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Phylogeny and Phylogenetic Taxonomy of Dipsacales, with Special Reference to Sinadoxa and Tetradoxa (Adoxaceae)
PHYLOGENY AND PHYLOGENETIC TAXONOMY OF DIPSACALES, WITH SPECIAL REFERENCE TO SINADOXA AND TETRADOXA (ADOXACEAE) MICHAEL J. DONOGHUE,1 TORSTEN ERIKSSON,2 PATRICK A. REEVES,3 AND RICHARD G. OLMSTEAD 3 Abstract. To further clarify phylogenetic relationships within Dipsacales,we analyzed new and previously pub- lished rbcL sequences, alone and in combination with morphological data. We also examined relationships within Adoxaceae using rbcL and nuclear ribosomal internal transcribed spacer (ITS) sequences. We conclude from these analyses that Dipsacales comprise two major lineages:Adoxaceae and Caprifoliaceae (sensu Judd et al.,1994), which both contain elements of traditional Caprifoliaceae.Within Adoxaceae, the following relation- ships are strongly supported: (Viburnum (Sambucus (Sinadoxa (Tetradoxa, Adoxa)))). Combined analyses of C ap ri foliaceae yield the fo l l ow i n g : ( C ap ri folieae (Diervilleae (Linnaeeae (Morinaceae (Dipsacaceae (Triplostegia,Valerianaceae)))))). On the basis of these results we provide phylogenetic definitions for the names of several major clades. Within Adoxaceae, Adoxina refers to the clade including Sinadoxa, Tetradoxa, and Adoxa.This lineage is marked by herbaceous habit, reduction in the number of perianth parts,nectaries of mul- ticellular hairs on the perianth,and bifid stamens. The clade including Morinaceae,Valerianaceae, Triplostegia, and Dipsacaceae is here named Valerina. Probable synapomorphies include herbaceousness,presence of an epi- calyx (lost or modified in Valerianaceae), reduced endosperm,and distinctive chemistry, including production of monoterpenoids. The clade containing Valerina plus Linnaeeae we name Linnina. This lineage is distinguished by reduction to four (or fewer) stamens, by abortion of two of the three carpels,and possibly by supernumerary inflorescences bracts. Keywords: Adoxaceae, Caprifoliaceae, Dipsacales, ITS, morphological characters, phylogeny, phylogenetic taxonomy, phylogenetic nomenclature, rbcL, Sinadoxa, Tetradoxa. -
Noteworthy Collections
48 THE MICHIGAN BOTANIST Vol. 52 NOTEWORTHY COLLECTIONS MICHIGAN Viburnum sieboldii Miq. Adoxaceae Siebold’s Arrowwood. Significance of the Report. First known naturalized occurrence in Michigan of a potentially invasive species. Previous Knowledge. Viburnum sieboldii is a large shrub or small tree native to Japan, where it occurs in thickets in lowlands and low mountains (Ohwi 1965). It was introduced to eastern North America in the late nineteenth century as a landscape ornamental, and is known to escape occasionally from cultivation in its introduced range (Kunstler 1993, Gleason and Cronquist 1991). Prior to this collection, naturalized populations of V. sieboldii were known in North America primarily from the Mid-Atlantic region of the United States, from Vir - ginia north to New York and Massachusetts (USDA NRCS 2014), although more recently it has also been reported from eastern and southern Ohio (Vincent et al. 2011). Escaped populations are known to establish in a variety of habitats, in - cluding mesic forests, stream edges, and suburban parks, and have been reported as abundant or invasive in New Jersey, New York, and Pennsylvania (Kunstler 1993, DeCandido and Lamont 2004, Pennsylvania Department of Conservation and Natural Resources 2014, Central New Jersey Invasive Species Strike Team 2014). On the other hand, Widrlechner and Iles (2002) found that V. sieboldii was at relatively low risk of naturalizing in Iowa because of climatic differences between there and the native habitat of V. sieboldii in Japan. However, it is un - known whether or how ornamental cultivars may differ from native genotypes in invasive potential. The genus Viburnum was long included in the Caprifoliaceae, but is now considered to belong to the Adoxaceae. -
Appendix 6: Invasive Plant Species
