An Inventory of Invertebrates recorded from

Three Hagges Wood-meadow

During 2018

Cercopis vulnerata Black-and-red Froghopper [a bug] adult Lithophane ornitopus Grey-shoulder Knot [a moth] adult

Ochlodes sylvanus Large Skipper [a butterfly] male Leucozona lucorum [a hoverfly] female

A Report to Hagge Woods Trust by Andrew Grayson

The Author is a lifelong amateur naturalist who has been a professional invertebrate specialist since 2009. He has been involved with invertebrate studies at Three Hagges Wood-meadow since 2014, when the site was then known as Three Hagges Jubilee Wood, and has been very pleased to be associated with The Wood Meadow Project and the wonderful group of people involved in management of the Three Hagges Wood-meadow site. This association was responsible for his nomination for the Gilbert White Adult Award at the 2018 UK Awards for Biological Recording and Information Sharing held in Nottingham, where he was honoured to make the final shortlist, and for his appearance in the BBC Wildlife Magazine. These were both unexpected achievements. Andrew very much looks forward to being involved with studies at Three Hagges Wood-meadow during 2019. These will not be on the scale of the studies of the past five years, but hopefully will continue to provide useful invertebrate records, including some interesting and unexpected species, as has been the case since 2014. Andrew Grayson at Three Hagges Wood-meadow

Invertebrates on the Front Cover

Cercopis vulnerata Black-and-red Froghopper [a bug] Single examples of this small but conspicuous and distinctive bug have been seen roughly half-way along the A19 Strip (Area 12) every year since 2014. Elsewhere at Three Hagges Wood-meadow; it was photographed in the Coups 11, 12 & 12a compartment (Area 7) by Lin Hawthorne in 2016, and in an unspecified compartment by Pat & Jim Bone in 2018. The individual on the front cover of this report was photographed in the A19 Strip (Area 12) on 3rd June 2018.

Lithophane ornitopus Grey-shoulder Knot [a moth] This moth flies during the autumn and spring: more specifically, it emerges as an adult during September and October, overwinters, and then flies again during the February to April period. The UK Moths website www.ukmoths.org.uk considers it to be ‘a woodland species which is found uncommonly in the southern part of England, Wales and Ireland’. This individual was encountered on 20th February 2018, whilst it was well-camouflaged at rest during the daytime on the bark of one of the tree-trunk pillars which support the roof of the Bodger’s Den (Area 8a).

Ochlodes sylvanus Large Skipper [a butterfly] This easily-identifiable skipper is very infrequently seen at Three Hagges Wood-meadow, but its numbers onsite are very probably increasing. Eleven individuals were recorded during 2018, compared with two in 2014, 2016 and 2017, and none during 2015. This pristine male was photographed in Nick’s Field (Area 16) on 3rd June 2018.

Leucozona lucorum [a hoverfly] This attractive and distinctive hoverfly was only seen once in each of the first three years of recording at Three Hagges Wood-meadow. It then became comparatively numerous during 2017 when twelve were counted. Only two were seen during 2018; however, the 2018 survey was carried out over far fewer days than previous surveys, so direct comparisons cannot be made. This female was photographed in the Coups 6, 8 & 10 survey compartment (Area 1) on 6th May 2018.

In memory of Jim Bone (1937-2019)

An Inventory of Invertebrates recorded from

Three Hagges Wood-meadow

During 2018

A REPORT TO

Hagge Woods Trust Little Annexe, Estate Office, Escrick, York, YO19 6EA www.haggewoodstrust.org.uk www.threehaggesjubileewood.org.uk Hagge Woods Trust is Registered Charity No. 1151831

BY

Andrew Grayson Invertebrate Consultant, ‘Scardale’, High Lane, Beadlam, Nawton, York, YO62 7SX Email: [email protected]

REPORT SUBMITTED ON

Thursday 24th January 2019

All PHOTOGRAPHS IN THIS REPORT WERE TAKEN BY ANDREW GRAYSON DURING 2018 UNLESS STATED OTHERWISE CONTENTS

INTRODUCTION ...... 1 SUMMARY ...... 1 THREE HAGGES WOOD-MEADOW ...... 1 INVERTEBRATE RECORDING ...... 1 SURVEY METHODOLOGY ...... 2 SURVEY METHODOLOGY FOR 2018 ...... 2 SURVEY DATES, WEATHER CONDITIONS AND NOTES ...... 3 SUPPLEMENTARY RECORDING OF INVERTEBRATES ...... 4 PRESERVATION OF SPECIMENS ...... 4 INCIDENTAL RECORDING OF PLANTS ...... 4 RESULTS FOR 2018 ...... 5 INTRODUCTION ...... 5 SPECIES OF NATIONAL CONSERVATION CONCERN ...... 5 SPIE [FORMERLY UK BAP PRIORITY SPECIES] ...... 6 JNCC-LISTED SPECIES ...... 6 SPECIES NEW TO VICE-COUNTY 61 ...... 7 DIPTERA NEW TO VICE-COUNTY 61 ...... 7 AN ICHNEUMON [HYMENOPTERA] NEW TO VICE-COUNTY 61 ...... 7 SUMMARY TABLES OF SURVEY DATA FOR 2018 ...... 8 ORDERS AND NUMBER OF SPECIES FOUND AT THREE HAGGES WOOD-MEADOW DURING 2018 ...... 8 FAMILIES AND NUMBER OF SPECIES FOUND AT THREE HAGGES WOOD-MEADOW DURING 2018 ...... 8 ACKNOWLEDGEMENTS ...... 13 REFERENCES ...... 13

APPENDICES

APPENDIX 1: PLANS ...... 14 THREE HAGGES WOOD-MEADOW AND ITS ENVIRONS ...... 14 EXTENTS OF SURVEY COMPARTMENTS AND LOCATIONS OF FEATURES ...... 15 APPENDIX 2: TABLES ...... 16 FULL INVERTEBRATE LIST (2014 TO 2018) ...... 16 INVERTEBRATE LIST FOR 2018 ...... 42 DISSECTION OF INVERTEBRATE DATA FOR 2018 ...... 53 PLANT LIST FOR 2018 ...... 62 APPENDIX 3: PHOTOGRAPHS ...... 63 SURVEY COMPARTMENT DETAILS ...... 63 SURVEY COMPARTMENT MONTHLY CHANGES ...... 79 PHOTOGRAPHS OF INVERTEBRATES ...... 175 PHOTOGRAPHS OF EVENTS AND FEATURES ...... 216 APPENDIX 4: SPREADSHEET ...... 223 INVERTEBRATE DATA FROM THREE HAGGES WOOD-MEADOW 2018 ...... 223

The spreadsheet is not paginated (pages 223 to 252)

1 INTRODUCTION

SUMMARY

Five years of continuous invertebrate recording have now been completed at Three Hagges Wood-meadow. During that time, nearly 1,000 invertebrates have been recorded, including some which are of national conservation interest. It seems appropriate that a special report should be produced to cover the first five years of invertebrate recording at Three Hagges Wood-meadow. Consequently, there will be two invertebrate reports produced during 2019, the first of which is the current report.

The current annual report divulges all 2018 invertebrate survey data, but does not give all the usual additional notes which were included in the four previous annual reports, which were: Grayson (2014) [the report for 2014], Grayson (2016) [the report for 2015], Grayson (2017) [the report for 2016], and Grayson (2018) [the report for 2017]. The usual additional notes are omitted from the current report, as they will form part of the special five-year report. The current annual report is therefore packed with tables and photographs from 2018, but is relatively short on text.

THREE HAGGES WOOD-MEADOW

Three Hagges Wood-meadow, formerly known as Three Hagges Jubilee Wood, is situated on the eastern side of the A19 road between the villages of Escrick and Riccall in the county of North Yorkshire. It is entirely within the National Grid 1km square SE6239, and its approximate central 1m square is SE6267039610. Given alternatively, its approximate central latitudinal and longitudinal geographic co-ordinates are 53°50’55.96”N 1°02’56.00”W.

For invertebrate recording purposes, the boundaries of the Three Hagges Wood-meadow site are as shown on the plan on page 15. This plan also shows the boundaries of the sixteen survey compartments, and the locations of the following structures: the Bee Hotel (Area 1a), Bodger’s Den (Area 8a), and Crombie Roundhouse (Area 9a).

The sixteen survey compartments basically comprise: a main meadow/plantation area (areas 1 to 9), which is enclosed within a high fence, and which was an arable field until as recently as summer 2012; a pond (Area 10), which was a feature created as part of The Wood Meadow Project; About Thyme Wood (Area 11), which is a small, well-established, managed woodland; strips of land, some with plantations (areas 12 to 15), which surround the main meadow/plantation area (areas 1 to 9); and part of a field known as Nick’s Field (Area 16), which was used as a ‘control’ area to enable comparisons to be made with the main part of the Three Hagges Wood-meadow site.

INVERTEBRATE RECORDING

Invertebrate studies have been carried out at Three Hagges Wood-meadow since 2014, leading to an overall site list of 990 species. This full invertebrate list is enumerated on pages 16 to 41.

Most invertebrate surveying has been carried out by Andrew Grayson, principally by sweep-sampling, ground-searching, and the recording of conspicuous and easily-identifiable species in the field. This surveying was carried out between April and September every year from 2014 to 2017; and, as a one-off exercise, was extended in 2018 to include all months of the year.

Other recording work at Three Hagges Wood-meadow has included: official visits by members of the Yorkshire Naturalists’ Union; light-trapping, principally carried out by Dave Chesmore and Lin Hawthorne; pond-dipping, principally carried out by a team led by Martin Hammond; and photography, principally by Pat and Jim Bone, Dave Tate and Mike Cowling.

The level of invertebrate studies and achievements to date at Three Hagges Wood-meadow would not have possible without funding support kindly provided by the Esmée Fairbairn Foundation. This funding support is a direct consequence of the considerable work and enthusiasm for the Three Hagges Wood-meadow project shown by Rosalind Forbes Adam [chair of the Hagge Woods Trust] and also Lin Hawthorne, who was heavily-involved with the project until early 2017. 2 SURVEY METHODOLOGY

SURVEY METHODOLOGY FOR 2018

Andrew Grayson has been responsible for continuous systematic invertebrate surveying at Three Hagges Wood-meadow since 2014. As the years have advanced, and the site has developed to provide habitats which support ever-increasing numbers of invertebrates, it was inevitable that the tasks involved in carrying out surveying and ultimately producing annual reports would become progressively more onerous. This resulted in the surveyor paying progressively more visits to Three Hagges Wood-meadow during the summer months each year, which created progressively more indoor work, and considerably delayed the production of reports, as happened with the reports for 2016 and 2017.

Hagge Woods Trust were keen that Andrew Grayson should amend the original principal survey requirement methodology in order to reduce the volume of his tasks, and allow the 2018 annual report to be produced in good time. This has obviously meant that fewer invertebrates were collected during 2018, which gives the false impression that fewer invertebrates were onsite, yet the opposite was true, particularly with arboreal species, which occurred in abundance.

Andrew Grayson’s basic amended survey requirements for 2018 were therefore as follows:

1. Each of the sixteen compartments, plus the features known as the Bee Hotel (Area 1a), Bodger’s Den (Area 8a) and Crombie Roundhouse (Area 9a), were to be individually investigated once each month during periods when weather conditions were favourable for recording invertebrates that are active during daytime. 2. Invertebrate samples required for critical examination via microscope were to be collected by a combination of spot-sampling, sweep-sampling, ground-searching and beating. 3. All conspicuous and easily-identifiable invertebrates were to be recorded in the field, and where practicable, a hand-held lens was to be used to identify some smaller invertebrates, which could then be released. 4. Particular attention was to be focussed on invertebrate assemblages which are important pollinators, e.g. solitary bees and hoverflies; therefore, these assemblages would always be collected or recorded in the field if possible. 5. Photographs were to be taken of each compartment during every survey visit, in order to provide good photographic coverage throughout 2018. 6. Supplementary records were to be obtained by liaising with specialists, and others, who visited Three Hagges Wood-meadow during 2018 for the purposes of invertebrate recording or photography etc.

SURVEY DATES, WEATHER CONDITIONS AND NOTES

JANUARY

The hard winter of 2017/2018 much reduced the number of days which were ideally suitable for invertebrate survey work, and particularly so in January, when the survey had to be postponed until nearly the end of the month, with the knock-on effect that surveys during the following months were also carried out during their latter parts, in order that the survey dates were not too close together.

The January survey was carried out on 28th January, which was an unseasonably mild day of 13°C. The survey period was afflicted by misty drizzle and moderate westerly breezes, with roughly equal periods of hazy sunshine and cloud. There were no flying insects present, but a reasonable quantity of subterranean and other invertebrates were to be found under debris, including a couple of additions to the site list. Furthermore, all Andrew Grayson’s monthly survey visits to Three Hagges Wood-meadow during 2018 provided additions to the site list, which was remarkable.

FEBRUARY

The February survey was carried out on 20th February, which was a reasonably pleasant day for the time of year. There had been heavy rainfall during the previous day and overnight, but 20th February began with sunshine and some light cloud. The temperature was a mild 10°C at the commencement of surveying, and 13°C at its cessation a few hours later. Cloud gradually increased, accompanied by cool, moderate, north-westerly wind. Some Diptera were out of hibernation. 3 MARCH

Surveying was carried out on 21st March, which was the second of two fine sunny days, following a prolonged period of snowy weather conditions. The temperature was initially a mild 9°C, rising to 13°C by the cessation of the survey. Breezes were cool, light-moderate in strength, and westerly in direction.

APRIL

Weather conditions on 22nd April were generally conducive to invertebrate activity, and many spring species were in evidence, including an abundance of water beetles in the Pond [Area 10]; albeit, these were not sampled, as per management protocol of general non-disturbance of the site’s aquatic fauna.

The survey commenced in sunny conditions, but these were soon replaced by high overcast, and there were a few spots of rain before the survey period ended. The temperature remained approximately 19°C throughout the day, despite changes from sunshine to high overcast, and from initial warm moderate southerly breezes, to cooler moderate westerly breezes.

MAY

From May onwards, invertebrate surveying was carried out near the start of each month, beginning with 6th May, which was a hot and sunny day under a mainly clear blue sky. Most invertebrates were to be found in the more-sheltered peripheral areas of Three Hagges Wood-meadow on 6th May, rather than the more-exposed central meadow/plantation area (areas 1 to 9). This general pattern of invertebrate distribution was repeated on similar days throughout the year.

The temperature during the survey period on 6th May rose from 18°C to 24°C. Winds were initially still or light south easterly, but later became a more regular light southerly breeze.

JUNE

The survey day of 3rd June coincided with a Big Nature Lunch event held at Three Hagges Wood-meadow. This was overall a very warm and humid day; initially being mainly cloudy with a few sunny breaks, then becoming mainly hot and sunny during mid-afternoon. There was quite a degree of moisture onsite initially, as it had rained the day before; however, this soon evaporated. Breezes were mainly light and westerly. The temperature initially rose slowly from 21°C at the start of the survey period, but then more rapidly to a high of 29°C during the mid-afternoon.

JULY

The July surveying was carried out on 1st July, which was a hot and sunny day with little or no wind. There were very few invertebrates to be found in the main meadow/plantation area (areas 1 to 9), as it was simply too hot and dry for most of them to survive out in the open. The average temperature was circa 27°C, but the maximum reached 30°C. Nick’s Field (Area 16) had been very-recently mown, and was devoid of invertebrates.

AUGUST

The August surveying was carried out on 4th August; which was mainly a hot and sunny day with half blue sky and half wispy clouds. There was either no breeze, or a light southerly one. The temperature when surveying commenced was 24°C; this rose to 30°C by late afternoon, but soon fell to 27°C.

All areas of open grassland had been recently-mown; and, following a prolonged period of drought, the level of water in the Pond (Area 10) was very low, and all seasonally-wet areas had dried out. Most of the seasonally-wet areas at Three Hagges Wood-meadow are on the MG4 Meadow (Area 8) and in About Thyme Wood (Area 11).

With the exception of grasshoppers, invertebrates were very scarce on the main meadow/plantation area (areas 1 to 9) on 4th August. Social wasp workers were noticeably abundant, but only a few were captured for identification purposes.

4 SEPTEMBER

The September surveying was carried out on 2nd September. This was a day of alternating periods of sunshine and light cloud, with winds generally easterly and light, but becoming mainly southerly and moderate towards the end of the survey period. The temperature rose from 19°C at the start of surveying at 09.23, to 25°C by the cessation in mid-afternoon.

OCTOBER

The October surveying took place on 1st October, which was a pleasant, if somewhat slightly cool, day. Most of the survey period was blessed by full sunshine, particularly during the morning; but this was replaced by gloomy cloudy conditions towards the end of the survey period in late afternoon. The temperature remained unexpectedly low, ranging only between 12°C and 15°C. Winds were variable: initially being mainly very light, cool, and westerly; but later turning to moderate, cool, and northerly; and then to cool, light moderate, and easterly. There were relatively-few flowers onsite on 1st October; particularly on areas of grassland that had been recently-mown, and the bulk of the MG5 Meadow (Area 9), which was being grazed by sheep. These areas were almost devoid of invertebrates.

NOVEMBER

The November survey was carried out on 1st November, which was a mild and pleasant autumnal day. There had been rainfall overnight, and during the early morning, but the survey period occurred after the rain had abated. Initial cloud slowly cleared to be replaced by full sunshine during the afternoon. Winds were either still, or rather consistently light, cool, and westerly. As might be expected for the time of year, the temperature was suppressed between 11°C and 14°C. Late-season Diptera were quite abundant, especially Muscidae and Calliphoridae.

DECEMBER

It is usually difficult to choose a day in December which is suitable for invertebrate recording, but 4th December was as pleasant as could be wished for. Inevitably, most of the few recordable invertebrates were to be found under debris. Upon commencement of surveying in the morning, the temperature was only 2°C, and the conditions were frosty. Although the temperature during the survey period never rose above 3°C, the actual conditions onsite felt much milder, as 4th December was a consistently sunny day, albeit the sunshine often became hazy. There was no wind throughout the survey period; at most an occasional slight movement of grasses was observable.

SUPPLEMENTARY RECORDING OF INVERTEBRATES

In addition to his systematic monthly surveys; Andrew Grayson also recorded some invertebrates on a fairly casual basis, during organised social events held at Three Hagges Wood-meadow on 8th July and 11th September 2018.

Richard Baker ran two light-traps on the MG4 Meadow area (Area 8) on 10th August 2018, in order to record moths.

Geoff Oxford recorded a list of spiders during a casual visit to Three Hagges Wood-meadow on 7th October 2018.

Pat and Jim Bone have made frequent summer visits to Three Hagges Wood-meadow since 2015 in order to photograph wildlife, count butterfly numbers and attend social events. 2018 produced their now customary butterfly lists, and albums of excellent photographs.

PRESERVATION OF SPECIMENS

As with previous years, good quantities of voucher specimens have been retained by Andrew Grayson as individually- labelled dry-pinned specimens. Ultimately, these will be deposited in an appropriate museum, or museums.

INCIDENTAL RECORDING OF PLANTS

Many invertebrates obtain nectar and pollen when visiting flowers. Pages 63 to 78 cite a few conspicuous plants which were in flower during the 2018 survey visits. Page 62 contains a summary table of these plant records from 2018. 5 RESULTS FOR 2018

INTRODUCTION

Parts of this section are reproduced verbatim from previous annual invertebrate reports, as it is not really possible to improve on the wording; hence, any alterations would be merely ‘window-dressing’.

It is customary for the results section of any invertebrate report to mention species which are of national conservation importance according to Government and similar organisations; therefore, such species are tabulated over the following pages. Far more details will be given in the forthcoming five-year summary report.

SPECIES OF NATIONAL CONSERVATION CONCERN

Species which are currently statutorily protected are of primary importance. Other species which may be considered are those currently listed in the UK BAP [United Kingdom Biodiversity Action Plan]; and, more particularly, those which are currently attributed a national conservation status by JNCC [Joint Nature Conservation Committee], or an international conservation status by IUCN [International Union for Conservation of Nature].

Statutorily protected species were listed in Schedule 5 of the Wildlife and Countryside Act 1981. The list is updated in quinquennial reviews, and is available via www.legislation.gov.uk .

UK BAP resulted from the UK Government signing the CBD [Convention on Biological Diversity] at the Rio Earth Summit in 1992. The current list of 1,150 UK BAP Priority Species is readily available via the UK BAP website www.ukbap.org.uk . The most recent UK BAP review was in August 2007. The UK BAP lists of species and habitats remain valid; however, there have been changes in terminology etc. since 2012, principally due to devolution. English UK BAP Priority Species are currently termed SPIE [Species of Principal Importance in England].

JNCC has allocated national conservation statuses to many British invertebrates, such statuses being based upon the collective expert opinion of many specialists. The publication of these national statuses began when invertebrates considered to be nationally rare and threatened were listed in the Red Data Books (Shirt, 1987; Bratton, 1991). These works were quickly superseded by national reviews such as Falk (1991), which used ‘Red Data Book’ categorization for nationally rare species, and ‘Notable’ categorization for those nationally scarce species which did not qualify for a ‘Red Data Book’ status. More recent JNCC reviews of species status have adopted IUCN categorization and terminology, and some recent JNCC reviews, such as Alexander (2014), have attributed national statuses to all species within their remit, not just those considered to be nationally scarce or rare. Much invertebrate conservation evaluation is based around species which have been categorized in JNCC publications. Explanations of JNCC conservation statuses, and listings of the species to which they are currently allocated, are readily available via the JNCC website www.jncc.gov.uk.

Internationally rare and threatened invertebrates are listed in The IUCN Red List of Threatened Species, which is readily accessible via www.iucnredlist.org .

The results of the 2018 invertebrate studies at Three Hagges Wood-meadow produced five SPIE invertebrates, and eight JNCC-listed invertebrates. This represents a very good result, given that the level of recording was reduced from that of previous years.

6 SPIE [FORMERLY UK BAP PRIORITY SPECIES]

Five SPIE were recorded during 2018, including Erynnis tages [Dingy Skipper], which had not been previously seen at Three Hagges Wood-meadow. The combined results from the 2014 to 2018 surveys have now produced fifteen SPIE, two being butterflies, and the remainder being moths.

Whilst it is of interest to learn that these insects are designated as SPIE, it is important to understand that, despite declines in some regions, these all remain widespread species that are either common or local in Britain, and do not have any overall tangible current conservation value.

The thirteen SPIE moths found at Three Hagges Wood-meadow between 2014 and 2018 were chosen by JNCC, along with many other common and widespread UK moths, to form a grouped action plan to investigate the causes of decline of widespread moth species. This research aims to investigate how moths are affected by changes in land use policy and management within both rural and urban environments.

