Getting to Grips with Skippers Jonathan Wallace Dingy Skipper Erynnis Tages
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Self-Repair and Self-Cleaning of the Lepidopteran Proboscis
Clemson University TigerPrints All Dissertations Dissertations 8-2019 Self-Repair and Self-Cleaning of the Lepidopteran Proboscis Suellen Floyd Pometto Clemson University, [email protected] Follow this and additional works at: https://tigerprints.clemson.edu/all_dissertations Recommended Citation Pometto, Suellen Floyd, "Self-Repair and Self-Cleaning of the Lepidopteran Proboscis" (2019). All Dissertations. 2452. https://tigerprints.clemson.edu/all_dissertations/2452 This Dissertation is brought to you for free and open access by the Dissertations at TigerPrints. It has been accepted for inclusion in All Dissertations by an authorized administrator of TigerPrints. For more information, please contact [email protected]. SELF-REPAIR AND SELF-CLEANING OF THE LEPIDOPTERAN PROBOSCIS A Dissertation Presented to the Graduate School of Clemson University In Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy ENTOMOLOGY by Suellen Floyd Pometto August 2019 Accepted by: Dr. Peter H. Adler, Major Advisor and Committee Co-Chair Dr. Eric Benson, Committee Co-Chair Dr. Richard Blob Dr. Patrick Gerard i ABSTRACT The proboscis of butterflies and moths is a key innovation contributing to the high diversity of the order Lepidoptera. In addition to taking nectar from angiosperm sources, many species take up fluids from overripe or sound fruit, plant sap, animal dung, and moist soil. The proboscis is assembled after eclosion of the adult from the pupa by linking together two elongate galeae to form one tube with a single food canal. How do lepidopterans maintain the integrity and function of the proboscis while foraging from various substrates? The research questions included whether lepidopteran species are capable of total self- repair, how widespread the capability of self-repair is within the order, and whether the repaired proboscis is functional. -
European Skipper Butterfly (Thymelicus Lineola) Associated
European Skipper Butterfly ( Thymelicus lineola ) Associated with Reduced Seed Development of Showy Lady’s-slipper Orchid (Cypripedium reginae ) PETER W. H ALL 1, 5, P AUL M. C ATLING 2, P AUL L. M OSQUIN 3, and TED MOSQUIN 4 124 Wendover Avenue, Ottawa, Ontario K1S 4Z7 Canada 2170 Sanford Avenue, Ottawa, Ontario K2C 0E9 Canada 3Research Triangle International, 3040 East Cornwallis Road, P.O. Box 12194, Raleigh, North Carolina 27709-2194 USA 43944 McDonalds Corners Road, Balderson, Ontario K0G 1A0 Canada 5Corresponding author: [email protected] Hall, Peter W., Paul M. Catling, Paul L. Mosquin, and Ted Mosquin. 2017. European Skipper butterfly ( Thymelicus lineola ) associated with reduced seed development of Showy Lady’s-slipper orchid ( Cypripedium reginae ). Canadian Field- Naturalist 131(1): 63 –68. https://doi.org/10.22621/cfn.v 131i 1. 1952 It has been suggested that European Skipper butterflies ( Thymelicus lineola ) trapped in the lips of the Showy Lady’s-slipper orchid ( Cypripedium reginae ) may interfere with pollination. This could occur through blockage of the pollinator pathway, facilitation of pollinator escape without pollination, and/or disturbance of the normal pollinators. A large population of the orchid at an Ottawa Valley site provided an opportunity to test the interference hypothesis. The number of trapped skippers was compared in 475 post-blooming flowers with regard to capsule development and thus seed development. The presence of any skippers within flowers was associated with reduced capsule development ( P = 0.0075), and the probability of capsule development was found to decrease with increasing numbers of skippers ( P = 0.0271). The extent of a negative effect will depend on the abundance of the butterflies and the coincidence of flowering time and other factors. -
General Methods Will Be Outlined in Chapter 2
