39 38 37 36 35 33 31 30 29 28 27 26 25 24 23 22 16 12 11 10 8 7 6 5 4 3 2 1 Sphaerius acaroides Sphaerius MICROSPORIDAE SPHAERIIDAE SPHAERIUSIDAE SuborderMYXOPHAGA Haliplus lineolatus Haliplus immaculatus Haliplus heydeni Haliplus furcatus Haliplus fluviatilis Haliplus lineatocollis Haliplus obliquus Haliplus confinis Haliplus caesus Peltodytes Brychius elevatus HALIPLIDAE villosus Orectochilus Gyrinus urinator Gyrinus suffriani Gyrinus substriatus Gyrinus marinus Gyrinus aeratus GYRININAE GYRINIDAE SuborderADEPHAGA Stephens Wehncke Illiger Gyllenhal (Duftschmid) Stephens Scriba Seidlitz (Panzer) (Fabricius) Aubé Mannerheim Stephens Waltl (Müller) (Müller) (Marsham) Gerhardt

Subfossil ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ●1998 ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare? [Canals, ponds, flooded gravel flooded pits]. ponds, Rare?[Canals, weedy Widespreadin ponds. VerysmallLocal ponds in detritus fen Rarepond in lakes, canals etc. ponds, Widespreadin streams, weedyWidespread in and ditches ponds. withChara of new coloniser ponds a rapid canbe Local,but lakescanals with Charophytes. and ponds, Local in increasing? LocalWeedy but ponds, flowing rarein water especiallysubmerged in gravel weirsnear and streams Largely rare,shade. on easily deep and oftenin overlooked.B nocturnal flowing Rarein water margins. fens Rarein lakes. and canals,ditches ponds, on oftenabundant Widespreadand water open especiallyLocal,largeon as lakeswatercanal such margins.on and bodies slow flowingCanalsand river margins

ecology 83 82 78 77 74 73 70 69 66 65 64 61 60 59 57 56 55 54 50 49 48 47 45 44 43 42 41 40 Haliplus fulvus Haliplus flavicollis Haliplus (= sibiricus Haliplus ruficollis Haliplus mucronatus Haliplus Ilybius chalconatus Ilybius ater Ilybius paludosus Agabus nebulosus Agabus guttatus Agabus didymus Agabus bipustulatus Agabus biguttatus Agabus uliginosus Agabus labiatus Agabus sturmii Agabus AGABINAE DYTISCIDAE hermanni Hygrobia HYGROBIIDAE PAELOBIIDAE clavicornis Noterus NOTERIDAE variegatus Haliplus laminatus Haliplus wehnckei (De Geer) ) (Gyllenhal in Schönherr) (Fabricius) (Brahm) (Paykull) (Olivier) Motschulsky (Olivier) (De Geer) (Forster) (Fabricius) (Linnaeus) Sturm (Schaller) (De Geer) Sturm (Panzer) (Fabricius) (Linnaeus) Stephens E.Ho Na Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ●1998 ● ● ● ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local. In ponds and rut pools in wooded areas, wooded in pools rut and ponds Local. In waters. ofstanding kinds all AlsoWidespread in comes light. to weedy Local,in margins moss. ofand flowing litter small flooded water, seepages in around also springheads and esecially.ponds Newsilt Widespread.In recently etc.or troughs cleared flier sites,a strong in frequent also fed spring Local streams in well-heads, outfalls of also man-made and drains. flowingLocal in water amongst marginalvegetation. flyer. a etc., strong troughs horse waterstagnant in including Ubiquitous fed waters, Rare.Calcareous ground heads spring pools mesotrophic Rarein ponds. detritus Widespreadin with silty esp. substrates. ponds Widespreadin lakes. canals,and common Widespreadand ponds, in extinct Weedystreams ponds. and rarepossiblyextinct canals. ditches, large in ponds, water. standing Local in water. standing Local in ponds. detritus Widespreadin record Rareone

ecology 140 138 137 136 135 130 127 126 124 121 120 117 111 109 108 107 106 105 104 101 99 95 94 93 92 89 87 85 84 Platambus maculatus Platambus quadriguttatus Ilybius montanus Ilybius fuliginosus Ilybius fenestratus Ilybius Hydroporus erythrocephalus Hydroporus discretus Hydroporus angustatus Hydroporus pictus Graptodytes granularis Graptodytes geminus Hydroglyphus HYDROPORINAE seminiger Hydaticus Dytiscus semisulcatus Dytiscus marginalis Dytiscus circumflexus sulcatus Acilius DYTISCINAE (= haemorrhoidalis Liopterus COPELATINAE suturalis Rhantus exsoletus Rhantus fuscus Colymbetes COLYMBETINAE Copelatus ) (Linnaeus) (Stephens) (MacLeay) (Forster) (Fabricius) (Fabricius) (Linnaeus) (Fabricius) Linnaeus (De Geer) Fairmaire & Brisout (Linnaeus) Müller, O.F. O.F. Müller, Fabricius (Lacordaire) (Linnaeus) (Fabricius) Sturm (Fabricius) (Linnaeus) E.Ho E.Ho Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ●2009 ● ● ● ● ● ● ● ● ● ●2005 ● ● ● Vc23 1939-2012 ● ● ● ● ● ●2011 ●2011 ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread and common in ponds and lakes, especially and ponds. commonacidic heath Widespreadand ponds in in pools. and Veryfed spring Local puddles in lakes, acidic waters canals,and avoiding common Widespread,and ponds, in ditches. and veryponds in locally abundant Rare.Richly weeded ponds pools. peat Rare,most and from records waterponds stagnant heath in situations open and shade in pools rich increasingly n colonist recntfound pools. peat Rare,most and from records waterponds stagnant heath in water standing especially of in kinds, all where as arelarvaeamphibians Ubiquitous breeding, tadpoles. specialise in silty withsubstrates. open Scarce,increasing?ponds In aLocal wide in variety of fish stocking. waters,ofintolerant still but pools. bog and weedyLocal. In ponds detritus of newrecently coloniser and Widespreadclearedgood pools. silt marshy and ponds areas. detritus Local in ofkinds. all species of ponds Frequent flowingLocal in water, streams, rivers. Tolerantof and shade. ponds. detritus Local in Localespeciallywatersacid in weedy and ponds margins Widespreadin offlowing water. Also comes light. to lakes Veryareas. especiallylargeLocal. In heathland ponds, in

ecology 200 199 197 196 194 188 187 184 181 180 179 178 175 170 165 164 161 160 159 156 150 148 145 143 141 Hydroporus neglectus Hydroporus memnonius Hydroporus marginatus Hydroporus incognitus Hydroporus gyllenhalii Hydroporus ferrugineus Hydroporus LACCOPHILINAE ovatus Hyphydrus versicolor (s.str.) Hygrotus inaequalis (s.str.) Hygrotus nigrolineatus (Coelambus) Hygrotus impressopunctatus (Coelambus) Hygrotus confluens (Coelambus) Hygrotus dorsalis Suphrodytes duodecimpustulatus Stictotarsus lepidus Stictonectes halensis Scarodytes lineatus Porhydrus Oreodytessanmarkii elegans Nebrioporus umbrosus Hydroporus tessellatus Hydroporus striola Hydroporus pubescens Hydroporus planus Hydroporus palustris Hydroporus obscurus Hydroporus nigrita Hydroporus (Linnaeus) (Gyllenhal) (Fabricius) (Fabricius) (Fabricius) (Olivier) (Fabricius) (Panzer) (Sahlberg, C.R.) sensu lato (Linnaeus) Sturm Schaum (Gyllenhal) (Drapiez) (Gyllenhal) Sharp Schiødte (Duftschmid) Stephens Nicolai (Schaller) (Fabricius) (Fabricius) H/Rom E.Ho Nb (Fabricius) Subfossil (Steven) ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH (Schaller) ● ● ● ● ● ● ● ● ● ●1998 ● ● ● ● ● ● ●1996 ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●1906 ●1998 ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread and common in ponds, canals, and lakes. canals,and common Widespreadand ponds, in canalmargins. gravelLocal in ponds, lakes. canals,and common Widespreadand ponds, in Recentimmigrant new increasing in ponds. field ponds, Widespread in withsilty of new substrates. coloniser ponds Widespreada rapid review recently under status split streams riversLocal and in submerged especiallyledges under etc.baresections in Extinct?? Local,increasing etc.] ditches Rare.Ponds, Rare.Gravelly tangles,tolerant. streams shade root in etc. permanent pits riversLocal streams. and in in Also ponds occasionally deep in mossy rarein fen pools puddles. water isolated of in newrecently also troughs, coloniser and Widespread.Rapid cleared ponds water. standing Local in water. standing acid Widespreadin water of in newrecently also troughs. coloniser and Widespreadrapid cleared ponds marshy and ponds areas bydetritus Widespreadin lakes flowing and water. Sphagnumpools] in [ aLocal variety in of small waterseasonalpools. including bodies acis waters Shaded seepages and leaf flooded litter. in pools woodland Widespreadin streams. winterbourne Semiin subterranean marshy and streams. beside pools pools shaded Local in ponds. RAREacidic heath in heads spring Subterranean rarein

ecology 251 250 247 246 245 244 243 241 240 239 238 236 232 231 230 229 225 224 223 219 217 216 215 209 207 206 205 202 201 Cicindela campestris Cicindela CICINDELINAE CARABIDAE minutus Laccophilus hyalinus Laccophilus Nebria salina Nebria brevicollis Nebria terminatus Leistus ferrugineus Leistus fulvibarbis Leistus spinibarbis Leistus rufomarginatus Leistus NEBRIINI Cychrus caraboides CYCHRINI violaceus Carabus problematicus Carabus nemoralis Carabus monilis Carabus granulatus Carabus arvensis Carabus inquisitor Calosoma CARABINI CARABINAE crepitans Brachinus BRACHININAE Fairmaire & Laboulbène Fabricius Herbst (Fabricius) Linnaeus Dejean (Hellwig in Panzer) (Hellwig (Fabricius) Müller, O.F. Müller, (Linnaeus) (Linnaeus) Linnaeus (Linnaeus) (Linnaeus) Linnaeus (Linnaeus) (De Geer) Herbst (Duftschmid)B215 Ir/Rom Rom Rom E.Hol Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ? Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ●1953 ●2003 ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ? modern Oxon Local on open sandy ground.. sandy ground.. open Localon common. Widespreadand dampLocal shadyin leaf places Aclimber. in litter. good heaths country and wooded Local in sites. ruderal disturbed hedgerows, leafLocal woods, and in etc.in litter gardens etc.woods, in Frequent 20 A districts. flightless wooded Widespreadpredator. in snail habitats. more parks,gardens,and common open woods, Widespreadand in occurring scarceprimarily Oxon. calcareous grasslandin on dry in areaslocal habitats. open Widespreadin records. dampLocal 1980 meadows,in post 10 oaks etc.] on [Arboreal Extinct? possiblyextinct? records, 1980 chalky post Rare on no soils, ground. disturbed and drywoodlands scattered in soils, open Localon lakes. canals,and common Widespreadand ponds, in slow Local,flowingin water, occasionally ponds, in extinct Specimen labelled Oxford in JJW coll. Oxford JJW in labelled Specimen extinct th century colonist (first noted in the 1940s), now widespread especially in woods in litter under trees. nowwidespread under especiallylitter in woods 1940s), the in in (first centurynoted colonist

ecology 302 301 299 298 293 288 286 284 281 277 276 275 274 273 272 271 269 267 266 265 264 263 262 261 260 258 257 Notiophilus palustris Notiophilus germinyi Notiophilus biguttatus Notiophilus aquaticus Notiophilus aesthuans Notiophilus NOTIOPHILINI Trechus quadristriatus Trechus obtusus Trechus secalis Trechus rivularis TRECHINI Broscus cephalotes BROSCINI Dyschirius politus Dyschirius luedersi Dyschirius globosus Dyschirius aeneus fossor Clivina collaris Clivina SCARITINI pilicornis Loricera LORICERINI riparius Elaphrus uliginosus Elaphrus cupreus Elaphrus ELAPHRINI substriatus Notiophilus rufipes Notiophilus quadripunctatus Notiophilus (Linnaeus) (Paykull) (Herbst) Erichson (Gyllenhal) Duftschmid (Linnaeus) (Dejean) (Dejean) (Fabricius) (Linnaeus) Wagner Curtis (Herbst) Fabricius (Duftschmid) Fauvel (Linnaeus) Motschulsky (Schrank) (Fabricius) Waterhouse, G.R. Dejean IrAge E.Hol Ir.Age E.Hol Ir.Age Br.Age E.Hol Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●1986 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ? ● ● ● ● ? modern Oxon Frequent in open country including gardens. country including open in Frequent fens and leaf woods in in local litter Rdampshady places mires] and [fens etc heaths sandpits, in bareground open Localon Local,wetlands, sandypartly in river subterranean etc. banks v.rarewetlands, sandypartly in river subterranean etc. banks largedamp especially litter in gardens.in also woods, Frequent in Localatmargins ofmarshywater in and ground. Veryfens rare in Localatmargins ofmarshywater in and ground. gardens. dry country including open in Frequent leafLocal moist,gardens. oftenshadyin in litter places,including Rare etc.leaf woods in in litter Frequent dry areas. open Rare in gardens. including situations ofopen kinds all in Frequent rare

ecology 352 351 350 349 348 347 346 345 341 337 334 330 329 328 327 323 322 320 318 316 314 310 308 307 306 305 303 Tachys (Paratachys Tachys BEMBIDIINI micros Trechoblemus discus Blemus longicornis Thalassophilus Trechus rubens Bembidion femoratum Bembidion (= deletum Bembidion decorum Bembidion (= bualei Bembidion bruxellense Bembidion tibiale Bembidion atrocaeruleum Bembidion varium Bembidion dentellum Bembidion punctulatum Bembidion properans Bembidion lampros Bembidion nigricorne Bembidion Ocys quinquestriatus Ocys harpaloides stierlini Asaphidion flavipes Asaphidion curtum Asaphidion parvulus Elaphropus micros (Paratachys) Tachys andreae nitidulum ) ) ) (Fabricius) (Fabricius) (Audinet-Serville) Jacquelin du Val (Duftschmid) (Olivier) (Heyden) Audinet-Serville ) (Herbst) (Heyden) (Linnaeus) (Zenker in Panzer) (Gyllenhal) (Dejean) bistriatus (Herbst) (Stephens) (Thunberg) Gyllenhal Sturm Wesmael Drapiez (Stephens) (Sturm) (Fischer von Waldheim) (Duftschmid) ● Nb Ir.Age Ir.Age Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ●1953 Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local in open sandy places, including sandpits and disturbed areas disturbed and sandpits sandy places,including open Local in sediments]. exposed waternear local [on baregravel on local marshy margins and Rareon ofponds places. etc ponds near Commonbareground on ?margins[Atwater]. ofrunning habitats. open Local in fields. including habitats open various Widespreadon walls Rare?old especially On ivy under (Hedera helix) nests. old including debris suspended in barkand trees, under on woods Widespreadin Local.Recently species, widespread split especially sites.chalky ruderal on clay in also and and soils, a varietyRare in primarily of habitats vicinity in of water. bareusually Localclayey on gardens. and rides woods, in soils gardens, including Rare? Increasing? ground Open mammal Rare?[In burrows] or runs mammal Rare?[In burrows] or runs ofmammals. runs Rarein margins] [Wetland

ecology 400 398 397 396 392 391 390 389 388 386 385 383 381 380 379 378 374 372 371 368 364 363 362 359 358 357 354 353 Bembidion tetracolum Bembidion stephensii Bembidion saxatile Bembidion monticola Bembidion lunatum Bembidion fluviatile Bembidion PTEROSTICHINI atrorufus Patrobus assimilis Patrobus PATROBINI chalceus Pogonus POGONINI (= mannerheimii Bembidion lunulatum Bembidion guttula Bembidion biguttatum Bembidion aeneum Bembidion obtusum Bembidion articulatum Bembidion doris Bembidion quadripustulatum Bembidion quadrimaculatum Bembidion fumigatum Bembidion clarkii Bembidion assimile Bembidion gilvipes Bembidion (= illigeri Bembidion genei unicolor ) ) (Panzer) (Dawson) (Marsham) Chaudoir (Fabricius) Netolitzky (Ström) Sturm Gyllenhal Germar Gyllenhal (Duftschmid) Dejean Audinet-Serville Crotch (Geoffroy in Fourcroy) Sturm Say Say (Fabricius) (Duftschmid) (Panzer) Sahlberg, C.R. (Linnaeus) Audinet-Serville E.Hol E.Hol E.Hol Ir.Age H/Ir.age Ir.Age E.Hol Ir.Age Ir.Age Subfossil ● * ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon widespread in wetwidespread in woodlands. river margins. and bogs, especiallyin woodlands dampLocal and litter in in dampwidespread habitats in wetlands,Local marginswaterlogged lakes,in arablein land. also and oflarge riverspasture ponds, and marshland.Local in widespreadoftenwaternear arable land. including dry, land, oftencultivated open Localon water.near damp Localbareground on marshes, Rarein wet woodland. Scarce.wetland sparsely open vegetated Widespreadon sandy areas fields including wetlands dampplaceswater r nea Scarcewetland quarries. and pits in ground barish open Localon by sand streams. rivers and and shingle, water,nearopen oftenon bareground Widespreadon clayey open Rareon ground. [On ? banks ofrivers andstreams].

ecology 439 438 437 436 435 434 433 432 431 430 428 427 426 425 422 421 420 419 418 415 413 412 410 409 405 404 402 401 Pterostichus madidus Pterostichus versicolor Poecilus lepidus Poecilus kugelanni Poecilus cupreus Poecilus pumicatus Stomis (= cinctus Calathus ambiguus Calathus (= rotundicollis Calathus (= vivalis Synuchus (= depressus Platyderus SPHODRINI parallelepipedus Abax strenuus Pterostichus diligens Pterostichus vernalis Pterostichus nigrita Pterostichus minor Pterostichus gracilis Pterostichus anthracinus Pterostichus melanarius Pterostichus oblongopunctatus Pterostichus niger Pterostichus macer Pterostichus longicollis Pterostichus mollis piceus nivalis ruficollis in part) part) in ) ) ) (Leske) (Illiger) (Illiger) Motschulsky (Linnaeus) (Panzer) (Schaller) (Sturm) (Gyllenhal) (Marsham) (Panzer) (Paykull) (Paykull) (Dejean) (Sturm) (Audinet-Serville) (Panzer) (Fabricius) (Panzer) Dejean &(Piller Mitterpacher) (Duftschmid) (Illiger) (Illiger) (Panzer) (Fabricius) E.Hol E.Hol Rom H/ax Rom Rom Rom Ir.Age Rom H/Sax Rom E.Hol H/Ir.Age Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ●1999 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare in open sandy places open Rarein gardens.Widespread etc.especiallylogs Also in areas,occurs wooded under in sites. dry disturbed areas open Local including in waste gardensLocal and in ground. wet in wetlands. frequent heath/boggyLocal. Most other areas, and marshes. in and heaths on habitats open Local in marshLocal wet in and woodlands. marshLocal carr. in and carr. and Marsh wetlands Rarein aLocal wide in variety of fromdamp marshes, habitats etc.woodland, bogs partly habitats. other wooded and parkland Widespreadin partly subterranean. Rare,though areas. ruderal including habitats open Widespreadin areas. ruderal including habitats open Widespreadin areas ruderal including habitats open Widespreadin soils?]. rich grasslands,Local gardens [on in

ecology 477 476 475 474 473 472 471 470 469 465 464 461 460 459 458 457 456 454 453 452 451 450 449 448 447 445 444 442 441 PLATYNINI terricola Laemostenus leucophthalmus Sphodrus mollis Calathus melanocephalus Calathus fuscipes Calathus Amara plebeja Amara tenebrioides Zabrus ZABRINI viduum Agonum versutum Agonum sexpunctatum Agonum nigrum Agonum muelleri Agonum marginatum Agonum (= emarginatum Agonum thoreyi Agonum scitulum Agonum piceum Agonum micans Agonum gracile Agonum fuliginosum Agonum livens Batenus assimilis Platynus dorsalis Anchomenus albipes Paranchus obscurus Oxypselaphus rotundatus Olisthopus moestum ) ) (Gyllenhal) (Gyllenhal) (Marsham) Sturm Dejean Nicolai Dejean (Panzer) (Linnaeus) (Herbst) (Goeze) Dejean (Paykull) (Fabricius) Sturm (Goeze) (Panzer) (Pontoppidan) (Linnaeus) (Herbst) (Paykull) (Gyllenhal) (Linnaeus) (Herbst) (Linnaeus) (Linnaeus) H/Ir.Age Ir.Age E.Hol E.Hol Rom E.Hol E.Hol E.Hol Rom Rom Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ●2000 ● ● ● ● ● ● ● ● ●2001 ● ●1988 ● ● ● ● ● ●1945 ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●2003 ●1976 ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local in dampLocal clayeyin areas. marshy Rarein areas. marshy Rarein wetlands. Westonfen rides. woodland open in also wet heathland open Local,on marshes. Rare in dryland Local cultivated in areas including Localatmargins etc.water stones ofbareareas standing in under marshyLocal in areas. marshesLocal in boggy and areas especiallyheathy commons.on marshes. Rarein marshes]. ?[In Rare[Wetlands]. marshes Rarebogs. in richer and tussocks. and litter marshlandLocal in in Rareby water moss. barkin etc.under woods Widespreadin etc.grassland, woods Widespreadin wetlands, especiallyWidespread in pools. carr areaswoodland and marsh. wetand dampLocal in litter woodland in areas. ruderal including habitats open Widespreadin come records Local.bark.Most from warrens, under occurs rabbit also near cellarsstables, etc.declining]. in ?[Synanthropic farmland areas gardens. common Widespread including and and open in

ecology 513 512 511 508 507 506 504 503 502 501 499 495 494 493 492 491 490 489 488 486 484 483 482 481 480 479 Amara eurynota Amara curta Amara convexior Amara communis Amara anthobia Amara aenea Amara Harpalus cupreus Harpalus attenuatus Harpalus anxius Harpalus affinis Harpalus rufipes Harpalus HARPALINI convexiusculus Curtonotus aulicus Curtonotus equestris Amara fulva Amara consularis Amara apricaria Amara praetermissa Amara bifrons Amara tibialis Amara spreta Amara similata Amara ovata Amara nitida Amara montivaga Amara lunicollis Amara lucida Amara familiaris Amara (Müller, O.F.) (Müller, Dejean (Fabricius) Sturm (De Geer) Dejean (Duftschmid) (Paykull) (Gyllenhal) (Gyllenhal) (Duftschmid (Panzer) Villa & Villa (Schrank) (Paykull) (Duftschmid) Schiødte (De Geer) (Duftschmid) Stephens (Duftschmid) Dejean (Panzer) Sturm (Panzer) (Sahlberg, C.R.) Stephens (Marsham) RD1 Ir.age Rom Rom Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread and common. Widespreadand dry sites especially disturbed sites.Also open ruderal comes Locallight. on to dry sites especially disturbed sites.Also open ruderal comes rarelight. on to composites. areas Widespreadespecially withabundant disturbed in downland. and heaths Rareon atlight. gardens.Also abundant sitescanbe and ruderal areas including disturbed Local in downland. on also Localbut primarily heathland, on dry Glebesites. open Local (JM in areas dry Widespread within crucifers. open areas. disturbed open Widespreadin chalk downs [Heathland] sites. ruderal dryLocal in areas disturbed oftenin gardens. and heathland areas including open Local in areas ruderal including habitats open Widespreadin areas vegetation. withruderal disturbed open Rarein sandy places. open Rarein gravelly Localon sites,& downland. commons. and downs Rare on dry soils. open Rareon areas. ruderal including habitats open Widespreadin

ecology 569 568 567 564 563 562 553 551 550 549 548 547 546 545 544 543 542 541 540 539 537 536 532 531 527 526 522 519 516 Harpalus smaragdinus Harpalus rufipalpis Harpalus rubripes Harpalus latus Harpalus honestus Harpalus dimidiatus Harpalus Bradycellus sharpi Bradycellus ruficollis Bradycellus harpalinus Bradycellus (= caucasicus Bradycellus placidus Trichocellus binotatus Anisodactylus schaubergerianus Ophonus rupicola Ophonus rufibarbis Ophonus puncticollis Ophonus puncticeps Ophonus parallelus Ophonus (= melletii Ophonus (= laticollis Ophonus cordatus Ophonus (= stictus Ophonus azureus Ophonus ardosiacus Ophonus tardus Harpalus collaris melleti punctatulus obscurus ) ) ) (Linnaeus) ) Stephens (Panzer) (Heer) (Fabricius) (Sturm) Mannerheim (Duftschmid) (Duftschmid) Joy Joy (Duftschmid) Sturm (Fabricius) (Dejean) Stephens Lutshnik (Rossi) (Paykull) (Stephens) (Gyllenhal) (Duftschmid) (Chaudoir) (Audinet-Serville) (Fabricius) (Puel) Rom SF Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●1984 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon rare in damp rareshadyin places. sandy especiallyRaresoils on heathland. on heather. oftenunder sandy Local especiallysoils on heathland on marshy In local areas. marshy Rarein areas. dry places. open Rarein Chalkgrassland] aLocal variety in especially of habitats sandy soils. on traps. light areas, atMV frequentruderal also dry sites,including open rarein Formerlychalk grassland. on dry Raresandyon chalk grasslands.. and chalk downland. open Rareon Local,traps light increasing? Grassland, sites,gardens etc,atMV frequent ruderal some on sandy heaths. abundant Localthough sandy soils. disturbed Rareon sandy rareground on especiallysoils sandydry Localheaths. open on on of many ground open kinds. Widespreadon chalk downland. Rareon

ecology 610 609 608 606 603 602 601 600 599 597 596 595 594 593 592 590 588 587 586 585 584 583 582 580 579 577 573 570 Acupalpus flavicollis Acupalpus exiguus Acupalpus dubius Acupalpus teutonus Stenolophus mixtus Stenolophus verbasciBradycellus Panagaeus cruxmajor Panagaeus bipustulatus Panagaeus PANAGAEINI dilatatus Badister sodalis Badister unipustulatus Badister meridionalis Badister (= Badister bullatus Badister punctatulus Licinus depressus Licinus LICININI helopioides Oodes OODINI vestitus Chlaenius nitidulus Chlaenius nigricornis Chlaenius CHLAENIINI consputus Anthracus (= parvulus Acupalpus meridianus Acupalpus bipustulatus dorsalis sp. ) ) (Duftschmid) in part) in (Schrank) Chaudoir (Paykull) Schilsky (Fabricius) (Paykull) Dejean (Schrank) (Herbst) (Fabricius) (Sturm) (Duftschmid) (Sturm) (Duftschmid) Puel (Schrank) (Fabricius) (Linnaeus) (Linnaeus) Bonelli (Fabricius) Rom E.Hol E.Hol E.Hol E,Hol Rom Rom Rom Subfossil ● ● ● ● ● ● ● ? ● ● ● ● ● ● ● ● Vc22 VCH ● ●2000 ● ● ●2002 ● ● ● ●2000 ●2012 ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare in open dry places in grasslands including ruderal sites. Shotover (VCH) sites.Shotover ruderal dry grasslands including places open in Rarein wetlands. Rare in marshy and wet in woodland, areas. fen carr and Marsh, Rarein large wastecommon and gardens. Widespreadand including ground calcareous ground Rareopen chalk downland. open Rareon wetlands Rarein well etc.avoiding stones Localvegetated atmargins under ground. pits ofwater old in margins Rarewetlands, pond marshy Rarein areas, comes also light. to boggyLocal in wetlands. ground]. cultivated clay areas including Rare?sand on [Open and boggy Rarein areas marshes. Rarein marshLocal wet in areas. and heath stones. under margins and litter in wetlands,Local pond in sandy Alsotraps. Local atlight frequentsoils. on

ecology 658 657 656 655 654 653 652 647 646 645 644 643 641 640 637 635 634 633 632 631 630 629 624 623 618 617 613 612 Demetrias atricapillus Demetrias imperialis Demetrias chlorocephala Lebia LEBIINI nigriceps Perigona PERIGONINI Helophorus porculus Helophorus nubilus Helophorus HELOPHORIDAE HYDROPHILOIDEA SuborderPOLYPHAGA melanura Odacantha ODACANTHINI truncatellus Syntomus obscuroguttatus Syntomus foveatus Syntomus Microlestesminutulus Microlestesmaurus quadrisignatus Philorhizus melanocephalus Philorhizus (= spilotus Calodromius quadrimaculatus Dromius meridionalis Dromius angustus Dromius agilis Dromius linearis Paradromius quadrinotatus (Fabricius) ) ) Brullé (Geoffroy in Fourcroy) (Dejean) B641 (Dejean) B641 (Sturm) Fabricius (Hoffmann, J.) (Germar) Bedel (Olivier) (Illiger) (Illiger) (Linnaeus) Dejean (Linnaeus) (Goeze) (Linnaeus) (Linnaeus) (Duftschmid) (Dejean) (Dejean) Rom E.Hol Rom Rom Rom H/Rom E.Hol Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Open, often cultivated ground, at roots of plants]. atroots ground, oftencultivated Open, of plants]. atroots ground, oftencultivated Open, sparsely open Local vegetated in areas. dampLocal areas.in sandy areas. disturbed dry and heaths areas open Widespread on in ground. Recentimmigrant,on disturbed bareand downland. open areas, dry including in open ground the Localon trees. deciduous ?On tussocks. in or commonground Widespreadthe and on bark. oftenunder branches, and boughs moribund and dead treeson esp. deciduous Widespreadon bark. oftenunder branches, and boughs moribund and dead on esp. pine trees and deciduous Widespreadon conifers. and Local.hardwoods Arboreal on pines. Rareon conifers. trees, including Widespreadon grass Widespreadin tussocks etc. grassy Widespreadrough amongst in litter areas. Rare.Grassland with Hypericum. Recentimmigrant. compost/manure Increasing in heaps.

ecology 710 707 706 705 701 700 699 698 697 696 695 688 684 683 682 679 677 676 675 672 671 669 668 667 666 664 662 659 Helophorus brevipalpis Helophorus arvernicus Helophorus grandis Helophorus glacialis Helophorus aequalis Helophorus rufipes Helophorus Chaetarthria seminulum Chaetarthria Berosus signaticollis Berosus luridus Berosus affinis lutescens Anacaena limbata Anacaena globulus Anacaena bipustulata Anacaena HYDROPHILINAE HYDROPHILIDAE elongatus Hydrochus angustatus Hydrochus HYDROCHIDAE obscurus Helophorus nanus Helophorus minutus Helophorus griseus Helophorus granularis Helophorus flavipes Helophorus dorsalis Helophorus Brullé (Linnaeus) (Fabricius) Sturm (Paykull) (Bosc d'Antic)(Bosc Herbst (Stephens) Illiger Illiger Fabricius (Marsham) (Charpentier) Fabricius Thomson, C.G. Villa (Schaller) Mulsant (Marsham) Germar (Linnaeus) Mulsant Bedel (Herbst) Rom Rom Rom E.Hol Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ? ? Vc23 1939-2012 ●1998 ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local in mud and moss in richer bogs and fen areas and moss bogs richer mudLocal and in in marshy Rarein pools marshy Rarein pools vegetation,especially with little gravel Local,claynew sand, increasing and in ponds in pits. pools. and ponds Widespreadin pools. and ponds Widespreadin larvae leaf in especially litter pools, woods. and in Commonponds in lakes,canals. atmargins litter and mudLocal and ofin ponds, weedy Rarein ponds. detritus especiallyweedyLocal heaths. in on ponds detritus especially very commonponds, avoiding smalland in woodland, water.acid leaf-filledin pools Widespreadin puddles fen marshy areas]Rare in [and pools A flier. strong puddles. and pools Commonponds, in vegetation. withVerylittle newLocal in ponds commons. and woods, in as winter activeshallow pools in under-recorded Rare,but heathland. on pools peat and pools acid in Local.Frequent pools. rut and muddy ponds Rare ) in woodland of newephemeral coloniser and A pools. flierrapid strong and puddles. and pools Commonponds, in atmargins silt mud Rare.and In of rivers. of newephemeral coloniser and A pools. flierrapid strong and puddles. and pools Commonponds, in of plants]. atroots dry On ground

ecology 761 760 759 758 757 756 754 750 749 748 745 744 742 741 740 738 735 733 731 730 728 727 726 724 723 717 713 712 711 Enochrus nigritus Enochrus melanocephalus Enochrus coarctatus Enochrus marginellus Cymbiodyta (= simillima Chaetarthria Cercyonlateralis Cercyonlaminatus (= Cercyonimpressus Cercyonhaemorrhoidalis Cercyongranarius Cercyonconvexiusculus orbiculare Coelostoma SPHAERIDIINAE striatulus Laccobius sinuatus Laccobius minutus Laccobius colon Laccobius bipunctatus Laccobius piceus Hydrophilus fuscipes Hydrobius Helochares punctatus Helochares lividus testaceus Enochrus quadripunctatus Enochrus ochropterus Enochrus (= atomarius isotae seminulum ) ) in part) part) in (Stephens) (Marsham) (Sharp) (Forster) Erichson (Sturm) Sharp (Linnaeus) (Linnaeus) Motschulsky (Fabricius) (Linnaeus) (Fabricius) (Gredler) Sharp (Fabricius) (Fabricius) Vorst &Cuppen (Marsham) Stephens (Fabricius) (Fabricius) (Olivier) (Herbst) Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread in dung, and carrion. and dung, Widespreadin dung. herbivore Widespreadin dung. herbivore Widespreadin Thames tussocksRare on In floodplain. atmarginsdetritus Local of ofwater kinds allin bodies. flowing atmargins and silt Localwater. in ofponds etc. newLocal in ponds scarceLocal and in. ponds Rarein aweedy variety in and Frequent of ponds flowing water. Extinct vegetation. withlittle new increasing in ponds Widespread and vegetation. withlittle new increasing in ponds Widespread and ponds. detritus Local in VeryweedyLocal in marshy areasmesotrophic very and ofavoiding bogs acidic conditions. ponds weedy Rarein base rich-fen areasweedy and mesotrophic and ponds. Veryincreasingly Local new but in frequentsilty ponds. weedyLocal in margins ponds. rich ofdetritus marshy and ponds lake margins.detritus Local in

ecology 801 798 796 795 794 793 789 788 786 785 784 782 779 778 777 776 775 774 773 772 771 770 769 768 767 766 765 764 763 (= Cercyonobsoletus (= Cercyonnigriceps Cercyonmelanocephalus Cercyonmarinus Acritus nigricornis Acritus dissectus Plegaderus perpusillus Abraeus granulum Abraeus ABRAEINAE HISTERIDAE scarabaeoides Sphaeridium marginatum Sphaeridium lunatum Sphaeridium bipustulatum Sphaeridium minutum Cryptopleurum immaculatum Megasternum (= concinnum Megasternum Cercyonustulatus Cercyonanalis Cercyonunipunctatus Cercyontristis Cercyonterminatus Cercyonsternalis Cercyonquisquilius Cercyonpygmaeus obscurum lugubris atricapillus ) ) (Illiger) (Illiger) (Paykull) ) Thomson, C.G. (Sharp) (Preyssler) (Gyllenhal) (Marsham) (Hoffmann, J.) (Illiger) (Illiger) Erichson (Marsham) (Marsham) (Linnaeus) Erichson (Linnaeus) Fabricius (Fabricius) (Linnaeus) (Marsham) Fabricius Fabricius (Linnaeus) (Stephens) Rom Rom Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local in rotting vegetation, haystacks rotting Local in etc. especially wood fungoid barkand beech. Localunder hardwoods. rotting on wood/parkland ancient Rarein especially dung ofherbivore Local cattle. that in especially dung ofherbivore Local cattle. that in especially dung ofherbivore Local cattle. that in especially dung ofherbivore Local cattle. that in compost. and dung Widespreadin common ofWidespreaddecaying fungi.kinds and all and in vegetation, dung, atmarginswater.detritus Local ofstanding in sewageAlso in filter beds. vegetation. rotting other and compost heaps, dung, Widespreadin especially dung herbivore occasionallycow,Widespread carrion. in and in pools acid avoiding ponds marshLocal in around litter compost heaps. and Rare?dung In VerymarshLocal in atmargins litter ofponds dung. Widespreadin dung. herbivore in carrion. and dung, Widespreadin Rare?dung. In dung. herbivore Widespreadin etc.oftenwater..near carrion wet Local dung, in litter,

ecology 854 853 849 847 846 844 843 839 838 837 836 834 833 832 830 829 828 827 826 825 822 821 820 813 811 807 806 805 Gnathoncus buyssoni Gnathoncus virescens Saprinus semistriatus Saprinus aeneus Saprinus SAPRININAE Margarinotus purpurascens Margarinotus neglectus Margarinotus merdarius Margarinotus [ (= brunneus Margarinotus HISTERINAE striatus Onthophilus punctatus Onthophilus ONTHOPHILINAE flavicornis Paromalus pumilio Carcinops Kissisterminimus pygmaeus Dendrophilus punctatus Dendrophilus DENDROPHILINAE (= paykulli Myrmetes nannetensis Gnathoncus schmidti Gnathoncus Margarinotus striola Margarinotus impressus piceus+B1308 ) (Fabricius) (Laporte) Kanaar (Erichson) B861 (Paykull) ) (Forster) Reitter Auzat (Sahlberg, C.R.)] (Scriba) (Herbst) (Müller, O.F.) O.F.) (Müller, (Germar) (Fabricius) (Herbst) (Linnaeus) (Hoffmann, J) (Marseul) (Herbst) Rom Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ? Vc22 VCH ● ● ● Vc23 1939-2012 ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare on ground especially sandy ground soils. on Rare on fungi, carrion. rotting and dung herbivore in Local especially dung Localhorse. on decaying in barkand timber Widespreadespeciallyunder beech. kitchens! Often compost entershot heaps. gardens etc.around in Local.Synanthropic dry in grassland. Often of Localplants under atroots formerly Extinct ofnests in Also etc.nests with ant old trees in hollow Local in , woodland. in nests bird old trees and hollow Rarein on [Parasitic carrion. and dung Localon carrion. and dung on Phaedon especially water-cress on Formica rufa Formica . Nasturtium Lasius fuliginosus Lasius