TABLE OF SPIE FOUND AT THREE HAGGES WOOD-MEADOW DURING 2018

SPECIES FAMILY ORDER VERNACULAR NAME Scotopteryx chenopodiata (Linnaeus) Geometridae Lepidoptera Shaded Broad-bar [a moth] Timandra comae Schmidt Geometridae Lepidoptera Blood-vein [a moth] Tyria jacobaeae (Linnaeus ) Erebidae Lepidoptera The Cinnabar [a moth] Hydraecia micacea (Esper) Noctuidae Lepidoptera Rosy Rustic [a moth] Erynnis tages (Linnaeus) Hesperiidae Lepidoptera Dingy Skipper [a butterfly]

JNCC-LISTED SPECIES

Eight JNCC-listed invertebrates were found at Three Hagges Wood-meadow during 2018, including the flies Lonchaea corusca, Pherbellia griseola and Conisternum decipiens; all which were new to the site list. This represents quite a good result, given that the level of surveying was reduced from that of previous years.

The combined results of the 2014 to 2018 surveys have produced eighteen JNCC-listed invertebrates; i.e. species which currently hold JNCC ‘nationally rare’ or ‘nationally scarce’ conservation statuses [or equivalent JNCC statuses which use different terminology].

It is fair to point out that many of the JNCC-listed species found at Three Hagges Wood-meadow since 2014 have become more frequent or better known in Britain since their British status was designated; therefore, based on current knowledge, any future revision would undoubtedly downgrade the national status of such species. Nevertheless; a few genuinely scarce, or otherwise interesting, invertebrates have been recorded from the site during the last five years.

TABLE OF JNCC-LISTED SPECIES FOUND AT THREE HAGGES WOOD-MEADOW DURING 2018

SPECIES FAMILY ORDER VERNACULAR NAME OR GROUP Agelastica alni (Linnaeus) Chrysomelidae Coleoptera Alder Leaf Beetle Polydrusus formosus (Mayer) Curculionidae Coleoptera a weevil Dolichovespula saxonica (Fabricius) Vespidae Hymenoptera Saxon Wasp [a social wasp] Bombus rupestris (Fabricius) Apidae Hymenoptera Hill Cuckoo-bee [a bumblebee] Lonchaea corusca Czerny Lonchaeidae Diptera a lance fly Pherbellia griseola (Fallén) Sciomyzidae Diptera a marsh fly Conisternum decipiens (Haliday) Scathophagidae Diptera a dung fly Zophomyia temula (Scopoli) Tachinidae Diptera a parasitic fly

7 SPECIES NEW TO VICE-COUNTY 61

DIPTERA NEW TO VICE-COUNTY 61

For the purposes of facilitating continuity in biological recording, the historic county of Yorkshire is conceptually sub- divided into five ‘Watsonian vice-counties’. The vive-county system was devised by Hewett Cottrell Watson (1804-1881) for botanical recording, but was quickly adopted by other naturalists, and soon became the standard method for all British and later Irish, biological recording. A species is said to be ‘new’ to a vice-county if it has not previously been recorded there.

Under the Watsonian system, Three Hagges Wood-meadow is situated in the far western part of vice-county 61 [VC61]. VC61 is named as South East Yorkshire under the Watsonian system.

The 2018 survey of Three Hagges Wood-meadow produced seven additions to the VC61 list, which is a very good result. The total number of additions to the VC61 list produced by the combined 2014 to 2018 surveys is a rather impressive thirty-three species. These comprise a mixture of widespread and common flies which have previously been overlooked, and some flies which are genuinely regionally scarce.

TABLE OF DIPTERA NEW TO VICE-COUNTY 61 FROM THREE HAGGES WOOD-MEADOW DURING 2018

SPECIES FAMILY VERNACULAR NAME OR GROUP Borophaga incrassata (Meigen) Phoridae a scuttle fly Lonchaea corusca Czerny Lonchaeidae a lance fly Scaptomyza flava (Fallén) Drosophilidae Turnip Leafminer [a vinegar fly] Fannia pallitibia (Rondani) Fanniidae a fanniid fly Coenosia perpusilla Meigen Muscidae a muscid fly Thricops simplex (Wiedemann) Muscidae a muscid fly Protophormia terraenovae (Robineau-Desvoidy) Calliphoridae Blackbottle [a bluebottle fly]

AN ICHNEUMON [HYMENOPTERA] NEW TO VICE-COUNTY 61

The ichneumon wasp Ichneumon stramentor Rasnitsyn is new to VC61 (Bill Ely, pers. comm.) based upon specimens taken at Three Hagges Wood-meadow. Two Ichneumon stramentor females were found together under a stone near the garage in About Thyme Wood (Area 11) on 28th January 2018, which was a mild day for the time of year.

8 SUMMARY TABLES OF SURVEY DATA FOR 2018

ORDERS AND NUMBER OF SPECIES FOUND AT THREE HAGGES WOOD-MEADOW DURING 2018

‘SPIE’ means Species of Principal Importance in England [formerly known as UK BAP Priority Species] ‘JNCC Species’ means a national categorization of Red Data Book or Notable status [or their equivalent statuses based on IUCN] The Bee Hotel (Area 1a), Bodger’s Den (Area 8a) and Crombie Roundhouse (Area 9a) are included in Wood-meadow (areas 1 to 9)

Order or Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Class* Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Gastropoda* 5 1 2 4 2 Araneae 8 8 1 1 Glomerida 1 1 1 Polydesmida 1 1 Lithobiomorpha 1 1 Julida 1 1 1 Isopoda 4 4 4 2 Odonata 10 6 10 1 5 Orthoptera 3 3 3 1 2 2 Dermaptera 1 1 1 Hemiptera 18 5 15 Neuroptera 2 2 1 Mecoptera 1 1 Lepidoptera 52 45 10 12 19 3 5 Coleoptera 39 30 3 6 20 4 2 Hymenoptera 50 28 4 17 32 8 2 Diptera 158 75 13 52 102 15 4 Species Total 355 209 45 103 203 32 13

FAMILIES AND NUMBER OF SPECIES FOUND AT THREE HAGGES WOOD-MEADOW DURING 2018

GASTROPODA – slugs and snails

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Arionidae 1 1 1 1 1 Helicidae 3 3 1 Lymnaeidae 1 1 Species Total 5 1 2 4 2 0 0

ARANEAE – spiders

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Araneidae 4 4 1 1 Philodromidae 1 1 Pisauridae 1 1 Tetragnathidae 1 1 Theridiidae 1 1 Species Total 8 8 0 1 1 0 0

9 GLOMERIDA – pill millipedes

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Glomeridae 1 1 1 Species Total 1 1 0 1 0 0 0

POLYDESMIDA – flat-backed millipedes etc

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Polydesmidae 1 1 Species Total 1 0 0 1 0 0 0

LITHOBIOMORPHA – stone centipedes

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Lithobiidae 1 1 Species Total 1 1 0 0 0 0 0

JULIDA – millipedes

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Julidae 1 1 1 Species Total 1 1 0 0 1 0 0

ISOPODA – woodlice and water-lice etc

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Oniscidae 2 2 2 Porcellionidae 1 1 1 1 Armadillidiidae 1 1 1 1 Species Total 4 4 0 4 2 0 0

ODONATA – damselflies and dragonflies

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Coenagriidae 3 1 3 1 Aeshnidae 3 3 3 2 Libellulidae 4 2 4 1 2 Species Total 10 6 10 1 5 0 0

10 ORTHOPTERA – grasshoppers, groundhoppers and crickets etc

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Tetrigidae 1 1 1 Acrididae 2 2 2 1 2 2 Species Total 3 3 3 1 2 2 0

DERMAPTERA – earwigs

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Forficulidae 1 1 1 Species Total 1 0 0 1 1 0 0

HEMIPTERA – bugs

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Acanthosomatidae 3 1 3 Pentatomidae 3 1 3 Lygaeidae 1 1 Miridae 5 1 4 Nabidae 2 1 1 Cercopidae 1 1 1 Aphrophoridae 1 1 Cicadellidae 2 1 1 Species Total 18 5 0 0 15 0 0

NEUROPTERA – lacewings and alder-flies etc

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Chrysopidae 2 2 1 Species Total 2 0 0 2 1 0 0

MECOPTERA – scorpion-flies

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Panorpidae 1 1 Species Total 1 1 0 0 0 0 0

11 LEPIDOPTERA – butterflies and moths

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Eriocraniidae 1 1 Yponomeutidae 1 1 Zygaenidae 1 1 1 Drepanidae 1 1 Geometridae 6 4 1 1 2 Notodontidae 1 1 Erebidae 5 3 1 1 2 1 Noctuidae 14 14 1 1 1 Hesperiidae 3 3 1 2 1 1 Pieridae 5 5 2 3 2 Nymphalidae 9 8 3 5 7 2 Lycaenidae 5 4 2 1 3 Species Total 52 45 10 12 19 3 5

COLEOPTERA – beetles

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Carabidae 12 9 2 1 2 Silphidae 2 2 Scarabaeidae 1 1 1 Elateridae 4 1 1 2 1 Cantharidae 4 2 1 3 2 Malachiidae 1 1 1 1 Nitidulidae 1 1 1 Coccinellidae 5 5 1 1 3 Oedemeridae 1 1 1 Pyrochroidae 1 2 Cerambycidae 2 2 2 Chrysomelidae 2 1 1 1 1 Curculionidae 3 3 3 1 Species Total 39 30 3 6 20 4 2

HYMENOPTERA – sawflies, ants, bees, wasps and ichneumons etc

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Cephidae 1 1 Tenthredinidae 7 2 1 5 Formicidae 1 1 1 1 Vespidae 4 3 3 2 3 1 1 Crabronidae 3 1 1 1 Andrenidae 8 4 1 6 Halictidae 4 2 1 1 Megachilidae 1 1 1 Anthophoridae 2 2 Apidae 10 10 1 6 8 6 1 Cynipidae 1 1 1 1 Ichneumonidae 8 3 3 3 Species Total 50 28 4 17 32 8 2 12 DIPTERA – flies

Family Species Wood- Pond & its About Thyme Dyke Strips Nick’s Field SPIE or Total meadow margins Wood etc. JNCC (1 to 9) (10) (11) (12 to 15) (16) Species Tipulidae 7 6 1 3 2 2 Pediciidae 1 1 Limoniidae 3 1 2 Bibionidae 5 3 2 3 1 Trichoceridae 3 3 Anisopodidae 1 1 Culicidae 3 1 3 2 Rhagionidae 3 3 1 1 Tabanidae 2 1 1 2 Stratiomyidae 1 1 1 Therevidae 1 1 Asilidae 3 3 2 2 Hybotidae 3 2 2 Empididae 11 7 3 3 2 Dolichopodidae 1 1 Platypezidae 1 1 Phoridae 1 1 Syrphidae 34 19 5 14 25 2 Psilidae 2 1 2 Lonchaeidae 1 1 1 Tephritidae 4 3 2 1 Lauxaniidae 2 2 Sciomyzidae 2 1 1 1 Opomyzidae 3 2 2 Heleomyzidae 1 1 Drosophilidae 1 1 Scathophagidae 5 2 1 5 1 Anthomyiidae 9 4 4 6 Fanniidae 1 1 Muscidae 19 7 2 5 12 3 Calliphoridae 10 6 1 4 5 1 Rhinophoridae 1 1 Sarcophagidae 5 1 1 3 4 Tachinidae 8 3 2 4 6 1 Species Total 158 75 13 52 102 15 4

13 ACKNOWLEDGEMENTS

Pat and Jim Bone deserve special credit for their wonderful photographs and butterfly-counting, and I was very saddened to hear from Pat that Jim had recently passed away. I found Jim to be a real gentleman who was consistently pleasant, enthusiastic and knowledgeable, and a pleasure to chat to – he will be sorely missed by all who knew him

Special mention must also be made of Bill Ely for once again being kind enough to identify parasitic wasps in the Ichneumonidae and Braconidae families. These provided quite a number of additions to the site list, many of which were from specimens captured during 2017.

Richard Baker helped fill a gap in the 2018 studies by running a light trap on 10th August, and kindly supplied a list of moths and photographs from his trapping exercise. Geoff Oxford also supplied a list of spiders from his visit to Three Hagges Wood-meadow on 7th October. Others who made a contribution were: Reijo Jussila, who identified a problematic ichneumon; and Rosalind Forbes Adam, who supplied a photograph of the dragonfly Libellula depressa [Broad-bodied Chaser].

REFERENCES

Alexander, K. N. A. (2014) A review of the beetles of Great Britain: The Soldier Beetles and their allies. Species Status 16. Natural England Commissioned Report NECR134. Natural England. 1-81.

Bratton, J. H. [ed.] (1991) Invertebrates other than insects. British Red Data Books 3. Peterborough: Joint Nature Conservation Committee. 1-253.

Falk, S. [J.] (1991) A review of the scarce and threatened bees, wasps and ants of Great Britain. Research & survey in nature conservation 35. Peterborough: Nature Conservancy Council. 1-344.

Grayson, A. (2014) An Inventory of Invertebrates recorded from Three Hagges Jubilee Wood during 2014. [Unpublished report to Hagge Woods Trust, Escrick, York]. 1-139.

Grayson, A. (2016) An Inventory of Invertebrates recorded from Three Hagges Jubilee Wood during 2015. [Unpublished report to Hagge Woods Trust, Escrick, York]. 1-161.

Grayson, A. (2017) An Inventory of Invertebrates recorded from Three Hagges Wood-meadow during 2016 and Updates on Previous Surveys. [Unpublished report to Hagge Woods Trust, Escrick, York]. 1-253.

Grayson, A. (2018) An Inventory of Invertebrates recorded from Three Hagges Wood-meadow during 2017 and Updates on Previous Surveys. [Unpublished report to Hagge Woods Trust, Escrick, York]. 1-313.

Shirt, D. B. [ed.] (1987) Insects. British Red Data Books 2. Peterborough: Nature Conservancy Council. 1-402.

14 APPENDIX 1: PLANS

THREE HAGGES WOOD-MEADOW AND ITS ENVIRONS

15 EXTENTS OF SURVEY COMPARTMENTS AND LOCATIONS OF FEATURES

16 APPENDIX 2: TABLES

FULL INVERTEBRATE LIST (2014 TO 2018)

This list is an inventory of all invertebrates thus far identified from surveys carried out at Three Hagges Wood-meadow between 2014 and 2018. The list contains 990 species (including a few that could not be identified beyond genus-level).

For simplification and ease of reference, genera and species are arranged in alphabetical order and subgenera are omitted. Current nomenclature is accepted, except in problematic cases. The appendages [s. l.] and [s. s.] indicate either a precise species from a species-complex [sensu stricto], or a species-complex [sensu lato]. This list only mentions sensu lato records when sensu stricto records are unavailable for any particular species.

‘C’ or ‘L’ are used for all species whose national [British] or regional [Yorkshire] statuses are common or local [or equivalent statuses], including common or irregular migrant species. ‘U’ is used where the national or regional status is unknown due to insufficient data being available; and ‘n/a’ is used where an identification is not down to species level, hence, a status allocation is not applicable. ‘RS’ denotes regionally scarce species, i.e. species which are known to be scarce in Yorkshire, or species which might be considered regionally scarce based on such factors as scarcity of available habitat, being on the extremity of their national range, or paucity of records etc. ‘SPIE’ denotes Species of Principal Importance in England [formerly known as UK BAP Species]. ‘JNCC’ indicates a current JNCC national status classification of Notable or higher status, e.g. Red Data Book, [or equivalent IUCN-based statuses].

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Class: Gastropoda Slugs & snails etc. Family: Agriolimacidae Terrestrial slugs Deroceras reticulatum (Müller) Grey Field Slug 08/09/2015 C C Family: Arionidae Terrestrial slugs Arion ater (Linnaeus) [s. l.] Black Slug 23/04/2014 C C Arion rufus (Linnaeus) Red Slug 05/05/2016 C C Family: Helicidae Terrestrial snails Cepaea hortensis (Müller) White-lipped Snail 11/04/2016 C C Cepaea nemoralis (Linnaeus) Brown-lipped Snail 05/05/2016 C C Cornu aspersum (Müller) Garden Snail 11/04/2016 C C Family: Hygromiidae Terrestrial snails Monacha cantiana (Montagu) Kentish Snail 05/05/2016 C C Family: Lymnaeidae Freshwater snails Lymnaea stagnalis (Linnaeus) Great Pond Snail 05/05/2016 C C Family: Planorbidae Ram’s-horn snails Bathyomphalus contortus (Linnaeus) Twisted Ram’s-horn Snail 04/08/2016 C C Planorbarius corneus (Linnaeus) Great Ram’s-horn Snail 05/05/2016 C C Order: Veneroida Clams & cockles etc. Family: Sphaeriidae Orb-mussels Sphaerium corneum (Linnaeus) European Fingernail-clam 04/08/2016 C C Order: Araneae Spiders Family: Araneidae Orb-weaver spiders etc. Araneus diadematus Clerck Garden Spider 28/08/2014 C C Araneus marmoreus Clerck var. pyramidatus Clerck Marled Orb-weaver Spider 07/10/2018 L L Araneus quadratus Clerck Four-spot Orb-weaver Spider 07/10/2018 C L Araniella cucurbitina (Clerck) Cucumber Green Spider 13/08/2015 C U Larinioides cornutus (Clerck) Furrow Spider 22/04/2017 C C Family: Philodromidae Slender Crab Spiders Tibellus species a slender crab spider 07/10/2018 n/a n/a 17 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Family: Pisauridae Nursery web spiders Pisaura mirabilis (Clerck) Nursery Web Spider 08/07/2015 C L Family: Tetragnathidae Long-jawed orb-weavers Metellina segmentata (Clerck) a long-jawed orb-weaver 25/08/2016 C U Family: Theridiidae Cobweb spiders Enoplognatha latimana Hippa & Oksala Scarce Candy-striped spider 07/10/2018 L U Enoplognatha ovata (Clerck) Candy-striped Spider 18/07/2015 C U Order: Amphipoda Sand-hoppers & scuds etc. Family: Crangonyctidae Amphipod shrimps Crangonyx pseudogracilis Bousfield an amphipod shrimp 04/08/2016 C C Order: Glomerida Pill millipedes Family: Glomeridae Pill millipedes Glomeris marginata (Villers) Pill Millipede 03/06/2015 C C Order: Polydesmida Flat-backed millipedes etc. Family: Polydesmidae Flat-backed millipedes Polydesmus angustus Latzel Common Flat-backed Millipede 28/01/2018 C C Polydesmus denticulatus C. L. Koch a flat-backed millipede 03/05/2017 C C Order: Lithobiomorpha Stone centipedes Family: Lithobiidae Stone centipedes Lithobius forficatus (Linnaeus) Brown Centipede 20/02/2018 C C Order: Julida Millipedes Family: Julidae Millipedes Tachypodoiulus niger (Leach) Black Snake Millipede 05/05/2016 C C Order: Isopoda Woodlice & water-lice etc. Suborder: Asellota Water-lice etc. Family: Asellidae Water-lice Asellus aquaticus (Linnaeus) Common Water-louse 04/08/2016 C C Suborder: Oniscidea Woodlice Family: Oniscidae Woodlice Oniscus asellus Linnaeus Common Woodlouse 26/03/2014 C C Philoscia muscorum (Scopoli) Common Striped Woodlouse 26/03/2014 C C Family: Porcellionidae Woodlice Porcellio scaber Latreille Common Rough Woodlouse 01/07/2015 C C Family: Armadillidiidae Pill woodlice Armadillidium vulgare (Latreille) Common Pill Woodlouse 01/07/2015 C C Order: Ephemeroptera Mayflies Family: Baetidae Mayflies Cloeon dipterum (Linnaeus) Pond Olive Mayfly 04/08/2016 C C Family: Ephemeridae Mayflies Ephemera vulgata Linnaeus Drake Mackerel Mayfly 09/05/2016 C C Order: Odonata Dragonflies & damselflies Suborder: Zygoptera Damselflies Family: Coenagriidae Blue damselflies etc. Coenagrion puella (Linnaeus) Azure Damselfly 15/05/2014 C C Enallagma cyathigerum (Charpentier) Common Blue Damselfly 15/05/2014 C C Ischnura elegans (Vander Linden) Blue-tailed Damselfly 12/06/2014 C C Pyrrhosoma nymphula (Sulzer) Large Red Damselfly 21/05/2015 C C

18 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Family: Lestidae Emerald damselflies Lestes sponsa (Hansemann) Emerald Damselfly 12/07/2017 C C Family: Calopterygidae Demoiselles Calopteryx splendens (Harris) Banded Demoiselle 13/07/2017 C L Suborder: Anisoptera Dragonflies Family: Aeshnidae Hawkers Aeshna cyanea (Müller) Southern Hawker 05/08/2014 C C Aeshna grandis (Linnaeus) Brown Hawker 08/07/2014 C C Aeshna mixta (Latreille) Migrant Hawker 01/09/2016 C C Anax imperator Leach Emperor Dragonfly 23/07/2017 C L Family: Libellulidae Skimmers & darters etc. Libellula depressa Linnaeus Broad-bodied Chaser 22/07/2014 C C Libellula quadrimaculata Linnaeus Four-spotted Chaser 12/06/2014 C C Orthetrum cancellatum (Linnaeus) Black-tailed Skimmer 06/06/2016 C L Sympetrum danae (Sulzer) Black Darter 04/08/2018 C L Sympetrum striolatum (Charpentier) Common Darter 09/07/2014 C C Order: Plecoptera Stoneflies Family: Nemouridae Stoneflies Nemoura cinerea (Retzius) a stonefly 01/06/2017 C C Order: Orthoptera Grasshoppers etc. Family: Tetrigidae Groundhoppers Tetrix subulata (Linnaeus) Slender Groundhopper 23/04/2014 C C Tetrix undulata (Sowerby) Common Groundhopper 15/05/2014 C C Family: Acrididae Grasshoppers Chorthippus brunneus (Thunberg) Common Field Grasshopper 22/07/2014 C C Chorthippus parallelus (Zetterstedt) Meadow Grasshopper 22/07/2014 C C Omocestus viridulus (Linnaeus) Common Green Grasshopper 22/07/2014 C C Order: Dermaptera Earwigs Family: Forficulidae Earwigs Forficula auricularia Linnaeus Common Earwig 08/07/2014 C C Order: Hemiptera True bugs etc. Suborder: Heteroptera True bugs Family: Acanthosomatidae Shieldbugs Acanthosoma haemorrhoidale (Linnaeus) Hawthorn Shieldbug 01/11/2018 C C Elasmostethus interstinctus (Linnaeus) Birch Shieldbug 09/08/2014 C L Elasmucha grisea (Linnaeus) Parent Bug 05/05/2016 C L Family: Coreidae Squashbugs etc. Coriomeris denticulatus (Scopoli) Denticulate Leatherbug 01/07/2015 C L Family: Cydnidae Shieldbugs Tritomegas bicolor (Linnaeus) Pied Shieldbug 05/05/2016 C L Family: Pentatomidae Shieldbugs Dolycoris baccarum (Linnaeus) Hairy Shieldbug 15/05/2014 C L Palomena prasina (Linnaeus) Green Shieldbug 23/04/2014 C L Pentatoma rufipes (Linnaeus) Red-legged Shieldbug 04/08/2016 C C Family: Rhopalidae Plant bugs Rhopalus subrufus (Gmelin) a plant bug 05/05/2016 L RS Stictopleurus punctatonervosus (Goeze) a plant bug 09/05/2016 L RS

