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19 3 153 188 Proszynski for Inet.P65 Arthropoda Selecta 19(3): 153188 © ARTHROPODA SELECTA, 2010 Description of some Salticidae (Araneae) from the Malay Archipelago. I. Salticidae of the Lesser Sunda Islands, with comments on related species Îïèñàíèå íåêîòîðûõ Salticidae (Araneae) èç Ìàëàéñêîãî Àðõèïåëàãà. I. Salticidae Ìàëûõ Çîíäñêèõ îñòðîâîâ ñ êîììåíòàðèÿìè î áëèçêèõ âèäàõ Jerzy Prószyñski*, Christa L. Deeleman-Reinhold** É. Ïðóøèíüñêèé*, Ê. Äèëåìàí-Ðåéíîëüä** * Museum and Institute of Zoology, Polish Academy of Sciences, ul. Wilcza 64, 00-679 Warszawa, Poland. E-mail: jerzy.Prószyñ[email protected] ** 4619GA Ossendrecht, the Netherlands. E-mail: [email protected] KEY WORDS: Salticidae, new species, diagnostic characters, geographical distribution, Indonesia, Bali, Flores, Lombok, Sumba, Sumbawa. ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: íîâûé âèä, äèàãíîñòè÷åñêèå ïðèçíàêè, ðàñïðîñòðàíåíèå, Èíäîíåçèÿ, î. Áàëè, î. Ôëîðåñ, î. Ëîìáîê, î. Ñóìáà, î. Ñóìáàâà. In memoriam Bohdan Pisarski, friend of J. Prószyñski and com- panion in the Java and Bali collecting trip in 1959, for many years the Director of the Institute of Zoology PAN. ABSTRACT. This paper provides preliminary ref- Myrmarachne MacLeay, 1839 is discussed. Comple- erence diagnostic drawings for selected Oriental gen- mentary diagnostic drawings are added for the fol- era and species, to complement the existing scanty lowing species: Artabrus erythrocephalus (C.L. Koch, literature. The following new taxa are described: new 1846), Harmochirus brachiatus (Thorell, 1877), genus Katya gen.n., new species: Burmattus Hasarius adansoni (Audouin, 1826), Myrmarachne pachytibialis sp.n., Carrhotus sundaicus sp.n., Chrysilla hirsutipalpi [?] Edmunds & Prószyñski, 2003, Spar- deelemani sp.n., Cosmophasis valerieae sp.n., Cytaea taeus spinimanus (Thorell, 1878), Thiania bhamoen- whytei sp.n., Euryattus [?] junxiae sp.n., Katya flore- sis Thorell, 1887. Several unidentified species are scens sp.n., Katya ijensis sp.n., Katya inornata sp.n., also mentioned, whenever they contribute to our Ligurra moniensis sp.n., Meata zabkai sp.n., Myrma- knowledge of the geographical distribution of their rachne balinese sp.n., Myrmarachne glavisi sp.n., Myr- respective genera. marachne jacksoni sp.n., Phaeacius azarkinae sp.n., Siler lewaense sp.n., Stergusa incerta sp.n., Thyene ÐÅÇÞÌÅ.  ñòàòüå ïðèâåäåíû èëëþñòðàöèè äëÿ gangoides sp.n. and Thyene benjamini sp.n. íåêîòîðûõ îðèåíòàëüíûõ ðîäîâ è âèäîâ â äîïîëíå- The following new combinations and synonyms are íèå ê íåïîëíûì ëèòåðàòóðíûì èñòî÷íèêàì. Îïèñà- established: Artabrus jolensis Simon, 1902 = Telamo- íû ñëåäóþùèå íîâûå òàêñîíû: íîâûé ðîä Katya nia jolensis (Simon, 1902) comb.n., Euophrys chiari- gen.n., íîâûå âèäû: Burmattus pachytibialis sp.n., atapuensis Tikader, 1977 = Thiania bhamoensis Carrhotus sundaicus sp.n., Chrysilla deelemani sp.n., Thorell, 1887 syn.n., Evarcha kochi Simon, 1902 (re- Cosmophasis valerieae sp.n., Cytaea whytei sp.n., instated as separate species), Gangus concinnus (Key- Euryattus [?] junxiae sp.n., Katya florescens sp.n., serling, 1881) = Thyene concinna (Keyserling, 1881) Katya ijensis sp.n., Katya inornata sp.n., Ligurra comb.n., Gangus decorus Simon, 1902 = Thyene deco- moniensis sp.n., Meata zabkai sp.n., Myrmarachne ra (Simon, 1902) comb.n., Gangus longulus Simon, balinese sp.n., Myrmarachne glavisi sp.n., 1902a = Thyene longula (Simon, 1902) comb.n., Gan- Myrmarachne jacksoni sp.n., Phaeacius azarkinae gus manipissus Barrion & Litsinger, 1995 = Thyene sp.n., Siler lewaense sp.n., Stergusa incerta sp.n., manipisa (Barrion & Litsinger, 1995). Original com- Thyene gangoides sp.n. è Thyene benjamini sp.n. Ïðåä- bination Emertonius exasperans Peckham & Peck- ëîæåí ðÿä íîâûõ êîìáèíàöèé è íîâàÿ ñèíîíèìèÿ ham, 1892 is reinstated. The subdivision of the genus äëÿ: Artabrus jolensis Simon, 1902 = Telamonia 154 J. Prószyñski & C. Deeleman-Reinhold Map 1. Lesser Sunda Islands. Êàðòà 1. Ìàëûå Çîíäñêèå îñòðîâà. jolensis (Simon, 1902) comb.n., Euophrys chiariata- sexes, so the remaining species without such documen- puensis Tikader, 1977 = Thiania bhamoensis Thorell, tation are hardly recognizable). Similarly, the Philip- 1887 syn.n., Evarcha kochi Simon, 1902 [âîññòàíîâ- pines have 95 species, but only 85 have diagnostic ëåí â ñòàòóñå âèäà], Gangus concinnus (Keyserling, drawings. In contrast Central America has 485 species 1881) = Thyene concinna (Keyserling, 1881) comb.n., (414 with diagnostic drawings) and North America has Gangus decorus Simon, 1902 = Thyene decora (Simon, 502 species (447 with diagnostic drawings) [Prószyñs- 1902) comb.n., Gangus longulus Simon, 1902a = ki, 2010 online]. It can be expected that the fauna of Thyene longula (Simon, 1902) comb.n., Gangus ma- tropical Indonesia will be more diverse than that of nipissus Barrion & Litsinger, 1995 = Thyene manipisa mainly temperate North America. (Barrion & Litsinger, 1995). Âîññòàíîâëåíà îðèãè- An additional complication with regard to our íàëüíàÿ êîìáèíàöèÿ Emertonius exasperans Peck- knowledge of Salticidae from the Malay Archipelago ham & Peckham, 1892. Îáñóæäàåòñÿ ïîäðàçäåëåíèÿ is that some species are described from a single, or a ðîäà Myrmarachne MacLeay, 1839. Ïðèâîäÿòñÿ íî- few locations, but their distributions are generalized to âûå ðèñóíêè äëÿ Artabrus erythrocephalus (C.L. Koch, include entire large islands or the whole archipelago. 