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Distribution maps and checklist of Namibian

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The user has requested enhancement of the downloaded file. ULA L LU L E B I L Suhling F. & A. Martens Namibian Dragonflies: Larval Key andDistribution ISSN 0723-6514 Zeitschrift der Gesellschaft deutschsprachiger Odonatologen (GdO) e.V. Preface Suhling F., O. Martens Müller&A. : Thedragonfly larvae ofNamibia(Odonata) ISSN 0723 - 65140723- ISSN Zeitschrift der Gesellschaft deutschsprachiger Odonatologen (GdO) e.V. (GdO) Odonatologen deutschsprachiger Gesellschaft der Zeitschrift M J W B P Libellulidae) and der aktuellen Temperatur und der Aufzuchtsbedingungen (Odonata: Libellulidae) M Neumark-Nord(Odonata: Coenagrionidae, Libellulidae) B Coenagrionidae) (Odonata: Schleswig-Holstein in Arealrand westlichen T nidia;Odonata: ) Coleoptera: Cicindelidae) H W nahmen(Odonata) UILLERAT ETZOLD RAUNER ILLMANNS ROCKHAUS OPE ÜHLE ILDERMUTH AUERSBERGER INKLER Sympetrum sinaiticum Sympetrum . Newrecords P.: of L., L. R

L.&C. M . ErstnachweisO.: von F.: Fund eines Fundvon Wassermilben F.: parasitierten Männchens von . C., H. N ...... O.: T., S. M . FörderungderLibellenfauna kleiner H.: Moorgräben durch einfache Naturschutzmaß - OHE ...... Nimmt R.: mixta Aeshna , I. F EUMANN ONNERAT ENG LENNER ...... Supplement 13 & O. M Leucorrhiniapectoralis & A. D :Odonata in southern Morocco, with first records of ecrhna caudalisLeucorrhinia ...... : Distribution maps and checklist of Namibian Odonata & F. S (Odonata: Libellulidae) ...... Ceriagriontenellum als Beute der Larve von Larve der Beute als ÜLLER REWS UHLING : Fossile Libellenlarven aus den interglazialen Seesedimenten von Verbreitung und Ökologie von : Atmungsverhalten: von inTurkey (Odonata: Libellulidae) m odse Dushad rzn z? (Odonata: zu? rezent Deutschlands Nordosten im inBrandenburg (Odonata: 25-29 Coenagrionidae) ...... Cicindela campestris Cicindela ...... Orthetrum cancellatum Coenagrion armatum 28 Anax parthenopeAnax ...... (Odonata: Aeshnidae; (Odonata: Orthetrumransonnetii ......

-Larven: Einfluss ......

...... (1/2) (Hydrach-

2009 am süd- 107-175 97-115 2014 59-68 93-96 31-48 69-84 49-58 85-88 89-91 5-106 1-24 1

Namibian Dragonflies 20092014 LibellulaLibellula Supplement 28 (1/2) 13 ULA L LU L E B I L ULA L LU L E B I L 28 ISSN 0723 - 65140723- ISSN Zeitschrift der Gesellschaft deutschsprachiger Odonatologen (GdO) e.V. (GdO) Odonatologen deutschsprachiger Gesellschaft der Zeitschrift ISSN 0723 - 65140723- ISSN Namibian Dragonflies: Larval Key andDistribution ISSN 0723-6514 Zeitschrift der Gesellschaft deutschsprachiger Odonatologen (GdO) e.V. Zeitschrift der Gesellschaft deutschsprachiger Odonatologen (GdO) e.V. (GdO) Odonatologen deutschsprachiger Gesellschaft der Zeitschrift M J W B P Libellulidae) and der aktuellen Temperatur und der Aufzuchtsbedingungen (Odonata: Libellulidae) M Neumark-Nord(Odonata: Coenagrionidae, Libellulidae) B Coenagrionidae) (Odonata: Schleswig-Holstein in Arealrand westlichen nidia;Odonata: Aeshnidae) T H Coleoptera: Cicindelidae) W nahmen(Odonata) UILLERAT ETZOLD RAUNER ILLMANNS ROCKHAUS OPE ÜHLE ILDERMUTH AUERSBERGER INKLER Sympetrum sinaiticum Sympetrum . Newrecords P.: of L., L. R L.&C. M . ErstnachweisO.: von F.: Fund eines Fundvon Wassermilben F.: parasitierten Männchens von C., H. N O.: T., S. M . FörderungderLibellenfauna kleiner H.: Moorgräben durch einfache Naturschutzmaß - OHE ...... Nimmt R.: (1/2) Aeshna mixta Aeshna , I. F EUMANN ONNERAT ENG LENNER ...... Supplement 13 & O. M Leucorrhiniapectoralis & A. D :Odonata in southern Morocco, with first records of ecrhna caudalisLeucorrhinia ...... & F. S (Odonata: Libellulidae) ...... Ceriagriontenellum als Beute der Larve von Larve der Beute als ÜLLER REWS UHLING : Fossile Libellenlarven aus den interglazialen Seesedimenten von Verbreitung und Ökologie von : Atmungsverhalten: von inTurkey (Odonata: Libellulidae) m odse Dushad rzn z? (Odonata: zu? rezent Deutschlands Nordosten im inBrandenburg (Odonata: 25-29 Coenagrionidae) ...... Cicindela campestris Cicindela ...... Orthetrum cancellatum Coenagrion armatum 28 Anax parthenopeAnax ...... (Odonata: Aeshnidae; (Odonata: Orthetrumransonnetii ...... -Larven: Einfluss (1/2) (Hydrach- 2009 2009 am süd- 97-115 2014 59-68 93-96 31-48 69-84 49-58 85-88 89-91 1-24

