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A New Species of Illacme Cook & Loomis, 1928
A peer-reviewed open-access journal ZooKeys 626: 1–43A new (2016) species of Illacme Cook and Loomis, 1928 from Sequoia National Park... 1 doi: 10.3897/zookeys.626.9681 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research A new species of Illacme Cook & Loomis, 1928 from Sequoia National Park, California, with a world catalog of the Siphonorhinidae (Diplopoda, Siphonophorida) Paul E. Marek1, Jean K. Krejca2, William A. Shear3 1 Virginia Polytechnic Institute and State University, Department of Entomology, Price Hall, Blacksburg, Virginia, USA 2 Zara Environmental LLC, 1707 W FM 1626, Manchaca, Texas, USA 3 Hampden-Sydney College, Department of Biology, Gilmer Hall, Hampden-Sydney, Virginia, USA Corresponding author: Paul E. Marek ([email protected]) Academic editor: R. Mesibov | Received 25 July 2016 | Accepted 19 September 2016 | Published 20 October 2016 http://zoobank.org/36E16503-BC2B-4D92-982E-FC2088094C93 Citation: Marek PE, Krejca JK, Shear WA (2016) A new species of Illacme Cook & Loomis, 1928 from Sequoia National Park, California, with a world catalog of the Siphonorhinidae (Diplopoda, Siphonophorida). ZooKeys 626: 1–43. doi: 10.3897/zookeys.626.9681 Abstract Members of the family Siphonorhinidae Cook, 1895 are thread-like eyeless millipedes that possess an astounding number of legs, including one individual with 750. Due to their cryptic lifestyle, rarity in natural history collections, and sporadic study over the last century, the family has an unclear phylogenetic placement, and intrafamilial relationships remain unknown. Here we report the discovery of a second spe- cies of Illacme, a millipede genus notable for possessing the greatest number of legs of any known animal on the planet. -
Short: New Species of Polyxenida in Israel ______Those Reported in Condé & Nguyen Duy- Abbreviations: ZMUC––Zoological Mu- Jacquemin (1971) Were Plotted (Fig
Opusc. Zool. Budapest, 2020, 51(Supplementum 2): 35–45 _____________________________________________________________________________ New species of Polyxenida in Israel (Diplopoda, Penicillata) M. SHORT Megan Short, Deakin University, 221 Burwood Highway, Burwood, Melbourne, Australia E-mail: [email protected] Abstract. The identification of 5 species from a recent collection of 23 specimens from two areas in Israel is given. Two new species of Polyxenida are recorded, including the first species in the family Synxenidae (Phryssonotus sp.) and the first species from the family Polyxenidae, subfamily Macroxeninae (Chilexenus sp.), from Israel. These spe- cies are identified only to genus due to the lack of undamaged adult material. Two species in the family Lophoprocti- dae are also identified, one previously recorded but not described, is now described as Lophoproctus israelensis sp. nov., and Lophoproctinus chichinii Condé, 1951is recorded from Israel for the first time. The fifth species is identified as possibly a new species in the genus Polyxenus and likely to be synonymous with the specimens identified as Polyx- enus lagurus Linnaeus, 1758 in a previous study. This is just the second collection from Israel to be identified and brings the total number of Polyxenida species found in Israel to 8. All 3 families of Polyxenida are represented in this list, together with all 3 subfamilies of the most numerous family, Polyxenidae. Keywords. Bristly millipedes, biodiversity, Mediterranean, millipedes. Israel are identified, including description of a INTRODUCTION new species. n the period 1962–1966, Dr G. Levy of The MATERIALS AND METHODS I Hebrew University of Jerusalem (Department of Entomology and Venomous Animals) made The millipedes examined for this study were the first recorded collection of Polyxenida from collected between 2014–2018 by Amir Wein- Israel, with Condé & Nguyen Duy-Jacquemin stein and stored in 75% ethanol. -
United States National Museum ^^*Fr?*5J Bulletin 212
