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Why Mushrooms Have Evolved to Be So Promiscuous: Insights from Evolutionary and Ecological Patterns
fungal biology reviews 29 (2015) 167e178 journal homepage: www.elsevier.com/locate/fbr Review Why mushrooms have evolved to be so promiscuous: Insights from evolutionary and ecological patterns Timothy Y. JAMES* Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, USA article info abstract Article history: Agaricomycetes, the mushrooms, are considered to have a promiscuous mating system, Received 27 May 2015 because most populations have a large number of mating types. This diversity of mating Received in revised form types ensures a high outcrossing efficiency, the probability of encountering a compatible 17 October 2015 mate when mating at random, because nearly every homokaryotic genotype is compatible Accepted 23 October 2015 with every other. Here I summarize the data from mating type surveys and genetic analysis of mating type loci and ask what evolutionary and ecological factors have promoted pro- Keywords: miscuity. Outcrossing efficiency is equally high in both bipolar and tetrapolar species Genomic conflict with a median value of 0.967 in Agaricomycetes. The sessile nature of the homokaryotic Homeodomain mycelium coupled with frequent long distance dispersal could account for selection favor- Outbreeding potential ing a high outcrossing efficiency as opportunities for choosing mates may be minimal. Pheromone receptor Consistent with a role of mating type in mediating cytoplasmic-nuclear genomic conflict, Agaricomycetes have evolved away from a haploid yeast phase towards hyphal fusions that display reciprocal nuclear migration after mating rather than cytoplasmic fusion. Importantly, the evolution of this mating behavior is precisely timed with the onset of diversification of mating type alleles at the pheromone/receptor mating type loci that are known to control reciprocal nuclear migration during mating. -
Radio 4 Listings for 2 – 8 May 2020 Page 1 of 14
Radio 4 Listings for 2 – 8 May 2020 Page 1 of 14 SATURDAY 02 MAY 2020 Professor Martin Ashley, Consultant in Restorative Dentistry at panel of culinary experts from their kitchens at home - Tim the University Dental Hospital of Manchester, is on hand to Anderson, Andi Oliver, Jeremy Pang and Dr Zoe Laughlin SAT 00:00 Midnight News (m000hq2x) separate the science fact from the science fiction. answer questions sent in via email and social media. The latest news and weather forecast from BBC Radio 4. Presenter: Greg Foot This week, the panellists discuss the perfect fry-up, including Producer: Beth Eastwood whether or not the tomato has a place on the plate, and SAT 00:30 Intrigue (m0009t2b) recommend uses for tinned tuna (that aren't a pasta bake). Tunnel 29 SAT 06:00 News and Papers (m000htmx) Producer: Hannah Newton 10: The Shoes The latest news headlines. Including the weather and a look at Assistant Producer: Rosie Merotra the papers. “I started dancing with Eveline.” A final twist in the final A Somethin' Else production for BBC Radio 4 chapter. SAT 06:07 Open Country (m000hpdg) Thirty years after the fall of the Berlin Wall, Helena Merriman Closed Country: A Spring Audio-Diary with Brett Westwood SAT 11:00 The Week in Westminster (m000j0kg) tells the extraordinary true story of a man who dug a tunnel into Radio 4's assessment of developments at Westminster the East, right under the feet of border guards, to help friends, It seems hard to believe, when so many of us are coping with family and strangers escape. -
Female Fellows of the Royal Society
Female Fellows of the Royal Society Professor Jan Anderson FRS [1996] Professor Ruth Lynden-Bell FRS [2006] Professor Judith Armitage FRS [2013] Dr Mary Lyon FRS [1973] Professor Frances Ashcroft FMedSci FRS [1999] Professor Georgina Mace CBE FRS [2002] Professor Gillian Bates FMedSci FRS [2007] Professor Trudy Mackay FRS [2006] Professor Jean Beggs CBE FRS [1998] Professor Enid MacRobbie FRS [1991] Dame Jocelyn Bell Burnell DBE FRS [2003] Dr Philippa Marrack FMedSci FRS [1997] Dame Valerie Beral DBE FMedSci FRS [2006] Professor Dusa McDuff FRS [1994] Dr Mariann Bienz FMedSci FRS [2003] Professor Angela McLean FRS [2009] Professor