USDA Forest Service Understanding i-Tree – Appendix 6: Invasive Plant Species APPENDIX 6 Invasive Plant Species The following is a list of invasive tree and shrub species by state that are included in i-Tree database (version 6). Each list of invasive species is followed by the reference of the source which were obtained circa 2014. Some of the Web addresses are no longer working; some have been relocated to alternative sites. State-specific invasive species lists will be updated in the future. Alabama Ailanthus altissima Lonicera japonica Poncirus trifoliate Albizia julibrissin Lonicera maackii Pyrus calleryana Ardisia crenata Lonicera morrowii Rosa bracteata Cinnamomum camphora Lonicera x bella Rosa multiflora Elaeagnus pungens Mahonia bealei Triadica sebifera Elaeagnus umbellata Melia azedarach Vernicia fordii Ligustrum japonicum Nandina domestica Wisteria sinensis Ligustrum lucidum Paulownia tomentosa Ligustrum sinense Polygonum cuspidatum Alabama Invasive Plant Council. 2007. 2007 plant list. Athens, GA: Center for Invasive Species and Ecosystem Health, Southeast Exotic Pest Plant Council. http://www.se-eppc.org/ alabama/2007plantlist.pdf Alaska Alnus glutinosa Lonicera tatarica Sorbus aucuparia Caragana arborescens Polygonum cuspidatum Cytisus scoparius Prunus padus Alaska National Heritage Program. 2014. Non-Native plant data. Anchorage, AK: University of Alaska Anchorage. http://aknhp.uaa.alaska.edu/botany/akepic/non-native-plant-species- list/#content Arizona Alhagi maurorum Rhus lancea Tamarix parviflora Elaeagnus angustifolia Tamarix aphylla Tamarix ramosissima Euryops multifidus Tamarix chinensis Ulmus pumila Arizona Wildland Invasive Plant Working Group. 2005. Invasive non-native plants that threaten wildlands in Arizona. Phoenix, AZ: Southwest Vegetation Management Association https:// www.swvma.org/wp-content/uploads/Invasive-Non-Native-Plants-that-Threaten-Wildlands-in- Arizona.pdf (Accessed Sept 3. -
Viburnum Sieboldii Siebold Viburnum1 Edward F
Fact Sheet ST-662 October 1994 Viburnum sieboldii Siebold Viburnum1 Edward F. Gilman and Dennis G. Watson2 INTRODUCTION Most often grown as a large multi-trunked shrub or small tree, Siebold Viburnum only reaches about 20 feet high (occasionally to 30 feet) and 10 to 12 feet wide, creating an upright silhouette with short, firm branches (Fig. 1). The two to six-inch-long by 1.5 to 3-inch-wide, dark green leaves give off a pungent odor when crushed, and have been known to give an occasional autumn display of red or purple color although typically there is no fall color. In late May, the plant is covered with three to six-inch-diameter clusters of off-white, tiny blossoms. These blooms are followed by small, half-inch, pinkish-red fruits which ripen to dark blue or black from August to October, and are held above the foliage, making them any easy target for the birds who find them a delectable treat. There are extremely showy for a period of about two weeks. Figure 1. Mature Siebold Viburnum. GENERAL INFORMATION Availability: generally available in many areas within its hardiness range Scientific name: Viburnum sieboldii Pronunciation: vye-BER-num see-BOLE-dee-eye DESCRIPTION Common name(s): Siebold Viburnum Family: Caprifoliaceae Height: 20 to 30 feet USDA hardiness zones: 4 through 7 (Fig. 2) Spread: 10 to 12 feet Origin: not native to North America Crown uniformity: irregular outline or silhouette Uses: hedge; wide tree lawns (>6 feet wide); Crown shape: upright medium-sized tree lawns (4-6 feet wide); Crown density: moderate recommended for buffer strips around parking lots or Growth rate: medium for median strip plantings in the highway; near a deck Texture: coarse or patio; narrow tree lawns (3-4 feet wide); specimen; sidewalk cutout (tree pit); residential street tree; no proven urban tolerance 1. -
Trees, Shrubs, and Perennials That Intrigue Me (Gymnosperms First
Big-picture, evolutionary view of trees and shrubs (and a few of my favorite herbaceous perennials), ver. 2007-11-04 Descriptions of the trees and shrubs taken (stolen!!!) from online sources, from my own observations in and around Greenwood Lake, NY, and from these books: • Dirr’s Hardy Trees and Shrubs, Michael A. Dirr, Timber Press, © 1997 • Trees of North America (Golden field guide), C. Frank Brockman, St. Martin’s Press, © 2001 • Smithsonian Handbooks, Trees, Allen J. Coombes, Dorling Kindersley, © 2002 • Native Trees for North American Landscapes, Guy Sternberg with Jim Wilson, Timber Press, © 2004 • Complete Trees, Shrubs, and Hedges, Jacqueline Hériteau, © 2006 They are generally listed from most ancient to most recently evolved. (I’m not sure if this is true for the rosids and asterids, starting on page 30. I just listed them in the same order as Angiosperm Phylogeny Group II.) This document started out as my personal landscaping plan and morphed into something almost unwieldy and phantasmagorical. Key to symbols and colored text: Checkboxes indicate species and/or cultivars that I want. Checkmarks indicate those that I have (or that one of my neighbors has). Text in blue indicates shrub or hedge. (Unfinished task – there is no text in blue other than this text right here.) Text in red indicates that the species or cultivar is undesirable: • Out of range climatically (either wrong zone, or won’t do well because of differences in moisture or seasons, even though it is in the “right” zone). • Will grow too tall or wide and simply won’t fit well on my property. -
