![Polypedates Leucomystax :Q39Q8 Rhacophorus Bisacculus :Q3DQA]DLOLTIQ8 Rhacophorus Bipunctatus :Q34T8]JDVL&](http://data.docslib.org/img/1b30416b65ebf81a7d4775b8f5db6914-1.webp)
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`8=86V ]4BT TA@'P34P&'P34P&LW6AQ8^J &)Q4]!QIS S9J Q)P8 'P&JF3'P P869WBT MORPHOLOGICAL CHARACTER OF SOME TADPOLE FROM KHAO SIP HA CHAN PROPOSED NATIONAL PARK, CHANTHABURI PROVINCE
=)BP 38PAIFI3P -S /, LQ'L& :BO64IP W86BIQB-/, ^DO F])./S #8*VL-/* Pachara Danaisawat-/, Art-ong Pradatsundarasan-/ & Wichase Khonsue-/*
ABSTRACT Descriptive morphology and dichotomous key of tadpole are an important information to gain the knowledge of amphibian diversity in Thailand, but still rare. To identify the amphibian species, these informations are needs. This study, we provided description data and key to species of tadpole from Khao Sip Ha Chan Proposed National Park, Chanthaburi Province. The tadpoles collected from aquatic habitat between January – December 008. To identify the species, morphological characters, mouth path structure and dental formula were compared with previous study. Some of tadpoles were reared until complete metamorphosis to confirm their species. We found 1 order, 6 family, 1 genus, and 9 species of tadpole. We described morphology and made dichotomous key for the tadpole in this area. Key word: tadpole, morphological character, mouth path structure, dental formulary, Khao Sip Ha Chan
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FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 6
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FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 66
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F&G Megophryidae Xenophrys lekaguli LU& BQA'869P XB2
F&G Bufonidae Bufo melanostictus #Q&# 9Q8 Bufo parvus #Q&# ^#BO Bufo macrotis #Q&# JFBQ9P
FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 67
F&G Microhylidae Calluella guttulata LU&DQA Glyphoglossus molossus LU&];Q Microhyla berdmorei LU&^@ J8QF Microhyla butleri LU&DQA]DLO Microhyla heymonsi LU&!Q&3 Q Microhyla fissipes LU&8Q]4 Q Microhyla pulchra LU&!Q#Q Micryletta inornata LU&JD&!P T3 Kaloula pulchra LU&L Q&9Q8 Kaloula mediolineata LU&L Q& 8! T3 F&G Dicroglossidae Fejervarya limnocharis 9J8L& Hoplobatrachus rugulosus 98Q Occidozyga lima ]!TA3'O8Q Occidozyga martensii ]!TA3JD&:P W@ Paa fasiculispina 9L J8Q@ F&G Ranidae Rana erythraea ]!TA3'S , ]!TA39FP Rana lateralis 9JD&a=DP Rana macrodactyla 9JD&!P T3 Rana milleti 9]!Q`J, Rana mortenseni 9LL&`J, F&G Rhacophoridae Chirixarus nongkhorensis :Q3'SFDQA^4@ Chirixarus hansenae :Q3'SFGBTBQ)Q Polypedates leucomystax :Q39Q8 Rhacophorus bisacculus :Q3DQA]DLOLTIQ8 Rhacophorus bipunctatus :Q34T8]JDVL&
^DO'36P QB X:FS7Q8]=VL`)' Q^8 )8S3!L&DX LL3`8= V86T a33 &4P La:8T
FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 68
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1. . DX LL3@ T_#B&IB Q&:Q 6T:BO L93FA'&LA:Q ^DO^5F>P8...... 2 !. _#B&IB Q&:Q a@ @T'&LA:Q ^DO^5F>P8 LQ'@TD P H2O]:'86 L]:S3LL 3Q8J8 Q JBVL@TBS@
^JF8...... ]!TA3'O8Q (Occidozyga lima) !. #BT9JQ&3Q898@ T!8Q3` D]# TA& 9#BP T9JQ&3Q8D Q& BL9) L&:Q ]:'8^;8J8 &BP X:] VL @Q 2 L8*P L8 8P .]!TA3JD&:P W@ (Occidozyga martensii) 4. . #BT9JQ&98]BS@48'Q 6QA4 FP ...... 5 !. #BT9JQ&98a@ a3]B S@48'Q 6QA4 FP ...... 12 . . BX 8]: S3LL `8^8F DQ&4FP ...... 6 !. BX 8]: S3LL 3Q8!FQ!L&4 FP ...... 9 6. . BX 8@ TD P H2O]:'86 L...... 7 !. BX 8@ TD P H2O]:'8) L&]:S3 DQ4 F^DOJQ&@P TIT3Q ...... #Q&# 9Q8 (Bufo melanostictus) 7. . '@X LA` X D:Q @Q FQ4Q ^;8J8 &BL9)P L&:Q @TBLAJA P ...... 8 !. '@X LA` X D4Q@Q FQ:Q ^;8J8 &BL9)P L&:Q 3Q8D Q&]BTA98X8...... 98Q (Hoplobatrachus rugulosus) 8. . ]@VLJ&QADX L L3!U8IQ@QB5@L&]J'8DQaI 'Q ?QA8L 3Q8D Q&!L&) L&:Q @T^;8J8 &6P T@TBLAJA P ^DO4W @J8&P - ^5F ]!TA8IX4B>P8a3]: '8 I : 5+5 // -+- : II...... :Q34T8]JDVL& (Rhacophorus bipunctatus) !. ]@VLJ&QADX L L3!U8a@ IQ@QB5@L&]J'8DQaI 'Q ?QA8L 3Q8D Q&!L&) L&:Q @T^;8J8 &6P T@TBLAJA P ^DO4W @J8&P 2 ^5F ]!TA8IX4B>P8a3]: '8 I : 0+0 // -+- : II...... :Q39Q8 (Polypedates leucomystax) 9. . ^;8J8 &BL9)P L&:Q 3Q8D Q&]:'8BLAJA P ^4 a@ A8AQFLLV ...... 10 !. ^;8J8 &BL9)P L&:Q 3Q8D Q&A8AQFLLV #DQAB X:8SF...... 11
FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 69
10. . ^;8J8 &BL9)P L&:Q !AQALL ]:'8^;8 FQ& '&LA:Q 98^DOD Q&]:'8BX: V ]!TA8IX4B>P8a3]: '8 II : 5+5 // -+- : II...... 9L J8Q@ (Paa fasiculispina) !. ^;8J8 &BL9)P L&:Q !AQALL ]D ' 8LA '&LA:Q 984B& '&LA:Q DQ&]: '8BX: V ]!TA8 IX4B>P8a3]: '8 I : 0+0 // -+- : II...... :Q3DQA]DLOLTIQ8 (Rhacophorus bisacculus) 11. . >P86W ^5F@T#FQ@AQFa@ ] S8'&LA:Q ]!TA8IX4B>P8a3]: '8 I // -+- : II...... 9]!Q`J, (Rana milletti) !. >P86W ^5F@T#FQ@AQF] S8'&LA:Q ]!TA8IX4B>P8a3]: '8 I : 1+1 // -+- : II...... 9LL&`J, (Rana mortenseni) 1. . #BT9JQ&98]BS@48]! Q@Q`8I F8JQ&...... 13 !. #BT9JQ&98]BS@48]! Q@Q`8I F8DQ4 FP ...... 14 13. . DX LL3@ TIT8Q4QD^3& ]@VLJ&QADX L L3!U8IQ@QB5@L&]J'8DQaI 'Q ?QA8L ...... #Q&# ^#BO (Bufo parvus) !. DX LL3@ TIT3QJB VL8Q4QD^3& ]@VLJ&QADX L L3!U8a@ IQ@QB5@L&]J'8DQaI 'Q ?QA8L ..#Q&# JFBQ9P (Bufo macrotis) 14. . 3Q8D Q&!L&) L&:Q 4B& DQ&a@ @T^;8J8 &P ...... 15 !. 3Q8D Q&!L&) L&:Q @T^;8J8 &_3A4DL3P ...... 17 1. . ]!TA8IX4B>P8a3]: '8 I : 1+1 // -+- : II...... 16 !. ]!TA8IX4B>P8a3]: '8 I : 3+3 // -+- : II...... :Q3'SFGBTBQ)Q (Chirixalus hansenae) 16. . ^5F>P8DQ&6 &J@3@P T!8Q3` D]# TA& 8P ...... :Q3'SFDQA^4@ (Chirixalus nongkhorensis) !. ^5F>P8DQ&^5FI W36QAAQF]: '8#BU&J8U&!L&^5FLV8... 