and US Geo-Explorer: Chinese Version of “China Map Library” is part of China Geo-Explorer. It offers about 8 million maps for the demographic and business data of China. Those maps provide comprehensive information of China at province, prefecture cities, county, and township levels. -
Year in Review
Year in review For the year ended 31 March 2017 Trustees2 Executive Director YEAR IN REVIEW The Trustees of the Society are the members Dr Julie Maxton of its Council, who are elected by and from Registered address the Fellowship. Council is chaired by the 6 – 9 Carlton House Terrace President of the Society. During 2016/17, London SW1Y 5AG the members of Council were as follows: royalsociety.org President Sir Venki Ramakrishnan Registered Charity Number 207043 Treasurer Professor Anthony Cheetham The Royal Society’s Trustees’ report and Physical Secretary financial statements for the year ended Professor Alexander Halliday 31 March 2017 can be found at: Foreign Secretary royalsociety.org/about-us/funding- Professor Richard Catlow** finances/financial-statements Sir Martyn Poliakoff* Biological Secretary Sir John Skehel Members of Council Professor Gillian Bates** Professor Jean Beggs** Professor Andrea Brand* Sir Keith Burnett Professor Eleanor Campbell** Professor Michael Cates* Professor George Efstathiou Professor Brian Foster Professor Russell Foster** Professor Uta Frith Professor Joanna Haigh Dame Wendy Hall* Dr Hermann Hauser Professor Angela McLean* Dame Georgina Mace* Dame Bridget Ogilvie** Dame Carol Robinson** Dame Nancy Rothwell* Professor Stephen Sparks Professor Ian Stewart Dame Janet Thornton Professor Cheryll Tickle Sir Richard Treisman Professor Simon White * Retired 30 November 2016 ** Appointed 30 November 2016 Cover image Dancing with stars by Imre Potyó, Hungary, capturing the courtship dance of the Danube mayfly (Ephoron virgo). YEAR IN REVIEW 3 Contents President’s foreword .................................. 4 Executive Director’s report .............................. 5 Year in review ...................................... 6 Promoting science and its benefits ...................... 7 Recognising excellence in science ......................21 Supporting outstanding science .....................