Ethnobotanical Importance of Some Highly Medicinal Plants of District Muzaffarabad, Pakistan with Special Reference to the Species of the Genus Viburnum
IOSR Journal of Pharmacy and Biological Sciences (IOSR-JPBS) e-ISSN: 2278-3008, p-ISSN:2319-7676. Volume 6, Issue 2 (May. – Jun. 2013), PP 53-66 www.iosrjournals.org Ethnobotanical Importance of Some Highly Medicinal plants of District Muzaffarabad, Pakistan with special reference to the Species of the Genus Viburnum Zahid Iqbal Awan1, Habib-ur-Rehman1, Ashfaq Ahmed Awan2, Fiaz Aziz Minhas1and Mohammad Nasir Khan2. 1(Department of Chemistry, University of Azad Jammu and Kashmir, Muzaffarabad 13100, Pakistan). 2(Department of Botany, University of Azad Jammu and Kashmir, Muzaffarabad 13100, Pakistan). Abstract: An ethnobotanical exploration was carried out in Muzaffarabad and its adjoining areas including Jhelum Valley of the District Muzaffarabad during 2010-2011. All the plants with the ethnobotanical importance were identified and segregated separately. The region is entirely mountainous, having sub-tropical to dry temperate climate with distinct seasonal variations. This study mainly focused on the information regarding traditional uses of plants over the years by local inhabitants. The informations were then confirmed by Hakims and the old people of the areas. During the survey informations were collected from various sites, i.e. Noon Bagla, Rahim Kot, Danna Kachilee, Kot Terhala, Sanwarrian, Chikar, Chikothi, Kathiee, Qazi Nag, Rashian, Daokhun, Mojee, Lamnian, Nardaggian, Pandu, Hatian Balla, Ghahi Dopatta, Chinnari, Rabanee, Bani Hafiz, Domel, Hattian Dopatta, Khanssian, Nandi Ka Sar, Sing Paharee,Nari Bela, Khalla Butt and Leepa. The plants were used medicinally and for other purposes. The investigations resulted that usually one plant or a mixture of two or more plant is used. The unplanned exploitation had resulted in the loss of medicinally important plant species. -
Chimonanthus Praecox: a Tions to Circulation Manager, Arnoldia, Arnold Arboretum, 125 Arborway, Boston, MA 02130- Redolence of China 3500
The Magazine of the Arnold Arboretum VOLUME 71 • NUMBER 3 The Magazine of the Arnold Arboretum VOLUME 71 • NUMBER 3 • 2014 CONTENTS Arnoldia (ISSN 0004–2633; USPS 866–100) 2 Untangling the Twisted Tale of is published quarterly by the Arnold Arboretum Oriental Bittersweet of Harvard University. Periodicals postage paid Peter Del Tredici at Boston, Massachusetts. Subscriptions are $20.00 per calendar year 19 Magnolia virginiana: Ephemeral domestic, $25.00 foreign, payable in advance. Courting for Millions of Years Remittances may be made in U.S. dollars, by Juan M. Losada check drawn on a U.S. bank; by international money order; or by Visa, Mastercard, or American 28 Wish You Were Here Express. Send orders, remittances, requests to Nancy Rose purchase back issues, change-of-address notices, and all other subscription-related communica- 36 Chimonanthus praecox: A tions to Circulation Manager, Arnoldia, Arnold Arboretum, 125 Arborway, Boston, MA 02130- Redolence of China 3500. Telephone 617.524.1718; fax 617.524.1418; David Yih e-mail [email protected] Arnold Arboretum members receive a subscrip- Front cover: Fruit of Oriental bittersweet (Celas- tion to Arnoldia as a membership benefit. To trus orbiculatus), a notoriously invasive vine. become a member or receive more information, Photo by Peter Del Tredici. please call Wendy Krauss at 617.384.5766 or email [email protected] Inside front cover: Read about the complex flowering and pollination process of sweetbay Postmaster: Send address changes to magnolia (Magnolia virginiana) starting on page Arnoldia Circulation Manager 19. Photo of M. virginiana × virginiana ‘Milton’, The Arnold Arboretum accession 779-87-A, by Michael S. -
Information to Users