Rumexacetosella ecology . Increasing?]. . . . 895 894 893 891 890 889 888 887 886 884 883 881 880 879 877 875 874 872 871 870 865 864 863 862 858 857 856 855 Atholus bimaculatus Atholus Histerunicolor Histerbissexstriatus marginatus Margarinotus (= ventralis Margarinotus Ochthebius aeneus Ochthebius dilatatus Ochthebius bicolon Ochthebius Enicocerus exsculptus OCHTHEBIINAE truncatellus Limnebius papposus Limnebius nitidus Limnebius aluta Limnebius testacea Hydraena riparia Hydraena pygmaea Hydraena pulchella [Hydraena nigrita Hydraena gracilis Hydraena flavipes Hydraena HYDRAENINAE HYDRAENIDAE STAPHYLINOIDEA duodecimstriatus Atholus (= minutissima carbonarius Linnaeus ) Bedel ) Germar Kugelann Germar (Marsham) Sturm Curtis Stephens Germar Waterhouse Fabricius Mulsant Germar ] Stephens (Linnaeus) (Germar) (Marseul) (Thunberg) (Schrank) (Erichson) E.Hol E.Hol E.Hol E.Hol E.Hol E.Hol Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● Vc23 1939-2012 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Extinct Verymargins. muddyLocal in tricklespond and VeryLocal atmargins ofstreams trickles and streams In mossy etc on boulders streams. Widespreadatmargins and ofponds marshfen and flooded litter In 1993) atmargins litter Early (Robinson, and ofstreamsE.Holocene Mud ponds. and puddles and pools leaf woodland flooded Localin litter in wetlands. and ponds in Frequent flowingLocal in water small especiallyin streams. and springheads near dung. on dung. on fungi, carrion. rotting and dung herbivore Widespreadin cow dung [In ) nests Mole carrion vegetation and rotting dung, in

ecology fscyd 952 951 950 949 946 945 944 943 936 931 928 927 926 923 922 921 920 919 918 916 912 911 910 909 908 907 904 898 PTENIDIINI PTILIINAE PTILIIDAE pusillus Ochthebius minimus Ochthebius Ptinella denticollis Ptinella cavelli Ptinella britannica Ptinella aptera Ptinella PTINELLINI minutissimum Millidium halidaii Micridium myrmecophilum Ptilium exaratum Ptilium foveolata Oligella spencei Ptiliolum fuscum Ptiliolum kunzei Ptiliola brevicollis Ptiliola PTILIINI nitidum Ptenidium pusillum Ptenidium punctatum Ptenidium longicorne Ptenidium fuscicorne Ptenidium intermedium Ptenidium gressneri Ptenidium pilosellum Nossidium (Heer) (Guérin-Méneville) (Broun) (Allibert) (Erichson) (Allibert) (Allibert) (Matthews, A.) (Matthews, (Fairmaire) Matthews, A. Matthews, (Heer) (Matthews, A.) (Matthews, (Gyllenhal) B949 Stephens Erichson (Fabricius) Erichson (Marsham) Fuss (Gyllenhal) Wankowicz (Allibert) (Ljungh)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● Vc23 1939-2012 ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon ? In grass ? In heaps. decaying on fungi silty Rarein margins ofnewly cleared ponds. moss. and litter, atmargins in Frequent ofponds

ecology 999 998 997 996 994 993 989 987 986 981 978 977 976 975 974 969 968 965 963 962 961 960 959 958 956 955 954 953 Pteryx suturalis Pteryx (= simsoni Ptinella limbata Ptinella errabunda Ptinella SOGDINI LEIODINAE LEIODIDAE grandicollis Acrotrichis thoracica Acrotrichis strandi Acrotrichis sericans Acrotrichis (= rosskotheni Acrotrichis montandonii Acrotrichis intermedia Acrotrichis insularis Acrotrichis henrici Acrotrichis fascicularis Acrotrichis dispar Acrotrichis brevipennis Acrotrichis atomaria Acrotrichis ACROTRICHINI filicornis Smicrus titan Nephanes NEPHANINI ACROTRICHINAE fraterna subvariolosa ) (Newman) (Heer) (Heer) (Matthews, A.) A.) (Matthews, ) ) (Fairmaire & Laboulbène) (Matthews, A.) (Matthews, Sundt (Matthews, A.) (Matthews, Johnson (Heer) (Mäklin) (De Geer) (Waltl) (Gillmeister) (Herbst) (Erichson) Sundt (Mannerheim) (Allibert) N Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● Vc23 1939-2012 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 1042 1040 1039 1038 1037 1035 1034 1033 1031 1030 1029 1028 1027 1023 1022 1021 1020 1019 1018 1017 1015 1014 1013 1012 1010 1009 1008 1004 1002 Leiodes cinnamomea Leiodes calcarata Leiodes badia Leiodes LEIODINI punctatus Hydnobius latifrons Hydnobius Liocyrtusa vittata Liocyrtusa triepkii Leiodes (= strigipennis Leiodes rugosa Leiodes (= rufipennis Leiodes oblonga Leiodes obesa Leiodes (= nigrita Leiodes (= longipes Leiodes litura Leiodes (= gyllenhalii Leiodes (= gallica Leiodes flavescens Leiodes (= ferruginea Leiodes (= (= flavicornis clavicornis dubia scita curta parvula brunnea ovalis pallens ) ) ) ) ) ) ) ) Stephens (Sturm) (Schmidt, W.L.E.) ) (Schmidt, W.L.E.) (Reitter) Stephens ) (Schmidt, W.L.E.) (Erichson) (Schmidt, W.L.E.) (Curtis) (Erichson) (Schmidt, W.L.E.) (Paykull) (Fabricius) Stephens Daffner (Curtis) (Panzer) (Sturm)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Themost frequentlyencountered Leiodes , frequent in flight interception and malaise traps, but hard to find by other meansby find other to malaise hard and but flight traps, interception in frequent ,

ecology 1080 1079 1078 1077 1076 1074 1070 1069 1068 1067 1064 1063 1062 1061 1060 1059 1057 1056 1054 1053 1052 1049 1046 1044 1043 Anisotoma humeralis Anisotoma AGATHIDIINI immunda Colenis cephalotes Agaricophagus PSEUDOLIODINI (= CHOLEVINAE viennense Colon serripes Colon zebei Colon dentipes Colon brunneum Colon latum Colon COLONINAE seminulum Agathidium pisanum Agathidium laevigatum Agathidium atrum Agathidium varians Agathidium rotundatum Agathidium nigripenne Agathidium marginatum Agathidium confusum Agathidium nigrinum Agathidium globus Amphicyllis orbicularis Anisotoma pauxilla Kraatz Kraatz ) (Sahlberg, C.R.) (Sahlberg, C.R.) Herbst (Latreille) (Sturm) (Paykull) (Fabricius) Beck Brisout (Fabricius) Sturm (Herbst) Brisout (Fabricius) Erichson (Linnaeus) (Gyllenhal) Sturm Schmidt, W.L.E.

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ●2000 ● ● ●1996 Vc23 1939-2012 ●1906 ●1907 ●1998 ●1998 ●2008 ● ● ● ●1999 ● ● ●1998 ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local in woods and open habitats in moss tussocks. in and habitats open and woods Local in sand] chalk, and the places on Open

ecology 1116 1115 1112 1111 1109 1108 1106 1105 1104 1103 1102 1101 1100 1099 1098 1095 1094 1093 1091 1090 1089 1088 1087 1085 1084 1083 1082 1081 Nargus anisotomoides Nargus wilkinii Nargus velox Nargus CHOLEVINI colonoides Nemadus ANEMADINI Ptomaphagus medius Ptomaphagus PTOMAPHAGINI tristis Catops nigrita Catops nigricans Catops morio Catops kirbii Catops grandicollis Catops fuscus Catops fuliginosus Catops coracinus Catops chrysomeloides Catops watsoni Sciodrepoides fumatus Sciodrepoides oblonga Choleva jeanneli Choleva glauca Choleva fagniezi Choleva cisteloides Choleva angustata Choleva agilis Choleva spadicea Choleva (Spence) (Panzer) (Spence) (Fabricius) (Panzer) (Illiger) (Illiger) Erichson Britten (Spence) Jeannel Britten (Spence) Latreille Kellner (Sturm) Erichson (Fabricius) (Frölich) Erichson B1133 (Kraatz) Rey (Spence) (Spence) (Spence) (Panzer) Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● Vc23 1939-2012 ●1906 ●1906 ●1998 ●1906 ●1998 ● ●1906 ●1998 ●1998 ● ● ●1906 ● ●2002 ●1906 ●1906 ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local in carrion Local in refuse, species. flood possiblya subterranean In

ecology 1163 1162 1161 1160 1156 1155 1153 1151 1149 1146 1142 1138 1137 1135 1134 1133 1132 1130 1128 1125 1124 1123 1122 1121 1120 1119 1118 1117 SCYDMAENIDAE SCYDMAENIDAE testaceus Leptinus PLATYPSILLINAE varicornis Ptomaphagus subvillosus Ptomaphagus Scydmaenus tarsatus Scydmaenus rufus Scydmaenus SCYDMAENINI scutellaris Stenichnus godarti Stenichnus collaris Stenichnus sparshalli Scydmoraphes praeteritus Neuraphes talparum Neuraphes angulatus Neuraphes maeklinii Euconnus hirticollis Euconnus fimetarius Euconnus denticornis Euconnus CYRTOSCYDMINI gallicum Cephennium CEPHENNIINI scydmaenoides Eutheia schaumii Eutheia plicata Eutheia formicetorum Eutheia EUTHEIINI SCYDMAENINAE (Gyllenhal) Kiesenwetter Kiesenwetter Müller, P.W.J. Müller, Müller, P.W.J. & P.W.J. Kunze Müller, (Latreille) (Müller, P.W.J. & P.W.J. Kunze) (Müller, (Mannerheim) (Illiger) (Müller, P.W.J. & P.W.J. Kunze) (Müller, (Illiger) Lokay Müller, P.W.J. & P.W.J. Kunze Müller, (Chaudoir) (Müller, P.W.J. & P.W.J. Kunze) (Müller, (Müller, P.W.J. & P.W.J. Kunze) (Müller, (Müller, P.W.J. & P.W.J. Kunze) (Müller, Rye Reitter Ganglbauer Stephens (Rosenhauer) (Denny) (Goeze)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● Vc23 1939-2012 ● ● ● ● ●1906 ●1998 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread in woods and parkland in litter, tussocks etc. litter, in parkland and woods Widespreadin Widespreadgrassyespecially open in sandy areas soils. on In rodent nests / runs also in nest boxes esp. amongst leaves, leaves, amongst boxes esp. nest in also runs / nests rodent In

ecology 1202 1201 1198 1197 1196 1194 1193 1192 1191 1189 1188 1187 1186 1185 1184 1183 1181 1180 1179 1175 1174 1172 1171 1170 1169 1167 1166 1165 1164 Thanatophilus sinuatus Thanatophilus rugosus Thanatophilus Necrodes littoralis SILPHINAE SILPHIDAE Eucnecosum brachypterum Eucnecosum tectum Deliphrum unicolor Anthobium atrocephalum Anthobium cruentata Acidota ANTHOPHAGINI OMALIINAE STAPHYLINIDAE vespilloides Nicrophorus vespillo Nicrophorus investigator Nicrophorus interruptus Nicrophorus humator Nicrophorus germanicus Nicrophorus NICROPHORINAE tristis Silpha obscura Silpha laevigata Silpha [ Dendroxena opaca Aclypea thoracicum Oiceoptoma Silpha atrata Silpha

Illiger Illiger quadrimaculata Linnaeus] (Linnaeus) Linnaeus Fabricius Mannerheim (Linnaeus) (Paykull) (Linnaeus) (Marsham) (Gleditsch) (Linnaeus) (Fabricius) (Linnaeus) Stephens Herbst Zetterstedt (Linnaeus) (Gyllenhal) (Gravenhorst) (Scopoli) Rom Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local on carrion. Localon carrion. Localon carrion. Localon

ecology 1256 1254 1251 1250 1249 1248 1243 1242 1241 1240 1234 1233 1232 1231 1228 1226 1225 1224 1223 1222 1220 1219 1217 1215 1210 1208 1206 1205 Philorinum sordidum Philorinum piceum Olophrum [ Lestevapubescens Lestevalongoelytrata Lestevahanseni Omalium rivulare Omalium oxyacanthae Omalium italicum Omalium exiguum Omalium excavatum Omalium caesum Omalium allardi Omalium vilis Dropephylla koltzei Dropephylla ioptera Dropephylla inflata Acrulia sulcula Acrolocha minuta Acrolocha OMALIINI torquatum Eusphalerum sorbi Eusphalerum primulae Eusphalerum minutum Eusphalerum luteum Eusphalerum EUSPHALERINI angusticolle Coryphium CORYPHIINI Lestevasicula (= vilis in part) in (Gyllenhal) Erichson] Lohse Fairmaire & Brisout (Erichson) Gravenhorst (Paykull) (Olivier) B1260 (Stephens) Mannerheim (Gyllenhal) Bernhauer (Gyllenhal) Jászay & Hlavac) Gyllenhal (Stephens) Stephens (Marsham) (Goeze) (Stephens) Gravenhorst (Stephens) (Fabricius) Stephens (Marsham)

Subfossil ● ● ● ● ● ● ● ● ● ?? ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● Vc23 1939-2012 ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon In woods at blossoms etc.] blossoms at woods In gorse on Local

ecology 1289 1288 1287 1286 1285 1284 1283 1282 1281 1280 1279 1278 1277 1276 1275 1273 1272 1270 1269 1268 1267 1266 1265 1264 1263 1262 1261 1260 1259 Phloeostiba plana Phloeostiba lapponica Phloeostiba pusillus Phloeonomus punctipennis Phloeonomus gayndahensis Paraphloeostiba septentrionis Omalium Proteinus laevigatus Proteinus crenulatus Proteinus brachypterus Proteinus atomarius Proteinus clypeata Metopsia prosseni Megarthrus hemipterus Megarthrus depressus Megarthrus denticollis Megarthrus bellevoyei Megarthrus PROTEININAE bosnica Xylostiba depressus Xylodromus concinnus Xylodromus salicis Phyllodrepa puberula Phyllodrepa nigra Phyllodrepa floralis Phyllodrepa (= (= (= (= (= retusa depressus sinuatocollis affinis monilicornis ) ) ) in part) in ) Bernhauer (Müller, P.W.J.) P.W.J.) (Müller, (Paykull) (Gravenhorst) (Gyllenhal) (Paykull) Hochhuth Erichson Schatzmayr Pandellé Bernhauer (Paykull) (Saulcy) B1304 (Zetterstedt) (Gravenhorst) (Gravenhorst) (Beck) Thomson, C.G. (Marsham) (Fabricius) (Illiger) (Illiger) Thomson, C.G. (MacLeay)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ?? ● ● ● ● Vc22 VCH ● Vc23 1939-2012 ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 1336 1335 1334 1333 1332 1326 1325 1323 1317 1315 1314 1310 1309 1308 1307 1306 1305 1304 1303 1300 1299 1298 1297 1296 1295 1293 1292 1291 1290 Micropeplus porcatus Micropeplus fulvus Micropeplus tesserula Arrhenopeplus MICROPEPLINAE ovalis Proteinus Bibloplectus delhermi Bibloplectus ambiguus Bibloplectus minutus Bibloporus bicolor Bibloporus TRICHONYCHINI signatus Euplectus sanguineus Euplectus piceus Euplectus karstenii Euplectus duponti Euplectus decipiens Euplectus bescidicus Euplectus EUPLECTINI EUPLECTITAE testaceus Claviger longicornis Claviger CLAVIGERINI CLAVIGERITAE venustus Batrisodes BATRISINI BATRISITAE PSELAPHINAE staphylinoides Micropeplus (= macropterus Stephens Motschulsky ) Preyssler Aubé (Denny) (Reichenbach) (Reichenbach) Erichson (Reichenbach) Raffray Raffray Müller, P.W.J. P.W.J. Müller, Reitter (Paykull) Guillebeau Denny (Reichenbach) (Curtis) (Marsham)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● Vc23 1939-2012 ●1998 ●1998 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon n subterranean nests of the ants LasiusL. ants of nests mixtus. the umbratus and subterranean n

ecology 1375 1374 1373 1371 1370 1369 1368 1367 1366 1364 1363 1362 1361 1360 1359 1358 1357 1348 1347 1346 1345 1344 1343 1342 1341 1340 1339 1338 1337 (= maerkelii Amauronyx tenebrosus Bibloplectus spinosus Bibloplectus pusillus Bibloplectus minutissimus Bibloplectus Pselaphaulax dresdensis Pselaphaulax PSELAPHINI PSELAPHITAE Tychus niger TYCHINI glabratus Tychobythinus macropalpus Bythinus burrellii Bythinus puncticollis Bryaxis curtisii Bryaxis bulbifer Bryaxis BYTHININI (= longicornis Rybaxis juncorum Reichenbachia (= impressa Fagniezia haematica Brachygluta fossulata Brachygluta BRACHYGLUTINI GONIACERITAE brevicorne Trimium sulcicollis Trichonyx laminata Trissemus maerkeli ) (Paykull) ) ) (Leach) (Reichenbach) Denny (Panzer) (Denny) (Leach) (Reichenbach) (Denny) (Reichenbach) Raffray (Aubé) (Reichenbach) Aubé (Reichenbach) (Reitter) (Herbst) (Rye) (Leach) (Aubé)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH

Vc23 1939-2012 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 1417 1416 1415 1414 1413 1412 1410 1409 1407 1406 1405 1404 1401 1399 1398 1395 1391 1390 1389 1388 1387 1386 1382 1381 1380 1379 1378 1377 1376 Bolitobius castaneus Bolitobius MYCETOPORINI TACHYPORINAE subtilissima Phloeocharis PHLOEOCHARINAE heisei Pselaphus Sepedophilus pedicularius Sepedophilus nigripennis Sepedophilus marshami Sepedophilus lusitanicus Sepedophilus littoreus Sepedophilus immaculatus Sepedophilus bipunctatus Sepedophilus saginatus Lamprinodes silphoides Cilea TACHYPORINI inclinans Parabolitobius rufescens Mycetoporus nigricollis Mycetoporus longulus Mycetoporus lepidus Mycetoporus clavicornis Mycetoporus trinotatus Lordithon thoracicus Lordithon lunulatus Lordithon exoletus Lordithon splendidum Ischnosoma longicorne Ischnosoma cingulatus Bolitobius (Linnaeus) Herbst (Erichson) (Linnaeus) (Erichson) (Gravenhorst) (Stephens) (Fabricius) (Mannerheim) (Linnaeus) Mannerheim (Stephens) (Gravenhorst) (Mäklin) Stephens (Stephens) (Stephens) (Gravenhorst) (Gravenhorst) Hammond (Stephens) Mannerheim (Gravenhorst) (Gravenhorst) (Stephens)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●2004 ● ● ●1990 ●2000 ●1984 ●1996 ● ●1984 ●1980 ●1991 ●2000 ●2000 ● Vc23 1939-2012 ● ● ●1998 ●1999 ●1994 ●1990 ●1970s ●1991 ●1949 ●1970s ●1998 ●1999 ●1965 ●2002 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 1459 1458 1457 1456 1452 1451 1450 1449 1448 1445 1442 1441 1440 1439 1438 1437 1435 1434 1433 1432 1431 1430 1426 1424 1423 1421 1419 1418 Tachinus marginellus Tachinus laticollis Tachinus humeralis Tachinus elongatus Tachinus bipustulatus Tachinus testaceus Sepedophilus Aleochara lata Aleochara curtula Aleochara brevipennis Aleochara ALEOCHARINI ALEOCHARINAE capillaricornis Habrocerus HABROCERINAE pilicornis Trichophya TRICHOPHYINAE solutus Tachyporus pusillus Tachyporus pallidus Tachyporus obtusus Tachyporus nitidulus Tachyporus hypnorum Tachyporus formosus Tachyporus dispar Tachyporus chrysomelinus Tachyporus atriceps Tachyporus subterraneus Tachinus scapularis Tachinus (= rufipes Tachinus signatus ) Gravenhorst (Linnaeus) (Goeze) Gravenhorst (Paykull) Erichson Gyllenhal (Linnaeus) Gravenhorst Stephens Gravenhorst Stephens) Sharp (Fabricius) (Gyllenhal) Matthews, A.H. Matthews, (Fabricius) (Fabricius) Gravenhorst (Fabricius) (Fabricius) (Linnaeus) (Gravenhorst) (Linnaeus)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ●1991 ●2004 ● ● ● ● ●2004 ● ● ●2001 ● ●2003 ●2003 ●2003 ●1980 Vc23 1939-2012 ● ●2001 ● ●1970s ● ● ● ● ● ●1994 ● ●1990 ●2003 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 1515 1514 1513 1512 1511 1503 1496 1495 1494 1493 1492 1491 1489 1487 1486 1485 1483 1477 1476 1475 1474 1466 1465 1464 1463 1462 1461 1460 Aleochara bipustulata Aleochara (= binotata Aleochara bilineata Aleochara ruficornis Aleochara intricata Aleochara Alaobia scapularis Alaobia pallidicornis Alaobia linderi Alaobia hybrida Alaobia gagatina Alaobia pygmaea Acrotona parvula Acrotona aterrima Acrotona ATHETINA ATHETINI morion Tinotus villosa Aleochara tristis Aleochara sparsa Aleochara moesta Aleochara moerens Aleochara maculata Aleochara lanuginosa Aleochara fumata Aleochara discipennis Aleochara cuniculorum Aleochara spadicea Aleochara verna Aleochara verna ) (Brisout) B1568 (Gravenhorst) (Sharp) Gravenhorst Say Say (Baudi) Mannerheim Heer (Mannerheim) Gravenhorst Gravenhorst (Gravenhorst) (Sahlberg, C.R.) (Gravenhorst, 1802) Kraatz Gyllenhal Mannerheim (Erichson) Gyllenhal Brisout Gravenhorst (Thomson, C.G.) Mulsant & Rey Gravenhorst (Linnaeus) Kraatz

Subfossil ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ●1995 ● ●2000 Vc23 1939-2012 ● ● ●1990 ● ● ●1946 ● ● ●1909 ● ● ● ● ● ●1965 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread in herbivore dung esp. cow, and carrion and on fungi. on and carrion cow, esp. and dung herbivore Widespreadin vicinity Rarewarrensin badgersetts] of rabbit [&

ecology 1559 1558 1557 1556 1555 1554 1553 1549 1548 1545 1543 1540 1537 1536 1534 1529 1528 1526 1525 1524 1523 1522 1521 1520 1519 1518 1517 1516 Alevonota egregia Alevonota aurantiaca Alevonota trinotata Alaobia taxiceroides Alaobia subglabra Alaobia sodalis Alaobia Atheta castanoptera Atheta brunneipennis Atheta aquatilis Atheta aquatica Atheta (= aeneicollis Atheta soedermani Anopleta puberula Anopleta (= corvina Anopleta nigrofusca Amischa decipiens Amischa analis Amischa longicollis Aloconota languida Aloconota sulcifrons Aloconota insecta Aloconota gregaria Aloconota currax Aloconota cambrica Aloconota incana Alianta rufotestacea Alevonota gracilenta Alevonota pertyi ellimani ) ) (Erichson) (Erichson) (Gravenhorst) (Thomson, C.G.) (Thomson, C.G.) (Kraatz) (Thomson, C.G.) (Kraatz) (Sharp) (Thomson, C.G.) (Rye) (Sharp) (Sharp) (Sharp) (Erichson) (Erichson) (Mannerheim) (Stephens) (Wollaston) Munster (Stephens) (Erichson) (Mulsant & Rey) (Bernhauer) Fauvel (Thomson, C.G.) (Kraatz)

Subfossil ● ? ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ●1992 ● ● ● ●1990 ●1992 ● ● Vc23 1939-2012 ●1998 ●1993 ● ●1908 ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local in fungLocal in fungi. rotting Rarein etc.woods in litter Local in

ecology 1608 1607 1604 1603 1596 1595 1594 1590 1589 1588 1587 1586 1585 1584 1583 1582 1581 1580 1578 1577 1576 1575 1573 1566 1561 Atheta crassicornis Atheta basicornis Atheta xanthopus Atheta triangulum Atheta hypnorum Atheta graminicola Atheta (= pervagata Ceritaxa rigidicornis Callicerus obscurus Callicerus occulta Bessobia monticola Bessobia fungivora Bessobia (= vaga Atheta strandiella Atheta ravilla Atheta procera Atheta pilicornis Atheta paracrassicornis Atheta oblita Atheta nigritula Atheta nidicola Atheta liturata Atheta harwoodi Atheta fungicola Atheta euryptera Atheta divisa Atheta dilaticornis nigricornis (Heer) (Erichson) (Märkel) (Erichson) (Stephens) (Kraatz) (Johansen) ) ) (Gravenhorst) (Thomson, C.G.) (Williams) (Stephens) (Erichson) (Thomson, C.G.) (Mulsant & Rey) (Kiesenwetter) (Kiesenwetter) (Thomson, C.G.) (Brundin) (Kraatz) (Benick) (Fabricius) (Thomson, C.G.) (Gravenhorst) (Thomson, C.G.) Gravenhorst (Erichson) Brundin

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●1992 ● ● ● ●1992 ●1992 ●1991 Vc23 1939-2012 ● ●1998 ●1930 ●1908 ●1998 ●1959 ●1990 ●1907 ●1998 ● ●1990 ● ● ●1970s ●1993 ● ● ●1998 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon In dreys and nests nests and dreys In

ecology 1658 1657 1656 1639 1638 1637 1636 1635 1634 1633 1632 1631 1629 1627 1626 1625 1622 1621 1619 1618 1617 1616 1615 1614 1613 1612 1611 1610 1609 Datomicra canescens Datomicra coriaria Dalotia immersa Dadobia melanaria Coprothassa longicornis Chaetida testaceipes Ceritaxa (= microptera Liogluta longiuscula Liogluta circellaris Geostiba hepatica Enalodroma clancula Dochmonota linearis Dinaraea angustula Dinaraea aequata Dinaraea setigera Dimetrota nigripes Dimetrota marcida Dimetrota laevana Dimetrota cinnamoptera Dimetrota cauta Dimetrota atramentaria Dimetrota vilis Dilacra luteipes Dilacra zosterae Datomicra sordidula Datomicra nigra Datomicra dadopora Datomicra celata Datomicra oblongiuscula (Erichson) (Erichson) (Kraatz) (Erichson) (Kraatz) (Erichson) (Gravenhorst) (Erichson) (Erichson) ) (Sharp) (Mulsant & Rey) (Thomson, C.G.) (Erichson) (Gravenhorst) (Thomson, C.G) (Heer) (Gyllenhal) Thomson, C.G. (Erichson) (Thomson, C.G.) (Gravenhorst) (Gravenhorst) (Sharp) (Erichson) (Erichson) (Mannerheim) (Gyllenhal) (Thomson, C.G.)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ●1993 ●1992 ●2003 ● ● Vc23 1939-2012 ●1990 ●1965 ● ●1907 ●1990 ● ●1990 ●1990 ●1907 ● ●1990 ●1910 ●1954 ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon ?Woodland moss litter] Wetlands in and bark. pine Localunder

ecology 1696 1695 1694 1693 1689 1687 1686 1685 1684 1683 1682 1681 1680 1679 1678 1677 1676 1675 1671 1670 1669 1664 1663 1662 Microdota atricolor Microdota atomaria Microdota amicula Microdota aegra Microdota anceps Lyprocorrhe pagana Liogluta Parameotica difficilis Parameotica mortuorum Pachyatheta cribrata Pachyatheta nigella Pachnida caesula Ousipalia flavipes Notothecta confusa Notothecta (= lividipennis Nehemitropia laticollis Mycetota fimorum Mycetota orphana Mocyta orbata Mocyta negligens Mocyta gilvicollis Mocyta fussi Mocyta fungi Mocyta clientula Mocyta amplicollis Mocyta subtilis Microdota liliputana Microdota inquinula Microdota indubia Microdota sordida ) (Bernhauer) (Gravenhorst) (Erichson) (Erichson) (Erichson) (Heer) (Erichson) (Erichson) (Sahlberg, J.) (Scheerpeltz) (Erichson) (Scriba) (Stephens) (Brisout) (Sharp) (Mulsant & Rey) (Stephens) (Gravenhorst) (Märkel) (Sharp) (Erichson) (Kraatz) (Brisout) (Gravenhorst) (Kraatz) (Brisout) (Thomson, C.G.) (Mannerheim)