2009 Libellula 28 (1/2) ULA L LU L E B I L 28 ISSN 0723 - 65140723- ISSN Zeitschrift der Gesellschaft deutschsprachiger Odonatologen (GdO) e.V. (GdO) Odonatologen deutschsprachiger Gesellschaft der Zeitschrift (1/2) 2009 LIBELLULA SUPPLEMENT Supplement 13, Bremen, 15. März 2014 Zeitschrift der Gesellschaft deutschsprachiger Odonatologen (GdO) e.V. LIBELLULA LIBELLULAISSN 0723-6514 Libellula 28 (1/2) LIBELLULA

Front cover: Larvae of Diplacodes levebvrii, Mastinogomphus dissimilis, and Pseudagrion massaicum Drawings:Zeitschrift Ole Müller der Gesellschaft deutschsprachiger Odonatologen (GdO) e.V. Zeitschrift der Gesellschaft deutschsprachiger Odonatologen (GdO) e.V. Colour: Dune 45 ISSN 0723 - 6514 2009 ISSN 0723 - 6514 2009 2009

Redaktion Klaus Guido Leipelt, Karlsruhe Graham Vick, Little London Mathias Lohr, Höxter Ida Suhling, Braunschweig WINKLER C., H. NEUMANN & A. DREWS: Verbreitung und Ökologie von Coenagrion armatum am süd- westlichen Arealrand in Schleswig-Holstein (Odonata: Coenagrionidae) ...... 1-24

BRAUNER O.: Erstnachweis von Ceriagrion tenellum in Brandenburg (Odonata: Coenagrionidae) 25-29

WILDERMUTH H.: Förderung der Libellenfauna kleiner Moorgräben durch einfache Naturschutzmaß - LIBELLULAnahmen (Odonata) SUPPLEMENT...... dient der Veröffentlichung umfangreicher Arbeiten 31-48 und der Zusammenstellung besonderer Themen auf dem Gebiet der Libellenkunde. BROCKHAUS T., S. MENG & O. MÜLLER: Fossile Libellenlarven aus den interglazialen Seesedimenten von EsNeumark-Nord erscheint unregelmäßig (Odonata: als Coenagrionidae, Ergänzungsband Libellulidae) der Zeitschrift...... 49-58

MÜHLE L., L. ROHE, I. FLENNER & F. SUHLING: Atmungsverhalten vonLIBELLULA Orthetrum cancellatum-Larven: Einfluss Schriftleitung:der aktuellen Temperatur und der Aufzuchtsbedingungen (Odonata: Libellulidae) ...... 59-68 Dr. Mathias Lohr, Hochschule Ostwestfalen-Lippe, Fachgebiete Landschaftsökologie und Tierökologie, AnM AUERSBERGERder WilhelmshöheR.: Nimmt 44, 37671Leucorrhinia Höxter caudalis im Nordosten Deutschlands rezent zu? (Odonata: Libellulidae) ...... 69-84