United States National Museum ^^*fr?*5j Bulletin 212 CHECKLIST OF THE MILLIPEDS OF NORTH AMERICA By RALPH V. CHAMBERLIN Department of Zoology University of Utah RICHARD L. HOFFMAN Department of Biology Virginia Polytechnic Institute SMITHSONIAN INSTITUTION • WASHINGTON, D. C. • 1958 Publications of the United States National Museum The scientific publications of the National Museum include two series known, respectively, as Proceedings and Bulletin. The Proceedings series, begun in 1878, is intended primarily as a medium for the publication of original papers based on the collections of the National Museum, that set forth newly acquired facts in biology, anthropology, and geology, with descriptions of new forms and revisions of limited groups. Copies of each paper, in pamphlet form, are distributed as published to libraries and scientific organizations and to specialists and others interested in the different subjects. The dates at which these separate papers are published are recorded in the table of contents of each of the volumes. The series of Bulletins, the first of which was issued in 1875, contains separate publications comprising monographs of large zoological groups and other general systematic treatises (occasionally in several volumes), faunal works, reports of expeditions, catalogs of type specimens, special collections, and other material of similar nature. The majority of the volumes are octavo in size, but a quarto size has been adopted in a few in- stances. In the Bulletin series appear volumes under the heading Contribu- tions from the United States National Herbarium, in octavo form, published by the National Museum since 1902, which contain papers relating to the botanical collections of the Museum. -
Arthropod Phylogeny Based on Eight Molecular Loci and Morphology
letters to nature melanogaster (U37541), mosquito Anopheles quadrimaculatus (L04272), mosquito arthropods revealed by the expression pattern of Hox genes in a spider. Proc. Natl Acad. Sci. USA 95, Anopheles gambiae (L20934), med¯y Ceratitis capitata (CCA242872), Cochliomyia homi- 10665±10670 (1998). nivorax (AF260826), locust Locusta migratoria (X80245), honey bee Apis mellifera 24. Thompson, J. D., Higgins, D. G. & Gibson, T. J. CLUSTALW: Improving the sensitivity of progressive (L06178), brine shrimp Artemia franciscana (X69067), water ¯ea Daphnia pulex multiple sequence alignment through sequence weighting, position-speci®c gap penalties and weight (AF117817), shrimp Penaeus monodon (AF217843), hermit crab Pagurus longicarpus matrix choice. Nucleic Acids Res. 22, 4673±4680 (1994). (AF150756), horseshoe crab Limulus polyphemus (AF216203), tick Ixodes hexagonus 25. Foster, P. G. & Hickey, D. A. Compositional bias may affect both DNA-based and protein-based (AF081828), tick Rhipicephalus sanguineus (AF081829). For outgroup comparison, phylogenetic reconstructions. J. Mol. Evol. 48, 284±290 (1999). sequences were retrieved for the annelid Lumbricus terrestris (U24570), the mollusc 26. Castresana, J. Selection of conserved blocks from multiple alignments for their use in phylogenetic Katharina tunicata (U09810), the nematodes Caenorhabditis elegans (X54252), Ascaris analysis. Mol. Biol. Evol. 17, 540±552 (2000). suum (X54253), Trichinella spiralis (AF293969) and Onchocerca volvulus (AF015193), and 27. Muse, S. V. & Kosakovsky Pond, S. L. Hy-Phy 0.7 b (North Carolina State Univ., Raleigh, 2000). the vertebrate species Homo sapiens (J01415) and Xenopus laevis (M10217). Additional 28. Strimmer, K. & von Haeseler, A. Quartet puzzlingÐa quartet maximum-likelihood method for sequences were analysed for gene arrangements: Boophilus microplus (AF110613), Euhadra reconstructing tree topologies. -
An Inventory of Endemic Leaf Litter Arthropods of Arkansas with Emphasis on Certain Insect Groups and Diplopoda Derek Alan Hennen University of Arkansas, Fayetteville
University of Arkansas, Fayetteville ScholarWorks@UARK Theses and Dissertations 12-2015 An Inventory of Endemic Leaf Litter Arthropods of Arkansas with Emphasis on Certain Insect Groups and Diplopoda Derek Alan Hennen University of Arkansas, Fayetteville Follow this and additional works at: http://scholarworks.uark.edu/etd Part of the Biology Commons, Entomology Commons, and the Terrestrial and Aquatic Ecology Commons Recommended Citation Hennen, Derek Alan, "An Inventory of Endemic Leaf Litter Arthropods of Arkansas with Emphasis on Certain Insect Groups and Diplopoda" (2015). Theses and Dissertations. 