DOCSLIB.ORG

(Eemian) Hydrographic Conditions in the Southeastern Baltic Sea, NE

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
(Eemian) Hydrographic Conditions in the Southeastern Baltic Sea, NE ARTICLE IN PRESS Quaternary International 130 (2005) 3–30 Last Interglacial (Eemian) hydrographic conditions in the southeastern Baltic Sea,NE Europe,based on dinoflagellate cysts Martin J. Heada,*,Marit-Solveig Seidenkrantz b,Zofia Janczyk-Kopikowa c, Leszek Marksc,d,Philip L. Gibbard a a Godwin Institute for Quaternary Research, Department of Geography, University of Cambridge, Downing Place, Cambridge CB2 3EN, UK b Department of Earth Sciences, University of Aarhus, DK-8000 Arhus( C, Denmark c Polish Geological Institute, Rakowiecka 4, PL-00-975 Warsaw, Poland d Institute of Geology, Warsaw University, Zwirki i Wigury 93, PL-02-089 Warsaw, Poland Available online 20 July 2004 Abstract A rich organic-walled dinoflagellate cyst and pollen record from the Licze borehole in northern Poland has been used to reconstruct the hydrographic history of the southeastern Baltic Sea during the Last Interglacial (Eemian Stage,Late Pleistocene). Warm,saline waters (ca. 10–15 psu) entered the site from the North Sea within the first few hundred years of the Eemian, corresponding to the Pinus–Betula (E1) or Pinus–Betula–Ulmus (E2) regional pollen assemblage zones (RPAZ). By about 300 years (beginning of RPAZ E3),dinoflagellate cyst assemblages were already indicating summer sea-surface salinities in excess of about 15 psu and temperatures that perhaps exceeded 27C. Warm and saline conditions of 15–20 psu or more,at least twice present levels, persisted throughout the early Eemian. A rise in sea level at Licze appears to correlate with a similar event in eastern Denmark,as both coincide with the increase in Corylus (ca. 750 years into the interglacial). This sea-level rise might therefore have a basinwide extent,and appears to correspond to an opening of the Danish Belts. There is little if any evidence of arctic waters throughout the sequence. Whereas dinoflagellate cysts reflect sustained high salinites within the upper water column,a concomitant increase in abundance of the chlorococcalean alga Pediastrum within the Carpinus–Corylus–Alnus (E5) RPAZ indicates an escalating freshwater input,presumably from the proto-Vistula whose mouth was nearby. This suggests the development of a thin,seasonal, low-salinity surface layer below which dinoflagellates lived in more saline waters. Increasing fluvial influence suggests shallowing through RPAZ E5. This study is the first to document dinoflagellate cysts from the Eemian of the southeastern Baltic Sea,and reveals a flora with distinctive Lusitanian/Mediterranean affinities. r 2004 Elsevier Ltd and INQUA. All rights reserved. 1. Introduction temperatures and salinities soon exceeded those of today,and in addition to a narrow seaway across The Last Interglacial (equivalent to the Eemian Stage, northern Germany,two main portals developed: one and approximately to Marine Isotope Substage 5e; connecting to the North Sea in the Danish area,and Kukla et al.,2002 ) began at about 130 ka with the another that linked the Baltic Sea to the White Sea by retreat of glaciers that had extended over much of means of a seaway across the Karelian Isthmus in northern Europe in Saalian times. As a consequence the Russia (Fig. 1). This Karelian seaway was until recently Baltic Sea region was left isostatically depressed,and thought to have permitted free exchange between the rising global sea level at the start of the Eemian led Baltic and White seas (e.g., Donner,1995,fig. 6.1 ),but is swiftly to the establishment of marine conditions now considered to have been too restricted for this to (Liivrand,1984,1991 ; Kalnina,2001 ). Surface water have happened (Funder et al.,2002 ). The present study focuses on marine,organic-walled *Corresponding author. Tel.: +44-1223-339751; fax: +44-1223- dinoflagellate cysts and other aquatic palynomorphs in the 0 00 0 00 333392. Licze borehole (19 07 45 E,53 44 45 N),located about E-mail address: [email protected] (M.J. Head). 