9J8L& (Fejervarya limnocharis) 17. . ]!TA8IX4B>P8a3]: '8 I : // -+- : II...... 18 !. ]!TA8IX4B>P8a3]: '8I :1+1 // -+- : II...... 9JD&a=DP (Rana lateralis) 18. . DQ4 F]=BP TAF I F86T FQ&6 TIW3!L&#BT9JQ&LA:BO@Q2 X U& DQ&#FQ@AQFJQ&...... 9JD&!P T3 (Rana macrodactyla) !. DQ4 F:P L@ I F86T FQ&6 TIW3!L&#BT9JQ&LA X :BO@Q2 2// !L&#FQ@AQFJQ&...... ]!TA3'S (Rana erythraea) 19 . DX LL3@ TDQ4 F]=BP TAF 6 L]:S3JL&]J& VL ]:S3LL DQ&4F3P Q8* QA BS@ FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 70 21 . 6 L]:S3]J&VL ]:S3LL L85U&BX 8]D ' 8LA BX 8]: '86 L`II8cP !8Q3]D ' ...... LU&L Q&9Q8 (Kaloula pulchra) !. ) L&]:S3]J&VL ]:S3LL `84Q^J8 &)S3 9BP X 8 BX 8]: '86 L`I A8AQFLLV 'Q DQ4 FP ...... LU&L Q& 8! T3 (Kaloula mediolineata) 22 . L4BQIP F8DQ4 F4P LJQ&:BO@Q2 - : 3 I F8...... 23 !. L4BQIP F8DQ4 F^DOJQ&@QP FQ - : 3 I F8...... 25 23 . I F8JF@P TD P H2O# L8!Q& D@...... 24 !. I F8JF@P TD P H2O# L8!Q&B T ]@VL@L&'Q 3Q8! Q&I F8#Q&])S3!U8LAQ&) 3]'8P ...... LU&!Q#Q (Microhyla pulchra) 24 . 6 L]:S3JL&]J& VL LA# X L8@Q6QA4 FP #BT9JQ&98]BS@]!Q@Q`8I F8JQ& BX 8_# &@Q3 Q8J8 Q ...... LU&^3& (Calluella guttulata) !. 6 L]:S3JL&]J& VL LA# X L8@Q6QA4 FP #BT9JQ&98]BS@]!Q@Q`8I F8DQ4 FP BX 8]: S3D&@Q4B&c JBVL_#&@Q3 Q8JD &P ...... LU&];Q (Glyphogolssus molossus) 25 . I F8JF#P L8!Q& D@ a@ @T^59IT]&S8=Q3BOJFQ&4Q ...... 26 !. I F8JFBP X:BT @T^59IT]&S8=Q3BOJFQ&4Q BS@ FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 71 F&G Megophryidae L&U BQA'P869XB2 Xenophrys lekaguli (25 - /0 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJFQ&#FQ@AQFD Q4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 F]BP TAFBX:a! IT8Q4QD ^3& `46 L&`I @T'W3:BOIT!QFa@ ]:'8BO]9TA9 BO'QA4 L]8VL&a:5U&I F8JQ& BX'@X LA` X D:DQA:Q @Q FQ4Q ) L&]:S3]J&VL FQ&4F`84P Q^J8 & DQ&4FP LA` X D:Q @Q F Q_#8JQ& _3A]:S3LL !Q& 4F3P Q8* QA:DQA6 L](TA&!U8]D ' 8LA a@ ]J'8DQaI 'Q ?QA8L BX 8]: '86 L`I]:S3LL 3Q86 QA`8 ^8F DQ&4FP I F8JQ&@TIT8Q4QD^3&L@]6Q@ T'W3IT!QF BO'QA_3A4DL3 #BT9JQ&3Q898I T8Q4QD ^3& #BT9JQ&]D' DQ@]8 VLJQ&!8Q3`J, #BT9JQ&3Q898^DO3 Q8D Q&]BS@53]!P Q@Q`8I F8JQ& ^DO@T!8Q3` D]# TA& 8P I F86T FQ&6 TIW3LA:BO@Q2#B X U&J8U&!L&#FQ@AQFJQ& (?Q=6T 1A) _#B&IB Q&:Q :Q LAXJ8 QI W3!L&JF@P TD P H2O]:'8 BFABX:IT]JDTA@!8@]:TA :X8]:S3LL 3Q898 a@ @T ^5F>P8^DO'&LA:Q ^; 8J8P&BL9) L&:Q @T4W @]D' c ]BTA&]:'8BO]9TA9:BO@Q2 2 - / ^5F (?Q=6T 1B) F&G Bufonidae #Q&# JPFBQ9 Bufo macrotis (15 – 20 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 1 DQ4 F:P L@BX:BT 6&DP Q4 FP ^DOJQ&@TIT3QJB VL8Q4QD ]@VLJ&QADX L L3!U8@L&]J'8DQaI a3'Q ?QA8L BX'@X LA3 X Q898` D 4Q@Q F Q:DQAIW3!L&JPF ) L&]:S 3]J&VL LA X U& DQ&BOJF Q&:Q ^DO_#8JQ& ]:S 3LL !Q&4 FP 3Q8* QA_3AFQ&](TA&!U8:BO@Q2 05 L&GQ # L8a:6Q&3Q8D Q&!L&DQ4 FP BX 8]: '86 LIP8c ]:S 3 LL DQ&DQ4 FP _3AI F898!L&6 L4S3 9#BP T9JQ&D Q& #BT9JQ&IT!QF! 8W JQ& D@ FQ& :DQAJQ& ]BTAF #BT9JQ&3Q898^DO3 Q8D Q&]BS@48]! Q@Q`8I F8JQ& #BT9JQ&3 Q898 FQ& F Q#BT9JQ& 3Q8D Q&]D ' 8LA I F86T FQ&6 TIW3LA:BO@Q2 X U& DQ&JQ& (?Q=6T 2A) _#B&IB Q&:Q :Q LA X3Q8D Q&!L&JPF BS@ FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 7 #Q&# 9 Q8 Bufo melanostictus (15 – 20 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 1 DQ4 F:P L@BX:BT :DQAJFP IL9]!Q]D ' 8 LA 6&DP Q4 F^DOJQ&@P TIT3Q ]@VLJ&QADX L L3!U8`46 L&`I^4 @L&a@ ]J'8DQaI 'Q ?QA8L BX'@X LA X3Q898` D4Q@Q F Q:DQAIW3!L&JFP ) L&]:S3]J&VL LA X U& DQ&BOJF Q&:Q ^DO_#8JQ& ]:S 3LL !Q&4 F3P Q8* QA_3AFQ&](TA&!U8:BO@Q2 05 L&GQ 4Q^J8 &# L8a:6Q& 3Q8D Q&!L&DQ4 FP BX 8]: '8) L&]:S3LL DQ&DQ4 FP _3AI F8984S3 9#BP T9JQ&D Q& #BT9JQ&IT!QF ! 8W @T'W3IT]6Q]D' c BO'QA6F#BP T9JQ& JQ&]BTAF :DQAJQ&]BTAF@8 #BT9JQ&3Q898^DO3 Q8D Q& ]BS@48'Q I F86QA!L&D Q4 FP _3A#BT9JQ&3Q898 FQ& F Q#BT9JQ&3Q8D Q&]D ' 8LA I F86T FQ& 6TIW3LA:BO@Q2 X U& DQ&JQ& (?Q=6T 3A) _#B&IB Q&:Q :Q LA3 X Q8D Q&!L&JFP BS@ #Q&# ^#BO Bufo parvus (15 - 20 @SDDS]@4B) D P H2ODX L L3 DQ4 F:P L@BX:BT 6&DP Q4 F^DOJQ&@P TIT8Q4QD]! @ ]@VLJ&QADX L L3!U8IQ@QB5@L&]J'8DQaI 'Q ?QA8L DQaI LA# X L8a:3Q8* QA!L& BX'@X LA X3Q898` D4Q@Q F Q:DQAIW3!L&JPF ) L& ]:S3]J&VL LA X U& DQ&BOJFQ&:Q ^DO_#8JQ& ]:S3LL !Q&D Q4 F3P Q8* QA_3AFQ&]( TA&!U8:BO@Q2 05 L&GQ 6Q&3Q8D Q&!L&DQ4 FP BX 8]: '86 L`IIP8c ]:S3LL DQ&DQ4 FP I F898!L&6 L4S3 9#BP T9 JQ&3Q8D Q& #BT9JQ&IT8Q4QD^3& JQ&]BTAF #BT9JQ&!8Q3]D ' :DQAJQ&_#&@8 #BT9JQ&3Q898 ^DO3Q8D Q&]BS@48]! Q@Q`8I F8JQ& @T!8Q3` D]# TA& 8P _3A#BT9JQ&3Q898 FQ& F Q#BT9JQ& 3Q8D Q&]D ' 8LA I F86T FQ&6 TIW3LA:BO@Q2 X - `8 0 !L&JQ& (?Q=6T 4A) _#B&IB Q&:Q :Q LA3 X Q8D Q&!L&JFP BS@ FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 73 F&G Dicroglossidae 9J8L& Fejervarya limnocharis (/0 - 00 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 F:P L@BX:a! IT 8Q4QD @T'W3IT]!@ BO'QA6FDP Q4 F^DOJQ&P :DQAJQ&LQ'@TITI @JBVL@T^593Q ]@VLJ&QADX L L3!U8 IQ@QB5@L&]J'8DQaI 'Q ?QA8L DQaI LA X DQ&DQ4 FP BX'@X LA3 X Q8! Q& U& DQ&BOJF Q&:DQAIW3 !L&JFP 94QP ) L&]:S3]J&VL LA` X D:Q @Q F Q_#8JQ& ]:S3LL !Q&D Q4 F3P Q8* QA_3AFQ&](TA& !U8:BO@Q2 05 L&GQ 4Q^J8 &# L8a:6Q&3 Q8D Q&!L&DQ4 FP BX 8]: '86 L`I]:S 3LL DQ&4FP 3Q8984 S3 9#BP T9JQ& #BT9JQ&IT!QF! 8@W T'W3IT]!