BP eETZOLD z u gF.: von Fund LIBELLULA eines von Wassermilben und Adressenänderung parasitierten Männchens über: von Anax parthenope (Hydrach- GdO,nidia; Traute Odonata: Fliedner, Aeshnidae) Louis-Seegelken-Straße...... 106, D-28717 Bremen 85-88

TILLMANNS O.: Aeshna mixta als Beute der Larve von Cicindela campestris (Odonata: Aeshnidae; FürColeoptera: Mitglieder Cicindelidae) der GdO ist ...... der Bezug von und LIBELLULA SUPPLEMENT im 89-91 JahresbeitragHOPE P.: New enthalten.records of DieserLeucorrhinia beträgt pectoralis 35,- €; aufin Turkey Antrag (Odonata: für Schüler, Libellulidae) Studenten (nach...... 2 Jahren 93-96 Neuantrag!) und Rentner 25,- €. Beiträge LIBELLULAbitte auf das Konto der GdO, z.Hd. Traute Fliedner überweisen:JUILLERAT L. & IBAN: C. M ONNERATDE32 3701: Odonata 0050 0330 in southern 9195 05 Morocco, (Konto Nr. with 03 30first 91 records 95 05) of bei Orthetrum der Postbank ransonnetii Köln, BIC:and PBNKDEFF Sympetrum (BLZ sinaiticum 370 100(Odonata: 50). Libellulidae) ...... 97-115

Die GdO im Internet:

© 2014 GdO, Bremen Printed in Germany, Druckerei Koch, Reutlingen Kein Teil dieses Heftes darf ohne vorherige Erlaubnis des Herausgebers – in welcher Form auch immer – reproduziert werden. 28(1/2) 28(1/2) Design, Layout und Satz: Birgit Rödder, Bad Münstereifel Distribution maps and checklist of Namibian Odonata 15. März 2014107 Distribution maps and checklist of Namibian Odonata

Frank Suhling 1 and Andreas Martens 2

1) Institut für Geoökologie, Technische Universität Braunschweig, Langer Kamp 19c, 38106 Braunschweig, Germany, 2) Institut für Biologie und Schulgartenentwicklung, Pädagogische Hochschule Karlsruhe, Bismarckstraße 10, 76133 Karlsruhe, Germany,

Abstract

This paper presents a checklist of the Odonata of Namibia and provides up-to-date dis- tribution maps of 130 species. Compared with the checklist from 2007, five species have been added to the list, four of them were new for science. Two species were omitted.

Zusammenfassung

Verbreitungskarten und Checkliste der Libellen Namibias (Odonata). – Aus Namibia sind derzeit 130 Libellenarten bekannt. Im Vergleich zur Checkliste von 2007 werden fünf Arten hinzugefügt, vier davon waren neu für die Wissenschaft. Zwei Arten werden von der bis- herigen Liste gestrichen. Für alle Arten werden aktualisierte Verbreitungskarten geliefert.

Introduction

Previously published species lists of the Odonata of Namibia are included in pub- lications on other topics, by Brinck (1955), Curtis (1991), and Pinhey (1984, 1985). Brinck’s (1955) list of Namibian Odonata refers to 32 species, whereas Pinhey’s South African Odonata checklist (Pinhey 1984, 1985) refers to 56 spe- cies for Namibia. Curtis (1991) included 70 valid species of dragonflies in a checklist of freshwater macro-invertebrates of Namibia (plus some taxa marked as doubtful; cf. Martens et al. 2003). With the activities of the entomologists of the Museum für Naturkunde, Humboldt-Universität, Berlin (ZMHB) and finally the BIOTA research project the survey activity in Namibia increased (Fig. 1). The revised dragonfly fauna recorded 102 species (Martens et al. 2003). The next update listed additional 25 species raising the total species number to 127 and presented preliminary distribution maps for all species as of 2006 (Suhling & Martens 2007). Libellula Supplement 13: 107-175 108 Frank Suhling & Andreas Martens

The aim of this contribution is to deliver an updated checklist and up-to-date dis- tribution maps for all species. We also critically examine the checklist of 2007 and suggest some few changes in the species list. We provide maps for a total of 130 species. Data acquisition