19 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Family: Anthocoridae Flower-bugs Anthocoris nemoralis (Fabricius) a flower bug 05/05/2016 C C Anthocoris nemorum (Linnaeus) Common Flower-bug 08/07/2014 C C Family: Berytidae Stilt bugs Berytinus minor (Herrich-Schäffer) a stilt bug 01/09/2016 C C Family: Lygaeidae Ground bugs Drymus ryei Douglas & Scott a ground bug 01/09/2016 C L Drymus sylvaticus (Fabricius) a ground bug 05/05/2016 C C Kleidocerys resedae (Panzer) Birch Catkin Bug 09/04/2017 C L Scolopostethus affinis (Schilling) a ground bug 05/05/2016 C C Stygnocoris fuligineus (Geoffroy) a ground bug 05/05/2016 C C Family: Miridae Grass-bugs etc. Adelphocoris lineolatus (Goeze) Lucerne Bug 07/09/2017 C L Blepharidopterus angulatus (Fallén) Black-kneed Capsid 04/08/2016 C C Campyloneura virgula (Herrich-Schäffer) a plant bug 04/08/2016 C C Capsus ater (Linnaeus) a plant bug 08/07/2014 L L Closterotomus norwegicus (Gmelin) Potato Capsid 08/07/2014 C C Deraeocoris flavilinea (Costa) a plant bug 09/07/2014 L L Deraeocoris lutescens (Schilling) a plant bug 01/09/2016 C L Deraeocoris ruber (Linnaeus) a plant bug 04/08/2016 C L Dicyphus stachydis J. Sahlberg a plant bug 01/09/2016 C L Heterotoma planicornis (Pallas) a plant bug 04/08/2016 C C Leptopterna dolabrata (Linnaeus) Meadow Plant Bug 08/07/2014 C C Liocoris tripustulatus (Fabricius) a plant bug 05/05/2016 C C Lopus decolor (Fallén) a grass bug 04/08/2016 C C Lygocoris pabulinus (Linnaeus) Common Green Capsid 08/07/2014 C C Lygocoris rugicollis (Fallén) Apple Capsid 04/08/2018 L U Lygus rugulipennis (Poppius) Tarnished Plant Bug 08/07/2014 C C Megaloceroea recticornis (Geoffroy) a grass bug 02/07/2017 C L Neolygus contaminatus (Fallén) a plant bug 04/08/2016 C C Neolygus viridis (Fallén) a plant bug 04/08/2016 C C Notostira elongata (Geoffroy) a grass bug 08/07/2014 C C Oncotylus viridiflavus (Goeze) a plant bug 04/08/2016 L U Orthops basalis (A. Costa) a plant bug 05/05/2016 L U Orthops campestris (Linnaeus) a plant bug 04/08/2018 C U Phytocoris tiliae (Fabricius) a plant bug 04/08/2016 C C Phytocoris varipes Boheman a plant bug 14/08/2016 C C Pinalitus cervinus (Herrich-Schäffer) a plant bug 04/08/2016 C C Pithanus maerkelii (Herrich-Schäffer) a plant bug 02/07/2017 L U Plagiognathus arbustorum (Fabricius) a plant bug 08/07/2014 C C Plagiognathus chrysanthemi (Wolff) a plant bug 04/08/2016 C C Psallus betuleti (Fallén) a plant bug 02/06/2017 L U Rhabdomiris striatellus (Fabricius) a plant bug 21/05/2017 C L Stenodema calcarata (Fallén) a grass bug 08/07/2014 C C Stenodema laevigata (Linnaeus) a grass bug 05/05/2016 C C Stenotus binotatus (Fabricius) a plant bug 04/08/2016 C C Family: Nabidae Damsel bugs Himacerus apterus (Fabricius) Tree Damsel Bug 04/08/2016 C L 20 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Himacerus major (A. Costa) Grey Damsel Bug 04/08/2016 C RS Himacerus mirmicoides (O. Costa) Ant Damsel Bug 04/08/2016 C L Nabis ferus (Linnaeus) Field Damsel Bug 02/07/2017 C L Nabis flavomarginatus Scholtz Broad Damsel Bug 04/08/2016 C C Nabis limbatus Dahlbom Marsh Damsel Bug 04/08/2016 C L Nabis rugosus (Linnaeus) Common Damsel Bug 05/05/2016 C L Family: Tingidae Lacebugs Tingis ampliata (Herrich- Schäffer) a lacebug 05/05/2016 C C Tingis cardui (Linnaeus) Spear Thistle Lacebug 05/05/2016 C RS Family: Corixidae Water boatmen Corixa punctata (Illiger) Common Water Boatman 08/07/2014 C C Sigara nigrolineata (Fieber) a water boatman 04/08/2016 C C Family: Gerridae Pond skaters Gerris lacustris (Linnaeus) Common Pond Skater 05/05/2016 C C Gerris odontogaster (Zetterstedt) Toothed Pond Skater 04/08/2016 C C Gerris thoracicus Schummel a pond skater 08/07/2014 C C Family: Notonectidae Backswimmers Notonecta glauca Linnaeus Common Backswimmer 04/08/2016 C C Notonecta maculata Fabricius a backswimmer 04/08/2016 L L Family: Pleidae Pygmy backswimmers Plea minutissima Leach Pygmy Backswimmer 04/08/2016 C C Family: Saldidae Shore bugs Saldula saltatoria (Linnaeus) Common Shore Bug 08/07/2014 C C Suborder: Auchenorrhyncha Planthoppers etc. Family: Cercopidae Froghoppers Cercopis vulnerata Illiger Black-and-red Froghopper 15/05/2014 C C Family: Aphrophoridae Froghoppers Aphrophora alni (Fallén) Alder Spittlebug 04/08/2016 C C Neophilaenus lineatus (Linnaeus) a froghopper 04/08/2016 C C Philaenus spumarius (Linnaeus) Common Froghopper 04/08/2016 C C Family: Cicadellidae Leafhoppers Adarrus ocellaris (Fallén) a leafhopper 01/09/2016 C C Allygus communis (Ferrari) a leafhopper 04/08/2016 U U Allygus modestus Scott a leafhopper 04/08/2016 C U Anoscopus flavostriatus (Donovan) a leafhopper 01/09/2016 C U Aphrodes makarovi Zakhvatkin a leafhopper 04/08/2016 C L Arthaldeus pascuellus (Fallén) a leafhopper 01/09/2016 C C Athysanus argentarius Metcalf a leafhopper 04/08/2016 C RS Balclutha punctata (Fabricius) a leafhopper 04/08/2016 C C Eupteryx aurata (Linnaeus) Potato Leafhopper 01/09/2016 C C Euscelis incisus (Kirschbaum) a leafhopper 01/06/2017 C C Euscelis lineolatus Brullé a leafhopper 05/05/2016 C L Kybos strigilifer (Ossiannilsson) a leafhopper 04/08/2016 C U Macrosteles ossiannilssoni Lindberg a leafhopper 01/09/2016 L U Megophthalmus scabripennis Edwards a leafhopper 01/09/2016 C L Metidiocerus rutilans (Kirschbaum) a leafhopper 01/09/2016 L L Mocydia crocea (Herrich-Schäffer) a leafhopper 01/09/2016 C C Oncopsis alni (Schrank) a leafhopper 04/08/2016 C C 21 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Oncopsis flavicollis (Linnaeus) a leafhopper 04/08/2016 C C Populicerus confusus (Flor) a leafhopper 04/08/2016 C C Zyginidia scutellaris (Herrich-Schäffer) a leafhopper 01/09/2016 C C Family: Cixiidae Planthoppers Cixius nervosus (Linnaeus) a planthopper 04/08/2016 C C Tachycixius pilosus (Olivier) a planthopper 02/06/2017 C C Family: Delphacidae Planthoppers Conomelas anceps (Germar) a planthopper 01/09/2016 C C Javesella dubia (Kirschbaum) a planthopper 04/08/2016 C C Javesella pellucida (Fabricius) a planthopper 05/05/2016 C C Stenocranus major (Kirschbaum) a planthopper 01/09/2016 L U Suborder: Sternorrhyncha Aphids & scale insects etc. Family: Aphididae Aphids Acyrthosiphon pisum (Harris) Pea Aphid 08/07/2014 C C Uroleucon jaceae (Linnaeus) an aphid 08/07/2014 C C Order: Neuroptera Lacewings etc. Family: Sialidae Alder flies Sialis lutaria (Linnaeus) an alder fly 04/08/2016 C C Family: Hemerobiidae Brown lacewings Hemerobius micans Olivier a brown lacewing 04/08/2016 C C Micromus variegatus (Fabricius) a brown lacewing 03/05/2017 C C Wesmaelius quadrifasciatus (Reuter) a brown lacewing 21/07/2016 L L Family Chrysopidae Green lacewings Chrysopa perla (Linnaeus) a green lacewing 02/06/2017 C C Chrysoperla carnea (Stephens) [s. s.] Common Green Lacewing 23/04/2014 C C Order: Mecoptera Scorpion flies etc. Family: Panorpidae Scorpion flies etc. Panorpa cognata Rambur a scorpion-fly 03/07/2017 L L Panorpa communis Linnaeus Common Scorpion-fly 01/07/2015 C C Order: Lepidoptera Butterflies & moths Suborder: Heterocera Moths Family: Eriocraniidae Micro moths Eriocrania sparrmannella (Bosc) a micro moth 22/04/2018 L RS Family: Hepialidae Swift moths Hepialus humuli (Linnaeus) Ghost Moth 18/06/2014 SPIE C Korscheltellus lupulina (Linnaeus) Common Swift 16/06/2015 C C Triodia sylvina (Linnaeus) Orange Swift 19/08/2014 C L Family: Adelidae Longhorn moths Cauchas rufimitrella (Scopoli) a longhorn moth 03/05/2017 C C Nemophora degeerella (Linnaeus) a longhorn moth 02/06/2017 C C Family: Yponomeutidae Ermine moths etc. Yponomeuta cagnagella (Hübner) Spindle Ermine 31/05/2018 C L Family: Glyphipterigidae Micro moths Glyphipterix simpliciella (Stephens) Cocksfoot Moth 22/07/2014 C C Family: Chimabachidae Micro moths Diurnea fagella (Denis & Schiffermüller) a micro moth 10/04/2015 C C Family: Gelechiidae Micro moths Syncopacma larseniella Gozmány a micro moth 02/07/2017 L U 22 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Family: Pterophoridae Plume moths Gillmeria pallidactyla (Howarth) Yarrow Plume moth 02/07/2017 C C Family: Tortricidae Leafroller moths etc. Aethes smeathmanniana (Fabricius) a micro moth 22/07/2014 L L Agapeta hamana (Linnaeus) a micro moth 18/06/2014 C C Apotomis turbidana Hübner White-shouldered Marble 01/06/2017 C C Celypha lacunana (Denis & Schiffermüller) a micro moth 25/06/2014 C C Eucosma campoliliana (Denis & Schiffermüller) a micro moth 05/07/2014 C C Eucosma cana (Haworth) Hoary Bell 18/06/2014 C C Grapholita compositella (Fabricius) Clover Seed Moth 09/08/2014 L L Hedya ochroleucana (Frölich) Buff-tipped Marble 09/08/2014 C RS Hedya pruniana (Hübner) Plum Tortrix 18/06/2014 C C Pammene aurana (Fabricius) a micro-moth 02/06/2017 C L Pandemis cerasana (Hübner) Barred Fruit-tree Tortrix 18/06/2014 C C Pandemis corylana (Fabricius) Chequered Fruit-tree Tortrix 18/06/2014 C C Spilonota ocellana (Denis & Schiffermüller) Bud Moth 05/07/2014 C C Family: Sesiidae Clearwing moths Synanthedon vespiformis (Linnaeus) Yellow-legged Clearwing 04/08/2017 L RS Family: Pyralidae Snout moths etc. Aphomia sociella (Linnaeus) Bee Moth 18/06/2014 C C Hypsopygia costalis (Fabricius) Gold Triangle 18/06/2014 C C Family: Zygaenidae Burnet moths Zygaena filipendulae (Linnaeus) Six-spot Burnet 25/06/2014 C L Zygaena lonicerae (Scheven) Narrow-bordered Five-spot Burnet 08/07/2014 C L Family: Crambidae Grass moths Acentria ephemerella (Denis & Schiffermüller) Water Veneer 09/08/2014 L C Agriphila straminella (Denis & Schiffermüller) a grass moth 09/08/2014 C C Agriphila tristella (Denis & Schiffermüller) a grass moth 09/08/2014 C C Catoptria falsella (Denis & Schiffermüller) a grass moth 09/08/2014 L L Catoptria pinella (Linnaeus) a grass moth 09/08/2014 L L Chrysoteuchia culmella (Linnaeus) Garden Grass-veneer 25/06/2014 C C Crambus lathoniellus (Zincken) a grass moth 18/06/2014 C C Crambus pascuella (Linnaeus) a grass moth 09/08/2014 C C Crambus perlella (Scopoli) a grass moth 18/06/2014 C C Pleuroptya ruralis (Scopoli) Mother of Pearl 09/08/2014 C C Scoparia ambigualis (Treitschke) a grass moth 18/06/2014 C C Family: Drepanidae Hook-tip moths Cilix glaucata (Scopoli) Chinese Character 09/08/2014 C L Drepana falcataria (Linnaeus) Pebble Hook-tip 11/09/2016 C L Falcaria lacertinaria (Linnaeus) Scalloped Hook-tip 19/08/2014 C L Habrosyne pyritoides (Hufnagel) Buff Arches 05/07/2014 C L Ochropacha duplaris (Linnaeus) Common Lutestring 05/07/2014 C C Family: Lasiocampidae Eggars etc. Euthrix potatoria (Linnaeus) The Drinker 18/06/2014 C C Family: Sphingidae Hawk-moths Deilephila elpenor (Linnaeus) Elephant Hawk-moth 18/06/2014 C C Smerinthus ocellata (Linnaeus) Eyed Hawk-moth 18/06/2014 L L

