DOCSLIB.ORG

Winged Seeds from the Early Miocene of the Mecsek Mts, W Hungary

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Winged Seeds from the Early Miocene of the Mecsek Mts, W Hungary Palaeobiodiversity and Palaeoenvironments https://doi.org/10.1007/s12549-020-00461-0 ORIGINAL PAPER Fossil Gordonia (s.l.)–like (Theaceae) winged seeds from the early Miocene of the Mecsek Mts, W Hungary Boglárka Erdei1 & Lilla Hably1 Received: 3 March 2020 /Revised: 5 May 2020 /Accepted: 21 September 2020 # The Author(s) 2021 Abstract Winged seeds were recovered from two sites of the late early Miocene (Karpatian) flora of Magyaregregy, Mecsek Mts, W Hungary. The seeds are assigned to the fossil-genus and species, Mecsekispermum gordonioides Hably and Erdei gen. nov. et sp. nov., and are tentatively related to the family Theaceae. Based on the overall character of the winged seeds and the isodiametric surface pattern of the seed coat, the seeds are most comparable with species of Gordonia J. Ellis (s.l.,) in Theeae (Laplacea Kunth or Polyspora Sweet). A comparison with winged seeds of other fossil genera, e.g. Saportaspermum Meyer and Manchester, and winged seeds of modern genera in various families is also given. The fossil flora is preserved in the fish scale-bearing clay marl belonging to the Feked Formation and Komló Claymarl Member and dated as Karpatian (late Burdigalian, standard chronostratigraphy). Keywords Fossil flora . Magyaregregy . Saportaspermum . Mecsekispermum . Fish scale-bearing clay marl . Karpatian Introduction Kentucky and Tennessee (Grote and Dilcher 1992; Martínez-Millán 2010). Winged fruits or seeds have frequently been reported in the The classification of the family Theaceae varied in many European fossil record. Disseminules having an apical or lat- authors, according to the characters emphasised (Prince and eral wing have been described variously as Cedrelospermum Parks 2001). In the Cronquist system, the family involves ~ 40 Saporta, Gordonia J. Ellis, Saportaspermum Meyer and genera and 600 species in four subfamilies (Cronquist 1981). Manchester, or Embothrites Unger and have been assigned Molecular data showed the family as non-monophyletic (e.g. to various plant groups, e.g. Ulmaceae, Theaceae, and Morton et al. 1996;Soltisetal.2000), and APG (1998)sug- Malvales (Unger 1850; Manchester 1987; Meyer and gested a narrow definition of the family with Theaceae s.str. Manchester 1997; Kvaček and Walther 1998; Kvaček and and a separate Ternstroemiaceae (Ternstroemieae in Wilde 2010). Fruits or seeds assigned to Theaceae are rare Pentaphylacaceae). Based on molecular data, three main line- not only in the European fossil record but also worldwide. ages of Theaceae s.str. were identified, i.e. Theeae, Grote and Dilcher (1989, 1992) gave a summary of fossil Stewartieae, and Gordonieae (Prince and Parks 2001). specimens reported as remains of Theaceae. The earliest fossil According to the same study, the monophyly of the genus fruits assigned to Gordonia (Gordonia lamkinensis Grote and Gordonia was not supported, and sequence data placed its Dilcher and Gordonia warmanensis Grote and Dilcher) were species in Gordonieae (Gordonia s. str.) and Theeae described from the middle Eocene Claiborne flora in (Laplacea Kunth and Polyspora Sweet). Although a common character of the seeds of Gordonia s.l. species is a prominent flattened apical wing, micromorphological study of the seed coat was used to corroborate the above placement of This article is a contribution to the special issue “Palaeobotanical contributions in honour of Volker Mosbrugger” Gordonia s.l. species (Gunathilake et al. 2015). Leaves of Theaceae have been reported from some locali- * Boglárka Erdei ties in the Northern Hemisphere fossil record during the [email protected] Cenozoic. Today