The distribution maps and the checklist presented here, base on previously pub- ● lished checklists (see above) and on additional sources: Namibian National Collection of , National Museum of Namibia, Wind- hoek (NMNW). This includes material collected by L. Coetzee, S.V. Green, J. Irish, A.H. Kirk-Spriggs, E. Marais, C.S. Roberts and H. & W. Rust. Virtually all the speci- mens dating from the period 1936-1963 lack a collector’s name on the attached labels; most of these have likely been collected by F. Gaerdes. The collection ● also includes material collected by R. Jödicke in 1999. For this study, we re- identified the majority of Odonata material held in the NMNW. Odonata held in the Museum für Naturkunde, Humboldt-Universität, Berlin ● (ZMHB) collected by J. Deckert, U. Göllner, F. Koch, and G. Peters (cf. Koch et al. 1995); F. Koch provided also more recently collected specimens. Odonata of the General Entomology Collection of the Transvaal Museum, Gauteng (TMSA), collected by B.I. Balinsky, F. Gaerdes, C. Koch, H. Strydom and others. W. Tarboton examined the specimens and provided a list.

Figure 1. Temporal distribution of dragonfly records in Namibia. First records are from 1887. Note the logarithmic scale of records.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 109

● Several photographic records by H.-M. Koch, H. Roland, and P. Reiner identified by the authors. Records of J. Kipping during several sampling trips since 2000. For some rare ● species we consider records made by him at the Chobe River near Kasane, Bo- tswana, which is just opposite Impalila Island, Namibia (Kipping 2010). Records generated from an extensive survey of the western ephemeral river catchments of Namibia and the upper Fish River catchment 2001 to 2003, and ● collecting trips to the Kunene in 2002 and the Okavango in 2003, by A. Martens and F. Suhling. Records generated from several excursions to different parts of the country, particularly the Caprivi, 2004 to 2013 by F. Suhling, accompanied by I. Suh- ling, K.G. Leipelt, E. Marais, A. Martens, E. Mosch, J. Ott, G. Sahlén, P. Schridde, C. Schütte, O. Richter and G. Rüppell, respectively, for some periods. All collected specimens are deposited in NMNW. All data were compiled in a data- base maintained by FS (cf. Fig. 2). Changes in the checklist since Suhling & Martens (2007)

Agriocnemis bumhilli Lestinogomphus silkae Five species are Trithemis newly recorded. morrisoni Among these, four are new to T. science,palustris namely Kipping, Martens & Suhling, 2012,Brachythemis wilsoni Kipping, 2010, Damm & Hadrys, 2009, and Agrio ­cnemisDamm bumhilli& Hadrys,T. 2009. morrisoni The fifth species newCrenigomphus to Namibia kavangoensis is Pinhey, 1952, which was found at the Kwando River in December 2010. , as well as Trithemis morrisoni T. palustris Suhling & Marais, 2010 have their type localities in Namibia (Damm & Hadrys 2009; Suhling & Marais 2010; Kipping et al. 2012). and can not be distinguished in the field so far. ● Notogomphus praetorius Two species have to be removed from the checklist: (Selys, 1878): the locality of the one single record from Namibia is unlikely to provide habitat for this species. The typical habitat is ● Trithemismountain pluvialis streams in woodland, which does not exist near Hardap Dam. We suppose wrong locality assignment. Förster, 1906 is based on observationT. pluvialis of one single male at the KwandoT. monardi River (Kappes & Nehls 2004); no specimens have been collected nor photos have been taken. Since then no records of were made. We Rhyothemisinfer that mariposa was observed, which is known from the Kwando River.

(Ris, 1913) is a special case: Pinhey (1984) assigned the type locality ‘Otiembora’ to Namibia. However, the locality cannot be localized with security since the collector, Axel W. Eriksson, did not leave notes. John Irish (in lit.) provided us with the best possible georeference, indicating that the local- ity is in Angola in close proximity to the Namibian border. The region is not well surveyed. Thus, although the species was not recorded it may occur in Namibia. We keep this species on the checklist. Libellula Supplement 13: 107-175 110 Frank Suhling & Andreas Martens