The Biodiversity of Abandoned Farmland Tom Fayle Gonville and Caius College April 2005 This dissertation is submitted for the degree of Master of Philosophy Mating Six-spot Burnet moths (Zygaena filipendulae) on the Roughs Declaration This dissertation is the result of my own work and includes nothing which is the outcome of work done in collaboration except where specifically indicated in the text. This dissertation does not exceed the limit of 15000 words in the main text, excluding figures, tables, legends and appendices. i Acknowledgements This work was carried out on the land of Miriam Rothschild, who sadly passed away before its completion. I would like to thank her for allowing me to stay at Ashton Wold during my fieldwork and making me feel welcome there. I would also like to thank the Eranda Foundation for funding this work. Various people have helped with the identification of my material and I am very grateful to them for their time. Brian Eversham was of great help in identifying my carabids and also took time out from his busy schedule to assist me for a day during my time in the field. Ray Symonds dedicated a great deal of time to identifying all the spiders I caught, a feat which would have undoubtedly taken me many weeks! Richard Preece identified all my gastropods, and I am grateful both to him and his student George Speller for passing on the material to him. Roger Morris verified the identification of voucher specimens of all the syrphids I caught, and Oliver Prŷs-Jones did the same for my bumblebees. -
Eastern Persius Duskywing Erynnis Persius Persius
COSEWIC Assessment and Status Report on the Eastern Persius Duskywing Erynnis persius persius in Canada ENDANGERED 2006 COSEWIC COSEPAC COMMITTEE ON THE STATUS OF COMITÉ SUR LA SITUATION ENDANGERED WILDLIFE DES ESPÈCES EN PÉRIL IN CANADA AU CANADA COSEWIC status reports are working documents used in assigning the status of wildlife species suspected of being at risk. This report may be cited as follows: COSEWIC 2006. COSEWIC assessment and status report on the Eastern Persius Duskywing Erynnis persius persius in Canada. Committee on the Status of Endangered Wildlife in Canada. Ottawa. vi + 41 pp. (www.sararegistry.gc.ca/status/status_e.cfm). Production note: COSEWIC would like to acknowledge M.L. Holder for writing the status report on the Eastern Persius Duskywing Erynnis persius persius in Canada. COSEWIC also gratefully acknowledges the financial support of Environment Canada. The COSEWIC report review was overseen and edited by Theresa B. Fowler, Co-chair, COSEWIC Arthropods Species Specialist Subcommittee. For additional copies contact: COSEWIC Secretariat c/o Canadian Wildlife Service Environment Canada Ottawa, ON K1A 0H3 Tel.: (819) 997-4991 / (819) 953-3215 Fax: (819) 994-3684 E-mail: COSEWIC/[email protected] http://www.cosewic.gc.ca Également disponible en français sous le titre Évaluation et Rapport de situation du COSEPAC sur l’Hespérie Persius de l’Est (Erynnis persius persius) au Canada. Cover illustration: Eastern Persius Duskywing — Original drawing by Andrea Kingsley ©Her Majesty the Queen in Right of Canada 2006 Catalogue No. CW69-14/475-2006E-PDF ISBN 0-662-43258-4 Recycled paper COSEWIC Assessment Summary Assessment Summary – April 2006 Common name Eastern Persius Duskywing Scientific name Erynnis persius persius Status Endangered Reason for designation This lupine-feeding butterfly has been confirmed from only two sites in Canada. -
Identifying the Garden Butterflies of the Upper Thames Region Summer Very Top Tip
Identifying the Garden Butterflies of the Upper Thames Region Summer Very Top Tip Identifying butterflies very often depends on details at the wing edges (including the margins), but this is less helpful with some summer species. Note the basic ground colour to get a clue to the family and then look at wing edges and eye spots for further identification. The butterfly’s size and actions, plus location and time of year are all helpful . Butterflies that deliberately chase others (and large flies etc.) are usually male. Butterflies spending a long time flying around and investigating an individual plant are probably female. Peacock Red Admiral & Painted Lady The wing tips of PL separate it from Fritillaries Red Admiral & Painted Lady Comma Painted Lady Small Tortoiseshell Speckled Wood Chequered margins andSpeckled large blotches of yellowy creamWood on chocolate background. Often seen basking in patches of sun in areas of dappled sunlight. Rarely visits flowers. Comma Small Tort Wavy outline Blue-ish margin Peacock Speckled Wood Usually less contrasty than the rest with various eyespots and With a straighter border between Blue-ish margin prominent veins Brimstone and Clouded Yellow B has pointy wing tips CY has a white centre to brown spot on hind wing Black tips on upper visible in flight. Small White m and Large White f Very small amounts of dark scales on leading edge of forewing of Small W rarely turn around tip, but Large has large L of black scales that extend half way down the wing Female Brimstone and Green-veined White Brimstone fem looks white in flight but slightly greenish-yellow when settled. -