the area of the family occupies subtropical- tropical Asia and America (Prince and Parks 2001); however, 1 Botanical Department, Hungarian Natural History Museum, palaeofloristic data indicate the presence of the family also in Könyves Kálmán krt. 40., Budapest 1087, Hungary Europe. Leaf records of Theaceae (s.l.) have been documented Palaeobio Palaeoenv in the European floras from the middle Eocene up to the late by the Budafa Formation, traditionally regarded as of Miocene. From the middle Eocene flora of Messel (Germany; Karpatian (late Burdigalian) age (Gyalog 1996;Gyalogand Wilde 1989) and from the early Oligocene flora of Flörsheim Budai 2004; Budai et al. 2015).TheBudafaFormation (Mainz Basin, Germany; Kvaček 2004), leaves of two genera, consisted of three members, the Komló Claymarl Member, Gordonia and Ternstroemites Berry, were mentioned. Leaves Pécsvárad Limestone Member, and the overlying Budafa preserved with cuticle were reported as Gordonia hradekensis Sandstone Members. The succession is overlain by (Kvaček and Bůžek) Bozukov and Palamarev, Gordonia Badenian (Langhian) Leitha limestones in the littoral zone pseudoknauensis Kvaček, and Ternstroemites floersheimensis and offshore sands and silts in the basins, locally with coal- Kvaček and Walther from the rich, thermophilous flora. From bearing swamp deposits along the shores. Based on the ab- the early Miocene flora of Hrádek (North Bohemia, sence of marine fossils in the Pécsvárad Limestone and in Czech Republic; Holy et al. 2012) and from the flora of the most part of the Komló Claymarl, Sebe et al. (2019)conclud- Cypris Shale (Western Bohemia, Czech Republic; Bůžek et al. ed that these rocks are lacustrine deposits. As suggested by the 1996), leaf remains of Gordonia hradekensis were mentioned. lake sediments and its fauna, instead of a system of smaller From the early Miocene (Ottnangian) flora of Köflach lakes, a contiguous water body, called “Lake Mecsek”,existed (Austria) ?Gordonia oberdorfensis Kovar-Eder and ?Cleyera in the area from the Karpatian up to the early Badenian ac- schilcheriana Kovar-Eder (Kovar-Eder and Meller 2001) cording to K/Ar age and biostratigraphic dating of the overly- were described based on leaf remains and preserved cuticle. ing marine deposits (Sebe et al. 2019). The Pécsvárad and From the late early/early middle Miocene flora of Parschlug Komló Members are both interpreted as lake deposits and (Austria; Kovar-Eder et al. 2004), there are also some leaves differ very much from the Budafa Sandstone Member; assigned to Theaceae, cf. ?Gordonia oberdorfensis and therefore, Sebe et al. (2019) united the two members as a Ternstroemites pereger (Unger)Kovar-EderandKvaček. separate formation, the Feked Formation. From the middle Miocene (Badenian) of the Satovcha Consequently, the “fish scale-bearing clay marl” Graben in the Western Rhodopes (Bulgaria), Bozukov and sediments preserving the plant fossils belong to the Feked Palamarev (1995) described a flora as a rich refuge of the Formation and Komló Claymarl Member, dated as Karpatian representatives of Theaceae (s.l.), e.g. Eurya aff. (late Burdigalian) (Sebe et al. 2019). It is composed of dark acuminatissima Merril and Chun, Gordonia hradekensis, grey, greenish-grey, massive, or laminated silty clay marl, Gordonia stefanovii Palamarev and Bozukov, Hartia calcareous silt, and fine sands, with numerous fish scales and palaeorhodopensis Bozukov and Palamarev, Stewartia bones, and sometimes with tuff interbeds. K/Ar dating mea- stefanovii Palamarev and Bozukov, and Stewartia surements from tuff interbeds within the Komló Claymarl submonadelpha Tanai and Onoe. In Europe, Theaceae pre- showed an age of 16.82 ± 0.65 Ma, i.e. Karpatian (late sumably survived up to the late Miocene or Pliocene based Burdigalian) (Sebe