@ BO'QA6FP :DQAJQ&]BTAF@8 #BT9JQ&3Q898 ^DO3Q8D Q&'Q ]BS@48I F86QA!L&D Q4 FP _3A#BT9JQ&3Q898 FQ& FQ#B T9JQ&3Q8D Q&]D ' 8LA I F86T FQ&6 TIW3!L&#BT9JQ&LA:BO@Q2 X U& DQ&JQ& (?Q=6T 5A) _#B&IB Q&:Q :Q LA3 X Q8J8 Q# L8a:3Q8D Q&!L&JFP BS@ 98Q Hoplobatrachus rugulosus (00 - 50 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 FLP F8: L@IT]JDVL& JBVL8Q4QD `46 L&I T!QF@L&a@ ]J'8DQaI 'Q ?QA8L BX'@X LA X3Q898` D4Q@Q F Q:DQAIW3 !L&JPF^DO@T'W3IT3QLA X` DB X'@X ) L&]:S3]J&VL LA X`84Q^J8 & U& DQ&DQ4 FP ]:S 3LL !Q&D Q4 FP 3Q8* QA _3AFQ&](TA&!U8 05 L&GQ # L8a:6Q&3Q8D Q&!L&DQ4 FP BX 8]: '86 L`I]:S3LL 3Q86 QA `8^8F DQ&DQ4 FP I F898!L&6 L4S3 9#BP T9JQ& JQ&@TD P H2O]BTAF FQ& FQ#FQ@ FQ&!L&D Q4 FP ]D ' 8LA I F8JQ&@TIT]JDVL&]) 8]3TAF 9DP Q4 FP LQ'@T'W3JBVL:V8IT3Q :DQAJQ&]BTAF^JD@ #BT9JQ& 3Q898^DO3 Q8D Q&]BS@'Q I F86QA!L&4 FP @T!8Q3` D]# TA& 8_3A#BP T9JQ&3Q898 FQ& FQ#B T9 JQ&3Q8D Q&]D ' 8LA I F86T FQ&6 TIW3LA# X L8a:6Q&:DQAJQ& (?Q=6T 6A) _#B&IB Q&:Q :Q LA X:DQAIW3!L&JPF^DO]:S3LL 6Q&3Q8D Q& ^; 8J8&BL9)P L&:Q 6&98^DODP Q&@T !8Q3`J, @TBLAJA P 6T!L9 6T@W@:Q a@ @T4W @J8&P ) L&:Q @T!8Q3`J, ]@VL]6TA9 9DP Q4 FP '&LA FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 74 :Q IT3Q!8Q3`J, @L&]J'8a3'Q ?QA8L !L9'&LA:Q 6&98^DODP Q&@TBLAJA P D P H2O'&LA :Q ]:'8BX:*S ^* =L3T 8P *T>P8J Q&c BX:JA38Q ]!TA8IX4B>P8a3]: '8 I : 3+3 // 2+2 : I (?Q=6T 6B) ]!A3'O8QT Occidozyga lima (/0 - 00 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 1 : 3 DQ4 F]BP TAFAQF :DQA JF^JD@P 6L&I T!QF@L&a@ ]J'8DQaI 'Q ?QA8L BX'@X ]:S3LL 3Q898 LABOJF X Q&:DQA:Q 9P 4Q ) L&]:S 3]J&VL LA X` D_#8JQ&@Q F Q:Q ]:S 3LL !Q&4 F3P Q8* QA:DQA6 L)Ta:3 Q8JD&P 4Q^J8 &#L8a:6Q&3 Q8D Q&!L&DQ4 FP BX 8]: '8) L&]:S3]D ' c ]:S3LL 3Q86 QA`8^8F DQ&4F^DOP a@ @TI F8`34S3 P9#BT9JQ& I F8JQ&IT8Q4QD^3& :DQAJQ&]BTAF^JD@ #BT9JQ&3 Q898@T 'W3]BS@48]! Q@Q`8D Q4 F]DP ' 8 LA^DO!AQALL LAQ&BF3]B 'F]:'8^; 8 FQ&#D QA BO_3& #BT9JQ& 3Q8D Q&@T!8Q3]D ' ]BS@'Q ) L&]:S3 8^DO]B TAFAQFa:'8IW3:DQAJQ& (?Q=6T 7A) _#B&IB Q&:Q :Q LA X3Q8J8 Q:DQAIW3!L&JPF _#B&IB Q&:Q D P H2O]:'86 L BL9) L&:Q ]:'8^; 8 J8&BP X:] VL @QA V3]!QLL a3#D QAD X IX9 3Q88L ]:'8^;8J8 &8P X8BX:F&^JF8 ) L&:Q ]:'8) L& ]:S3 D@@T!8Q3]D ' ]@VL]6TA9 9DP Q4 FP '&LA:Q IT3Q LA3 X Q8`8) L&:Q !L9'&LA:Q 6&98^DOP D Q&`84Q^J8 &6TI9 8@P TBLAJA P ]D ' c '&LA:Q 98BX:_#&J8Q AQF]DA'&LA:Q D Q&]D ' 8LA^DO LA3 X Q88L '&LA:Q DQ&D P H2O#DQAL P HB U LAD X U ]!Qa:3 Q8`8 a@ @T*T>P8 (?Q=6T 7B) ]!A3JDT &:P W@ Occidozyga martensii (/0 - 00 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 1 : 3 DQ4 F]BP TAFAQF :DQA JF^JD@P @T^59IT3Q#Q3; Q84Q^D F# LAc 'Q&JQAa: !L9:Q ^DO!Q&D Q4 F@P T^59IT!QF FQ& 6L& IT!QF@L&a@ ]J'8DQaI 'Q ?QA8L BX'@X LA` X D4Q@Q F Q:DQA:Q ) L&]:S3]J&VL LA` X D_#8 JQ&@Q FQ:Q 6 L]:S3LL !Q&D Q4 F3P Q8* QA :DQA6 L)Ta:3Q8JD &P 4Q^J8 &# L8a:6Q&3Q8D Q& !L&DQ4 FP BX 8]: '8) L&]:S3]D' c ]:S3LL 3Q86 QA`8^8F DQ&4F^DOa@P @TI F8`34S3 9#BP T9JQ& I F8JQ&IT8Q4QD^3& :DQAJQ&]BTAF^JD@ #BT9JQ&3Q898^DOD Q&!8Q3]D ' @Q #BT9JQ&3Q898 @T'W3]BS@485 3]!P Q@Q`8I F8JQ&]D ' 8LA I F86T FQ&6 TIW3LA# X L8a:6Q&:DQAJQ& (?Q=6T 8A) FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 7 _#B&IB Q&:Q :Q LA X3Q8J8 Q:DQAIW3!L&JPF _#B&IB Q&:Q D P H2O]:'86 L BL9) L&:Q ]:'8^; 8 J8&BP X:] VL @Q 2 L8*P L8 8P ) L&:Q ]:'8) L&]:S3 D@@T!8Q3]D ' ]@VL]6TA9 9DP Q4 FP '&LA:Q IT 3Q LA3 X Q8`8) L&:Q !L9'&LA:Q 6&98^DODP Q&`84Q^J8 &6TI9 8@P TBLAJA P ]D ' c '&LA:Q 98BX:_#&J8Q AQF]DA'&LA:Q D Q&]D ' 8LA^DOLA 3 X Q88L '&LA:Q D Q&D P H2O#DQAL P HB U ^DOLAD X U ]!Qa:3 Q8`8 a@ @T*T>P8 (?Q=6T 8B) 9L J8Q@ Paa fasiculispina (00 - 50 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 F]BP TAFBX:a! IT#BT@ `4 6 L&`I @T'W3IT!QFDO]LTA3 BO'QA6F4DL3DP Q4 F`4P 6 L&^DO_#8JQ& ]@VLJ&QADX L L3!U8 IQ@QB5@L&]J'8DQaI 'Q ?QA8L DQaI # L8a:6Q&3Q8* QA!L&D Q4 FBP X'@X LA3X Q898` D:DQA IW3!L&JPF@Q F Q4Q ) L&]:S3]J&VL LA` X D:Q @Q F Q_#8JQ& ]:S3LL !Q&D Q4 F3P Q8* QA _3A FQ&](TA&!U8 05 L&GQ 4Q^J8 &# L8a:6Q&3 Q8D Q&!L&DQ4 FP BX 8]: '86 L`IBX: BFA]:S 3LL 3Q8!FQ!L&#B T9JQ& I F8JQ&IT!QF! 8W @T'W3 D@IT3Q!8Q3]D ' 6T_#8JQ&^DF# LAc !AQALL ]:'8 'W3 D@!8Q3`J, ) F& DQ&!L&JQ&^DO'W3 D@3&P D QF'O# LAc 'Q&D&]@VL5U&:DQAJQ& :DQAJQ& ]BTAF@8 #BT9JQ&3Q898^DO3 Q8D Q&]BS@'Q I F86QA!L&D Q4 FP @T!8Q3` D]# TA& 8_3A#BP T9JQ& 3Q898 FQ& FQ#B T9JQ&3Q8D Q&]D ' 8LA I F86T FQ&6 TIW3LA# X L8a:6Q&:DQAJQ& (?Q=6T 9A) _#B&IB Q&:Q :Q LA3 X Q8D Q&!L&JFP BL9) L&:Q @T^;8J8 &!8Q3`J,P BS@ F&G Ranidae ]!A3'T S Rana erythraea (00 - 50 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2.5 I F8JPF# L8!Q& ^JD@ IT8Q4QD `46 L&`I @T'W3:BOIT!QF BO'QALA XJ Q&c ^DO4 L]8VL&a:5U&I F8JQ& ]@VLJ&QA DX L L3!U8IQ@QB5@L&]J'8DQaI 'Q ?QA8L DQaI !3LA X DQ&4FP BX'@X ` D:DQAI W3!L&JPF FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 76 @Q F Q4Q ) L&]:S3]J&VL LA` X D_#8JQ&@Q F Q:Q 6 L]:S3LL !Q&D Q4 F3P Q8* QA`84Q^J8 & DQ&DQ4 FP _3AFQ&](TA&!U88 LA F Q 05 L&GQ BX 8]: '86 L`IBX: BFAIP8c ]:S3LL 3Q8!FQ!L& DQ4 F^DO]:P '8I F8J8U&!L&#BT9JQ& I F8 DQ@]8 VL^DO#BT9JQ&IT8Q4QD@ T'W3DO]LTA3IT!QF6T BO'QA4 L]8VL&@Q'Q 6L& :DQAJQ&^JD@ #BT9JQ&98]BS@]!Q@Q`8I F8DQ4 FP #BT9JQ&D Q&]BS@ 'Q I F86QA!L&4 FP #BT9JQ&3Q898 FQ& FQ#B T9JQ&3Q8D Q&]D ' 8LA I F86T FQ&6 TIW3LA# X L8a: 6Q&_#8 (?Q=6T 10A) _#B&IB Q&:Q :Q LA X3Q8D Q&!L&JPF BS@ 9JD&a=DP Rana lateralis (40 - 50 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJFQ&#FQ@AQFD Q4 F4P L#FQ@AQFJQ&:BO@Q2 - : 1 DQ4 F3P Q898# L8!Q& BQ93X#DQA]: '8]JDTA@]D ' 8LA DQ4 FIP T8Q4QD@ T'W3:BODO]LTA3IT]6Q BO'QA6F4DL3DP Q4 F^DOJQ&P `46 L&I T!QF ]@VLJ&QADX L L3!U8'Oa@ ]J'8DQaI 'Q ?QA8L BX'@X LA# X L8@Q6Q&:DQAIW3!L&JFP ) L&]:S 3]J&VL LA X` D:Q @Q F Q_#8JQ& ]:S 3LL !Q&D Q4 F3P Q8* QA:DQA6 L)Ta:3Q8JD &P 4Q^J8 &# L8a:6Q&3Q8D Q&!L&DQ4 FP BX 8]: '86 L`IBX: BFAIP8c]:S3LL 3Q86 QA`8^8F DQ& 4FP 3Q898!L&6 L4S3 9#BP T9JQ&] VL94DL3#FQ@AQF :DQAJQ&]BTAF@8 #BT9JQ&3Q898]B S@48 ]!Q@Q`8D Q4 FP 3Q8D Q&]BS@'Q I F86QA!L&D Q4 F^DO@P T!8Q3` D]# TA& 8_3A#BP T9JQ&3Q898 FQ& FQ#B T9JQ&3Q8D Q& I F86T FQ&6 TIW3LA# X L8a:6Q&:DQAJQ& (?Q=6T 11A) _#B&IB Q&:Q :Q LA X3Q8J8 Q# L8a:3Q8D Q&!L&JPF BS@ FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 77 9JD&!P 3T Rana macrodactyla (30 - 40 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2.5 I F8JF^DODP Q4 FBP X:BT IT]JDVL&JBVL8Q4QD @T^59IT8Q4QD#Q3; Q84Q ]@VL@L&'Q 3Q898@ T^59IT8Q4QD FQ&#D QAL P HB Y =Q3'Q BX'@X 6&IL&!P Q&@Q5 U& DQ&DQ4 FP `46 L&`I @T'W3:BOIT!QF BO'QALAJ X Q&c^DO4 L]8VL& a:5U&I F8JQ& ]@VLJ&QADX L L3!U8IQ@QB5@L&]J'8DQaI 'Q ?QA8L DQaI LA X# L8a:3Q8* QA BX'@X LAX3 Q898` D:DQAI W3!L&JPF@Q F Q4Q ) L&]:S3]J&VL LA X U& DQ&BOJF Q&:Q ^DO_#8 JQ& ]:S3LL !Q&4 F3P Q8* QA_3AFQ&4F](P TA&!U88LA F Q 05 L&GQ 6Q&3Q8D Q&!L&DQ4 FP BX 8]: '8 6 L`IIP8c]:S3LL 3Q86 QA`8^8F DQ&4F^DO]:P '8I F8J8U&!L&#BT9JQ& I F8JQ&# L8!Q&`I @T 'W3IT!QF^DO8Q4QD BO'QALA#D X QA^594Q@!FQ& JQ&@TD P H2O]=BTAF :DQAJQ&^JD@ #BT9JQ& 3Q898^DO3 Q8D Q&]BS@'Q I F86QA!L&4 F^DO@P T!8Q3` D]# TA& 8_3A#BP T9JQ&3Q898 FQ& F Q #BT9JQ&3Q8D Q&]D ' 8LA I F86T FQ&6 TIW3LA:BO@Q2 X U& DQ&JQ& (?Q=6T 12A) _#B&IB Q&:Q :Q LA3 X Q8D Q&!L&JFP BS@ 9L L& Rana mortenseni (30 - 40 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2.5 DQ4 F:P L@]:'8IP8 IT 8Q4QD @T'W3IT8Q4QD]! @^DO!QF BO'QA6FP `46 L&`I DQaI LA X DQ&DQ4 FP BX'@X LA X3Q898` D :DQAIW3!L&JF@QP F Q4Q ) L&]:S3]J&VL LABOJF X Q&:Q ^DO_#8JQ& ]:S3LL !Q&4 F3P Q8* QA_3A FQ&](TA&!U88LA F Q 05 L&GQ 4Q^J8 &# L8a:6Q&3Q8D Q&!L&DQ4 FP BX 8]: '86 L`I!8Q3`J, ]:S 3LL 3Q8!FQ!L&#B T9JQ& I F8JQ&`I @TDQA'W3]D' IT8Q4QD]! @ BO'QA6FP :DQAJQ&]BTAF@8 #BT9JQ&3Q898]B S@48]! Q@Q`8I F8JQ& #BT9JQ&3Q8D Q&]BS@'Q I F86QA!L&4 F_3A#BP T9JQ& 3Q898 FQ& FQ#B T9JQ&3Q8D Q&]D ' 8LA I F86T FQ&6 TIW3LA# X L8a:6Q&:DQAJQ& (?Q=6T 13A) FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 78 _#B&IB Q&:Q :Q LA3 X Q8J8 Q# L8a:3Q8D Q&!L&JFP BS@ 9]!Q`J, Rana milletti (30 - 40 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 F:P L@ IT8Q4QD @T ^59IT]!@#Q3; Q84Q `46 L&`I @T'W3:BOIT!QF BO'QALAJ X Q&c ^DO4 L]8VL&a:5U&I F8JQ& ]@VLJ&QA DX L L3!U8IQ@QB5@L&]J'8DQaI 'Q ?QA8L DQaI LA# X L8a:3Q8* QA!L&DQ4 FP BX'@X LA3 X Q898 ` D4Q@Q F Q:DQAIW3!L&JPF ) L&]:S 3]J&VL LA X` D_#8JQ&@Q F Q:Q ]:S 3LL DQ&DQ4 FP 3Q8* QA_3AFQ&]( TA&!U8:BO@Q2 05 L&GQ BX 8]: S3LL 3Q8!FQ!L&D Q4 F^DO]:P '8I F8J8U&!L&#BT9 JQ& DQ@]8 VL#BT9JQ&IT8Q4QD I F8JQ&@T'W3IT!QF^DO8Q4QD BO'QALAJ8Q^8 X 8 :DQAJQ&^JD@ #BT9JQ&3Q898^DO3 Q8D Q&]BS@'Q I F86QA!L&D Q4 F^DO@P T!8Q3` D]# TA& 8_3A#BP T9JQ&3Q898 FQ& FQ#B T9JQ&3Q8D Q&]D ' 8LA I F86T FQ&6 TIW3LA:BO@Q2 X U& DQ&!L&JQ& (?Q=6T 14A) _#B&IB Q&:Q :Q LA X3Q8J8 Q# L8@Q6Q&3Q8D Q&!L&JPF BS@ F&G Rhacophoridae :Q3'SFGBTBQ)Q Chirixalus hansenae (25- 30 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJFQ&#FQ@AQFD Q4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 FP : L@BX:a! IT]JDVL& 9BS]F2BX'@X @T^4@I T3Q D L&I@L&@TIT]!@B X:IT]JDTA@!8@]:TA :X84 L3FA]I 84B&@Q5U&_#8JQ& `46 L&`I ]@VLJ&QADX L L3!U8IQ@QB5@L&]J'8DQaI 'Q ?QA8L DQaI LA X# L8a:3Q8* QA!L& DQ4 FP BX'@X LA3 X Q898` D:DQAI W3!L&JF@QP F Q4Q ) L&]:S3]J&VL LABOJF X Q&:Q ^DO_#8JQ& FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 79 ]:S 3LL !Q&D Q4 F3P Q8* QA _3AFQ&](TA&!U8:BO@Q2 05 L&GQ 4Q^J8 &# L8a:6Q&3Q8D Q&!L& DQ4 FP BX 8]: '86 L`I6L3; Q8#BT9JQ&D Q&^DO]:S3LL `8^8F3S&J Q&'Q _#8JQ&]D' 8 LA I F8 JQ&IT!QF#L8! Q&`I @T'W3IT3Q]D ' c BO'QA6F_3A](=QO9BP S]F2 DQ@]8 VLJQ& JQ&]BTAF :DQAJQ& ]BTAF@8 #BT9JQ&3Q898 FQ& F Q#BT9JQ&3Q8D Q&]D ' 8LA _3A#BT9JQ&3Q898]B S@48]! Q@Q`8 I F8DQ4 F]DP ' 8 LA]@VL=8) F&DQ4 Fa:'O!AQALLP LA Q&BF3]B'FI F86T FQ&6 TIW3LA X:BO@Q2 U& DQ&JQ& (?Q=6T 15A) _#B&IB Q&:Q :Q LA3 X Q8J8 Q# L8a:3Q8D Q&!L&JFP BS@ :Q3'SFDQA^4 @ Chirixalus nongkhorensis (25 - 30 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJFQ&#FQ@AQFD Q4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 F:P L@BX:a! IT8Q4QD @T'W3IT8Q4QD BO'QA6FP D L&I@L&@TIT]!@#D QAL P HB Y :BQ .'Q JD&4Q5P U&] VL96QA4 FP `4 6L&I T!QF ]@VLJ&QADX L L3!U8IQ@QB5@L&]J'8DQaI 'Q ?QA8L DQaI LA X DQ&DQ4 FP BX'@X LA X 3Q898` D:DQAI W3!L&JPF@Q F Q4Q ) L&]:S3]J&VL LA XBOJF Q&:Q ^DO_#8JQ& ]:S3LL !Q& DQ4 F3P Q8* QA _3AFQ&](TA&!U8:BO@Q2 05 L&GQ 4Q^J8 &# L8a:6Q&3Q8D Q&!L&DQ4 FP BX 8]: '8 6 L`I!8Q3`J, ]:S3LL 3Q86 QA`8^8F DQ&DQ4 F_3A3P Q8984 S3 9#BP T9JQ& I F8JQ&IT!QF # L8!Q&`I @TDQA'W3IT8Q4QD]D ' c BO'QA6F]J@P VL8 9IP F8DQ4 FP JQ&@TD P H2O]BTAF :DQAJQ& ]BTAF^JD@ #BT9JQ&3Q898]B S@48]! Q@Q`8I F8DQ4 F]DP ' 8LA #BT9JQ&3Q8D Q&]BS@'Q I F86QA !L&DQ4 F_3A#BP T9JQ&3Q898 FQ& F Q#BT9JQ&3Q8D Q&]D' 8 LA I F86T FQ&6 TIW3LA X:BO@Q2 U& DQ&JQ& (?Q=6T 16A) _#B&IB Q&:Q :Q LA X3Q8J8 Q# L8a:3Q8D Q&!L&JPF BS@ FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 80 :Q39 Q8 Polypedates leucomystax (40 - 50 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : / DQ4 F:P L@BX:a! IT]JDVL& ) F&6L&IL9]! Q]D ' 8LA :DQA:Q @T'W3IT]&S8]D ' c `46 L&I T!QF a@ ]J'8DQaI 'Q ?QA8L BX'@X LA X 3Q898`84 Q^J8 &` D:DQAI W3!L&JF@QP FQ4Q ) L&]:S3]J&VL LA6Q&3 X Q8D Q&!L&D Q4 FP 6 LLA` X D :Q @Q F Q_#8JQ& :DQA6 L)Ta:3Q8JD &]AP VL&LL 3Q8! Q&]D ' 8 LA BX 8]: '86 L`IBX: BFA]:S 3 LL 3Q86 QA`8^8F DQ&4F^DO]:P '8I F8J8U&!L&#BT9JQ& I F8JQ&IT!QF! 8a@W @TDQA :DQAJQ&]BTAF ^JD@ #BT9JQ&3Q898^DO3 Q8D Q&]BS@'Q I F86QA!L&D Q4 F^DO@P T!8Q3` D]# TA& 8_3A#BP T9JQ& 3Q898 FQ& FQ#B T9JQ&3Q8D Q&]D ' 8LA I F86T FQ&6 TIW3LA:BO@Q2 X U& DQ&!L&JQ& (?Q=6T 17A) _#B&IB Q&:Q :Q LA3 X Q8J8 Q# L8a:3Q8D Q&!L&JFP BS@ :Q348]JDT L&V Rhacophorus bipunctatus (40 - 50 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2.5 DQ4 F:P L@BX:a! IT ]JDVL&I F8JPF# L8!Q&: Q8 `46 L&`I ]@VLJ&QADX L L3!U8IQ@QB5@L&]J'8DQaI 'Q ?QA8L DQaI LAX DQ&DQ4 FP BX'@X LAX3 Q898`84 Q^J8 & U& DQ&BOJF Q&:DQAIW3!L&JPF 94QP ) L&]:S3 ]J&VL LAX` D:Q @Q F Q_#8JQ& ]:S3LL !Q&D Q4 F3P Q8* QA_3AFQ&4F](P TA&!U8@Q F Q 05 L&GQ 4Q^J8 &# L8a:6Q&3Q8D Q&!L&DQ4 FP BX 8]: '86 L`IBX: BFAIP8c ]:S3LL 3Q86 QA`8 ^8F DQ&DQ4 F^DO]:P '8I F8J8U&!L&#BT9JQ& I F8JQ&IT!QF!W 8# L8!Q&`Ia@ @TDQA JQ&@T D P H2O]BTAF :DQAJQ&]BTAF@8 #BT9JQ&3Q898^DO3 Q8D Q&]BS@'Q I F86QA!L&D Q4 F^DO@P T !8Q3` D]# TA& 8P _3A#BT9JQ&3Q898 FQ& F Q#BT9JQ&3Q8D Q&]D' 8 LA I F86T FQ&6 TIW3LA X # L8a:6Q&:DQAJQ& (?Q=6T 18A) _#B&IB Q&:Q :Q LAX3 Q8J8 Q# L8a:3Q8D Q&!L&JPF BS@ FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 81 :Q 6&98^DODP Q&@TBLAJA P ]D' c '&LA:Q 98BX:_#& AQF]DA'&LA:Q D Q&]D' 8LA ]!TA8IX4B >P8a3]: '8 I : 5+5 // -+- : II (?Q=6T 18B) :Q3DQA]DLOLIQ8T Rhacophorus bisacculus (40 - 50 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 F:P L@BX:a! IT8Q4QD JQ&IT8Q4QD]JD VL& `46 L&I T!QF ]@VLJ&QADX L L3!U8IQ@QB5@L&]J'8DQaI 'Q ?QA8L DQaI LA X DQ&DQ4 FP BX'@X LA3 X Q8! Q& U& DQ&BOJFQ&:DQAI W3!L&JFP 94QP ) L&]:S3]J&VL ]:'86 L]D ' IP8AQF :BO@Q2 -`8 -, !L&#FQ@AQFDQ4 F^DO]:P '8I F8J8U&!L&;8&4P FP 6 LLA` X D_#8JQ&@Q F Q:Q ]:S3LL !Q&4 F3P Q8* QA_3AFQ&]( TA&!U8:BO@Q2 05 L&GQ 4Q^J8 &# L8a:6Q&3Q8D Q&!L&DQ4 FP BX 8]: '86 L`I]:S 3LL 3Q8!FQ!L&4 F^DO]:P '8I F8J8U&!L&#BT9JQ& #BT9JQ&IT!QF! W8a@ @TDQA :DQAJQ&]BTAF@8 #BT9JQ&3Q898^DO3 Q8D Q&]BS@48I F86QA!L&D Q4 FP _3A#BT9JQ&3Q898 FQ& FQ#B T9JQ&3Q8D Q&]D ' 8LA I F86T FQ&6 TIW3LA:BO@Q2 X U& DQ&JQ& (?Q=6T 19A) _#B&IB Q&:Q :Q LA3 X Q8J8 Q# L8a:3Q8D Q&!L&JFP BS@ F&G Microhylidae L&^3&U Calluella guttulata (00 - 50 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : / DQ4 FP D@ FQ& `I ]@VL @L&'Q 98=9F QI F86L&#L3# LAc#L3]!QI X6QA4 FP `46 L&`I@L&]J '8DQaI 'Q ?QA8L BX'@X LA3 X Q898` D:DQAI W3!L&JPF@Q F Q4Q BX'@X @T'W3IT3Q D L&I@L&@TIT]!@B X: ^DO@T^59IT ]!@AQF4 L]8VL&@Q'85U&I F86QA!L&D Q4 FP ) L&]:S3]J&VL ]:'86 L`I!8Q3`J, ]:S3LL 3Q86 L& 6QAI W3!L&4FP L85U&_#8JQ&]=TA&]D ' 8LA BX 8]: '86 L`I6L3;Q8#B T9JQ&D Q& :DQA6 L)TD&^DO _#&a:3 Q8J8 Q #BT9JQ&`IJBVLLQ'@TIT3Q 9BS]F2 DQ@]8 VLJQ&3Q898@ P @TIT!QF^DO3Q8D Q&@ P @TIT3Q :DQAJQ&^JD@ #BT9JQ&3Q898@ T'W3]BS@48]! Q@Q`8I F8JQ&]D ' 8LA #BT9JQ&3Q8D Q&]BS@ FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 8 'Q I F86 QA!L&4FP _3A#BT9JQ&3 Q8D Q& FQ& F Q#BT9JQ&3 Q898@Q I F86T FQ&6 TIW3LA X6T 'W3]BS@48!L&#B T9JQ& (?Q=6T 20A) _#B&IB Q&:Q :Q LA:DQAI X W3!L&JF^DO]:P S3LL 6Q&3Q8J8 Q a@ @T4W @J8&BL9)P L&:Q a@ @T*T>P8^DO '&LA:Q BS@ L&];U Q Glyphoglossus molossus (00 - 50 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : / DQ4 FP D@ FQ& `I ]@VL @L&'Q 98=9F QI F86L&#L3]! Q6 86P T `46 L&`I@L&]J '8DQaI 'Q ?QA8L BX'@X LA X3Q898 BOJF Q&4Q^DO:DQAIW3!L&JPF BX'@X @T'W3IT3Q D L&I@L&@TIT]!@B X: ^DO@T^59IT]!@AQF 4 L]8VL&@Q'85U&I F86QA!L&D Q4 FP ) L&]:S3]J&VL ]:'86 L`I!8Q3`J, ]:S3LL 3Q86 L& 6QAI W3 !L&DQ4 FP L85U&_#8JQ&]=TA&]D ' 8LA BX 8]: '86 L`I6L3;Q8#B T9JQ&D Q&:DQA6 L)TD&3Q8D Q& 4&(QP 9#BP T9JQ&JBVLLQ'_#&a:3 Q8JD &P #BT9JQ&`I JBVLLQ'@TIT3Q9B S]F2 DQ@]8 VLJQ& I F8 JQ&]BTAF :DQAJQ&^JD@ #BT9JQ&3Q898@ T'W3]BS@48]! Q@Q`8I F8DQ4 F]DP ' 8LA #BT9JQ&3Q8D Q& ]BS@'Q I F86QA!L&D Q4 FP _3A#BT9JQ&3Q8D Q& FQ& F Q#BT9JQ&3Q898@Q I F86T FQ&6 TIW3LA X 6T'W3]BS@48!L&#B T9JQ& (?Q=6T 21A) _#B&IB Q&:Q :Q LA:DQAI X W3!L&JF^DO]:P S3LL 6Q&3Q8J8 Q a@ @T4W @J8&BL9)P L&:Q a@ @T*T>P8^DO '&LA:Q BS@ L&LU Q& 8!3T Kaloula mediolineata (/0 - 00 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 F:P L@^9898D Q& I F8JFDQ3D&]DP ' 8LA @TIT3QJB VL8Q4QD]! @ @T^59IT8Q4QDL L8=Q3BOJF Q&4Q ]@VL]BS@@T QB]'BS, !L&!QJD&IP F86QA4 F^DO#BP T9JQ&4L848'O]B S@:BQ .!T34Q@AQFIT8QL L8^DO'O)3]'8!P U8]@VL !QJD&]'BP S,]4@6' T `46 L&@ TDQA]DLOIT3Q^DO!QF @L&a@ ]J'8DQaI 'Q ?QA8L BX'@X LA3 X Q898` D 4Q@Q F Q:DQAIW3!L&JFP ) L&]:S3]J&VL ]:'8) L&!8Q3`J, ]:S3LL 3Q86 L&) S3 9BP X 8 BX 8]: '8 6 L`I!8Q3]D' A8AQFLLV 'Q DQ4 F3P Q8984 S3 9#BP T9JQ&4DL3#FQ@AQF #BT9JQ&IT!QF! W8^DO@T :V8IT]6Q LQ'@TIT]JDVL&I @JBVLa@ @T 9BS]F2 DQ@]8 VLJQ&LQ'@T!T34Q@AQFJBVLa@ @T 'a3 I F8JQ& FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 83 ]BTAF :DQAJQ&@8 #BT9JQ&3Q898]B S@485 3]!P Q@Q`8I F8JQ&]D ' 8LA^DO3 Q8D Q&]BS@'Q I F86QA !L&4FP _3A#BT9JQ&3Q8D Q& FQ& FQ#B T9JQ&3Q898]D ' 8LA I F86T FQ&6 TIW3LA# X L8a:6Q&:DQA JQ& (?Q=6T 22A) _#B&IB Q&:Q :Q LA:DQAI X W3!L&JF^DO]:P S3LL 6Q&3Q8J8 Q a@ @T4W @J8&BL9)P L&:Q a@ @T*T>P8^DO '&LA:Q BS@ L&LU Q&9 Q8 Kaloula pulchra (30 - 40 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 F:P L@^9898D Q& I F8JPFDQ3D&]@VL@L&'Q 3Q8! Q& @TIT3QJB VL8Q4QD]! @ @T^59IT8Q4QDL L8=Q3BOJF Q&4Q `4 6L&@ TDQA]DLOIT3Q^DO!QF @L&a@ ]J'8DQaI 'Q ?QA8L BX'@X LA3X Q898` D4Q@Q FQ:DQAI W3 !L&JFP ) L&]:S3]J&VL ]:'8) L&!8Q3`J, ]:S3LL 3Q86 L&`84 Q^J8 & L85U&BX 8]D ' 8LA BX 8 ]:'86 L`IIP8c !8Q3]D ' 3Q8984 S3 9#BP T9JQ&] VL94DL3#FQ@AQF I F8JQ&]BTAF :DQAJQ&@8 #BT9JQ&IT!QF! 8^DO@W T:V8IT]6Q #BT9JQ&3Q898]B S@485 3]!P Q@Q`8I F8JQ&]D ' 8LA^DO3 Q8D Q& ]BS@'Q I F86QA!L&4 FP _3A#BT9JQ&3Q8D Q& FQ& FQ#B T9JQ&3Q898]D ' 8LA I F86T FQ&6 TIW3LA X # L8a:6Q&:DQAJQ& (?Q=6T 23A) _#B&IB Q&:Q :Q LA:DQAI X W3!L&JF^DO]:P S3LL 6Q&3Q8J8 Q a@ @T4W @J8&BL9)P L&:Q a@ @T*T>P8^DO '&LA:Q BS@ L&^@U J8QF Microhyla berdmorei (/0 – /5 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 -: 1.5 DQ4 FP D@: L@`I `46 L&`I@L&]J '8DQaI 'Q ?QA8L !3LA3 X Q8* QA BX'@X LA3 X Q898` D4Q@Q F Q:DQAIW3!L& JFP BX'@X @T'W3IT3Q D L&I@L&IT]!