INFORMATION TO USERS This manuscript has been reproduced from the microfilm master UMI films the text directly from the original or copy submitted. Thus, some thesis and dissertation copies are in typewriter face, while others may be from any type of computer printer. The quality of this reproduction is dependent upon the quality of the copy submitted. Broken or indistinct print, colored or poor quality illustrations and photographs, print bleedthrough, substandard margins, and improper alignment can adversely affect reproduction. In the unlikely event that the author did not send UMI a complete manuscript and there are missing pages, these will be noted. Also, if unauthorized copyright material had to be removed, a note will indicate the deletion. Oversize materials (e.g., maps, drawings, charts) are reproduced by sectioning the original, beginning at the upper left-hand comer and continuing from left to right in equal sections with small overlaps. Photographs included in the original manuscript have been reproduced xerographically in this copy. Higher quality 6" x 9" black and white photographic prints are available for any photographs or illustrations appearing in this copy for an additional charge. Contact UMI directly to order. ProQuest Information and Learning 300 North Zeeb Road, Ann Arbor, Ml 48106-1346 USA 800-521-0600 Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. Reproduced with permission of the copyright owner. Further reproduction prohibited without permission. SODIUM CHLORIDE TOLERANCE OF SELECTED HERBACEOUS PERENNIALS A N D THE EFFECTS OF SODIUM CHLORIDE O N OSM OTIC ADJUSTMENT A N D IO NIC UPTAKE IN THREE SPECIES OF HERBACEOUS PERENNIALS DISSERTATION Presented in Partial Fulfillment of the Requirements of the Degree Doctor of Philosophy in the Graduate School of The Ohio State University By Laura M. -
Design and Development Manual (Adobe PDF)
Design and Development Manual Updated 2021 Native and Adaptive Plant Lists Section Page # Landscaping 1 Native and Adaptive Plant List 1-23 Large Trees 1-4 Medium Trees 5 Small Trees 6-8 Large Shrubs 9-11 Small Shrubs 12-14 Parking Lot Shade Trees 15 Parking Lot Screening Shrubs 16 Trees Under Power Lines 17-18 Suggested Type A Buffer Trees & Shrubs 19-21 High-Quality Shade Trees 22-23 RCA Establishment Requirements 24-25 Tree Planting Guidelines 26 Planting in Special Situations 27 Tree Pruning 28 Recommended Plant List for BMPs 29-31 Mulch Standards 32-33 Protection Fencing Standard 34 Transportation 35-40 Typical Asphalt Greenway 35 Typical Keyed Concrete 36 Safety Railing Details 37 Locking Bollard Detail 38 Accessible Parking Spaces 39 Bicycle Parking 40 Site Details 41 Dumpster Enclosure Standard 41 Small Town Character Residential Design Standards 42-49 Architectural Features 43 Decorative Features 44 Residential Standards-Roofs 45 Residential Standards-Facades 46 Residential Standards-Windows 47 Residential Standards-Sides facing public streets 48 Residential Standards-Entryways & Materials 49 Native and Adaptive Plant Lists The Native and Adaptive Plant Lists are in chart form and include descriptions of trees and shrubs. Native species shown are native to the eastern half of the United States. These plant lists are not intended to be all-inclusive. The intent of these lists is to encourage the use of landscape plants that are hardy in Apex and exhibit tolerance of urban conditions. Plants not on the native and adaptive plant list may be used with approval from the Planning Department. -
A Bayesian Approach for Evaluating the Impact of Historical Events on Rates of Diversification
A Bayesian approach for evaluating the impact of historical events on rates of diversification Brian R. Moorea,1 and Michael J. Donoghueb,1 aDepartment of Integrative Biology, University of California, Berkeley, CA 94720; and bDepartment of Ecology and Evolutionary Biology, Yale University, New Haven, CT 06520 Contributed by Michael J. Donoghue, November 29, 2008 (sent for review June 22, 2008) Evolutionary biologists often wish to explore the impact of a rates that relies on cross-validation predictive densities, which particular historical event (e.g., the origin of a novel morphological effectively asks ‘‘How diverse would the effected lineage be if the trait, an episode of biogeographic dispersal, or the onset of an inferred event had no impact on rates of diversification?’’ The ecological association) on rates of diversification (speciation minus cross-validation predictive densities approach provides a versatile extinction). We describe a Bayesian approach for evaluating the inference framework (under both hypothesis-testing and data- correlation between such events and differential rates of diversi- exploration scenarios) for investigating the influence of diverse fication that relies on cross-validation predictive densities. This types of historical events (whether unique or replicated), while approach exploits estimates of the marginal posterior probability simultaneously accommodating various sources of uncertainty. We for the rate of diversification (in the unaffected part of the tree) illustrate this approach with two empirical analyses, first exploring and the marginal probability for the timing of the event to the impact of fruit evolution on rates of diversification in the plant generate a predictive distribution of species diversity that would group Adoxaceae and then investigating the influence of biogeo- be expected had the event not occurred. -