et al. 2019). on reports from the Sessenheim flora in France (Geissert et al. Palaeontological research of the Mecsek Mts dates back to 1990), Fritzlar (Hessen) in Germany (Hottenrott et al. 1996), decades, attested by numerous studies, e.g. nannoplankton and refuge areas of the Pannonian Basin. A rich late Miocene (Báldi-Beke 1963), foraminifers (Korecz-Laky 1968), and thermophilous flora was described from Mataschen (Austria, palynology (Nagy 1969). After floristic works published by Kovar-Eder and Hably 2006) with four species assigned to Staub (1882), Andreánszky (1955), and Pálfalvy (1953, 1961, Theaceae. Gordonia emanuelii Kovar-Eder, Gordonia 1964, 1967), Hably L. and her colleagues have contributed pannonica Kovar-Eder, Gordonia styriaca Kovar-Eder, and with studies to the knowledge on the Miocene flora of the Schima mataschensis Kovar-Eder were described based on Mecsek Mts (Hably 1992, 2001, 2002; Hably and Thiébaut leaf remains with cuticle. The flora of Magyaregregy also 2002). A monographic study of the plant fossils of yielded leaf remains described as Ternstroemites pereger, Magyaregregy is going to be published by Hably (2020). which may be related to the Theaceae (s.l.) family (Hably The Magyaregregy flora is extremely diverse with more than 2020). 100 taxa hitherto described, including Glyptostrobus europaeus (Brongniart) Unger, Zizyphus paradisiaca (Unger) Heer, Daphnogene cinnamomifolia (Brongniart) Geology Unger,Laurophyllumsp., Myrica lignitum (Unger) Saporta, Engelhardia orsbergensis (P.Wessel and C. O. Weber) The area of the Mecsek Mts is situated in the SW Pannonian Jähnichen, Mai and Walther, Cedrelospermum flichei Basin, on the Tisza-Dacia Megaunit, a tectonic block south of (Saporta) Hably and Thiébaut, C. aquense Saporta, Zelkova the mid-Hungarian Shear Zone. The Neogene stratigraphy of zelkovifolia (Unger) Bůžek and Kotlaba, legumes the Mecsek area was summarised by Hámor (1970), Chikán (Podocarpium podocarpum (A. Braun) Herendeen, (1991), and Barabás (2010, 2011). Widespread lower Leguminosae gen. et sp.), Ailanthus confucii Unger, and Miocene fluvial clastics (Szászvár
Load more

Recommended publications
  • O Babarc Község Önkormányzata 40 80 0 0 2 0 0 Liptód Község
    SZOCIÁLIS ÉS GYERMEKVÉDELMI Brado- Lily Brado- Kesz- Brado- Lily FŐIGAZGATÓSÁG Maszk man gél life tyű lin 1L 1L BARANYA MEGYEI KIRENDELTSÉG 1L 500ml 300ml O Babarc Község Önkormányzata 40 80 0 0 2 0 0 Liptód Község Önkormányzata 40 80 0 0 2 0 0 Versend Község Önkormányzata Kerekítve 100 200 0 0 4 0 0 Összesen 80 160 0 0 4 0 0 SZOCIÁLIS ÉS GYERMEKVÉDELMI Brado- Lily Brado- Kesz- Brado- Lily FŐIGAZGATÓSÁG Maszk man gél life tyű lin 1L 1L BARANYA MEGYEI KIRENDELTSÉG 1L 500ml 300ml O Baksa Község Önkormányzata 400 400 0 0 20 0 0 Adorjás Község Önkormányzata 40 80 0 0 2 0 0 Baranyahídvég Községi Önkormányzat 40 80 0 0 2 0 0 Bogádmindszent Község Önkormányzata 152 232 0 0 7 0 9 Hegyszentmárton Község Önkormányzata 152 232 0 0 7 0 9 Kisszentmárton Község Önkormányzata 40 80 0 0 2 0 0 Ózdfalu Község Önkormányzata 76 116 0 0 3 0 0 Sámod Község Önkormányzata 40 80 0 0 2 0 0 Tengeri Község Önkormányzata Téseny Község Önkormányzata 40 80 0 0 2 0 0 Kerekítve 1000 1400 0 0 47 0 18 Összesen 980 1380 0 0 47 0 18 SZOCIÁLIS ÉS GYERMEKVÉDELMI Brado- Lily Brado- Kesz- Brado- Lily FŐIGAZGATÓSÁG Maszk man gél life tyű lin 1L 1L BARANYA MEGYEI KIRENDELTSÉG 1L 500ml 300ml O Beremendi Gyermekjóléti és Szociális Társulás 1778 1941 14 38 77 10 36 Kistapolca Község Önkormányzata 40 80 0 0 2 0 0 Kerekítve 1800 2000 14 38 79 10 36 Összesen 1818 2021 14 38 79 10 36 SZOCIÁLIS ÉS GYERMEKVÉDELMI Brado- Lily Brado- Kesz- Brado- Lily FŐIGAZGATÓSÁG Maszk man gél life tyű lin 1L 1L BARANYA MEGYEI KIRENDELTSÉG 1L 500ml 300ml O Bicsérd Községi Önkormányzata 40 80 0 0 2 0 0
    [Show full text]