Subfossil ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ●2000 ●1995 ● ● ●1965 ●1992 Vc23 1939-2012 ● ●1905 ● ●1970s ●1990 ● ● ●1990 ●1990 ● ●1990 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare in wetland in marsh wetland Rarein in litter. formerly Extinct atnests ant wood in Lasius ant with fuliginosus. ?the Hollowtrees and litter] Wetlands in moss? and litter ?In Widespread. haystack and Rare?[Moss refuse] brackets] birch [in woodlands In

ecology 1740 1739 1737 1736 1735 1734 1733 1732 1731 1730 1728 1726 1725 1719 1718 1717 1716 1711 1709 1708 1707 1706 1704 1703 1702 1701 1700 1699 1697 Philhygra fallaciosa Philhygra elongatula Philhygra deformis Philhygra debilis Philhygra britteni Philhygra inhabilis Paranopleta DEINOPSINI rivularis Autalia longicornis Autalia impressa Autalia AUTALIINI hospita Thamiaraea cinnamomea Thamiaraea THAMIARAEINA myrmecobia Xenota immigrata Trichiusa vestita Thinobaena gemina Schistoglossa curtipennis Schistoglossa paradoxa Pycnota brunnea Plataraea volans Philhygra terminalis Philhygra obtusangula Philhygra malleus Philhygra luridipennis Philhygra hygrotopora Philhygra hygrobia Philhygra gyllenhalii Philhygra (Gravenhorst) (Olivier) (Scriba) (Erichson) Joy Joy (Mulsant & Rey) (Joy) (Joy) (Fabricius) (Gravenhorst) Scheerpeltz (Kraatz) (Thomson, C.G.) (Kraatz) (Märkel) (Sharp) (Gravenhorst) Lohse B1759 (Thomson, C.G.) (Gravenhorst) (Erichson) (Kraatz) (Kraatz) (Joy) (Joy) (Mannerheim) (Sharp) (Gravenhorst)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●2002 ●1954 ● ●1992 ●1992 Vc23 1939-2012 ●1990 ●1953 ●1907 ● ● ●1990 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare in woodlands [At goat moth Cossus trees] trees] goatmoth [AtCossus woodlands Rarein nests] )?[Mole wetlands. ?In wet Rarein areas marshes, river-banksin litter Local and in moss wetlands] in [In )

ecology 1792 1791 1790 1789 1787 1786 1785 1784 1783 1782 1777 1776 1775 1774 1770 1768 1767 1766 1765 1764 1759 1758 1754 1753 1751 1750 1746 1741 Falagrioma thoracica Falagrioma sulcatula Falagria caesa Falagria obscura Cordalia FALAGRIINI erosa Deinopsis Gyrophaena congrua Gyrophaena bihamata Gyrophaena affinis Gyrophaena complicans Encephalus latissima Agaricochara GYROPHAENINA williamsi Thecturota marchii Thecturota gracilis Tachyusida clavigera Rhopalocerina ruficollis Leptusa pulchella Leptusa norvegica Leptusa fumida Leptusa obliqua Bolitochara lucida Bolitochara bella Bolitochara BOLITOCHARINA HOMALOTINI (= sulcata Myrmecopora concinnus Myrmecocephalus (= simillima reyi ) ) Erichson (Erichson) (Stephens) Märkel (Gravenhorst) (Erichson) (Mannerheim) Strand, A. (Gravenhorst) (Gravenhorst) Mannerheim (Dodero) (Erichson) Erichson (Bernhauer) B1814 (Bernhauer) B1814 Erichson (Kiesenwetter) (Kiesenwetter) (Stephens) Thomson, C.G. (Stephens) Stephens (Scriba) (Erichson)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ●2000 ● ●1954 ●1975 ●2004 Vc23 1939-2012 ●1907 ● ● ●1980s ●1990 ● ●1954 ●1959 ●1999 ●1970s VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 1828 1827 1825 1824 1823 1822 1821 1820 1819 1818 1817 1816 1815 1814 1813 1811 1810 1808 1803 1801 1800 1799 1798 1797 1796 1795 1794 1793 Gyrophaena lucidula Gyrophaena joyioides Gyrophaena joyi Gyrophaena hanseni Gyrophaena gentilis Gyrophaena fasciata Gyrophaena Cypha punctum Cypha pulicaria Cypha longicornis Cypha laeviuscula Cypha discoidea Cypha (= aprilis Cypha (= apicalis Cypha HYPOCYPHTINI dimidiata Hygronoma HYGRONOMINI rubiginosa Silusa SILUSINA plana Homalota cuspidatus Anomognathus HOMALOTINA strictula Gyrophaena pulchella Gyrophaena nana Gyrophaena minima Gyrophaena (= manca Gyrophaena imitator nitida angustata ) ) (Rey) ) (Brisout) (Motschulsky) (Erichson) (Gyllenhal) (Erichson) Wendeler (Paykull) Erichson (Mannerheim) (Paykull) Erichson Erichson Erichson (Marsham) Strand, A. Erichson Erichson Wüsthoff (Gravenhorst) Heer (Erichson.) RDK Subfossil ● ● ● ● ● ● ● ● ● ● ● ●1915 ● ? Vc22 VCH ● ● ● ●1955 ● ● ●1980 ●1978 Vc23 1939-2012 ●1961 ● ● ●1962 ●1967 ●1914 ●1970s ● ●1920 ●1963 ●1987 ●1970s VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare in maze-gill Rarein oak & sweet Daedaleaon quercina chestnut squamosus.W bracket RarefungiPolyporus esp. on wet fungi in woods] [On )

ecology 1867 1865 1864 1863 1860 1859 1858 1857 1855 1854 1853 1852 1851 1850 1849 1847 1846 1843 1842 1841 1840 1839 1836 1835 1834 1833 1832 1831 Oligota parva Oligota inflata Oligota granaria Oligota apicata Oligota flavicornis Holobus seminulum Cypha Myllaena masoni Myllaena kraatzi Myllaena intermedia Myllaena infuscata Myllaena gracilis Myllaena elongata Myllaena dubia Myllaena brevicornis Myllaena MYLLAENINI collaris Zyras lugens Pella limbata Pella laticollis Pella humeralis Pella funesta Pella cognata Pella canaliculata Drusilla MYRMEDONIINA emarginata Lomechusa LOMECHUSINA LOMECHUSINI pusillima Oligota punctulata Oligota (Gravenhorst) (Gravenhorst) (Paykull) (Märkel) (Märkel) Kraatz B1957 (Märkel) (Mannerheim) (Erichson) (Gravenhorst) (Gravenhorst) Erichson Sharp, 1871 Sharp, 1871 Matthews, A. Matthews, (Gravenhorst) (Matthews, A.H.) (Matthews, (Erichson) (Matthews, A.H.) (Matthews, Heer Kraatz (Lacordaire) Erichson (Matthews, A.H.) (Matthews, (Fabricius) (Paykull)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ●2000 ●2001 ●1994 ● Vc23 1939-2012 ●1998 ●1970s ●1970s ●1998 ●1907 ●2003 ● ● ●1954 ●1907 ●1990 ●1970s ●1910 ●1951 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare [With Myrmica Rare[With , ants] withMyrmica ants etc compostLocal heaps in

ecology 1911 1910 1909 1908 1907 1906 1903 1902 1901 1900 1898 1891 1890 1888 1887 1886 1884 1881 1880 1878 1876 1875 1874 1873 1870 1869 1868 Homoeusa acuminata Homoeusa pygmaea Dinarda maerkeli Dinarda DINARDINA OXYPODINI minuta Myllaena Ilyobates nigricollis Ilyobates (= bennetti Ilyobates cunctans Hygropora (= villosula Haploglossa nidicola Haploglossa marginalis Haploglossa gentilis Haploglossa corticina Dexiogyia suturalis Crataraea nigrata Cousya rufescens Calodera riparia Calodera protensa Calodera aethiops Calodera umbrosa Amarochara forticornis Amarochara OXYPODINA apicalis Meotica exillima Meotica exilis Meotica MEOTICINA subopacus pulla ) (Knoch in Gravenhorst) ) (Fairmaire & Laboulbène) Benick Erichson Sharp (Gravenhorst) Donisthorpe Kiesenwetter Kiesenwetter Wasmann (Gravenhorst) Mannerheim Kraatz (Mannerheim) (Erichson) (Paykull) (Märkel) (Erichson) (Fairmaire) (Stephens) (Erichson) (Märkel) (Boisduval & Lacordaire) (Gravenhorst)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●1992 ●1992 Vc23 1939-2012 ●1945 ●1906 ●1970 ●1908 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon in and around sand martin nests in sandpits sandpits martin sand in nests around and in Rare trees in Hollow inold nests. mammal in [Subterranean runs] Subterranean downland. on ? Withants

ecology 1947 1942 1941 1939 1938 1934 1933 1932 1931 1930 1929 1928 1927 1926 1925 1924 1923 1922 1921 1920 1919 1917 1916 1914 1913 1912 Ocalea picata Ocalea badia Ocalea incrassata Mniusa prolixa Ischnoglossa obscura Ischnoglossa propinquus Ilyobates Oxypoda nigrocincta Oxypoda longipes Oxypoda lentula Oxypoda induta Oxypoda haemorrhoa Oxypoda formiceticola Oxypoda ferruginea Oxypoda exoleta Oxypoda elongatula Oxypoda (= carbonaria Oxypoda (= brevicornis Oxypoda brachyptera Oxypoda annularis Oxypoda alternans Oxypoda (= acuminata Oxypoda Ocyusapicina ?? Ocyusanitidiventris Ocyusamaura rivularis Ocalea sericea umbrata lividipennis ) ) Erichson (Stephens) (Aubé) (Erichson) ) Mulsant & Rey Miller, L. L. Miller, Erichson Erichson Mulsant & Rey (Mulsant & Rey) (Gravenhorst) Mannerheim Erichson Aubé (Stephens) (Heer) ?* (Stephens) (Gravenhorst) Mulsant & Rey (Aubé) (Stephens) Wunderle (Mannerheim) Märkel

Subfossil ● ● ● ● ● ● ● ● ? ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ●1994 ●1995 ● ● ● ● ●2003 Vc23 1939-2012 ● ●1990 ● ● ● ●1990 ●1998 ●1998 ● ● ●1951 ●1990 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local in wetlands.Local in bark. Rare,under

ecology 1992 1991 1985 1984 1983 1980 1979 1977 1976 1975 1974 1971 1970 1969 1967 1964 1963 1962 1961 1960 1959 1958 1957 1955 1954 1953 1948 Phloeopora concolor Phloeopora vittata Oxypoda (= tarda Oxypoda spectabilis Oxypoda opaca Oxypoda Placusa complanata Placusa PLACUSINI atra Thinonoma objecta Tachyusa constricta Tachyusa scitula Ischnopoda leucopus Ischnopoda rubrior Gnypeta ripicola Gnypeta carbonaria Gnypeta caerulea Gnypeta velata Dasygnypeta concolor Brachyusa TACHYUSINA inquilina Thiasophila angulata Thiasophila longitarsis Tetralaucopora testacea Phloeopora (= scribae Phloeopora nitidiventris Phloeopora (= corticalis Phloeopora teres angustiformis brachyptera ) Sharp (Gravenhorst) Märkel Tottenham in part) part) in (Gravenhorst) (Kiesenwetter) B2006 (Kiesenwetter) (Mulsant & Rey) (Sahlberg, C.R.) ) (Erichson) (Erichson) Eppelsheim Märkel (Mannerheim) (Erichson) (Erichson) (Mannerheim) Erichson B2021 Kraatz (Marsham) (Gravenhorst) (Erichson) (Märkel) Fauvel (Erichson)

Subfossil ●1910s ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●1991 ●1965 ● ● ●1992 Vc23 1939-2012 ●1970s ●1998 ● ●1998 ● ● ●1906 ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare on bare sand and silt atmargins silt and ofbaresand rivers. Rareon withLasius fuliginosus nests Mole rarein Edges of water of Edges

ecology 2025 2024 2023 2016 2015 2014 2013 2012 2011 2010 2009 2008 2007 2006 2005 2004 2003 2002 2001 2000 1999 1998 1997 1996 1995 1994 1993 Scaphidium quadrimaculatum Scaphidium SCAPHIDIINI SCAPHIDIINAE tachyporoides Placusa pumilio Placusa depressa Placusa Anotylus saulcyi Anotylus rugosus Anotylus nitidulus Anotylus insecatus Anotylus fairmairei Anotylus complanatus Anotylus clypeonitens Anotylus OXYTELINI aeneum Syntomium Deleaster dichrous DELEASTERINI striatulus Coprophilus COPROPHILINI OXYTELINAE quadricorne Siagonium PIESTINAE boleti Scaphisoma assimile Scaphisoma agaricinum Scaphisoma SCAPHISOMATINI immaculatum Scaphium SCAPHIINI (Gravenhorst) (Pandellé) Mäklin (Fabricius) (Panzer) (Gravenhorst) (Gravenhorst) (Gravenhorst) (Pandellé) (Müller, P.W.J.) P.W.J.) (Müller, Erichson (Pandellé) (Erichson) (Waltl) (Fabricius ) Kirby Kirby (Linnaeus) (Olivier) Olivier Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● Vc23 1939-2012 ● ●1998 ●1998 ● ● ●1919 ●1909 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 2076 2074 2073 2069 2068 2066 2065 2063 2062 2046 2045 2044 2043 2042 2041 2040 2039 2038 2037 2036 2035 2034 2033 2032 2030 2029 2028 2027 2026 Oxytelus sculptus Oxytelus migrator Oxytelus laqueatus Oxytelus fulvipes Oxytelus tetracarinatus Anotylus sculpturatus Anotylus Carpelimus elongatulus Carpelimus corticinus Carpelimus bilineatus Carpelimus pallipes Bledius opacus Bledius longulus Bledius gallicus Bledius filipes Bledius femoralis Bledius ssp. [ (= limicola Bledius subterraneus Bledius caelatus Aploderus THINOBIINI arenarius Platystethus nodifrons Platystethus nitens Platystethus degener Platystethus cornutus Platystethus capito Platystethus alutaceus Platystethus Bledius spectabilis Bledius frisius frisius germanicus Lohse Sharp (Block) (Gravenhorst) ) ) (Gravenhorst) Tottenham Erichson Erichson Gravenhorst B2069 (Gyllenhal) Fauvel (Sahlberg, C.R.) Heer (Gravenhorst) (Marsham) Kraatz] Mulsant & Rey Erichson (Gravenhorst) (Gravenhorst) Stephens (Gravenhorst) Thomson, C.G. (Fourcroy) Mannerheim (Block) (Erichson) Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●1953 ● ● ● Vc23 1939-2012 ● ● ●1998 ●1998 ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 2138 2137 2136 2135 2134 2132 2130 2128 2127 2125 2124 2123 2120 2117 2116 2115 2114 2113 2112 2101 2100 2097 2093 2091 2090 2087 2086 2081 2078 Carpelimus zealandicus Carpelimus subtilis Carpelimus rivularis Carpelimus pusillus Carpelimus impressus Carpelimus fuliginosus Carpelimus Stenus binotatus Stenus bifoveolatus Stenus tarsalis Stenus solutus Stenus similis Stenus latifrons Stenus fulvicornis Stenus cicindeloides Stenus subaeneus Stenus pallipes Stenus ossium Stenus ochropus Stenus impressus Stenus fuscicornis Stenus aceris Stenus coerulescens Dianous STENINAE rufus Oxyporus OXYPORINAE unicolor Teropalpus palpalis Plaeustomus aureus Ochthephilus mandibularis Manda Stephens (Herbst) Stephens Erichson Ljungh Gravenhorst (Linnaeus) Erichson Kiesenwetter Kiesenwetter Ljungh Germar Stephens Erichson Erichson (Erichson) Gyllenhal (Gravenhorst) (Schaller) (Sharp) (Motschulsky) (Fauvel) (Gyllenhal) (Erichson) (Gyllenhal) (Boisduval & Lacordaire) (Gravenhorst) (Sharp)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ●1991 Vc23 1939-2012 ● ● ● ● ● ● ● ●1906 ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Raremoss water beside falls ,streams. fungi rotting in declining Rareand

ecology 2178 2177 2176 2175 2174 2173 2170 2168 2166 2164 2163 2162 2161 2160 2158 2157 2156 2155 2154 2153 2151 2149 2148 2147 2146 2144 2143 2140 2139 Stenus picipennis Stenus pallitarsis Stenus nitidiusculus Stenus flavipes Stenus butrintensis Stenus brevipennis Stenus Stenus melanopus Stenus melanarius Stenus lustrator Stenus longitarsis Stenus juno Stenus incrassatus Stenus guttula Stenus fuscipes Stenus europaeus Stenus comma Stenus clavicornis Stenus circularis Stenus carbonarius Stenus canaliculatus Stenus boops Stenus bimaculatus Stenus biguttatus Stenus atratulus Stenus ater Stenus assequens Stenus argus Stenus pubescens Stenus picipes Stenus Mannerheim (Paykull) Gravenhorst Ljungh Stephens Müller, P.W.J. P.W.J. Müller, LeConte Stephens Gravenhorst Erichson Erichson Gravenhorst Stephens (Linnaeus) Rey Stephens, 1833 Erichson, 1840 Puthz Thomson, C.G. (Scopoli) (Marsham) Erichson Stephens Thomson Smetana Gyllenhal Gyllenhal Stephens Gyllenhal

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ? ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Fens, and mesotrophic wetlands]mesotrophic and Fens, Wetlandspecies]

ecology 2218 2217 2216 2215 2214 2213 2212 2210 2209 2208 2207 2204 2202 2201 2200 2199 2198 2196 2195 2194 2193 2191 2189 2188 2186 2185 2184 2181 2180 Stenus brunnipes Stenus (= pusillus Stenus providus Stenus nitens Stenus nanus Stenus Lathrobium pallidipenne Lathrobium longulum Lathrobium impressum Lathrobium geminum Lathrobium fulvipenne Lathrobium fovulum Lathrobium elongatum Lathrobium brunnipes Lathrobium debilicornis Hypomedon lyonessius Astenus immaculatus Astenus pulchellus Astenus humile Achenium depressum Achenium PAEDERINAE sulcata Pseudopsis PSEUDOPSINAE ruficapillus Euaesthetus laeviusculus Euaesthetus bipunctatus Euaesthetus EUAESTHETINAE nigritulus Stenus crassus Stenus exiguus ) Stephens Stephens Stephens Stephens Erichson Gyllenhal Stephens (Nicolai) (Joy) (Joy) Newman (Heer) B2233 (Heer) B2233 Stephens Stephens (Gravenhorst) Kraatz Gravenhorst (Fabricius) (Gravenhorst) (Linnaeus) Heer, 1841 Heer, 1841 Lacordaire (Wollaston) (Ljungh) Hochhuth Mannerheim

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ●1953 ● ● ● ● ● ● Vc23 1939-2012 ● ●1998 ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 2256 2255 2254 2253 2252 2251 2250 2249 2248 2247 2246 2243 2242 2239 2238 2237 2236 2234 2233 2232 2230 2229 2227 2224 2231 2220 2219 Tetartopeus terminatus Tetartopeus quadratus Tetartopeus angustatus Tetartopeus multipunctum Lobrathium pallidum Lathrobium (= Rugilus rufipes Rugilus orbiculatus Rugilus geniculatus Rugilus erichsoni Rugilus (= angustatus Rugilus obsoletus Pseudomedon obscurellus Pseudomedon riparius Paederus littoralis Paederus fuscipes Paederus jacquelini Ochthephilum fracticorne Ochthephilum ripicola Medon pocofer Medon piceus Medon fusculus Medon dilutus Medon castaneus Medon brunneus Medon apicalis Medon ochracea Lithocharis nigriceps Lithocharis fragilis ripicola ) ) [Check] (Kraatz) (Erichson) (Peyron) (Kraatz) (Kraatz) Germar (Mannerheim) (Erichson) (Gravenhorst) (Fauvel) (Linnaeus) Curtis Gravenhorst Geoffroy) (Paykull) (Erichson) (Gravenhorst) Kraatz Nordmann (Paykull) (Gravenhorst) (Nordmann) (Lacordaire) (Boieldieu) (Paykull) (Erichson) B2293 (Gravenhorst)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● Vc23 1939-2012 ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 2298 2296 2295 2294 2292 2291 2287 2286 2284 2283 2282 2281 2280 2279 2278 2277 2276 2275 2274 2272 2271 2270 2269 2268 2267 2266 2265 2264 2257 STAPHYLININAE propinquus Sunius melanocephalus Sunius bicolor Sunius sulcicollis Scopaeus similis Rugilus Gabrius breviventer Gabrius astutoides Gabrius (= appendiculatus Gabrius signaticornis Erichsonius cinerascens Erichsonius subuliformis Bisnius sordidus Bisnius puella Bisnius pseudoparcus Bisnius parcus Bisnius nigriventris Bisnius fimetarius Bisnius cephalotes Bisnius PHILONTHINA STAPHYLININI (= subuliformis Othius punctulatus Othius laeviusculus Othius angustus Othius Atrecus affinis OTHIINI subnigritulus myrmecophilus (Nordmann) (Olivier) (Paykull) (Sharp) ** (Erichson) Stephens (Gravenhorst) ) (Gravenhorst) (Gravenhorst) (Strand, A.) (Brisout) (Goeze) ) (Thomson, C.G.) Stephens (Stephens) Stephens (Sperk) (Gravenhorst) (Brunne) Sharp (Fabricius) (Gravenhorst) (Mulsant & Rey)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●2004 ● ● ● Vc23 1939-2012 ●1998 ●1998 ●1956 ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 2339 2338 2337 2336 2335 2334 2333 2332 2331 2329 2328 2327 2322 2321 2320 2319 2316 2315 2314 2311 2310 2309 2308 2307 2306 2305 2304 2303 2299 Gabrius trossulus Gabrius splendidulus Gabrius piliger Gabrius osseticus Gabrius nigritulus Gabrius (= Philonthus longicornis Philonthus lepidus Philonthus laminatus Philonthus jurgans Philonthus intermedius Philonthus furcifer Philonthus fumarius Philonthus ebeninus Philonthus discoideus Philonthus decorus Philonthus debilis Philonthus cruentatus Philonthus concinnus Philonthus cognatus Philonthus (= carbonarius Philonthus albipes Philonthus addendus Philonthus villosulus Neobisnius (= lathrobioides Neobisnius thermarum Gabronthus velox Gabrius varius cerrutii pennatus ) ) ) Sharp Mulsant & Rey (Kolenati) (Nordmann) (Gravenhorst) (Gravenhorst) (Gravenhorst) (Gravenhorst) Tottenham (Gravenhorst) Renkonen (Gravenhorst) Stephens (Gravenhorst) Sharp (Stephens) (Gravenhorst) (Creutzer) (Gravenhorst) (Gravenhorst) (Gmelin in Linnaeus) Stephens (Lacordaire) (Gravenhorst) (Aubé) (Baudi)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ●2000 ● ●1992 Vc23 1939-2012 ● ● ● ● ● ●2000 ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 2378 2377 2376 2375 2372 2369 2366 2365 2364 2363 2362 2361 2360 2359 2358 2357 2356 2355 2354 2353 2350 2349 2348 2346 2344 2343 2342 2341 2340 Philonthus nigrita Philonthus (= micantoides Philonthus micans Philonthus marginatus Philonthus mannerheimi Philonthus Heterothops praevius Heterothops niger Heterothops (= minutus Heterothops picipes Euryporus QUEDIINA ventralis Philonthus varians Philonthus umbratilis Philonthus tenuicornis Philonthus succicola Philonthus splendens Philonthus spinipes Philonthus sanguinolentus Philonthus (= rufipes Philonthus rubripennis Philonthus rotundicollis Philonthus rectangulus Philonthus quisquiliarius Philonthus politus Philonthus (= parvicornis Philonthus dissimilis immundus agilis micans ) in part) part) in in major in part) ) Kraatz (Paykull) (Linnaeus) (Gravenhorst) (Stephens) (Gravenhorst) (Paykull) Sharp (Gravenhorst) Thomson, C.G. Wollaston Erichson (Fabricius) (Gravenhorst) (Gravenhorst) Mulsant & Rey (Müller, O.F.) O.F.) (Müller, Stephens Sharp Benick & Lohse (Ménétriés) Fauvel (Gyllenhal) (Gravenhorst)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ●1998 ● ● ● ● Vc23 1939-2012 ●1954 ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local? in compostLocal? heaps in

ecology 2414 2413 2411 2410 2409 2407 2406 2403 2402 2401 2400 2399 2396 2395 2394 2393 2392 2390 2389 2388 2387 2386 2385 2384 2383 2382 2381 2380 2379 Quedius fulgidus Quedius cruentus Quedius brevis Quedius brevicornis Quedius aetolicus Quedius cinctus Quedius Quedius nitipennis Quedius nigriceps Quedius maurorufus Quedius humeralis Quedius fumatus Quedius boops Quedius boopoides Quedius molochinus Quedius (= levicollis Quedius fuliginosus Quedius curtipennis Quedius xanthopus Quedius (= truncicola Quedius scitus Quedius puncticollis Quedius microps Quedius mesomelinus Quedius maurus Quedius longicornis Quedius lateralis Quedius invreae Quedius tristis ventralis ) ) ) (Gravenhorst) (Gravenhorst) Erichson (Paykull) Gridelli (Sahlberg, C.R.) Gravenhorst, J.L.C. (Gravenhorst) (Fabricius) (Stephens) (Brullé) (Olivier) Kraatz Kraatz (Stephens) Munster Stephens Fairmaire & Laboulbène Erichson (Thomson, C.G.) (Gravenhorst) Bernhauer Kraatz (Thomson, C.G.) (Gravenhorst) (Gravenhorst) (Marsham)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ●1997 ●2003 ●1994 ●1981 ●1996 ● ● ● ● ●1970s ●1978 ●2012 ● ● ● ●2004 Vc23 1939-2012 ●1990 ●1990 ●1999 ● ● ● ● ●1908 ● ●1952 ● ●1965 ● ●2004 ●1907 ●1952 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare in litter and moss and water.litter near Rarein litter. Rarein carr areas and woods Local in ofnests mammals. subterranean Rare.In

ecology 2447 2446 2445 2444 2442 2441 2439 2438 2437 2436 2435 2434 2433 2432 2431 2430 2427 2426 2425 2424 2423 2422 2421 2420 2419 2418 2417 2416 2415 Quedius schatzmayri Quedius riparius Quedius plancus Quedius picipes Quedius (= persimilis Quedius Tasgius melanarius Tasgius globulifer Tasgius erythropterus Staphylinus dimidiaticornis Staphylinus stercorarius Platydracus latebricola Platydracus tessellatus Ontholestes murinus Ontholestes Ocypus fuscatus Ocypus fortunatarum Ocypus aeneocephalus Ocypus ophthalmicus Ocypus olens (= Ocypus nitens Ocypus brunnipes maxillosus Creophilus STAPHYLININA Velleius dilatatus umbrinus Quedius semiobscurus Quedius semiaeneus Quedius scintillans Quedius nero aridulus ) ) (Müller, O.F.) O.F.) (Müller, (Schrank) (Mannerheim) (Gravenhorst) Erichson Kellner (Fabricius) (Fabricius) (Geoffroy) Mulsant & Rey Erichson (Gravenhorst) (Heer) (Stephens) (Linnaeus) Gridelli (Wollaston) (Scopoli) (Geoffroy) (Linnaeus) (Marsham) (De Geer) (Gravenhorst) (Olivier) Linnaeus Gemminger Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ●2004 ● ●2003 ● ●1984 ●1998 ● ● ● ●2008 ● ●2001 ● ●1996 ●1996 ● ● ### Vc23 1939-2012 ● ●2004 ● ● ●2004 ●2001 ●2002 ●2003 ● ●2004 ● ● ●1970s ● ●1970s ●2001 ●1970s ● ●2000 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon In hornet nests hornet In

ecology 2478 2477 2476 2475 2474 2472 2471 2470 2469 2468 2467 2466 2465 2464 2463 2462 2461 2460 2459 2458 2457 2456 2455 2454 2453 2451 2450 2449 2448 Gauropterus fulgidus Gauropterus XANTHOLININI ater Tasgius winkleri Tasgius (= morsitans Tasgius Xantholinus longiventris Xantholinus linearis Xantholinus gallicus Xantholinus (= elegans Xantholinus laevigatus Xantholinus (= pallidipennis Phacophallus parumpunctatus Phacophallus lentus Nudobius glabratus Megalinus pusillus Leptacinus intermedius Leptacinus formicetorum Leptacinus batychrus Leptacinus angularis Hypnogyra wagneri Gyrohypnus punctulatus Gyrohypnus fracticornis Gyrohypnus atratus Gyrohypnus angustatus Gyrohypnus jarrigei tricolor compressus ) ) (Gravenhorst) ) (Gravenhorst) Bernhauer (Rossi) (Stephens) (Heer) (Olivier) (Olivier) (Gravenhorst) Coiffait Scheerpeltz B2488 Ganglbauer (Gyllenhal) (Fabricius) Jacobsen Donisthorpe Stephens (Müller, O.F.) O.F.) (Müller, (Paykull) Heer Märkel (Motschulsky) B2517 (Gyllenhal) Rom Rom Rom Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ●2002 ● Vc23 1939-2012 ● ● ● ● ● ●1987 ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Extinct formerly with wood ants Extinctformerly etc. heaps manure In

ecology 2514 2512 2511 2505 2504 2503 2501 2500 2499 2498 2497 2496 2495 2494 2493 2492 2491 2489 2488 2487 2486 2485 2484 2483 2482 2481 2480 2479 Lucanus cervus Lucanus LUCANINAE cylindricum Sinodendron SYNDESINAE LUCANIDAE SCARABAEOIDEA Aphodius luridus Aphodius APHODIINI APHODIINAE SCARABAEIDAE vernalis Trypocopris stercorarius Geotrupes spiniger Geotrupes mutator Geotrupes stercorosus Anoplotrupes typhoeus Typhaeus GEOTRUPIDAE armiger Odonteus BOLBOCERATIDAE scaber Trox sabulosus Trox TROGIDAE parallelipipedus Dorcus DORCINAE Platycerus caraboides (Linnaeus) (Linnaeus) (Linnaeus) (Fabricius) (Scopoli) (Marsham) (Marsham) (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) (Scriba) (Linnaeus) Med Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon In dung esp. cow] esp. dung In comes Rare,light. subterranean, to compost heaps in also and nests, bird old HollowtreesLocal in in dry carrion] ?(Na)[Under Extinct

ecology 2555 2554 2552 2548 2547 2546 2545 2543 2539 2538 2537 2536 2535 2534 2533 2532 2531 2529 2528 2527 2526 2525 2524 2523 2520 2519 2518 2515 Aphodius fimetarius Aphodius sordidus Aphodius rufus Aphodius constans Aphodius ater Aphodius rufipes Aphodius Aphodius fossor Aphodius porcus Aphodius borealis Aphodius haemorrhoidalis Aphodius obliteratus Aphodius contaminatus Aphodius sphacelatus Aphodius prodromus Aphodius zenkeri Aphodius lividus Aphodius subterraneus Aphodius coenosus Aphodius pusillus Aphodius merdarius Aphodius erraticus Aphodius paykulli Aphodius distinctus Aphodius granarius Aphodius ictericus Aphodius (= pedellus Aphodius foetidus Aphodius foetens Aphodius fimetarius (De Geer) in part)) part)) in (Moll) (Moll) (Linnaeus) (Olivier) (Fabricius) (Linnaeus) (Fabricius) Germar (Herbst) (Herbst) Gyllenhal Bedel (De Geer) (Laicharting) (Fabricius) Duftschmid (Linnaeus) (Panzer) (Müller, O.F.) O.F.) (Müller, (Linnaeus) (Fabricius) (Linnaeus) Sturm (Brahm) (Panzer) (Linnaeus) (Herbst) (Linnaeus) Rom Rom Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ●1998 ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Extinct? In dung, once widespread once dung, In Extinct?

ecology 2606 2603 2602 2601 2600 2598 2597 2596 2595 2594 2590 2588 2587 2586 2583 2582 2581 2580 2579 2577 2576 2575 2574 2573 2561 2559 2557 2556 SCARABAEINAE sylvestris Oxyomus villosus Euheptaulacus (= sticticus Aphodius Gnorimus nobilis Gnorimus aurata Cetonia CETONIINAE horticola Phyllopertha dubia Anomala RUTELINAE brunnea Serica ruricola Omaloplia solstitiale Amphimallon melolontha Melolontha philanthus Hoplia MELOLONTHINAE vacca Onthophagus similis Onthophagus nuchicornis Onthophagus (= joannae Onthophagus coenobita Onthophagus taurus Onthophagus lunaris Copris ovatus equestris ) ) (Linnaeus) (Linnaeus) (Linnaeus) (Scopoli) (Linnaeus) (Panzer) (Füessly) (Scopoli) (Fabricius) (Linnaeus) (Schreber) (Scriba) Goljan (Linnaeus) (Gyllenhal in Schönherr) (Linnaeus) (Linnaeus) (Herbst) (Linnaeus) Rom Rom Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ●1992 ● ●1904 ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 2648 2647 2646 2644 2643 2642 2641 2637 2636 2635 2632 2631 2630 2629 2628 2627 2626 2621 2620 2619 2618 2617 2616 2615 2614 2613 2612 2611 2607 Eucinetus meridionalis Eucinetus EUCINETIDAE SCIRTOIDEA variabilis Gnorimus Cyphon ochraceus Cyphon (= laevipennis Cyphon hilaris Cyphon coarctatus Cyphon testacea Microcara Odelesmarginata (= pseudominuta Elodes minuta Elodes elongata Elodes SCIRTIDAE simsoni Clambus punctulum Clambus pubescens Clambus pallidulus Clambus evae Clambus armadillo Clambus CLAMBINAE dubius Calyptomerus CALYPTOMERINAE CLAMBIDAE phragmiteticola minuta in part) part) in Endrödy-Younga (Linnaeus) Nyholm Tournier Blackburn B2658 (Fabricius) Stephens Paykull (Linnaeus) (De Geer) Reitter Tournier Redtenbacher (Linnaeus) ) (Beck) (Marsham) Klausnitzer (Laporte) Rom Subfossil ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● Vc23 1939-2012 ● ● ●1998 ●1998 ● ●1998 ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare. Shaded streams Rare.Shaded

ecology 2692 2691 2690 2689 2688 2686 2684 2683 2679 2678 2677 2676 2675 2667 2666 2665 2663 2662 2661 2660 2658 2656 2654 2653 2651 2650 2649 Scirteshemisphaericus serricornis Prionocyphon variabilis Cyphon pubescens Cyphon palustris Cyphon padi Cyphon Trachys subglaber Rey subglaber Trachys scrobiculatus Trachys (= minutus Trachys TRACHYINI emarginatus Aphanisticus pusillus Aphanisticus APHANISTICINI sinuatus Agrilus laticornis Agrilus biguttatus Agrilus angustulus Agrilus viridis Agrilus AGRILINI AGRILINAE BUPRESTINAE BUPRESTIDAE BUPRESTOIDEA cervinus Dascillus DASCILLIDAE DASCILLOIDEA minuta ) (Linnaeus) (Linnaeus) (Olivier) (Linnaeus) Thomson, C.G. (Illiger) (Illiger) (Thunberg) (Fabricius) (Linnaeus) (Fabricius) (Illiger) (Illiger) (Olivier) Kiesenwetter Kiesenwetter (Linnaeus) (Müller, P.W.J.) P.W.J.) (Müller, (Olivier) Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon On sallows, On hazeletc fen areasmarshes wetlands,richer poor and Rare in

ecology 2728 2726 2725 2724 2721 2718 2717 2716 2715 2714 2711 2710 2709 2708 2707 2706 2705 2704 2702 2700 2699 2698 2697 2696 2695 2694 2693 BYRRHINAE BYRRHIDAE BYRRHOIDEA troglodytes Trachys Riolus subviolaceus Riolus cupreus Riolus tuberculatus Oulimnius major Oulimnius quadrituberculatus Macronychus volckmari Limnius parallelepipedus Esolus aenea Elmis ELMIDAE (= maritima Curimopsis spinosa Chaetophora SYNCALYPTINAE murinus Porcinolus Byrrhus pustulatus Byrrhus pilula Byrrhus fasciatus sericeusCytilus semistriata Simplocaria maculosa Simplocaria striatopunctata (Müller, P.W.J.) P.W.J.) (Müller, (Linnaeus) (Müller, P.W.J.) P.W.J.) (Müller, (Forster) (Rey) (Forster) (Panzer) (Forster) ) Gyllenhal in Schönherr (Müller, P.W.J.) P.W.J.) (Müller, (Fabricius) (Rossi) (Marsham) Erichson (Fabricius) (Müller, P.W.J.) P.W.J.) (Müller, (Müller, P.W.J.) P.W.J.) (Müller, RD3 Müller, P.W.J. P.W.J. Müller, Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ●190 ●190 ●190 ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare in stream Rare in streams In formerly chalk nd Rareon sandy grasslandon locallycommon on sandy ground primarily species. a winter

ecology 2779 2778 2775 2774 2772 2771 2767 2763 2762 2761 2760 2759 2755 2754 2753 2752 2750 2746 2743 2742 2741 2740 2739 2736 2735 2734 2733 2732 2729 Dryops similaris Dryops luridus Dryops griseus Dryops ernestiDryops auriculatus Dryops DRYOPIDAE Trixagus obtusus Trixagus dermestoides Trixagus carinifrons Trixagus THROSCIDAE cornutus Epiphanis buprestoides Melasis MELASINAE EUCNEMIDAE ELATEROIDEA palustris Eubria PSEPHENIDAE Heterocerus marginatus Heterocerus fenestratus HETEROCERIDAE pygmaeus Limnichus LIMNICHIDAE des Gozis (Erichson) (Erichson) Germar Bollow Bollow (Curtis) (Fourcroy) Eschscholtz (Linnaeus) (de Bonvouloir) (Sturm) (Linnaeus) (Thunberg) (Fabricius) Rom Subfossil ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ●1998 ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon veryfen ponds and Fens rare in rare saproxylic saproxylic rare oaks wood especially dead on seepages fen in rare by water sedients and mud on local by water sedients and mud on local

ecology 2820 2817 2816 2815 2814 2813 2812 2809 2807 2806 2805 2803 2802 2801 2798 2797 2796 2794 2792 2791 2790 2788 2787 2786 2785 2784 2782 2781 2780 DENTICOLLINAE riparius Hypnoidus HYPNOIDINAE murinus Agrypnus AGRYPNINAE ELATERIDAE AGRIOTINI pallens Adrastus ADRASTINI ELATERINAE hirtus Hemicrepidius (= rhombeus Stenagostus bicolor Athous vittatus Athous haemorrhoidalis Athous linearis Denticollis minuta Kibunea aeruginosus Cydnopus DENTICOLLINI aeneus Selatosomus tessellatum Prosternon nigricornis Paraphotistus incanus Aplotarsus bipustulatus Calambus Ctenicerapectinicornis Cteniceracuprea Actenicerus sjaelandicus CTENICERINI villosus ) (Goeze) (Fabricius) (Linnaeus) (Fabricius) (Fabricius) (Linnaeus) (Linnaeus) (Gyllenhal) (Fabricius) (Linnaeus) (Herbst) (Olivier) (Olivier) (Linnaeus) (Linnaeus) (Linnaeus) (Fabricius) (Müller, O.F.) O.F.) (Müller, (Panzer) Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ●2000 ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon common on herbage countywide herbage on common by water ground chalky and sadny

ecology 2877 2876 2875 2868 2867 2864 2859 2858 2857 2855 2854 2853 2850 2848 2847 2845 2844 2841 2832 2830 2829 2828 2827 2826 2824 2823 2822 2821 Dalopius marginatus Dalopius sputator Agriotes pallidulus Agriotes obscurus Agriotes lineatus Agriotes acuminatus Agriotes Drilus flavescensDrilus DRILIDAE asellus Cardiophorus CARDIOPHORINAE quadripustulatus Oedostethus NEGASTRIINAE (= villosus Melanotus (= castanipes Melanotus MELANOTINAE Sericus brunneus ELATERINI tibialis Procraerus MEGAPENTHINI sanguinicollis Ischnodes sanguinolentus Ampedus rufipennis Ampedus elongantulus Ampedus cardinalis Ampedus balteatus Ampedus AMPEDINI erythropus villosus in part) part) in ) ) (Linnaeus) (Fourcroy) (Linnaeus) (Linnaeus) (Linnaeus) (Boisduval & Lacordaire) (Geoffroy in Fourcroy) (Illiger) (Illiger) (Linnaeus) (Schiødte) (Stephens) (Stephens) (Linnaeus) Erichson (Paykull) (Fabricius) (Panzer) (Schrank) (Fabricius) RD2 Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ●2002 ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ●2001 ● ● ● ● ● ● ● ● ● VC22 v ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 2916 2915 2914 2913 2912 2911 2910 2908 2907 2906 2905 2903 2901 2900 2899 2898 2897 2896 2895 2894 2892 2891 2890 2886 2885 2884 2883 2879 2878 Lampyris noctiluca Lampyris LAMPYRIDAE minutus Platycis LYCIDAE Rhagonycha translucida Rhagonycha testacea Rhagonycha lutea Rhagonycha (= limbata Rhagonycha lignosa Rhagonycha fulva Rhagonycha thoracica Cantharis rustica Cantharis rufa Cantharis pellucida Cantharis pallida Cantharis nigricans Cantharis nigra Cantharis livida Cantharis lateralis Cantharis fusca Cantharis figurata Cantharis decipiens Cantharis cryptica Cantharis alpinus Podabrus CANTHARINAE CANTHARIDAE femoralis ) ) Linnaeus Linnaeus (De Geer) Linnaeus (Fabricius) Fallén Goeze (Paykull) (Müller, O.F.) O.F.) (Müller, (Scopoli) Ashe Linnaeus Mannerheim (Linnaeus) Fabricius Baudi (Olivier) O.F.) (Müller, (Müller, O.F.) O.F.) (Müller, Thomson, C.G. (Linnaeus) (Krynicki) Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●(1997) ● ●(1992) Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●(1999) VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 2955 2954 2953 2950 2945 2944 2943 2942 2941 2940 2939 2938 2937 2936 2934 2933 2932 2930 2929 2928 2927 2926 2923 2922 2921 2920 2919 2918 2917 Malthinus frontalis Malthinus flaveolus Malthinus balteatus Malthinus MALTHININAE ruficollis Silis SILINAE Dermestes undulatus Dermestes murinus Dermestes maculatus Dermestes DERMESTINAE DERMESTIDAE BOSTRICHOIDEA erichsonii Laricobius DERODONTIDAE DERODONTOIDEA pumilus Malthodes minimus Malthodes maurus Malthodes marginatus Malthodes guttifer Malthodes fuscus Malthodes flavoguttatus Malthodes fibulatus Malthodes dispar Malthodes seriepunctatus Malthinus (Fabricius) (Germar) (Waltl) Kiesenwetter Kiesenwetter (Laporte) (Brébisson) Linnaeus (Marsham) (Linnaeus) (Herbst) Kiesenwetter Kiesenwetter Suffrian Brahm Rosenhauer De Geer (Latreille) Kiesenwetter Kiesenwetter Kiesenwetter Kiesenwetter