5 km to the west of Prabuty in northern Poland (Figs. 1 1040-6182/$ - see front matter r 2004 Elsevier Ltd and INQUA. All rights reserved. doi:10.1016/j.quaint.2004.04.027 ARTICLE IN PRESS 4 M.J. Head et al. / Quaternary International 130 (2005) 3–30 GulfGulf ofof GdaGGdañskdañsksk n oo ag a l GdaGGda–skda–sksk tul VistulaVis lagoon Szkar pawa g n Vistula a io l t lb a Elblag ElblagE v le ElevationE Nogat Tczew Sztum Nowiny N Obrzynowo Kwidzyn Prabuty Licze Fig. 1. Eemian hydrography and geography of the Baltic Sea region during maximum inundation in the early Eemian (based on Funder km et al.,2002 ). The present study area is marked,along with other 0 5 10 15 20 Eemian sites mentioned in the text. RK=Ristinge Klint,M=Mom- mark. Fig. 2. Sketch of the Lower Vistula region,northern Poland with the marine limit of the Eemian sea and location of the Licze,Nowiny and Obrzynowo boreholes (solid dots) and nearby towns (open dots). The extent of marine Eemian deposits (horizontal shading) and coeval fluvial deposits (stippling) is also shown. and 2). The specific aims of our study are to understand the nature and timing of marine transgression into the southeastern Baltic Sea during the early Eemian,and to assess whether the Karelian seaway influenced surface water conditions by facilitating the inflow of arctic waters. Summer upper-water-column salinity and temperature are reconstructed accordingly,but because the proto-Vistula 2. Dinoflagellate cysts entered the sea nearby during Eemian times,it has been necessary to differentiate as far as possible between local Dinoflagellates are almost exclusively unicellular and regional influences on hydrography. Pollen stratigra- protists,and include both autotrophic and heterotrophic phy and its correlation to the well-dated pollen record at species (Fensome et al.,1993 ). Dinoflagellates are BispingeninnorthernGermany(Muller,1974. ; Field et al., mostly planktonic,but unlike diatoms have a motile 1994) provides a tentative chronostratigraphic control for stage in their life cycle that allows them to control and the Licze borehole. optimise their position in the water column (Levan- This study is part of a larger investigation into the dowsky and Kaneta,1987 ). Many dinoflagellates Eemian of both the Licze core and a nearby core at produce a resting cyst (as a strategy for over-wintering Obrzynowo (Fig. 2). The mollusc faunas have been in mid to high latitudes) whose wall is generally very published by Makowska (2001) whereas publications on resistant to degradation,and the geological record of the foraminifera,ostracods and diatoms are forth- dinoflagellates is based on these species (Head,1996 ). coming. Previous studies of dinoflagellate cysts from Owing to the sensitivity of dinoflagellates to salinity, the Eemian of the Baltic Sea are limited to the Gulf of temperature,and nutrient levels over a wide range of Bothnia (Eriksson et al.,1999 ),Denmark ( Head and environmental parameters,and to the high preserva- Gibbard,2000 ) and to a brief general account (Head bility of the cysts,the Quaternary record of dino- et al.,2001 ) and short taxonomic study (Head,2003 ). flagellate cysts is becoming increasingly used for The present work is therefore the first to document reconstructing sea-surface conditions (Dale,1996 ; Eemian dinoflagellates in detail from the southeastern Rochon et al.,1999 ; Matthiessen and de Vernal,2001 ; Baltic. Dale and Dale,2002 ). ARTICLE IN PRESS M.J. Head et al. / Quaternary International 130 (2005) 3–30 5 3. Present hydrography and Rewal (Krzyszkowski et al.,1999 ). These deposits consist of fine-grained sand to sandy silt and shell The present day Baltic Sea is one of the world’s largest detritus,and represent a beach or lagoonal facies. bodies of brackish water,with surface salinities gen- However,it is in the Lower Vistula region that erally not exceeding 10 psu. Summer sea-surface tem- Eemian marine sequences are most complete. Here, peratures are generally above 19C over most of the evidence from shoreface and deeper-water molluscs, southern and eastern Baltic Sea (Bundesamt fur. particularly from Elblag and the Nowiny borehole Seeschiffahrt und Hydrographie,2002 ),largely due to (Makowska,1986 ; Funder et al.,2002 ; Fig. 2),depicts water stratification. The Gulf of Gdansk is characterised a protected embayment in which sea-surface salinities by a highly dynamic hydrography (Pastuszak,1995 ), and temperatures were considerably higher than today, and is strongly influenced by freshwater outflow from resulting from an early connection to the North Sea. In the Vistula,the largest freshwater source entering the the Nowiny borehole,marine incursion began at the Baltic Sea in the area. Surface water salinity reaches onset of the Quercus–Fraxinus–Ulmus regional pollen about 8 psu,but lower salinities averaging about 4.0 psu assemblage zone (RPAZ E3) of Mamakowa (1988, are found close to the mouth of