Elizabeth Blackburn AC FRS [1992] Professor Anne Mills FMedSci FRS [2013] Professor Andrea Brand FMedSci FRS [2010] Professor Brenda Milner CC FRS [1979] Professor Eleanor Burbidge FRS [1964] Dr Anne O'Garra FMedSci FRS [2008] Professor Eleanor Campbell FRS [2010] Dame Bridget Ogilvie AC DBE FMedSci FRS [2003] Professor Doreen Cantrell FMedSci FRS [2011] Baroness Onora O'Neill * CBE FBA FMedSci FRS [2007] Professor Lorna Casselton CBE FRS [1999] Dame Linda Partridge DBE FMedSci FRS [1996] Professor Deborah Charlesworth FRS [2005] Dr Barbara Pearse FRS [1988] Professor Jennifer Clack FRS [2009] Professor Fiona Powrie FRS [2011] Professor Nicola Clayton FRS [2010] Professor Susan Rees FRS [2002] Professor Suzanne Cory AC FRS [1992] Professor Daniela Rhodes FRS [2007] Dame Kay Davies DBE FMedSci FRS [2003] Professor Elizabeth Robertson FRS [2003] Professor Caroline Dean OBE FRS [2004] Dame Carol Robinson DBE FMedSci -
Trafalgar Day Dr Liz Sidwell Our Eco
This issue: Victory Open Evening | Sixth Form Trip | Council Visit |<U 3UR¿OHV November 2011 Vol 2 No 1 Trafalgar Day Celebrations Galore over 2 days Dr Liz Sidwell Full visit report Our Eco Day We show our true green colours www.ormistonvictoryacademy.co.uk November 2011 Victory Flag: Vol 1 No. 6 Principal Points P3 News in Brief P4 Costessey News/Beauty Blog P5 Celebrating Trafalgar Day P6 - 11 Reach For The Stars P12 - 13 Beauty Transformational A-Levels P14 Eton @ Norfolk P15 School Commissioner Visit P16 - 17 ISSUE Pen to Paper P18 VIP Roll Of Honour P19 Victory Goes Green P20 - 21 @Victory Green Academy Update P22 - 23 VIP Visit - Peter Swift P24 Lights, Camera, Action P25 7KH¿UVWEHDXW\VDORQZLWKLQDQ$FDGHP\ Welcome Year 6s Victory Open Evening P26 from across the county. It was a Victory In The Stars P27 Principal Points brilliant day. State of the art facilities, The Stars @ Victory P27 Sixth Form Sojourn Trip P28 - 29 So much has happened I said from the outset that my goal was Family Memories P30 WRPDNH9LFWRU\$FDGHP\WKHÀDJVKLS latest technologies in the last few months. school of the Eastern Region. By any Sixth Form Makeover P31 measure, we are well on the way to Open Thursday 1pm-4pm and Friday 9.30am – 2.30pm Inside Science P32 Our GCSE results were achieving this. Beauty @ Victory P33 We offer a full range of beauty treatments. Full price list available outstanding with a Above all, everything we do is for County Council Visit P34 Ring for appointments on 01603 742310 ex 3312 our students. -
Women in Engineering Fixing the Talent Pipeline
REPORT WOMEN IN ENGINEERING FIXING THE TALENT PIPELINE Amna Silim and Cait Crosse September 2014 © IPPR 2014 Institute for Public Policy Research ABOUT IPPR IPPR, the Institute for Public Policy Research, is the UK’s leading progressive thinktank. We are an independent charitable organisation with more than 40 staff members, paid interns and visiting fellows. Our main office is in London, with IPPR North, IPPR’s dedicated thinktank for the North of England, operating out of offices in Newcastle and Manchester. The purpose of our work is to conduct and publish the results of research into and promote public education in the economic, social and political sciences, and in science and technology, including the effect of moral, social, political and scientific factors on public policy and on the living standards of all sections of the community. IPPR 4th Floor 14 Buckingham Street London WC2N 6DF T: +44 (0)20 7470 6100 E: [email protected] www.ippr.org Registered charity no. 800065 This paper was first published in September 2014. © 2014 The contents and opinions in this paper are the authors’ only. NEW IDEAS for CHANGE CONTENTS Summary ............................................................................................................1 Introduction: Why should we care about the lack of women in engineering? .....2 1. The scale of the challenge in the UK .............................................................3 2. The choices girls make in education ..............................................................5 2.1 Choices at school ............................................................................................ 5 2.2 Choices in higher education ............................................................................. 6 2.3 Choices in employment.................................................................................... 7 3. Why do girls reject the idea of a career in engineering?..............................10 3.1 Perception of STEM subjects and engineering careers .................................. -