Subfossil ●(1982) ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●(1984) ●(1982) ●(1997) ● ● ● ●(1994) ● ● ● ● ● ● ● ● Vc23 1939-2012 ●(1970s) ● ● ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 3029 3025 3023 3018 3014 3006 3005 3004 3003 3002 3001 3000 2994 2993 2992 2991 2990 2989 2987 2981 2980 2977 2976 2969 2968 2965 2961 2958 MEGATOMINAE MEGATOMINAE unicolor Attagenus pellio Attagenus ATTAGENINAE lardarius Dermestes Ptinus sexpunctatus Ptinus lichenum Ptinus fur Ptinus unicolor Tipnus PTININI imperialis Hedobia EUCRADINAE ANOBIIDAE Lyctus linearis Lyctus brunneus LYCTINAE BOSTRICHIDAE hirtus Trinodes TRINODINAE verbasci Anthrenus fuscus Anthrenus museorum Anthrenus ANTHRENINAE serraCtesias undata Megatoma nigripes Globicornis (Linnaeus) (Fabricius) (Goeze) (Fabricius) (Piller &(Piller Mitterpacher) (Linnaeus) (Stephens) Marsham Olivier (Linnaeus) (Linnaeus) (Brahm) (Linnaeus) Linnaeus Panzer (Fabricius) (Linnaeus) Rom/Med Med Subfossil ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ●(1985) ● ●(1993) ●(2001) Vc23 1939-2012 ● ●1998 ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 3072 3069 3067 3066 3065 3064 3063 3062 3061 3059 3057 3056 3055 3053 3052 3050 3049 3047 3046 3045 3044 3043 3040 3038 3037 3036 3034 3033 3031 Ochina ptinoides Ochina ERNOBIINAE pusillus Dryophilus planus Grynobius DRYOPHILINAE tectus Ptinus Caenocara bovistae bovistae Caenocara (= substriata Dorcatoma flavicornis Dorcatoma dresdensis Dorcatoma chrysomelina Dorcatoma ambjoerni Dorcatoma DORCATOMINAE longitarsis Xyletinus XYLETININAE pectinicornis Ptilinus PTILININAE denticollis Hadrobregmus punctatum Anobium inexspectatum Anobium fulvicorne Anobium paniceum Stegobium ANOBIINAE pini Ernobius nigrinus Ernobius mollis Ernobius rufovillosum Xestobium serra ) Boieldieu (Sturm) (Linnaeus) (Marsham) (Fabricius) (Sturm) (Gyllenhal) (Sturm) (Hoffmann, J.J.) Jansson (De Geer) (Linnaeus) (Linnaeus) Baranowski Herbst Hummel (Fabricius) (De Geer) Lohse Sturm (Creutzer in Panzer) Rom Med Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ●(1973) ●(1973) ● ● ● ● ●(1961) ● ●(2003) ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ●(1922) ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon

ecology 3122 3121 3120 3119 3118 3117 3116 3115 3113 3112 3106 3105 3104 3103 3102 3100 3099 3096 3095 3094 3093 3092 3085 3084 3083 3082 3081 3080 (= edwardsii Phloiophilus PHLOIOPHILIDAE CLEROIDEA (= impressus Aplocnemus RHADALINAE DASYTIDAE violacea Necrobia rufipes Necrobia ruficollis Necrobia Korynetescaeruleus KORYNETINAE formicarius Thanasimus (= femoralis Thanasimus mollis Opilo CLERINAE elongatus Tillus TILLINAE CLERIDAE elongatum Nemozoma mauritanicus Tenebroides TROGOSSITINAE pini rufipes edwardsi ) ) ) (Linnaeus) ) (Linnaeus) (De Geer) (Linnaeus) B3146 (Fabricius) (De Geer) (Zetterstedt) (Linnaeus) (Marsham) Stephens (Linnaeus) (Linnaeus) Med Med Subfossil ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●(1990) ●(1980) ●(2003) ● ●(1989) ●(2002) Vc23 1939-2012 ●(1990) ●(1970s) ●(1960) ●(1998) ●1998 ●(1970s) ●(1957) ● VC22 ● ● ● ● ● ● ● ● ● ● ● modern Oxon Cosmopolitan. Local in carrion, old bones old carrion, Local in Cosmopolitan.

ecology 3167 3166 3165 3164 3163 3162 3161 3160 3159 3158 3155 3154 3149 3146 3145 3144 3139 3136 3135 3132 3131 3130 3129 3128 3125 3124 Dasytesvirens Dasytesplumbeus Dasytesniger Dasytesaeratus DASYTINAE Kateretespusillus Kateretespedicularius KATERETIDAE orbiculatus Aspidiphorus dubius Sphindus SPHINDIDAE CUCUJOIDEA rufus Anthocomus fasciatus Anthocomus marginellus Clanoptilus bipustulatus Malachius aeneus Malachius viridis Cordylepherus marginalis Axinotarsus MALACHIIDAE viridicoeruleus Psilothrix (= puncticollis (Linnaeus) (Marsham) Stephens ) (Gyllenhal) (Thunberg) (Müller, O.F.) O.F.) (Müller, (Linnaeus) (Herbst) (Fabricius) (Linnaeus) (Linnaeus) (Linnaeus) (Laporte) (Olivier) (Gyllenhal) (Geoffroy in Fourcroy) Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●(2000) ●(2001) ● ●(1970s) ●(2012) ●(2012) ●(2012) ●(2012) Vc23 1939-2012 ●(1998) ●(1998) ●1998 ● ●(1998) ●(2001) ●(2012) ●(2012) ●(2012) ● ●(1997) ●(2012) VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Saproxylic. s and in FITs, rarelySaproxylic. in means.by s and other encountered slimefeeding wood, on fungoid Local,fungus. In vegetationlocallytall water near in reedPhragmites esp. beds flowers Widespreadherbage.on and 20 rapidlyWidespread, spreading in common on blossoms blossoms on common th C on grass and herbage, breeding in dead wood esp. oak. esp. wood dead in grass herbage,breeding on and C

ecology 3219 3218 3210 3209 3207 3206 3205 3203 3202 3201 3199 3198 3197 3196 3195 3187 3185 3184 3180 3179 3178 3177 3175 3171 3170 3169 3168 Brachypterolus pulicarius pulicarius Brachypterolus urticae Brachypterus glaber Brachypterus Kateretesrufilabris Epuraea melanocephala Epuraea limbata Epuraea (= pallescens Epuraea melina Epuraea (= marseuli Epuraea guttata Epuraea fuscicollis Epuraea distincta Epuraea (= biguttata Epuraea angustula Epuraea aestiva Epuraea EPURAEINI marginellus Carpophilus ligneus Carpophilus hemipterus Carpophilus CARPOPHILINI CARPOPHILINAE NITIDULIDAE ( = bipustulatus = florea pusilla unicolor ) ) ) (Linnaeus) Erichson ) (Olivier) (Fabricius) (Grimmer) Reitter (Thunberg) Sturm (Latreille) (Stephens) (Stephens) Murray B3214 Murray (Newman) (Fabricius) (Linnaeus) (Marsham) Motschulsky (Linnaeus)

Subfossil ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●(2012) ●(2012) ●(1999) ● ●(2012) ●(2012) ●(1990) Vc23 1939-2012 ●(2012) ●(1995) ●(2012) ●(1980s ●(1980s ●(1922) ●(2012) ●(2012) ●(2001) VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread on flowers of trees and shrubs, esp. rowan, hawthorn and Gorse. and flowers rowan, esp. hawthorn Widespreadon shrubs, of trees and synanthropic. Cosmopolitan, synanthropic. Cosmopolitan, Widespread, synanthropic, in houses and gardens, also at sap in the wild! wild! the in sap at also gardens, and houses in synanthropic, Widespread, wild! the in sap at also gardens, and houses in synanthropic, Widespread,

ecology 3262 3260 3259 3258 3257 3256 3255 3253 3252 3251 3250 3249 3247 3246 3245 3241 3240 3239 3235 3232 3231 3230 3227 3226 3224 3223 3222 3221 3220 Meligethes atratus Meligethes atramentarius Meligethes aeneus Meligethes dulcamarae Pria MELIGETHINAE Meligethes solidus solidus Meligethes (= ruficornis Meligethes rotundicollis Meligethes planiusculus Meligethes (= persicus Meligethes pedicularius Meligethes ovatus Meligethes ochropus Meligethes obscurus Meligethes nigrescens Meligethes morosus Meligethes lugubris Meligethes kunzei Meligethes gagathinus Meligethes fulvipes Meligethes flavimanus Meligethes difficilis Meligethes (= carinulatus Meligethes brunnicornis Meligethes bidens Meligethes flavipes pedicularius erythropus ) ) ) pre-1993) pre-1993) (Scopoli) Sturm Brisout de Barneville Erichson (Fabricius) (Olivier) (Kugelann) Brisout de Barneville (Faldermann) (Heer) Sturm Erichson Erichson Sturm (Marsham) Stephens Erichson Stephens Förster (Gyllenhal) Brisout de Barneville (Heer) Sturm Förster

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ●(2004) ● ● ● ● ●(2012) ●(2012) ●(2001) Vc23 1939-2012 ● ● ● ●(2004) ●(1996) ●(1996) ● ●(2012) ● ● ● ●(2000 ●(2000 ●(1996) ●(2012) ●(2012) ●(2012) ●(1995) VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Very local on rock-rose Helianthemum Verydowns on the local on black Localhorehound on crucifers, Rarecharlock on viper’s bugloss Veryon local betony Stachys Localon officinalisKirk-Spriggs common Localon hemp-nettle ground-ivy Local on marsh Rareon woundwort wood-sage Widespreadon white cloverWidespread on white dead-nettle Widespread on wild thyme Rareon yellow Localon archangel mint corn Rareon dog-rose. roses esp. Widespreadon white dead-nettle Widespread on wild basil Rareon Localon mostof the yellowone commonvarious on clothing, of beetles! crucifers, bright attracted also to Ubiquitous bittersweet Localon on Lotus corniculatus Lotus on sylvatica woundwortStachys hedge on Widespread Rosa especially dog-rose Mentha arvensis. arvensis. Mentha Clinopodium vulgare vulgare Clinopodium Thymus Solanum dulcamarae. Solanum Glechoma hederacea T hederacea Glechoma Ballota nigra Ballota Sinapis arvensis Sinapis Teucrium scorodonia S scorodonia Teucrium see Kkitrk Spriggs galeobdolon Lamiastrum Stachys palustris Stachys Trifolium repens Trifolium Echium vulgare Echium Galeopsis tetrahit tetrahit Galeopsis Lamium album Lamium album Lamium R.canina. Rosa canina Bagley canina Rosa on the downs the on & hedgemustard .

hotover ecology Sisymbrium officinale. officinale. Sisymbrium 3298 3297 3296 3292 3290 3289 3287 3286 3285 3284 3283 3282 3281 3280 3279 3277 3276 3275 3273 3272 3270 3269 3268 3267 3266 3265 3264 NITIDULINAE viridescensMeligethes umbrosus Meligethes symphyti Meligethes serripesMeligethes RHIZOPHAGINAE MONOTOMIDAE quadriguttatus Glischrochilus hortensis Glischrochilus undata Cryptarcha strigata Cryptarcha CRYPTARCHINAE fervida Thalycra ferrugineus Pocadius (= adustus Pocadius luteus Cychramus punctatissima Soronia grisea Soronia discoidea Omosita colon Omosita rufipes Nitidula carnaria Nitidula bipunctata Nitidula lanuginosus (Linnaeus) (Linnaeus) (Linnaeus) ) (Olivier) Reitter (Schaller) (Fabricius (Fabricius) (Olivier) (Gyllenhal) (Fabricius) (Linnaeus) (Heer) Sturm (Fabricius) (Fabricius) (Illiger) (Illiger) (Geoffroy in Fourcroy) (Fabricius)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●(2000) ●(1999) ●(1965) ●(2001) ●(2001) ● Vc23 1939-2012 ●(1980s ●(2001) ●(1980s ●(1905) ●(2002) ●(1980s ●(1970s) ●1998 ●(1998) ●(2001) ● ● ●(1950) ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local at sap, in fungi, etc. Local in atsap, fungiLocalpuffballs esp. in fungiLocal esp. in Localattrees also attackedatsap, by cossus. goatmoths Cossus barkof ash. Under carrion Local in carrion Local in carrion in carrion in vulgaris. self-heal Rare.on Prunella Comphrey On Kirk Spriggs in Oxon hemp red nettle on Lycoperdon spp.

ecology 3338 3335 3332 3331 3327 3326 3325 3324 3323 3322 3320 3319 3318 3317 3316 3312 3308 3307 3306 3305 3304 3303 3302 3300 3299 Rhizophagus parallelocollis Rhizophagus oblongicollis Rhizophagus nitidulus Rhizophagus ferrugineus Rhizophagus dispar Rhizophagus bipustulatus Rhizophagus Silvanus bidentatus Silvanus surinamensis Oryzaephilus Ahasverus advena SILVANINAE bipunctatus Psammoecus planata Uleiota BRONTINAE SILVANIDAE testacea Monotoma spinicollis Monotoma quadricollis Monotoma picipes Monotoma longicollis Monotoma brevicollis Monotoma bicolor Monotoma conicicollis Monotoma angusticollis Monotoma MONOTOMINAE cribratus Rhizophagus picipes Rhizophagus perforatus Rhizophagus (Linnaeus) (Waltl) Herbst Villa & Villa Motschulsky (Fabricius) (Paykull) (Olivier) Aubé (Gyllenhal) Aubé Gyllenhal (Fabricius) Aubé Guérin-Méneville Erichson (Gyllenhal) (Paykull) (Fabricius) (Fabricius) Blatch & Horner (Linnaeus) Gyllenhal RD3 Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●(2000) ● ●(2003) ●(1970s ● ● ●(2000) ●(2000) Vc23 1939-2012 ●(1998 ●(2001) ●(1999 ● ● ●(2001) ●(1986 ●(1986 ●(1986 ●(2000) ●(2001) ●(2001) VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local in compost and manure compostLocal heaps. and in marshsedgewetlands. Local,tussocks.Also and in in in litter compost &grassin etc.heaps decaying Widespreadin vegetable matter.s decaying other and vegetable compost heaps, Widespreadin matter.Regularly comes light. to decaying other and vegetable compost heaps, Widespreadin matter. grassLocal? compost In heaps. and ?Compost heaps. . treesbark of on manyLocal of old under atroots oftree kinds Also abundant Rare,saproxylic,sapy under bark of treesvarious woodlan gardens and in traps large subterranean in numbers in compost and in bark,also hardwood dead Under oftensubterranean and carrion bark,in hardwood dead Localunder Rare,saproxylic, subterranean?. barkof many Rareunder oftree kinds trees on of barkofold many atroots under Frequent ofabundant tree, also kinds barkof manyLocal under oftree kinds barkof many under Frequent of tree. Alsokinds atroots

ecology 3379 3378 3377 3376 3375 3374 3371 3370 3368 3367 3366 3364 3362 3359 3358 3357 3356 3355 3352 3350 3349 3346 3345 3344 3342 3341 3339 LAEMOPHLOEIDAE dermestoides Pediacus CUCUJIDAE unidentatus Silvanus CRYPTOPHAGINI CRYPTOPHAGINAE CRYPTOPHAGIDAE testaceus Stilbus oblongus Stilbus millefolii Olibrus liquidus Olibrus corticalis Olibrus affinis Olibrus aeneus Olibrus fimetarius Phalacrus corruscus Phalacrus caricis Phalacrus PHALACRIDAE clematidis Leptophloeus unifasciatus Notolaemus pusillus Cryptolestes spartii Cryptolestes ferrugineus Cryptolestes duplicatus Cryptolestes (Sturm) (Fabricius) Erichson (Panzer) (Erichson) (Paykull) (Panzer) Sturm (Curtis) (Schönherr) B3381 (Olivier) (Panzer) (Fabricius) (Fabricius) (Waltl) (Erichson) (Stephens) (Latreille) Na Subfossil ● ● ● ● ● ● ● ● ● ●? ● ● ● ● ● ● Vc22 VCH ●(2004) ●(1995) ●(2004) ●(2004) ●(2004) ●(2003) ●(2002) ●(2002) ? Vc23 1939-2012 ●(2004 ● ●(1980s ●(2004 ●(2004 ●(2011 ●(2002 ●(1999 ●(1950 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Frequent in dry in grassy Frequent debris Rare?reedmace On ( Rare?[On yellow & goat’s-beard, other Rare [mainly on composites. coastal] mayweeds, on Frequent tansy mugwortgrassy and rough in places smutted Localgrasses on smutted cereal on crops] smutted Local on Rare,saproxylic softwoods and barkof hard Local under bark,especially hardwood Local,beech. under softwoods. and barkLocal.of hard Under Nb ) ? [On yarrow [On ? ) Matricaria Carex Typha latifolia Typha Achillea ] ] atmargins ofwater.

ecology 3444 3442 3441 3440 3439 3429 3428 3425 3424 3423 3422 3421 3420 3419 3416 3415 3410 3409 3400 3394 3391 3390 3389 3388 3386 3385 3382 3381 Cryptophagus acutangulus Cryptophagus serratus Henoticus Paramecosoma typhae Telmatophilus schonherrii Telmatophilus caricis Telmatophilus Antherophagus pallens Antherophagus (= woodroffei Micrambe (= ulicis Micrambe Cryptophagus Cryptophagus scanicus Cryptophagus Cryptophagus Cryptophagus (= Cryptophagus (= Cryptophagus Cryptophagus Cryptophagus Cryptophagus Cryptophagus Cryptophagus Cryptophagus Cryptophagus Cryptophagus villosa villosa vini pallidus pilosus ) pre-2007) pre-2007) pre-2007) pre-2007) ) )

setulosus scutellatus saginatus ruficornis reflexus punctipennis pubescens populi lycoperdi laticollis distinguendus dentatus cellaris badius

melanocephalum (Stephens) (Gyllenhal) Johnson (Olivier) (Fallén) Paykull (Scopoli) Sturm Rey (Linnaeus) Lucas (Herbst) (Linnaeus) Sturm (Scopoli) Sturm Stephens Sturm Newman Gyllenhal (Gyllenhal) Brisout de Barneville Sturm Med (Herbst) Subfossil ● ● ● ● ●? ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●(1989) ●(2004) ●(1944) ●(2004) ●(2004) ●(1965) Vc23 1939-2012 ●(2004 ●(2011 ●(1907 ●(1956 ●(1907 ●(1909 ●(1954 ●(1907 ●(1982 ●(2001 ●(2011 ●(1980s ●(1970s ●(1989 ●(1984 ●(2004 ●(2004 ## VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local, breeds in nests of nests Local, in breeds commongorse on Bumblebee nests and burrows rabbit [in litter heathland in haystacks] and nests, vegetable in birds refuse.Synanthropic in [Also decayingLocal in matterorganic haystacks, in wasp in an nests especially wood fungoid and in dead Localon fungidecaying ?In and wood dead matter organic including Localhaystack usually in synanthropic refuse, hotbeds and grass clippings, decaying ?In matter.organic decaying and litter matter]fensriverorganic in banks and [Woods, ) Local in haystack fungiLocal mouldyand in other vegetable and matter. buildings in fungi and wood, dead Localon granaries, warehouse in etc.] Synanthropic granaries] and nests birds in nests, dreys, bird’s squirrel and in woodland [In cellars, warehousesmouldy in etc.in matteorganic Synanthropic, Rare,saproxylic. bark,atblossoms Under trees, bridgework in wire suspended fences debris etc1984 flood In on frequent Localbut Localon Scleroderma Carex in wetlands in Typha fungi Bombus on flower heads and hiding in leaf in sheaths. flower hiding on and heads Daldini. W Daldini.

ecology 3490 3489 3485 3484 3483 3475 3472 3471 3470 3468 3467 3464 3462 3461 3459 3457 3455 3454 3451 3450 3449 3448 3447 3446 3445 CAENOSCELINI (= similis Antherophagus (= silaceus Antherophagus (= Atomaria (Anchicera) lewisi (Anchicera) Atomaria fuscipes (Anchicera) Atomaria fuscata (Anchicera) Atomaria atricapilla (Anchicera) Atomaria atra (Anchicera) Atomaria apicalis (Anchicera) Atomaria (= vespertina (s.str.) Atomaria punctithorax (s.str.) Atomaria (= pulchra (s.str.) Atomaria nigriventris (s.str.) Atomaria nigrirostris (s.str.) Atomaria (= linearis (s.str.) Atomaria fimetarii (s.str.) Atomaria diluta (s.str.) Atomaria (= ATOMARIINI HYPOCOPRIN1 ATOMARIINAE ferruginea Caenoscelis pulchra barani procerula sahlbergi pallens canescens nigricornis ) ) pre-2006) pre- 2007) pre- ) ) ) ) ) Curtis Erichson (Sahlberg, C.R.) (Herbst) Stephens Erichson (Fabricius) (Herbst) Mäklin Stephens Reitter Stephens (Schönherr) Erichson Reitter (Gyllenhal) Stephens Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ●? ● Vc22 VCH ● ● ● ●(1994) ● Vc23 1939-2012 ●(1970s ●(1996 ●(1922 ●1998 ●(1998 ● ●(1907 ●(1980 ●(1996 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare [In dung and decaying and dung vegetable Rare matter][In decaying other compostvegetable and Widespreadin matter wetlands] Rare?[In decaying other compostvegetable and in frequent Widespread matterand decaying Rare? [In vegetable matter litter] wetlands in Rare?[In haystack heaps, [Compost refuse.] moss and litter in woodland Widespreadin compost etc dung, Local in grasslandfarmland, esp. In shaggyRare, declining? cornatus on fungiink-cap Coprinus of nests Local, in breeds of nests Local, in breeds Bombus Bombus

ecology 3537 3536 3535 3534 3533 3531 3528 3527 3526 3525 3524 3520 3519 3518 3517 3516 3514 3513 3510 3509 3508 3505 3502 3498 3497 3496 3494 Atomaria (Anchicera) peltata (Anchicera) Atomaria nitidula (Anchicera) Atomaria nigripennis (Anchicera) Atomaria munda (Anchicera) Atomaria mesomela (Anchicera) Atomaria Byturus tomentosus tomentosus Byturus ochraceus Byturus BYTURIDAE bipustulata Tritoma russica Triplax aenea Triplax rufifrons Dacne bipustulata Dacne EROTYLIDAE reitteri Ephistemus globulus Ephistemus globosus Ootypus zetterstedti(Anchicera) Atomaria (= testacea(Anchicera) Atomaria rubida (Anchicera) Atomaria (= rubella (Anchicera) Atomaria pusilla (Anchicera) Atomaria ruficornis berolinensis (Schaller ) ) (Linnaeus) (Fabricius ) ) (Waltl) (Thunberg ) (Scriba) Casey Fabricius (DeGeer) (Paykull) Reitter (Paykull) Kraatz Erichson Heer (Marsham) Stephens (Herbst) Rom (Zetterstedt) (Kugelann) Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ●(2001) ●(2003) ● ● ● ● ●(1998) ●2000 Vc23 1939-2012 ●2001 ●1970s ● ●(1998 ● ● ● ●(1997 ●(1987 ●(1987 ●(1910 ●(1970s ● ●(2001 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread on blossoms,larvae fruits of Widespreadon in spring the blossoms in Localon esp. Wytham 1980s Wood hardwoods] fungi on bracketsother and ? Saproxylicold on fungi. Local,saproxylic.on barkand Under fungi. Local.on Saproxylic. barkand Under fungi. Local.on Saproxylic. barkand Under marsh wetlandsLocal in in litter Rubus Rubus

and a ofraspberry pest cultivated and etc. ecology 3572 3571 3570 3569 3568 3567 3566 3565 3564 3562 3561 3560 3559 3558 3557 3556 3555 3554 3553 3552 3551 3550 3548 3543 3542 3541 3540 3539 3538 BOTHRIDERIDAE fagi Diplocoelus lunatus Biphyllus BIPHYLLIDAE Holoparamecus depressus Holoparamecus caularum Holoparamecus MEROPHYSIINAE latus Symbiotes ANAMORPHINAE coccineus Endomychus ENDOMYCHINAE ENDOMYCHIDAE pilosum Sphaerosoma ALEXIIDAE histeroides Cerylon ferrugineum Cerylon fagi Cerylon CERYLONINAE segregatus Murmidius ovalis Murmidius MURMIDIINAE CERYLONIDAE duodecimstriatus Anommatus ANOMMATINAE Brisout de Barneville Redtenbacher (Chevrolat in Guérin-Méneville) ?* (Beck) (Fabricius) (Fabricius) Stephens (Panzer) Waterhouse, C.O. (Linnaeus) Curtis B29 (Aubé) (Müller, P.W.J.) P.W.J.) (Müller,

Subfossil ● ● ● ● ● ● ● ● Vc22 VCH ● ●(2003) ● ● Vc23 1939-2012 ●(1970s) ●1998 ●1998 ●1998 ●(1998 ● VC22 ● ● ● ● ● ● ● ● ● ● modern Oxon Local. Synanthropic in manure and compost heaps etc.manureFormerly compost heaps in and Local. Merophysidae Synanthropic in placed wood fungoid Rarein birch. and beech esp. wood fungoid Localon etc compost heaps in Local.Synanthropic oak and beech esp. barkof hardwoods under oak . and beech esp. barkof hardwoods Rare,under Rare,subterranean fungi the Local in under bark of hardwoods esp. beech and oak oak and beech esp. hardwoods of bark under Daldina concentrica Daldina especially ash. on

ecology 3607 3606 3604 3603 3601 3600 3599 3598 3597 3596 3595 3592 3591 3590 3586 3585 3584 3583 3582 3581 3580 3579 3578 3577 3576 3575 3574 3573 Mycetaea subterranea Mycetaea MYCETAEINAE succincta Lycoperdina bovistae Lycoperdina LYCOPERDININAE singularis Holoparamecus CHILOCORINAE pseudopustulata Hyperaspis redtenbacheri Nephus quadrimaculatus Nephus limbatus (Neopullus) Scymnus haemorrhoidalis (Neopullus) Scymnus suturalis (Pullus) Scymnus auritus (Pullus) Scymnus schmidti (s.str.) Scymnus rubromaculatus (s.str.) Scymnus nigrinus (s.str.) Scymnus frontalis (s.str.) Scymnus femoralis (s.str.) Scymnus punctillum Stethorus litura Rhyzobius chrysomeloides Rhyzobius scutellata Coccidula rufa Coccidula COCCIDULINAE COCCINELLIDAE (= hirta ) ) (Herbst) (Fabricius) (Herbst) (Weise) (Fabricius) (Mulsant) (Fabricius) (Linnaeus) Kugelann (Fabricius) Fürsch (Herbst) Thunberg (Gyllenhal) (Beck) (Herbst) Thunberg Mulsant Stephens Rom Rom/Med (Goeze)B3625 Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ●?? ?? Herbst Vc22 VCH ● ● ● ● ● ●1994 ● ● Vc23 1939-2012 ●(1914 ●(1914 ●(1996 ●2012 ●2004 ●(2002 ●(1984 ● ●(1991 ● ● ●(1988) ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare on sallowswillows Rare and on grasslandLocal in pines Localon foliage Localon of oak trees young Verypines on local shrubs trees and on .local usually frequent, Widespreadterrestrial and Recentimmigrant, bushes. trees and Increasing on wetlands, oftenamongst in local emergenttall wetlands, on Widespread in waterside vegetation hay/straw rotting compost heaps wood, fungoid in in puffballs in Typha

ecology 3639 3638 3637 3636 3635 3634 3633 3632 3631 3627 3626 3625 3624 3623 3622 3621 3620 3619 3618 3617 3616 3615 3614 3612 3611 3610 3609 3608 Anisostica novemdecimpunctata Anisostica COCCINELLINAE quadripustulatus Exochomus renipustulatus Chilocorus bipustulatus Chilocorus luteorubra Platynaspis Henosepilachna argus Henosepilachna EPILACHNINAE quadripunctata Harmonia axyridis Harmonia decempunctata Adalia bipunctata Adalia undecimpunctata Coccinella septempunctata Coccinella quinquepunctata Coccinella hieroglyphica Coccinella variegata Hippodamia tredecimpunctata Hippodamia obliterata Aphidecta ocellata Anatis vigintiduopunctata Psyllobora sedecimguttata Halyzia (Poda) duodecimguttata Vibidia quattuordecimguttata Calvia quattuordecimpunctata Propylea octodecimguttata Myrrha oblongoguttata Myzia sedecimpunctata Tytthaspis (Linnaeus) (Linnaeus) (Pallas) (Linnaeus) (Linnaeus) (Linnaeus) (Goeze) (Geoffroy in Fourcroy) (Goeze) (Linnaeus) (Linnaeus) Linnaeus (Linnaeus) (Scriba) (Pontoppidan) Linnaeus (Linnaeus) Linnaeus Linnaeus (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) Rom Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ●? ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ● ●1976 ● ● ● ● ● ● ●(1984 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local, increasing in ruderal areas, esp. on light soils light areas, on Local,ruderal esp. increasing in Extinct? firsoccasionallypines. larch, Localand on on pine scot’s Localon Wychwood UK in Last in Extinct. 1905 forest recorded in Ash, LocalLimetrees on broadleaved other and pine scot’s Localon sycamore ash, trees including on confiers. and Ubiquitous sallow. and alder treesash, various esp. Widespread,on downs the on junipers and heathland Localon grassy Rare?open In placesant withthe on white bryony. on pines on rare Lasius niger Lasius

ecology sens str. sens str. 3686 3685 3684 3682 3680 3679 3676 3674 3673 3670 3669 3668 3667 3665 3664 3663 3661 3660 3659 3654 3653 3651 3648 3646 3645 3643 3642 3641 3640 Corylophus cassidoides Corylophus lateralis Sericoderus CORYLOPHINAE CORYLOPHIDAE vigintiquattuorpunctata Subcoccinella Dienerellaclathrata transversus Enicmus testaceus Enicmus histrio Enicmus brevicornis Enicmus bergrothi Thes minutus Latridius (= assimilis Latridius nodifer (Aridius) Cartodere bifasciata (Aridius) Cartodere constricta (s.str.) Cartodere lardarius Stephostethus (= angusticollis Stephostethus alternans Stephostethus LATRIDIINAE LATRIDIIDAE (= corticalis Orthoperus atomus Orthoperus ORTHOPERINAE pseudominutus kokujevi mundus ) ) ) (Reitter) Joy & Tomlin Joy Mannerheim (Stephens) (Linnaeus) ) (Gyllenhal) (Mannerheim) (Gyllenhal) (Mannerheim) (Olivier) (Redtenbacher) (Marsham) (De Geer) (Mannerheim) (Gyllenhal) (Gyllenhal) (Westwood) (Reitter) Rom Rom Subfossil ● ● ● ●? ● ● ● ● ● ● (Linnaeus) Vc22 VCH ● ●1980) ●(1969) ● ● ● ● ● ● ● ●(1978) ● Vc23 1939-2012 ●1998 ●1998 ●(1970s ●(1998) ●(1998) ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread in grasslands Widespreadin woodlands Local in haystacks. grassheapsold and including litter in sycamore and beech, bark,esp. Localunder with sooty bark disease. trees. ?LocalWoodland. on decaying Raredebris in plant decaying Raredebris in plant

ecology 3741 3739 3738 3736 3735 3734 3733 3732 3727 3726 3724 3722 3721 3720 3714 3711 3705 3704 3703 3701 3699 3698 3697 3694 3690 3689 3688 3687 CORTICARIINAE Dienerellafilum (= Dienerellavincenti Dienerellaruficollis (= Mycetophagus multipunctatus Mycetophagus atomarius Mycetophagus connexus Litargus bicolor Triphyllus suturalis Pseudotriphyllus MYCETOPHAGIDAE TENEBRIONOIDEA (= suturalis Melanophthalma gibbosa Cortinicara similata Corticarina (= minuta Corticarina serrata Corticaria punctulata Corticaria inconspicua Corticaria impressa Corticaria fulva Corticaria ferruginea Corticaria elongata Corticaria crenulata Corticaria alleni Corticaria transversalis fuscula elongata separanda ) ) ) (Comolli) Johnson in part) part) in (Aubé) (Paykull) (Fabricius) (Geoffroy in Fourcroy) Johnson (Fabricius) (Gyllenhal) (Herbst) (Marsham) (Olivier) (Gyllenhal) (Gyllenhal) Marsham Marsham Wollaston (Fabricius) (Fabricius) (Mannerheim) Fabricius N Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ●?? ● ? Vc22 VCH ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ●1998 ●(2001) ● ● ● ● ●(1998) ●(1945) VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local in fungi on trees, and under bark. under trees, and fungiLocal on in hardwoods. fungoid fungi Rareand ) in hardwoods. fungoid fungiLocal and in Riparian

ecology 3780 3779 3778 3777 3774 3770 3768 3766 3765 3764 3763 3762 3760 3759 3758 3757 3756 3754 3753 3752 3751 3750 3748 3747 3746 3745 3743 3742 Eulagius filicornis Eulagius stercorea Typhaea haagi Typhaea quadripustulatus Mycetophagus populi Mycetophagus piceus Mycetophagus Cis vestitus Cis submicans Cis pygmaeus Cis micans Cis hispidus Cis fagi Cis castaneus Cis boleti Cis bilamellatus Cis bidentatus alni Orthocis bicornis Strigocis nitidus Sulcacis CIINI glabriculus Octotemnus OROPHIINI CIINAE CIIDAE (=micans) (=nitidus) (=affinis) Waltl (Scopoli) (Fabricius) Mellié Mellié (Paykull) (Gyllenhal) (Herbst) (Marsham) Abeill de Perrin (Olivier) Reitter (Fabricius) Wood (Mellié) (Mellié) (Linnaeus) (Reitter) (Fabricius) Fabricius (Gyllenhal) (Linnaeus) Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●(1996) ●(1980) ● ● ●(1982) ● ● ● ● Vc23 1939-2012 ●(1956) ●(1998) ●(1998) ●(1964) ● ●(2001) ●(1950 ● ● ● ●(2000) VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local on dead oak branches [also in Polypore fungi on elm beech] fungi on and Polypore in [also oak branches dead Localon oak. esp. hardwoods, fungi on Local on oak esp. hardwoods, fungi on Local on bracket Localfungi on oak. rotten red esp. wood fungoid Rarein hardwoods fungi on Local on soft brackets.in esp wood fungoid fungi and in Frequent Australasian, Naturalised, 20 appeared bracket Rare,fungi.in jew’s on Fungus esp. Ear hardwood, fungoid in fungi and Local on soft Rarebrackets) in fungiLocalTrametes esp. in versicolor bracket fungiin esp. Frequent hardwood fungoid fungi and on Naturalised, bark. haystack Widespreadhardwood in occasionally and under debris bark. under trees, and fungiLocal on in newly recognised in UK in newly recognised Trametes Trametes Trametes etc.Sunnymead spp. on hardwoods. on spp. th Century. Local in Birch Polypore Polypore Century. Birch Local in esp. in beech woods. beech in esp.