the Vistula. Sea-surface 1989) and lasted until some time into the Picea–Abies– temperatures in the Gulf of Gdansk reach between 19C Alnus zone (RPAZ E6) (Makowska,1986 ). Molluscs, and 24C during summer (Nowacki and Jarosz,1998 ). including the foreshore/tidal zone species Retusa umbi- In the outer part of the Gulf of Gdansk during summer, licata,indicate that salinity culminated in RPAZ E3 a thermocline is established at about 20 m and halocline where it periodically rose above 30 psu,which is more at about 70 m,but in the inner parts of the bay a than 20 psu higher than present,although surface-water freshened surface layer 1 or 2 m deep results from salinity was in general probably not higher than 25 psu. discharge from the Vistula (Nowacki and Jarosz,1998 ). High salinities persisted from RPAZ E3 until the end of the Carpinus–Corylus–Alnus zone (RPAZ E5; Makows- ka,1986 ; Funder et al.,2002 ). It is also noteworthy that 4. Geological and palaeoenvironmental background the occurrence of Lusitanian mollusc species indicates a winter sea-surface temperature of about 9C,which is An embayment of the Baltic Sea occupied the Lower 6C warmer than today (Funder et al.,2002 ).
Load more

Recommended publications
  • Kenai National Wildlife Refuge Species List, Version 2018-07-24
    Kenai National Wildlife Refuge Species List, version 2018-07-24 Kenai National Wildlife Refuge biology staff July 24, 2018 2 Cover image: map of 16,213 georeferenced occurrence records included in the checklist. Contents Contents 3 Introduction 5 Purpose............................................................ 5 About the list......................................................... 5 Acknowledgments....................................................... 5 Native species 7 Vertebrates .......................................................... 7 Invertebrates ......................................................... 55 Vascular Plants........................................................ 91 Bryophytes ..........................................................164 Other Plants .........................................................171 Chromista...........................................................171 Fungi .............................................................173 Protozoans ..........................................................186 Non-native species 187 Vertebrates ..........................................................187 Invertebrates .........................................................187 Vascular Plants........................................................190 Extirpated species 207 Vertebrates ..........................................................207 Vascular Plants........................................................207 Change log 211 References 213 Index 215 3 Introduction Purpose to avoid implying
    [Show full text]
  • Late Weichselian and Holocene Shore Displacement History of the Baltic Sea in Finland
    Late Weichselian and Holocene shore displacement history of the Baltic Sea in Finland MATTI TIKKANEN AND JUHA OKSANEN Tikkanen, Matti & Juha Oksanen (2002). Late Weichselian and Holocene shore displacement history of the Baltic Sea in Finland. Fennia 180: 1–2, pp. 9–20. Helsinki. ISSN 0015-0010. About 62 percent of Finland’s current surface area has been covered by the waters of the Baltic basin at some stage. The highest shorelines are located at a present altitude of about 220 metres above sea level in the north and 100 metres above sea level in the south-east. The nature of the Baltic Sea has alter- nated in the course of its four main postglacial stages between a freshwater lake and a brackish water basin connected to the outside ocean by narrow straits. This article provides a general overview of the principal stages in the history of the Baltic Sea and examines the regional influence of the associated shore displacement phenomena within Finland. The maps depicting the vari- ous stages have been generated digitally by GIS techniques. Following deglaciation, the freshwater Baltic Ice Lake (12,600–10,300 BP) built up against the ice margin to reach a level 25 metres above that of the ocean, with an outflow through the straits of Öresund. At this stage the only substantial land areas in Finland were in the east and south-east. Around 10,300 BP this ice lake discharged through a number of channels that opened up in central Sweden until it reached the ocean level, marking the beginning of the mildly saline Yoldia Sea stage (10,300–9500 BP).