Radio 4 Listings for 21 – 27 August 2021 Page 1 of 16 SATURDAY 21 AUGUST 2021 SAT 06:07 Open Country (M000ytzz) Jay Rayner Hosts the Culinary Panel Show
Radio 4 Listings for 21 – 27 August 2021 Page 1 of 16 SATURDAY 21 AUGUST 2021 SAT 06:07 Open Country (m000ytzz) Jay Rayner hosts the culinary panel show. Sophie Wright, Tim A Fabric Landscape Anderson, Asma Khan and Dr Annie Gray share delectable SAT 00:00 Midnight News (m000yvbc) ideas and answer questions from the audience. The latest news and weather forecast from BBC Radio 4. Fashion designer and judge of The Great British Sewing Bee, Patrick Grant, has a dream: he wants to create a line of jeans This week, the panellists tell us their favourite recipes for that made in Blackburn. It sounds simple, but Patrick wants to go classic savoury nibble, the cheese straw. They also delve into SAT 00:30 Hello, Stranger by Will Buckingham (m000yvbf) the whole hog - growing the crop to make the fabric in the world of fresh peas and, when it comes to cooking with this Episode 5 Blackburn, growing the woad to dye it blue in Blackburn and small green vegetable, our panellists are not quite peas in a pod! finally processing the flax into linen and sewing it all When Will Buckingham's partner died, he coped with his grief together...in Blackburn. Nigerian food writer Yemisi Aribisala explains the significance by throwing his doors open to new people, and travelling alone of soup in Nigerian cuisine, and tells us what goes into the to far-flung places among strangers. 'Strangers are unentangled In this programme, the writer and broadcaster Ian Marchant perfect jollof rice. in our worlds and lives,' he writes, 'and this lack can lighten our travels to a tiny field of flax on the side of the Leeds and own burdens.' Starting from that experience of personal grief, Liverpool Canal, where Patrick and a group of passionate local Producer: Hannah Newton he draws on his knowledge as a philosopher and anthropologist, people are trying to make this dream a reality, and bring the Assistant Producer: Aniya Das as well as a keen and wide-roaming traveller, to explore the textile industry back to Blackburn. -
Radio 4 Listings for 29 February – 6 March 2020 Page 1 of 14
Radio 4 Listings for 29 February – 6 March 2020 Page 1 of 14 SATURDAY 29 FEBRUARY 2020 Series 41 SAT 10:30 The Patch (m000fwj9) Torry, Aberdeen SAT 00:00 Midnight News (m000fq5n) The Wilberforce Way with Inderjit Bhogal National and international news from BBC Radio 4 The random postcode takes us to an extraordinary pet shop Clare Balding walks with Sikh-turned-Methodist, Inderjit where something terrible has been happening to customers. Bhogal, along part of the Wilberforce Way in East Yorkshire. SAT 00:30 The Crying Book, by Heather Christle Inderjit created this long distance walking route to honour Torry is a deprived area of Aberdeen, known for addiction (m000fq5q) Wilberforce who led the campaign against the slave trade. They issues. It's also full of dog owners. In the local pet shop we Episode 5 start at Pocklington School, where Wilberforce studied, and discover Anna who says that a number of her customers have ramble canal-side to Melbourne Ings. Inderjit Bhogal has an died recently from a fake prescription drug. We wait for her Shedding tears is a universal human experience, but why and extraordinary personal story: Born in Kenya he and his family most regular customer, Stuart, to help us get to the bottom of it how do we cry? fled, via Tanzania, to Dudley in the West Midlands in the early - but where is he? 1960s. He couldn’t find anywhere to practice his Sikh faith so American poet Heather Christle has lost a dear friend to suicide started attending his local Methodist chapel where he became Producer/presenter: Polly Weston and must now reckon with her own depression. -
SPRING 2010 Dearwheaton