ecology Auricularia auricula-judae auricula-judae Auricularia Piptoporus betulinus Piptoporus . Elder on esp. 3813 3812 3811 3810 3809 3804 3803 3802 3801 3800 3798 3797 3796 3795 3794 3793 3792 3790 3789 3788 3787 3786 3785 3784 3783 3782 3781 HALLOMENINAE TETRATOMIDAE cornutum Ennearthron (= Cis villosulus Osphya bipunctata Osphya testaceus Conopalpus OSPHYINAE caraboides Melandrya vaudoueri Phloiotrya triguttata Abdera quadrifasciata Abdera flexuosa Abdera biflexuosa Abdera fuscula Anisoxya Orchesiaundulata Orchesiaminor Orchesiamicans MELANDRYIDAE fungorum Tetratoma desmarestii Tetratoma ancora Tetratoma TETRATOMINAE binotatus Hallomenus setiger ) (Marsham) Walker (Paykull) (Gyllenhal) (Illiger) (Illiger) (Panzer) (Curtis) Kraatz Fabricius (Fabricius) Fabricius Mulsant (Olivier) (Quensel) (Linnaeus) (Curtis) Latreille (Gyllenhal)

Subfossil ● ● ● ● ● ● ● ● ● Vc22 VCH ●(1991 ●(1992 ●(1990 ●(1978) ● ●2008 ● ● ●(2002 ● ● Vc23 1939-2012 ●(1998) ●(1998) ●(1998) ● ●1998 ● ● ●(1960 ● ●(1970s ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local on dead and moribund oak boughs on large trees. on oak boughs moribund and dead Localon oak. esp. hardwoods on oak. of esp and beech boughs hardwood fungoid on V.local decaying and wood fungoid bracket ApparentlyfungiLocal fungi. and in very 19 rare in wood fungoid fungi and in V.local including wood fungoid fungi and in V.local polypore birch including wood, fungoid fungi and Local on Saproxylic,onoak winter active. wood fungoid fungi and in V.local wood. fungoid fungi and Widespreadon wood. bracket fungoid fungiin and On oak hazel on canker canker on hazel On oak Inonotus

ecology

radiatus Piptoporus betulinus. betulinus. Piptoporus on Alder. on th C!! 3866 3863 3862 3858 3857 3856 3855 3851 3850 3849 3848 3847 3846 3843 3841 3839 3835 3833 3829 3828 3827 3826 3825 3824 3823 3816 3815 3814 MORDELLISTENINI villosa Variimorda Tomoxiabucephala MORDELLINI MORDELLINAE MORDELLIDAE COLYDIINI crenata Bitoma variegatus Cicones SYNCHITINI Orthocerus clavicornis ORTHOCERINI COLYDIINAE COLYDIIDAE paradoxus Metoecus RIPIPHORIDAE abdominalis Mordellochroa (= acuticollis Mordellistena variegata Mordellistena pumila Mordellistena pseudopumila Mordellistena pseudoparvula Mordellistena parvula Mordellistena neuwaldeggiana Mordellistena humeralis Mordellistena brevicauda Mordellistena imitatrix ) (Fabricius) (Schrank) (Hellwig) (Hellwig) (Linnaeus) (Gyllenhal) (Linnaeus) (Gyllenhal) (Fabricius) Schilsky (Linnaeus) (Boheman) (Fabricius) Ermisch Ermisch (Panzer)

Subfossil ● ● ● ● ● ● Vc22 VCH ● ● ●(2000 ●(1999) ●(2001) ●1980 ●?? Vc23 1939-2012 ● ● ●(2004) ●(1999) ●(2004) ●(2002) ●(1998) ●(2003) ●(1998) ●(2002) ●(1998) ●(1970s) ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread under bark of hardwoods, and conifers. and barkof hardwoods, Widespreadunder dry fungi and deaying Rare on hardwoods. fungoid ofnests Local waspssocial in muchrecords with confused Old hedgerowswoods. Rare and on chalk downland Rareon local on mugwort on local Vespidae , sometimes , houses. in

ecology variegata 3917 3915 3914 3910 3908 3905 3904 3900 3903 3899 3898 3894 3892 3891 3888 3886 3884 3883 3881 3880 3879 3878 3876 3875 3873 3872 3871 3869 TENEBRIONINAE hirta Lagria LAGRIINAE TENEBRIONIDAE trisulcus Aulonium Gonodera luperus luperus Gonodera ater Prionychus ALLECULINAE cornutus Gnatocerus metallicum Scaphidema fraxini Corticeus bicolor Corticeus quisquilius Crypticus vaporariorum Myrmechixenus DIAPERINAE mucronata Blaps laevioctostriatus Nalassus sabulosum Opatrum tibialis Melanimon subdepressus Palorus oryzae Latheticus confusum Tribolium castaneum Tribolium diaperinus Alphitobius obscurus Tenebrio molitor Tenebrio agricola Eledona (Linnaeus) (Fabricius) Linnaeus (Kugelann) (Olivier) (Herbst) Latreille Waterhouse, C.O. Fabricius (Herbst) (Fabricius) (Geoffroy in Fourcroy) (Linnaeus) Jacquelin du Val (Fabricius) (Linnaeus) (Wollaston) (Herbst) (Panzer) (Fabricius) (Goeze) Guérin-Méneville

Subfossil ● ● ● ● ● ●? ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ●(1984) ●(1993) ● ● ● ● ●1980 Vc23 1939-2012 ● ● ● ● ● ● ● ●1998 ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Saproxylic. Adults on vegetation in rides and woodland edge habitats. woodland and rides vegetationSaproxylic. in Adults on oak. red-rotten esp. wood rotting Local in grainetc.] stored in Synanthropic stables. old Also elder. esp. in wood fungoid Localon elm and ash bark withRare Scolytids under esp. locally Frilford abundant grass and compostcuttings. Rare heaps in grainetc.] stored in Synanthropic grainetc. stored in Synanthropic comeswild, also fungi in light readilyon also MV heaps to Manure Local,in stables. in etc.also flour stored in Synanthropic etc.Oftenlight. flour attracted to stored In Synanthropic. fungi,Local usually in Chicken-of-the-woods elm Rareon attacking Scolytids etc. Laetiporus sulphureus Laetiporus Scolytus multistriatus Scolytus

ecology esp. on oak and cherry. oak and on esp. & S. scolytu S. s in elm s in bark. 3962 3961 3960 3959 3953 3952 3951 3950 3948 3947 3940 3939 3938 3936 3935 3933 3932 3930 3929 3928 3926 3925 3924 3921 3919 3918 Nacerda melanura Nacerda OEDEMERIDAE sulphureus Cteniopus murina Isomira ceramboides Pseudocistela Pyrochroa serraticornis Pyrochroa coccinea Pyrochroa PYROCHROIDAE Mycterus curculioides MYCTERIDAE muralis Apalus NEMOGNATHINAE violaceus Meloe rugosus Meloe proscarabaeus Meloe MELOINAE MELOIDAE nobilis Oedemera lurida Oedemera femoralis Oedemera sanguinicollis Ischnomera cyanea Ischnomera cinerascens Ischnomera caerulea Ischnomera Marsham (Forster) (Linnaeus) Marsham (Marsham) (Scopoli) (Linnaeus) (Fabricius) (Linnaeus) (Olivier) (Linnaeus) Linnaeus (Fabricius) (Linnaeus) (Scopoli) (Pandellé) (Fabricius) (Linnaeus) Rom Subfossil ● ● ● ● ● ● ● ● ● ● ? Vc22 VCH ● ● ● ● ● ● ) ●(19 ●(1985) ● ●?? ●(1986) ● Vc23 1939-2012 ● ●? ● ● ● ● ●(1998 ● ●(1980s ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●( modern Oxon Widespread, larvae under bark of hardwoods and conifers and barkWidespread,oflarvae hardwoods under wallsolitary nesting [Parasitizes Extinct? bees] common saproxylic. woods, Rare in Comes light to soft elm.blossoms. old Adults on in develops , records. 1980 post 5 atblossoms. decaying adults in .local, hardwoods, decaying Rare,blossoms. in on elm oak,adults and Rare,casual import?by Primarilytimberor saltwater in coastal on decaying Rare,blossoms. in on elm oak,adults and commons flowers. and woods Widespread on in comes saproxyliclight. hardwoods, to on common increasing common

ecology 4002 4001 4000 3999 3998 3997 3996 3995 3992 3991 3984 3982 3981 3980 3979 3978 3977 3975 3974 3970 3969 3968 3967 3966 3965 3964 3963 SALPINGINAE brunneus Aglenus AGLENINAE SALPINGIDAE pectinicornis Schizotus SCRAPTIIDAE brevicornis Vanonus oculatus Euglenes populneus Aderus ADERIDAE tobias Stricticomus formicarius Omonadus floralis Omonadus bifasciatus Omonadus antherinus Anthicus monoceros Notoxus ANTHICIDAE ruficollis Salpingus planirostris Salpingus Vincenzellus ruficollis castaneus Sphaeriestes ater Sphaeriestes denticolle Lissodema cursor Lissodema (Paykull) (Gyllenhal) (Paykull) (Gyllenhal) (Creutzerin Panzer) (Linnaeus) (Marseul) (Linnaeus) (Linnaeus) (Linnaeus) (Perris) (Gyllenhal) (Fabricius) (Rossi) (Panzer) (Panzer) (Linnaeus) (Goeze) Rom Med Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ●(1998 ●2000 ●(1998 ●(2004 ●(1998 ●(1998 ●(1998 ●(1998 ●(1998 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local on oaks, larvae in red-rot, adults on foliage on adults oaks, larvaeLocal red-rot, on in foliage on adults oaks, larvaeLocal red-rot, on in haystack in and grass compost heaps, debris. in frequent and Widespreadand manure compost heaps. Rare and in etc.compost heaps in Widespread.Synanthropic sandy areas soils. on disturbed open Rarein hardwoods moribund and dead Localon hardwoods moribund and dead Widespreadon especially beech hardwoods, and hawthorn moribund and dead Localon conifers. moribund and dead on trees. on up dying high ofbranches in Ashtips Saproxylic, Fraxinus, on Synanthropic.

ecology 4052 4051 4046 4044 4043 4039 4037 4036 4035 4033 4032 4031 4029 4026 4025 4024 4023 4022 4020 4017 4016 4015 4014 4013 4012 4011 4010 4009 4007 Anaspis lurida Anaspis garneysi Anaspis frontalis Anaspis (= fasciata Anaspis ANASPIDINAE Pseudovadonia livida Pseudovadonia tabacicolor Alosterna scutellata Stictoleptura sanguinolenta Anastrangalia quadrifasciata Leptura ustulata Grammoptera ruficornis Grammoptera (= abdominalis Grammoptera meridianus Stenocorus mordax Rhagium bifasciatum Rhagium LEPTURINAE CERAMBYCIDAE CHRYSOMELOIDEA rufilabris Anaspis costai Anaspis thoracica Anaspis regimbarti Anaspis pulicaria Anaspis maculata Anaspis variegata humeralis ) ) ) Emery Stephens (Forster) Fowler Fowler (Linnaeus) Costa, A. (De Geer) (Geoffroy in Fourcroy) (Gyllenhal) (Linnaeus) Schilsky (Fabricius) (De Geer) Fabricius (Schaller) (Fabricius) Linnaeus (Linnaeus) (Fabricius) (Stephens) (Linnaeus)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ●(1996) ● ● ● ●(1975) ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ●(1998 ●(1998 ●(1998 ●(1998 ●(1998 ●(1998 ●(1998 ●(1998 ●(1998 ●(1998 ●(1998 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon V.local, adults atflowersadults umbels. esp. V.local, flowers on widespreadumbels esp. Localadults but Rare,increasing? Larvae birch. oak,and beech, in deaying stumps birch. in esp. and Local,develops wood commonest heaths, wooded on a variety size. Breedsdespite in illusive woodlands, Local of hardwoods. in ash. oak and esp. hardwoods Widespreadin pine. in esp. heaths wooded and woods in Rare,saproxylic, atblossoms. adults etc sloe hawthorn, blossoms esp. on Local,adults etc sloe hawthorn, blossoms esp. on Local,adults etc sloe hawthorn, blossoms esp. on Local,adults etc. sloe hawthorn, blossoms esp. on oftenabundant Widespreadand etc. sloe hawthorn, blossoms Local, esp. saproxylic, on adults etc sloe hawthorn, blossoms esp. on Local,adults etc sloe hawthorn, blossoms esp. saproxylic, frequent, on Widespreadand adults etc. sloe hawthorn, blossoms esp. saproxylic, frequent, on Widespreadand adults

ecology 4097 4093 4091 4090 4089 4088 4085 4084 4083 4082 4080 4079 4078 4076 4074 4073 4072 4071 4067 4063 4062 4061 4058 4057 4056 4055 4054 Tetropium gabrieli Tetropium striatum Asemum SPONDYLIDINAE nigra Stenurella melanura Stenurella maculata Rutpela Phytoecia cylindrica Phytoecia populnea Saperda nebulosus Leiopus linnei Leiopus hispidus Pogonocherus hispidulus Pogonocherus villosoviridescens Agapanthia nebulosa Mesosa LAMIINAE mysticus Anaglyptus arietis Clytus alni Poecilium testaceus Phymatodes violaceum Callidium bajulus Hylotrupes moschata Aromia umbellatarum Glaphyra minor Molorchus minuta Gracilia CERAMBYCINAE rusticus Arhopalus ferus Arhopalus (Linnaeus) (Linnaeus) (Fabricius) (Mulsant) (Linnaeus) (Fabricius) (Linnaeus) (Poda) (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) Weise (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) (von Schreber) (Piller &(Piller Mitterpacher) (De Geer)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ? ● ● ● Vc22 VCH ● ? ● ● ●(2004 ● ● ● ● ● ● ● ● Vc23 1939-2012 ●1998 ● ● ●(2000 ●(1996 ●(1998 ● ● ● ●(1998 ● ●(1988 ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon V.local on aspen on damp heathland and woodland rides woodland and damp heathland on aspen on V.local oflargebranches attached trees but dead / oak preferringusually on moribund oflargebranches attached trees but dead / oak preferringusually on moribund a wide Localvariety on Holly on esp. of trees, adults umbellifers, and thistles increasing ?in Rare,declining. a variety [In ? of trees] willows Rarein water near dying and woody dead RosaceaeRare, in typically breeds hawthorn, esp. sickly on specimens. looking a varietyRare Naturalised, in of treesspruce? Adultsvisit blossom. esp. largely and much replaced increased since1950s conifers pine, Local, esp. in woodland. ancient arein commonestheaths widespread wooded perhaps Local, but on hardwoods in atblossoms,breeding Frequent

ecology A.tristis 4135 4134 4132 4131 4130 4125 4124 4123 4121 4120 4119 4118 4117 4115 4114 4113 4112 4111 4110 4109 4108 4107 4105 4103 4102 4101 4100 4099 4098 ZEUGOPHORINAE MEGALOPODIDAE starkii Tetrops praeustus Tetrops oculata Oberea Donacia cinerea Donacia bicolora Donacia mutica Macroplea appendiculata Macroplea DONACIINAE villosus Bruchidius varius Bruchidius olivaceus Bruchidius cisti Bruchidius rufipes Bruchus rufimanus Bruchus pisorum Bruchus loti Bruchus atomarius Bruchus BRUCHINAE CHRYSOMELIDAE (= humeralis Orsodacne cerasi Orsodacne ORSODACNIDAE subspinosa Zeugophora lineola ) Paykull Chevrolat (Linnaeus) (Fabricius) Herbst Herbst (Linnaeus) Zschach (Linnaeus) (Linnaeus) (Olivier) (Fabricius) (Fabricius) (Linnaeus) Boheman (Germar) Latreille (Fabricius) (Panzer)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ?? Vc22 VCH ● ● ● ● ● ●2001 ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ●1990 ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon on Typha.V.local on reeds PhragmitesTypha.V.local atmargins on on oflakes ponds. and formerly Extinct Sparganium erectum. on usually wherever Local,present but of Broom stands decent Recentimmigrant, clover. red wellnowwidespread on and established rock-rose on Rare,adults vetchesvarious Widespreadon legumesbeans cultivated various Localon including pulses. and beans imported ?On Meadow LocalVetchling on vetches On Rosaceae. allied and Hawthorn Rareon Rare Aspen on (TDH) UK atWidley in first ash 1991 found Copse on treesrosaceous on local Populus tremulae Populus Vicia, Lathyrus. Vicia, Helianthemum , and Willows and , Lathyrus pratensis Lathyrus Vicia , and possiblyclovers and , on Downs, larvae on in Salix. Salix.

ecology Trifolium Cytisus Lotus ?? grow. spp. 4168 4165 4164 4163 4162 4160 4159 4158 4157 4155 4154 4153 4152 4151 4150 4149 4148 4147 4146 4145 4144 4143 4142 4140 4139 4138 4137 4136 Donacia semicuprea Donacia marginata Donacia impressa Donacia dentata Donacia crassipes Donacia clavipes Donacia Clytra quadripunctata Clytra CLYTRINI CRYPTOCEPHALINAE lilii Liliocerus Crioceris asparagi (= rufocyanea Oulema (= obscura Oulema melanopus Oulema erichsoni Oulema cyanella Lema CRIOCERINAE sericea Plateumaris discolor Plateumaris braccata Plateumaris affinis Plateumaris vulgaris Donacia versicolorea Donacia thalassina Donacia sparganii Donacia simplex Donacia melanopus lichenis ) (Scopoli) (Linnaeus) in part) part) in Hoppe (Stephens) Fabricius Fabricius Zschach (Suffrian) Paykull (Linnaeus) Ahrens Fabricius (Kunze) Germar (Linnaeus) Hoppe (Linnaeus) (Suffrian) Panzer (Panzer) (Brahm) (Scopoli) (Linnaeus)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ? ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ●2009 ● ● ●1997 ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ● ● ●2011 ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local, increasing, on various gardenlilies various Local,increasing,on very gardens,neglectedasparagusWidespread allotmentsbut in etc. on local dampwetlands Widespreadgrassland,in and acidic wetlands in local sedges byon local streams on pondweed broad-leaved Localon Carex Rareon Sparganium] [On Extinct? Widespreadon on typha etc on local Rare,on arrowhead [on Extinct? white Rarelily on Nymphaea O alba reeds Phragmites atmarginson ofV.local lakes ponds. and local in fens and rich wetlands rich and fens in local Typha, Sparganium Typha, Glyceriamaxima Scirpus Sparganium at margins of lakes and ponds. atmargins oflakes ponds. and atwater marginsflowing beside esp. water. etc. Sagittaria at margins of ponds, lakes,atmargins canals rivers and ofponds, Potamogeton natans natans Potamogeton ]

ecology in ponds and pools. and ponds in 4215 4214 4213 4208 4206 4203 4200 4199 4198 4197 4196 4195 4192 4191 4187 4185 4184 4183 4182 4181 4180 4179 4178 4174 4173 4172 4171 4170 4169 Cryptocephalus bipunctatus Cryptocephalus bilineatus Cryptocephalus biguttatus Cryptocephalus aureolus Cryptocephalus CRYPTOCEPHALINI affinis Smaragdina Chrysolina sanguinolenta Chrysolina polita Chrysolina oricalcia Chrysolina hyperici Chrysolina herbacea Chrysolina fastuosa Chrysolina brunsvicensis Chrysolina banksi Chrysolina americana Chrysolina tenebricosa Timarcha goettingensis Timarcha CHRYSOMELINAE concolor Oomorphus LAMPROSOMATINAE pusillus Cryptocephalus primarius Cryptocephalus parvulus Cryptocephalus nitidulus Cryptocephalus moraei Cryptocephalus labiatus Cryptocephalus hypochaeridis Cryptocephalus fulvus Cryptocephalus frontalis Cryptocephalus (Linnaeus) (Fabricius) (Illiger) (Illiger) (Forster) (Scopoli) (Müller, O.F.) O.F.) (Müller, (Duftschmid) (Sturm) (Fabricius) (Linnaeus) (Goeze) (Linnaeus) (Linnaeus) Fabricius (Linnaeus) Marsham Müller, O.F. Müller, Suffrian Fabricius (Gravenhorst) (Linnaeus) (Linnaeus) Harold (Scopoli) (Linnaeus) (Linnaeus) Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ●2001 ● ● ● ● ● ● ●(1986) ● ● ● ● ? Vc23 1939-2012 ●1998 ●2002 ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare on common Rareon toadflax Linaria vulgaris. damp labiates Widespread areasin on cow Widespreadparsley on S Anthriscus. Localon Localon damp areas] in Rarelabiates [hemp-nettle On plantains nigraand Ballota black horehound ?Rare[On bedstraws Local Galiumon spp. bedstraws Local on and birch Widespreadon areas heath on birch on V.local downs. hazel on and young birch Rare on Worts St.John’s Localon oak. and birch Localon yellowvarious on compositesadults downland, on V.local sorrel sheep’s Localon hawthorn. On treesvarious on downs the on and birch?, on heathland on V.local ) kidney on vetch Anthyllis downland vulneraria on V.Rare ) yellowvarious on compositesadults downland, on V.local EXTINCT?Wychwood (JWL) NNR only Ukenigmatic 1954 site for this lastseen in spp. Rare, on wet heathland on on wet Rare, on heathland Hypericum & aquatica Mentha Galium Galium esp. on sandy chalky and on esp. soils. Rumexacetosella Salix Hypericum Erica tetralix Erica spp. spp. hybrids in wetlands, increasingly occurring in gardens on cultivated mints.cultivated wetlands, increasingly gardenshybrids in on in occurring . spp. esp. on sandy chalky and on esp. soils. spp. on sandy heathland. Also on Also sandyon heathland. on Oxford dist (JJW)Oxford dist

ecology Lotus corniculatus, corniculatus, Lotus esp. on the chalk the on esp. 4248 4247 4246 4245 4244 4243 4242 4241 4240 4238 4237 4235 4234 4233 4232 4230 4229 4228 4227 4225 4224 4222 4221 4220 4219 4218 4217 4216 Phaedon armoraciae Phaedon viridula Gastrophysa polygoni Gastrophysa violacea Chrysolina varians Chrysolina staphylaea Chrysolina Galerucella sagittariae Galerucella pusilla Galerucella nymphaeae Galerucella lineola Galerucella calmariensis Galerucella GALERUCINI GALERUCINAE vulgatissima Phratora vitellinae Phratora laticollis Phratora viminalis Gonioctena olivacea Gonioctena decemnotata Gonioctena tremula Chrysomela populi Chrysomela versicolora Plagiodera phellandrii Prasocuris junci Prasocuris marginella Hydrothassa glabra Hydrothassa tumidulus Phaedon cochleariae Phaedon (Brahm) Suffrian (Linnaeus) Linnaeus (Duftschmid) (Schaller) (Fabricius) (Germar) (Herbst) (Müller, O.F.) O.F.) (Müller, (Forster) Fabricius (De Geer) (Linnaeus) (Fabricius) (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) (Linnaeus) (Laicharting) (Gyllenhal) (Linnaeus) (Linnaeus) (Marsham) (Linnaeus) Med Rom Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ●1998 ● ● ● ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ●1906 ● ● ● ● ● ● ● ● ● ● ●1987 ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread on various wetland plants wetlandvarious plants Widespreadon precise locality no Ox dist waterlillies Localon esp. sallows Local on loosestrife purple Widespreadon Localon Localon Localon sallowother Localand on broom Rareon commons, and heaths on aspen rareon ( low growing on V.rare hogweed Widespread on Localon brooklime Local on docks increasing on Localbut knotgrass Local on ground-ivy Rareon Rareon Rare?buttercups? On RDB1 ) Extinct in UK? in Extinct ) Hypericum Salix Populus Populus officinale Nasturtium and and Salix esp. aspen aspen esp. aspen esp. Polygonum Populus Glechoma hederacea hederacea Glechoma Veronica beccabungae Veronica esp. on sandy chalky and on esp. soils. . Populus tremula tremula Populus Nymphaea alba alba Nymphaea Heracleum sphondylium Salix P.tremulae P.tremulae Rumex, and and in disturbed open areas. open disturbed in esp. on damp heathy on esp. areas. Lythrum salicaria Lythrum Rorippa Rumexhydrolapathi and and and and Persicaria, Persicaria, in ponds and streams and ponds in in wetlands including streams wetlands including in Salix repens repens Salix

atriver, stream canal margins. and ecology esp. on damp heaths and rides. damp and heaths on esp. 4280 4279 4278 4277 4276 4275 4274 4273 4272 4271 4270 4269 4268 4266 4265 4263 4262 4261 4260 4258 4257 4256 4255 4254 4253 4251 4250 Galeruca yarrow form yarrow Galeruca tanaceti Galeruca (= laticollis Galeruca viburni Pyrrhalta tenella Galerucella Phyllotreta striolata Phyllotreta (= punctulata Phyllotreta ochripes Phyllotreta nodicornis Phyllotreta nigripes Phyllotreta nemorum Phyllotreta flexuosa Phyllotreta exclamationis Phyllotreta diademata Phyllotreta cruciferae Phyllotreta consobrina Phyllotreta atra Phyllotreta ALTICINI halensis Sermylassa circumfusus Calomicrus longicornis Luperus flavipes Luperus quadrimaculata Phyllobrotica suturalis Lochmaea crataegi Lochmaea caprea Lochmaea aerea interrupta ) ) ) ) (Fabricius) (Linnaeus) (Paykull) (Linnaeus) (Linnaeus) (Sahlberg, C.R.) (Linnaeus) (Forster) (Fabricius) (Fabricius) (Thomson, C.G.) (Curtis) (Illiger) (Illiger) (Linnaeus) (Fabricius) (Linnaeus) (Goeze) Foudras (Marsham) (Marsham) (Curtis) (Marsham) (Thunberg) (Linnaeus) Rom Rom Subfossil ●? ● ● ● ● ● ● ●? ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ●2000 ● ● ● ● ● ● ● ● ● ●1995 ● ● ● ● ● Vc23 1939-2012 ● ● ● ●1991 ● ● ●2002 ●1906 ● ● ● ● ● ● ## VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Oxdist Armoracia horse-radish and rusticana Raphanus, radish Rareon crucifers Localon wild Scarcemignonette on crucifers Localweld on and crucifers on Frequent Ox dist crucifersvarious ?On cuckooflower Rareon Cardamine pratense crucifersvarious on local crucifersvarious Localon crucifersvarious on Frequent etc [oak] birch, aspen, Localon skullcap on V.local hawthorn Widespreadon sallowsother Widespread and on ‘breck dryform/sp.?’the yarrow and soils damp rides, on disturbed grassland in and on cabious devil’s-bit Localon Meadow On rue Extinct wayfaring Localon tree meadowsweet Localon Scutellaria galericulata galericulata Scutellaria Filipendula ulmaria Filipendula Viburnum lantana lantana Viburnum Reseda luteola Reseda Reseda lutea Reseda Salix on the chalk the on esp. on the chalk, and guelder-rose chalk, and the on esp. esp in damp meadowsin esp in damp wetlands. grasslandin and

ecology V.opulus. 4313 4312 4310 4309 4308 4307 4306 4305 4304 4303 4302 4301 4300 4299 4298 4296 4292 4291 4290 4289 4288 4287 4286 4285 4284 4283 4282 4281 (= ?atratula Aphthona vittula Phyllotreta undulata Phyllotreta tetrastigma Phyllotreta (= Longitarsus gracilis Longitarsus ganglbaueri Longitarsus flavicornis Longitarsus ferrugineus Longitarsus exoletus Longitarsus dorsalis Longitarsus curtus Longitarsus brunneus Longitarsus ballotae Longitarsus atricillus Longitarsus anchusae Longitarsus agilis Longitarsus aeruginosus Longitarsus (= aeneicollis Longitarsus nonstriata Aphthona nigriceps Aphthona melancholica Aphthona lutescens Aphthona herbigrada Aphthona euphorbiae Aphthona atrocaerulea Aphthona suturalis atrovirens vittata ) ) ) ) (Redtenbacher) (Rye) (Allard) (Gyllenhal) (Redtenbacher) Allard Kutschera (Fabricius) (Marsham) Kutschera (Linnaeus) (Goeze) (Linnaeus) (Curtis) (Schrank) (Duftschmid) (Paykull) (Comolli) (Stephens) (Faldermann) (Stephens) Weise (Foudras) Heikertinger (Foudras) Med Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ●1998 ● ●1996 ● ● ● ● ● ●2000 Vc23 1939-2012 ● ● ● ● ● ● ● ● ● ●1996 ● ● ●1999 ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon ? On ragwort ?On Ragwort] ? [On ragworts on equent water On mint hound’s-tongue and viper’s bugloss Localon layer woodlands. ragwort,, Local,ground winteringin increasing also on in Boraginaceae. Rareon meadow-rue ?[On Thalictrum black horehound Rare on legumes Widespreadon esp. comfrey? Localon figworts ?On yellow Widespreadon flag Geranium esp. cranesbill ?On Extinct ) pratense spurges especially ?On spurge wood loosestrife purple ?On withdowns rock-rose. the Localon evergreens wintering frequent, on Widespreadand spurges Euphorbia On marjoram Rareon crucifers ?Local on crucifers Widespreadon on Cardamine Cardamine Scrophularia Scrophularia Mentha aquatica Mentha and and Origanum vulgare Origanum Senecio Senecio Nasturtium Ballota nigra Ballota Iris pseudacorus Trifolium spp. sp. ] Euphorbia amygdaloides Euphorbia primarily Downs the on spp.