  • Act Cciii of 2011 on the Elections of Members Of
    Strasbourg, 15 March 2012 CDL-REF(2012)003 Opinion No. 662 / 2012 Engl. only EUROPEAN COMMISSION FOR DEMOCRACY THROUGH LAW (VENICE COMMISSION) ACT CCIII OF 2011 ON THE ELECTIONS OF MEMBERS OF PARLIAMENT OF HUNGARY This document will not be distributed at the meeting. Please bring this copy. www.venice.coe.int CDL-REF(2012)003 - 2 - The Parliament - relying on Hungary’s legislative traditions based on popular representation; - guaranteeing that in Hungary the source of public power shall be the people, which shall pri- marily exercise its power through its elected representatives in elections which shall ensure the free expression of the will of voters; - ensuring the right of voters to universal and equal suffrage as well as to direct and secret bal- lot; - considering that political parties shall contribute to creating and expressing the will of the peo- ple; - recognising that the nationalities living in Hungary shall be constituent parts of the State and shall have the right ensured by the Fundamental Law to take part in the work of Parliament; - guaranteeing furthermore that Hungarian citizens living beyond the borders of Hungary shall be a part of the political community; in order to enforce the Fundamental Law, pursuant to Article XXIII, Subsections (1), (4) and (6), and to Article 2, Subsections (1) and (2) of the Fundamental Law, hereby passes the following Act on the substantive rules for the elections of Hungary’s Members of Parliament: 1. Interpretive provisions Section 1 For the purposes of this Act: Residence: the residence defined by the Act on the Registration of the Personal Data and Resi- dence of Citizens; in the case of citizens without residence, their current addresses.
    [Show full text]
  • Differential Resistance of Gordonieae Trees to Phytophthora Cinnamomi
    HORTSCIENCE 44(5):1484–1486. 2009. Successful crosses of Franklinia · Schima produced the intergeneric hybrid ·Schimlinia (Ranney et al., 2003) and crosses of Frank- Differential Resistance of Gordonieae linia · Gordonia produced the intergeneric hybrid ·Gordlinia (Ranney and Fantz, 2006). Trees to Phytophthora cinnamomi However, little is known about the resistance 1 2,5 3 of related species and potential parents to Elisabeth M. Meyer , Thomas G. Ranney , and Thomas A. Eaker P. cinnamomi. The objective of this study Department of Horticultural Science, Mountain Horticultural Crops was to evaluate a collection of species, Research and Extension Center, North Carolina State University, 455 clones, and hybrids of Franklinia, Gordonia, Research Drive, Fletcher, NC 28732 and Schima for resistance to P. cinnamomi. 4 Kelly Ivors Materials and Methods Department of Plant Pathology, Mountain Horticultural Crops Research and Extension Center, North Carolina State University, 455 Research Drive, During the summer of 2008, seven taxa of Gordonieae trees were inoculated with Mills River, NC 28759 P. cinnamomi at the North Carolina State Additional index words. host plant resistance, disease resistance, Abies fraseri, Franklinia University Mountain Horticultural Crops alatamaha, Gordonia lasianthus, ·Gordlinia grandiflora, ·Schimlinia floribunda, Schima Research Station in Mills River, NC. These taxa included F. alatamaha, G. lasianthus, S. wallichii, Schima khasiana, Phytophthora cinnamomi khasiana, S. wallichii, ·Gordlinia H2004- Abstract. Trees in the Theaceae tribe Gordonieae are valuable nursery crops, but some of 024-008, ·Schimlinia H2002-022-083, and these taxa are known to be highly susceptible to root rot caused by Phytophthora ·Schimlinia H2002-022-084. The plants of cinnamomi Rands. The objective of this study was to evaluate a collection of Gordonieae the selected Gordonieae taxa were 5-month- taxa for resistance to this pathogen.