Campbell Et Al. 2020 Peerj.Pdf
A novel curation system to facilitate data integration across regional citizen science survey programs Dana L. Campbell1, Anne E. Thessen2,3 and Leslie Ries4 1 Division of Biological Sciences, School of STEM, University of Washington, Bothell, WA, USA 2 The Ronin Institute for Independent Scholarship, Montclair, NJ, USA 3 Center for Genome Research and Biocomputing, Oregon State University, Corvallis, OR, USA 4 Department of Biology, Georgetown University, Washington, DC, USA ABSTRACT Integrative modeling methods can now enable macrosystem-level understandings of biodiversity patterns, such as range changes resulting from shifts in climate or land use, by aggregating species-level data across multiple monitoring sources. This requires ensuring that taxon interpretations match up across different sources. While encouraging checklist standardization is certainly an option, coercing programs to change species lists they have used consistently for decades is rarely successful. Here we demonstrate a novel approach for tracking equivalent names and concepts, applied to a network of 10 regional programs that use the same protocols (so-called “Pollard walks”) to monitor butterflies across America north of Mexico. Our system involves, for each monitoring program, associating the taxonomic authority (in this case one of three North American butterfly fauna treatments: Pelham, 2014; North American Butterfly Association, Inc., 2016; Opler & Warren, 2003) that shares the most similar overall taxonomic interpretation to the program’s working species list. This allows us to define each term on each program’s list in the context of the appropriate authority’s species concept and curate the term alongside its authoritative concept. We then aligned the names representing equivalent taxonomic Submitted 30 July 2019 concepts among the three authorities. -
Urban Indicators for UK Butterflies
Ecological Indicators 76 (2017) 184–193 Contents lists available at ScienceDirect Ecological Indicators jo urnal homepage: www.elsevier.com/locate/ecolind Original Article Urban indicators for UK butterflies a,b,∗ a c b Emily B. Dennis , Byron J.T. Morgan , David B. Roy , Tom M. Brereton a School of Mathematics, Statistics and Actuarial Science, University of Kent, Canterbury, UK b Butterfly Conservation, Manor Yard, East Lulworth, Wareham, UK c Centre for Ecology & Hydrology, Benson Lane, Crowmarsh Gifford, Wallingford, UK a r t i c l e i n f o a b s t r a c t Article history: Most people live in urban environments and there is a need to produce abundance indices to assist Received 26 October 2016 policy and management of urban greenspaces and gardens. While regional indices are produced, with Received in revised form the exception of birds, studies of the differences between urban and rural areas are rare. We explore 19 December 2016 these differences for UK butterflies, with the intention to describe changes that are relevant to people Accepted 10 January 2017 living in urban areas, in order to better connect people with nature in support of conservation, provide a Available online 3 February 2017 measure relevant to human well-being, and assess the biodiversity status of the urban environment. Transects walked under the UK Butterfly Monitoring Scheme are classified as urban or rural, using Keywords: Abundance a classification for urban morphological zones. We use models from the Generalised Abundance Index Biodiversity family to produce urban and rural indices of relative abundance for UK butterfly species. -
Mardon Skipper Site Management Plans