● Mastigogomphus In addition there are some taxonomicNeurogomphus changes: Due to distinct larval characters Cammaerts, 2004 formerly ● regarded as a subgenus of by Cammaerts (2004)Aeshna have been raised to genus level (Suhling et al.Zosteraeschna 2014). Aeshna minusculaPeters & Theischinger (2011) revised the former genus in Africa ● and created the monotypic genus Trithetrum, including the former T. navasi. Sympetrum navasi ● AethiothemisDijkstra & Pilgrim discrepans (2007) created the genus which is represented A.by solitaria (Lacroix, 1921), formerly . Lieftinck, 1969 is considered to be a junior synonym of Martin, 1908 (see Kipping 2010). Pinheyagrion angolense The changes reported abovePseudagrion bring the checklist helenae toP. 130fisheri species.Trithemis Some more brydeni spe- cies may be expected, particularly , which occurs in An- gola as well as in Botswana. , , and all occur in the Okavango Delta (Kipping 2010).Pseudagrion At the Orange citricola River thereTrithemis may be twodorsalis additional species, which have both been recorded at Augrabies Falls, 50 km upstream ofP. theangolense Namibian border, namely and . Finally, in streams in the lower Kunene catchment (e.g. Otjihipa Mts) one may expect , which is found in S Angola.

Figure 2. Distribution of all dragonfly records in Namibia (N = 7,488 records). The map indicates elevations from sea level (white) to highest summits (black, max. 2,573 m a.s.l.). Dark lines are ephemeral river courses, thick black lines perennial rivers.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 111

Distribution maps: Calopterygidae

Phaon iridipennis (Burmeister, 1839) 29 records, 14 localities.

Distribution maps: Chlorocyphidae

Platycypha caligata (Selys, 1853) 32 records, 7 localities.

Libellula Supplement 13: 107-175 112 Frank Suhling & Andreas Martens

Distribution maps: Lestidae

Lestes dissimulans Fraser, 1955 10 records, 9 localities.

Lestes pallidus Rambur, 1842 156 records, 121 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 113

Lestes pinheyi Fraser, 1955 14 records, 10 localities.

Lestes tridens (McLachlan, 1895) 2 records, 1 locality.

Libellula Supplement 13: 107-175 114 Frank Suhling & Andreas Martens

Distribution maps: Platycnemididae

Mesocnemis singularis Karsch, 1891 54 records, 33 localities.

Elattoneura cellularis (Grünberg, 1902) (white dot) 2 records, 1 locality; and Elattoneura glauca(Selys, 1860) (black dots) 46 records, 22 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 115

Distribution maps: Coenagrionidae

Aciagrion heterosticta Fraser, 1955 1 record, 1 locality.

Africallagma glaucum (Burmeister, 1839) 50 records, 32 localities.

Libellula Supplement 13: 107-175 116 Frank Suhling & Andreas Martens

Africallagma subtile(Ris, 1921) 6 records, 5 localities.

Agriocnemis angolensis Longfield, 1947 25 records, 10 localities.

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Agriocnemis bumhilli Kipping, Martens & Suhling, 2012 4 records, 1 locality.

Agriocnemis exilis Selys, 1872 61 records, 38 localities.

Libellula Supplement 13: 107-175 118 Frank Suhling & Andreas Martens

Agriocnemis gratiosa Gerstäcker, 1891 17 records, 9 localities.

Agriocnemis ruberrima albifrons Balinsky, 1961 6 records, 2 localities.

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Agriocnemis victoria Fraser, 1928 10 records, 4 localities.

Azuragrion nigridorsum (Selys, 1876) 32 records, 19 localities.

Libellula Supplement 13: 107-175 120 Frank Suhling & Andreas Martens

Ceriagrion corallinum Campion, 1914 11 records, 4 localities.

Ceriagrion glabrum (Burmeister, 1839) 105 records, 61 localities.

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Ceriagrion katamborae Pinhey, 1961 6 records, 4 localities.

Ceriagrion suave Ris, 1921 48 records, 26 localities.

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Ischnura senegalensis (Rambur, 1842) 306 records, 178 localities.

Pseudagrion (A) kersteni (Gerstäcker, 1869) 43 records, 13 localities.

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Pseudagrion (A) salisburyense Ris, 1921 36 records, 17 localities.

Pseudagrion (B) acaciae Förster, 1906 64 records, 37 localities.

Libellula Supplement 13: 107-175 124 Frank Suhling & Andreas Martens

Pseudagrion (B) assegaii Pinhey, 1950 10 records, 7 localities.

Pseudagrion (B) coeleste Longfield, 1947 28 records, 21 localities.