23 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Family: Geometridae Carpet moths etc. Alcis repandata (Linnaeus) Mottled Beauty 05/07/2014 C C Archiearis parthenias (Linnaeus) Orange Underwing 11/04/2016 L L Biston betularia (Linnaeus) Peppered Moth 18/06/2014 C C Cabera exanthemata (Scopoli) Common Wave 16/06/2015 C C Cabera pusaria (Linnaeus) Common White Wave 18/06/2014 C C Campaea margaritaria (Linnaeus) Light Emerald 18/06/2014 C C Camptogramma bilineata (Linnaeus) Yellow Shell 25/06/2014 C C Colostygia pectinataria (Knoch) Green Carpet 16/06/2015 C C Dysstroma truncata (Hufnagel) Common Marbled Carpet 09/08/2014 C C Ectropis bistortata (Goeze) The Engrailed 05/07/2014 C C Ennomos alniaria (Linnaeus) Canary-shouldered Thorn 19/08/2014 C C Epirrhoe alternata (Müller) Common Carpet 18/06/2014 C C Eupithecia abbreviata Stephens Brindled Pug 10/04/2015 C L Eupithecia centaureata (Denis & Schiffermüller) Lime-speck Pug 09/08/2014 C L Eupithecia vulgata (Haworth) Common Pug 05/07/2014 C C Hydriomena furcata (Thunberg) July Highflyer 09/08/2014 C C Idaea aversata (Linnaeus) Riband Wave 18/06/2014 C C Idaea biselata (Hufnagel) Small Fan-footed Wave 05/07/2014 C L Idaea dimidiata (Hufnagel) Single-dotted Wave 05/07/2014 C L Lampropteryx suffumata (Denis & Schiffermüller) Water Carpet 10/04/2015 C L Lomaspilis marginata (Linnaeus) Clouded Border 05/07/2014 C C Lomographa temerata (Denis & Schiffermüller) Clouded Silver 18/06/2014 C L Macaria liturata (Clerck) Tawny-barred Angle 16/06/2015 C L Mesotype didymata (Linnaeus) Twin-spot Carpet 09/08/2014 C C Ourapteryx sambucaria (Linnaeus) Swallow-tailed Moth 02/10/2014 C C Peribatodes rhomboidaria (Denis & Schiffermüller) Willow Beauty 18/06/2014 C C Petrophora chlorosata (Scopoli) Brown Silver-line 18/06/2014 C C Scotopteryx chenopodiata (Linnaeus) Shaded Broad-bar 05/07/2014 SPIE C Pennithera firmata (Hübner) Pine Carpet 02/10/2014 L L Thera obeliscata (Hübner) Grey Pine Carpet 02/10/2014 C C Timandra comae Schmidt Blood-vein 09/08/2014 SPIE L Xanthorhoe ferrugata (Clerck) Dark-barred Twin-spot Carpet 09/08/2014 SPIE L Xanthorhoe montanata (Denis & Schiffermüller) Silver-ground Carpet 18/06/2014 C C Xanthorhoe quadrifasiata (Clerck) Large Twin-spot Carpet 18/06/2014 L L Family: Notodontidae Kitten & prominent moths Furcula furcula (Clerck) Sallow Kitten 19/08/2014 C L Notodonta dromedarius (Linnaeus) Iron Prominent 09/08/2014 C L Phalera bucephala (Linnaeus) Buff-tip 18/07/2015 C C Pheosia gnoma (Fabricius) Lesser Swallow Prominent 09/08/2014 C C Ptilodon capucina (Linnaeus) Coxcomb Prominent 19/08/2014 C C Family: Erebidae Tiger & footman moths etc. Arctia caja (Linnaeus) Garden Tiger 18/07/2015 SPIE L Callistege mi (Clerck) Mother Shipton Moth 03/06/2015 L L Calliteara pudibunda (Linnaeus) Pale Tussock 16/06/2015 C L Eilema depressa (Esper) Buff Footman 09/08/2014 L RS Eilema griseola (Hübner) Dingy Footman 05/08/2014 C RS Eilema lurideola (Zincken) Common Footman 05/07/2014 C C 24 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Euclidia glyphica (Linnaeus) Burnet Companion 03/06/2018 L RS Euproctis similis (Fuessly) Yellow-tail 05/07/2014 C C Herminia tarsipennalis Treitschke The Fan-foot 18/06/2014 C L Hypena proboscidalis (Linnaeus) The Snout 02/10/2014 C C Leucoma salicis (Linnaeus) White Satin Moth 05/07/2014 L L Orgyia antiqua (Linnaeus) The Vapourer 15/07/2014 C C Phragmatobia fuliginosa (Linnaeus) Ruby Tiger 03/10/2014 C L Spilosoma lubricipeda (Linnaeus) White Ermine 22/05/2016 SPIE C Spilosoma lutea (Hufnagel) Buff Ermine 18/07/2015 SPIE C Tyria jacobaeae (Linnaeus ) The Cinnabar 12/06/2014 SPIE C Family: Noctuidae Owlet moths Acronicta leporina (Linnaeus) The Miller 18/06/2014 L L Acronicta rumicis (Linnaeus) Knot Grass 25/08/2016 C C Agrotis exclamationis (Linnaeus) Heart and Dart 18/06/2014 C C Agrotis puta (Hübner) Shuttle-shaped Dart 10/08/2018 C L Amphipoea oculea (Linnaeus) Ear Moth 09/08/2014 SPIE RS Amphipyra tragopoginis (Clerck) Mouse Moth 09/08/2014 SPIE C Anarta trifolii (Hufnagel) The Nutmeg 18/06/2014 L L Anorthoa munda (Denis & Schiffermüller) Twin-spotted Quaker 10/04/2015 L L Apamea monoglypha (Hufnagel) Dark Arches 05/07/2014 C C Autographa gamma (Linnaeus) Silver Y 15/05/2014 C C Axylia putris (Linnaeus) The Flame 18/06/2014 C L Deltote pygarga (Hufnagel) Marbled White Spot 18/06/2014 L RS Diarsia mendica (Fabricius) Ingrailed Clay 05/07/2014 C C Diarsia rubi (Vieweg) Small Square-spot 16/06/2015 SPIE C Enargia paleacea (Esper) Angle-striped Sallow 10/08/2018 L L Euplexia lucipara (Linnaeus) Small Angle Shades 18/06/2014 C L Hoplodrina blanda (Denis & Schiffermüller) The Rustic 18/07/2015 SPIE L Hoplodrina octogenaria (Goeze) The Uncertain 18/06/2014 C L Hydraecia micacea (Esper) Rosy Rustic 21/08/2016 SPIE C Lithophane ornitopus (Dadd) Grey-shoulder Knot 20/02/2018 L RS Luperina testacea (Denis & Schiffermüller) Flounced Rustic 19/08/2014 C C Mesapamea secalis (Linnaeus) [s. l.] Common Rustic 09/08/2014 C C Mythimna ferrago (Fabricius) The Clay 18/06/2014 C C Mythimna impura (Hübner) Smoky Wainscot 05/07/2014 C C Noctua comes (Hübner) Lesser Yellow Underwing 09/08/2014 C C Noctua interjecta Hübner Least Yellow Underwing 09/08/2014 L L Noctua janthe (Borkhausen) Lesser Broad-bordered Yellow Underwing 09/08/2014 C C Noctua pronuba (Linnaeus) Large yellow Underwing 09/08/2014 C C Ochropleura plecta (Linnaeus) Flame Shoulder 18/06/2014 C C Oligia fasciuncula (Haworth) Middle-barred Minor 18/06/2014 C C Oligia strigilis (Linnaeus) [s. l.] Marbled Minor 18/06/2014 C C Omphaloscelis lunosa (Howarth) Lunar Underwing 10/10/2015 C L Orthosia cerasi (Fabricius) Common Quaker 10/04/2015 C C Orthosia gothica (Linnaeus) Hebrew Character 10/04/2015 C C Orthosia gracilis (Denis & Schiffermüller) Powdered Quaker 06/05/2016 L L Panemeria tenebrata (Scopoli) Small Yellow Underwing 15/05/2014 L L Panolis flammea (Denis & Schiffermüller) Pine Beauty 10/04/2015 C L 25 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Phlogophora meticulosa (Linnaeus) Angle Shades 19/08/2014 C C Plusia festucae (Linnaeus) Gold Spot 26/09/2015 L L Rivula sericealis (Scopoli) Straw Dot 18/06/2014 L L Tiliacea aurago (Denis & Schiffermüller) Barred Sallow 02/10/2014 L L Xestia baja (Denis & Schiffermüller) Dotted Clay 09/08/2014 C L Xestia c-nigrum (Linnaeus) Setaceous Hebrew Character 09/08/2014 C C Xestia sexstigata (Howarth) Six-striped Rustic 09/08/2014 C C Xestia triangulum (Hufnagel) Double Square-spot 18/06/2014 L C Xylocampa areola (Esper) Early Grey 10/04/2015 C L Family: Nolidae Owlet moths Pseudoips prasinana (Linnaeus) Green Silver-lines 18/06/2014 C L Suborder: Rhopalocera Butterflies Family: Hesperiidae Skippers Erynnis tages (Linnaeus) Dingy Skipper 03/06/2018 SPIE L Ochlodes sylvanus (Esper) Large Skipper 09/07/2014 C C Thymelicus sylvestris (Poda) Small Skipper 05/08/2014 C C Family: Pieridae Whites & yellows etc. Anthocharis cardamines (Linnaeus) Orange Tip 23/04/2014 C C Colias crocea (Geoffroy) Clouded Yellow 09/08/2014 L RS Gonepteryx rhamni (Linnaeus) Brimstone 23/04/2014 C C Pieris brassicae (Linnaeus) Large White 15/05/2014 C C Pieris napi (Linnaeus) Green-veined White 23/04/2014 C C Pieris rapae (Linnaeus) Small White 23/04/2014 C C Family: Nymphalidae Brush-footed butterflies etc. Aglais io (Linnaeus) Peacock 23/04/2014 C C Aglais urticae (Linnaeus) Small Tortoiseshell 26/03/2014 C C Aphantopus hyperantus (Linnaeus) Ringlet 08/07/2014 C C Coenonympha pamphilus (Linnaeus) Small Heath 26/06/2015 SPIE L Maniola jurtina (Linnaeus) Meadow Brown 12/06/2014 C C Melanargia galathea (Linnaeus) Marbled White 06/07/2018 L L Pararge aegeria (Linnaeus) Speckled Wood 23/04/2014 C C Polygonia c-album (Linnaeus) Comma 23/08/2015 C C Pyronia tithonus (Linnaeus) Gatekeeper 05/08/2014 C C Vanessa atalanta (Linnaeus) Red Admiral 12/06/2014 C C Vanessa cardui (Linnaeus) Painted Lady 05/08/2014 C C Family: Lycaenidae Blues & hairstreaks etc. Aricia agestis (Denis & Schiffermüller) Brown Argus 12/06/2014 L L Celastrina argiolus (Linnaeus) Holly Blue 08/08/2015 C L Favonius quercus (Linnaeus) Purple Hairstreak 07/07/2018 C L Lycaena phlaeas (Linnaeus) Small Copper 15/05/2014 C C Polyommatus icarus (Rottemburg) Common Blue 06/06/2014 C C Order: Coleoptera Beetles Suborder: Adephaga Ground beetles etc. Family: Gyrinidae Whirligig beetles Gyrinus substriatus (Stephens) a whirligig beetle 08/07/2014 C C Family: Haliplidae Crawling water beetles Haliplus heydeni Wehncke a crawling water beetle 04/08/2016 L L Haliplus lineatocollis (Marsham) a crawling water beetle 04/08/2016 C C 26 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Haliplus ruficollis (De Geer) [s. s.] a crawling water beetle 04/08/2016 C C Family: Dytiscidae Water beetles Acilius sulcatus (Linnaeus) a water beetle 08/04/2015 C C Agabus bipustulatus (Linnaeus) a water beetle 04/08/2016 C C Agabus didymus (Olivier) a water beetle 04/08/2016 C C Agabus nebulosus (Forster) a water beetle 04/08/2016 C C Agabus sturmii (Gyllenhal) a water beetle 04/08/2016 C C Colymbetes fuscus (Linnaeus) a water beetle 04/08/2016 C C Dytiscus species a water beetle 04/08/2016 n/a n/a Hydroglyphus geminus (Fabricius) a water beetle 04/08/2016 L L Hydroporus melanarius Sturm a water beetle 04/08/2016 L L Hydroporus palustris (Linnaeus) a water beetle 04/08/2016 C C Hydroporus planus (Fabricius) a water beetle 04/08/2016 C C Hydroporus tristis (Paykull) a water beetle 04/08/2016 L L Hygrotus inaequalis (Fabricius) a water beetle 04/08/2016 C C Hyphydrus ovatus (Linnaeus) a water beetle 04/08/2016 C C Family: Carabidae Ground beetles Agonum marginatum (Linnaeus) a ground beetle 27/06/2015 C C Amara aenea (De Geer) a ground beetle 06/05/2018 C C Amara communis (Panzer) a ground beetle 05/05/2016 C L Amara convexior Stephens a ground beetle 05/05/2016 L RS Amara eurynota (Panzer) a ground beetle 08/07/2014 L L Amara familiaris (Duftschmid) a ground beetle 12/06/2014 C C Amara lunicollis Schiödte a ground beetle 07/07/2016 C L Amara ovata (Fabricius) a ground beetle 12/06/2014 C L Amara plebeja (Gyllenhal) a ground beetle 15/05/2014 C C Amara similata (Gyllenhal) a ground beetle 01/09/2017 C C Anchomenus dorsalis (Pontoppidan) a ground beetle 11/04/2016 C C Bembidion articulatum (Panzer) a ground beetle 02/06/2016 L RS Bembidion bruxellense Wesmael a ground beetle 08/07/2014 C L Bembidion illigeri Netolitzky a ground beetle 15/05/2014 L L Bembidion lampros (Herbst) a ground beetle 12/06/2014 C C Bembidion properans (Stephens) a ground beetle 08/04/2015 L L Bembidion quadrimaculatum (Linnaeus) a ground beetle 12/06/2014 L L Bembidion tetracolum Say a ground beetle 08/04/2015 C C Calathus fuscipes (Goeze) a ground beetle 02/07/2016 C C Calathus melanocephalus (Linnaeus) a ground beetle 26/09/2015 C C Calathus rotundicollis Dejean a ground beetle 02/07/2017 L L Cicindela campestris Linnaeus Green Tiger Beetle 08/04/2015 L L Curtonotus aulicus (Panzer) a ground beetle 04/08/2016 C C Cychrus caraboides (Linnaeus) a ground beetle 04/08/2018 L L Elaphrus riparius (Linnaeus) a ground beetle 15/05/2014 C C Harpalus affinis (Schrank) a ground beetle 09/05/2016 C L Harpalus rubripes (Duftschmid) a ground beetle 05/05/2016 L RS Harpalus rufipes (De Geer) a ground beetle 01/09/2017 C C Harpalus tardus (Panzer) a ground beetle 05/05/2016 L RS Loricera pilicornis (Fabricius) a ground beetle 12/06/2014 C C Nebria brevicollis (Fabricius) a ground beetle 21/05/2015 C C 27 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Nebria salina Fairmaire & Laboulbéne a ground beetle 23/04/2014 C L Notiophilus biguttatus (Fabricius) a ground beetle 15/05/2014 C C Ocys harpaloides (Audinet-Serville) a ground beetle 08/04/2015 C C Poecilus versicolor (Sturm) a ground beetle 23/04/2014 C C Pterostichus madidus (Fabricius) a ground beetle 21/07/2016 C C Pterostichus melanarius (Illiger) a ground beetle 08/09/2015 C C Pterostichus niger (Schaller) a ground beetle 07/09/2014 C C Pterostichus nigrita (Paykull) a ground beetle 04/08/2016 C C Pterostichus strenuus (Panzer) a ground beetle 11/04/2016 C C Pterostichus vernalis (Panzer) a ground beetle 01/09/2017 C L Trechus obtusus Erichson a ground beetle 28/01/2018 C C Suborder: Polyphaga Beetles and weevils Family: Helophoridae Water scavenger beetles Helophorus minutus [s. s.] a water scavenger beetle 04/08/2016 C C Family: Hydrophilidae Scavenger water beetles Anacaena lutescens (Stephens) a scavenger water beetle 04/08/2016 C C Enochrus species a scavenger water beetle 04/08/2016 n/a n/a Helochares lividus (Forster) a scavenger water beetle 04/08/2016 C C Hydrobius fuscipes (Linnaeus) a scavenger water beetle 04/08/2016 C C Laccobius bipunctatus (Fabricius) a scavenger water beetle 04/08/2016 C C Family: Hydraenidae Moss beetles Ochthbius minimus (Fabricius) a moss beetle 04/08/2016 C C Family: Silphidae Carrion beetles etc. Nicrophorus vespillo (Linnaeus) a sexton beetle 29/07/2015 C C Nicrophorus vespilloides (Herbst) a sexton beetle 05/07/2017 C C Silpha atrata (Linnaeus) a carrion beetle 09/05/2016 C C Silpha tristis (Illiger) a carrion beetle 21/05/2015 L RS Family: Staphylinidae Rove beetles Ocypus olens (Müller) Devil’s Coach Horse 08/09/2015 C C Stenus comma (LeConte) a rove beetle 03/06/2015 L RS Family: Scarabaeidae Dung beetles etc. Phyllopertha horticola (Linnaeus) Garden Chafer 02/06/2017 C L Family: Elateridae Click beetles Adrastus pallens (Fabricius) a click beetle 08/07/2014 L U Agriotes acuminatus (Stephens) a click beetle 03/06/2018 C L Athous haemorrhoidalis (Fabricius) a click beetle 21/05/2017 C C Cidnopus aeruginosus (Olivier) a click beetle 05/05/2016 L U Denticollis linearis (Linnaeus) a click beetle 02/06/2015 C U Kibunea minuta (Linnaeus) a click beetle 06/05/2018 C U Family: Cantharidae Soldier beetles Cantharis cryptica Ashe a soldier beetle 02/06/2016 C C Cantharis livida Linnaeus a soldier beetle 03/06/2015 C C Cantharis nigra (De Geer) a soldier beetle 01/07/2015 C C Cantharis nigricans (Müller) a soldier beetle 03/06/2015 C C Cantharis pallida Goeze a soldier beetle 06/06/2016 L C Cantharis pellucida Fabricius a soldier beetle 02/06/2016 C C Cantharis rufa Linnaeus a soldier beetle 03/06/2015 C C Cantharis rustica Fallén a soldier beetle 15/05/2014 C C 28 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Rhagonycha fulva (Scopoli) a soldier beetle 08/07/2014 C C Family: Malachiidae Soft-wing flower beetles Malachius bipustulatus (Linnaeus) Malachite Beetle 15/05/2014 C C Family: Nitidulidae Pollen beetles etc. Glischrochilus hortensis (Geoffroy) a picnic beetle 08/04/2015 C C Meligethes aeneus (Fabricius) Rape Fly 08/07/2014 C C Family: Coccinellidae Ladybirds Adalia bipunctata (Linnaeus) 2-spot Ladybird 21/05/2015 C C Adalia decempunctata (Linnaeus) 10-spot Ladybird 09/04/2017 C C Calvia quattuordecimguttata (Linnaeus) Cream-spot Ladybird 20/06/2016 C C Coccinella septempunctata (Linnaeus) 7-spot Ladybird 26/03/2014 C C Exochomus quadripustulatus (Linnaeus) Pine Ladybird 20/06/2016 C C Halyzia sedecimguttata (Linnaeus) Orange Ladybird 09/08/2018 L C Harmonia axyridis (Pallas) Harlequin Ladybird 23/04/2014 C C Propylea quatuordecimpunctata (Linnaeus) 14-spot Ladybird 23/04/2014 C C Psyllobora vigintiduopunctata (Linnaeus) 22-spot Ladybird 12/06/2014 C C Family: Oedemeridae False blister beetles Oedemera lurida (Marsham) a false blister beetle 06/06/2016 L L Family: Pyrochroidae Cardinal beetles etc. Pyrochroa serraticornis (Scopoli) Red-headed Cardinal Beetle 19/05/2018 C L Family: Cerambycidae Longhorn beetles Clytus arietis (Linnaeus) Wasp Beetle 12/06/2014 C L Grammoptera ruficornis (Fabricius) a longhorn beetle 03/05/2017 C C Pogonocherus hispidus (Linnaeus) Lesser Thorn-tipped Longhorn Beetle 21/05/2017 C L Rutpela maculata (Poda) Black and Yellow Longhorn Beetle 09/07/2014 C L Tetropium gabrieli Weise Larch Longhorn Beetle 09/05/2016 L L Family: Chrysomelidae Leaf beetles Agelastica alni (Linnaeus) Alder Leaf Beetle 04/05/2017 JNCC L Cassida rubiginosa Müller Thistle Tortoise Beetle 06/06/2016 C C Oulema melanopus (Linnaeus) [s. l.] Cereal Leaf Beetle 05/05/2016 C C Psylliodes chrysocephala (Linnaeus) Cabbage Stem Flea Beetle 01/10/2018 C C Family: Apionidae Weevils Oxystoma cerdo (Gerstäcker) a weevil 08/07/2014 JNCC L Protapion apricans (Herbst) a clover seed weevil 08/07/2014 C C Protapion fulvipes (Geoffroy) a weevil 08/07/2014 C C Family: Curculionidae Weevils Phyllobius argentatus (Linnaeus) a weevil 03/05/2017 C L Phyllobius maculicornis Germar a weevil 21/05/2015 C C Phyllobius pyri (Linnaeus) Common Leaf Weevil 05/05/2016 C C Phyllobius virideaeris (Laicharting) a weevil 03/06/2015 C RS Polydrusus formosus (Mayer) a weevil 01/07/2015 JNCC U Sitona lineatus (Linnaeus) Pea and Bean Weevil 08/07/2014 C C Strophosoma melanogrammum (Forster) Nut Leaf Weevil 02/06/2017 C C Tychius picirostris (Fabricius) a clover seed weevil 08/07/2014 C L Order: Hymenoptera Sawflies, bees & ichneumons etc. Suborder: Symphyta Sawflies Family: Argidae Sawflies Arge cyanocrocea (Förster) Bramble Sawfly 02/06/2016 L L 29 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Arge ustulata (Linnaeus) a sawfly 03/06/2015 C C Family: Cephidae Sawflies Cephus spinipes (Panzer) a sawfly 03/06/2015 C U Family: Cimbicidae Club-horned sawflies Cimbex connatus (Schrank) a club-horned sawfly 10/09/2017 L U Family: Tenthredinidae Sawflies Ametastegia tenera (Fallén) a sawfly 09/04/2017 L U Athalia ancilla Serville a sawfly 21/05/2015 L U Athalia bicolor Serville a sawfly 02/07/2016 L U Athalia cordata Lepeletier a sawfly 03/06/2015 C L Athalia rosae (Linnaeus) Turnip Sawfly 07/08/2017 L L Craesus septentrionalis (Linnaeus) a sawfly 23/08/2014 L U Dineura stilata (Klug) a sawfly 03/06/2015 C U Dolerus aeneus Hartig a sawfly 21/05/2015 C C Dolerus ferrugatus Serville a sawfly 04/05/2017 L U Dolerus gonager (Fabricius) a sawfly 21/05/2015 C C Dolerus madidus (Klug) a sawfly 06/06/2016 L L Dolerus nigratus (Müller) a sawfly 08/04/2015 C C Dolerus picipes (Klug) a sawfly 21/05/2015 C C Dolerus sanguinicollis (Klug) a sawfly 08/04/2015 C C Dolerus yokonensis Norton a sawfly 09/05/2016 L U Eriocampa ovata (Linnaeus) Woolly Alder Sawfly 27/08/2016 C U Eutomostethus luteiventris (Klug) a sawfly 06/05/2018 C U Fenusa dohrnii (Tischbein) a sawfly 23/08/2014 C U Pachynematus extensicornis (Norton) a sawfly 01/06/2017 L U Pachynematus kirbyi (Dahlbom) a sawfly 17/08/2015 L U Pachyprotasis antennata (Klug) a sawfly 02/06/2017 L U Pachyprotasis rapae (Linnaeus) a sawfly 03/06/2015 C C Platycampus luridiventris (Fallén) a sawfly 03/05/2017 L U Rhogogaster chlorosoma (Benson) a sawfly 02/06/2017 C U Rhogogaster viridis (Linnaeus) Green Sawfly 22/06/2016 C C Selandria melanosterna (Serville) a sawfly 01/07/2015 L U Selandria serva (Fabricius) a sawfly 17/08/2015 C C Tenthredo arcuata Förster a sawfly 21/05/2015 C C Tenthredo atra Linnaeus a sawfly 03/06/2015 C C Tenthredo brevicornis (Konow) a sawfly 21/05/2015 C C Tenthredo celtica Benson a sawfly 03/06/2015 C C Tenthredo mesomela Linnaeus a sawfly 01/06/2017 C C Tenthredo notha Klug a sawfly 07/09/2014 C C Tenthredo velox Fabricius a sawfly 21/07/2016 L U Tenthredopsis nassata (Linnaeus) a sawfly 11/04/2016 C C Suborder: Apocrita Bees, wasps & ichneumons etc. Superfamily: Aculeata Bees, wasps & ants Family: Chrysididae Ruby-tailed wasps Chrysis corusca Valkeila a ruby-tailed wasp 07/07/2016 U U Chrysis ignita Linnaeus a ruby-tailed wasp 02/07/2016 C L Family: Formicidae Ants Lasius niger (Linnaeus) [s. s.] Black Garden Ant 08/04/2015 C C 30 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Myrmica rubra (Linnaeus) Common Red Ant 23/04/2014 C C Family: Mutillidae Velvet ants etc. Myrmosa atra Panzer Black-headed Velvet Ant 17/08/2015 C L Family: Pompilidae Spider-hunting wasps Dipogon variegatus (Linnaeus) a spider-hunting wasp 01/07/2017 L L Priocnemis perturbator (Harris) a spider-hunting wasp 15/05/2014 L L Family: Tiphiidae Solitary wasps Tiphia minuta Vander Linden a solitary wasp 07/07/2016 JNCC RS Family: Eumenidae Potter wasps Ancistrocerus gazella (Panzer) a potter wasp 01/07/2015 C L Ancistrocerus trifasciatus (Müller) a potter wasp 07/09/2017 C L Symmorphus bifasciatus (Linnaeus) a potter wasp 03/08/2017 L L Family: Vespidae Social wasps Dolichovespula norwegica (Fabricius) Norwegian Wasp 07/07/2016 C C Dolichovespula saxonica (Fabricius) Saxon Wasp 03/06/2015 JNCC L Dolichovespula sylvestris (Scopoli) Tree Wasp 05/08/2014 C C Vespa crabro Linnaeus Hornet 15/05/2014 L L Vespula germanica (Fabricius) German Wasp 23/04/2014 C C Vespula vulgaris (Linnaeus) Common Wasp 23/04/2014 C C Family: Crabronidae Solitary wasps Crabro cribrarius (Linnaeus) a solitary wasp 01/07/2015 C C Crossocerus cetratus (Shuckard) a solitary wasp 06/07/2017 L L Crossocerus megacephalus (Rossi) a solitary wasp 01/09/2017 C C Crossocerus quadrimaculatus (Fabricius) a solitary wasp 17/08/2015 C C Ectemnius continuus (Fabricius) a solitary wasp 17/08/2015 C C Mellinus arvensis (Linnaeus ) Field Digger Wasp 17/08/2015 C C Pemphredon inornata Say a solitary wasp 03/08/2017 C C Pemphredon lugubris (Fabricius) a solitary wasp 06/07/2017 C C Psenulus pallipes (Panzer) a solitary wasp 12/06/2014 C C Trypoxylon attenuatum Smith a solitary wasp 01/06/2017 C C Family: Colletidae Mining bees Colletes daviesanus Smith Davies’ Mining Bee 08/07/2014 L L Hylaeus communis Nylander Common Yellow-face Bee 01/06/2017 C C Family: Andrenidae Mining bees Andrena bicolor Fabricius Gwynne’s Mining Bee 23/04/2014 C C Andrena denticulata (Kirby) Grey-banded Mining Bee 05/08/2014 L L Andrena fulva (Müller) Tawny Mining Bee 23/04/2014 C C Andrena haemorrhoa (Fabricius) Early Mining Bee 23/04/2014 C C Andrena helvola (Linnaeus) Coppice Mining Bee 03/05/2017 L RS Andrena minutula (Kirby) Common Mini-miner 08/04/2015 C L Andrena nigroaenea (Kirby) Buffish Mining Bee 15/05/2014 C C Andrena scotica Perkins Chocolate Mining Bee 15/05/2014 C C Andrena subopaca Nylander Impunctate Mini-miner 23/04/2014 C C Andrena synadelpha Perkins Broad-margined Mining Bee 06/05/2018 L RS Andrena wilkella (Kirby) Wilke’s Mining Bee 03/06/2015 L L Family: Halictidae Mining bees etc. Halictus rubicundus (Christ) Orange-legged Furrow Bee 12/06/2014 L L Halictus tumulorum (Linnaeus) Bronze Furrow Bee 09/04/2017 C C 31 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Lasioglossum albipes (Fabricius) Bloomed Furrow Bee 12/06/2014 C C Lasioglossum calceatum (Scopoli) Common Furrow Bee 07/09/2014 C C Lasioglossum leucopus (Kirby) White-footed Furrow Bee 09/05/2016 L L Lasioglossum leucozonium (Schrank) White-zoned Furrow Bee 12/06/2014 C L Lasioglossum minutissimum (Kirby) Least Furrow Bee 12/06/2014 L RS Lasioglossum rufitarse (Zetterstedt) Rufous-footed Furrow Bee 23/04/2014 L L Lasioglossum villosulum (Kirby) Shaggy Furrow Bee 07/09/2014 C L Sphecodes ephippius (Linnaeus) Bare-saddled Blood Bee 23/04/2014 L L Sphecodes gibbus (Linnaeus) Dark-winged Blood Bee 05/08/2014 L L Family: Megachilidae Mason bees etc. Anthidium manicatum (Linnaeus) Wool Carder Bee 05/08/2014 L L Coelioxys inermis (Kirby) Shiny-vented Sharp-tail Bee 06/07/2017 L RS Coelioxys rufescens Lepeletier & Audinet-Serville Rufescent Sharp-tail Bee 01/07/2015 L L Megachile centuncularis (Linnaeus) Patchwork Leaf-cutter Bee 07/07/2016 C L Megachile ligniseca (Kirby) Wood-carving Leaf-cutter Bee 02/07/2016 C L Megachile willughbiella (Kirby) Willughby’s Leaf-cutter Bee 07/08/2017 C L Osmia bicornis (Linnaeus) Red Mason Bee 12/06/2014 C C Family: Anthophoridae Nomad bees Nomada flava Panzer Flavous Nomad Bee 09/05/2016 C L Nomada fulvicornis Fabricius Orange-horned Nomad Bee 09/04/2017 JNCC RS Nomada goodeniana (Kirby) Gooden’s Nomad Bee 21/05/2015 C C Nomada marshamella (Kirby) Marsham’s Nomad Bee 12/06/2014 C C Nomada panzeri Lepeletier Panzer’s Nomad Bee 03/05/2017 C C Family: Apidae Bumblebees etc. Apis mellifera Linnaeus Honey Bee 12/06/2014 C C Bombus bohemicus (Seidl) Gypsy Cuckoo-bee 23/04/2014 C C Bombus campestris (Panzer) Field Cuckoo-bee 15/05/2014 C C Bombus hortorum (Linnaeus) Small Garden Bumblebee 23/04/2014 C C Bombus hypnorum (Linnaeus) Tree Bumblebee 23/04/2014 C C Bombus lapidarius (Linnaeus) Red-tailed Bumblebee 23/04/2014 C C Bombus lucorum (Linnaeus) White-tailed Bumblebee 23/04/2014 C C Bombus pascuorum (Scopoli) Common Carder-bee 23/04/2014 C C Bombus pratorum (Linnaeus) Early Bumblebee 12/06/2014 C C Bombus rupestris (Fabricius) Hill Cuckoo-bee 23/04/2014 JNCC C Bombus sylvestris (Lepeletier) Forest Cuckoo-bee 23/04/2014 C C Bombus terrestris (Linnaeus) Buff-tailed Bumblebee 26/03/2014 C C Bombus vestalis (Geoffroy) Vestal Cuckoo-bee 23/04/2014 C C Superfamily: Parasitica Ichneumon wasps etc. Family: Cynipidae Gall Wasps Andricus kollari (Hartig) Oak Marble Gall Wasp 08/07/2014 C C Cynips divisa Hartig Red Pea Gall Wasp 24/07/2016 C C Family: Braconidae Braconid wasps Meteorus pendulus (Müller) a braconid wasp 03/05/2017 U U Microplitis mediator (Haliday) a braconid wasp 03/05/2017 U U Synelix semirugosa (Haliday) a braconid wasp 04/05/2017 U U Therophilus clausthalianus (Ratzeburg) a braconid wasp 02/06/2017 U U Zele deceptor (Wesmael) a braconid wasp 01/06/2017 U U