Mardon Skipper (Polites mardon) Site Management Plans Gifford Pinchot National Forest Service Cowlitz Valley Ranger District Prepared by John Jakubowski North Zone Wildlife Biologist Reviewed by Rich Hatfield, The Xerces Society for Invertebrate Conservation October 2, 2015 Cowlitz Valley Ranger District Mardon Skipper Sites Group Meadow Longitude Latitude Area (acres) Elevation (ft.) Midway Midway 121 32.0 46 21.2 8 4,313 Midway PCT 121 31.1 46 21.1 2 4,530 Midway 115 Spur 121 30.9 46 21.0 3 4,494 Midway Grapefern 121 30.9 46 21.5 3 4,722 Midway 7A North 121 31.4 46 21.5 2 4,657 Midway 7A South 121 31.5 46 21.4 2 4,625 Midway 7A 121 31.1 46 21.4 7 4,676 Muddy Muddy 121 32.2 46.18.5 4 4,450 Muddy Lupine 121 31.8 46 18.7 3 4,398 Spring Cr Spring Cr. 121 33.5 46 20.4 unknown 3,900 Goal of the Management Plans Maintain and improve grassland/forb habitat at known occupancy meadow sites to ensure continued occupancy by mardon skipper butterfly as well as other important pollinator species such as western bumble bee. 1 Introduction On the Gifford Pinchot National Forest (GPNF), mardon skippers were first detected on the Mt. Adams Ranger District (MTA) in 2000 and on Cowlitz Valley Ranger District (CVRD) in 2002. Mardon skippers are known to inhabit ten, upland dry grassy meadows on the CVRD. Portions of the meadows are mesic and are unsuitable mardon skipper habitat. -
The Life History and Ecology of Hesperia Nabokovi in the Dominican Republic (Lepidoptera: Hesperiidae)
Vol. 1 No. 2 1990 Hesperia nabokovi: EMMEL and EMMEL 77 TROPICAL LEPIDOPTERA, 1(2): 77-82 THE LIFE HISTORY AND ECOLOGY OF HESPERIA NABOKOVI IN THE DOMINICAN REPUBLIC (LEPIDOPTERA: HESPERIIDAE) THOMAS C. EMMEL and JOHN F. EMMEL Department of Zoology, University of Florida, Gainesville, FL 32611, and 720 East Latham Avenue, Hemet, CA 92343, USA ABSTRACT.— The life history and ecology of Hesperia nabokovi (Bell & Comstock), the only species of this genus outside the Holarctic Region, is described; egg, larval and pupal characters confirm its generic assignment. The complete life cycle takes 94 days. KEY WORDS: Atalopedes, Calisto, Haiti, Gramineae, Hesperia comma, Hispaniola, immature stages, Nymphalidae, Oarisma stillmani, Satyrinae. to collect living females in the Dominican Republic, confine one of these and obtain an egg which was reared through by the second co-author (JFE) in California on a substitute grass foodplant. The purpose of this paper is to describe the elements of the life history and present some ecological notes on the biology of this fascinating Caribbean member of the genus Hesperia. DISTRIBUTION AND HABITAT The known distribution of Hesperia nabokovi is restricted to three general regions on the island of Hispaniola (Schwartz, 1989). The three areas are widely separated geographically and arc divided topographically from each other by high mountain ridges. These regions are; 1) the Cul de Sac-Valle de Neiba plain and the extreme western edge of the Llanos de Azua, from Fig. 1. Habitat for Hesperia nabokovi adults, 1km SE Monte Cristi (50ft elev.), Thomazeau and Fon Parisien in Haiti in the west to Canoa in the near the northern Dominican Republic coast, (photo by T. -
The C-SCOPE Marine Plan (Draft)
The C-SCOPE Marine Plan (Draft) C-SCOPE Marine Spatial Plan Page 1 Contents List of Figures & Tables 3 Chapter 5: The Draft C-SCOPE Marine Plan Acknowledgements 4 5.1 Vision 67 Foreword 5 5.2 Objectives 67 The Consultation Process 6 5.3 Policy framework 68 Chapter 1: Introduction 8 • Objective 1: Healthy Marine Environment (HME) 68 Chapter 2: The international and national context for • Objective 2: Thriving Coastal Communities marine planning (TCC) 81 2.1 What is marine planning? 9 • Objective 3: Successful and Sustainable 2.2 The international policy context 9 Marine Economy (SME) 86 2.3 The national policy context 9 • Objective 4: Responsible, Equitable and 2.4 Marine planning in England 10 Safe Access (REA) 107 • Objective 5: Coastal and Climate Change Chapter 3: Development of the C-SCOPE Marine Plan Adaptation and Mitigation (CAM) 121 3.1 Purpose and status of the Marine Plan 11 • Objective 6: Strategic Significance of the 3.2 Starting points for the C-SCOPE Marine Plan 11 Marine Environment (SS) 128 3.3 Process for producing the C-SCOPE • Objective 7: Valuing, Enjoying and Marine Plan 16 Understanding (VEU) 133 • Objective 8: Using Sound Science and Chapter 4: Overview of the C-SCOPE Marine Plan Area Data (SD) 144 4.1 Site description 23 4.2 Geology 25 Chapter 6: Indicators, monitoring 4.3 Oceanography 27 and review 147 4.4 Hydrology and drainage 30 4.5 Coastal and marine ecology 32 Glossary 148 4.6 Landscape and sea scape 35 List of Appendices 151 4.7 Cultural heritage 39 Abbreviations & Acronyms 152 4.8 Current activities 45 C-SCOPE -