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Pseudagrion (B) commoniae (Förster, 1902) 2 records, 2 localities.

Pseudagrion (B) deningi Pinhey, 1961 23 records, 13 localities.

Libellula Supplement 13: 107-175 126 Frank Suhling & Andreas Martens

Pseudagrion (B) glaucescens Selys, 1876 52 records, 28 localities.

Pseudagrion (B) hamoni Fraser, 1955 15 records, 13 localities.

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Pseudagrion (B) massaicum Sjöstedt, 1909 118 records, 53 localities.

Pseudagrion (B) nubicum Selys, 1876 8 records, 4 localities.

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Pseudagrion (B) rufostigma Longfield, 1947 2 records, 1 locality.

Pseudagrion (B) sjoestedtiFörster, 1906 24 records, 16 localities.

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Pseudagrion (B) sublacteum (Karsch, 1893) 104 records, 58 localities.

Pseudagrion (B) sudanicum rubroviride Pinhey, 1955 19 records, 13 localities.

Libellula Supplement 13: 107-175 130 Frank Suhling & Andreas Martens

Pseudagrion (B) vaalense Chutter, 1962 6 records, 6 localities.

Distribution maps: Gomphidae

Ceratogomphus pictus Hagen in Selys, 1854 20 records, 14 localities.

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Crenigomphus cornutus Pinhey, 1956 6 records, 3 localities.

Crenigomphus hartmanni (Förster, 1898) 1 record, 1 locality.

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Crenigomphus kavangoensis Suhling & Marais, 2010 18 records, 8 localities.

Gomphidia quarrei (Schouteden, 1934) 15 records, 10 localities.

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Ictinogomphus dundoensis (Pinhey, 1961) 6 records, 4 localities.

Ictinogomphus ferox (Rambur, 1842) 105 records, 54 localities.

Libellula Supplement 13: 107-175 134 Frank Suhling & Andreas Martens

Lestinogomphus angustus Martin, 1911 42 records, 22 localities.

Lestiogomphus silkeae Kipping, 2010 1 record, 1 locality.

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Mastigogomphus dissimilis (Cammaerts, 2004) 6 records, 4 localities.

Neurogomphus cocytius Cammaerts, 2004 5 records, 4 localities.

Libellula Supplement 13: 107-175 136 Frank Suhling & Andreas Martens

Neurogomphus zambeziensis Cammaerts, 2004 3 records, 2 localities.

Phyllogomphus selysi Schouteden, 1933 12 records, 10 localities.

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Paragomphus cataractae Pinhey, 1963 10 records, 6 localities.

Paragomphus cognatus (Rambur, 1842) 5 records, 5 localities.

Libellula Supplement 13: 107-175 138 Frank Suhling & Andreas Martens

Paragomphus elpidius (Ris, 1921) 15 records, 11 localities.

Paragomphus genei (Selys, 1841) 192 records, 107 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 139

Paragomphus sabicus Pinhey, 1950 24 records, 12 localities.

Distribution maps: Aeshnidae

Anax bangweuluensis Kimmins, 1955 3 records, 2 localities.

Libellula Supplement 13: 107-175 140 Frank Suhling & Andreas Martens

Anax ephippiger (Burmeister, 1839) 98 records, 80 localities.

Anax imperator Leach, 1815 290 records, 165 localities.

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Anax speratus Hagen, 1867 51 records, 20 localities.

Anax tristis Hagen, 1867 35 records, 32 localities.

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Gynacantha manderica Grünberg, 1902 4 records, 4 localities.

Gynacantha villosa Grünberg, 1902 1 record, 1 locality.

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Zosteraeshna minuscula (McLachlan, 1896) 14 records, 8 localities.

Distribution maps: Macromiidae

Phyllomacromia contumax Selys, 1879 48 records, 33 localities.

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Phyllomacromia overlaeti (Schouteden, 1934) 4 records, 4 localities.

Phyllomacromia picta (Hagen in Selys, 1871) 39 records, 27 localities.

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Distribution maps: Libellulidae

Acisoma panorpoides ascalaphoides Rambur, 1842 53 records, 28 localities.

Aethiothemis solitaria Martin, 1908 17 records, 9 localities.

Libellula Supplement 13: 107-175 146 Frank Suhling & Andreas Martens

Aethriamanta rezia Kirby, 1889 16 records, 9 localities.