32 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Family: Ichneumonidae Ichneumon wasps Achaius oratorius (Fabricius) an ichneumon wasp 01/09/2016 U U Alomya debellator (Fabricius) an ichneumon wasp 21/05/2015 U U Atractodes croceicornis Haliday an ichneumon wasp 01/06/2017 U U Azelus erythropalpus (Gmelin) an ichneumon wasp 03/05/2017 U U Banchus falcatorius (Fabricius) an ichneumon wasp 08/07/2014 U U Chorinaeus brevicalcar Thomson an ichneumon wasp 03/06/2018 U U Cosmoconus meridionator Aubert an ichneumon wasp 01/09/2017 U U Ctenichneumon panzeri (Wesmael) an ichneumon wasp 08/07/2014 U U Cylloceria melancholica (Gravenhorst) an ichneumon wasp 17/08/2015 U U Diplazon laetatorius (Fabricius) an ichneumon wasp 02/09/2018 U U Dusona terebrator (Förster) an ichneumon wasp 21/05/2015 U U Eurylabus torvus Wesmael an ichneumon wasp 01/09/2016 U U Glyphicnemis profligator (Fabricius) an ichneumon wasp 01/07/2015 U U Glypta nigrotrochanterata Strobl an ichneumon wasp 05/07/2017 U U Glypta vulnerata Gravenhorst an ichneumon wasp 05/07/2017 U U Hadrodactylus fugax (Gravenhorst) an ichneumon wasp 04/05/2017 U U Ichneumon sarcitorius Linnaeus an ichneumon wasp 02/07/2016 C C Ichneumon stramentor Rasnitsyn an ichneumon wasp 28/01/2018 U U Itoplectis maculator (Fabricius) an ichneumon wasp 03/06/2018 U U Lagarotis debitor (Thunberg) an ichneumon wasp 01/09/2017 U U Lissonota biguttata Holmgren an ichneumon wasp 02/07/2017 U U Lissonota clypeator (Gravenhorst) an ichneumon wasp 01/06/2017 U U Lissonota coracina (Gmelin) an ichneumon wasp 03/08/2017 U U Macrocentrus pallipes (Nees) an ichneumon wasp 01/09/2017 U U Mesoleptus laevigatus (Gravenhorst) an ichneumon wasp 03/08/2017 U U Mesoleptus vigilatorius (Förster) an ichneumon wasp 01/06/2017 U U Oiorhinus pallipalpis Wesmael an ichneumon wasp 09/04/2017 U U Polyblastus varitarsus (Gravenhorst) an ichneumon wasp 02/06/2016 U U Scambus buolianae (Hartig) an ichneumon wasp 01/09/2016 U U Scambus inanis (Schrank) an ichneumon wasp 03/08/2017 U U Scambus signatus (Pfeffer) an ichneumon wasp 02/09/2018 U U Sussaba flavipes (Lucas) an ichneumon wasp 03/05/2017 U U Sussaba pulchella (Holmgren) an ichneumon wasp 03/05/2017 U U Syrphoctonus tarsatorius (Panzer) an ichneumon wasp 17/08/2015 U U Trychosis legator (Thunberg) an ichneumon wasp 01/07/2015 U U Tryphon latrator (Fabricius) an ichneumon wasp 02/06/2017 U U Tryphon rutilator (Linnaeus) an ichneumon wasp 01/06/2017 U U Tryphon trochanteratus Holmgren an ichneumon wasp 06/06/2016 U U Tymmophorus obscuripes (Holmgren) an ichneumon wasp 02/07/2016 U U Order: Diptera True flies Suborder: Nematocera Thread-horns Family: Tipulidae Crane-flies Nephrotoma appendiculata (Pierre) a crane-fly 15/05/2014 C C Nephrotoma flavescens (Linnaeus) a crane-fly 12/06/2014 C C Nephrotoma scurra (Meigen) a crane-fly 12/06/2014 L L Tipula couckei Tonnoir a crane-fly 04/08/2016 L L Tipula fascipennis Meigen a crane-fly 07/07/2016 C C 33 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Tipula lateralis Meigen a crane-fly 07/09/2014 C C Tipula lunata Linnaeus a crane-fly 02/06/2016 C C Tipula oleracea Linnaeus a crane-fly 15/05/2014 C C Tipula pagana Meigen a crane-fly 01/11/2018 C C Tipula paludosa Meigen a crane-fly 07/09/2014 C C Tipula varipennis Meigen a crane-fly 06/05/2018 C C Tipula vernalis Meigen a crane-fly 15/05/2014 C C Family: Pediciidae Crane-flies Tricyphona immaculata (Meigen) a crane-fly 03/05/2017 C C Family: Limoniidae Crane-flies Dicranomyia modesta (Meigen) a crane-fly 02/06/2016 C C Epiphragma occelare (Linnaeus) a crane-fly 15/05/2014 L L Limonia nubeculosa Meigen a crane-fly 09/04/2017 C C Limonia phragmitidis (Schrank) a crane-fly 21/05/2015 C C Metalimnobia quadrinotata (Meigen) a crane-fly 06/05/2018 C L Ormosia hederae (Curtis) a crane-fly 03/05/2017 C C Symplecta hybrida (Meigen) a crane-fly 08/07/2014 L C Symplecta stictica (Meigen) a crane-fly 02/06/2016 C C Family: Bibionidae March flies Bibio johannis (Linnaeus) a March fly 23/04/2014 C C Bibio leucopterus (Meigen) a March fly 21/05/2015 C C Bibio marci (Linnaeus) St. Mark’s Fly 05/05/2016 C C Bibio nigriventris Haliday a March fly 21/05/2015 C C Dilophus febrilis (Linnaeus) Fever Fly 23/04/2014 C C Dilophus femoratus Meigen a March fly 11/04/2016 C C Family: Sciaridae Dark-winged fungus gnats Sciara hemerobioides (Scopoli) a dark-winged fungus gnat 17/08/2015 C C Family: Trichoceridae Winter gnats Trichocera annulata Meigen a winter gnat 01/11/2018 C C Trichocera hiemalis (De Geer) a winter gnat 01/11/2018 L C Trichocera regelationis (Linnaeus) a winter gnat 01/10/2018 C C Trichocera saltator (Harris) a winter gnat 09/04/2017 C C Family: Anisopodidae Wood gnats Sylvicola fenestralis (Scopoli) a wood gnat 07/09/2014 C C Sylvicola punctatus (Fabricius) a wood gnat 09/08/2014 C C Family: Culicidae Mosquitoes Culiseta annulata (Schrank) a mosquito 01/09/2017 C C Ochlerotatus annulipes (Meigen) a mosquito 03/06/2018 C C Ochlerotatus punctor (Kirby) a mosquito 17/08/2015 C C Ochlerotatus rusticus (Rossi) a mosquito 06/05/2018 C C Suborder: Brachycera Short-horns Family: Rhagionidae Snipe-flies Chrysopilus cristatus (Fabricius) Black Snipe-fly 12/06/2014 C C Rhagio lineola Fabricius Small Fleck-winged Snipe-fly 08/07/2014 C C Rhagio scolopaceus (Linnaeus) Down-looker Fly 15/05/2014 C C Rhagio tringarius (Linnaeus) Marsh Snipe-fly 08/07/2014 C C Family: Tabanidae Horseflies Chrysops caecutiens (Linnaeus) Splayed Deerfly 01/07/2017 L L 34 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Chrysops relictus Meigen Twin-lobed Deerfly 08/07/2014 L L Haematopota pluvialis (Linnaeus) Notch-horned Cleg 12/06/2014 C C Hybomitra bimaculata (Macquart) Hairy-legged Horsefly 03/06/2018 L RS Family: Stratiomyidae Soldier-flies Beris chalybata (Forster) Murky-legged Black Legionnaire 23/04/2014 C C Beris vallata (Forster) Common Orange Legionnaire 09/07/2014 C C Chloromyia formosa (Scopoli) Broad Centurion 12/06/2014 C C Chorisops tibialis (Meigen) Dull Four-spined Legionnaire 09/07/2014 L L Microchrysa polita (Linnaeus) Black-horned Gem 21/05/2015 C C Pachygaster leachii Stephens Yellow-legged Black 05/07/2017 L L Sargus bipunctatus (Scopoli) Twin-spot Centurion 26/09/2015 L L Family: Bombyliidae Bee-flies Bombylius major Linnaeus Dark-edged Bee-fly 23/04/2014 C C Family: Therevidae Stiletto-flies Thereva nobilitata (Fabricius) Common Stiletto-fly 08/07/2014 C C Family: Asilidae Robber-flies Dioctria atricapilla Meigen Violet Black-legged Robber-fly 12/06/2014 L L Dioctria rufipes (De Geer) Common Red-legged Robber-fly 15/05/2014 L C Leptogaster cylindrica (De Geer) Striped Slender Robber-fly 08/07/2014 C C Leptogaster guttiventris Zetterstedt Dashed Slender Robber-fly 03/06/2015 L L Family: Hybotidae Dance flies Bicellaria vana Collin a dance fly 15/05/2014 C C Hybos culiciformis (Fabricius) a dance fly 21/07/2016 C C Ocydromia glabricula (Fallén) a dance fly 03/06/2018 C C Platypalpus agilis (Meigen) a dance fly 03/06/2015 C C Platypalpus albicornis (Zetterstedt) a dance fly 03/05/2017 L L Platypalpus articulatus Macquart a dance fly 08/07/2014 JNCC U Platypalpus longicornis (Meigen) a dance fly 09/05/2016 C C Platypalpus minutus (Meigen) [s. s.] a dance fly 01/07/2015 C C Platypalpus pallidiventris (Meigen) a dance fly 01/06/2017 C C Trichinomyia flavipes (Meigen) a dance fly 01/11/2018 C C Family: Empididae Dance flies Dolichocephala guttata (Haliday) a dance fly 23/04/2014 C C Dolichocephala irrorata (Fallén) a dance fly 03/05/2017 C C Empis albinervis Meigen a dance fly 09/07/2014 L C Empis caudatula Loew a dance fly 01/06/2017 L C Empis chioptera Meigen a dance fly 09/05/2016 C C Empis grisea Fallén a dance fly 03/06/2018 C C Empis livida Linnaeus a dance fly 12/06/2014 C C Empis nigripes Fabricius a dance fly 15/05/2014 C C Empis nuntia Meigen a dance fly 15/05/2014 C C Empis praevia Collin a dance fly 02/06/2016 L C Empis stercorea Linnaeus a dance fly 03/06/2015 C C Empis tessellata Fabricius a dance fly 15/05/2014 C C Empis trigramma Wiedemann a dance fly 21/05/2015 C C Hilara albipennis von Roser a dance fly 03/05/2017 L L Hilara brevistyla Collin a dance fly 03/05/2017 U C Hilara chorica (Fallén) a dance fly 03/05/2017 C C 35 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Hilara cornicula Loew a dance fly 21/05/2015 L C Hilara fulvibarba Strobl a dance fly 02/06/2017 L U Hilara interstincta (Fallén) a dance fly 03/06/2015 C C Hilara litorea (Fallén) a dance fly 07/07/2016 C C Hilara manicata Meigen a dance fly 08/07/2014 C C Hilara maura (Fabricius) a dance fly 15/05/2014 C C Hilara nigrina (Fallén) a dance fly 03/06/2018 L L Hilara quadrifasciata Chvála a dance fly 21/05/2015 C C Rhamphomyia albipennis (Fallén) a dance fly 04/05/2017 L L Rhamphomyia albohirta Collin a dance fly 05/05/2016 C C Rhamphomyia nigripennis (Fabricius) a dance fly 03/06/2018 C C Rhamphomyia subcinerascens Collin a dance fly 09/05/2016 L L Rhamphomyia sulcata (Meigen) a dance fly 23/04/2014 C C Rhamphomyia tarsata Meigen a dance fly 21/05/2015 C C Family: Dolichopodidae Long-legged flies Campsicnemus curvipes (Fallén) a long-legged fly 02/07/2016 C C Chrysotimus molliculus (Fallén) a long-legged fly 14/08/2016 L U Chrysotus gramineus (Fallén) a long-legged fly 17/08/2015 C C Chrysotus neglectus (Wiedemann) a long-legged fly 12/06/2014 L C Dolichopus claviger Stannius a long-legged fly 21/05/2015 C C Dolichopus festivus Haliday a long-legged fly 09/07/2014 C C Dolichopus longicornis Stannius a long-legged fly 12/06/2014 L L Dolichopus plumipes (Scopoli) a long-legged fly 12/06/2014 C C Dolichopus popularis Wiedemann a long-legged fly 02/06/2017 C C Dolichopus subpennatus d’Assis-Fonseca a long-legged fly 01/07/2015 C L Dolichopus trivialis Haliday a long-legged fly 08/07/2014 C C Dolichopus ungulatus (Linnaeus) a long-legged fly 08/07/2014 C C Medetera saxatilis Collin a long-legged fly 05/08/2014 L L Medetera truncorum Meigen a long-legged fly 02/07/2016 C C Rhaphium caliginosum Meigen a long-legged fly 03/07/2017 C C Sympycnus desoutteri Parent a long-legged fly 09/08/2014 C C Syntormon pallipes (Fabricius) a long-legged fly 15/05/2014 C C Xanthochlorus galbanus Chandler & Negrobov a long-legged fly 05/07/2017 L L Xanthochlorus tenellus (Wiedemann) a long-legged fly 01/07/2015 L L Family: Platypezidae Flat-footed flies Protoclythia modesta (Zetterstedt) a flat-footed fly 01/10/2018 U U Family: Phoridae Scuttle flies Borophaga incrassata (Meigen) a scuttle fly 04/08/2018 C U Family: Lonchopteridae Pointed-winged flies Lonchoptera bifurcata (Fallén) a pointed-winged fly 05/08/2014 C C Lonchoptera lutea Panzer a pointed-winged fly 23/04/2014 C C Family: Syrphidae Hoverflies Baccha elongata (Fabricius) a hoverfly 02/09/2018 C C Cheilosia albitarsis (Meigen) [s. s.] a hoverfly 21/05/2015 C C Cheilosia bergenstammi Becker a hoverfly 23/04/2014 L C Cheilosia illustrata (Harris) a hoverfly 12/06/2014 C C Cheilosia impressa Loew a hoverfly 03/08/2017 L L Cheilosia lasiopa Kowarz a hoverfly 03/05/2017 L L 36 FULL INVERTEBRATE LIST (2014 TO 2018)

VERNACULAR FIRST NAT. REG. SCIENTIFIC NAME NAME RECORD STATUS STATUS Cheilosia latifrons (Zetterstedt) a hoverfly 17/08/2015 L L Cheilosia pagana (Meigen) a hoverfly 23/04/2014 C C Cheilosia proxima (Zetterstedt) a hoverfly 03/08/2017 L C Cheilosia ranunculi Doczkal a hoverfly 21/05/2015 U U Cheilosia variabilis (Panzer) a hoverfly 15/05/2014 C C Cheilosia vernalis (Fallén) a hoverfly 23/04/2014 C C Chrysogaster solstitialis (Fallén) a hoverfly 05/08/2014 C C Chrysotoxum bicinctum (Linnaeus) a hoverfly 05/07/2017 L L Chrysotoxum festivum (Linnaeus) a hoverfly 12/06/2014 L L Dasysyrphus albostriatus (Fallén) a hoverfly 29/07/2014 C C Dasysyrphus pinastri (De Geer) a hoverfly 03/06/2015 L L Dasysyrphus tricinctus (Fallén) a hoverfly 17/08/2015 C C Epistrophe eligans (Harris) a hoverfly 23/04/2014 C C Epistrophe grossulariae (Meigen) a hoverfly 05/08/2014 L C Episyrphus balteatus (De Geer) a hoverfly 12/06/2014 C C Eristalis arbustorum (Linnaeus) a hoverfly 12/06/2014 C C Eristalis interruptus (Poda) a hoverfly 09/07/2014 C C Eristalis intricarius (Linnaeus) a hoverfly 08/07/2014 C C Eristalis pertinax (Scopoli) a hoverfly 23/04/2014 C C Eristalis tenax (Linnaeus) Drone-fly 23/04/2014 C C Eumerus funeralis Meigen a lesser bulb fly 02/06/2017 C L Eumerus strigatus (Fallén) a lesser bulb fly 04/08/2018 C L Eupeodes corollae (Fabricius) a hoverfly 02/07/2016 C C Eupeodes luniger (Meigen) a hoverfly 12/06/2014 C C Helophilus hybridus Loew a hoverfly 08/07/2014 L L Helophilus pendulus (Linnaeus) a hoverfly 12/06/2014 C C Helophilus trivittatus (Fabricius) a hoverfly 08/07/2014 L L Leucozona lucorum (Linnaeus) a hoverfly 23/04/2014 C C Melangyna lasiophthalma (Zetterstedt) a hoverfly 22/04/2018 L C Melanostoma mellinum (Linnaeus) a hoverfly 23/04/2014 C C Melanostoma scalare (Fabricius) a hoverfly 23/04/2014 C C Meliscaeva auricollis (Meigen) a hoverfly 08/09/2015 C C Meliscaeva cinctella (Zetterstedt) a hoverfly 12/06/2014 C C Merodon equestris (Fabricius) Greater Bulb Fly 15/05/2014 C C Myathropa florea (Linnaeus) a hoverfly 04/08/2018 C C Paragus haemorrhous Meigen a hoverfly 07/08/2017 L L Pipiza noctiluca (Linnaeus) a hoverfly 03/06/2015 C C Pipizella viduata (Linnaeus) a hoverfly 21/05/2015 C L Platycheirus albimanus (Fabricius) a hoverfly 23/04/2014 C C Platycheirus angustatus (Zetterstedt) a hoverfly 21/05/2015 C C Platycheirus clypeatus (Meigen) a hoverfly 08/07/2014 C C Platycheirus manicatus (Meigen) a hoverfly 23/04/2014 C C Platycheirus occultus Goeldlin de Tiefenau, Maibach & Speight a hoverfly 01/07/2015 L U Platycheirus rosarum (Fabricius) a hoverfly 05/07/2017 L L Platycheirus scutatus (Meigen) [s. s.] a hoverfly 01/09/2016 C C Platycheirus splendidus Rotheray a hoverfly 22/04/2018 U U Rhingia campestris Meigen a hoverfly 08/07/2014 C C Scaeva pyrastri (Linnaeus) a hoverfly 02/07/2017 C C 37 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Sphaerophoria interrupta (Fabricius) a hoverfly 15/05/2014 L L Sphaerophoria scripta (Linnaeus) a hoverfly 08/07/2014 C C Syritta pipiens (Linnaeus) a hoverfly 12/06/2014 C C Syrphus ribesii (Linnaeus) a hoverfly 23/04/2014 C C Syrphus torvus Osten Sacken a hoverfly 12/06/2014 C C Syrphus vitripennis Meigen a hoverfly 23/04/2014 C C Volucella bombylans (Linnaeus) a hoverfly 09/07/2014 C C Volucella pellucens (Linnaeus) a hoverfly 12/06/2014 C C Xylota segnis (Linnaeus) a hoverfly 03/06/2015 C C Family: Pipunculidae Big-headed flies Eudorylas inferus Collin a big-headed fly 07/07/2016 JNCC U Eudorylas obliquus Coe a big-headed fly 01/07/2015 JNCC U Eudorylas subterminatus Collin a big-headed fly 09/05/2016 C U Pipunculus campestris Latreille a big-headed fly 01/07/2015 C C Pipunculus zugmayeriae Kowarz a big-headed fly 02/06/2016 JNCC L Family: Micropezidae Stilt-legged flies Micropeza corrigiolata (Linnaeus) a stilt-legged fly 06/06/2016 L L Neria ephippium (Fabricius) a stilt-legged fly 01/07/2015 L L Family: Psilidae Rust flies Chamaepsila nigricornis (Meigen) a rust fly 03/05/2017 U C Imantimyia albiseta (Schrank) a rust fly 14/08/2016 C L Psila merdaria Collin a rust fly 06/05/2018 U C Family: Conopidae Thick-headed flies Conops flavipes Linnaeus a thick-headed fly 09/07/2014 C C Conops quadrifasciatus De Geer a thick-headed fly 05/08/2014 L C Physocephala rufipes (Fabricius) a thick-headed fly 05/08/2014 L L Sicus ferrugineus (Linnaeus) a thick-headed fly 12/06/2014 L C Family: Platystomatidae Picture-winged flies Rivellia syngenesiae (Fabricius) a picture-winged fly 12/06/2014 L L Family: Lonchaeidae Lance flies Lonchaea corusca Czerny a lance fly 01/10/2018 JNCC U Family: Tephritidae Picture-winged flies Anomoia purmunda (Harris) a picture-winged fly 17/08/2015 L C Campiglossa misella (Loew) a picture-winged fly 06/06/2016 C C Chaetorellia jaceae (Robineau-Desvoidy) a picture-winged fly 06/06/2016 L L Chaetostomella cylindrica (Robineau-Desvoidy) a picture-winged fly 01/06/2017 C C Tephritis formosa (Loew) a picture-winged fly 12/06/2014 L L Tephritis hyoscyami (Linnaeus) a picture-winged fly 06/05/2018 L L Tephritis neesii (Meigen) a picture-winged fly 23/04/2014 L L Tephritis vespertina (Loew) a picture-winged fly 15/05/2014 C C Terellia serratulae (Linnaeus) a picture-winged fly 08/07/2014 C C Terellia tussilaginis (Fabricius) a picture-winged fly 03/07/2017 C C Urophora jaceana (Hering) a picture-winged fly 06/06/2016 C C Urophora quadrifasciata (Meigen) a picture-winged fly 01/06/2017 L L Urophora stylata (Fabricius) a picture-winged fly 01/06/2017 C C Family: Lauxaniidae Lauxaniid flies Calliopum aeneum (Fallén) a lauxaniid fly 02/09/2018 C C Meiosimyza rorida (Fallén) a lauxaniid fly 02/09/2018 C C 38 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Family: Dryomyzidae Dryomyzid flies Dryomyza anilis Fallén a dryomyzid fly 01/09/2016 C C Family: Sciomyzidae Marsh flies Coremacera marginata (Fabricius) a marsh fly 14/08/2016 L L Elgiva cucularia (Linnaeus) a marsh fly 23/04/2014 L L Ilione albiseta (Scopoli) a marsh fly 21/05/2015 C C Pherbellia cinerella (Fallén) a marsh fly 07/09/2014 C C Pherbellia griseola (Fallén) a marsh fly 01/07/2018 JNCC RS Tetanocera elata (Fabricius) a marsh fly 03/06/2015 C C Tetanocera ferruginea Fallén a marsh fly 03/05/2017 C C Tetanocera robusta Loew a marsh fly 15/05/2014 C C Family: Sepsidae Black scavenger flies Nemopoda nitidula (Fallén) a black scavenger fly 14/08/2016 C C Sepsis cynipsea (Linnaeus) a black scavenger fly 17/08/2015 C C Sepsis flavimana Meigen a black scavenger fly 09/04/2017 C C Sepsis fulgens Meigen a black scavenger fly 12/06/2014 C C Sepsis orthocnemis Frey a black scavenger fly 15/05/2014 C C Sepsis violacea Meigen a black scavenger fly 12/06/2014 C C Themira annulipes (Meigen) a black scavenger fly 15/05/2014 C C Themira leachi (Meigen) a black scavenger fly 15/05/2014 L U Family: Agromyzidae Leaf-mining flies Agromyza alnivora Spencer a leaf-mining fly 09/08/2014 U U Cerodontha denticornis (Panzer) a leaf-mining fly 01/06/2017 C C Liriomyza strigata (Meigen) a leaf-mining fly 19/10/2016 U U Phytomyza leucanthemi Hering a leaf-mining fly 27/08/2016 U U Family: Opomyzidae Cereal flies Geomyza balachowskyi Mesnil a cereal fly 01/09/2016 C C Geomyza tripunctata Fallén a cereal fly 08/04/2015 C C Opomyza florum (Fabricius) Yellow Cereal Fly 07/09/2014 C C Opomyza germinationis (Linnaeus) a cereal fly 07/09/2014 C C Opomyza petrei Mesnil a cereal fly 02/07/2016 C C Family: Heleomyzidae Heleomyzid flies Suillia affinis (Meigen) a heleomyzid fly 21/05/2015 L L Suillia pallida (Fallén) a heleomyzid fly 02/09/2018 C C Suillia variegata (Loew) a heleomyzid fly 05/08/2014 C C Family: Drosophilidae Vinegar flies Scaptomyza flava (Fallén) Turnip Leafminer 02/09/2018 C C Family: Scathophagidae Dung flies etc. Conisternum decipiens (Haliday) a dung fly 04/12/2018 JNCC RS Nanna armillata (Zetterstedt) a Timothy fly 09/05/2016 C U Nanna fasciata (Meigen) a Timothy fly 11/04/2016 C C Nanna flavipes (Fallén) a Timothy fly 06/06/2016 L U Nanna inermis (Becker) a Timothy fly 09/05/2016 L U Norellia spinipes (Meigen) a daffodil leaf-mining fly 11/04/2016 JNCC U Norellisoma spinimanum (Fallén) a dung fly 09/05/2016 C C Scathophaga furcata (Say) a dung fly 01/06/2017 C C Scathophaga stercorea (Linnaeus) Yellow Dung Fly 23/04/2014 C C