Contribution to the Knowledge of the Fauna of Bombyces, Sphinges And
driemaandelijks tijdschrift van de VLAAMSE VERENIGING VOOR ENTOMOLOGIE Afgiftekantoor 2170 Merksem 1 ISSN 0771-5277 Periode: oktober – november – december 2002 Erkenningsnr. P209674 Redactie: Dr. J–P. Borie (Compiègne, France), Dr. L. De Bruyn (Antwerpen), T. C. Garrevoet (Antwerpen), B. Goater (Chandlers Ford, England), Dr. K. Maes (Gent), Dr. K. Martens (Brussel), H. van Oorschot (Amsterdam), D. van der Poorten (Antwerpen), W. O. De Prins (Antwerpen). Redactie-adres: W. O. De Prins, Nieuwe Donk 50, B-2100 Antwerpen (Belgium). e-mail: [email protected]. Jaargang 30, nummer 4 1 december 2002 Contribution to the knowledge of the fauna of Bombyces, Sphinges and Noctuidae of the Southern Ural Mountains, with description of a new Dichagyris (Lepidoptera: Lasiocampidae, Endromidae, Saturniidae, Sphingidae, Notodontidae, Noctuidae, Pantheidae, Lymantriidae, Nolidae, Arctiidae) Kari Nupponen & Michael Fibiger [In co-operation with Vladimir Olschwang, Timo Nupponen, Jari Junnilainen, Matti Ahola and Jari- Pekka Kaitila] Abstract. The list, comprising 624 species in the families Lasiocampidae, Endromidae, Saturniidae, Sphingidae, Notodontidae, Noctuidae, Pantheidae, Lymantriidae, Nolidae and Arctiidae from the Southern Ural Mountains is presented. The material was collected during 1996–2001 in 10 different expeditions. Dichagyris lux Fibiger & K. Nupponen sp. n. is described. 17 species are reported for the first time from Europe: Clostera albosigma (Fitch, 1855), Xylomoia retinax Mikkola, 1998, Ecbolemia misella (Püngeler, 1907), Pseudohadena stenoptera Boursin, 1970, Hadula nupponenorum Hacker & Fibiger, 2002, Saragossa uralica Hacker & Fibiger, 2002, Conisania arida (Lederer, 1855), Polia malchani (Draudt, 1934), Polia vespertilio (Draudt, 1934), Polia altaica (Lederer, 1853), Mythimna opaca (Staudinger, 1899), Chersotis stridula (Hampson, 1903), Xestia wockei (Möschler, 1862), Euxoa dsheiron Brandt, 1938, Agrotis murinoides Poole, 1989, Agrotis sp. -
The Butterfly Drawings by John Abbot in the Hargrett Rare Book and Manuscript Library, University of Georgia
VOLUME 61, NUMBER 3 125 Journal of the Lepidopterists’ Society 61(3), 2007, 125–137 THE BUTTERFLY DRAWINGS BY JOHN ABBOT IN THE HARGRETT RARE BOOK AND MANUSCRIPT LIBRARY, UNIVERSITY OF GEORGIA. JOHN V. C ALHOUN1 977 Wicks Dr., Palm Harbor, FL 34684 ABSTRACT. Artist-naturalist John Abbot completed 105 drawings of insects that are now deposited in the Hargrett Rare Book and Manu- script Library, University of Georgia. The provenance of these drawings is unknown, but available evidence dates them to ca. 1820–1825. The adults in the 32 butterfly drawings are identified and the figures of larvae and pupae are assessed for accuracy. The illustrated plants are also identified and their status as valid hosts is examined. Abbot’s accompanying notes are transcribed and analyzed. Erroneous figures of larvae, pupae, and hostplants are discussed using examples from the Hargrett Library. At least four of the butterfly species portrayed in the drawings were probably more widespread in eastern Georgia during Abbot’s lifetime. Additional key words: Larva, Lepidoptera, pupa, watercolors In 1776, the English artist-naturalist John Abbot METHODS (1751–ca.1840) arrived in Georgia, where he I visited the Hargrett Rare Book and Manuscript documented species of animals and plants for the next Library (University of Georgia) in April, 2005. Digital six decades. Living in Burke, Bullock, Chatham, and photographs were taken of John Abbot’s butterfly Screven Counties of eastern Georgia, he explored a drawings and their accompanying notes. The adult region roughly bound by the cities of Augusta and butterflies were identified and the figures compared Savannah, between the Oconee, Altamaha, and with those in other sets of Abbot’s drawings that are Savannah Rivers.