    [Show full text]
  • A Study on the Biological and Physiological Traits of Bactrocera Dorsalis, with Special Reference to Its Invasion Potential Into the Western Cape of South Africa
    A study on the biological and physiological traits of Bactrocera dorsalis, with special reference to its invasion potential into the Western Cape of South Africa. by Welma Pieterse Dissertation presented for the degree of Doctor of Philosophy (Agricultural Sciences) at Stellenbosch University Department of Conservation Ecology and Entomology, Faculty of AgriSciences Supervisor: Dr Pia Addison Co-supervisors: Prof John Terblanche Dr Aruna Manrakhan March 2018 Stellenbosch University https://scholar.sun.ac.za Declaration By submitting this dissertation electronically, I declare that the entirety of the work contained therein is my own, original work, that I am the sole author thereof (save to the extent explicitly otherwise stated) that reproduction and publication thereof by Stellenbosch University will not infringe any third party rights and that I have not previously in its entirety or in part submitted it for obtaining any qualification. Welma Pieterse Date: 26 February 2018 Copyright © 2018 Stellenbosch University All rights reserved Stellenbosch University https://scholar.sun.ac.za Summary Bactrocera dorsalis (Hendel) is of Asian origin and is present in the northern and north-eastern parts of South Africa, but is still absent in other areas of the country including the Western Cape Province. The Western Cape Province is the largest producer of deciduous fruit in South Africa, exporting 41% of the deciduous fruit grown in the province. South Africa earned about R7 billion in export revenue from deciduous fruit exports in 2015. Currently, Ceratitis capitata (Wiedemann) and Ceratitis rosa s.l. Karsch are economically the most important fruit fly species on deciduous fruit in the Western Cape Province of South Africa.