@6 U9BX: ) L&]:S3]J&VL ]:'86 L`I]:S3LL 3Q86 L& # L8 a:6Q&6QA4 FP BX 8]: '86 L`I6L3;Q8#B T9JQ&DQ& :DQA6 L)Ta:3Q8JD &JP Q&'Q _#8JQ&]D ' 8LA #BT9JQ&`I :DQAJQ&^JD@ #BT9JQ&3Q898^DO3 Q8D Q&]BS@'Q I F86QA!L&4 FP _3A#BT9JQ& 3Q8D Q& FQ& FQ#B T9JQ&3Q898@Q I F86T FQ&6 TIW3LA6X T U& DQ&!L&#BT9 (?Q=6T 24A) FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 84 _#B&IB Q&:Q :Q LA:DQAI X W3!L&JF^DO]:P S3LL 6Q&3Q8J8 Q a@ @T4W @J8&BL9)P L&:Q a@ @T*T>P8^DO '&LA:Q BS@ L&DQA]DLOU Microhyla butleri (-5 – 25 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 1.5 DQ4 FP D@`I ]@VL@L& 'Q 3Q8! Q&]J '8I F8#Q&])S3!U8]D' 8 LA `46 L&`I@L&]J '8DQaI 'Q ?QA8L BX'@X LA X` D4Q @Q F Q:DQAIW3!L&JPF BX'@X @T'W3IT3Q D L&I@L&@TIT]!@B X: ) L&]:S3]J&VL ]:'86 L`I!8Q3 `J, ]:S3LL 3Q86 L& 6QAI W3!L&4FP BX 8]: '86 L`I6L3;Q8#B T9JQ&D Q& :DQA6 L)Ta:3Q8JD &P J Q&'Q _#8JQ&]D ' 8LA #BT9JQ&`I:DQAJQ&@ P @TIT^3& I F8JQ&]BTAF :DQAJQ&^JD@ #BT9JQ& 3Q898^DO3 Q8D Q&]BS@'Q I F86QA!L&4 FP _3A#BT9JQ&3Q8D Q& FQ& F Q#BT9JQ&3Q898@Q I F86T FQ&6 TIW3LA6 X T'W3]BS@48!L&#B T9JQ& :DQAJQ&]BTAF]:'8]I 8]D ' (?Q=6T 25A) _#B&IB Q&:Q :Q LA:DQAI X W3!L&JF^DO]:P S3LL 6Q&3Q8J8 Q a@ @T4W @J8&BL9)P L&:Q a@ @T*T>P8^DO '&LA:Q BS@ L&8U Q]4 Q Microhyla fissipes (20 - 30 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJFQ&#FQ@AQFD Q4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 FP D@`I ]@VL@L&'Q 3Q8! Q&]J '8I F8#Q&])S3!U8]D ' 8LA `46 L&`I@L&]J '8DQaI 'Q ?QA8L BX'@X LA` X D4Q@Q F Q :DQAIW3!L&JFP D L&I@L&@TBX: \ / ) L&]:S3]J&VL ]:'86 L`I!8Q3`J, ]:S3LL 3Q86 L&# L8a: 3Q86 QA4 FP L85U&_#8JQ&]=TA&]D' 8LA AQF:BO@Q2 - `8 5 !L&#FQ@AQFDQ4 FP BX 8]: '86 L`I 6L3; Q8#BT9JQ&D Q&:DQA6 L)TD&3Q8D Q&4&(QP 9#BP T9JQ& #BT9JQ&`I :DQAJQ&^JD@ #BT9 JQ&3Q898^DO3 Q8D Q&]BS@'Q I F86QA!L&4 FP _3A#BT9JQ&3Q8D Q& FQ& F Q#BT9JQ&3Q898 @Q I F86T FQ&6 TIW3LA6 X T'W3]BS@48!L&#B T9JQ& (?Q=6T 26A) FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 8 _#B&IB Q&:Q :Q LA:DQAI X W3!L&JF^DO]:P S3LL 6Q&3Q8J8 Q a@ @T4W @J8&BL9)P L&:Q a@ @T*T>P8^DO '&LA:Q BS@ L&!U Q&3Q Microhyla heymonsi (25 - /0 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 2 DQ4 FP D@`I ]@VL@L& 'Q 3Q898@ T]I 8IT3Q]D ' c 4Q@AQF'Q :DQA:Q 5U&_#8JQ& ^DO@T:V8IT]!@6 T4Q^J8 & D L&I@L& BOJF Q&4Q@T^59IT]&S8 `46 L&`I@L&]J '8DQaI IT]&S8 BX'@X LA3 X Q898` D4Q@Q FQ:DQAI W3!L& JPF ) L&]:S3]J&VL ]:'86 L`IIP8c ]:S36QAD Q4 F3P Q86 L& L85U&'W3]BS@48#B T9JQ&D Q& BX 8]: '8 6 L`I6L3;Q8#B T9JQ&D Q& :DQA6 L)TD&J Q&'Q _#8JQ&]D ' 8LA #BT9JQ&`I ^DOLQ'@T^59IT]&S8 - ^59 JBVLa@ @T a3' 9BS]F2 DQ@]8 VLJQ&@TIT]JDVL&L L8 :DQAJQ&^JD@ #BT9JQ&3Q898]B S@48 53]!P Q@Q`8I F8JQ&]D' 8 LA #BT9JQ&3Q8D Q&]BS@'Q I F86QA!L&D Q4 FP _3A#BT9JQ&3Q8D Q& FQ& FQ#B T9JQ&3Q898]D ' 8LA I F86T FQ&6 TIW3LA:BO@Q2 X U& DQ&JQ& (?Q=6T 27A) _#B&IB Q&:Q :Q LA:DQAI X W3!L&JF^DO]:P S3LL 6Q&3Q8J8 Q a@ @T4W @J8&BL9)P L&:Q a@ @T*T>P8^DO '&LA:Q BS@ L&!Q#U Q Microhyla pulchra (30 - 40 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJFQ&#FQ@AQFD Q4 F4P L#FQ@AQFJQ&:BO@Q2 - : 4 DQ4 F`IP ^98`8^8FBQ9 ]@VL@L&'Q 3Q8! Q&]J '8I F8#Q&])S3!U8#DQA:DQ]8 VLL L8 `46 L&`I@L&]J '8DQaI 'Q ?QA8L BX '@X LA X3Q898` D4Q@Q F Q:DQAIW3!L&JFP 6QA4Q@ TDQABX:IQ@]JDTA@#FQ^DO@ T]I 8=Q3; Q8 'Q BX'@X @Q'85U&:DQAIW3!L&JFP ) L&]:S3]J&VL ]:S3LL 3Q86 L&# L8a:3Q86 QAD Q4 FP L85U& BX 8]= TA&]D ' 8LA BX 8B X: BFA]:S3LL `8^8F DQ&DQ4 FP ^DO]:'8I F8J8U&!L&#BT9JQ&D Q& #BT9 JQ&`I^4 @T'W3:BODO]LTA3 BO'QALA6 X FP @T^59IT]JDVL&^DO3Q#Q34 &^4P _#8'85U&:DQAJQ& I F8 JQ&]BTAF :DQAJQ&^JD@ #BT9JQ&3Q898^DO3 Q8D Q&]BS@'Q I F86QA!L&D Q4 FP _3A#BT9JQ& 3Q8D Q& FQ& FQ#B T9JQ&3Q898@Q I F86T FQ&6 TIW3LA6X T'W3]BS@48!L&#B T9JQ& (?Q=6T 28A) FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 86 _#B&IB Q&:Q :Q LA:DQAI X W3!L&JF^DO]:P S3LL 6Q&3Q8J8 Q a@ @T4W @J8&BL9)P L&:Q a@ @T*T>P8^DO '&LA:Q BS@ L&JDU &!P 3T Micryletta inornata ( 20 – 25 @SDDS]@4B) D P H2ODX L L3 L4BQIP F8BOJF Q&#FQ@AQFDQ4 F4P L#FQ@AQFJQ&:BO@Q2 - : 1.5 DQ4 FP D@IT8Q4QD @T ^59ITL L86TJQ& 6L&`I@L&]J '8DQaI 'Q ?QA8L BX'@X LA3 X Q898BOJF Q&4Q^DO:DQAIW3!L& D L&I@L&@TBX:F&]D9' ) L&]:S3]J&VL ]:'86 L`I]:S3LL 3Q86 L&# L8a:3Q86 QA4 FP L85U&_#8 JQ&]=TA&]D ' 8LA BX 8]: '86 L`I6L3;Q8#B T9JQ&D Q&_#&4Q@^8FD Q4 F]:P S3LL 6Q&3Q8J8 Q #BT9 JQ&`I I F8JQ&]BTAF:DQAJQ&^JD@ #BT9JQ&3Q898^DO3 Q8D Q&]BS@]!Q@Q`8I F8DQ4 F]DP ' 8LA@ T !8Q3` D]# TA& 8_3A#BP T9JQ&3Q8D Q& FQ& F Q#BT9JQ&3Q898]D ' 8 LA I F86T FQ&6 TIW3LA X6T 'W3]BS@48!L&#B T9JQ& #BT9JQ&'OIL9]!Q@Q^DO]B TAF]:'8]I 8]D ' c (?Q=6T 29A) _#B&IB Q&:Q :Q LA:DQAI X W3!L&JF^DO]:P S3LL 6Q&3Q8J8 Q a@ @T4W @J8&BL9)P L&:Q a@ @T*T>P8^DO '&LA:Q BS@ IBW:^DOF'QB2S ;D QBGU HQ 'Q QBGU HQD P H2O6Q&IP2/Q8FS6AQ!L&DX L L3`8 QBGU HQ#BP&8T@T)8S36T9BBAQA D P H2O]:'8#BP&^B 3 )8S3 a3^ DX L L3!L& 9]!Q`J, (Rana milletti) :Q3DQA]DLOLTIQ8 (Rhacophorus bisacculus) ^DOLU&^3& (Calluella guttulata) D P H2O6Q&IP2/Q8FS6AQ!L&DX L L3I F8`J, `8 QBGU HQ#BP&8T IL3#DL& 9]LP IQB6T @T QBGU HQ@Q L8J8 Q ^4 @TD P H2O!L&DX L L39Q&)8S36T^4 4 Q&a:a3^ DX L L3!L& 9JD&P a=D (Rana lateralis) *U&`8 QBGU HQ#BP&8T@TD P H2O!L&^5F>P84 Q&'Q QBGU HQ!L& _ FS6 (2505) `8BOAO6QA!L& QB]:DTA8^:D&BX:B Q& (Late metamorphosis) !L&:Q3DQA]DLOLTIQ8 (Rhacophorus bisacculus) 'Q QBGU HQ#BP&8T=9FQ9B S]F2!LGL ^DOJF]!P Q'O:BQ .