1423. http://scholarworks.uark.edu/etd/1423 This Thesis is brought to you for free and open access by ScholarWorks@UARK. It has been accepted for inclusion in Theses and Dissertations by an authorized administrator of ScholarWorks@UARK. For more information, please contact [email protected], [email protected]. An Inventory of Endemic Leaf Litter Arthropods of Arkansas with Emphasis on Certain Insect Groups and Diplopoda A thesis submitted in partial fulfillment of the requirements for the degree of Master of Science in Entomology by Derek Hennen Marietta College Bachelor of Science in Biology, 2012 December 2015 University of Arkansas This thesis is approved for recommendation to the Graduate Council. ___________________________________ Dr. Ashley P.G. Dowling Thesis Director ___________________________________ Dr. Frederick M. Stephen Committee Member ___________________________________ Dr. John David Willson Committee Member Abstract Endemic arthropods of Arkansas were sampled and their nomenclature and distributions were updated. The Arkansas endemic species list is updated to 121 species, including 16 species of millipedes. A study of the millipedes of Arkansas was undertaken, and resulted in the first checklist and key to all millipede species in the state. -
Millipede Defense
Proc. Natl. Acad. Sci. USA Vol. 93, pp. 10848-10851, October 1996 Ecology Millipede defense: Use of detachable bristles to entangle ants* (predation/mechanical defense/Diplopoda/Polyxenida/Formicidae) THOMAS EISNERtI, MARIA EISNERt, AND MARK DEYRUP§ tSection of Neurobiology and Behavior, Cornell University, Ithaca, NY 14853; and §Archbold Biological Station, Lake Placid, FL 33852 Contributed by Thomas Eisner, July 24, 1996 ABSTRACT The millipede Polyxenus fasciculatus (Dip- lopoda; Polyxenida) defends itself against ants by use ofa pair of bristle tufts at its rear. When attacked, it wipes the tufts against the ants, thereby causing these to become encumbered by bristles that detach from the tufts. Ants contaminated with bristles desist from their assault. The bristles have grappling hooks at the tip by which they lock onto setae of the ants and barbs along their length bywhich they interlinlk In attempting to rid themselves of bristles, ants may succeed only in further entangling themselves by causing the bristles to become enmeshed. Ants heavily contaminated may remain entangled and die. Most millipedes have chemical defenses; polyxenids, instead, have a mechanical weapon. FIG. 1. Schematic representation of a polyxenid-ant encounter. Millipedes (Arthropoda; Diplopoda) are an ancient group. They are also a hardy group, for they have survived since form what appears to be a single caudal projection (Fig. 2A). Silurian times (1) despite the evolutionary onslaught of insects. The bristles within these tufts are extremely fine in conparison Insects are among their chief enemies, and millipedes are to the body setae, and barely resolved at lower magnifications. well-protected against insect attack. Most possess defensive Because they glisten in light, the tufts appear brighter than the glands, in the form of integumental sacs, from which they body itself. -
Morphological Comparison of Bisexual and Parthenogenetic Polyxenus Lagurus (Linne, 1758) (Diplopoda, Polyxenidae) in Denmark
Morphological comparison of bisexual and parthenogenetic Polyxenus lagurus (Linne, 1758) (Diplopoda, Polyxenidae) in Denmark and Southern Sweden, with notes on taxonomy, distribution, and ecology by HENRIK ENGHOFF Abstract: A morphological comparison of the bisexual and the parthenogenetic (thelyto kous) form of Polyxenus lagurus, based on several metric and meristic characters, is pre sented. Both forms are rather variable, and cannot be separated from each other by the measured characters. A few samples are constituted by exceptionally large or small indi viduals. The two forms were distinguished entirely on the basis of sex ratio, which is not fully satisfactory, since it cannot be excluded that the thelytokous form produces a few males. It is recommended to distinguish neither the thelytokous form nor the "giant" and "dwarf" populations as separate species or subspecies. On the other hand, Polyxenus fasci culatus Say may be a subspecies of P. lagurus. The distribution of the two forms of P. lagurus in Denmark and southern-most Sweden is mapped, and new records from other countries are given. No difference in choice of habitat between the two forms could be demonstrated. A parasitoid gallmidge, presumably Chiliodiplosis vasta, attacks both forms and is recorded for the first time from Denmark and Sweden. INTRODUCTION Parthenogenesis in Polyxenus lagurus has been studied by rather many authors. Vandel (1926, 1928) discussed the geographic variation in sex ratio and hypothesized that the rarity or lack of males in the northeastern parts of Europe might be attributed to thelytoky (i.e. parthenogenesis in which unfertilized females produce female offspring). Vandel's data were mapped by Udvardy (1969) who mentioned P. -