This version of Wheaton magazine does not contain the Class News section. s p r i n g 2 0 1 0 WHEATON The Litfin Legacy Continuity Amid Growth President Duane Litfin retires after 17 years Inside: Science Station Turns 75 • Remembering President Armerding • The Promise Report 150.WHEATON.EDU Wheaton College exists to help build the church and improve society worldwide by promoting the development of whole and effective Christians through excellence in programs of Christian higher education. This mission expresses our commitment to do all things “For Christ and His Kingdom.” volume 14 i s s u e 2 s PR i N G 2 0 1 0 6 a l u m n i n e w s departments 32 A Word with Alumni 2 Letters Open letter from Tim Stoner ’82, 5 News president of the Alumni Board 10 Sports 33 Wheaton Alumni Association News Association news and events 27 The Promise Report 37 Alumni Class News 56 Authors Books by Wheaton’s faculty; thoughts from published alumnus Walter Wolfram ’63 Cover photo: President Litfin enjoys the lively bustle of the Sports and A Sentimental Journey Recreation Complex that was built in 2000 as a result of the New 58 Century Challenge. The only “brick-and-mortar” part of that campaign, An archival reflection from an alumna the SRC features a large weight room, three gyms, a pool, elevated Faculty Voice running track, climbing wall, dance and fitness studio, and wrestling 60 room, as well as classrooms, conference rooms, and a physiology lab. Dr. Nadine Folino-Rorem mentors biology Dr. -
Deep Carbon Emissions from Volcanoes Michael R
Reviews in Mineralogy & Geochemistry Vol. 75 pp. 323-354, 2013 11 Copyright © Mineralogical Society of America Deep Carbon Emissions from Volcanoes Michael R. Burton Istituto Nazionale di Geofisica e Vulcanologia Via della Faggiola, 32 56123 Pisa, Italy [email protected] Georgina M. Sawyer Laboratoire Magmas et Volcans, Université Blaise Pascal 5 rue Kessler, 63038 Clermont Ferrand, France and Istituto Nazionale di Geofisica e Vulcanologia Via della Faggiola, 32 56123 Pisa, Italy Domenico Granieri Istituto Nazionale di Geofisica e Vulcanologia Via della Faggiola, 32 56123 Pisa, Italy INTRODUCTION: VOLCANIC CO2 EMISSIONS IN THE GEOLOGICAL CARBON CYCLE Over long periods of time (~Ma), we may consider the oceans, atmosphere and biosphere as a single exospheric reservoir for CO2. The geological carbon cycle describes the inputs to this exosphere from mantle degassing, metamorphism of subducted carbonates and outputs from weathering of aluminosilicate rocks (Walker et al. 1981). A feedback mechanism relates the weathering rate with the amount of CO2 in the atmosphere via the greenhouse effect (e.g., Wang et al. 1976). An increase in atmospheric CO2 concentrations induces higher temperatures, leading to higher rates of weathering, which draw down atmospheric CO2 concentrations (Ber- ner 1991). Atmospheric CO2 concentrations are therefore stabilized over long timescales by this feedback mechanism (Zeebe and Caldeira 2008). This process may have played a role (Feulner et al. 2012) in stabilizing temperatures on Earth while solar radiation steadily increased due to stellar evolution (Bahcall et al. 2001). In this context the role of CO2 degassing from the Earth is clearly fundamental to the stability of the climate, and therefore to life on Earth. -
My Main Research Interests Centre on the Science Behind Volcanoes and Volcanic Behaviour
PROF. TAMSIN A. MATHER Department of Earth Sciences, University of Oxford, South Parks Road, Oxford OX1 3AN, UK. Tel.: +44 (0)1865 282125 [email protected] My main research interests centre on the science behind volcanoes and volcanic behaviour. My motivation is to understand volcanoes as (a) natural hazards, (b) a key planetary scale process throughout geological time, playing a key role in change but vital for maintaining habitability and (c) natural resources (e.g., geothermal power and the development of ore deposits). I use techniques including satellite Earth Observation, remote and direct measurements of volcanic gas/aerosol, field mapping and petrological and geochemical analysis. I have also studied the emissions from an oil depot fire (Buncefield 2005) and am generally interested in the global mercury cycle as well as other biogeochemical cycles. Post-doctoral employment (Maternity leave: June-December 2007 and January-July 2010) 2014–present: Professor of Earth Sciences, University of Oxford. 