    [Show full text]
  • Agrosilvopastoral Systems in Northern Thailand and Northern Laos: Minority Peoples’ Knowledge Versus Government Policy
    Land 2014, 3, 414-436; doi:10.3390/land3020414 OPEN ACCESS land ISSN 2073-445X www.mdpi.com/journal/land/ Article Agrosilvopastoral Systems in Northern Thailand and Northern Laos: Minority Peoples’ Knowledge versus Government Policy Chalathon Choocharoen 1, Andreas Neef 2,*, Pornchai Preechapanya 3 and Volker Hoffmann 1 1 Institute for Social Sciences of the Agricultural Sector, Rural Communication and Extension (430a), University of Hohenheim, 70593 Stuttgart, Germany; E-Mails: [email protected] (C.C.); [email protected] (V.H.) 2 Center for Development Studies, School of Social Sciences, Faculty of Arts, University of Auckland, Auckland 1142, New Zealand 3 Queen Sirikit Botanic Garden, Mae Rim, Chiang Mai 50180, Thailand; E-Mail: [email protected] * Author to whom correspondence should be addressed; E-Mail: [email protected]; Tel.: +64-9-9233486; Fax: +64-9-3737439. Received: 28 January 2014; in revised form: 2 May 2014 / Accepted: 13 May 2014 / Published: 20 May 2014 Abstract: Traditional agrosilvopastoral systems have been an important component of the farming systems and livelihoods of thousands of ethnic minority people in the uplands of Mainland Southeast Asia. Drawing on a combination of qualitative and participatory inquiries in nine ethnic minority communities, this study emphasizes the complex articulation of local farmers’ knowledge which has been so far excluded from governmental development and conservation policies in the northern uplands of Thailand and Laos. Qualitative analysis of local knowledge systems is performed using the Agroecological Knowledge Toolkit (AKT5) software. Results show that ethnic minorities in the two countries perceive large ruminants to be a highly positive component of local forest agro-ecosystems due to their contribution to nutrient cycling, forest fire control, water retention, and leaf-litter dispersal.
    [Show full text]
  • Schima Wallichii (DC.) Korth
    Schima wallichii (DC.) Korth. Theaceae simartolu, schima, samak, needlewood, mang tan, Chinese guger tree, chilauni LOCAL NAMES Bengali (cheloni,mukria sal,makrisal); Burmese (laukya); Chinese (he,muhe,heshu,hemu,haozi haozi,gaobei,Chinese guger tree); English (needlewood,schima); Hindi (makusal,kanak,dieng-shyr-nagan,chilauni,nogabhe); Indonesian (seru,madang gatal,puspa); Javanese (medang gatal,seru); Lao (Sino- Tibetan) (‘mi,boun nak,‘khai sou); Malay (kelinchi padi,gegatal,medang gatal,gatal-gatal,samak); Nepali (sule-chilauni,aule-chilaune,chilaune,goe- chassi); Thai (thalo,champa dong,bunnak); Trade name (simartolu,Chinese guger tree,samak,needlewood,schima,mang tan,chilauni); Vietnamese (v[oos]i thu[oos]c) BOTANIC DESCRIPTION Schima wallichii is an evergreen, medium-sized to large tree growing to 47 m in height; bole cylindrical, branchless for up to 25 m, diameter up to 125 (max. 250) cm, with a steep buttresses rarely up to 1.8 m high; bark surface ruggedly cracked into small, thick, angular pieces, red-brown to dark grey; inner bark with skin-irritating fibres, bright red in colour. Leaves spiral, oblong to broadly elliptic, 6-13 x 3-5 cm; base wedge shaped; apex acute to acuminate; margin toothed; secondary veins 6-8 pairs; petioles about 3 mm long. Flowers solitary in axils at the apices of twigs, with 2 bracteoles, pentamerous; sepals subequal, persistent in fruit; petals connate at base, white, with a rosy flush; stamens many, adnate to the corolla base; anthers versatile; ovary superior, 5-locular with 2-6 ovules in each cell; style simple. Fruit a woody subglobose capsule, 2-3 cm in diameter, silky, opening by 5 valves; seeds winged all around.
    [Show full text]
  • Forest Structures, Composition, and Distribution on a Pacific Island, with Reference to Ecological Release and Speciation!