^4@I T!QF *U&IL3#DL& 9P Ming (2004) 6T6Q QB9BBAQAD P H2O!L&DX L L3:Q3DQA]DLO`4 (R. appendiculatus) =9FQBOAO6 QA!L& QB]:DTA8^:D&BX:B Q& :BQ' .D P H2O3&P DQF]) 8 8P FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 87 DX LL36 T=9`8^JD &8Q^994 Q&c 'O@TBX:B Q&^DOD P H2O_#B&IB Q&:Q ^4 4 Q& 8^DOP 'O@T=C4S BB@ QB S8LQJQB4 Q& P83 FA IL3#D L& P9 QBGU HQ!L&'P86B6S=A (250/), _ FS6 (2505), FW1S ( 2546), Duellman and Trueb (1994), Khan (1998) ^DO Pough et al., 2001 *U& BX:B Q&^DO_#B&IB Q&:Q ^I3&3P&?Q#;8F 5S86TLA XLQGA6P T=9DX L L3)8S34 Q&c^I3&3P& 4QBQ&6T1 DX L L36TLQGA`8^JDP &8Q8 S&@ P @TDQ4 FP D@: L@ #BT9JQ&`J, DQ@]8 VLJQ&!8Q3]D' :Q LA X3Q8J8 Q_#B&IB Q&:Q ]:'8) L&]:S 3 a@ @T'&LA:Q ^DO*T>P8 a3^ DX L L3!L&LU&^3& (Calluella guttulata) LU&];Q (Glyphoglossus molossus) LU&DQA]DLO (Microhyla butleri) LU&!Q& 3Q (M. heymonsi) LU&8Q]4 Q (M. fissipes) LU&!Q#Q (M. pulchra) LU&JD&!P T3 (Micryletta inornata) DX L L3`8 DW @8TJQ S8`8BO39]J8P VL8Q^DO DQ&8Q_3A QB BL& S8L8W?Q#LQJQB!8Q3]D ' ^DO ^=D& 4L8= )6V TD L&DLALA`88 X QD P H2O3&P D QF'U&]:'8D P H2O6T]J@QOI@ 9P QB]#DVL86T!U8D& `88Q8 S& 9Q&)8S3@T:DQA#BT9JQ&D P H2O]BTAF]D' ]:'8]I 86Q`J IQ@QB5F QA8Qa3 LA Q&BF3]B'F ]) 8 DX L L3!L&LU&DQA]DLO ^DOLU&JDP&!T3 ^DOBF@5U&DX L L3!L&LU&^@ J8QF (Microhyla berdmorei) *U&LQGALAP `8D X QJ FA6 T8QaJD]L VLA3FA]) 8 8P DX L L3!L&LU&!Q&3 Q (Microhyla heymonsi) _#B&IB Q&:Q ]:'8) L&]:S36TBS@ FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 88 Leimberger (1997) 6T=9FQ DX LL3!L& Hyla chrysoscelis `8^JD&8 Q6 T@T4FLP L8!L&^@D&:LLA X 'O@T'W3IT3QJ8Q^8 86TI F8JQ&@Q FQD X LL3)8 S3]3TAF 86P TLQGA`8^JDP &8Q6 Ta@ @T;DX Q DX L L3#Q&# 9Q8 (Bufo melanostictus) #Q&# ^#BO (B. parvus) ^DO#Q&# JPFBQ9 (B. macrotis) ^@'O=9`8^L &8Q! &^4P DX L L3#Q&# 9Q8^DO#Q&# JFBQ9'O=9a3P ](=QO^L & 8Q!P&`8=V86TBQ94Q 5U&^@ IQ@QB5=94Q@DQJ FAa3 9 Q&^4 ']:'8#W&8 Q# L8! Q&8S& 3P&8P8 DQ@]8 VLJQ&!L&DX L L36&IL&)8P S3'U&@T!8Q3]D ' F Q DQ@]8 VLJQ&!L&DX L L3#Q&# ^#BO*U& =9`8^L &8Q! &BP S@DQJ FA `8D P H2O6T] S3'Q 8Q`8D QJ FAaJD8LAD&^DOaJD!Q34L8' U&] S3 ]:'8^L &8Q! &a@P `) ^L &8Q6 T] S3'Q 8Q<8! &P ^DO^L &8Q9Q&^J &#&A&@P T8QaJD; Q8LAX 3&8P P8 DX LL3#Q&# ^#BO'U&@T DQ@]8 VL#BT9JQ&6T^!&^B&^DO#B' T96T@T!8Q3]D ' ]BTAF@8@Q F QDX LL3 #Q&# 9Q8^DO#Q&# JFBQ9P DX LL36 TLQGA`8JP FA8 QaJDI F8`J, @ P 'O@T DQ@]8 VLJQ&!8Q3`J, #BT9JQ&!8Q3]D ' ^DO@TD P H2O6T]:DTA8^:D&a:`J ]J@QO 9P QBLQGA`8P BO^I8Q6 TaJD^B& ]) 8 DX L L3!L& 9L J8Q@ (Paa fasiculispina) 6T_#B&IB Q&:Q :BO L93FA^5F> P8'Q8F8@Q @TBS@ DX L L3!L&LU& BQA'869P XB2 (Xenophrys lekaguli) LQGA`8^JDP &8Q6 TaJD# L8!Q&^B& JQ S8_3A QB BL&L8W?Q#!8Q3]D ' 6T;SF8Q _#B&IB Q&:Q 'U&@TBS@ `8DQJ FA6 T8QaJD]L VLAIQ@QB5=9DX L L3!L&LU&^@ J8QF (Microhyla berdmorei) ^@F Q 'O@T#BT9JQ&6T FQ&^DO DQ@]8 VLJQ&!8Q3]D' ^4 DX L L3!L&LU&^@ J8QF 'IQ@QB5LQGA`8P DQJ FA6 T8QaJD]L VLAa3 ]=BQO@T=C4S BB@ QBJQ S8^99BF@<&X _3A'O=9FQA8 Q] QO DW @BF@ 8P ]:'8 L8 D@LA X DQ&8Q 6Q`J D3^B&]I TA36Q8!L& BO^I8Qa3 @Q FQLA ^99 X BO'QAJBVLLA4X FP ]3TAF *U&=C4S BB@^998TIQ@QB5=9a3`8D X L L3#Q&# 9Q8 (Bufo melanostictus) 6TLA X`8 DQJ FA6 T8QaJD]L VLA]) 8 8P ^4 QBBF@<&!L&#Q&#X 9Q8'O]: '8`8D P H2O]:'8 DW @6T^;^98LL @Q FQ6 T'O]:'8 DW@ D@]J@VL8LU&^@ J8QF FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 89 ?Q#;8F ?Q=DP H2O6Q&IP2/Q8^DO_#B&IB Q&:Q !L&DX L L36T=9`8=V86T]4BTA@'P34P& LW6AQ8^J &)Q4]!QIS S9J Q)P8 '&JFP 3'P 869P WBT A B ?Q=6 T 1 DX LL3^DO_#B&IB Q&:Q !L&LU& BQA'869P XB2 (Xenophrys lekaguli) Gosner Stage A B ?Q=6 T DX LL3^DO_#B&IB Q&:Q !L&#Q&# JFBQ9P (Bufo macrotis) Gosner Stage 3 A B ?Q=6 T / DX LL3^DO_#B&IB Q&:Q !L&#Q&# 9Q8 (Bufo melanostictus) Gosner Stage 33 A B ?Q=6 T 0 DX LL3^DO_#B&IB Q&:Q !L&#Q&# ^#BO (Bufo parvus) Gosner Stage 3 FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 90 A B ?Q=6 T 1 DX LL3^DO_#B&IB Q&:Q !L& 9J8L& (Fejervarya limnocharis) Gosner Stage 38 A B ?Q=6 T 6 DX LL3^DO_#B&IB Q&:Q !L& 98Q (Hoplobatrachus rugulosus) Gosner Stage 37 A B ?Q=6 T 3 DX LL3^DO_#B&IB Q&:Q !L&]!TA3'O8Q (Occidozyga lima) Gosner Stage 38 B A ?Q=6 T 4 DX LL3^DO_#B&IB Q&:Q !L&]!TA3JD&:P W@ (Occidozyga martensii) Gosner Stage FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 91 B A ?Q=6 T 9 DX LL3^DO_#B&IB Q&:Q !L& 9L J8Q@ (Paa fasiculispina) Gosner Stage A B ?Q=6 T 10 DX LL3^DO_#B&IB Q&:Q !L&]!TA3'S (Rana erythraea) Gosner Stage 39 A B ?Q=6 T -- DX L L3^DO_#B&IB Q&:Q !L& 9JD&a=DP (Rana lateralis) Gosner Stage 38 A B ?Q=6 T -. DX LL3^DO_#B&IB Q&:Q !L& 9JD&!P T3 (Rana macrodactyla) Gosner Stage 37 FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 9 B A ?Q=6 T -/ DX LL3^DO_#B&IB Q&:Q !L& 9LL&`J, (Rana mortenseni) Gosner Stage 3 A B ?Q=6 T -0 DX LL3^DO_#B&IB Q&:Q !L& 9]!Q`J, (Rana milletti) Gosner Stage 30 A B ?Q=6 T -1 DX LL3^DO_#B&IB Q&:Q !L&:Q3'SFGBTBQ)Q (Chirixalus hansenae) Gosner Stage 3 A B ?Q=6 T -6 DX LL3^DO_#B&IB Q&:Q !L&:Q3'SFDQA^4@ (Chirixalus nongkhorensis) Gosner Stage 37 FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 93 A B ?Q=6 T 17 DX LL3^DO_#B&IB Q&:Q !L&:Q39Q8 (Polypedates leucomystax) Gosner Stage 37 A B ?Q=6 T 18 DX LL3^DO_#B&IB Q&:Q !L&:Q34T8]JDVL& (Rhacophorus bipunctatus) Gosner Stage 37 A B ?Q=6 T 19 DX LL3^DO_#B&IB Q&:Q !L&:Q3DQA]DLOLTIQ8 (Rhacophorus bisacculus) Gosner Stage 38 A B ?Q=6 T 0 DX LL3^DO_#B&IB Q&:Q !L&LU&^3& (Calluella guttulata) Gosner Stage 37 FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 94 A B ?Q=6 T .