39 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Family: Anthomyiidae Root-maggot flies etc. Anthomyia liturata (Robineau-Desvoidy) an anthomyiid fly 01/10/2018 C C Anthomyia procellaris Rondani an anthomyiid fly 01/09/2017 C C Botanophila brunneilinea (Zetterstedt) an anthomyiid fly 02/07/2017 U U Botanophila fugax (Meigen) an anthomyiid fly 22/04/2018 C C Botanophila striolata (Fallén)/discreta (Meigen) an anthomyiid fly 22/04/2018 L U Delia coarctata (Fallén) an anthomyiid fly 03/07/2017 C C Delia florilega (Zetterstedt) an anthomyiid fly 03/06/2015 C C Delia platura (Meigen) Seedcorn Maggot Fly 03/06/2015 C C Delia radicum (Linnaeus) an anthomyiid fly 04/05/2017 C C Hydrophoria lancifer (Harris) an anthomyiid fly 02/06/2016 C C Hylemya urbica van der Wulp an anthomyiid fly 01/06/2017 L L Hylemya vagans (Panzer) an anthomyiid fly 07/09/2017 C C Pegomya solennis (Meigen) an anthomyiid fly 11/04/2016 L L Pegoplata aestiva (Meigen) an anthomyiid fly 07/09/2017 C C Phorbia fumigata (Meigen) Spring Fly 06/05/2018 U U Family: Fanniidae Lesser Housefly etc. Fannia mollissima (Haliday) a fanniid fly 03/05/2017 U C Fannia pallitibia (Rondani) a fanniid fly 01/10/2018 L L Fannia serena (Fallén) a fanniid fly 03/05/2017 C C Fannia sociella (Zetterstedt) a fanniid fly 21/05/2017 C C Family: Muscidae Housefly & sweat flies etc. Coenosia mollicula (Fallén) a muscid fly 03/05/2017 C C Coenosia infantula Rondani a muscid fly 02/09/2018 C U Coenosia pedella (Fallén) a muscid fly. 01/06/2017 L U Coenosia perpusilla Meigen a muscid fly 06/05/2018 L U Coenosia testacea (Robineau-Desvoidy) a muscid fly 09/05/2016 C C Coenosia tigrina (Fabricius) a muscid fly 21/05/2015 C C Eudasyphora cyanella (Meigen) a muscid fly 11/04/2016 C C Graphomya maculata (Scopoli) a muscid fly 05/08/2014 C C Hebecnema vespertina (Fallén) a muscid fly 03/05/2017 C C Helina calceata (Rondani) a muscid fly 21/07/2016 JNCC RS Helina depuncta (Fallén) a muscid fly 03/06/2015 C C Helina evecta (Harris) a muscid fly 08/04/2015 C C Helina impuncta (Fallén) a muscid fly 21/05/2015 C C Helina lasiophthalma (Macquart) a muscid fly 01/06/2017 C L Helina obscurata (Meigen) a muscid fly 02/07/2017 L L Helina quadrum (Fabricius) a muscid fly 17/08/2015 C C Helina reversio (Harris) a muscid fly 17/08/2015 C C Helina setiventris Ringdahl a muscid fly 21/07/2016 C C Helina trivittata (Zetterstedt) a muscid fly 04/08/2016 C C Hydrotaea irritans (Fallén) a sweat fly 07/07/2016 C C Hydrotaea meteorica (Linnaeus) a sweat fly 04/05/2017 C L Hydrotaea militaris (Meigen) a sweat-fly 03/06/2015 C L Hydrotaea palaestrica (Meigen) a sweat fly 21/05/2017 L RS Hydrotaea parva Meade a sweat fly 07/08/2017 JNCC RS Mesembrina meridiana Linnaeus Noon Fly 03/06/2015 C C Morellia aenescens Robineau-Desvoidy a muscid fly 03/08/2017 C C 40 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Morellia hortorum (Fallén) a muscid fly 01/06/2017 C C Morellia simplex (Loew) a muscid fly 02/06/2017 C C Musca autumnalis De Geer a muscid fly 04/08/2018 C C Muscina levida (Harris) a muscid fly 14/08/2016 C C Mydaea urbana (Meigen) a muscid fly 01/09/2016 C C Myospila meditabunda (Fabricius) a muscid fly 01/09/2016 C C Phaonia angelicae (Scopoli) a muscid fly 01/09/2016 C C Phaonia errans (Meigen) a muscid fly 12/06/2014 C C Phaonia fuscata (Fallén) a muscid fly 03/06/2015 L L Phaonia halterata (Stein) a muscid fly 03/05/2017 C L Phaonia incana (Wiedemann) a muscid fly 21/05/2015 C C Phaonia serva (Meigen) a muscid fly 03/06/2015 C C Phaonia subventa (Harris) a muscid fly 15/05/2014 C C Phaonia trimaculata (Bouché) a muscid fly 03/05/2017 C C Phaonia tuguriorum (Scopoli) a muscid fly 02/06/2016 C C Polietes lardarius (Fabricius) a muscid fly 01/10/2018 C C Thricops simplex (Wiedemann) a muscid fly 01/10/2018 U L Family: Calliphoridae Bluebottles etc. Bellardia viarum (Robineau-Desvoidy) Dark-veined Emerald-bottle 14/08/2016 C C Bellardia vulgaris (Robineau-Desvoidy) Pale-veined Emerald-bottle 23/04/2014 C C Calliphora vicina Robineau-Desvoidy Common Bluebottle 23/04/2014 C C Calliphora vomitoria (Linnaeus) Orange-bearded Bluebottle 03/06/2015 C C Cynomya mortuorum (Linnaeus) Yellow-faced Blowfly 21/05/2015 C L Lucilia ampullacea Villeneuve Streakless Greenbottle 14/08/2016 L RS Lucilia caesar (Linnaeus) Common Greenbottle 15/05/2014 C C Lucilia illustris (Meigen) Illustrious Greenbottle 12/06/2014 C C Lucilia silvarum (Meigen) Marsh Greenbottle 21/05/2015 L L Melanomya nana (Meigen) Little Black Blowfly 01/06/2017 C C Melinda viridicyanea (Robineau-Desvoidy) Dark-palped Melinda 03/05/2017 C C Pollenia angustigena Wainwright Narrow-cheeked Clusterfly 02/07/2017 C C Pollenia pediculata Macquart Tufted Clusterfly 01/06/2017 C U Pollenia rudis (Fabricius) Common Clusterfly 12/06/2014 C C Protocalliphora azurea (Fallén) Bird Blowfly 02/09/2018 U L Protophormia terraenovae (Robineau-Desvoidy) Blackbottle 03/06/2018 C L Family: Rhinophoridae Woodlouse flies Rhinophora lepida (Meigen) a woodlouse fly 04/08/2018 U U Family: Sarcophagidae Flesh flies Sarcophaga carnaria (Linnaeus) a flesh fly 03/06/2015 C C Sarcophaga crassimargo Pandellé a flesh fly 02/07/2017 C U Sarcophaga dissimilis Meigen a flesh fly 03/07/2017 U U Sarcophaga haemorrhoa Meigen a flesh fly 17/08/2015 C L Sarcophaga nigriventris Meigen a flesh fly 21/05/2017 C U Sarcophaga subvicina Rohdendorf a flesh fly 01/06/2017 C C Sarcophaga vagans Meigen a flesh fly 02/06/2016 C C Family: Tachinidae Parasitic flies Blepharomyia pagana (Meigen) a parasitic fly 08/07/2014 C RS Carcelia lucorum (Meigen) a parasitic fly 01/09/2017 C U Dexiosoma caninum (Fabricius) a parasitic fly 14/08/2016 C L 41 FULL INVERTEBRATE LIST (2014 TO 2018)

FIRST NAT. REG. SCIENTIFIC NAME VERNACULAR NAME RECORD STATUS STATUS Epicampocera succincta (Meigen) a parasitic fly 03/08/2017 C U Eriothrix rufomaculata (De Geer) a parasitic fly 05/08/2014 C C Eurithia anthophila (Robineau-Desvoidy) a parasitic fly 05/08/2014 C C Exorista rustica (Fallén) a parasitic fly 08/07/2014 C C Gonia picea (Robineau-Desvoidy) a parasitic fly 11/04/2016 L RS Gymnocheta viridis (Fallén) a parasitic fly 09/04/2017 C C Lydella stabulans (Meigen) a parasitic fly 01/06/2017 U L Lypha dubia (Fallén) a parasitic fly 21/05/2015 C L Pales pavida (Meigen) a parasitic fly 06/06/2016 C L Phasia hemiptera (Fabricius) a parasitic fly 05/08/2014 L L Phasia obesa (Fabricius) a parasitic fly 01/06/2017 L L Phryxe vulgaris (Fallén) a parasitic fly 15/05/2014 C C Phyllomya volvulus (Fabricius) a parasitic fly 05/07/2017 U U Phytomyptera cingulata (Robineau-Desvoidy) a parasitic fly 01/09/2017 U U Ramonda spathulata (Fallén) a parasitic fly 03/05/2017 C U Siphona geniculata (De Geer) a parasitic fly 15/05/2014 C C Tachina fera (Linnaeus) a parasitic fly 07/09/2014 C C Thelaira nigrina (Fallén) a parasitic fly 07/07/2016 C L Thelaira solivaga (Harris) a parasitic fly 02/06/2016 C RS Trixa conspersa (Harris) a parasitic fly 08/09/2015 C L Zophomyia temula (Scopoli) a parasitic fly 07/07/2016 JNCC RS

42 INVERTEBRATE LIST FOR 2018

This list is an inventory of all invertebrates thus far identified from surveys carried out at Three Hagges Wood-meadow during 2018.

For simplification and ease of reference, genera and species are arranged in alphabetical order and subgenera are omitted. Current nomenclature is accepted, except in problematic cases. The appendages [s. l.] and [s. s.] indicate either a precise species [sensu stricto] or a species-complex [sensu lato]: var. = variety.

355 species were recorded during 2018. The list contains a few invertebrates that could not be confidently identified beyond genus-level.

National Status Codes Numbers of Individuals Codes Determiner Codes C = Common A = 1 AG = Andrew Grayson L = Local B = 2 to 5 GSO = Geoff Oxford U = Unknown C = 6 to 24 PJB = Pat & Jim Bone JNCC = JNCC-listed D = 25 to 99 RHAB = Richard Baker SPIE = SPIE-listed E = 100 to 499 RJ = Reijo Jussila n/a = not applicable F = 500 to 1,000 WAE = Bill Ely G = over 1,000

* = an addition or amendment to previous invertebrate lists for Three Hagges Wood-meadow

NAT. NOS. SCIENTIFIC NAME VERNACULAR NAME DET. STATUS CODE Class: Gastropoda Slugs & snails etc. Family: Arionidae Terrestrial slugs Arion ater (Linnaeus) [s. l.] Black Slug C D AG Family: Helicidae Terrestrial snails Cepaea hortensis (Müller) White-lipped Snail C B AG Cepaea nemoralis (Linnaeus) Brown-lipped Snail C B AG Cornu aspersum (Müller) Garden Snail C F AG Family: Lymnaeidae Freshwater snails Lymnaea stagnalis (Linnaeus) Great Pond Snail C E AG Order: Araneae Spiders Family: Araneidae Orb-weaver spiders etc. Araneus diadematus Clerck Garden Spider C B AG, GSO Araneus marmoreus Clerck var. pyramidatus Clerck * Marled Orb-weaver Spider L A GSO Araneus quadratus Clerck * Four-spot Orb-weaver Spider C A GSO Larinioides cornutus (Clerck) Furrow Spider C A GSO Family: Philodromidae Slender Crab Spiders Tibellus species * a slender crab spider n/a A GSO Family: Pisauridae Nursery web spiders Pisaura mirabilis (Clerck) Nursery Web Spider C A GSO Family: Tetragnathidae Long-jawed orb-weavers Metellina segmentata (Clerck) a long-jawed orb-weaver C A GSO Family: Theridiidae Cobweb spiders Enoplognatha latimana Hippa & Oksala * Scarce Candy-striped spider L A GSO Order: Glomerida Pill millipedes Family: Glomeridae Pill millipedes Glomeris marginata (Villers) Pill Millipede C B AG

43 INVERTEBRATE LIST FOR 2018

NAT. NOS. SCIENTIFIC NAME VERNACULAR NAME DET. STATUS CODE Order: Polydesmida Flat-backed millipedes etc. Family: Polydesmidae Flat-backed millipedes Polydesmus angustus Latzel * Common Flat-backed Millipede C A AG Order: Lithobiomorpha Stone centipedes Family: Lithobiidae Stone centipedes Lithobius forficatus (Linnaeus) * Brown Centipede C D AG Order: Julida Millipedes Family: Julidae Millipedes Tachypodoiulus niger (Leach) Black Snake Millipede C B AG Order: Isopoda Woodlice & water-lice etc. Suborder: Oniscidea Woodlice Family: Oniscidae Woodlice Oniscus asellus Linnaeus Common Woodlouse C E AG Philoscia muscorum (Scopoli) Common Striped Woodlouse C D AG Family: Porcellionidae Woodlice Porcellio scaber Latreille Common Rough Woodlouse C E AG Family: Armadillidiidae Pill woodlice Armadillidium vulgare (Latreille) Common Pill Woodlouse C C AG Order: Odonata Dragonflies & damselflies Suborder: Zygoptera Damselflies Family: Coenagriidae Blue damselflies etc. Coenagrion puella (Linnaeus) Azure Damselfly C C AG Ischnura elegans (Vander Linden) Blue-tailed Damselfly C B AG Pyrrhosoma nymphula (Sulzer) Large Red Damselfly C A PJB Suborder: Anisoptera Dragonflies Family: Aeshnidae Hawkers Aeshna grandis (Linnaeus) Brown Hawker C C AG Aeshna mixta (Latreille) Migrant Hawker C C AG Anax imperator Leach Emperor Dragonfly C B AG Family: Libellulidae Skimmers & darters etc. Libellula depressa Linnaeus Broad-bodied Chaser C D AG, PJB Libellula quadrimaculata Linnaeus Four-spotted Chaser C B AG, PJB Sympetrum danae (Sulzer) * Black Darter C B AG, PJB Sympetrum striolatum (Charpentier) Common Darter C D AG, PJB Order: Orthoptera Grasshoppers etc. Family: Tetrigidae Groundhoppers Tetrix undulata (Sowerby) Common Groundhopper C B AG Family: Acrididae Grasshoppers Chorthippus brunneus (Thunberg) Common Field Grasshopper C D AG Omocestus viridulus (Linnaeus) Common Green Grasshopper C E AG Order: Dermaptera Earwigs Family: Forficulidae Earwigs Forficula auricularia Linnaeus Common Earwig C B AG Order: Hemiptera True bugs etc. Suborder: Heteroptera True bugs Family: Acanthosomatidae Shieldbugs Acanthosoma haemorrhoidale (Linnaeus) * Hawthorn Shieldbug C A AG Elasmostethus interstinctus (Linnaeus) Birch Shieldbug C B AG 44 INVERTEBRATE LIST FOR 2018

NAT. NOS. SCIENTIFIC NAME VERNACULAR NAME DET. STATUS CODE Elasmucha grisea (Linnaeus) Parent Bug C F AG Family: Pentatomidae Shieldbugs Dolycoris baccarum (Linnaeus) Hairy Shieldbug C B AG Palomena prasina (Linnaeus) Green Shieldbug C C AG Pentatoma rufipes (Linnaeus) Red-legged Shieldbug C A AG Family: Lygaeidae Ground bugs Kleidocerys resedae (Panzer) Birch Catkin Bug C A AG Family: Miridae Grass-bugs etc. Leptopterna dolabrata (Linnaeus) Meadow Plant Bug C A AG Lygus rugulipennis (Poppius) * Tarnished Plant Bug C A AG Orthops basalis (A. Costa) a plant bug L A AG Orthops campestris (Linnaeus) * a plant bug C A AG Psallus betuleti (Fallén) a plant bug L A AG Family: Nabidae Damsel bugs Himacerus mirmicoides (O. Costa) Ant Damsel Bug C A AG Nabis rugosus (Linnaeus) Common Damsel Bug C A AG Suborder: Auchenorrhyncha Planthoppers etc. Family: Cercopidae Froghoppers Cercopis vulnerata Illiger Black-and-red Froghopper C B AG Family: Aphrophoridae Froghoppers Philaenus spumarius (Linnaeus) Common Froghopper C B AG Family: Cicadellidae Leafhoppers Allygus modestus Scott a leafhopper C A AG Balclutha punctata (Fabricius) a leafhopper C A AG Order: Neuroptera Lacewings etc. Family Chrysopidae Green lacewings Chrysopa perla (Linnaeus) a green lacewing C A AG Chrysoperla carnea (Stephens) [s. s.] Common Green Lacewing C A AG Chrysoperla carnea (Stephens) [s. l.] Common Green Lacewing n/a B AG Order: Mecoptera Scorpion flies etc. Family: Panorpidae Scorpion flies etc. Panorpa communis Linnaeus Common Scorpion-fly C B AG Order: Lepidoptera Butterflies & moths Suborder: Heterocera Moths Family: Eriocraniidae Micro moths Eriocrania sparrmannella (Bosc) * a micro moth L A AG Family: Yponomeutidae Ermine moths etc. Yponomeuta cagnagella (Hübner) * Spindle Ermine C E PJB Family: Zygaenidae Burnet moths Zygaena lonicerae (Scheven) Narrow-bordered Five-spot Burnet C B AG Family: Drepanidae Hook-tip moths Drepana falcataria (Linnaeus) Pebble Hook-tip C A RHAB Family: Geometridae Carpet moths etc. Alcis repandata (Linnaeus) Mottled Beauty C A RHAB Epirrhoe alternata (Müller) Common Carpet C B RHAB Eupithecia centaureata (Denis & Schiffermüller) Lime-speck Pug C B RHAB Scotopteryx chenopodiata (Linnaeus) Shaded Broad-bar SPIE B RHAB Timandra comae Schmidt Blood-vein SPIE A AG 45 INVERTEBRATE LIST FOR 2018