    Pacific Science (1991), vol. 45, no. 1: 28-49 © 1991 by University of Hawaii Press. All rights reserved Forest Structures, Composition, and Distribution on a Pacific Island, with Reference to Ecological Release and Speciation! YOSHIKAZU SHIMIZU2 AND HIDEO TABATA 3 ABSTRACT: Native forest and scrub of Chichijima, the largest island in the Bonins, were classified into five types based on structural features: Elaeocarpus­ Ardisia mesic forest, 13-16 m high, dominated by Elaeocarpus photiniaefolius and Ardisia sieboldii; Pinus-Schima mesic forest, 12-16 m high, consisting of Schima mertensiana and an introduced pine , Pinus lutchuensis; Rhaphiolepis­ Livistonia dry forest, 2-6 m high, mainly occupied by Rhaphiolepis indica v. integerrima; Distylium-Schima dry forest, 3-8 m high, dominated by Distylium lepidotum and Schima mertensiana; and Distylium-Pouteria dry scrub, 0.3­ 1.5 m high, mainly composed of Distylium lepidotum. A vegetation map based on this classification was developed. Species composition and structural features of each type were analyzed in terms of habitat condition and mechanisms of regeneration. A group of species such as Pouteria obovata, Syzgygium buxifo­ lium, Hibiscus glaber, Rhaphiolepis indica v. integerrima, and Pandanus boninen­ sis, all with different growth forms from large trees to stunted shrubs, was subdominant in all vegetation types. Schima mertensiana , an endemic pioneer tree, occurred in both secondary forests and climax forests as a dominant canopy species and may be an indication of "ecological release," a characteristic of oceanic islands with poor floras and little competitive pressure. Some taxonomic groups (Callicarpa, Symplocos, Pittosporum, etc.) have speciated in the under­ story of Distylium-Schima dry forest and Distylium-Pouteria dry scrub.
    [Show full text]
  • Vegetation Changes from the Late Pleistocene Through the Holocene from Three Areas of Archaeological Significance in Thailand Joyce C
    ARTICLE IN PRESS Quaternary International 113 (2004) 111–132 Vegetation changes from the late Pleistocene through the Holocene from three areas of archaeological significance in Thailand Joyce C. Whitea,*, Daniel Pennyb, Lisa Kealhoferc, BernardMaloney d,{ a University of Pennsylvania Museum, 3260 South Street, Philadelphia, PA 19104-6324, USA b School of Geosciences, Department of Archaeology and the Australian Key Centre for Microscopy and Microanalysis, University of Sydney, Madsen Building FO9, 2006 NSW, Australia c Department of Anthropology and Sociology/Environmental Studies Program, Santa Clara University, Santa Clara, CA 95053, USA d School of Archaeology and Palaeoecology, The Queen’s University, Belfast BT7 1NN, Northern Ireland, UK Abstract Reconstruction of the environmental history of mainlandSoutheast Asia from the late Pleistocene is a relatively recent endeavor. Beginning in the mid-1990s, lacustrine sediments in Thailand with deposits dating from the late Pleistocene have been cored and analyzedfor palaeoenvironmental indicators.The three cores reportedhere were extractedby the ThailandPalaeoenvironment Project, whose objective was to retrieve empirical data on vegetation and sedimentary sequences that can in turn be related to the growing archaeological record from this part of monsoonal Asia. This evidence, along with data from other recently analyzed cores, is beginning to develop a picture of regionally diverse environmental/cultural trajectories. Possible relationships between the environmental changes and cultural and/or climatic impacts are discussed. r 2003 Elsevier LtdandINQUA. All rights reserved. 1. Introduction cene hadbeen extractedandanalyzedfor palaeovegeta- tion andother environmental changes. Those records For research on the evolution of Asia’s monsoonal that hadbeen publishedwere only a few millennia in system, Thailandholdsa key geographic position.
    [Show full text]
  • The Karpatian (Late Early Miocene) Flora of the Mecsek Area
    Acta Palaeobotanica 60(1), 51–122, 2020 e-ISSN 2082-0259 https://doi.org/10.35535/acpa-2020-0003 ISSN 0001-6594 The Karpatian (late early Miocene) flora of the Mecsek area LILLA HABLY Botanical Department, Hungarian Natural History Museum, H-1431 Budapest, P.O. box 137, Hungary; e-mail: [email protected] Received 9 September 2019; accepted for publication 28 January 2020 In remembrance of the beautiful summers spent collecting with my family in the Mecsek Mountains ABSTRACT. A rich macroflora has been collected from Karpatian (late early Miocene) layers of the Mecsek Mts during recent decades. The bulk of the fossil assemblage consists of leaves and also fruits of angiosperms. Among the more than a hundred taxa, several endemic species were described: Leguminocarpum mecsekense Andreánszky, Ailanthus mecsekensis Hably, Nyssa gyoergyi sp. nov., Nyssa gergoei sp. nov., Nyssa sp. 1, Clematis csabae sp. nov., Gordonia sp. and Carpolithes gergoei Hably et Erdei sp. nov. Many taxa were last recorded in the Carpathian Basin, e.g. Cedrelospermum, Ziziphus. Other taxa appeared in this flora, e.g. Quercus kubinyii, Podocarpium podocarpum, Liquidambar europaea and Populus populina, and later became dominant in the middle Miocene (Sarmatian) floras or even in the late Miocene (Pannonian) floras. Four main vegetation types were determined. The most significant types are subxerophytic vegetation showing high diversity, swamp veg- etation, riparian vegetation, and a vegetation type growing in habitats with higher rainfall. Thermophilous flora elements are dominant in the assemblage, although “arctotertiary” species also appear. The floristic character of the flora supports the results of an earlier quantitative climate analysis of the Magyaregregy flora, accord- ing to which mean annual temperature was 15.6–16.6°C and coldest-month and warmest-month temperatures were 5–6.2°C and 24.7–27.9°C, respectively.