ecology 4348 4347 4346 4345 4342 4341 4340 4339 4338 4337 4336 4335 4334 4333 4332 4330 4329 4328 4327 4323 4322 4321 4320 4319 4316 4315 4314 Longitarsus melanocephalus Longitarsus lycopi Longitarsus luridus Longitarsus kutscherae Longitarsus jacobaeae Longitarsus holsaticus Longitarsus Altica oleracea Altica lythri Altica (= pusilla) helianthemi Altica ericeti Altica carinthiaca Altica brevicollis Altica tabidus Longitarsus suturellus Longitarsus succineus Longitarsus rutilus Longitarsus rubiginosus Longitarsus reichei Longitarsus quadriguttatus Longitarsus (= pratensis Longitarsus pellucidus Longitarsus parvulus Longitarsus ochroleucus Longitarsus obliteratus Longitarsus nasturtii Longitarsus membranaceus Longitarsus bearei ) ) Aubé (Allard) (Linnaeus) Foudras Weise (Foudras) (Allard) (Illiger) (Illiger) (Allard) (Scopoli) (Fabricius) (Fabricius) (Paykull) (Panzer) (Foudras) (Duftschmid) (Waterhouse, G.R.) (Linnaeus) (Foudras) (Rosenhauer) (Rye) (Foudras) (Marsham) (Foudras) (Pontoppidan) (De Geer)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ●2002 ● ● ● ● ●1999 ● ● ● ●1994 ●1994 ● ● ● Vc23 1939-2012 ● ● ● ● ● ●1906 ● ● ● ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon ? on heathers heathers ?on loosestrife willowherbspurple and on Frequent saladburnet rock-rose and on downs the Localon heather on Recentlywidespread probably on Britain, speciesrecognised in hazel] ?[On mulleinsgreat esp. Rare mulleinon Verbascum thapsus. ragworts on Frequent bindweed Widespreadon Plantains Widespreadon flax On ragwort] ?[On thyme, chalk on the Localmarjoram. on and comfrey Rareon viper’s bugloss an sage wood Local on plantains Localon gipsywort Rareon compositesvarious on frequent Widespreadand plantains. ?On Pedicularis] louseworts ?[On Widespread on a variety of herbs esp. yarrow and hemp agrimony Eupatorium cannabinum Eupatorium agrimony yarrow hemp and esp. herbs varietyof a on Widespread On ragworts Linum. Linum. Calluna, Erica Calluna, Oncerare nowvery countyacrossfrequent the Lycopus europaeus Ox dist HSD wher>?? HSD Ox dist europaeus Lycopus Teucrium scorodonia scorodonia Teucrium Calystegia, riddling the leaves the withsmall riddling E.Hoes

ecology Lathyris 4386 4385 4384 4383 4379 4378 4377 4376 4375 4374 4372 4370 4367 4366 4365 4364 4363 4362 4361 4360 4359 4358 4357 4355 4352 4351 4350 4349 Neocrepidodera transversa Neocrepidodera ferruginea Neocrepidodera rubi Batophila aerata Batophila mercurialis Hermaeophaga palustris Altica Apteropeda orbiculata Apteropeda globosa Apteropeda testaceum Sphaeroderma rubidum Sphaeroderma (= picipes Chaetocnema hortensis Chaetocnema confusa Chaetocnema concinna Chaetocnema arida Chaetocnema rustica Mantura matthewsii Mantura fuscicornis Podagrica pubescens Epitrix atropae Epitrix plutus Crepidodera nitidula Crepidodera fulvicornis Crepidodera aurea Crepidodera aurata Crepidodera modeeri Hippuriphila Derocrepisrufipes concinna in part) part) in (Paykull) Weise Foudras (Linnaeus) (Marsham) (Koch, J.D.W.) (Linnaeus) (Fourcroy) (Latreille) (Marsham) Foudras (Illiger) (Illiger) (Curtis) (Linnaeus) Stephens (Linnaeus) (Linnaeus) (Marsham) (Boheman) (Fourcroy) (Marsham) (Fabricius) (Graëlls) (Fabricius) (Marsham) (Scopoli) (Fabricius) Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●1982 ● ● ● ● ● ●1972 ● ● ●2002 ●2002 ● ● ● ● ● ● ● ● ● ●1991 ● ● Vc23 1939-2012 ● ●1998 ● ● ● ● ●1970 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●1906 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread in woods and scrubby places and woods Widespreadin Rubus? Rareon thistles Widespreadon thistles Widespreadon grasslands open Widespreadin Rare.Grassy rides. sorrel sheep’s on .local grasslands.Ox Rare dist in rock-rose Rareon bittersweet. Widespread on chalky disturbed sites on deadlyAtropa bell-donna nightshade on V.local Localon ( Widespreadon Widespreadon Widespreadon horsetails Localon Vetches ?On thistles Widespreadon thistles Widespreadon Widespreadon Rareon mercurydog’s hedgebanks. on and woods in oftenabundant Widespreadand loosestrife willowherbspurple Local and on on musk on mallow Nb ) Rare on aspen on damp heath and commons. dampand heath on aspen Rare) on Rubus Salix Salix Vicia sp. and Salix Salix Salix Salix Rubus Malva moschata. moschata. Malva Helianthemum nummularium Helianthemum Equisetum Equisetum Populus and and and sp. Rumexacetosella Populus Populus Populus Populus esp. in damp grasslandin esp. in dry in grasslands. acid

ecology 4429 4428 4427 4423 4421 4420 4419 4418 4416 4415 4414 4412 4411 4410 4407 4406 4404 4403 4401 4399 4398 4397 4396 4395 4393 4392 4390 Psylliodes hyoscyami Psylliodes dulcamarae Psylliodes cuprea Psylliodes chrysocephala Psylliodes chalcomera Psylliodes affinis Psylliodes Anthribus fasciatus Anthribus ANTHRIBINAE ANTHRIBIDAE attelaboides Cimberis NEMONYCHIDAE CURCULIONOIDEA vittata Cassida viridis Cassida vibex Cassida rubiginosa Cassida prasina Cassida nobilis Cassida murraea Cassida hemisphaerica Cassida flaveola Cassida fastuosa Pilemostoma CASSIDINAE picina Psylliodes napi Psylliodes luteola Psylliodes (= laticollis Psylliodes weberi ) ) Linnaeus Linnaeus de Villers Linnaeus (Fabricius) Illiger Illiger Thunberg (Marsham) (Paykull) Linnaeus (Koch, J.D.W.) (Müller, O.F.) O.F.) (Müller, Kutschera Müller, O.F. Müller, (Forster) (Linnaeus) (Fabricius) (Schaller) (Illiger) (Illiger) (Koch, J.D.W.) Herbst (Linnaeus)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ●1992 ● ● Vc23 1939-2012 ●2003 ● ● ● ● ● ● ● ●1922 ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Oxdist water Localon mint knapweeds and thistles Widespreadon knapweeds and thistles Widespreadon verges,sites. ruderal road and yarrow Localgrassland including on open in sandy soils disturbed spurreys Rareon campion Rareon unknown? oftendamp plant(s) places.Host open in V.local, spikenard ploughman’s Rareon loosestrife. purple ?On crucifersvarious Localon Alsoautumn) trees comes in light. on cereal also to Widespreadcrops, on ?On AssociatedExtinct? with henbane bittersweet Widespreadon crucifersvarious on crucifersvarious on thistles Localon bittersweet Widespreadon Rorippa Silene Carduus Mentha aquatica Mentha . and possibly and Solanum dulcamarae Solanum Inula conyza Inula Hyoscyamus niger. Last recorded in 1930 in recorded Last Hyoscyamus niger. Cirsium Cirsium on downland downland on sp

. ecology 4465 4464 4463 4462 4461 4458 4456 4455 4454 4453 4450 4449 4448 4447 4446 4445 4444 4442 4441 4440 4439 4438 4436 4435 4432 4431 4430 URODONTINAE sheppardi Choragus CHORAGINAE albinus Platystomos resinosus Platyrrhinus nebulosus Anthribus Apoderus coryli Apoderus nitens Attelabus ATTELABIDAE betulae Deporaus Byctiscus betulae tomentosus Temnocerus nanus Temnocerus longiceps Temnocerus pauxillus Neocoenorrhinus interpunctatus Neocoenorrhinus germanicus Neocoenorrhinus aequatus Neocoenorrhinus aeneovirens Neocoenorrhinus olivaceus Lasiorhynchites cavifrons Lasiorhynchites caeruleus Involvulus RHYNCHITIDAE rufipes Bruchela (Linnaeus) (Scopoli) (Olivier) B4500 (Linnaeus) (Linnaeus) (Paykull) (Linnaeus) Kirby Kirby (Forster) (De Geer) (Thomson, C.G.) (Scopoli) (Gyllenhal) (Gyllenhal) (Gyllenhal) (Linnaeus) (Germar) (Herbst) (Marsham) (Stephens) Subfossil ● ● ● ● ● ● ? ● ● ● ●? ● ● ● ● ? Vc22 VCH ●1985 ● ●1980 ●1990 ●1993 ●2007 ● ● ●1980 ● Vc23 1939-2012 ●2004 ●2004 ●2004 ●1998 ●1980s ●1990 ● ●1996 ●1998 ●1998 ●200 ●2002 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local on hazel Localon oak foliage on especially oaks.V.local young ‘scrub’ on on Frequent hazel. and birch On Rare, declining? Localon birch Localon birch, on V.local wide Rosaceae,On herbaceus range ofand esp. shrubby oak. Rare on wide Rosaceae.herbaceus Local,range on and of shrubby, Rosaceae other on also hawthorn, on Frequent oak Localon on V.local trees rosaceuos esp. on V.local ivy dead old Rareon etc. beech alder, hardwoods, Localon Salix Quercus Betula , and and also also Salix less common on Populus Populus in one-yeartwigs. in old Salix Helixhederae and and and and Pyrus. 1993 Pyrus. esp. Alnus Cratageus, Prunus Cratageus, P. tremulae P. . Alnus, Corylus, Alnus, Malus, Prunus. Malus, and and

etc.. ecology Malus. Malus. Cratageus, Prunus Cratageus, ] 4511 4510 4509 4508 4506 4504 4502 4498 4495 4494 4493 4491 4488 4486 4485 4483 4480 4479 4478 4477 4476 4473 4472 4470 4468 4467 4466 Apion rubiginosum Apion rubens Apion haematodes Apion frumentarium Apion cruentatum Apion APIONIDAE Squamapion flavimanum Squamapion cineraceum Squamapion atomarium Squamapion minimum Melanapion pallipes Kalcapion ulicis Exapion fuscirostre Exapion hookerorum Omphalapion stolidum Diplapion confluens Diplapion onopordi Ceratapion gibbirostre Ceratapion carduorum Ceratapion brunnipes Acentrotypus aeneum Aspidapion radiolus Aspidapion brevirostre Pseudoperapion violaceum Perapion marchicum Perapion hydrolapathi Perapion curtirostre Perapion affine Perapion Stephens (Forster) (Kirby) (Kirby) Walton (Kirby) (Kirby) Kirby Kirby (Germar) Grill (Fabricius) (Fabricius) (Kirby) (Kirby) (Marsham) (Germar) (Kirby) (Kirby) (Kirby) (Kirby) (Linnaeus) (Herbst) (Gyllenhal) (Herbst) (Marsham) (Kirby) (Kirby) (Boheman) (Kirby) (Kirby) (Wencker) (Gyllenhal) (Kirby) (Herbst)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● ● ●1998 ● ● ● ●1994 Vc23 1939-2012 ●1980s ● ● ● ● ● ● ●2002 ● ● ● ● ● ●2012 ●2004 VC22 ● ● ● ● ● ● ●2003 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon ? [On Sallows in sawfly Sallows in ? [On galls madeby mercurydog’s Localon mayweeds Localon Tripleurospermumspp ox-eye on daisy V.local mayweeds ?[On chamomile] and thistles ?Rareon mallows Locallyon abundant Localon Localon Localon sorrels and docks various on Frequent sorrel Rareon sandy Rumex soils. Rare on disturbed acetosella on sandy Rumex Localsoils. on disturbed acetosella on Localon leaved docks broad Localon Rumex grasslands.on unimproved acetosa in V.local esp. Rumex Rumexacetosella Rumex Rumexacetosella Rumexacetosa incl. incl. R. acetosa R. acetosa R. Mercurialis perennis, Mercurialis perennis, on disturbed sandy soils disturbed on sandy soils disturbed on Rumex Malva spp. Malva . in grassland on verges grassland on in etc. Pontania in woodlands in spp.] spp.] ecology 4546 4545 4543 4542 4540 4539 4538 4537 4535 4534 4533 4532 4531 4530 4529 4528 4527 4526 4525 4524 4523 4522 4517 4516 4515 4514 4513 4512 Cyanapion spencii Cyanapion longirostre Rhopalapion rufirostre Pseudapion malvae Malvapion urticarium Taeniapion vicinum Squamapion Stenopterapion tenue Stenopterapion meliloti Stenopterapion atratulum Protopirapion virens Ischnopterapion modestum Ischnopterapion loti Ischnopterapion seniculus Catapion pubescens Catapion immune Pirapion subulatum Oxystoma pomonae Oxystoma craccae Oxystoma cerdo Oxystoma aethiops Holotrichapion pisi Holotrichapion ononis Holotrichapion waltoni Eutrichapion reflexum Eutrichapion punctigerum Eutrichapion vorax Eutrichapion viciae Eutrichapion ervi Eutrichapion (Gerstäcker) (Kirby) (Kirby) (Linnaeus) (Kirby) (Kirby) (Kirby) (Kirby) (Fabricius) (Kirby) (Kirby) (Fabricius) (Herbst) (Paykull) (Fabricius) (Kirby) (Kirby) (Kirby) (Kirby) (Kirby) (Kirby) (Kirby) (Kirby) (Stephens) (Kirby) (Kirby) (Fabricius) (Kirby) (Kirby) (Herbst) (Gyllenhal) (Herbst) (Kirby) (Kirby) (Herbst) (Olivier) (Germar) (Paykull) (Germar) Rom Rom Rom Subfossil ● ● ● ● ● ? ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●2001 ● ●2001 ● ● ●1953 ● ● ● ● ● ●1999 ●2004 ● ● ● ●2004 ●2002 Vc23 1939-2012 ●1907 ● ● ● ● ● ● ● ● ● ● ● ● ● ●2004 ●1998 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local on medicks Localon Localon broom Localon a wide Localvariety on of clovers grassy Trifoliumopen in places. Widespread,on Trifolium Medicago[and hybridium Rare] on broom Localon meadow widespread on but vetchling vetchesvarious Local on vetchesvarious Local on vetches Local esp. on ?[On Medicks Widespreadon restharrows on V.local vetch horseshoe Rareon Rare? On vetchesvarious on V.local vetchesvarious on V.local vetchesvarious on V.local Vicia Vicia Melilotus. Melilotus. Onobrychis vicifolia Onobrychis Lotus corniculatus Lotus Cytisus Cytisus Medicago V.cracca Ononis repens Ononis esp. on sandy soils on esp. sandy soils. on esp. Medicago Hippocrepis comosa Hippocrepis Vicia Vicia Vicia esp. on calcareous grassland. on and lupulina Lathyrus and Lathyrus pratensis Lathyrus O. spinosa, O. on the chalk. the on

ecology esp. on the chalk. the on esp. 4588 4586 4582 4580 4579 4578 4576 4575 4574 4570 4569 4567 4566 4565 4564 4563 4562 4560 4559 4558 4555 4554 4553 4552 4551 4550 4549 4548 Protapion dissimile Protapion difforme Protapion assimile Protapion apricans Protapion simile Betulapion ebeninum Synapion Thryogenes nereisThryogenes festucae Thryogenes scirpi Notaris acridulus Notaris equiseti Grypus ERIRHINIDAE oryzae Sitophilus granarius Sitophilus RHYNCHOPHORINAE marmoratus Nanophyes NANOPHYIDAE astragali Pseudoprotapion varipes Protapion trifolii Protapion schoenherri Protapion (= ononidis Protapion nigritarse Protapion fulvipes Protapion filirostre Protapion ononicola (Fabricius) ) (Fabricius) (Linnaeus) (Linnaeus) (Linnaeus) (Kirby) (Kirby) (Germar) (Paykull) (Geoffroy in Fourcroy) (Kirby) (Kirby) (Kirby) (Kirby) (Herbst) (Gyllenhal) (Germar) (Germar) (Kirby) (Kirby) (Kirby) (Kirby) (Herbst) (Linnaeus) (Boheman) (Goeze) (Paykull) E.Ho E.Ho Med Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ●2004 ●2004 ●1970s ● ●1942 ● ● ● ● ● ●2001 ● Vc23 1939-2012 ● ● ● ●1998 ●2000 ●2012 ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local on spike-rush Localon sedges? wetlandsLocal on in large pond-sedge on grasses wetlands on in esp. damp (arvenseplacesEquisetum palustre) horsetail and on V.local grainoccasionally outdoors. in found Syananthropic occasionallygrain,outdoors. in found Syananthropic 1942 clovers] ?[On white cloverand red on Frequent cloversvarious on Frequent clovers esp. on oftenabundant Widespreadand black medick ?[On hare’s-footclover on V.local ?Usually on found various Localon clover red on Frequent Betula. on commonheaths the Local but on trefoil uliginosus. Lotus large Rare,bird’s-foot on Trifolium Trifolium Polygonum Polygonum Eleocharis palustris Eleocharis Medicago lupulina Medicago Carexacutiformis Glyceria maxima spp. spp. T.arvense T.arvense spp.

] Typha on sandy soils. on Trifolium repens Trifolium ecology 4630 4628 4627 4626 4622 4620 4619 4617 4614 4613 4612 4611 4610 4609 4607 4606 4605 4604 4602 4601 4600 4599 4598 4592 4591 4590 CURCULIONINAE CURCULIONIDAE rufinasus Stenopelmus bimaculatus Tournotaris scirrhosus Thryogenes Cionus nigritarsis Cionus hortulanus Cionus alauda Cionus CIONINI ulmi Anthonomus rubi Anthonomus pomorum Anthonomus pedicularius Anthonomus chevrolati Anthonomus brunnipennis Anthonomus bituberculatus Anthonomus ANTHONOMINI carpini Acalyptus ACALYPTINI venosus Curculio rubidus Curculio nucum Curculio glandium Curculio salicivorus Archarius pyrrhoceras Archarius CURCULIONINI (Herbst) Linnaeus (Gyllenhal) (Gravenhorst) (Fabricius) (Herbst) Reitter (De Geer) (Fourcroy) Marsham (Paykull) (Marsham) (Gyllenhal) Gyllenhal (Linnaeus) Desbrochers (Fabricius) (Linnaeus) (Curtis) Thomson, C.G. E.Ho E.Ho E.Ho E.Ho Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ●1970s ●2002 ● ● ● ● ●1986 ● ● ● ● ●2004 Vc23 1939-2012 ● ● ● ● ● ●1970s ●1970s ● ● ●2004 ● ● ● ● ●1948 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare on dark mullein Rare on figworts mulleins and Buddleja] Widespreadon [occasionallyon figworts Widespreadon elm Rareon Rosaceae Widespreadon Apple Localon hawthorn Widespreadon possibly and hawthorn On erecta. tormentilPotentilla Rareon hawthorn Rareon Rareon oak Localon birch. Rareon with greycompetition to ?? paucitydue squirrel to of nuts, due hazel.Declining? Possibly Localon oak Widespreadon Widespreadon Cynipidae). galls madecurrant in byoak,developing wasp Widespreadthe on water Veryon local fern large wetland grasses [On sedges] and Salix in wetlands. in Salix, Salix, Malus Verbascum nigrum Verbascum galls madein developing by the sp. Scrophularia nodosa Scrophularia Azolla, Azolla, Sorbus but usually present on good colonies of the host plant. plant. host of the colonies good usually on present but ] on the chalk. the on and and S .

auriculata ecology Pontania . Also on Mulleins including including Mulleins Also . on saw-flies. Neuroterus quercusbaccarum quercusbaccarum Neuroterus V.blattaria (Hym., 4669 4668 4667 4666 4665 4660 4659 4657 4655 4652 4651 4650 4649 4648 4645 4644 4642 4641 4640 4639 4637 4635 4634 4633 4632 4631 Ellescus bipunctatus ELLESCINI pulchellus Cleopus tuberculosus Cionus scrophulariae Cionus Rhinusa linariae Rhinusa collina Rhinusa antirrhini Rhinusa campanulae Miarus pyraster Mecinus pascuorum Mecinus beccabungae Gymnetron villosulum Gymnetron veronicae Gymnetron melanarium Gymnetron plantarum Cleopomiarus graminis Cleopomiarus MECININI tremulae Dorytomus tortrix Dorytomus taeniatus Dorytomus rufatus Dorytomus melanophthalmus Dorytomus longimanus Dorytomus (= ictor Dorytomus hirtipennis Dorytomus dejeani Dorytomus validirostris (Gyllenhal) (Herbst) ) ) (Panzer) (Herbst) (Linnaeus) (Bedel) Faust (Paykull) (Herbst) (Linnaeus) (Gyllenhal) (Scopoli) (Fabricius) (Linnaeus) (Fabricius) (Germar) (Linnaeus) Gyllenhal Bedel (Forster) (Gyllenhal) (Germar) (Linnaeus) (Germar) (Paykull)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ●2002 ●1995 ● ● ● ● ●2001 ● ●1996 ●2004 ● ● Vc23 1939-2012 ● ●1970s ● ● ● ● ●1970s ● ● ● ● ● ● ●1963 ●2001 ● ●1996 ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon rare on common rareon toadflax common on toadflax Rare on breeds dry areas. open in plantains Localon plantains Widespreadon water Local hybrid]. on speedwells [and beccabungae. brooklime Veronica Rare on germander Rareon chamaedrys.speedwell Veronica Campanula] [On Rareon aspen Rareon Widespreadon Localon Rare?on poplars Localon blackhybrid esp. poplars. poplars Rareon willows On wet in woodsKings Weir 1909 aspen. Localon willows ?On damp areas in heaths. esp. figworts Localon mulleins. and figworts mulleins and Buddleja] Widespreadon [occasionallyon V.blattaria figworts Widespreadon Campanula Campanula Salix Salix. spp. Campanula 1990 ?? 1990 Salix Populus. Linaria vulgaris Linaria on the downs the on Scrophularia nodosa Scrophularia Plantago Linaria vulgaris MFVCorley vulgaris Linaria esp. esp. C. glomerata C. spp. and and ] on the downs, the on ] S .

auriculata ecology

[harebell [harebell . Also on Mulleins including including Mulleins Also . on C. rotundifolia C. ], Adults also atyellow Adults also ], composites 4722 4720 4718 4716 4708 4706 4705 4704 4701 4699 4697 4696 4695 4690 4689 4688 4687 4686 4685 4683 4682 4681 4680 4679 4676 4673 4672 4671 Orchestes quercus Orchestes pilosus Orchestes alni sequensi Isochnus foliorum Isochnus RHAMPHINI Tychius pusillus Tychius picirostris Tychius meliloti Tychius junceus Tychius primita Sibinia TYCHIINI pillumus Pseudostyphlus setiger Orthochaetes STOREINI jungermanniae Smicronyx SMICRONYCHINI Tachyerges stigma Tachyerges salicis pulicarius Rhamphus oxyacanthae Rhamphus Pseudorchestes pratensis Orchestes fagi (= Orchestes testaceus (= Orchestes signifer Orchestes rusci alni avellanae ) ) (Linnaeus) (Linnaeus) (Herbst) (Herbst) Stephens (Reich) Germar (Fabricius) (Creutzer) (Linnaeus) (Linnaeus) (Stierlin) (Fabricius) (Müller, O.F.) O.F.) (Müller, (Germar) (Müller, O.F.) O.F.) (Müller, (Beck) (Herbst) (Marsham) (Gyllenhal) (Germar) (Reich) E.Hol E.Hol E.Hol E.Hol Med Subfossil ● ● ● ● ● ● ● ● ●?? ● ● ●? ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ●1998 ● ● ● ●2002 ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local on disturbed groun on small on groun disturbed Localon clovers on frequent Widespreadand melilots Localon smallvarious Localon legumes sandy grounds. disturbed spurreys Localon on etc.esp. mayweedscented [On Extinct? withbluebells] woodlands in also [but grasslanddownland, esp. open litter/mossLocal in on etc.Lotus grasslandsricher on and downs on and Calluna, on Cuscuta dodder on V.local willows Localalder on and Localon Rosaceae other and hawthorn Widespreadon common Rareon knapweed C crevices and trees.Holes wintering beech, in in on frequent Widespreadand alder] [On alder] [On Salix entaurea Matricaria recutita Matricaria Trifolium

nigra. nigra. sp

] ecology 4778 4777 4776 4775 4774 4772 4771 4770 4769 4768 4767 4766 4765 4763 4762 4761 4757 4756 4755 4754 4747 4745 4744 4733 4729 4728 4727 4725 Bagous tempestivus Bagous collignensis Bagous BAGOINAE tibialis Tychius stephensi Tychius squamulatus Tychius Ceutorhynchus contractus Ceutorhynchus constrictus Ceutorhynchus cochleariae Ceutorhynchus (= chalybaeus Ceutorhynchus atomus Ceutorhynchus (= assimilis Ceutorhynchus alliariae Ceutorhynchus terminatus Calosirus melanarius Amalorrhynchus CEUTORHYNCHINI CEUTORHYNCHINAE [ (= dolorosa Limnobaris picicornis Aulacobaris lepidii Aulacobaris BARIDINAE alismatis Bagous lutulentus Bagous glabrirostris Bagous Limnobaris t-album Limnobaris timidus pleurostigma pilistriata ) ) Boheman (Marsham) ) Gyllenhal (Gyllenhal) (Germar) (Herbst) (Herbst) (Herbst) (Goeze) (Linnaeus)] (Herbst) Gyllenhal (Marsham) Boheman Brisout, H. (Paykull) (Marsham) (Marsham) Germar (Gyllenhal) (Stephens) E.Hol E.Hol Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ●1995 ●2002 ●2004 ● ● ●1997 Vc23 1939-2012 ● ● ● ● ● ●2004 ●1984 ●1994 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon ? On hedgemustard ?On garlic Rare mustard on Thale On Cress garlic Localvergeshedgebanks,mustard on Alliaria on etc petiolata wild carota Daucus carrot Rareon water-cress on sedges Local Carexwetlandson in sedges Local Carexwetlandson in wildchalky mignonette RareResedaon on lutea soils. crucifersvarious wetland Rareareas in on esp. water on Plantain hornworts On repens Ranunculus creeping buttercup Rareon water-milfoil Rare,on Trifolium?] on soils ?[Sandy Rareon possiblyKidney trefoil,and Vetch bird’s-foot on downs the Rare) on Trifoliums. Trifoliums. Ceratophyllum Nasturtium Alisma plantago-aquatica Alisma Sisymbrium officinale Sisymbrium Myriophyllum Alliaria petiolata Alliaria spp. esp. on plants growing plants over flowing on esp. water spp. sp.] sp.] sp. sp.

ecology 4823 4822 4821 4820 4819 4818 4816 4814 4811 4809 4808 4807 4806 4805 4802 4801 4800 4799 4798 4796 4795 4793 4790 4786 4785 4784 4783 4781 4779 (= pallidactylus Ceutorhynchus (= obstrictus Ceutorhynchus hirtulus Ceutorhynchus erysimi Ceutorhynchus Hadroplontus litura Hadroplontus punctiger Glocianus pilosellus Glocianus moelleri Glocianus (= distinctus Glocianus nasturtii Drupenatus melanostictus Datonychus rubicundus Coeliodinus (= transversealbofasciatus Coeliodes ruber Coeliodes (= rana Coeliodes (= typhae Ceutorhynchus turbatus Ceutorhynchus thomsoni Ceutorhynchus syrites Ceutorhynchus resedae Ceutorhynchus rapae Ceutorhynchus pyrrhorhynchus Ceutorhynchus picitarsis Ceutorhynchus quadridens marginatus erythroleucos dryados floralis assimilis ) ) ) ) (Fabricius) ) ) ) ) (Marsham) ) ) (Thomson, C.G.) (Fabricius) (Gyllenhal) (Gyllenhal) (Brisout, Ch.) (Germar) Gyllenhal Germar (Herbst) (Fabricius) (Marsham) Germar (Herbst) Schultze Gyllenhal Kolbe (Marsham) (Marsham) (Marsham) (Marsham) (Goeze) Subfossil ● ● ● ● ● ● ● ● ● ? ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ●1980 ● ● ● ● ● ● ●1996 Vc23 1939-2012 ● ● ● ● ● ● ●1995 ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local on small Localon yellow grassland open composites in water-cress Localon gypsywort Localon birch Rareon oak Localon oak Rare on oak Localon crucifers. recentlyLocal as Until on Britain known in garlic mustard ?[On Ox1970 pre weld On Reseda luteola] crucifers.various Rareon hedgemustard Localon brassicas] On crucifers Widespreadon Lycopus europaeus, Lycopus Nasturtium spp. Nasturtium Alliaria petiolata Alliaria Sisymbrium officinale Sisymbrium esp. on plants growing plants over flowing on esp. water ] pre 1970 pre ] and water and mint

ecology C.floralis Mentha aquatica Mentha and hybrids near waterhybrids near and 4855 4854 4849 4848 4847 4845 4844 4842 4841 4839 4838 4837 4836 4835 4834 4833 4832 4831 4830 4829 4828 4827 4826 4825 4824 Mogulones asperifoliarum Mogulones triangulum Microplontus rugulosus Microplontus campestris Microplontus Micrelus ericae trimaculatus Hadroplontus Eubrychius velutus Eubrychius PHYTOBIINI scortillum Amalus AMALINI geranii Zacladus exiguus Zacladus troglodytes Trichosirocalus rufulus Trichosirocalus horridus Trichosirocalus barnevillei Trichosirocalus viduatus Thamiocolus (= ruficornis Stenocarus quercicola Sirocalodes mixtus Sirocalodes depressicollis Sirocalodes sisymbrii Poophagus pollinarius Parethelcus Nedyus quadrimaculatus geographicus Mogulones euphorbiae Mogulones umbrinus ) (Gyllenhal) (Paykull) (Olivier) (Herbst) (Mulsant & Rey) (Beck) (Fabricius) (Stephens) (Gyllenhal) (Paykull) (Brisout, Ch.) (Forster) (Dufour) (Herbst) (Gyllenhal) (Panzer) (Linnaeus) (Goeze) (Gyllenhal) (Boheman) (Gyllenhal) (Grenier) (Fabricius) (Fabricius) Rom Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ? ● ● ● ● ● Vc22 VCH ●1953 ●2004 ● ●1965 ●2001 ● ● ●2001 ●2004 ● ●1998 Vc23 1939-2012 ● ●2004 ● ●1999 ●1907 ● ● ●2001 ●1918 ●2001 ●1997 ●1907 ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare, aquatic, submerged on pond-weeds. submergedpond-weeds. Rare, aquatic, on Geraniums on small Rareon flowered plantains on habitats open disturbed Widespreadin plantains on spear thistle on yarrow on marsh Stachyswoundwort ?[On Extinct palustris] poppies Rareon common On fumitory Fumaria Fumaria officinalispossiblyother sp.] and climbing corydalis on fumitorys on water water-cress Localon Common nettles on whereverfrequent Widespreadand nettles viper’s bugloss Localon ground-ivy. on esp. Rare,polyphagous, Boraginaceae Localon esp. yarrow Rare) on chamomile Rareon ox-eye Rareon daisy Erica.spp. on also but Calluna esp. heathers Widespreadon natans] musk On Carduus thistle Achillea millefolium Achillea Plantago Fumaria Cirsium vulgare Cirsium Papaver Achillea. Achillea. Urtica Nasturtium spp. and and spp. Nasturtium Leucanthemum vulgare vulgare Leucanthemum Corydalis clavicula Corydalis spp. on disturbed sandy chalky and disturbed sites. on spp. sp. in opne grassland, and ruderal sites. ruderal grassland, and opne in sp. Geranium spp. spp. Cynoglossum officinale Cynoglossum in open grassy verges. open road in places including and musk and thistle spp. spp. Urtica on wooded former heathland and commons.former wooded and on heathland yellocress spp. grow. spp. Carduus nutans Carduus Plantago , and and ,

Rorippa spp. spp. Rorippa ecology Echium vulgare Echium esp. on plants growing plants over flowing on esp. 4904 4903 4901 4900 4899 4898 4897 4896 4895 4894 4891 4890 4888 4887 4878 4876 4875 4872 4871 4870 4869 4868 4866 4863 4859 4858 4857 Rhinoncus bruchoides Rhinoncus leucogaster Phytobius quadrituberculatus Pelenomus comari Pelenomus canaliculatus Pelenomus quadrinodosus Neophytobius ENTIMINAE ENTIMINAE lunatus Gronops CYCLOMINAE roboris Kyklioacalles ptinoides Acalles (= misellus Acalles lapathi Cryptorhynchus CRYPTORHYNCHINAE lignarius Phloeophagus RHYNCOLINI huttoni Pentarthrum confine Euophryum PENTARTHRINI uncipes Cotaster parallelepipedus Cossonus COSSONINI COSSONINAE perpendicularis Rhinoncus pericarpius Rhinoncus inconspectus Rhinoncus castor Rhinoncus turbatus ) ) Boheman (Fabricius) (Marsham) (Boheman) (Fabricius) (Herbst) (Broun) Wollaston (Curtis) (Marsham) (Herbst) (Linnaeus) (Linnaeus) (Marsham) (Herbst) (Fåhraeus) (Herbst) (Reich) (Gyllenhal) (Fabricius) E.Hol Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ●1995 ●1980 ●2002 ● ● ● ● ● ● ● ●1996 Vc23 1939-2012 ● ● ● ● ●1995 ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare on spurreys Rareon wood. dead oak leaf in in woods and Rare litter in nests ant and wood rotting litter, in heaths on and woodland in V.local hedgerows. small twigsesp. and wood woods and dead branches in Widespreadon willows] [On Sycamore also ash, oak esp. and wood dead Local in possiblyatexpense of declined abovespp. timber. Much structural esp. wood, rotting Rareon timber. Immigrant frequent. Widespread and Naturalised. building and wood from rotting in New found Zealand, ofRare, saproxylic,heartwood hardwoods. in developing on Localon bistort amphibious on sorrel sheep’s Localon redshank Localon Myriophyllum on ponds Rare, aquatic in On palustris. marsh Potentila Rare on cinquefoil on Rare, aquatic Polygonum?] grassland [on Rare in Polygonums Polygonums Rumex. Spp. Rumex. including amphibious bistort bistort amphibious including [in dry grassland] [in Myriophyllum Spergularia. Polygonum persicaria Polygonum Polygonum amphibium Polygonum Rumexacetosella esp. sand spurrey on disturbed sandy soils. disturbed spurrey sand esp. on in ponds. ponds. in in rough grasslandrivers.near esp. rough in P.amphibium

ecology 4960 4957 4956 4954 4952 4950 4949 4946 4944 4943 4942 4938 4936 4934 4932 4926 4922 4921 4920 4918 4917 4916 4913 4912 4911 4910 4907 4906 4905 Neliocarus sus Neliocarus nebulosus Neliocarus faber Neliocarus BRACHYDERINI triguttatus Graptus ALOPHINI Phyllobius argentatus Phyllobius pyri Phyllobius PHYLLOBIINI waltoni Caenopsis PERITELINI sulcatus Otiorhynchus singularis Otiorhynchus rugosostriatus Otiorhynchus raucus Otiorhynchus porcatus Otiorhynchus ovatus Otiorhynchus ligneus Otiorhynchus desertus Otiorhynchus crataegi Otiorhynchus clavipes Otiorhynchus OTIORHYNCHINI obscurus Barynotus moerens Barynotus GEONEMINI plumbeus Attactogenus CNEORHININI melanogrammum Strophosoma capitatum Strophosoma (Stephens) (Linnaeus) (Herbst) (Boheman) (Fabricius) (Fabricius) (Fabricius) (Linnaeus) (Fabricius) (Olivier) (Stephens) (Linnaeus) (Bonsdorff) Germar Rosenhauer (Fabricius) (Herbst) (Marsham) (De Geer) (Linnaeus) (Goeze) (Forster) Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ●1999 ● ● ●2003 ● ● ● ●1998 ● ●2002 ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ●2005 ● ● ●2002 ●1991 ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Widespread and frequent on hedges and in woods on a variety on woods trees in of hedgeshardwood and on frequent Widespreadand a variety on frequent trees Widespreadand ofon herbage and ling, and heather under heathlands dwelling esp.on Rareground largely and A gardenpest. Frequent synanthropic. serious gardenpest Widespread,a serious many gardens in including Polyphagous habitats gardens, Localparks and oftenin gardens including sandy ground disturbed on chalk and sand areas on open disturbed in V.local downland. on esp. ground the on trees and Local, on polyphagous grassy Open places mercurywith esp dog’s woods in oftensandy ground open Rareon commons aheathsand wide variety on on VeryFrequent ofherbs. abundant trees and a variety heathy on in habitats esp. oftrees bushes and heathers rareon a wide heathers variety on including ofherbs trees and Corallian Local on plantains. on sandy ground disturbed on grassland with Plantains. in rare on disturbed chalky and sandy ground sandy and chalky disturbed on rare Mercurialis perennis Mercurialis perennis

ecology esp. esp. 4997 4996 4995 4993 4991 4989 4987 4983 4982 4981 4980 4979 4978 4977 4976 4975 4974 4972 4971 4970 4968 4967 4966 4965 4964 4963 4962 4961 Phyllobius viridicollis Phyllobius roboretanus Phyllobius oblongus Phyllobius pomaceus Phyllobius (= glaucus Phyllobius Andrion regensteinense Andrion SITONINI asperatus Sciaphilus echinatus Brachysomus pellucidus Barypeithes araneiformis Barypeithes SCIAPHILINI cervinus Polydrusus marginatus Polydrusus pterygomalis Polydrusus flavipes Polydrusus confluens Polydrusus mollis Polydrusus (= formosus Polydrusus (= tereticollis Polydrusus lethierryi Pachyrhinus tessulatus Liophloeus POLYDRUSINI virideaeris Phyllobius maculicornis Phyllobius sericeus undatus calcaratus ) ) ) ) ) (Ström) (Scopoli) (De Geer) (Linnaeus) (Linnaeus) (Bonsdorff) (Mayer) (Mayer) Gyllenhal Stephens (Müller, O.F.) O.F.) (Müller, (Laicharting) (Fabricius) (De Geer) (Desbrochers) (Boheman) Stephens Gredler (Herbst) (Bonsdorff) Germar Boheman (Schrank)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ● ● ●1988 ● ●1974 ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ● ● ● ●1986 ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Locallybroom gorse and on frequent many trees hernage in Localand on habitats grassy open dwelling in Rare,places. ground common gardens,and grassy in atbase ofalso in trees. sand, ground on places esp. Widespreadon grassy in ground places Widespreadon a variety on woods trees in of hedgeshardwood and on frequent Widespreadand shrubs trees On and a variety on woods trees in of hedgeshardwood and on frequent Widespreadand a variety Rareon of treesaspen esp. a variety On ofbushes. trees and a variety on woods treesbirch in esp. of hedgeshardwood and on frequent and Ivy Localon trees,clad larvae Umbellifers. in a variety on frequent trees Widespreadand ofon herbage and a variety on frequent trees Widespreadand ofon herbage and ??? nettles on frequent Widespreadand a variety on woods trees in of hedgeshardwood and on frequent Widespreadand nettles on frequent Widespreadand a variety on woods treesalder in esp. of hedgeshardwood and on

ecology 5042 5041 5040 5039 5037 5036 5034 5030 5029 5028 5026 5025 5019 5018 5017 5015 5014 5013 5012 5010 5008 5007 5006 5005 5003 5002 4999 4998 Sitona humeralis Sitona hispidulus Sitona cylindricollis Sitona ambiguus Sitona cambricus Coelositona griseus Charagmus Hyperafuscocinerea Hyperaarator HYPERINAE terricola Tropiphorus elevatus Tropiphorus TROPIPHORINI scabriculus Trachyphloeus bifoveolatus Trachyphloeus asperatus Trachyphloeus aristatus Trachyphloeus angustisetulus Trachyphloeus alternans Trachyphloeus TRACHYPHLOEINI palliatus Tanymecus TANYMECINI suturalis Sitona sulcifrons Sitona striatellus Sitona puncticollis Sitona macularius Sitona lineatus Sitona lepidus Sitona (Linnaeus) Gyllenhal (Linnaeus) Stephens Gyllenhal (Thunberg) Stephens (Fabricius) Gyllenhal (Marsham) Stephens (Fabricius) (Fåhraeus) (Marsham) (Fabricius) (Newman) (Herbst) (Stephens) (Gyllenhal) Gyllenhal Boheman (Linnaeus) (Beck) Hansen