1846), Harmochirus brachiatus (Thorell, 1877), The broader distributions of even better defined Hasarius adansoni (Audouin, 1826), Myrmarachne species have not been documented by diagnostic draw- hirsutipalpi [?] Edmunds & Prószyñski, 2003, Spar- ings, and often represent summaries of misidentified taeus spinimanus (Thorell, 1878) è Thiania bhamoensis related species. Thorell, 1887. Óïîìèíàåòñÿ ðÿä íåîïðåäåëåííûõ The current insufficiency of data for Salticidae pre- âèäîâ, â òîì ñëó÷àå åñëè ñâåäåíèÿ î íèõ ðàñøèðÿ- clude the investigation of several interesting biological þò äàííûå î ðàñïðîñòðàíåíèè òåõ èëè èíûõ ðîäîâ. questions. For example, what is the geographical speciation Introduction pattern of Salticidae in the Malay Archipelago? One can expect that each small island may harbor its own The Malay Archipelago is politically divided into species, and larger islands should have chains of relat- several countries, and harbors one of the richest faunae ed species but do they? What is the transition pattern of the world. It is a classic area of zoogeographic and of Salticidae between the Asiatic mainland and Austra- evolutionary research, initiated by the memorable book lia? To what extent is the distribution of Salticidae of Wallace [1881]. Strangely, relatively few publica- influenced by the classic Wallaces Line? tions have dealt Salticidae from the Archipelago, and This paper cannot answer these or similar ques- our knowledge of that fauna is particularly incomplete. tions, but it defines a number of new or poorly known For example, Indonesia has 330 described species of species, in some cases extending significantly the known Salticidae, but only 215 of these having any diagnostic geographic ranges. The faunal relationships between drawings available (only 83 have drawings for both larger islands and groups of smaller islands cannot be Description of some Salticidae from the Malay Archipelago 155 resolved yet, although some hints are discernible even rialize. The relationships of the described species are from the preliminary data. illustrated by drawings of relevant species taken from the Identifications and definitions of species in Salti- literature [Prószyñski, 1984b, 1987, and others]. All orig- cidae are based on comparison with existing diagnostic inal drawings for this paper are made by J. Prószyñski. drawings of Salticidae in the literature, as summarized Specimens were studied under a stereomicroscope, in the Internet database Monograph of Salticidae (Ara- with magnification up to 100x. Palpal organs were neae) of the World by Prószyñski [2010 online, a detached, fixed in sand in an ethanol filled Petri dish. summary of all previous versions since 1995]. The After examination they were put in microvials together basis of identification is highly insufficient for Salti- with the original specimens. The epigynes were drawn cidae of the Oriental Region: many species previously in situ. For study of the internal structures the epigynes described in the worlds literature are in fact single were dissected, soaked in 1020% solution of KOH examples of larger clades, the existing diagnostic doc- (under controlled conditions for 25 hours), stained in umentation is incomplete, and there are a number of alcohol solution of Chlorazol Black E and mounted in genera for which only one sex was described. Clove oil for examination under a compound micro- The main aim of this paper is to provide prelimi- scope. Subsequently, the epigynes were deposited in a nary reference diagnostic drawings to complement the microvial with ethanol and stored together with the scanty literature data for certain genera and species. original specimen. All drawings were made using a grid The genera and species are classified provisionally, system. Species are defined in this paper by pictures of pending revisions of related genera, especially their their genital organs and body features, studied in single insufficiently studied type specimens. An untapped specimens and compared with drawings of type species source of taxonomic information are photographs of and all other species of each genus, shown in Prószyñski Salticidae, available now on the internet, some of which [2010 online]. There was no
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