    [Show full text]
  • A Brief Nomenclatural Review of Genera and Tribes in Theaceae Linda M
    Aliso: A Journal of Systematic and Evolutionary Botany Volume 24 | Issue 1 Article 8 2007 A Brief Nomenclatural Review of Genera and Tribes in Theaceae Linda M. Prince Rancho Santa Ana Botanic Garden, Claremont, California Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons, and the Ecology and Evolutionary Biology Commons Recommended Citation Prince, Linda M. (2007) "A Brief Nomenclatural Review of Genera and Tribes in Theaceae," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 24: Iss. 1, Article 8. Available at: http://scholarship.claremont.edu/aliso/vol24/iss1/8 Aliso 24, pp. 105–121 ᭧ 2007, Rancho Santa Ana Botanic Garden A BRIEF NOMENCLATURAL REVIEW OF GENERA AND TRIBES IN THEACEAE LINDA M. PRINCE Rancho Santa Ana Botanic Garden, 1500 North College Ave., Claremont, California 91711-3157, USA ([email protected]) ABSTRACT The angiosperm family Theaceae has been investigated extensively with a rich publication record of anatomical, cytological, paleontological, and palynological data analyses and interpretation. Recent developmental and molecular data sets and the application of cladistic analytical methods support dramatic changes in circumscription at the familial, tribal, and generic levels. Growing interest in the family outside the taxonomic and systematic fields warrants a brief review of the recent nomenclatural history (mainly 20th century), some of the classification systems currently in use, and an explanation of which data support various classification schemes. An abridged bibliography with critical nomen- clatural references is provided. Key words: anatomy, classification, morphology, nomenclature, systematics, Theaceae. INTRODUCTION acters that were restricted to the family and could be used to circumscribe it.
    [Show full text]
  • 127 Acta Biol. Debr. Oecol. Hung 20: 127–144, 2009 CONTRIBUTION
    127 Acta Biol. Debr. Oecol. Hung 20: 127–144, 2009 CONTRIBUTION TO THE RIFFLE BEETLE FAUNA OF HUNGARY (COLEOPTERA: ELMIDAE) Z. KÁLMÁN – A. KÁLMÁN – Z. CSABAI* University of Pécs, Department of General and Applied Ecology, Ifjúság útja 6., Pécs H-7624, Hungary, *Corresponding author, e-mail: [email protected] ADATOK MAGYARORSZÁG KARMOSBOGÁR-FAUNÁJÁHOZ (COLEOPTERA: ELMIDAE) KÁLMÁN ZOLTÁN – KÁLMÁN ANDRÁS – CSABAI ZOLTÁN Pécsi Tudományegyetem, Általános és Alkalmazott Ökológia Tanszék, 7624 Pécs, Ifjúság útja 6. ABSTRACT: Altogether 147 new faunistical records of 11 riffle beetle species are given originated from 61 sampling sites in Hungary. A comprehensive view on Hungarian distribution of riffle beetle species is given by summarizing all available locality data on UTM grid maps. Key words: distribution, faunistics, UTM grid maps, Hungary KIVONAT: A közleményben Magyarország 61 mintavételi helyéről előkerült 11 karmosbogárfaj 147 faunisztikai adatát adjuk közre. UTM hálótérképek segítségével bemutatjuk a hazai karmosbogárfajok ismert hazai előfordulásait. Kulcsszavak: elterjedés, faunisztika, UTM hálótérképek, Magyarország Introduction Riffle beetle fauna of Hungary is poorly known due to the identification problems. External morphological characters are slightly different and strongly variable, such species level identification could be safely done based on checking male genitalia only. Hungarian checklist of the riffle beetle fauna was published by KOVÁCS & MERKL (2005) and a new species was added by CSABAI & SÁR (2007). Based on these works, family Elmidae is represented with 17 species of 8 genera in Hungary. ENTZ et al. (1954) mentioned Normandia nitens (P.J.W.Müller, 1817) (as Riolus nitens), but voucher specimens are not available in the Hungarian collections, so we cannot take this species into consideration in the Hungarian fauna.