Millipedes of Ohio Field Guide [Pdf]
MILLIPEDES OF OHIO field guide OHIO DIVISION OF WILDLIFE This booklet is produced by the Ohio Division of Wildlife as a free publication. This booklet is not for resale. Any unauthorized repro- duction is prohibited. All images within this booklet are copyrighted by the Ohio Division of Wildlife and its contributing artists and INTRODUCTION photographers. For additional information, please call 1-800-WILDLIFE (1-800-945-3543). Text by: Dr. Derek Hennen & Jeff Brown Millipedes occupy a category of often seen, rarely identified bugs. Few resources geared towards a general audience exist HOW TO VIEW THIS BOOKLET for these arthropods, belying their beauty and fascinating biol- Description & Overview ogy. There is still much unknown about millipedes and other Order Name myriapods, particularly concerning specific ecological informa- Family Name tion and detailed species ranges. New species await discovery Common Name and description, even here in North America. This situation Scientific Name makes species identification difficult for anyone lucky enough Range Map to stumble upon one of these animals: a problem this booklet indicates distribution and counties intends to solve. Here we include information on millipede life were specimen was collected Size history, identification, and collecting tips for all ~50 species denotes the range of length of Ohio’s millipedes. Millipede species identification often common for the species depends on examining the male genitalia, but to make this Secondary Photo (when applicable) booklet accessible as a -
Biodiversity Studies of Six Traditional Orchards in England
Natural England Research Report NERR025 Biodiversity studies of six traditional orchards in England www.naturalengland.org.uk Natural England Research Report NERR025 Biodiversity studies of six traditional orchards in England M. Lush1, H. J. Robertson, K. N. A. Alexander, V. Giavarini, E. Hewins1, J. Mellings1, C. R. Stevenson, M. Storey & P.F. Whitehead 1Just Ecology Environmental Consultency Limited Published on 23 April 2009 The views in this report are those of the authors and do not necessarily represent those of Natural England. You may reproduce as many individual copies of this report as you like, provided such copies stipulate that copyright remains with Natural England, 1 East Parade, Sheffield, S1 2ET ISSN 1754-1956 © Copyright Natural England 2009 Project details This report results from research commissioned by English Nature and completed after the successor organisation, Natural England, was set up in October 2006. The work was undertaken under English Nature contract CPAU03/02/189 by the following team: Mike Lush, Eleanor Hewins and Jon Mellings of Just Ecology, Heather Robertson, of Natural England during the project, now retired, and individual consultants Keith Alexander, Vince Giavarini, Robin Stevenson and Malcolm Storey. Results from the report were used from 2005 to 2007 to support the proposal to list traditional orchards as a national priority habitat in the Biodiversity Action Plan. Since 2007, the report has been expanded to incorporate previous work by Paul Whitehead on one study site. The study site results are now being made more widely available, in the form of a permanent record, in this Natural England Research Report. -
Diversity and Distribution of the Millipedes (Diplopoda) of Georgia, Caucasus
A peer-reviewed open-access journal ZooKeys 930: 199–219 (2020)Diversity and distribution patterns of the millipedes of Georgia 199 doi: 10.3897/zookeys.930.47490 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research Diversity and distribution of the millipedes (Diplopoda) of Georgia, Caucasus Mzia S. Kokhia1, Sergei I. Golovatch2 1 Institute of Zoology, Ilia State University, K. Cholokashvili Ave., 3/5, Tbilisi 0162, Georgia 2 Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia Corresponding author: Sergei I. Golovatch ([email protected]) Academic editor: Z. Korsós | Received 21 October 2019 | Accepted 10 January 2020 | Published 28 April 2020 http://zoobank.org/3D90FD4C-DC71-4F68-987D-8F24DE2141A8 Citation: Kokhia MS, Golovatch SI (2020) Diversity and distribution of the millipedes (Diplopoda) of Georgia, Caucasus. In: Korsós Z, Dányi L (Eds) Proceedings of the 18th International Congress of Myriapodology, Budapest, Hungary. ZooKeys 930: 199–219. https://doi.org/10.3897/zookeys.930.47490 