    [Show full text]
  • Y of the Eemian Baltic Sea Discoveries in GEOLOGIAN TUTKIMUSKESKUS GEOLOGICAL SURVEY of FINLAND Tutkimusraportti 102 Report of Investigation 102
    TUTKIMUSRAPORTTI REPORT OF INVESTIGATION 102 y of the Eemian Baltic Sea discoveries in GEOLOGIAN TUTKIMUSKESKUS GEOLOGICAL SURVEY OF FINLAND Tutkimusraportti 102 Report of Investigation 102 Tuulikki Gränlund THE DIATOM STRATIGRAPHY OF THE EEMIAN BALTIC SEA ON THE BASIS OF SEDIMENT DISCOVERIES IN OSTROBOTHNIA, FINLAND Espoo 1991 Grönlund, T. 1991. The diatom stratigraphy of the Eemian Baltic Sea on the basis of sediment discoveries in Ostrobothnia, Finland. Geological Survey of Finland, Report of Investigation 102, 26 pages and 16 figures. Sediments from interglacial sites in Ostrobothnia, western Finland have been in­ vestigated for their diatom content. All the sites are correlated with the Eemian interglacial stage. The diatom successions of two of the sites, Norinkylä (Teuva) and Viitala (Perä• seinäjoki), begin with freshwater diatoms indicating deposition in a freshwater (great lake) basin. A rich marine diatom flora is found overlying the sediments with fresh­ water diatoms. The other sites Evijärvi, Ollala (Haapavesi) and Ukonkangas (Kär• sämäki), have also a similar marine flora with Grammatophora oceanica, Hyalodiscus scoticus, Paralia sulcata and Rhabdonema arcuatum as dominant species . The flora also includes many diatoms that mainly thrive in highly saline water and have not been found in Finland earlier. In the Ollala and Norinkylä deposits, there is a tran­ sition from marine to freshwater diatoms implying isolation from the sea. The marine deposits at the sites are interpreted as having been deposited in the Eemian Baltic Sea, and the freshwater sediments found beneath the marine deposits in a lake which occupied the Baltic basin and preceded the Eemian Baltic Sea. The history of the Eem!an Baltic Sea is discussed and a comparison is also made with the Holocene Baltic basin.
    [Show full text]
  • Integrated Ocean Drilling Program Expedition 347 Preliminary Report
    Integrated Ocean Drilling Program Expedition 347 Preliminary Report Baltic Sea Basin Paleoenvironment Paleoenvironmental evolution of the Baltic Sea Basin through the last glacial cycle Platform operations 12 September–1 November 2013 Onshore Science Party 22 January–20 February 2014 Expedition 347 Scientists Published by Integrated Ocean Drilling Program Publisher’s notes Material in this publication may be copied without restraint for library, abstract service, educational, or personal research purposes; however, this source should be appropriately acknowledged. Core samples and the wider set of data from the science program covered in this report are under moratorium and accessible only to Science Party members until 20 February 2015. Published by International Ocean Discovery Program for the Integrated Ocean Drilling Program and prepared by the European Consortium of Ocean Research Drilling (ECORD) Science Operator. Funding for the program is provided by the following agencies: National Science Foundation (NSF), United States Ministry of Education, Culture, Sports, Science and Technology (MEXT), Japan European Consortium for Ocean Research Drilling (ECORD) Ministry of Science and Technology (MOST), People’s Republic of China Korea Institute of Geoscience and Mineral Resources (KIGAM) Australian Research Council (ARC) and GNS Science (New Zealand), Australian/New Zealand Con- sortium Ministry of Earth Sciences (MoES), India Coordination for Improvement of Higher Education Personnel, Brazil Disclaimer Any opinions, findings, and conclusions or recommendations expressed in this publication are those of the author(s) and do not necessarily reflect the views of the participating agencies, IODP, British Geological Survey, European Petrophysics Consortium, University of Bremen, or the authors’ institutions. Portions of this work may have been published in whole or in part in other International Ocean Discovery Program documents or publications.
    [Show full text]
  • The Baltic Sea Basin, Edited by Jan Harff, Svente Björck & Peer Hoth
    Sedimentary Record Book Review Accepted February 28, 2013 The Baltic Sea Basin, edited by Jan Harff, Svente Björck & Peer Hoth, 2011. Central and eastern European development studies. Springer-Verlag Berlin, Heidelberg, Germany. Hardcover, xiii + 449 pages. Price US$ 179.00, € 139.05. ISBN 978-3-642-17219-9. E-book price € 119.99. E-ISBN 978-3-642-17220-5. In many ways the Baltic Sea, situated in northern Europe, is a unique area. It is a shallow (55 m average depth), semi-enclosed, marginal sea. However, like fjords, it is composed of several basins separated by shallow sills. Consequently, the water exchange between the various parts, and between the Baltic and the North Sea is limited. It makes the Baltic the world’s largest brackish-water basin. There is still a debate going on about the origin of the Baltic Sea Basin; some consider it to be mostly an erosional structure, whereas others tend to see it as a tectonic depression. Its latest history encompasses the time-span after the Last Glacial Maximum when a large ice-marginal lake developed through a number of fresh-water and brackish stages into the modern brackish sea. During this evolution, large sea (lake) level changes occurred, which were partly due to glacio- isostatic movements. Observations of their impacts may serve as a kind of laboratory for the analysis of the effects of sea-level changes on various types of coast. Additionally, Baltic Sea sediments provide high-resolution records of climate and environmental changes during the Late Quaternary at a regional scale. Around 15,000 years ago, humans settled around the Baltic Sea and used the local ecosystem resources; they were also exposed, however, to site-specific natural hazards.