Early Detection of Invasive Species: Surveillance, Monitoring, and Rapid Response Eastern Rivers and Mountains Network and Northeast Temperate Network
National Park Service U.S. Department of the Interior Natural Resource Program Center Early Detection of Invasive Species: Surveillance, Monitoring, and Rapid Response Eastern Rivers and Mountains Network and Northeast Temperate Network Natural Resource Report NPS/ERMN/NRR—2010/196 ON THE COVER Spotted knapweed (Centaurea stoebe ssp. micranthos). Photograph by: Jennifer Stingelin Keefer. Early Detection of Invasive Species: Surveillance, Monitoring, and Rapid Response Eastern Rivers and Mountains Network and Northeast Temperate Network Natural Resource Data Series NPS/ERMN/NRR—2010/196 Jennifer Stingelin Keefer1, Matthew R. Marshall2, and Brian R. Mitchell3 1The Pennsylvania State University School of Forest Resources 309 Forest Resources Laboratory University Park, Pennsylvania 16802 2National Park Service Eastern Rivers and Mountains Network 420 Forest Resources Building University Park, Pennsylvania 16802 3National Park Service Northeast Temperate Network Marsh-Billings-Rockefeller NHP 54 Elm Street Woodstock, Vermont 05091 May 2010 U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The National Park Service, Natural Resource Program Center publishes a range of reports that address natural resource topics of interest and applicability to a broad audience in the National Park Service and others in natural resource management, including scientists, conservation and environmental constituencies, and the public. The Natural Resource Report Series is used to disseminate high-priority, current natural resource management information with managerial application. The series targets a general, diverse audience, and may contain NPS policy considerations or address sensitive issues of management applicability. All manuscripts in the series receive the appropriate level of peer review to ensure that the information is scientifically credible, technically accurate, appropriately written for the intended audience, and designed and published in a professional manner. -
Correlates of Diversification in the Plant Clade Dipsacales
vol. 170, supplement the american naturalist august 2007 Correlates of Diversification in the Plant Clade Dipsacales: Geographic Movement and Evolutionary Innovations Brian R. Moore* and Michael J. Donoghue† Department of Ecology and Evolutionary Biology, Yale University, including morphological, behavioral, and physiological New Haven, Connecticut 06520 novelties—and extrinsic events—including episodes of biogeographic or climatic change—have been hypothe- sized to promote shifts in diversification rate by altering the probability of speciation and/or extinction. A com- abstract: We explore patterns of diversification in the plant clades prehensive understanding of the causes of diversification Adoxaceae and Valerianaceae (within Dipsacales), evaluating corre- therefore requires incorporation of both elements. Sur- lations between biogeographic change (i.e., movements into new prisingly, the relative impact of both biotic and abiotic areas), morphological change (e.g., the origin of putative key in- factors on diversification rates has seldom been evaluated novations associated with vegetative and reproductive characters), in an explicitly phylogenetic context. and shifts in rates of diversification. Our findings indicate that rates of diversification in these plants tend to be less tightly correlated In plants, the search for intrinsic correlates of diversi- with the evolution of morphological innovations but instead exhibit fication has tended to focus on a few putative key inno- a pronounced correlation with movement into new geographic areas, vations relating to floral morphology, growth form, and particularly the dispersal of lineages into new mountainous regions. fruit type/dispersal mechanism. The role of extrinsic fac- The interdependence among apparent novelties (arising from their tors in plant diversification has received less attention; nested phylogenetic distribution) and the correlation between mor- however, an important class of abiotic factors pertains to phological and biogeographic change suggests a complex history of diversification in Dipsacales.