Abstract The diplopod fauna of Georgia, Transcaucasia, is very rich given the country’s relatively small territory; it presently comprises 103 species from 44 genera, 12 families, and 7 orders. Most of the Diplopoda known from Georgia (86 species, or 83%) demonstrate Caucasian distribution patterns, 36 and 46 species, as well as 8 and 9 genera being endemic or subendemic to the country, respectively. A single Holarctic family, Anthroleucosomatidae (order Chordeumatida), contains 44 Caucasian species and 20 genera, of which 27 species and 14 genera are endemic or subendemic to Georgia. Likewise, all species from the orders Polyzoniida, Siphonocryptida, Glomerida and Chordeumatida, as well as most species of Julida and Polydesmida are native, also endemic or subendemic to the Caucasus, but the genera and families they represent are widely distributed at least across the Euro-Mediterranean Realm. -
Millipede (Diplopoda) Distributions: a Review
SOIL ORGANISMS Volume 81 (3) 2009 pp. 565–597 ISSN: 1864 - 6417 Millipede (Diplopoda) distributions: A review Sergei I. Golovatch 1* & R. Desmond Kime 2 1 Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia; e-mail: [email protected] 2 La Fontaine, 24300 La Chapelle Montmoreau, France; e-mail: [email protected] *Corresponding author Abstract In spite of the basic morphological and ecological monotony, integrity and conservatism expressed through only a small number of morphotypes and life forms in Diplopoda, among which the juloid morphotype and the stratobiont life form are dominant, most of the recent orders constituting this class of terrestrial Arthropoda are in a highly active stage of evolution. This has allowed the colonisation by some millipedes of a number of derivative, often extreme and adverse environments differing from the basic habitat, i.e. the floor of temperate (especially nemoral), subtropical or tropical forests (in particular, humid ones). Such are the marine littoral, freshwater habitats, deserts, zonal tundra, high mountains, caves, deeper soil, epiphytes, the bark of trees, tree canopies, ant, termite and bird nests. Most of such difficult environments are only marginally populated by diplopods, but caves and high altitudes are often full of them. To make the conquest of ecological deviations easier and the distribution ranges usually greater, some millipedes show parthenogenesis, periodomorphosis or morphism. Very few millipede species demonstrate vast natural distributions. Most have highly restricted ranges, frequently being local endemics of a single cave, mountain, valley or island. This contrasts with the remarkable overall diversity of the Diplopoda currently estimated as exceeding 80 000 species, mostly confined to tropical countries. -
LESOPS 23 Daws Millipedes
LEICESTERSHIRE ENTOMOLOGICAL SOCIETY The Millipedes of Leicestershire and Rutland Jonathan Daws* & Helen Ikin Polyxenus lagurus LESOPS 23 (2010) ISSN 0957 - 1019 *Corresponding author: 'Ysildor ', 177 Featherstone Drive, Leicester, LE2 9RF VC55 Millipedes 2 Acknowledgements The authors have only been able to compile this report with the help of a large group of people and we would like to thank them for their help. The maps were produced by Gareth Price who also helped sift many of the erroneous records from the database. Paul Lee, Steve Gregory and Tony Barber verified records and identified specimens. Julia Daws and Tony Barber proof-read the report picking out the typographical and technical mistakes respectively. Technical support and sources of information were provided by Jan Dawson, Ivan Pedley, Anona Finch, Ray Morris, Steve Woodward, Jean Harvey, Peter Brown (Biological Records Centre), Elizabeth Peat, Darwyn Sumner and Ian Evans. Finally, this report would not have been possible without the group of enthusiasts who, for over a century, have recorded these invertebrates, the most notable of whom is Dr Adrian Rundle. We are also indebted to Steve Woodward who kindly allowed reproduction of photographs of millipedes he has recorded in VC55. Introduction Millipedes are a diverse group of invertebrates. They can be told apart from the superficially similar groups of centipedes and pill woodlice by possessing two pairs of legs per body segment whereas the other two groups have only one pair of legs per segment. Currently, there are sixty two species of millipedes on the British list - six of these are unable to survive our winter climate and are confined to heated glasshouses.