    [Show full text]
  • BARANYA Megye Autóbusz-Hálózata
    Jelmagyarázat / Legend 5707 Lengyel Országos autóbuszjáratok útvonala Mekényes BARANYA megye autóbusz-hálózata 5707 Route of long-distance bus services Dombóvár 5707 Nagyhajmás Regionális autóbuszjáratok útvonala Bus network of Baranya county 5707 Route of regional bus services 5611 Pécs 5707 Komló 2021. április 11-től Dombóvár 1652 5924 Mohács térségi központtal Dombóvár 5611 5707 Siklós és Harkány with regional hub Pápa Szárász 1564 Sásd 5706 Mágocs 1652 Vajszló és Sellye Veszprém 1740 5623 5708 5611 5611 Szigetvár Siófok 5608 5623 5707 Magyaregregy 5702 5706 1134 Budapest Egyéb (szomszédos megyék gócpontjairól induló) vonalak Dombóvár Dombóvár 5708 5707 Egyházaskozár 1566 Kecskemét Other services (of the neighbouring counties) 5516 5702 Budapest 50 Bikal 5924 1568 Kiskőrös A vastagabb vonallal jelölt szakaszon az autóbuszok sűrűbben, 1652 1707 Győr a vékonyabb vonallal jelölt jelölt szakaszon ritkábban közlekednek az autóbuszok 5708 5611 1843 Tatabánya Thicker line represents more frequent services, Vásárosdombó Hegyhátmaróc Tófű 1136 Budapest Alsómocsolád 5623 5625 Bonyhád Thinner line represents less frequent services Jágónak Gerényes 1652 Szeged 5706 5626 Siófok 5605 5740 Vásárosdombó 5710 5707 5434 5624 Szekszárd 5887 Autóbuszvonal száma Keszthely 1576 5702 Mágocs- 5630 Szekszárd Kiskeresztúr 5702 Köblény 5924 5485 5710 Bonyhád Bus line number Kaposvár 5606 5930 5931 5932 40 Alsómocsolád Szalatnak 5631 Nagydorog Gödre 5924 Mohács Autóbuszvonal száma a járat végállomásánál 1578 Meződ 1652 Ág Szászvár 5634 Paks 5887 Kárász- 5856
    [Show full text]
  • (Theaceae) Species: Insights Into DNA Barcoding and Phylogeny
    RESEARCH ARTICLE Comparative chloroplast genomes of eleven Schima (Theaceae) species: Insights into DNA barcoding and phylogeny Xiang-Qin Yu1,2, Bryan T. Drew3, Jun-Bo Yang1, Lian-Ming Gao2*, De-Zhu Li1* 1 Germplasm Bank of Wild Species in Southwest China, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan, China, 2 Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan, China, 3 Department of a1111111111 Biology, University of Nebraska, Kearney, NE, United States of America a1111111111 a1111111111 * [email protected] (DZL); [email protected] (LMG) a1111111111 a1111111111 Abstract Schima is an ecologically and economically important woody genus in tea family (Thea- ceae). Unresolved species delimitations and phylogenetic relationships within Schima limit OPEN ACCESS our understanding of the genus and hinder utilization of the genus for economic purposes. Citation: Yu X-Q, Drew BT, Yang J-B, Gao L-M, Li In the present study, we conducted comparative analysis among the complete chloroplast D-Z (2017) Comparative chloroplast genomes of (cp) genomes of 11 Schima species. Our results indicate that Schima cp genomes possess eleven Schima (Theaceae) species: Insights into DNA barcoding and phylogeny. PLoS ONE 12(6): a typical quadripartite structure, with conserved genomic structure and gene order. The size e0178026. https://doi.org/10.1371/journal. of the Schima cp genome is about 157 kilo base pairs (kb). They consistently encode 114 pone.0178026 unique genes, including 80 protein-coding genes, 30 tRNAs, and 4 rRNAs, with 17 dupli- Editor: Genlou Sun, Saint Mary's University, cated in the inverted repeat (IR).
    [Show full text]