NAT. NOS. SCIENTIFIC NAME VERNACULAR NAME DET. STATUS CODE Xanthorhoe montanata (Denis & Schiffermüller) Silver-ground Carpet C A AG Family: Notodontidae Kitten & prominent moths Pheosia gnoma (Fabricius) Lesser Swallow Prominent C B RHAB Family: Erebidae Tiger & footman moths etc. Callistege mi (Clerck) Mother Shipton Moth L B AG Eilema griseola (Hübner) Dingy Footman C A AG Euclidia glyphica (Linnaeus) * Burnet Companion L B AG Phragmatobia fuliginosa (Linnaeus) Ruby Tiger C A AG Tyria jacobaeae (Linnaeus ) The Cinnabar SPIE C AG Family: Noctuidae Owlet moths Agrotis puta (Hübner) * Shuttle-shaped Dart C A RHAB Autographa gamma (Linnaeus) Silver Y C C AG, PJB Enargia paleacea (Esper) * Angle-striped Sallow L A RHAB Hydraecia micacea (Esper) Rosy Rustic SPIE B RHAB Lithophane ornitopus (Dadd) * Grey-shoulder Knot L A AG Luperina testacea (Denis & Schiffermüller) Flounced Rustic C A RHAB Mesapamea secalis (Linnaeus) [s. l.] Common Rustic C A RHAB Noctua janthe (Borkhausen) Lesser Broad-bordered Yellow Underwing C B RHAB Noctua pronuba (Linnaeus) Large yellow Underwing C B RHAB Ochropleura plecta (Linnaeus) Flame Shoulder C C RHAB Rivula sericealis (Scopoli) Straw Dot L B RHAB Xestia baja (Denis & Schiffermüller) Dotted Clay C A RHAB Xestia c-nigrum (Linnaeus) Setaceous Hebrew Character C D RHAB Xestia sexstigata (Howarth) Six-striped Rustic C B RHAB Suborder: Rhopalocera Butterflies Family: Hesperiidae Skippers Erynnis tages (Linnaeus) * Dingy Skipper SPIE B AG, PJB Ochlodes sylvanus (Esper) Large Skipper C C AG, PJB Thymelicus sylvestris (Poda) Small Skipper C E AG, PJB Family: Pieridae Whites & yellows etc. Anthocharis cardamines (Linnaeus) Orange Tip C D AG, PJB Gonepteryx rhamni (Linnaeus) Brimstone C B PJB Pieris brassicae (Linnaeus) Large White C C AG, PJB Pieris napi (Linnaeus) Green-veined White C C AG, PJB Pieris rapae (Linnaeus) Small White C E AG, PJB Family: Nymphalidae Brush-footed butterflies etc. Aglais io (Linnaeus) Peacock C C AG, PJB Aglais urticae (Linnaeus) Small Tortoiseshell C D AG, PJB Aphantopus hyperantus (Linnaeus) Ringlet C D AG, PJB Maniola jurtina (Linnaeus) Meadow Brown C E AG, PJB Melanargia galathea (Linnaeus) * Marbled White L B PJB Pararge aegeria (Linnaeus) Speckled Wood C D AG, PJB Pyronia tithonus (Linnaeus) Gatekeeper C A AG Vanessa atalanta (Linnaeus) Red Admiral C C AG, PJB Vanessa cardui (Linnaeus) Painted Lady C B AG, PJB Family: Lycaenidae Blues & hairstreaks etc. Aricia agestis (Denis & Schiffermüller) Brown Argus L C AG, PJB Celastrina argiolus (Linnaeus) Holly Blue C B PJB 46 INVERTEBRATE LIST FOR 2018

NAT. NOS. SCIENTIFIC NAME VERNACULAR NAME DET. STATUS CODE Favonius quercus (Linnaeus) * Purple Hairstreak C B PJB Lycaena phlaeas (Linnaeus) Small Copper C E AG, PJB Polyommatus icarus (Rottemburg) Common Blue C C AG, PJB Order: Coleoptera Beetles Suborder: Adephaga Ground beetles etc. Family: Carabidae Ground beetles Amara aenea (De Geer) * a ground beetle C A AG Bembidion illigeri Netolitzky a ground beetle L A AG Calathus melanocephalus (Linnaeus) a ground beetle C C AG Curtonotus aulicus (Panzer) a ground beetle C B AG Cychrus caraboides (Linnaeus) * a ground beetle L A AG Elaphrus riparius (Linnaeus) a ground beetle C A AG Poecilus versicolor (Sturm) a ground beetle C B AG Pterostichus melanarius (Illiger) a ground beetle C B AG Pterostichus niger (Schaller) a ground beetle C C AG Pterostichus strenuus (Panzer) a ground beetle C C AG Pterostichus vernalis (Panzer) a ground beetle C A AG Trechus obtusus Erichson * a ground beetle C B AG Suborder: Polyphaga Beetles and weevils Family: Silphidae Carrion beetles etc. Silpha atrata (Linnaeus) a carrion beetle C A AG Silpha tristis (Illiger) a carrion beetle L A AG Family: Scarabaeidae Dung beetles etc. Phyllopertha horticola (Linnaeus) Garden Chafer C C AG Family: Elateridae Click beetles Agriotes acuminatus (Stephens) * a click beetle C A AG Athous haemorrhoidalis (Fabricius) a click beetle C A AG Cidnopus aeruginosus (Olivier) a click beetle L B AG Kibunea minuta (Linnaeus) * a click beetle C A AG Family: Cantharidae Soldier beetles Cantharis nigricans (Müller) a soldier beetle C A AG Cantharis rufa Linnaeus a soldier beetle C B AG Cantharis rustica Fallén a soldier beetle C B AG Rhagonycha fulva (Scopoli) a soldier beetle C C AG Family: Malachiidae Soft-wing flower beetles Malachius bipustulatus (Linnaeus) Malachite Beetle C B AG Family: Nitidulidae Pollen beetles etc. Meligethes aeneus (Fabricius) Rape Fly C D AG Family: Coccinellidae Ladybirds Coccinella septempunctata (Linnaeus) 7-spot Ladybird C D AG, PJB Halyzia sedecimguttata (Linnaeus) * Orange Ladybird L A RHAB Harmonia axyridis (Pallas) Harlequin Ladybird C C AG Propylea quatuordecimpunctata (Linnaeus) 14-spot Ladybird C B AG Psyllobora vigintiduopunctata (Linnaeus) 22-spot Ladybird C A AG Family: Oedemeridae False blister beetles Oedemera lurida (Marsham) a false blister beetle L B AG Family: Pyrochroidae Cardinal beetles etc. Pyrochroa serraticornis (Scopoli) * Red-headed Cardinal Beetle C B AG 47 INVERTEBRATE LIST FOR 2018

NAT. NOS. SCIENTIFIC NAME VERNACULAR NAME DET. STATUS CODE Family: Cerambycidae Longhorn beetles Clytus arietis (Linnaeus) Wasp Beetle C A AG Rutpela maculata (Poda) Black and Yellow Longhorn Beetle C B AG Family: Chrysomelidae Leaf beetles Agelastica alni (Linnaeus) Alder Leaf Beetle JNCC C AG Psylliodes chrysocephala (Linnaeus) * Cabbage Stem Flea Beetle C A AG Family: Curculionidae Weevils Phyllobius maculicornis Germar a weevil C B AG Phyllobius pyri (Linnaeus) Common Leaf Weevil C B AG Polydrusus formosus (Mayer) * a weevil JNCC C AG Order: Hymenoptera Sawflies, bees & ichneumons etc. Suborder: Symphyta Sawflies Family: Cephidae Sawflies Cephus spinipes (Panzer) a sawfly C A AG Family: Tenthredinidae Sawflies Athalia cordata Lepeletier a sawfly C A AG Eutomostethus luteiventris (Klug) * a sawfly C A AG Pachyprotasis rapae (Linnaeus) a sawfly C A AG Tenthredo arcuata Förster a sawfly C A AG Tenthredo brevicornis (Konow) a sawfly C B AG Tenthredo mesomela Linnaeus a sawfly C B AG Tenthredo notha Klug a sawfly C A AG Suborder: Apocrita Bees, wasps & ichneumons etc. Superfamily: Aculeata Bees, wasps & ants Family: Formicidae Ants Lasius niger (Linnaeus) [s. s.] Black Garden Ant C E AG Family: Vespidae Social wasps Dolichovespula saxonica (Fabricius) Saxon Wasp JNCC B AG Vespa crabro Linnaeus Hornet L B AG, PJB Vespula germanica (Fabricius) German Wasp C D AG Vespula vulgaris (Linnaeus) Common Wasp C C AG Family: Crabronidae Solitary wasps Crabro cribrarius (Linnaeus) a solitary wasp C A AG Ectemnius continuus (Fabricius) a solitary wasp C A AG Mellinus arvensis (Linnaeus ) Field Digger Wasp C A AG Family: Andrenidae Mining bees Andrena denticulata (Kirby) Grey-banded Mining Bee L A AG Andrena haemorrhoa (Fabricius) Early Mining Bee C C AG Andrena helvola (Linnaeus) Coppice Mining Bee L A AG Andrena nigroaenea (Kirby) Buffish Mining Bee C B AG Andrena scotica Perkins Chocolate Mining Bee C B AG Andrena subopaca Nylander Impunctate Mini-miner C B AG Andrena synadelpha Perkins * Broad-margined Mining Bee L A AG Andrena wilkella (Kirby) Wilke’s Mining Bee L A AG Family: Halictidae Mining bees etc. Lasioglossum albipes (Fabricius) Bloomed Furrow Bee C A AG Lasioglossum leucozonium (Schrank) White-zoned Furrow Bee C A AG Lasioglossum villosulum (Kirby) Shaggy Furrow Bee C A AG 48 INVERTEBRATE LIST FOR 2018

NAT. NOS. SCIENTIFIC NAME VERNACULAR NAME DET. STATUS CODE Sphecodes gibbus (Linnaeus) Dark-winged Blood Bee L A AG Family: Megachilidae Mason bees etc. Osmia bicornis (Linnaeus) Red Mason Bee C B AG Family: Anthophoridae Nomad bees Nomada flava Panzer Flavous Nomad Bee C A AG Nomada panzeri Lepeletier Panzer’s Nomad Bee C A AG Family: Apidae Bumblebees etc. Apis mellifera Linnaeus Honey Bee C E AG Bombus bohemicus (Seidl) Gypsy Cuckoo-bee C B AG Bombus hortorum (Linnaeus) Small Garden Bumblebee C B AG Bombus lapidarius (Linnaeus) Red-tailed Bumblebee C E AG Bombus lucorum (Linnaeus) White-tailed Bumblebee C A AG Bombus pascuorum (Scopoli) Common Carder-bee C D AG Bombus pratorum (Linnaeus) Early Bumblebee C C AG Bombus rupestris (Fabricius) Hill Cuckoo-bee JNCC C AG Bombus terrestris (Linnaeus) Buff-tailed Bumblebee C C AG Bombus vestalis (Geoffroy) Vestal Cuckoo-bee C C AG Superfamily: Parasitica Ichneumon wasps etc. Family: Cynipidae Gall Wasps Andricus kollari (Hartig) Oak Marble Gall Wasp C C AG Family: Ichneumonidae Ichneumon wasps Chorinaeus brevicalcar Thomson * an ichneumon wasp U A WAE Diplazon laetatorius (Fabricius) * an ichneumon wasp U A WAE Glypta nigrotrochanterata Strobl an ichneumon wasp U A WAE Ichneumon stramentor Rasnitsyn * an ichneumon wasp U B WAE Itoplectis maculator (Fabricius) * an ichneumon wasp U B WAE Mesoleptus vigilatorius (Förster) * an ichneumon wasp U A RJ Scambus buolianae (Hartig) an ichneumon wasp U A WAE Scambus signatus (Pfeffer) * an ichneumon wasp U A WAE Order: Diptera True flies Suborder: Nematocera Thread-horns Family: Tipulidae Crane-flies Nephrotoma appendiculata (Pierre) a crane-fly C A AG Nephrotoma flavescens (Linnaeus) a crane-fly C A AG Tipula oleracea Linnaeus a crane-fly C B AG Tipula pagana Meigen * a crane-fly C B AG Tipula paludosa Meigen a crane-fly C C AG Tipula varipennis Meigen * a crane-fly C A AG Tipula vernalis Meigen a crane-fly C B AG Family: Pediciidae Crane-flies Tricyphona immaculata (Meigen) a crane-fly C A AG Family: Limoniidae Crane-flies Dicranomyia modesta (Meigen) a crane-fly C A AG Metalimnobia quadrinotata (Meigen) * a crane-fly C A AG Ormosia hederae (Curtis) a crane-fly C B AG Family: Bibionidae March flies Bibio johannis (Linnaeus) a March fly C A AG Bibio leucopterus (Meigen) a March fly C A AG 49 INVERTEBRATE LIST FOR 2018

NAT. NOS. SCIENTIFIC NAME VERNACULAR NAME DET. STATUS CODE Bibio marci (Linnaeus) St. Mark’s Fly C F AG Dilophus febrilis (Linnaeus) Fever Fly C B AG Dilophus femoratus Meigen a March fly C A AG Family: Trichoceridae Winter gnats Trichocera annulata Meigen * a winter gnat C A AG Trichocera hiemalis (De Geer) * a winter gnat L A AG Trichocera regelationis (Linnaeus) * a winter gnat C B AG Family: Anisopodidae Wood gnats Sylvicola punctatus (Fabricius) a wood gnat C A AG Family: Culicidae Mosquitoes Ochlerotatus annulipes (Meigen) * a mosquito C B AG Ochlerotatus punctor (Kirby) a mosquito C C AG Ochlerotatus rusticus (Rossi) * a mosquito C C AG Suborder: Brachycera Short-horns Family: Rhagionidae Snipe-flies Rhagio lineola Fabricius Small Fleck-winged Snipe-fly C B AG Rhagio scolopaceus (Linnaeus) Down-looker Fly C C AG Rhagio tringarius (Linnaeus) Marsh Snipe-fly C B AG Family: Tabanidae Horseflies Haematopota pluvialis (Linnaeus) Notch-horned Cleg C B AG Hybomitra bimaculata (Macquart) * Hairy-legged Horsefly L A AG Family: Stratiomyidae Soldier-flies Chloromyia formosa (Scopoli) Broad Centurion C B AG Family: Therevidae Stiletto-flies Thereva nobilitata (Fabricius) Common Stiletto-fly C A AG Family: Asilidae Robber-flies Dioctria atricapilla Meigen Violet Black-legged Robber-fly L C AG Dioctria rufipes (De Geer) Common Red-legged Robber-fly L C AG Leptogaster cylindrica (De Geer) Striped Slender Robber-fly C B AG Family: Hybotidae Dance flies Bicellaria vana Collin a dance fly C B AG Ocydromia glabricula (Fallén) * a dance fly C A AG Trichinomyia flavipes (Meigen) * a dance fly C B AG Family: Empididae Dance flies Empis chioptera Meigen a dance fly C B AG Empis grisea Fallén * a dance fly C A AG Empis livida Linnaeus a dance fly C B AG Empis nigripes Fabricius a dance fly C A AG Empis nuntia Meigen a dance fly C A AG Empis praevia Collin a dance fly L A AG Empis tessellata Fabricius a dance fly C B AG Hilara maura (Fabricius) a dance fly C C AG Hilara nigrina (Fallén) * a dance fly L A AG Rhamphomyia nigripennis (Fabricius) * a dance fly C A AG Rhamphomyia tarsata Meigen a dance fly C A AG Family: Dolichopodidae Long-legged flies Dolichopus ungulatus (Linnaeus) a long-legged fly C A AG

50 INVERTEBRATE LIST FOR 2018

NAT. NOS. SCIENTIFIC NAME VERNACULAR NAME DET. STATUS CODE Family: Platypezidae Flat-footed flies Protoclythia modesta (Zetterstedt) * a flat-footed fly U A AG Family: Phoridae Scuttle flies Borophaga incrassata (Meigen) * a scuttle fly C B AG Family: Syrphidae Hoverflies Baccha elongata (Fabricius) * a hoverfly C A AG Cheilosia albitarsis (Meigen) [s. s.] a hoverfly C B AG Cheilosia bergenstammi Becker a hoverfly L B AG Cheilosia lasiopa Kowarz a hoverfly L B AG Cheilosia pagana (Meigen) a hoverfly C A AG Cheilosia proxima (Zetterstedt) a hoverfly L B AG Dasysyrphus albostriatus (Fallén) a hoverfly C A AG Dasysyrphus tricinctus (Fallén) a hoverfly C A AG Epistrophe eligans (Harris) a hoverfly C C AG Episyrphus balteatus (De Geer) a hoverfly C D AG Eristalis interruptus (Poda) a hoverfly C B AG Eristalis intricarius (Linnaeus) a hoverfly C B AG Eristalis pertinax (Scopoli) a hoverfly C D AG Eristalis tenax (Linnaeus) Drone-fly C B AG Eumerus strigatus (Fallén) * a lesser bulb fly C A AG Eupeodes corollae (Fabricius) a hoverfly C A AG Eupeodes luniger (Meigen) a hoverfly C A AG Helophilus pendulus (Linnaeus) a hoverfly C C AG Leucozona lucorum (Linnaeus) a hoverfly C B AG Melangyna lasiophthalma (Zetterstedt) * a hoverfly L A AG Melanostoma mellinum (Linnaeus) a hoverfly C B AG Melanostoma scalare (Fabricius) a hoverfly C B AG Merodon equestris (Fabricius) Greater Bulb Fly C B AG Myathropa florea (Linnaeus) * a hoverfly C B AG Platycheirus albimanus (Fabricius) a hoverfly C B AG Platycheirus rosarum (Fabricius) a hoverfly L A AG Platycheirus splendidus Rotheray * a hoverfly U A AG Scaeva pyrastri (Linnaeus) a hoverfly C B AG Sphaerophoria scripta (Linnaeus) a hoverfly C B AG Syritta pipiens (Linnaeus) a hoverfly C C AG Syrphus ribesii (Linnaeus) a hoverfly C C AG Volucella bombylans (Linnaeus) a hoverfly C C AG Volucella pellucens (Linnaeus) a hoverfly C C AG Xylota segnis (Linnaeus) a hoverfly C B AG Family: Psilidae Rust flies Chamaepsila nigricornis (Meigen) a rust fly U B AG Psila merdaria Collin * a rust fly U A AG Family: Lonchaeidae Lance flies Lonchaea corusca Czerny * a lance fly JNCC A AG Family: Tephritidae Picture-winged flies Chaetorellia jaceae (Robineau-Desvoidy) a picture-winged fly L A AG Tephritis hyoscyami (Linnaeus) * a picture-winged fly L A AG Tephritis neesii (Meigen) a picture-winged fly L C AG 51 INVERTEBRATE LIST FOR 2018

NAT. NOS. SCIENTIFIC NAME VERNACULAR NAME DET. STATUS CODE Tephritis vespertina (Loew) a picture-winged fly C A AG Family: Lauxaniidae Lauxaniid flies Calliopum aeneum (Fallén) * a lauxaniid fly C B AG Meiosimyza rorida (Fallén) * a lauxaniid fly C A AG Family: Sciomyzidae Marsh flies Pherbellia griseola (Fallén) * a marsh fly JNCC A AG Tetanocera elata (Fabricius) a marsh fly C A AG Family: Opomyzidae Cereal flies Opomyza florum (Fabricius) Yellow Cereal Fly C C AG Opomyza germinationis (Linnaeus) a cereal fly C A AG Opomyza petrei Mesnil a cereal fly C B AG Family: Heleomyzidae Heleomyzid flies Suillia pallida (Fallén) * a heleomyzid fly C A AG Family: Drosophilidae Vinegar flies Scaptomyza flava (Fallén) * Turnip Leafminer C A AG Family: Scathophagidae Dung flies etc. Conisternum decipiens (Haliday) * a dung fly JNCC A AG Nanna fasciata (Meigen) a Timothy fly C B AG Nanna inermis (Becker) a Timothy fly L B AG Scathophaga furcata (Say) a dung fly C A AG Scathophaga stercorea (Linnaeus) Yellow Dung Fly C B AG Family: Anthomyiidae Root-maggot flies etc. Anthomyia liturata (Robineau-Desvoidy) * an anthomyiid fly C A AG Anthomyia procellaris Rondani an anthomyiid fly C A AG Botanophila brunneilinea (Zetterstedt) an anthomyiid fly U B AG Botanophila fugax (Meigen) * an anthomyiid fly C B AG Botanophila striolata (Fallén)/discreta (Meigen) * an anthomyiid fly L A AG Delia florilega (Zetterstedt) an anthomyiid fly C B AG Delia platura (Meigen) Seedcorn Maggot Fly C D AG Delia radicum (Linnaeus) an anthomyiid fly C B AG Phorbia fumigata (Meigen) * Spring Fly U A AG Family: Fanniidae Lesser Housefly etc. Fannia pallitibia (Rondani) * a fanniid fly L A AG Family: Muscidae Housefly & sweat flies etc. Coenosia infantula Rondani * a muscid fly C A AG Coenosia perpusilla Meigen * a muscid fly L A AG Coenosia testacea (Robineau-Desvoidy) a muscid fly C B AG Coenosia tigrina (Fabricius) a muscid fly C B AG Hebecnema vespertina (Fallén) a muscid fly C A AG Helina depuncta (Fallén) a muscid fly C A AG Helina evecta (Harris) a muscid fly C A AG Helina impuncta (Fallén) a muscid fly C B AG Helina reversio (Harris) a muscid fly C B AG Hydrotaea irritans (Fallén) a sweat fly C B AG Hydrotaea meteorica (Linnaeus) a sweat fly C A AG Mesembrina meridiana Linnaeus Noon Fly C B AG Musca autumnalis De Geer * a muscid fly C B AG Muscina levida (Harris) a muscid fly C B AG 52 INVERTEBRATE LIST FOR 2018

NAT. NOS. SCIENTIFIC NAME VERNACULAR NAME DET. STATUS CODE Phaonia angelicae (Scopoli) a muscid fly C C AG Phaonia incana (Wiedemann) a muscid fly C C AG Phaonia tuguriorum (Scopoli) a muscid fly C C AG Polietes lardarius (Fabricius) * a muscid fly C B AG Thricops simplex (Wiedemann) * a muscid fly U B AG Family: Calliphoridae Bluebottles etc. Calliphora vicina Robineau-Desvoidy Common Bluebottle C C AG Cynomya mortuorum (Linnaeus) Yellow-faced Blowfly C B AG Lucilia ampullacea Villeneuve Streakless Greenbottle L A AG Lucilia caesar (Linnaeus) Common Greenbottle C A AG Melanomya nana (Meigen) Little Black Blowfly C B AG Pollenia angustigena Wainwright Narrow-cheeked Clusterfly C B AG Pollenia pediculata Macquart Tufted Clusterfly C B AG Pollenia rudis (Fabricius) Common Clusterfly C A AG Protocalliphora azurea (Fallén) * Bird Blowfly U A AG Protophormia terraenovae (Robineau-Desvoidy) * Blackbottle C A AG Family: Rhinophoridae Woodlouse flies Rhinophora lepida (Meigen) * a woodlouse fly U A AG Family: Sarcophagidae Flesh flies Sarcophaga carnaria (Linnaeus) a flesh fly C B AG Sarcophaga haemorrhoa Meigen a flesh fly C B AG Sarcophaga nigriventris Meigen a flesh fly C A AG Sarcophaga subvicina Rohdendorf a flesh fly C C AG Sarcophaga vagans Meigen a flesh fly C B AG Family: Tachinidae Parasitic flies Eriothrix rufomaculata (De Geer) a parasitic fly C C AG Eurithia anthophila (Robineau-Desvoidy) a parasitic fly C A AG Exorista rustica (Fallén) a parasitic fly C B AG Gymnocheta viridis (Fallén) a parasitic fly C B AG Phytomyptera cingulata (Robineau-Desvoidy) a parasitic fly U A AG Siphona geniculata (De Geer) a parasitic fly C B AG Tachina fera (Linnaeus) a parasitic fly C C AG Zophomyia temula (Scopoli) a parasitic fly JNCC B AG

53 DISSECTION OF INVERTEBRATE DATA FOR 2018

The following table dissects the numbers of each species into the compartments in which they were recorded. There is an additional column headed ‘U’ to accommodate records which could not be allocated to a particular compartment. The numbers given usually refer to the number of adults, but occasionally refer to larvae, galls or mines. Data for the Bee Hotel is included in the column for Area 1, data for the Bodger’s Den is included in the column for Area 8, and data for the Crombie Roundhouse is included in the column for Area 9. With field records, it is quite possible for the same individuals to be counted more than once, e.g. the same individual can be seen on different days, or in different survey compartments on the same day. In many cases therefore, the overall number of individuals may have been fewer than the total recorded. In cases where the same snail shells were recorded on different visits, the overall total has been adjusted to compensate for this.