2006–2014: Lecturer/Academic Fellow in Physics & Chemistry of the Earth and Environment, Dept of Earth Sciences, University of Oxford & Fellow, University College. 2005–2009: Royal Society Dorothy Hodgkin research fellow, awarded at Cambridge and moved to Oxford (Volcanic volatile emissions: from lithosphere to atmosphere) 2005: NERC fellow Parliamentary Office of Science and Technology (Report title: Carbon capture and storage) Education and Qualifications 2001–2004: Ph.D., Dept of Earth Sciences, University of Cambridge, ‘Near-source chemistry of tropospheric volcanic plumes’. NERC funded. Awarded December 2004. 1999–2000: M.Phil. (History and Philosophy of Science), University of Cambridge. Distinction. 1995–1999: M.Sci. -
Royal Society, 1985
The Public Understanding of Science Report of a Royal Society adhoc Group endorsed by the Council of the Royal Society The Royal Society 6 Carlton House Terrace London SWlY 5AG CONTENTS Page Preface 5 Summary 6 1. Introduction 7 2. Why it matters 9 3. The present position 12 4. Formal education 17 5. The mass media 2 1 6. ' The scientific community 24 7. Public lectures, children's activities, museums and libraries 27 8. Industry 29 9. Conclusions and recommendations 31 Annexes A. List of those submitting evidence B. Visits and seminars C. Selected bibliography PREFACE This report was prepared by an ad hoc group under the chairmanship of Dr W.F. Bodmer, F.R.S.; it has been endorsed by the Council of the Royal Society. It deals with an issue that is important not only, or even mainly, for the scientific community but also for the nation as a whole and for each individual within it. More than ever, people need some understanding of science, whether they are involved in decision-making at a national or local level, in managing industrial companies, in skilled or semi-skilled employment, in voting as private citizens or in making a wide range of personal decisions. In publishing this report the Council hopes that it will highlight this need for an overall awareness of the nature of science and, more particularly, of the way that science and technology pervade modern life, and that it will generate both debate and decisions on how best they can be fostered. The report makes a number of recommendations. -
Department of Physics Review
The Blackett Laboratory Department of Physics Review Faculty of Natural Sciences 2008/09 Contents Preface from the Head of Department 2 Undergraduate Teaching 54 Academic Staff group photograph 9 Postgraduate Studies 59 General Departmental Information 10 PhD degrees awarded (by research group) 61 Research Groups 11 Research Grants Grants obtained by research group 64 Astrophysics 12 Technical Development, Intellectual Property 69 and Commercial Interactions (by research group) Condensed Matter Theory 17 Academic Staff 72 Experimental Solid State 20 Administrative and Support Staff 76 High Energy Physics 25 Optics - Laser Consortium 30 Optics - Photonics 33 Optics - Quantum Optics and Laser Science 41 Plasma Physics 38 Space and Atmospheric Physics 45 Theoretical Physics 49 Front cover: Laser probing images of jet propagating in ambient plasma and a density map from a 3D simulation of a nested, stainless steel, wire array experiment - see Plamsa Physics group page 38. 1 Preface from the Heads of Department During 2008 much of the headline were invited by, Ian Pearson MP, the within the IOP Juno code of practice grabbing news focused on ‘big science’ Minister of State for Science and (available to download at with serious financial problems at the Innovation, to initiate a broad ranging www.ioppublishing.com/activity/diver Science and Technology Facilities review of physics research under sity/Gender/Juno_code_of_practice/ Council (STFC) (we note that some the chairmanship of Professor Bill page_31619.html). As noted in the 40% of the Department’s research Wakeham (Vice-Chancellor of IOP document, “The code … sets expenditure is STFC derived) and Southampton University). The stated out practical ideas for actions that the start-up of the Large Hadron purpose of the review was to examine departments can take to address the Collider at CERN.