    [Show full text]
  • 3. Eemian Interglacial: 130 - 117 Ka Bp
    SE9800205 SKB rapport R-98-02 January 1998 Compilation of information on the climate and evaluation of the hydrochemical and isotopic composition during Late Pleistocene and Holocene Cecilia Andersson Intern KB Svensk Karnbranslehantering AB Swedish Nuclear Fuel and Waste Management Co SKB, Box 5864, S-102 40 Stockholm, Sweden Tel 08-665 28 00 Fax 08-661 57 19 Tel+46 8 665 28 00 Fax+46 8 661 57 19 ISSN 1402-3091 SKB Rapport R-98-02 COMPILATION OF INFORMATION ON THE CLIMATE AND EVALUATION OF THE HYDROCHEMICAL AND ISOTOPIC COMPOSITION DURING LATE PLEISTOCENE AND HOLOCENE Cecilia Andersson Intera KB January 1998 This report concerns a study which was conducted for SKB. The conclusions and viewpoints presented in the report are those of the author(s) and do not necessarily coincide with those of the client. COMPILATION OF INFORMATION ON THE CLIMATE AND EVALUATION OF THE HYDROCHEMICAL AND ISOTOPIC COMPOSITION DURING LATE PLEISTOCENE AND HOLOCENE 98-01-07 Cecilia Andersson Intera KB ABSTRACT This report summarises and evaluates some of the existing information on the Late Pleistocene and Holocene climates, i.e. the last 130 000 years. An estimation of the conditions at the Aspo island (southeast Sweden) has also been made during this time span. The knowledge about Late Pleistocene (Eemian Interglacial and Weichselian glacial) is not yet fully understood. There are still a lot of assumptions concerning this period and more information is needed to be able to establish the climatic conditions. This is not the case for the Weichselian deglaciation and the present interglacial, Holocene, for which the environmental conditions are quite certain.
    [Show full text]
  • Effects of Standard Management Practices On, and Faunistics of Native Prairies
    Conservation Biology Research Grants Program Division of Ecological Services Minnesota Department of Natural Resources EFFECTS OF STANDARD MANAGEMENT PRACTICES ON, AND FAUNISTICS OF NATIVE PRAIRIES: A study of three sites in western Minnesota. Report submitted to: Minnesota and Wisconsin Departments of Natural Resources, The Nature Conservancy, and the R. J. Kose foundation. David Rider, Systematic Entomologist Gerald Fauske, Research specialist Paul Tinerella, Graduate student Department of Entomology North Dakota State University Fargo, ND May 2000 ABSTRACT From 1995 through 1999 an ongoing study was conducted to examine the effects of standard prairie management practices (burning, grazing, haying) and unmanaged prairie, with regard to arthropod communities. Additional objectives were to gather baseline data on the arthropod species present on remnant prairies of western Minnesota, to identify rare species found on theses sites, and if possible, to determine prairie indicator species which might be used to identify dry, mesic, or wet prairies types in a manner similar to the plant indicator species used in botanical studies. At present, nearly 35,000 insect specimens have been pinned, labeled, and identified representing more than 750 species. As additional material is processed and identified, we expect the number of species found on these sites to more than double, perhaps even triple. This report provides a summary of work completed as of May, 2000, and includes species lists, information on species distributions (state records and extensions of known species ranges), presence of rare or state endangered species on these prairies, diversity indices for prairie sites, types, and management practices, and management histories of the study areas.