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ? Vc22 VCH ● ● ● ● ●2002 ● ● ● ● ● ● ● ● ● Vc23 1939-2012 ● ● ● ● ●2001 ● ● ●1951 ● ● ● ●1980s ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon on Caryophyllaceae on ( mercurydog’s oftennear Woodlands perennis] Mercurialis 1935 in at Rare, lastrecorded locallycommon Corallian on a variety on ofground dry substrates. disturbed open Rareon a variety on ofground dry substrates. disturbed open reon a variety on ofground dry substrates. disturbed open Rareon a variety on esp. of ground soils. disturbed open on chalk the areas on esp. open disturbed in ground ?On Rare?Knapweedsburdocks. On and clovers On clovers On Locallybroom gorse and on frequent clovers on legumes] [On gardens including Veryvirtually in frequent situations all clovers on legumes chalk on and medicks esp. sand dryLocal in grasslandon esp. legumes other clovers and grasslands Widespread on in melilots On trefoil large damp wetlands bird’s-foot grasslandin on and Lotus dryrestharrow broom and sadny on local on soils Stellaria, Cerastium, Spergularia Cerastium, Stellaria,

ecology etc.) Lotus uliginosus Lotus 5104 5100 5096 5094 5091 5089 5088 5085 5084 5083 5082 5081 5080 5076 5071 5064 5062 5061 5057 5055 5053 5051 5050 5048 5047 5046 5045 5043 Hyperazoilus Hyperavenusta Hyperasuspiciosa Hyperapostica Hyperaplantaginis Hyperanigrirostris OROBITIDINAE castaneus Pissodes abietis Hylobius plantaris Anoplus (= oblongulum Leiosoma deflexum Leiosoma MOLYTINAE cerasi Magdalis barbicornis Magdalis carbonaria Magdalis armigera Magdalis ruficornis Magdalis MESOPTILIINAE pigra Cleonis paraplecticus Lixus planus Larinus LIXINAE Hyperarumicis Hyperapollux Hyperameles (= pyrenaeum punctata ) ) (Scopoli) (Fabricius) (Scopoli) (Fabricius) (Gyllenhal) (Fabricius) ) ) (Fabricius) (Linnaeus) (Linnaeus) (Linnaeus) (Naezen) (Herbst) (De Geer) (Fabricius) (De Geer) (Linnaeus) (Panzer) (Fourcroy) (Linnaeus) (Linnaeus) (Latreille) Boheman E.Hol Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ●1998 ● ● ● ● ●1997 ●2012 ● ●1998 ● ● ● Vc23 1939-2012 ●1995 ●1999 ● ●1909 ● ● ●1998 ● ● ● ● ● ● ● ● ● ● ● VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Local on pine. An important pest species in commercial An species pest in important pine. Localon plantations pine scot’s on birch Widespreadon ( Buttercups. damp grasslandsanemonein on wood and on woods in Ranunculacae Widespreadon Local,saproxylic Rosaceae, oak. in and developing twigs Rareon ofRosaceae. B birch Rareon elm Localon wych esp. elm Localon wetland umbellifers] [In Extinct docks Widespreadon umbellifers Localon white cloverincreasing on Rarebut sites ruderal dwelling) including cloversvarious TrifoliumgrasslandLocal (ground In on spp. gorse Rareon Lathyrus Lotus, Vicia] On and cloversvarious Widespreadon trefoil bird’s-foot Widespreadon cloverclovers red esp. on Nb ) Rare on Ranunculacae (wood anemone?). (wood Ranunculacae Rare) on Prunus Pinus sylvestris Pinus Ulex etula, esp. Sloe. Early E.Holocene, Mingies Ditch (Robinson ,1993) (Robinson EarlyDitch Mingies Sloe. esp. E.Holocene, esp Oenanthe, Apium Oenanthe, boring in dead trunks and branches. and trunks dead in boring Rumexsp . U.minor. . T.pratense Ulmus glabra Ulmus Trifolium p. Lotus corniculatus corniculatus Lotus T.repens in grassland including roadside verges. roadside grassland including in in wetlands in spp. and medicks and spp. M in hedgerows in

ecology in grassland including roadside verges. roadside grassland including in edicago spp. in grassland including ruderal sites. ruderal grassland including in spp. 5154 5153 5152 5151 5150 5146 5138 5137 5136 5134 5131 5128 5127 5126 5121 5120 5117 5116 5115 5112 5111 5109 5108 5107 5106 5105 Scolytus mali Scolytus intricatus Scolytus SCOLYTINA SCOLYTINI SCOLYTINAE cyaneus Orobitis Xyleborus dryographus Xyleborus dispar Xyleborus (= saxesenii Xyleborinus XYLEBORINA bidentatus Pityogenes laricis Orthotomicus IPINA bispinus Xylocleptes bicolor Taphrorychus Dryocoetesvillosus DRYOCOETINA (= binodulus Trypophloeus tiliae Ernoporus asperatus Cryphalus CRYPHALINA pubescens Pityophthorus CORTHYLINA scolytus Scolytus rugulosus Scolytus multistriatus Scolytus saxeseni asperatus ) (Bechstein & Scharfenberg) in part) part) in (Panzer) (Linnaeus) (Fabricius) (Fabricius) (Ratzeburg) (Müller, P.W.J.) P.W.J.) (Müller, (Fabricius) (Gyllenhal) (Fabricius) (Duftschmid) (Marsham) (Herbst) (Ratzeburg) (Herbst) (Ratzeburg) (Ratzeburg) (Marsham)

Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● ● ●1978 ● ● ● ● ● ● ●1980 Vc23 1939-2012 ● ● ● ●1998 ● ● ● ● ● ● ● ● ●1906 VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon Rare in hardwoods esp. oak esp. hardwoods Rarein oaksappyLocal esp. heartwood. in exposed Beech oakLocal and in bark spruce and pine in local traveller’sLocal in joy hardwoods. other oak and beech, in V.local oak Local in ?[In bark of small-leavedIn lime elmLocal in twigs and branches Local of woody in rosaceae sloe. esp. elmLocal in Rare in recently offoftenon beech broken esp.oakand boughs ofhardwoods branches Widespreadin easily violets Rare(but on overlooked?) Populus Prunus Ulmus Ulmus and and and and Salix Malus Clematis vitalba, Clematis ] ] including orchard trees. orchard including Tilia cordata. cordata. Tilia Viola. Viola. frequent on the chalk. the on frequent

ecology 5195 5194 5193 5192 5191 5189 5186 5182 5178 5177 5176 5173 5172 5170 5169 5168 5167 5166 5165 5164 5163 5162 5161 5160 5157 Hylesinus toranio Hylesinus orni Hylesinus crenatus HYLESININA HYLESININI domesticum Trypodendron 1851 aterrimus Stylops Stylopidae cylindrus Platypus PLATYPODIDAE lemnae Tanysphyrus TANYSPHYRINAE piniperda Tomicus TOMICINA rhododactylus Phloeotribus PHLOEOTRIBINA palliatus Hylurgops Hylastes opacus Hylastes attenuatus Hylastes ater HYLASTINA vittatus Pteleobius hederae Kissophagus obscurus Hylastinus Hylesinus varius (= oleiperda (Paykull) ) Fuchs Erichson (Fabricius) (Danthoine in Bernard) Newport, Newport, (Fabricius) (Fabricius) (Linnaeus) Erichson (Fabricius) (Gyllenhal) (Marsham) (Paykull) (Schmitt) (Linnaeus) (Marsham) Mdv Mdv Subfossil ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● Vc22 VCH ● ● ● ● Vc23 1939-2012 ●1963 ●1963 ● ● ● ● ●1970s VC22 ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● modern Oxon V.local on duckweeds on V.local occasionally and Norway pine, on spruce conifers pine Local esp. on elm. and occasionallyash and on pine on pine ?On pine on twigs in ofelm V.local ivy dead in V.local broom dead in branches. slender in ash, on twigs.recently and in ash, branches Localdead on twigs small in ash, Maybe and Rarebranches. on conspecific with walnut oak and on [Also mainlytrunks, ash, Locallarger and on branches bark on in beech oak and in local in bark on dead gorse and broom stems broom and gorse dead on bark in local Cytisus Hedera helix. Hedera and occasionally gorse stemsand in Ulmus. Picea.

ecology L.. varius L.. (Fabricius, 1775) (Fabricius, Juglans ] Stylops dalii dalii Stylops 1918 R.C.L., bimaculatae Stylops Curtis,1828 Perkins,

Subfossil ● ● Vc22 VCH

Vc23 1939-2012 VC22 ● ● modern Oxon

ecology RDB2) Fringford Rd (Pond Action (Pond Rd Fringford RDB2) (CMD) Otmoor 2009 . (CMD) Otmoor 2009 Bayswater . (JJW) 1907 Brook Bayswater(JJW) 1907 Brook R.Cherwell? JSD) . det (JMC R.Evenlode 1998 (JSD) (JSD) 1998 R.Evenlode etal )

vc23 Cothill (D.Bilton) Cothill

vc22 Nb Nb Nb Nb RD3 Nb RDK ↓ Local, in weedy Local,in margins moss. ofand flowing litter small flooded water, seepages in around also springheads and esecially.ponds Newsilt Widespread.In recently etc.or troughs cleared flier sites,a strong in frequent also . Weston 1907 (J.Collins) Weston 1907 . . Sparsey . Blecthingdon Bridge, JJW in Marston record old One C 19th in Yarnton RDB3) Shotover (CMD) Otmoor 2009 Action 1988 Pond

vc23 Hutchin’s Copse Copse Hutchin’s Copse Hutchin’s

vc22 Nb Nb Nb Nb Nb RD3 Nb ↓ Ubiquitous in standing water standing especially of in kinds, all where as arelarvaeamphibians Ubiquitous breeding, tadpoles. specialise in Localy abundant at Otmoor in ponds and ditches. and ponds LocalyatOtmoorin abundant . (VCH) Yarnton Yarnton Marston, (PNicolete) 2007 Standlake (CMD) Otmoor 2009 Otmoor JJW in Marston (JSD) 2005 Stoke South Ambrosden in area ponds Field . Otmoor (CMD)Otmoor

vc23 Hutchins Copse 1997 (JSD) 1997 Copse Hutchins (JSD) 2011 Brook Sandford (JSD) 2011 Brook (VCH Sandford Wytham.

vc22 Nb Nb Nb Nb Nb Nb ↓ ↓ ↓ ↓ Field ponds near Ambrosden near ponds Field (CMD) Otmoor 2009 . VCH (R.Angus) 1963-9 Cothill Common Nettlebed (A.Hine) Angus ActionRob Ogley Bog(H) Pond Smith) (Matt Common Nettlebed 6.10.1999 (W) St.John Near . Straton

vc23 Tubney((WH) Tubney etc (JJW) Hill Boars Wood Bagley Extinct.

vc22 [Na] Nb Nb Nb Nb ↓ ↓ Oxford Uni Parks 1953 (P.J.Osborne) (P.J.Osborne) 1953 OxfordParks Uni extinct Specimen labelled Oxford in JJW coll. Oxford JJW in labelled Specimen extinct

vc23 Bagley Wood 1952 (H.G.Champion) (H.G.Champion) 1952 Wood Bagley

vc22 Nb Na Nb ↓ dampshady places ,1993) only from Sub-fossil from Ditch Mingies (Robinson late E.Holocene (PW) water.near baresand Horsehays 8.4.1989 Rare on pits (VCH) sand BogShotover Bullingdon (PW) 19.5.1989 WestonFen (DC) rareClattercote Reservoir 18.5.1985 (GLF) Northmoor 1953 . Local,wetlands, sandypartly in river subterranean etc. banks (PW) LowerOtmoor Water 5.5.1989 Eaton. Windrush (PW) 1989 Weston Fen Oxford (Hope) (Shipp) Stadhampton rare.Aston Rowant 1982) materialOx dist JJW

vc23 Hitchcopse Pit (DJM) Pit Hitchcopse Hitchopse Pit 1986 (PW) 1986 Pit Hitchopse

vc22 RD3 Nb Nb Nb Rare in damplitter. Rareplacesin in Cowley shingle Rare,on (JWS) Cowleydates no (J.W>Shipp) (PW) Gravel 1986 Windrush Pits etc ponds near Commonbareground on Oxford (VCH) Headington, . PW) (DC, 1988 Lower1977, Windrush mammal Rare?[In burrows] or runs (GLF) HytheBablock modern records 4 1953

vc23 Wytham park 1913 (HGC) 1913 park Wytham

vc22 Nb Nb Na↑ Nb Na Nb ↓ Subfossil only. A northern species now found from species nowfound only. A Subfossil Staffordshire northern northwards. Bankof Thames (Hope) (KNAA) 2007 Great (SU2796) faringdon (WS) 1945 Wolvercote(DC) Ponds RareClattercote Res 1984 Otmoor(W) dampplaceswater r nea 1896 Oxford(JWShipp) (PW) Gravel 1989 Windrush , Pits 1986 Cherwell R. Northbrook Mesopotamia (JWS) Mesopotamia(DC) VCH Rare Banbury 1984

vc23

vc22 Nb Nb Nb Nb Nb↑ Nb Nb Nb ↓ ↓ Windrush 1985 (PW) (PW) 1985 Windrush Chimney (PW) Otmoor1989 Meadows 1988 common atOtmoor. ? partly subterranean. Rare,though Marsham,VCH Waterin Eaton (FWH) onlyrecord Oxford1819

vc23 Tubney <1906 (JJW) (JJW) Tubney <1906 Hill Boars Tubney (WH) 1904 & Tubney 1900 (JSD) 1998 Copse Hutchins

vc22 Nb Nb Nb Nb Nb Nb Nb RD1 ↓ ↓ . Clattercote Reservoir 1988 (PW) Clattercote Reservoir . 1988

vc23 Hitchcopse Pit , Frilford Frilford , Pit Hitchcopse (M.Amphlett) 1976 Drayton (DJM)1998 Pit Hitchcopse

vc22 Na Nb Na Nb Na Nb LWGPP.W pit (L.Jones-Walters) AstonRowant 1982 areas. ruderal including habitats open Widespreadin (L.Jones –Walters) Aston Rowant. 1982

vc23 Tubney 1906 (HSD) Tubney1906 Frilford (PW) Tubney1984-86 (PW) 1986 Pit Hitchcopse Cothill, Tubney (VCH) Cothill WythamUnconformed Survey 1951 Hitchcopse Pit 1999 (D.Newman) 1999 Pit Hitchcopse (PW) Pit Tubney (VCH)Hitchcopse Cothill Corallian. (JMCJSD) Glebe det

vc22 Nb Nb Nb Nb Nb↑ Na Nb Nb Tadmarton G.C. 2000 (JMC det PW) det (JMC Tadmarton G.C. 2000 Gibraltar . Quarries1903 aLocal variety in especially of habitats sandy soils. on cholsey Ardley (PW) QuarryFields 1988 Horsehays (PW) Quarrycoll.) (Hope 1987 1902 Shotover Wychwood (PW) (VCH) 1988 Harcourt chalky sandyStanton [Open and sites] RDB3 Gibraltar Weston Quarries 1907 Emslow 1909 Shotover RDB1 Hardwicke 1896 ) ? ) AAA) det Cholsey (Holland & Streatley Shothover ) 1905

vc23 Shotover Hitchcopse 1990 1990 Hitchcopse Shotover Dry Sandford 1986 (PW) (PW) 1986 Dry Sandford (PW) Tubney , 1986 Cothill (JJW) Tubney 1904 (AGC) Tubney(VCH) 1906 Cothill Quarry 199 (VCH)Shillingford Cumnor

vc22 Nb Nb RD3 RD3 Na Na RD3 RD1 Nb Nb Nb RD1 Na ↓ Pre 1938. Cowley 1880s 1938. (JW Shipp) Pre (VCH) Shotover (PW) Whitecross . Green 1988 Wood large wastecommon and gardens. Widespreadand including ground (PW) Quarry Eaton 1986 Wood . PW) det (JMC 1987 Hill Watlington Water(VCH) , Eaton Yarnton . (JSD) Ambrosden 2012 JJW in Ox dist (GLF) Otmoors. etcHythe Bablock 1953 marshy Rarein areas, comes also light. to PW) det (KP 1987 Weston Fen ground]. cultivated clay areas including Rare?sand on [Open and boggy Rarein areas marshes. Rarein marshLocal wet in areas. and heath stones. under margins and litter in wetlands,Local pond in

vc23 HitchcopseDJM) 2001 Pit Tubney Frilford (VCH)etc Tubney Frilford (JEC) Park Wytham

vc22 RD1 Nb Nb Nb RDI Na Nb Nb ?↑ RD1 Nb Nb Na Nb Nb ↓ ↓ Widespread on deciduous trees and pine esp. on dead and moribund boughs and branches, often under bark. oftenunder branches, and boughs moribund and dead on esp. pine trees and deciduous Widespreadon Subfossil only [wetlands] Early E.Holocene (Robinson, 1993) Early (Robinson, only[wetlands] E.Holocene Subfossil PW) det (JMC Wychwood 1989 Forest (PSH) Waterperry 1988 Woods (JSD) Ambrosden 2012

vc23

vc22 Nb RD1 Nb Nb Widespread in pools and ponds, especially very commonponds, avoiding smalland in woodland, water.acid leaf-filledin pools Widespreadin puddles Otmoor 2009 (CMD) (CMD) Otmoor 2009 (JJW) Banks of.CherwellR 6.5.1907 especiallyweedyLocal heaths. in on ponds detritus (JJW) Marston King’s Weir VCH

vc23 Wytham Park 22.4.1907 (JJW) 22.4.1907 Park Wytham

vc22 Nb Nb Nb Nb Nb RD3 Nb Nb Nb Nb Nb ↓ South Stoke (JMC det JSD). det (JMC Stoke South atmarginsdetritus Local of ofwater kinds allin bodies. flowing atmargins and silt Localwater. in ofponds (JSD) Bicester 2002 scarceLocal and in. ponds Rarein aweedy variety in and Frequent of ponds flowing water. 1980) Alchester (Robinson, vegetation. withlittle new increasing in ponds Widespread and (MCD) Otmoor 2009 atOtmoor Frequent (CMD) Otmoor 2009

vc23 ??Tubney (JJW) Hutchins Copse 1998 (JSD) (JSD) 1998 Copse Hutchins (JSD) 2011 fen Barrow farm

vc22 RD3 Nb Nb RD3 Nb Nb Nb Nb RD3 Nb ↓ ↓ ↓ ↓ Kings Weir (WH) (JSD) 2005 Stoke South , Otmoor 2003 . (CMD) Otmoor 20909 ..

vc23 Wytham Park 1907 (JJW) 1907 Park Wytham (TR-SJSD) det 2012 Wytham beech. especially bark under Tubney (WH) (JSD) 1998 Copse Hutchins Tubney (VCH) (JSD) 1998 Copse Hutchins

vc22 Nb Na Nb Nb Nb N [N] ↓ ↓ ↓ ↓ Hardwick(VCH) Duke’s Cut (JEC) elm 1907 hollow starlingnest Waterin Eaton Hartslock 1991 (TDH) Hartslock 1995 EynshamWater Eaton (VCH) ?

vc23 [In mole nests Frilford JMC det JSD Hitchopse Pit 1993 (TDH) 1993 Pit Hitchopse JSD det JMC mole Frilford nests [In (JEC) Tubney ants. wood With Extinct? 1907 local on ground especially on sandy soils. Frilford soils. sandy on especially ground on local

vc22 RDK Nb Na RDK ↓ SparseyBridge (CMD) Otmoor 2009 . (AME) R.Thames, . 1946 Shillingford cGosford Bridges (W) (CMD) Otmoor 2009 . 1993) Early (Robinson, only[wetlands] E.Holocene Subfossil 1993) Early (Robinson, only[wetlands] E.Holocene Subfossil , Extinct? (GLF) BablockHythe 1953 1993) Early (Robinson, [wetlands] Holocene ApplefordWater Eaton ] Caversham Cowley Warren, Shotover Water(VCH) Eaton Caversham . Cowley Warren, Shotover Water(VCH Eaton S ubfossil only [wetlands] Early E.Holocene (Robinson, 1993) Early (Robinson, only[wetlands] E.Holocene ubfossil

vc23

vc22 RD1 Nb Nb Nb Nb RD3 Nb RD3 RD3 Nb Nb Nb↑ Nb↑

↓ ↓ Under barktreesof dead especiallyUnder VCH Oak. WaterYarnton Eaton Green the Westonon JJW Weston Godstow (VCH

vc23

vc22 N RDK N RDK N N RDK N N RD3 Local? [In compost etc.], Local?[In South Stoke 2002 (JSD) 2002 Stoke South ??

vc23

vc22 N N RDK RDK Themost frequentlyencountered Leiodes , frequent in flight interception and malaise traps, but hard to find by other meansby find other to malaise hard and but flight traps, interception in frequent , Marcham Park 2003 (JMC det JSD) det (JMC Marcham2003 Park Summertown (JJW) chalk]. Wytham Beech on woods 1904 1906 (TH) Hartslock 1994 downland. and Woods Summertown,Water (VCH) Eaton, Yarnton

vc23 Woodlands?]. as Wytham parvula Woodlands?]. (JJW) Chalky(JEC) sandy Tubney and places]. 1928 Wytham Park (JJW) (JJW) Park Wytham (JJW) park ?TubneyWytham 1906 Wytham 1906 Tubney 1904 (JMCJSD) det 2009 , 1999 Glebe (JMCJSD) det 1999 Glebe ?Wytham curta as (JJ Tubney 19.10.1907 (JMCJSD) 2005 det 199, Frilford (JEC) Tubney 1928

vc22 RDK RDK N RDK RDI RDK RDK RDI N N RDK OgleyFerry Marston (JJW)Bog, Yarnton (AME) R.Thames, 1946 Shillingford Wychwood1966 Wychwood Ascot on – (VCH) Otmoor 2000,(JMC det JSD) det 2000,(JMC Otmoor Tubney Park Wytham 1996 Nettlebed

vc23 Tubney (WH) (HSD) WythamPark13.9.1907 Tubney , Wytham 1906 (JJW) (JJW) 1906 Tubney Wytham , (JJW) Tubney 1906 (JSD) 1998 Copse Drayton (JSD) 1998 Copse Drayton (JMCJSD) det 2008 Frilford 2000, Otmoor (JMCJSD) det 2008 Frilford (JSD) 1998 Copse Drayton Copse/ Hutchin's (VCH) Wychwood

vc22 RDK RDK RDK N RDK N RDK . Hartslock 1991 (SH) Hartslock . 1991 OxfordElsfield (JJW) (VCH) Yarnton WaterIslip, Eaton Ogley (JJW) Yarnton Bog, Yarnton , Elsfield, (VCH) Elsfield, , (JMCJSD det 2000 Taynton Fen (JJW) Tubney 1906 (JSD) 2003 Otmoor

vc23 Tubney Cothill (JJW) (JJW) Tubney Cothill 1998 Copse Hutchins (WH) Tubney<1906 MC ( JSD 1999 det Frilford (JJW) Tubney,Wytham Hill, Boar's (JMCJSD det 2000 House Dog (JJW) Tubney 1906 MC ( JSD 2002 det Frilford 2002-4 Tubney Wood (JJW) Tubney 1906 (JJW) Tubney 1906

vc22 N RDK Elsfield, Yarnton, Brasenose (VCH) Elsfield,Yarnton, Yarnton Sunnymead (VCH) Yarnton Elsfield downland moss oftentreeon clad trunks. woods, Local in Wolvercote,Summertown, (JJW) Botley 1908 Wolvercote SummertownMarston (VCH) Wolvercote (JJW) VCH Wood Prattle Eaton Wood ?

vc23 Local in compost/manureLocal Wythamin heaps. etc (VCH) Tubney woodlands, Rarein Hutchins Copse, Cow Common 1998 (JSD) (JSD) 1998 Cow Common Copse, Hutchins (JJW) Wytham (JJW) Wytham Bagley,

vc22 RD2 RD3 RDK N RDI N N RDK RD1 RDK RDI Local on carrion. Localon (TH) Hartslock 1989 carrion. Local in Extinct taken in carrion at Witham in 19th C 1822(FWH) C1822(FWH) 19th in Witham at carrion in Extinct taken (JMCJSD) det 12.9.2000 Shotover

vc23 Tubney, Frilford ect Tubney,Frilford Downs etc East

vc22 Nb Nb RD2 Nb Na Summertown 7.6.1907 (JJE) 7.6.1907 Summertown (JJW) Yarnton 227.2.1907

vc23 Bagley Wood JD) det (JMC 2008 Frilford 1906 Hill Cumnor Tubney 25.5.1907

vc22 N N N N vc23 Hutchins Copse 1998 (JSD) 1998 Copse Hutchins Cothill

vc22 Nb Na RDK Na RDI Summertown( Yarnton Quarry 1920 Blechingdon I (VCH) Kidlington Bablock Hythe 1953 (GLF) 1953 Hythe Bablock

vc23 Hutchins Copse 1998 (JSD) 1998 Copse Hutchins

vc22 RDK Nb N RDI RDK N RD1 Na Yarnton 1907 (JJW) 1907 Yarnton Beckley,(VCH) Yarnton Wolvercote (VCH)

vc23

vc22 N RDI RDK RD2 RD3 RDK N N RDK Shotover 2000 (IW det JSD) det (IW 2000 Shotover (JMCJSD) det 1984 Burford (VCH) Ogley Wytham

vc23 Tubney 1905 & 1906 (JJW) & 1906 Tubney1905 Drayton Copse 1998 (JSD) 1998 Copse Drayton KG 1990 Wytham Survey Wytham Survey Wytham

vc22 N Nb Na N Yarnton (WH) Otmoor (JJW) 20.6.1907 Summertown Oxford (VCH) Oxon VCH Uni (Creid) 1980 Wood Waterperry

vc23

vc22 N Nb Na RDI RD1 N Ellisfield Meadow (VCH) Church ?Christ Water Eaton, (VCH) Yarnton Eaton Wood ?Downland Wytham?Water Eaton, (JEC) Hardwick. . Wytham survey Wytham review see JMCVG det 1989 Wychwood (JJW) 16.3.1907 Wood Hen Survey Wytham (WH) Bog Ogley (JEC) Eaton Wood Shotover (JMCJSD) det 2000 Shotover

vc23 Wytham, Tubney (JJW) Wytham, (RCW) 1981 Wytham cotes dove In (JSD) 1998 Copse Hutchins JSD 1998 Cow JMCJSD Common det 1999 Frilford (JMCJSD) det 2009 central Glebe TDH 1996 Pit Hitchcopse Copse Hutchins Survey Wytham

vc22 N RDK RDK RDK N RDI RDK N RDK RDK N i. Wychwood 1965 (GEW)Wychwood i. 1965 Ent dte (JMC 1992 Vicarage Pit ?Downland ) Wychwood 1938 Forest Yarnton Taynton JMC Taynton Entwhistle Yarnton det JMCEntwhistle 1992 det Pit Vicarage common 1990 Pit Vicarage VCH Bridge Enslow

vc23 Hutchin's Copse 1998 (JSD) 1998 Copse Hutchin's 1908 Tubney Wytham (JEC) 1907 Besselsleigh Kraatz splendens Homolota (GCC) as 28.v.1906 Hill umnor

vc22 N N RDK N N N RDK RDI RDI common in rotting fungicommon rotting in Water Eaton JSD) det (JMC 1992 Vicarage dampPit litter] ?[In Water Eaton Water JMCEntwhistle 1992 det Pit Vicarage (JJW) Eaton Water

vc23 Tubney(JJW) Tubney(JJW) Wytham1950 Wood Rare in damp woods. Tubney (JJW) woods. damp in Rare (JSD) 1998 Copse Hutchin's Dr.V.Gusarov Wytham (JSD) 1998 Copse Hutchin's Dr.V.Gusarov Wytham Tubney (JJW) (JSD) 1998 Copse Hutchin's

vc22 RDK N RDK N RDK N N . Yarnton . Caversham ?Downland. marshes? Rare in carr. Islip and Water(VCH) Bisney Eaton Eaton, Wood Ent det (JMC 1992 ?Vicarage Pit Godstow Bladon Wychwood 1965 Shotover Hill 21.5.1907 (AJC) 21.5.1907 Hill Shotover Yarnton (TDH) 1992 SU7676 R.Thames (JSD) 1998 R.Evenlode (JEC) Elsfield Eaton, Water 1907 pre Elsfield

vc23 Wytham Dr.V.Gusarov Wytham Wytham Dr.V.Gusarov Wytham Dr.V.Gusarov Wytham Dr.V.Gusarov Wytham (JEC) Tubney 1907 Dr.V.Gusarov Wytham 1907 pre Tubney (JJW)

vc22 N N RDK Rare? in wetlands. R.Thames, Shillingford 1946 (AME) wetlands. Rare? R.Thames, in 1946 Shillingford in Shirley and (JJW) Otmoor (JSD) Yarnton (AHH) 1908 Shotover Cherwell on Hogley , Shipton Bog, (VCH) Yarnton (TDH) Hartslock 1995 (JJW) dung Waterin Eaton Wychwood (GEW) forest 1965 Ent) det (JMC 1992 Vicarage Pit Water Eaton (JEC) Eaton Water Oxford Dr.V.Gusarov Wytham

vc23 Tubney formerly Extinct (JEC) atTubneynests ant wood before in 1907 Cothill 30.9.1907 (AJC) as laticepes laticepes as (AJC) 30.9.1907 Cothill (JEC) 1907 Tubney before (JJW) 1907 Park Wytham Dr.V.Gusarov Wytham Dr.V.Gusarov Wytham Dr.V.Gusarov Wytham

vc22 N RDK N N RDK N RDK N N N Cowley Summertown (JEC) Water (HSD) Eaton 1907 Ent) det (JMC Vicarage1992 Water . Eaton Yarnton . moss wetland and areas. litter in RareBayswater in 1917 Wood Prattle Shotover 2002 (J.Wright det JSD) JSD) det (J.Wright 2002 Shotover Yarnton 1910 Yarnton 1908 Yarnton Yarnton

vc23 Wytham Dr.V.Gusarov Wytham Wytham 1998 Hutchins

vc22 N N N RDK N N N RDK Vicarage Pit 1992 JMCEntwhistle 1992 det Pit Vicarage W.Wint Dr. Wood Bagley Howe Wood (JMCJSD) det 2004 Wendlebury (JJW) King'sWeir

vc23 Howe Wood Wytham Survey Wytham Howe Wood Wytham Dr.V.Gusarov Wytham (JSD) 1998 Copse Hutchins

vc22 N RDK RD1 RDK N N Ox before 1969 see Allen 1975 Johnson 1967 as nitida 1967 Johnson see Allen Ox1975 before 1969 1955 . (VCH) Eaton Wood (JJW) LowerCaversham1907 Yarnton (VCH) as convexicollis VCH in Godstow

vc23 Cothill 1905 (JJW) 1905 Cothill 1914 Cothill

vc22 N Nb RDK RDK N N N N N N N EXTINCT Weston-on-the-Green 1853 remains Record! only EXTINCTBritish the Weston-on-the-Green 1853 (JJW) Yarnton (AHH) Wytham 1907 Wick Sydlings(modern) (JJW) & Copse BagleyShotover Headington Copse 1954. 1907 Wood sites Widespread,3 Headington . Yarnton 1908 ?????? Yarnton 1908 (WH) Grandpont (JJW) Park Wytham Elsfiedl, Hills, Chilswell with ants. (DC) 1994 Alkerton (JEC) Tubney 9.2.1908 nests ant In with Myrmica ants common 1910 1911

vc23 Tubney (JJW) Hill Boar's

vc22 EXT N N N RDK N N N RDK Water Eaton Yarnton Wolvercote Yarnton Water Eaton Summertown, Woodstock nr Wood, Water Otmoor Eaton, Wooton Water Eaton 1920s in Water Eaton Yarnton, W Eaton Wood Sparsey Bridge(W, R.Thame(TDH) C) 1992 Onslow Bridge(TC) Gibraltar Quarries Yarnton (JJW) <1906 Yarnton (JJW) Yarnton 1908

vc23 Tubney (JJW) Tubney (JJW) <1906 Tubney (JJW) Cumnor, (JJW) 25.5.1907 Cothill

vc22 RDK N RDK N N RDK RDK N RDI N RDK RDK RDI Yarnton Swamp (CD) 1924 Swamp Yarnton (CD) 1924 Elsfield WaterIslip Eaton Eaton Wood Yarnton Eaton Wood Ferry Marston Lane W Blenheim 1978 Clanfield VCH, Yarnton, Wateras collinsi Eaton Wateras waterhousei) Eaton Waterhouse as nigrina Yarnton Water Eaton Kidlington, Shipton-on-Cherwell, woods. etc.in litter Local in EmmaGreen VCH green. the Weston on (G) Bladon sandy ?Open ground? Way Icknield (TDH) VCH 1996 Yarnton Bagley Wychwood 1980s VG VG 1980s Wychwood Bagley Shotoevr Bagley

vc23 Wytham (JJW) Wytham (JEC) Tubney Novwith 1907 wood ants (JEC) Tubney 7.8.1907 (JSD) 1998 Copse Hutchin's (JSD) 1998 Copse Hutchin's Wytham 1920 Wytham (JJW) park Tubney Wytham (JJW) 1906 Park Wytham