- DX LL3^DO_#B&IB Q&:Q !L&LU&];Q (Glyphoglossus molossus) Gosner Stage 37 A B ?Q=6 T DX LL3^DO_#B&IB Q&:Q !L&LU&LQ& 8! T3 (Kaloula mediolineata) Gosner Stage 37 A B ?Q=6 T 3 DX LL3^DO_#B&IB Q&:Q !L&LU&LQ&9 Q8 (Kaloula pulchra) Gosner Stage 39 A B ?Q=6 T 4 DX LL3^DO_#B&IB Q&:Q !L&LU&^@ J8QF (Microhyla berdmorei) Gosner Stage 37 FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 9 A B ?Q=6 T DX LL3^DO_#B&IB Q&:Q !L&LU&DQA]DLO (Microhyla butleri) Gosner Stage 31 A B ?Q=6 T 6 DX LL3^DO_#B&IB Q&:Q !L&LU&8Q]4 Q (Microhyla fessipes) Gosner Stage A B ?Q=6 T 7 DX LL3^DO_#B&IB Q&:Q !L&LU&!Q&3 Q (Microhyla heymonsi) Gosner Stage 3 A B ?Q=6 T 8 DX LL3^DO_#B&IB Q&:Q !L&LU&!Q#Q (Microhyla pulchra) Gosner Stage 31 FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 96 A B ?Q=6 T 9 DX LL3^DO_#B&IB Q&:Q !L&LU&JD&!P T3 (Micryletta inornata) Gosner Stage 30 FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 97 ?Q#;8F (4 L) 4QBQ&6-T ^I3&5S86TLALQGX A!L&DP X LL3^4 DO)8S3 ^L &8Q! &P (8Q8 S&) ^8FDQJ FA (8QaJD ) _: &8Q] Q 8QaJD *L JS8` D 8Q4 )8S3 D P H2O6LAT XLQGPA `86BQ9T )QA: Q BLAD LB5 BS@DQJ FA 8QaJD]L LAV # L8! Q& 6@T 8T Q^(O 6@T T8QaJD; Q8 ^B& F&G Megophryidae Xenophrys lekaguli * * F&G Bufonidae Bufo melanostictus * Bufo parvus * Bufo macrotis * F&G Microhylidae Calluella guttulata * Glyphoglossus molossus * Microhyla berdmorei * Microhyla butleri * * * FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 98 4QBQ&6-T (4 L) ^I3&5S86TLALQG X A!L&DP X LL3^4 DO)8S3 ^L &8Q! &P (8Q8 S&) ^8FDQJ FA (8QaJD ) _: &8Q] Q 8QaJD *L JS8` D 8Q4 )8S3 D P H2O6TLA XLQGPA `86BQ9T )QA: Q BLAD LB5 BS@DQJ FA 8QaJD]L LAV # L8! Q& 6@T 8T Q^(O 6@T T8QaJD; Q8 ^B& Microhyla heymonsi * * * Microhyla fissipes * * * Microhyla pulchra * * * Micryletta inornata * * Kaloula pulchra * Kaloula mediolineata * F&G Dicroglossidae Fejervarya limnocharis * * * Hoplobatrachus rugulosus * * * Occidozyga lima * * * Occidozyga martensii * * * FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 99 4QBQ&6-T (4 L) ^I3&5S86TLALQG X A!L&DP X LL3^4 DO)8S3 ^L &8Q! &P (8Q8 S&) ^8FDQJ FA (8QaJD ) _: &8Q] Q 8QaJD *L JS8` D 8Q4 )8S3 D P H2O6LAT XLQGPA `86BQ9T )QA: Q BLAD LB5 BS@DQJ FA 8QaJD]L LAV # L8! Q& 6@T 8T Q^(O 6@T T8QaJD; Q8 ^B& Paa fasiculispina * F&G Ranidae Rana erythraea * Rana lateralis * Rana macrodactyla * Rana milleti * Rana mortenseni * * * F&G Rhacophoridae Chirixarus nongkhorensis * * Chirixarus hansenae * * Polypedates leucomystax * * * FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 100 4QBQ&6-T (4 L) ^I3&5S86TLALQG X A!L&DP X LL3^4 DO)8S3 ^L &8Q! &P (8Q8 S&) ^8FDQJ FA (8QaJD ) _: &8Q] Q 8QaJD *L JS8` D 8Q4 )8S3 D P H2O6LAT XLQGPA `86BQ9T )QA: Q BLAD LB5 BS@DQJ FA 8QaJD]L LAV # L8! Q& 6@T 8T Q^(O 6@T T8QaJD; Q8 ^B& Rhacophorus bisacculus * Rhacophorus bipunctatus * FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 101 4QBQ&6 T . J@QA]D!4FLAP Q&L Q&L S& )8S3 J@QA]D!4PFLA Q&L Q&L&S Xenophrys lekaguli CUMZ A 5237 Bufo melanostictus CUMZ A 5238 Bufo parvus CUMZ A 5239 Bufo macrotis CUMZ A 5240 Calluella guttulata CUMZ A 5241 Glyphoglossus molossus CUMZ A 5242 Microhyla berdmorei CUMZ A 5243 Microhyla butleri CUMZ A 5244 Microhyla heymonsi CUMZ A 5245 Microhyla fissipes CUMZ A 5246 Microhyla pulchra CUMZ A 5247 Micryletta inornata CUMZ A 5248 Kaloula pulchra CUMZ A 5249 Kaloula mediolineata CUMZ A 5250 Fejervarya limnocharis CUMZ A 5251 Hoplobatrachus rugulosus CUMZ A 5252 Occidozyga lima CUMZ A 5253 Occidozyga martensii CUMZ A 5254 Paa fasiculispina CUMZ A 5255 Rana erythraea CUMZ A 5256 Rana lateralis CUMZ A 5257 Rana macrodactyla CUMZ A 5258 Rana milleti ZMKUAMP-000301 Rana mortenseni CUMZ A 5259 Chirixarus nongkhorensis CUMZ A 5260 Chirixarus hansenae CUMZ A 5261 Polypedates leucomystax CUMZ A 5262 Rhacophorus bisacculus CUMZ A 5263 Rhacophorus bipunctatus CUMZ A 5264 FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 10 44S S BB@:BO QG !L!L9=BO#W2 F Q6TB LA_6 #Q8 U& 886_=7P S]3) JFJ8P Q= V86T]4BTA@'34P &LP W6AQ8^J &)Q4S ]!QIS9J Q)8#P W2)W8 LQ3FSa) #W2FS])H/ ](TA&]J8VL^DO]'QJ8 Q6 T=S6 P H: Q6W 6 Q86Ta3L Q8FA #FQ@IO3F `8 QB] 9!' 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AW=3T ]I4=BB2. 2500. ?X@SD P H26T@T;D4 LI?Q=^F3DL@:BO]6Ga6A . `8 )TFS4 P9IS&^F3D L@ (Human being and environment), J8Q 69-88. BW&]6=N : =SGSH/ QB=S@=. F1W S 6 P HS27BB@. 2502. #FQ@JDQ )8S3!L& 94PF]4@F/ A^DODP X L M3`8=86V ]!4BT P HQ=P87WIP4F: Q #DL&^I&. FS6AQ8S=87:BS,,Q@JQ9P20S4. IQ!QIP4FFS6AQ #2OFS6AQGQI4B @JQFS6AQDA]P H4BGQI4B. aIF F&J&HQP ^DO#8LV8c. 2550. DW @: QBLA4 L 5 '&JFP 3?Q#4OFP 8LLP : 4L86T 1 #FQ@IQ# ,P ])S&?X@SGQI4B^DO:BOF4P SGQI4B. `8 ;D&Q8FS'PA ^DOBQA&Q8#FQ@ QFJ8 Q&Q8FS'PA :BO'Q:T 2550,J8Q -52. BW&]6=N: IQ8 P L8WBP HI P4F: Q B@LW6AQ8^J &)Q4S IP4F: Q ^DO=87P W=)V . Buskirk, J. V., Aschwanden, J., Buckelmüller, I., Reolon, I., and Silvan Rüttiman. 2004. Bold tail coloration protects tadpoles from dragonfly strikes. Copeia. 2004(3):599-602. Duellman, W.E. and Trueb, L. 1986. Biology of Amphibian. NewYork: McGraw-Hill. FQBIQBI4FP : Q]@VL&a6A :T6T 17 (996P T - =.G. 253 Journal of Wildlife in Thailand Vol.17 No.1 010 103 Inger, R.F. 2005. 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