    [Show full text]
  • Proceedings of a Seminar on Sea Level Displacement and Bedrock Uplift, 10-11 June 2010, Pori, Finland
    Working Report 2011-07 Proceedings of a Seminar on Sea Level Displacement and Bedrock Uplift, 10-11 June 2010, Pori, Finland Editors: Ari T. K. Ikonen Tarmo Lipping January 2011 POSIVA OY Olkiluoto FI-27160 EURAJOKI, FINLAND Tel +358-2-8372 31 Fax +358-2-8372 3709 Working Report 2011-07 Proceedings of a Seminar on Sea Level Displacement and Bedrock Uplift, 10-11 June 2010, Pori, Finland Editors: Ari T. K. Ikonen Posiva Oy Tarmo Lipping Tampere University of Technology Pori Unit January 2011 Base maps: ©National Land Survey, permission 41/MML/11 Working Reports contain information on work in progress or pending completion. The conclusions and viewpoints presented in the report are those of author(s) and do not necessarily coincide with those of Posiva. ABSTRACT This working report is the proceedings of a seminar on Sea level displacement and bedrock uplift held on 10-11 June 2010 in Pori, Finland. The seminar included invited oral presentations, as well as poster presentations, addressing the causes and mechanisms, observations, modelling and implications of the sea level change and crustal uplift still continuing after the last glaciation in the Baltic Sea region. In the proceedings, a total of 14 papers are included, in addition to foreword and a summary of seminar discussions. Keywords: land uplift, post-glacial crustal rebound, global isostatic adjustment, sea level change, GIS modelling, radiocarbon dating, historical maps, archaeological findings, landscape reconstruction. Seminaari merenpinnan muutoksesta ja kallioperän kohoamisesta TIIVISTELMÄ Tähän työraporttiin on koostettu Porissa 10.-11. kesäkuuta 2010 pidetyn seminaarin "Sea level displacement and bedrock uplift" kirjallinen anti 14 seminaaripaperin, alkusanojen ja keskustelujen yhteenvedon muodossa.
    [Show full text]
  • WO 2016/092376 Al 16 June 2016 (16.06.2016) W P O P C T
    (12) INTERNATIONAL APPLICATION PUBLISHED UNDER THE PATENT COOPERATION TREATY (PCT) (19) World Intellectual Property Organization International Bureau (10) International Publication Number (43) International Publication Date WO 2016/092376 Al 16 June 2016 (16.06.2016) W P O P C T (51) International Patent Classification: HN, HR, HU, ID, IL, IN, IR, IS, JP, KE, KG, KN, KP, KR, A61K 36/18 (2006.01) A61K 31/465 (2006.01) KZ, LA, LC, LK, LR, LS, LU, LY, MA, MD, ME, MG, A23L 33/105 (2016.01) A61K 36/81 (2006.01) MK, MN, MW, MX, MY, MZ, NA, NG, NI, NO, NZ, OM, A61K 31/05 (2006.01) BO 11/02 (2006.01) PA, PE, PG, PH, PL, PT, QA, RO, RS, RU, RW, SA, SC, A61K 31/352 (2006.01) SD, SE, SG, SK, SL, SM, ST, SV, SY, TH, TJ, TM, TN, TR, TT, TZ, UA, UG, US, UZ, VC, VN, ZA, ZM, ZW. (21) International Application Number: PCT/IB20 15/002491 (84) Designated States (unless otherwise indicated, for every kind of regional protection available): ARIPO (BW, GH, (22) International Filing Date: GM, KE, LR, LS, MW, MZ, NA, RW, SD, SL, ST, SZ, 14 December 2015 (14. 12.2015) TZ, UG, ZM, ZW), Eurasian (AM, AZ, BY, KG, KZ, RU, (25) Filing Language: English TJ, TM), European (AL, AT, BE, BG, CH, CY, CZ, DE, DK, EE, ES, FI, FR, GB, GR, HR, HU, IE, IS, IT, LT, LU, (26) Publication Language: English LV, MC, MK, MT, NL, NO, PL, PT, RO, RS, SE, SI, SK, (30) Priority Data: SM, TR), OAPI (BF, BJ, CF, CG, CI, CM, GA, GN, GQ, 62/09 1,452 12 December 201 4 ( 12.12.20 14) US GW, KM, ML, MR, NE, SN, TD, TG).