+ = the number of individuals were not recorded, therefore the number given is only the minimum number * = the total includes estimated numbers of individuals seen in the field

SPECIES 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 U Class: Gastropoda Family: Arionidae Arion ater [s. l.] 20* 1 3 10* 1 2 Family: Helicidae Cepaea hortensis 2 Cepaea nemoralis 3 1 Cornu aspersum 500* Family: Lymnaeidae Lymnaea stagnalis 100* Order: Araneae Family: Araneidae Araneus diadematus 2 1 1+ Araneus marmoreus var. 1+ Araneus quadratus 1+ Larinioides cornutus 1+ Family: Philodromidae Tibellus species 1+ Family: Pisauridae Pisaura mirabilis 1+ Family: Tetragnathidae Metellina segmentata 1+ Family: Theridiidae Enoplognatha latimana 1+ Order: Glomerida Family: Glomeridae Glomeris marginata 3 1 Order: Polydesmida Family: Polydesmidae Polydesmus angustus 1 Order: Lithobiomorpha Family: Lithobiidae Lithobius forficatus 2 19* 5 Order: Julida Family: Julidae Tachypodoiulus niger 2 1 Order: Isopoda Family: Oniscidae Oniscus asellus 6 120* 1 Philoscia muscorum 2 4 21* 1 Family: Porcellionidae Porcellio scaber 11 11 77* 1 1 Family: Armadillidiidae Armadillidium vulgare 1 2 2 1

54 DISSECTION OF INVERTEBRATE DATA FOR 2018

SPECIES 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 U Order: Odonata Suborder: Zygoptera Family: Coenagriidae Coenagrion puella 1 8 2 Ischnura elegans 4 Pyrrhosoma nymphula 1 Suborder: Anisoptera Family: Aeshnidae Aeshna grandis 1 3 1 1 1 Aeshna mixta 1 3 2 Anax imperator 2 1 Family: Libellulidae Libellula depressa 7 1 Libellula quadrimaculata 1 4 Sympetrum danae 3 Sympetrum striolatum 2 1 2 2 1 52* 1 1 1 1 Order: Orthoptera Family: Tetrigidae Tetrix undulata 3 1 Family: Acrididae Chorthippus brunneus 4 2 1 3 2 6 3 1 15* 3 23* 24* 1 Omocestus viridulus 2 10* 1 1 11* 21* 1 20* 1 15* 7 43* 11* Order: Dermaptera Family: Forficulidae Forficula auricularia 3 2 Order: Hemiptera Suborder: Heteroptera Family: Acanthosomatidae Acanthosoma haemorrhoidale 1 Elasmostethus interstinctus 3 Elasmucha grisea 1 508* 3 Family: Pentatomidae Dolycoris baccarum 1 1 Palomena prasina 14* 1 Pentatoma rufipes 1 Family: Lygaeidae Kleidocerys resedae 1 Family: Miridae Leptopterna dolabrata 1 Lygocoris rugicollis 1 Orthops basalis 1 Orthops campestris 1 Psallus betuleti 1 Family: Nabidae Himacerus mirmicoides 1 Nabis rugosus 1 Suborder: Auchenorrhyncha Family: Cercopidae Cercopis vulnerata 1 1 Family: Aphrophoridae Philaenus spumarius 1 1 Family: Cicadellidae Allygus modestus 1 Balclutha punctata 1 Order: Neuroptera Family Chrysopidae Chrysopa perla 1 Chrysoperla carnea [s. s.] 1 Chrysoperla carnea [s. l.] 2 1 1

55 DISSECTION OF INVERTEBRATE DATA FOR 2018

SPECIES 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 U Order: Mecoptera Panorpidae Panorpa communis 1 1 Order: Lepidoptera Suborder: Heterocera Family: Eriocraniidae Eriocrania sparrmannella 1 Family: Yponomeutidae Yponomeuta cagnagella 105* Family: Zygaenidae Zygaena lonicerae 1 1 1 Family: Drepanidae Drepana falcataria 1 Family: Geometridae Alcis repandata 1 Epirrhoe alternata 2 Eupithecia centaureata 2 Scotopteryx chenopodiata 2 Timandra comae 1 Xanthorhoe montanata 1 Family: Notodontidae Pheosia gnoma 3 Family: Erebidae Callistege mi 1 1 Eilema griseola 1 Euclidia glyphica 1 1 Phragmatobia fuliginosa 1 Tyria jacobaeae 20* 3 Family: Noctuidae Agrotis puta 1 Autographa gamma 1 1 1 2 4 1 1 2 Enargia paleacea 1 Hydraecia micacea 2 Lithophane ornitopus 1 Luperina testacea 1 Mesapamea secalis [s. l.] 1 Noctua janthe 5 Noctua pronuba 2 Ochropleura plecta 6 Rivula sericealis 3 Xestia baja 1 Xestia c-nigrum 47 Xestia sexstigata 3 Suborder: Rhopalocera Family: Hesperiidae Erynnis tages 1 1 Ochlodes sylvanus 1 2 1 2 Thymelicus sylvestris 2 5* 3 1 2 1 2 5* 2 2 170 Family: Pieridae Anthocharis cardamines 1 2 2 1 1 1 4 4 49 Gonepteryx rhamni 5 Pieris brassicae 1 1 2 19 Pieris napi 1 5 2 1 2 3 Pieris rapae 1 1 2 114 Family: Nymphalidae Aglais io 1 1 1 1 5 Aglais urticae 1 1 2 2 2 1 28 Aphantopus hyperantus 3 1 1 1 2 1 2 4 2 5* 2 2 23 Maniola jurtina 4 1 3 1 1 1 2 11* 6 1 2 1 12* 34* 2 1 277 Melanargia galathea 2 56 DISSECTION OF INVERTEBRATE DATA FOR 2018

SPECIES 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 U Pararge aegeria 1 1 9 9* 5 8 Pyronia tithonus 1 Vanessa atalanta 1 1 1 1 2 Vanessa cardui 1 1 Family: Lycaenidae Aricia agestis 2 3 1 2 Celastrina argiolus 3 Favonius quercus 3 Lycaena phlaeas 2 1 2 4 1 1 133 Polyommatus icarus 1 1 1 2 1 1 9 Order: Coleoptera Suborder: Adephaga Family: Carabidae Amara aenea 1 Bembidion illigeri 1 Calathus melanocephalus 9 1 Curtonotus aulicus 1 1 Cychrus caraboides 1 Elaphrus riparius 1 Poecilus versicolor 1 2 2 Pterostichus melanarius 3 2 Pterostichus niger 7 7 3 Pterostichus strenuus 1 11 1 Pterostichus vernalis 1 Trechus obtusus 2 Suborder: Polyphaga Family: Silphidae Silpha atrata 1 Silpha tristis 1 Family: Scarabaeidae Phyllopertha horticola 4 2 Family: Elateridae Agriotes acuminatus 1 Athous haemorrhoidalis 1 Cidnopus aeruginosus 1 2 Kibunea minuta 1 Family: Cantharidae Cantharis nigricans 1 Cantharis rufa 1 1 Cantharis rustica 1 1 Rhagonycha fulva 3 2 2 1 7* Family: Malachiidae Malachius bipustulatus 1 1 2 Family: Nitidulidae Meligethes aeneus 3 1 40* 1 1 Family: Coccinellidae Coccinella septempunctata 1 2 3 9 3 2 2 17 3 1 3 2 Halyzia sedecimguttata 1 Harmonia axyridis 1 1 3 5 1 1 Propylea quatuordecimpunctata 2 2 Psyllobora vigintiduopunctata 1 Family: Oedemeridae Oedemera lurida 1 1 1 Family: Pyrochroidae Pyrochroa serraticornis 2 Family: Cerambycidae Clytus arietis 1 Rutpela maculata 1 1 Family: Chrysomelidae Agelastica alni 6 2 9 57 DISSECTION OF INVERTEBRATE DATA FOR 2018

SPECIES 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 U Psylliodes chrysocephala 1 Family: Curculionidae Phyllobius maculicornis 1 2 Phyllobius pyri 1 1 2 Polydrusus formosus 1 2 1 1 1 Order: Hymenoptera Suborder: Symphyta Family: Cephidae Cephus spinipes 1 Family: Tenthredinidae Athalia cordata 1 Eutomostethus luteiventris 1 Pachyprotasis rapae 1 Tenthredo arcuata 1 Tenthredo brevicornis 1 3 Tenthredo mesomela 2 Tenthredo notha 1 Suborder: Apocrita Superfamily: Aculeata Family: Formicidae Lasius niger [s. s.] 3 2 21* 30* 100* 22* Family: Vespidae Dolichovespula saxonica 2 Vespa crabro 1 1 1 1 1 Vespula germanica 1 1 2 3 2 1 2 2 18* Vespula vulgaris 1 1 4 1 3 3 Family: Crabronidae Crabro cribrarius 1 Ectemnius continuus 1 Mellinus arvensis 1 Family: Andrenidae Andrena denticulata 1 Andrena haemorrhoa 1 1 1 3 1 Andrena helvola 1 Andrena nigroaenea 1 1 Andrena scotica 1 1 Andrena subopaca 1 1 Andrena synadelpha 1 Andrena wilkella 1 Family: Halictidae Lasioglossum albipes 1 Lasioglossum leucozonium 1 Lasioglossum villosulum 1 Sphecodes gibbus 1 Family: Megachilidae Osmia bicornis 3 1 Family: Anthophoridae Nomada flava 1 Nomada panzeri 1 Family: Apidae Apis mellifera 20* 30* 10* 30* 1 61* 30* 1 2 4 1 30* 3 11* 1 Bombus bohemicus 1 3 Bombus hortorum 1 1 Bombus lapidarius 1 21* 12* 14* 7* 4 5 2 35* 3 1 2 3 4 2 Bombus lucorum 1 Bombus pascuorum 2 1 1 2 5 1 5 25* 5 6 2 7 6 2 1 Bombus pratorum 2 1 1 4 Bombus rupestris 2 1 1 1 1 Bombus terrestris 1 1 1 1 1 2 1 2 2 Bombus vestalis 1 1 2 11* 58 DISSECTION OF INVERTEBRATE DATA FOR 2018

SPECIES 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 U Superfamily: Parasitica Family: Cynipidae Andricus kollari 1 6 1 2 Family: Ichneumonidae Chorinaeus brevicalcar 1 Diplazon laetatorius 1 Glypta nigrotrochanterata 1 Ichneumon stramentor 2 Itoplectis maculator 1 1 Mesoleptus vigilatorius 1 Scambus buolianae 1 Scambus signatus 1 Order: Diptera Suborder: Nematocera Family: Tipulidae Nephrotoma appendiculata 1 Nephrotoma flavescens 1 Tipula oleracea 1 2 Tipula pagana 1 1 1 Tipula paludosa 1 1 2 10* 1 1 1 4 3 1 2 Tipula varipennis 1 Tipula vernalis 1 1 Family: Pediciidae Tricyphona immaculata 1 Family: Limoniidae Dicranomyia modesta 1 Metalimnobia quadrinotata 1 Ormosia hederae 2 Family: Bibionidae Bibio johannis 1 Bibio leucopterus 1 Bibio marci 1 20* 50* 2 502* 200* 10* 20* 2 Dilophus febrilis 1 1 Dilophus femoratus 1 Family: Trichoceridae Trichocera annulata 1 Trichocera hiemalis 1 Trichocera regelationis 4 Family: Anisopodidae Sylvicola punctatus 1 Family: Culicidae Ochlerotatus annulipes 4 1 Ochlerotatus punctor 5 2 Ochlerotatus rusticus 1 7 Suborder: Brachycera Family: Rhagionidae Rhagio lineola 2 Rhagio scolopaceus 1 2 1 14* 1 Rhagio tringarius 2 1 Family: Tabanidae Haematopota pluvialis 1 1 1 1 1 Hybomitra bimaculata 1 Family: Stratiomyidae Chloromyia formosa 1 2 Family: Therevidae Thereva nobilitata 1 Family: Asilidae Dioctria atricapilla 1 1 1 1 1 1 Dioctria rufipes 4 1 1 1 2 Leptogaster cylindrica 2 59 DISSECTION OF INVERTEBRATE DATA FOR 2018

SPECIES 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 U Family: Hybotidae Bicellaria vana 2 1 Ocydromia glabricula 1 Trichinomyia flavipes 2 Family: Empididae Empis chioptera 1 1 2 1 Empis grisea 1 Empis livida 1 1 2 Empis nigripes 1 Empis nuntia 1 Empis praevia 1 Empis tessellata 1 3 1 Hilara maura 1 4 4 Hilara nigrina 1 Rhamphomyia nigripennis 1 Rhamphomyia tarsata 1 Family: Dolichopodidae Dolichopus ungulatus 1 Family: Platypezidae Protoclythia modesta 1 Family: Phoridae Borophaga incrassata 2 Family: Syrphidae Baccha elongata 1 Cheilosia albitarsis [s. s.] 1 1 Cheilosia bergenstammi 2 Cheilosia lasiopa 3 Cheilosia pagana 1 Cheilosia proxima 4 Dasysyrphus albostriatus 1 Dasysyrphus tricinctus 1 Epistrophe eligans 1 3 3 Episyrphus balteatus 1 1 2 2 21* 3 11* Eristalis interruptus 2 Eristalis intricarius 1 1 1 Eristalis pertinax 1 5 1 1 2 20* 2 3 3 Eristalis tenax 1 1 1 1 1 Eumerus strigatus 1 Eupeodes corollae 1 Eupeodes luniger 1 Helophilus pendulus 1 1 1 1 1 1 4 2 1 Leucozona lucorum 1 1 Melangyna lasiophthalma 1 Melanostoma mellinum 1 1 Melanostoma scalare 1 1 1 1 Merodon equestris 2 1 1 Myathropa florea 2 1 Platycheirus albimanus 1 1 1 2 Platycheirus rosarum 1 Platycheirus splendidus 1 Scaeva pyrastri 1 1 Sphaerophoria scripta 1 3 Syritta pipiens 1 1 1 1 3 Syrphus ribesii 1 2 3 1 1 1 8 4 2 Volucella bombylans 2 11* 1 Volucella pellucens 10* 1 Xylota segnis 2 1 Family: Psilidae Chamaepsila nigricornis 3 2 Psila merdaria 1 60 DISSECTION OF INVERTEBRATE DATA FOR 2018

SPECIES 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 U Family: Lonchaeidae Lonchaea corusca 1 Family: Tephritidae Chaetorellia jaceae 1 Tephritis hyoscyami 1 Tephritis neesii 3 1 1 2 1 Tephritis vespertina 1 Family: Lauxaniidae Calliopum aeneum 1 1 Meiosimyza rorida 1 Family: Sciomyzidae Pherbellia griseola 1 Tetanocera elata 1 Family: Opomyzidae Opomyza florum 1 12 2 Opomyza germinationis 1 Opomyza petrei 2 Family: Heleomyzidae Suillia pallida 1 Family: Drosophilidae Scaptomyza flava 1 Family: Scathophagidae Conisternum decipiens 1 Nanna fasciata 1 2 Nanna inermis 1 2 Scathophaga furcata 1 Scathophaga stercoraria 2 1 1 Family: Anthomyiidae Anthomyia liturata 1 Anthomyia procellaris 1 Botanophila brunneilinea 2 1 Botanophila fugax 1 2 Botanophila striolata/discreta 1 Delia florilega 3 1 Delia platura 1 3 17 1 2 2 2 2 Delia radicum 1 2 Phorbia fumigata 1 Family: Fanniidae Fannia pallitibia 1 Family: Muscidae Coenosia infantula 1 Coenosia perpusilla 1 Coenosia testacea 1 1 Coenosia tigrina 1 1 Hebecnema vespertina 1 Helina depuncta 1 Helina evecta 1 Helina impuncta 1 1 1 2 Helina reversio 1 3 Hydrotaea irritans 1 2 2 Hydrotaea meteorica 1 Mesembrina meridiana 1 3 Musca autumnalis 1 1 Muscina levida 2 Phaonia angelicae 3 3 1 Phaonia incana 1 1 2 2 1 Phaonia tuguriorum 2 1 3 1 2 Polietes lardarius 2 1 1 1 Thricops simplex 2

61 DISSECTION OF INVERTEBRATE DATA FOR 2018

SPECIES 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 U Family: Calliphoridae Calliphora vicina 1 1 2 1 1 1 2 Cynomya mortuorum 1 1 Lucilia ampullacea 1 Lucilia caesar 1 Melanomya nana 1 1 Pollenia angustigena 1 1 Pollenia pediculata 1 1 2 Pollenia rudis 1 Protocalliphora azurea 1 Protophormia terraenovae 1 Family: Rhinophoridae Rhinophora lepida 1 Family: Sarcophagidae Sarcophaga carnaria 2 1 Sarcophaga haemorrhoa 1 1 Sarcophaga nigriventris 1 Sarcophaga subvicina 4 1 1 1 1 Sarcophaga vagans 2 1 Family: Tachinidae Eriothrix rufomaculata 2 2 1 1 Eurithia anthophila 1 Exorista rustica 1 1 1 Gymnocheta viridis 1 1 Phytomyptera cingulata 1 Siphona geniculata 2 Tachina fera 1 1 3 1 Zophomyia temula 1 1 1 1

62 PLANT LIST FOR 2018

The table below summarises plants mentioned within the report. The plant list is neither extensive nor inclusive, but merely serves to record a few conspicuous flowering plants which provide sources of nectar and pollen for invertebrates.

TAXA COMMON NAME AREAS Apiaceae Parsley family [unspecified] 3, 5, 11, 12, 13, 14, 15 Caltha palustris Marsh Marigold 10 Cardamine pratensis Lady’s Smock 14 Centaurea nigra Common Knapweed 1, 2, 3, 4, 5, 6, 7, 8, 9, 11, 13, 14, 15, 16 Cirsium Thistles [unspecified] 2, 6, 11, 12, 13 Echium vulgare Viper’s Bugloss 11 Epilobium angustifolium Rose-bay Willow-herb 2, 7, 10, 11 Ficaria verna Lesser Celandine 11 Galanthus nivalis Snowdrop 11 Heracleum sphondylium Hogweed 11, 12, 14 Hyacinthoides non-scripta Bluebell 7, 11, 14 Hypochaeris radicata Cat’s Ear 15 Lamium album White Deadnettle 11, 12 Lamium purpureum Purple Deadnettle 3, 7 Leucanthemum vulgare Ox-eye Daisy 1, 2, 3, 4, 6, 7, 8, 9, 10, 11, 13, 14, 15, 16 Lotus corniculatus Bird’s-foot Trefoil 8, 9, 13 Lotus uliginosus Greater Bird’s-foot Trefoil 1, 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 13, 15 Lychnis flos-cuculi Ragged Robin 6, 7, 8 Malva moschata Musk Mallow 1, 2, 3, 4, 5, 6, 7, 9, 13, 15 Myosotis Forget-me-nots [unspecified] 16 Narcissus Daffodils [unspecified] 11 Papaver rhoeas Field Poppy 13 Primula veris Cowslip 6, 7, 8, 9, 14 Ranunculus Buttercups [unspecified] 1, 2, 3, 4, 5, 6, 7, 8, 9, 11, 12, 13, 14, 15, 16 Rhinanthus minor Yellow Rattle 9 Rubus fruticosus Bramble 11, 15 Senecio jacobaea Common Ragwort 2, 3, 4, 6, 7, 8, 10, 11, 12, 13, 14, 15 Senecio vulgaris Groundsel 11, 13 Silene dioica Red Campion 11 Sonchus arvensis Perennial Sow-thistle 14 Taraxacum officinale Dandelion 1, 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 12, 13, 14, 15, 16 Trifolium pratense Red Clover 1, 4, 6, 7, 8, 9, 10, 11, 14 Trifolium repens White Clover 5, 6, 7, 8, 9, 10, 13, 14, 15