Brachythemis lacustris (Kirby, 1889) 83 records, 45 localities.

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Brachythemis leucosticta (Burmeister, 1839) 187 records, 109 localities.

Brachythemis wilsoni Pinhey, 1952 2 records, 1 locality.

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Bradinopyga cornuta Ris, 1911 8 records, 8 localities.

Chalcostephia flavifrons Kirby, 1889 7 records, 5 localities.

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Crocothemis divisa Karsch, 1898 1 record, 1 locality.

Crocothemis erythraea (Brullé, 1832) 446 records, 241 localities.

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Crocothemis sanguinolenta (Burmeister, 1839) 52 records, 21 localities.

Diplacodes deminuta Lieftinck, 1969 4 records, 4 localities.

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Diplacodes lefebvrii (Rambur, 1842) 240 records, 140 localities.

Diplacodes luminans (Karsch, 1893) 92 records, 78 localities.

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Hemistigma albipunctum (Rambur, 1842) 73 records, 43 localities.

Nesciothemis farinosa (Förster, 1898) 67 records, 39 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 153

Olpogastra lugubris Karsch, 1895 84 records, 47 localities.

Orthetrum abbotti Calvert, 1892 9 records, 7 localities.

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Orthetrum brachiale (Palisot de Beauvois, 1817) 102 records, 78 localities.

Orthetrum caffrum (Burmeister, 1839) 8 records, 7 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 155

Orthetrum chrysostigma (Burmeister, 1839) 340 records, 180 localities.

Orthetrum guineense Ris, 1910 1 record, 1 locality.

Libellula Supplement 13: 107-175 156 Frank Suhling & Andreas Martens

Orthetrum hintzi Schmidt, 1951 1 record, 1 locality.

Orthetrum icteromelas Ris, 1910 18 records, 13 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 157

Orthetrum julia falsum Longfield, 1955 69 records, 35 localities.

Orthetrum machadoi Longfield, 1955 22 records, 10 localities.

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Orthetrum robustum Balinsky, 1965 9 records, 6 localities.

Orthetrum stemmale (Burmeister, 1839) 12 records, 8 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 159

Orthetrum trinacria (Selys, 1841) 168 records, 105 localities.

Palpopleura deceptor (Calvert, 1899) 13 records, 13 localities.

Libellula Supplement 13: 107-175 160 Frank Suhling & Andreas Martens

Palpopleura jucunda Rambur, 1842 22 records, 15 localities.

Palpopleura lucia (Drury, 1773) 16 records, 15 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 161

Palpopleura portia (Drury, 1773) 3 records, 3 localities.

Pantala flavescens (Fabricius, 1798) 431 records, 291 localities.

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Parazyxomma flavicans (Martin, 1908) 11 records, 7 localities.

Rhyothemis fenestrina (Rambur, 1842) 24 records, 12 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 163

Rhyothemis mariposa Ris, 1913 1 record, 1 locality (the record is likely in Angola, but very close to the Namibian border)

Rhyothemis semihyalina (Desjardins, 1832) 52 records, 32 localities.

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Sympetrum fonscolombii (Selys, 1840) 216 records, 160 localities.

Tholymis tillarga (Fabricius, 1798) 96 records, 56 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 165

Tramea basilaris (Palisot de Beauvois, 1817) 120 records, 96 localities.

Tramea limbata (Desjardins, 1832) 3 records, 2 localities.

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Trithemis aconita Lieftinck, 1969 14 records, 8 localities.

Trithemis aequalis Lieftinck, 1969 3 records, 3 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 167

Trithemis annulata (Palisot de Beauvois, 1807) 295 records, 161 localities.

Trithemis arteriosa (Burmeister, 1839) 200 records, 100 localities.

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Trithemis donaldsoni (Calvert, 1899) 24 records, 12 localities.

Trithemis furva Karsch, 1899 18 records, 11 localities.

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Trithemis hecate Ris, 1912 63 records, 42 localities.

Trithemis kirbyi ardens (Gerstäcker, 1891) 462 records, 248 localities.

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Trithemis monardi Ris, 1931 28 records, 19 localities.

Trithemis morrisoni Damm & Hadrys, 2009 and T. palustris Damm & Hadrys, 2009 These two species can only be distin­ guished by molecular methods. Most records are therefore T. morrisoni/ T. palustris. 78 records, 29 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 171

Trithemis stictica (Burmeister, 1839) 24 records, 7 localities.