vc22 RDI N N Headington Headington R.Thame(TDH) 1991 wetlandsSparsey BridgeWolvercote (VCH) (AHH) Shotover JD) det (PW Otmoor 2000 Wood. Prattle . Ogley Bog, Yarnton Bog, Ogley Shotover (D.Steel) 1970 Shotover (DC) 1994 Hitchcopse auct teres elm on fungus (GCC) in 1905 Eaton Wood

vc23 Hutchin's Copse 1998 (JSD) (JSD) 1998 Copse Hutchin's (JEC) Tubney Novwith 1907 wood ants (JSD) 1998 Copse Hutchin's (JEC) 1907 Wytham (DC) 1994 Hitchcopse auct teres 1906 Wytham (JEC) 23.3.1907 Wytham

vc22 RDK N N N N N N N RDI N Gibraltar Quarries pre 1907 (JJW) Gibraltar1907 Quarries pre In moles nest (JJW JEC) (JJW nest moles In (JJW) 17.6.1907 Elsfield

vc23 Hutchins Copse 1998 (JSD) (JSD) 1998 Copse Hutchins (JSD) 1998 Copse Drayton 1919 Wood Wytham

vc22 N N RDK Nb Nb RDI RDK N N ↓ North Moor 1953 (GLF) MoorNorth 1953 (JJW) 18.4.1907 Yarnton

vc23 Cothill, Cumnor, Tubney Cumnor, Cothill, (JSD) 1998 Copse Hutchins (JSD) 1998 Copse Hutchins

vc22 RD1 Na N Na Bayswater (JC) Mill (SH) Hartslock . 1991 Otmoor JMC 2003 Hays Horse (KP) 1987 Lane Lashford JMC Court Friars Otmoor

vc23 [Wet woodlands] Drayton Copse 1998 (JSD) Drayton 1998 Copse woodlands] [Wet Barrow farm fen Barrow farm Wytham 1956 Wytham Tubney (JJW)

vc22 Nb RDK RD1 N N Wood Eaton Eaton Wood (VCH) Dukes . Court Sparsey Wytham Wetlands] WestonBridge, (VCH) 1951 (VCH) Yarnton (VCH) Yarnton Beckley,(WH)Water Eaton Yarnton, (TDH) Otmoor1998 Water(VCH) Eaton (VCH) Stow Wood Yarnton, Yarnton, (VCH) Yarnton, Medley (WH) (TDH) 1999 Otmoor Yarnton Wolvercote vch green the on Weston (WH) Ogley Bog Stonesfield Otmoor Burford Yarnton

vc23 WythamGusarov 1989 Woods Tubney (VCH)

vc22 RDI Nb Nb Nb Nb N Nb N Kidlington, Wood Eaton Eaton Wood Kidlington, Bagley Wood, King's Weir, Yarnton (VCH) Weir, King's Wood, Bagley Bicester (JEC) Eaton Water Weir, King's (JSD) 2000 Otmoor (GLF) 1953 Hythe Bablock (JSD) 2005 Stoke South Wytham Eaton, Water

vc23 Wytham (VCH) Wytham (JSD) 1998 Copse Hutchin's

vc22 N N Nb N Nb Otmoor Wendlebury 2003 (JMC det JSD) Bagley Yarnton (VCH) Yarnton Bagley (JMCJSD) det 2003 Wendlebury Otmoor Yarnton (VCH)

vc23 Tubney (VCH) (CEC) Tubney 1899 (JMCJSD) det 2002 Frilford (JJW) Tubney 1905 Hill, Boar's (JEC) 1905 Wytham

vc22 RDI N RDI N Nb RDI N RDI RDI RDI RDI N RDK Eurytopic] Hartslock 1995 (TDH) Hartslock 1995 Eurytopic] R.Evenlode 1998 (JSD) Wendlebury 2004 (JMC det JSD) (JMCJSD) det 2004 Wendlebury (JSD) 1998 R.Evenlode Eaton. Yarnton Wood etc. heaps manure In (WH) Bagley

vc23 Hutchin's Copse 1998 (JSD) (JSD) 1998 Copse Hutchin's (JEC) 15.9.1907 Wytham (JSD) 1998 Copse Hutchin's Tubney(VCH)

vc22 RD1 Nb N N RDK N Elsfield (JEC) Elsfield Stoke South (HSD) Stoke South (JEC) Eaton Water

vc23 Frilford (JJW)? 1907 Besselsleigh (JJW) Tubney 1907

vc22 RDK RDK Nb N Nb Nb ? Water Eaton, Weston on the Green Oddington (VCH) Green the Oddington WestonWater on ? Eaton, JJW etc in as dissimilisYarnton Water Eaton (JEC) Eaton Water

vc23 Hutchins Copse 1998 (JSD) (JSD) 1998 Copse Hutchins

vc22 RDK Nb Rare in litter and moss and water.litter near Rarein (DC) Hartslock1997 JSD) det (JMC MeadowBurford RarePort Wytham etc.??Brazenose (D.Steel??) Wood M.Shaw (M.Jacoby) 1970s ExladeSt. Chilswell1907 Rare,etc. saproxylicmould Park wood Hollowtrees in in ‘fortresses’!] JSD) breeding more det the (JMC nests, Wychwood appropriatelyin Mole 1989 Blenheim/ Wychwood det) (JMC Saproxylic. Summertown 1907 (JJW) Wytham 2012 ( det JSD) det ( 2012 Wytham (JJW) 1907 Summertown Saproxylic. (Holland) 1898 Shotover 9AHH) Ogley Bog (JEC) Eaton Wood ,

vc23 Besselsleigh Wytham (VG) Wytham (VG) Wytham (TR-SJSD) det 2012 Wytham (VCH) Wytham (JMCJD) Downs det 2008 East (JEC) Tubney 1907 at ants with Wood Extinctformerly

vc22 Nb Nb Nb Nb Nb Nb Nb Nb Nb Na ○ oak farm (SP5610) 2008 oakfarm (SP5610) Brighthampton . Glebe N 2002 (JMCJSD) det N2002 Glebe Tubney (JMCJSD) det Frilford Tubney (JJW) 1907 pre (JEC) Eaton Wood

vc23 Drayton Copse 1998 (JSD) (JSD) 1998 Copse Drayton (JMCJSD) det Frilford (D.Bilton) 1990 Hitchcopse (JSD) 1998 Copse Drayton

vc22 Nb Nb Nb Na Na RD1 RDK Na ↓ Ogley Bog Yarnton etc Summertown, etc. heaps manure In Yarnton etc Summertown,

vc23 Tubney 1907 (JEC) Tubney 1907 (JEC) Tubney 1907 with wood ants Extinctformerly

vc22 Na Nb Rare?Hartslock CowleySummertown . (JEC) Rd 1936 (M.Harvey/Chris Hartslock Raper) 1995 15 . caversham , Warren 1938 pre earlyOxford in in 19 species. EXTINCTFound Ancient woodland Oxford JJW Tubney etc th Century (Robinson, 1986) Oxford (JJW) 1986) Century (Robinson,

vc23 Wytham (VCH) Wytham (JJW) Tubney 1906 (JJW) Tubney 1906 th C on several on C occasions.

vc22 Nb Nb Na Na EXT Nb . Marston Ferry Wood Eaton 1906 Ferry Eaton Marston Wood . Islip (WH) pre 1907) Bagley Wood (JJW) Wood Bagley 1907) pre (WH) Islip VCH Hardwick (JJW) Elsfield Oxon (FWH) 1919 (JJW) 1906 Radley 1977

vc23 Hutchin's Copse 1998 (JSD) 1998 Copse Hutchin's (WH) Hill Boars abundant locally still Tubney area Frilford (JJW) hill Boars (VCH) Tubney Cothill

vc22 Nb Nb RD1 RD1 Nb Nb Nb Na Peppard Common 1998 Common 1998 Peppard (GEW) Rare,Wychwood 1966 Forest Bagley Wood (JWS) (JWS) Wood Bagley

vc23 Dry Sandford S.Grove 1992) Dry Sandford (F.J.Parker) 1942 farm Sheepstead Heath Frilford Tubney 1913. (JJW) 1904 heath Frilford

vc22 RD2 Nb Nb Na EXT RD1 Na . Sarsgrove 1994 (JMC det JSD) det (JMC Sarsgrove . 1994

vc23 Hutchins Copse 1998 (JSD) Barrow Farm Fen 2012 (JSD) 2012 Fen Barrow (JSD) Farm 1998 Copse Hutchins (JSD) 2012 Fen Barrow (JSD) Farm 1998 Copse Hutchins (JSD) 1998 Copse Hutchins (JSD) 1998 Copse Drayton (JSD) 1998 Copse Drayton Abingdon

vc22 RDK RD1 .] Stanton St.John 1919 JEC 1919 St.John Stanton .] Entwhistle det JMC Wychwood . 1992 Bagley Wood (JW Shipp) Shipp) (JW Wood Bagley

vc23 Wytham Park 1907 (HSD) 1907 Park Wytham (JSD) GCCarswell 2004 (JSD 1998 Copse Hutchins

vc22 Na RD2 RD1 Nb Na Nb Na Nb Na Nb Nb ○ s. Bicester 1909 Weston Vc23 2002 Vc23 Weston Bicester s. 1909 2004 JEC), Bayswater Marston . Weston 1924 1910 R.Thames inReading on OxonSide. 1988 Gosford BridgesVCH JSD det Aston Rowant. 2002 L.Jane-Walters AstonRowant NNR1982 Great Brooks 1990 1990 Brooks Great

vc23 Frilford Tubney Frilford (JJW) Tubney 1906 (JJW) Tubney 1906 (JJW) Tubney 1906 Tubney Wytham Cothill 2001

vc22 Nb Nb Na RD3 Nb RDI Shotover 1915 Shotover (CMD) Otmoor 2009 1993-2011 Road, Fringford Otmoor 2009 (CMD) 2009 Otmoor 2001 Vc23 2002 Vc22 1975) Robinson, & (Giorgi Remains Roman Alcehster 1938 pre SP41

vc23 Drayton Copse 1998 (JSD) (JSD) 1998 Copse Drayton (JSD) 1998 Copse Hutchin's Cothill

vc22 Nb RD3 Na RD3 RD3 Nb

On Corallian eas tof Oxford at Wheatley and Garsington pre 1938 (VCH) 1938 pre Garsington and Oxford tof Wheatley at eas On Corallian by water Pixey Mead 1992 (S.Grove) Bagley Pixey1992 Mead (JWS) Wood

vc23

vc22 RD3 Nb Na BagleyC (FWH) 19th Wood HM) det (JMC park 1993 Cornbury (KNA) (DC) 2002 Blenheim1985 Park Pucketty farm 1970s MVCorley 1970s farm Pucketty 2007 vc23 2003 vc22 2001 (JWS) Dorchester Atlas) Click (see records Old

vc23

vc22 Na Nb Na RD3 Na Na Na RD2 ○ ○ Warren farm 1985 JMCKNAA det 1985 farm Warren vc22

vc23

vc22 Nb Nb RD3 Nb ○ ○ Oxford 1821 (FWH) Oxford (FWH) 1821 West W Sr 1980s Wood Bagley JMCKNAA det 1989 Wychwood

vc23

vc22 Nb Nb Nb Nb Nb Nb Thame in 15th C (Smith, 1999) Water Eaton pre 1938 pre Eaton Water 1999) C(Smith, 15th in Thame (WH) Wood bagley Summertown ? (Britten) 1916 Park Thame (JSD) 2005 Stoke South KNAA Woodstock JMCJSD Burford det

vc23 Hutchins Copse 1998 (JSD) 1998 Copse Hutchins (JSD) 1998 Copse Drayton

vc22 Nb RD3 Nb Nb RD3 Nb Nb RD1 Tubney (KNAA) 1986 Park Cornbury (KNAA) 1986 Park Cornbury 2000 Park Ashdown DW.Wint 1980s Bagley (JEC) 1914 Thame

vc23 Marcham Park 1995 JMCJSD) det 1995 Park Marcham

vc22 RD3 Na Nb Na RDK RD2 Nb Nb Oxford 1819, 1822 (FWH) 1822 Oxford1819, Thame in 15th C (Smith, 1999) 1999) C(Smith, 15th in Thame 1986) C(Robinson, Oxford 15th in Priory (M.Harvey) 1998 Hartslock

vc23 Cow Common 1998 (JSD) (JSD) 1998 Cow Common

vc22 Nb Nb RD3 Nb Nb RD3 Nb Local on sedges and rushes in damp areas, in sedgesrushes marshes Local and on etc. flowersmeadowsweet esp. sedges Local on (JJW) Water1906 Eaton (JMC) Stoke South . gardens.Saproxylic. rural parks and woods, Local,in Pixey Summertown Mead GrasslandsEaton, Wood . Garsington (VCH) ?Hardwick Wychwood(VCH) 1953 Carex in bogs, marshes bogs, in etc.Also on Filipendula ulmaria Filipendula

vc23 Wytham1921 Wood Drayton Copse 1998 (JSD) 1998 Copse Drayton (JSD) 1998 Copse Hutchin's (JJW) Cothill

vc22 Nb Nb RD3 Nb Nb Na Rare on Rareon Commonnettles on Commonnettles on damp areas, in sedgesrushes marshes Local and on etc. ! Taynton Fen 1990 JMCEntwhistle det 1990 Taynton Fen ! Linaria . Urtica. Urtica. Urtica. Spp Spp. .

vc23 Bagley Wood 1980s DrW.Wint 1980s Wood Bagley DrW.Wint 1980s Wood Bagley

vc22 Nb Nb Na Nb Ubiquitous on various yellow crucifers, also attracted to bright clothing, one of the mostof the yellowone commonvarious on clothing, of beetles! crucifers, bright attracted also to Ubiquitous . Wood Eaton, Shotover (WH) Shotover Eaton, Wood . black Localhorehound on crucifers, Rarecharlock on viper’s bugloss Veryon local Weston- on the Green the on Weston- Ballota nigra Ballota Sinapis arvensis Sinapis Echium vulgare Echium & hedgemustard .

vc23 Tubney (JEC) (JJW) (1920 Tubney Wytham Tubney (JJW) Frilford Tubney (JJW) (JJW) Tubney Wytham Tubney (JJW) (JJW) Wytham (JMCJSD) det 2000 Besselsleigh Sisymbrium officinale. officinale. Sisymbrium

vc22 N N N N N N N Local at sap, in fungi, etc. Local in atsap, Rare,Summertown atsap. (JEC) 1907 (JEC Wick 1907 Headington Copse (JEC) Wick 1907 Headington Copse crucifers Widespreadon (JJW W.Holland 1907) Shotover Comphrey On Wytham CowleySummertown(HSD) (JEC) 1907

vc23 Rare, at sap.Bagley Wood 1980s DrW.Wint 1980s Wood sap.Bagley at Rare, DrW.Wint 1980s Wood sap.Bagley at Rare, DrW.Wint 1980s Wood Bagley (JJW) Cothill

vc22 Nb Nb Nb N Under dead hardwood bark, also in compost and in large numbers in subterranean traps in gardens and woodlan gardens and in traps large subterranean in numbers in compost and in bark,also hardwood dead Under Ox canal D.Copestake 1984 L det LositoJMC AscottWychwood . in 1982 EN date Blenheimno Park

vc23 Hutchin's Copse 1998 (JSD) 1998 Copse Hutchin's (JJW) Rare?Cothill formerly Extinct (JEC) atTubney ants with wood 1907 Tubney 1907 (JJW) Tubney 1907

vc22 Nb Na Na RD1 Nb . Summertown . VCH SummertownVCH (JJW) Chawley broom scolytids in Predates 19.4.1907 EnslowBridge . WaterVCH Eaton

vc23 Tubney (HSD) pre 1907 Tubney (HSD)pre

vc22 Nb RD1 Na Local, breeds in nests of nests Local, in breeds Yarnton Bayswater Mill fungidecaying ?In and wood dead matter organic including Localhaystack usually in synanthropic refuse, hotbeds and grass clippings, decaying ?In matter.organic decaying and litter matter]fensriverorganic in banks and [Woods, ) Oxford Cellar(WH)in asaffinis Oxford (WH) (JJW) 1907 Yarnton (Copestake)Wychwood . 1989 Bombus

vc23 Drayton Copse 1998 (JSD) 1998 Copse Drayton (JJW) 1907 Park Wytham (JJW) 1907 Cothill stems reedmace lesser In

vc22 N N RDI RDK BicesterThame , (JJW) Shotover park (HBritten) Chilswell . Water(WH) Eaton Bicester, Oxford, Wolvercote (JJW)Sparsey Bridge(ETaylor) (JC) Yarnton I.McClenaghanWychwood 1989 Hardwick ] (CJ) Oxford (JJW) Oxford (JJW) Enslow(JC Gatehampton . TDHarrison CharlburyMDDarby Oxford, Reid) (C WaterperryWood 1980 of nests Local, in breeds Hardwick (Barnes) Bombus

vc23 Drayton Copse 1998 (JSD) Drayton1998 Copse Drayton Copse 1998 (JSD) (JSD) 1998 Copse Drayton (JJW) Tubney 1907

vc22 N N N N N RDK Nb Widespread in decaying Widespreadhay,in compost heaps. and vegetation including (HB) (AME) (JJW) OxfordShotover dist. Shillingford dung Rare?herbivore In ( decaying in vegetable matter decaying in vegetable matter farmyard compost and ?[In debris] farmyard compost and (JJW)?Oxford[In dist debris] most in frequent habitats. Widespreadand ?farmland? Oxford (JJW) (JC) Oxford (JJW)straw debris. stable ?In Water and Eaton (JSD) 2003 Stoke South RDBK ) Extinct? [In wetlands] Oxford dist. (JJW) Oxford (AAA) wetlands] dist. Play hatch [In Extinct? ) Weston(JC)

vc23 Cow Common 1998 (JSD) 1998 Cow Common

vc22 Na RDK RDK N . 1991 1991 . (SH) Hartslock 1991 Dorchester . (TDH) Gatehampton , 1995 Hartslock 1995

vc23 Drayton Copse 1998 (JSD) 1998 Copse Drayton

vc22 Nb Nb EXT Na Nb ?? JSD) det (JMC Alkerton . 1996

vc23 Rare on downs, and heaths, usually on the ground. Frilford 2002 JMCJSD det 2002 Frilford ground. the on usually heaths, and downs, on Rare etc. Tubney at Frilford frequent still dry ground open on puffballs in Tubney 1908

vc22 Nb RD2 Nb Nb Nb RD2 RD3 ↓ ↓ ) Oxford 1820 FWH Oxford ) 1820 FWH Oxford 1891 2010 in Oxon in 2010 1970s Porter K Otmoor

vc23 Hitchcopse Pit 1984 (Copestake 1984 Pit Hitchcopse

vc22 RD3 Nb RDK RDK Na WaterperryC.Reid 1980 ?Blenheim Park Swyncombe (TDH) 1991 Water C.Reid perry1980 Bagley Water C.Reid perry1980

vc23 Drayton Copse 1998 (JSD) 1998 Copse Drayton Bagley Hutchins

vc22 N ↓ Clattercote Resv. 1984 (DC) Clattercote Resv. 1984 Bladon. Summertown Islip (VCH) Water Eaton AstonRowant, Waterperry Godstow Wood (VCH) (DJM) 1998 Standlake ] OxfordUniversity Museum Clattercote Res. D Copestake 1984. 1984. D Copestake Res. Clattercote

vc23 Hutchins Copse, 1998 in red rotten oak with Lasius brunneus (JSD) oak with Lasius rotten red brunneus in 1998 Copse, Hutchins Tubney (WH) Hutchin's Copse 1998 (JSD) (JSD) 1998 Copse Hutchin's Tubney (WH) (VCH) Tubney,Wytham DrW.Wint 1980s Wood Wytham Survey 1962 Wytham

vc22 N Local on fungi and in fungoid hardwood, esp. on jew’s on Fungus esp. Ear hardwood, fungoid in fungi and Local on Auricularia auricula-judae auricula-judae Auricularia esp. on Elder on esp. Hartslock 1996 (TH) Hartslock 1996 (TH) Hartslock . 1996 N ofReading Thames (JSD) 2004 WaterVCH nr Eaton (SH) Hartslock 1991 1982 vc23 Drayton Copse 2011 (PMH) 2011 Copse Drayton

vc22 Nb Na Nb (KNAA) 1995 EstatesRctory Buscot JSD) farm det (JMC 2008 (VCH) Stow Wood Oxford . Water(VCH) Eaton

vc23 Drayton Drayton & Hutchin's Copse 1998 (JSD) 1998 Copse Hutchin's & Drayton

vc22 RD3 Nb Nb Nb Nb Na Nb Nb Na Nb Nb Na Nb Nb Hartslock 1995 (TdH) Hartslock1995 (M.Harvey) Bank8.7.1998 (JJW) WickUnhill Hedington 1907 . local on mugwort on local

vc23 t? [Sandy ground with Peltigera lichens] Hitchcopse Pit 1998 1998 Pit Hitchcopse lichens] with Peltigera ground [Sandy t? (JMCJSD) det 2002 Frilford (JMCJSD) det 2003 Frilford

vc22 Na Nb RDK RDK RDK RDK RDK RDK RDK Nb ↓ ↓ Oxford (WH) and in 1946 in Oxford and (WH) DJM (WH) Shotover OxfordMuseums JJW) (IN Eynsham . Mill Bone JJW houses Oxford in 1980 Oxford 1938 Caversham Warren Warren Caversham

vc23 Wytham Drayton Copse 1998 (JSD) (JSD) 1998 Copse Drayton Tubney (JSD) 1998 lane Hutchin's (JSD) 1998 lane Hutchin's

vc22 Nb Nb Nb RD3 Nb Nb Na ↓ Wheatley(Kitchen) 1969 (TH) Hartslock1998 (FWH) Hartslock ) ( Oxford . 1891 WychwoodCornbury (JJW) Oxford 1909 WaterVCH Eaton Hartslock Oxford9M.Gunning) nr 1882

vc23 )OxfordVCH Frilford Dry Sandford 1976 (PW) Hitchcopse 1996 (DJM) Lamb & Flag 2005 (JMC) 2005 Flag & (DJM) 1996 Lamb Hitchcopse (PW) 1976 Dry Sandford Frilford (JSD) 1998 Copse Drayton sands Corallian on common Still

vc22 Nb EXT RD1 Nb RD3 Nb Nb Nb RD2 RD3 Nb Watlington 1980 (BRC) 1980 Watlington (2010) telnov in Oxon (HSD) Oxford 1935 Cowley (OWR) 1924 Summertown (JJW) ,1986) C(Robinson Oxford 15th Priory (KNAA Grey1986 & AF) s Court

vc23 common on Corallian in Vc22 in Corallian on common (JSD) 1998 Copse Drayton

vc22 RD2 Nb Nb Nb Nb Na Na vc23

vc22 Na RD3 RD3 Na Na . Headington Wick 1907 (JJW) Wick Headington . 1907 Rare onlyfrom (C.Raper) modern record Hartslock 1999 Wychwood Forest Summertown . (JMC) Stoke South . (PSH) Wychwood 1985 (JJW) 25.6.1905 Elsfield rd to Marston Caversham Warren (H) Buckland Warren 1988 (MVC) 1988 Warren Buckland

vc23 Drayton Copse 1998 (JSD) Drayton1998 Copse Bagley, Boars Hill (JJW) (JJW) Hill Boars Bagley, (JSD) 1998 Cow Common

vc22 Nb Nb RD3 Nb Nb Nb Na RD2 Na ↓ ↓ Kings Weir on the Thames 1915 (WS). Marlborough Pool 1985 (JMC) 1985 Pool Thames (WS).the Marlborough Kings Weir on 1915 1938 Mapledurham Henleyfrom Thames Sharpe1908 Gosford&by 1913 Cherwell 1910 (MVB) AstonRowant 2000 Wychwood Wood Marston (KNAA) park1986 Wychwood Cornbury (TDH) Widley1991 Copse (FWH) Oxford1819

vc23 (RDB1) Last Uk record Cothill in 1923 in Last (RDB1) Cothill Uk record (JSD) Drayton1998 Copse

vc22 Nb RD2 Na RD3 RD1 Nb Nb RDK RD1 Spartum fen JMC (CMD) Otmoor 2009 Lock 1987 JMC Pinkhill Yarnton fromrecords ld Cherwell Oxford in (JMC) 1985 Pool Marlborough

vc23 Tubney 2.6.1901 (Hope coll.) its host the wood ant has disappeared from disappeared has area.ant the wood the host its coll.) (Hope Tubney2.6.1901 (M.Corley) 1983 Cothill reeds rareon

vc22 RD1 Na Nb Nb Na Na Na Nb Nb ↓ Local on sheep’s sorrel sorrel sheep’s Localon Rumexacetosella on sandy heathland. Also on Also sandyon heathland. on Lotus corniculatus, corniculatus, Lotus esp. on the chalk the on esp. Stones field 1990 (M.Gascoigne-Pees) field1990 Stones JD) det (JMC 2008 Pits Shepherds (S.Lane) 1992 (FWH) Rollright Oxford 1819 Oxford parksUniv 1980 ] (D.Murray)Warburg . Reserve 1984 Wychwood . Woodstock 1905 Yarnton

vc23 Cholsey Downs 1927 & 1955 (BM HobbY (BM & 1955 CholseyDowns 1927

vc22 Na Nb RD1 Nb RD1 Na Nb Nb RD2 Nb RD1 ↓ ↓ Last seen in 1946 atBrazenose Wood Last1946 in seen

vc23 Frilford 1952 nd 1987 nd 1952 Frilford

vc22 Nb RD1 Nb Local on devil’s-bit cabious in damp grassland and rides, and the ‘breck dryform/sp.?’the yarrow and soils damp rides, on disturbed grassland in and on cabious devil’s-bit Localon Ascott in Wychwood 1982 JMC MLC det 1982 Wychwood in Ascott JMC MC det 1998 Minster Lovell M>Cox) det JMC 1995 Burford . Brazenose Shiplake Oxford(JJW)Dist Wheatley

vc23 Cothill 1986 (MCorley) 1986 Cothill

vc22 Na Nb Nb Na Nb RD1 ↓ R.Thames, Shillingford 1946 (AME) R.Thames, 1946 Shillingford ChimneyMC t de meadowsJMC 1991 EnslowQuarry SUB in .Thame 15 th C SmithC (1999)

vc23 Wytham(J.Collins) 1912 ● Hitchcopse 1913 & 1996 & 1996 1913 Hitchcopse

vc22 Na RD1 Nb Na Nb Nb Nb Na RD1 Nb Na ↓ Finstock VCH Cherwell 1984 (DC) Cherwell 1984 VCH Finstock VCH Hardwick ANHS SparseyBridge1929 Beckley(WH) Blenheim Ewelme Summertown ANHS Radley 1906 1944 Cowley1904 Bullingdon Bog (WH) (WH) Bog Bullingdon

vc23 Tubney 1906 Tubney 1906 2008 Glebe auriculata. waterScrophularia figwort on Rare Witt 1980s Wood Wytham

vc22 Nb Na Nb Na Na [ Nb Nb Nb Na ↓ ↓] Swyncombe Downs 1989 (PSH) Swyncombe (PSH) Downs 1989 Wychwood

vc23 Cothill

vc22 Nb Nb Nb Nb Nb Nb Bred from Bred coll.) (Hope larvae taken vii.1899 atChilswell in GEW Wychwood1965

vc23 on pine trees Tubney 1907 (JJW) (JJW) Tubney 1907 trees pine on Tubney Woods Wytham

vc22 Na Nb Nb Na Na RDK RD1 Nb ↓ C.Reid Waterperry 1980 Wood Oxdist Waterperry Wood 1980 C.Reid Waterperry1980 Wood

vc23 Bagley1922-7 Wood Rare,wild increasing.mignonette On Hutchin's Copse 1998 (JSD) (JSD) 1998 Copse Hutchin's (JSD) 1998 Cow Common Reseda lutea. lutea. Reseda vc22 Nb Nb Nb RD3 Nb Na Nb Na Nb Nb Nb East Downs 2008 (JMC det JD) det (JMC EastDowns 2008 . Otmoor 1998 JMC det JSD det JMC Otmoor 1998 . (DJM) 1998 Standlake S.Lane det 1998 JMC Wootton (DC) Alkerton1994 Shotover 1907 (THB) 1907 Shotover (WH) St.John Stanton

vc23 Tubney 1903 (W.Holland 1905 (HSD) 1905 (W.Holland Tubney1903 JSD det JMC Tadmarton 2001 Frilford JMCJSD det Frilford

vc22 Nb Na RD3 Nb RD1 Na RD3 Roman 1975) , Alchester (Giorgi &Robinson ?? (GLW) HytheBablock 1953

vc23 ? [On ?[On . Tubney 1907 Tubney. 1907 JMCJSD det Frilford Sainfoin on (JEC) 1906 Cumnor (JSD) 1998 Copse Hutchin's Trifolium especially small yellow flowered Tubney species] (JJW)

vc22 Nb Nb Na Na ↓ . Shotover 1970s. Shotover . (VCH) Norton Chipping Stonesfield, (WH) 1907 Wood Hen (TDH) 1990 Shotover

vc23 Na Abingdon common 2004 (JMCJSD) det 2004 common Abingdon (JJW) Tubney 1907 (JSD) 2008 Lane Hutchins (JSD) 2008 Lane Hutchins ) ? [On hares-foot clover] Tubney (JJW) Frilford 1952 Tubney hares-footclover] (JJW) 1952 ?[On ) Frilford

vc22 Nb Nb Na Nb Na Nb Nb Nb Widespread on figworts Widespreadon Scrophularia nodosa Scrophularia and and S . auriculata . Also on Mulleins including including Mulleins Also . on V.blattaria Pucketty farm MvC Kidlington .

vc23 Cothill 1906 (JJW) 1906 Cothill Tubney 1907 (JJW) Tubney 1907

vc22 Na Nb RDK Nb Nb Nb Nb Rare on breeds Campanula esp. esp. C. glomerata C. ] on the downs, the on ]

[harebell [harebell C. rotundifolia C. ], Adults also atyellow Adults also ], composites Islip, Marston Ferry, Marston Summertown Islip, (JJW) (GLF) HytheBablock 1953 Swyncombe(PSH) Downs 1989 (JJW) Yarnton r (JEC) Islip Park Prattlewood Bagley (JJW) Wood

vc23 Cothill & Tubney 1907 (JJW) &Tubney 1907 Cothill Cothill, Cumnor, Tubney (VCH), Cumnor, Cothill, (JJW) 1907 Cothill

vc22 Na Na Na Nb Nb Nb RDK Nb Nb Nb Na Nb Hartslock 1994 (TH) Hartslock1994 ,1993) (Robinson EarlyDitch Mingies E.Holocene,

vc23

vc22 Nb Nb Na Nb Nb Nb RD2 RDK Na ↓ . Summertown, . Ferry Marston (VCH) . ,1993) (Robinson Early Ditch . Mingies E.Hoocene, ,1993) (Robinson Early Ditch Mingies E.Hoocene, (MVB) Aston Rowant 2000 hornworts On (JJW) 1907 & Kennington Yarnton . Aston Rowant 1999 (JMC det JSD) (JMCJSD) Rowant1999 det Aston Aston Rowant 2000 (MVB) (MVB) AstonRowant 2000 Ceratophyllum sp.] sp.]

vc23 Tubney (JEC) Tubney (JEC) (JJW) Cothill (DC) 1984 Hitchopse Hollier A J Wytham

vc22 Nb Na Nb Na Nb Nb Nb Nb Na RD3 Na Nb Shotover 1918 (JEC) (JEC) 1918 Shotover (MM) Aston Rowant 1971 Rare.1970 (TDH) 1990 Shotover Elsfield (JJW) Sydlings1938 Copse brassicas] On

vc23 Tubney 1907 (JJW) Hitchcopse Pit Glebe (JJW) Pit TubneyHitchcopse 1907 Wytham Survey 1954? Wytham etc Frilford

vc22 Nb RD2 RDK Nb Nb Nb RDK RD1 Nb Nb Nb Bablcock Hythe 1953 (GLF) Bablcock Hythe 1953 JD) det (JMC Taynton 2008 Godstow (JJW) Water1906 Eaton 1965 (JEC) Water1907 Eaton JSD det JMC 1997 RMCS . (TDH) 1990 Shotover Roman1975) , Alchester (Giorgi &Robinson (DJM) (GLF) HytheBablock 1998 Standlake 1953

vc23 Tubney etc Hitchcopse 1995-2004 (JMC) 1995-2004 Hitchcopse Tubney etc (JJW) 1907 Cothill (JJW) park Wytham Tubney (JEC) Wytham(VCH )Cothill, , Wood Bagly

vc22 Nb Nb Na Na Nb Nb Nb Na Nb Nb Na Nb Nb Nb Naturalised. Widespread and frequent. Immigrant from New Zealand, found in rotting wood and building timber. Immigrant frequent. Widespread and Naturalised. building and wood from rotting in New found Zealand, Wychwood 1985 (PSH) (PSH) Wychwood 1985 JSD) det JMC Common Nettlebed . 1996 (JJW) Ferry Marston 1907 1980 (PMH) WarburgNR new 6.6.2004 for Britain Yarnton Ferry Marston 1906 (DS) 1996 Park Cornbury

vc23 Frilford Hitchcopse (JMC) Hitchcopse Frilford

vc22 Nb Nb Nb Nb Nb Nb Nb Nb Na ↓ Nettlebed (JMC det JSD det (JMC Nettlebed (FWH) Oxford . 1820 Bablock Hythe 1953 (GLF) no specimen?? no (GLF) 1953 Hythe Bablock

vc23 rare low open ground. Hatford (SP3395) 7-9.2005 (JMC det JSD) det (JMC 7-9.2005 Hatford (SP3395) ground. rarelow open Tubney (JJW) Tubney (JJW)

vc22 Nb Nb Nb Nb Nb Nb Peppard, Hardwick Goring (VCH) and Peppard, (JEC) 1906 Eaton Wood

vc23

vc22 Nb RD2 Nb Nb Nb Na Oxford 1820 (FWH) <1938 Sydlings Copse Oxford 1820 Ogley . Bog JSD det ChimneyJMC 1991

vc23 Frilford 2000 JMC det JSD det JMC 2000 Frilford Wytham park Hitchcopse Hitchcopse park Wytham Corallian on common still etc Frilford Tubney (JJW)

vc22 Nb Nb Nb Nb Nb Nb Nb Nb Widespread on various cloversvarious Widespreadon Trifolium spp. and medicks and spp. M edicago spp. in grassland including ruderal sites. ruderal grassland including in spp. . Ambrosden 2012 (JSD) Ambrosden . 2012 JSD) det (JMC Otmoor 1998

vc23 Tubney,(JJW) Hilll Boars 2008 Frilford Cow Common 1998 (JSD) 1998 Cow Common (JSD) 1998 Copse Drayton

vc22 Nb Nb Na Nb Nb RD1 Nb Na ↓ Hartslock 1997 (DC) Hartslock 1997 (C.Reid) Waterperry 1980 Wood JSD) det (JMC Lovell Minster 2001

vc23 Boars Hill (WH) Hill Boars bark,Tubney spruce and pine in Marcham Park 2011 (JSD) 2011 Park Marcham (JSD) 1998 Copse Drayton (VCH) Tubney Wytham

vc22 Nb Nb Na Na RD1 Nb Otmoor 2000 (JSD) Otmoor 2000 JSD det JMC Wychwood 1992

vc23 Tubney 1907 (AHH) Tubney 1907 (JSD) 1998 Copse Drayton

vc22 Nb Nb vc23

vc22