    [Show full text]
  • Butterflies and Moths of the Yukon
    Butterflies and moths of the Yukon FRONTISPIECE. Some characteristic arctic and alpine butterflies and moths from the Yukon. Upper, males of the nymphalid butterflies Oeneis alpina Kurentzov (left) and Boloria natazhati (Gibson) (right), normally encountered on rocky tundra slopes; Middle, males of the alpine arctiid moths Pararctia yarrowi (Stretch) (left), typically on dry rocky slopes with willow, and Acsala anomala Benjamin (right), confined to the Yukon and Alaska and shown here on the characteristic dry rocky habitat of the lichen-feeding larvae; Lower, (left) female of the arctiid moth Dodia kononenkoi Chistyakov and Lafontaine from dry rocky tundra slopes, and (right) a mated pair of the noctuid moth Xestia aequeva (Benjamin), showing the reduced wings of the female. All species were photographed at Windy Pass, Ogilvie Mountains (see book frontispiece), except for B. natazhati (Richardson Mountains). Forewing length of these species is about 2 cm (first 3 species) and 1.5 cm (last 3). 723 Butterflies and Moths (Lepidoptera) of the Yukon J.D. LAFONTAINE and D.M. WOOD Biological Resources Program, Research Branch, Agriculture and Agri-Food Canada K.W. Neatby Bldg., Ottawa, Ontario, Canada K1A 0C6 Abstract. An annotated list of the 518 species of Lepidoptera known from the Yukon is presented with a zoogeographic analysis of the fauna. Topics discussed are: historical review of Yukon collecting and research; the expected size of the Yukon fauna (about 2000 species); zoogeographic affinities; special features of Yukon fauna (endemic species, disjunct species, biennialism, flightless species). There are 191 species of Lepidoptera (37% of the fauna) in the Yukon that occur in both Nearctic and Palaearctic regions.
    [Show full text]
  • Kenai National Wildlife Refuge Species List, Version 2017-10-17
    Kenai National Wildlife Refuge Species List, version 2017-10-17 Kenai National Wildlife Refuge biology staff October 17, 2017 2 Cover images represent changes to the checklist. Top left: Lemna mi- nor observed at Watson Lake, June 28, 2017 (http://www.inaturalist. org/observations/6903405). Image CC BY Matt Bowser. Top right: Ephemerella aurivillii from Skilak Lake Road, July 21, 2017 (http: //arctos.database.museum/media/10570401). Image CC0 Todd Eske- lin. Bottom left: Andromeda polifolia infected by the fungus Exoba- sidium karstenii, Headquarters Lake, July 18, 2017. (http://www. inaturalist.org/observations/7120482). Image CC BY Matt Bowser. Bottom right: Matteuccia struthiopteris near the Finger Lakes, Au- gust 16, 2017 (https://www.inaturalist.org/observations/7524787). Im- age CC BY Matt Bowser. Contents Contents 3 Introduction 5 Purpose............................................................ 5 About the list......................................................... 5 Acknowledgments....................................................... 5 Native species 7 Vertebrates .......................................................... 7 Invertebrates ......................................................... 24 Vascular Plants........................................................ 50 Bryophytes .......................................................... 65 Other Plants ......................................................... 69 Chromista........................................................... 70 Fungi ............................................................
    [Show full text]