Trithemis werneri Ris, 1912 3 records, 3 localities.

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Trithetrum navasi (Lacroix, 1921) 11 records, 9 localities.

Urothemis assignata (Selys, 1872) 14 records, 3 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 173

Urothemis edwardsii (Selys, 1849) 78 records, 44 localities.

Zygonoides fuelleborni (Grünberg, 1902) 27 records, 16 localities.

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Zygonyx natalensis (Martin, 1900) 23 records, 11 localities.

Zygonyx torridus (Kirby, 1889) 56 records, 24 localities.

Libellula Supplement 13: 107-175 Distribution maps and checklist of Namibian Odonata 175

References

Brinck P. (1955) Odonata. In: Hanström, Koch F., J. Deckert & M. Uhlig (1995) Die B., P. Brinck & G. Rudebeck (eds) South entomologischen Afrika-Expeditionen des African Life: Results of the Lund Museums für Naturkunde Berlin von 1992 University Expedition in 1950-1951. Vol. II, bis 1995 als Grundlage für das Forschungs- Almqvist & Wiksell, Stockholm projekt “Zur Biodiversität von Insekten in der afrotropischen Region”. Mitteilungen Cammaerts R. (2004) Taxonomic studies des Zoologischen Museums Berlin 71: 189- on African Gomphidae (Odonata, Aniso­ 211 ptera). 2. A revision of the genus Neuro- gomphus Karsch, with the description of Martens A., R. Jödicke & F. Suhling (2003) some larvae. Belgian Journal of Entomolo- An annotated checklist of the Odonata of gy 6: 91-239 Namibia. Cimbebasia 18: 139-160 Curtis B.A. (1991) Freshwater macro- Peters G. & G. Theischinger (2011) The ge- invertebrates of Namibia. Madoqua 17: nera of the Afrotropical “Aeshnini”: Afro­ 163-187 aeschna gen. nov., Pinheyschna gen. nov. and Zosteraeschna gen. nov., with the de- Damm S. & H. Hadrys (2009) Trithemis scription of Pinheyschna waterstoni spec. morrisoni sp. nov. and T. palustris sp. nov. nov. (Anisoptera: Aeshnidae). Odonatolo- from the Okavango and Upper Zambezi gica 40: 227-249 Floodplains previously hidden under T. stic- tica (Odonata: Libellulidae). International Pinhey E. (1984) A survey to the dragonflies Journal of Odonatology 12: 131-145 (Odonata) of south Africa. Part 1. Journal of the entomological Society of Southern Af- Dijkstra K.-D.B. & E.M. Pilgrim (2007) Tri- rica 47: 147-188 thetrum, a new genus of African dragon- flies formerly placed in Sympetrum (Odo- Pinhey E. (1985) A survey to the dragonflies nata, Libellulidae). Journal of Afrotropical (Odonata) of south Africa. Part 2. Journal of Zoology 3: 77-81 the entomological Society of Southern Af- rica 48: 1-48 Kappes W. & H.W. Nehls (eds) (2004) Nami- bia-Botswana-Simbabwe. Naturkundliche Suhling F. & E. Marais (2010) Creni­ Reisenotizen 23.10.-20.11.2003. Natur- gomphus kavangoensis sp. nov. from kundliche Reiseberichte 24: 88–100 the Okavango River, Namibia (Odonata: Gomphi­dae). International Journal of Odo- Kipping J. (2010) The dragonflies and dam- natology 13: 267-276 selflies of Botswana – an annotated check- list with notes on distribution, phenology, Suhling F. & A. Martens (2007) Dragon- habitats and Red List status of the species flies and damselflies of Namibia. Gamsberg (Insecta: Odonata). Mauritiana 21: 126- Macmillan, Windhoek 204 Suhling F., O. Müller & A. Martens (2014) Kipping J., A. Martens & F. Suhling (2012) The dragonfly larvae of Namibia (Odonata). Africa’s smallest damselfly – a new Agrio­ Libellula Supplement 13: 5-106 cnemis from Namibia (Odonata: Coen­ agrio­nidae). Organisms Diversity & Evolu- tion 12: 301-306

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Libellula Supplement 13: 107-175

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