Subfossil Vc22 VCH Vc23 1939-2012 VC22 modern Oxon ecology 1 Suborder MYXOPHAGA 2 3 SPHAERIUSIDAE 4 SPHAERIIDAE 5 MICROSPORIDAE 6 Sphaerius acaroides Waltl ● ● 7 8 Suborder ADEPHAGA 10 GYRINIDAE 11 GYRININAE 12 Gyrinus aeratus Stephens ● ● Canals and slow flowing river margins 16 Gyrinus marinus Gyllenhal ● ● ● ● Local, on open water especially on large water bodies such as lakes and canal margins. 22 Gyrinus substriatus Stephens ● ● ● ● Widespread and often abundant on ponds, canals, ditches and lakes. 23 Gyrinus suffriani Scriba ● ● Rare in fens 24 Gyrinus urinator Illiger ●1998 ● Rare in flowing water margins. 25 Orectochilus villosus (Müller) ● ● ● ● rare, on streams often in deep shade. Largely nocturnal and easily overlooked.B 26 27 HALIPLIDAE 28 Brychius elevatus (Panzer) ● ● ● ● rare in flowing water especially in submerged gravel and near weirs 29 Peltodytes caesus (Duftschmid) ● ● ● Local but increasing? Weedy ponds, 30 Haliplus confinis Stephens ● ● ● ● Local in ponds, lakes and canals with Charophytes. 31 Haliplus obliquus (Fabricius) ● ● ● ● Local, but can be a rapid coloniser of new ponds with Chara 33 Haliplus lineatocollis (Marsham) ● ● ● ● Widespread in streams, ditches and weedy ponds. 35 Haliplus fluviatilis Aubé ● ● ● Widespread in ponds, lakes, canals etc. 36 Haliplus furcatus Seidlitz ● ● Rare in fen pond 37 Haliplus heydeni Wehncke ● ● Very Local in small detritus ponds 38 Haliplus immaculatus Gerhardt ● ● ● ● Widespread in weedy ponds. 39 Haliplus lineolatus Mannerheim ● ● ● Rare? [Canals, ponds, flooded gravel pits]. Subfossil Vc22 VCH Vc23 1939-2012 VC22 modern Oxon ecology Haliplus mucronatus Stephens Na ● ● Rare one record 40 Haliplus ruficollis (De Geer) ● ● ● Widespread in detritus ponds. 41 Haliplus sibiricus Motschulsky ● ● ● Local in standing water. 42 (= wehnckei ) Local in standing water. 43 Haliplus flavicollis Sturm ● ● ● ● in large ponds, ditches, canals. 44 Haliplus fulvus (Fabricius) ● ● ● rare possibly extinct 45 Haliplus laminatus (Schaller) ● ● ● Weedy streams and ponds. 47 Haliplus variegatus Sturm ● ● extinct 48 49 NOTERIDAE 50 Noterus clavicornis (De Geer) ● ● ● ● Widespread and common in ponds, canals, and lakes. 54 55 PAELOBIIDAE 56 HYGROBIIDAE 57 Hygrobia hermanni (Fabricius) ● ● ● ● Widespread in ponds esp. with silty substrates. 59 60 DYTISCIDAE 61 AGABINAE 64 Agabus sturmii (Gyllenhal in Schönherr)● ● ● ● Widespread in detritus ponds. 65 Agabus labiatus (Brahm) ● ● 66 Agabus uliginosus (Linnaeus) ● ● Rare in mesotrophic pools 69 Agabus biguttatus (Olivier) ● ● Rare.Calcareous ground fed waters, spring heads 70 Agabus bipustulatus (Linnaeus) E.Ho● ● ● ● Ubiquitous in stagnant water including horse troughs etc., a strong flyer. 73 Agabus didymus (Olivier) ● ● ● ● Local in flowing water amongst marginal vegetation. 74 Agabus guttatus (Paykull) ● ● ● ● Local in spring fed streams and well-heads, also outfalls of man-made drains. 77 Agabus nebulosus (Forster) ● ● ● ● Widespread. In silt ponds esecially. New or recently cleared sites, a strong flier also frequent in troughs etc. 78 Agabus paludosus (Fabricius) ● ● ● ● Local, in weedy margins of flowing water, also around small seepages and springheads in flooded litter and moss. 82 Ilybius ater (De Geer) ● ● ● ● Widespread in all kinds of standing waters. Also comes to light. 83 Ilybius chalconatus (Panzer) ● ● ●1998● Local. In ponds and rut pools in wooded areas, Subfossil Vc22 VCH Vc23 1939-2012 VC22 modern Oxon ecology 84 Ilybius fenestratus (Fabricius) ● ● ● ● Very Local. In large ponds, lakes especially in heathland areas. 85 Ilybius fuliginosus (Fabricius) ● ● ● ● Widespread in ponds and weedy margins of flowing water. Also comes to light. 87 Ilybius montanus (Stephens) ● ● ● Localespecially in acid waters 89 Ilybius quadriguttatus (Lacordaire) ● ●2005 ● Local in detritus ponds. 92 Platambus maculatus (Linnaeus)E.Ho ● ● ● Local in flowing water, streams, and rivers. Tolerant of shade. 93 94 COLYMBETINAE 95 Colymbetes fuscus (Linnaeus) E.Ho● ● ● ● Frequent species of ponds of all kinds. 99 Rhantus exsoletus (Forster) ● ● ● ● Local in detritus ponds and marshy areas. 101 Rhantus suturalis (MacLeay) ● ● ● ● Widespread good coloniser of new and recently cleared silt pools. 104 105 COPELATINAE 106 Liopterus haemorrhoidalis (Fabricius)● ● ● ● Local. In weedy detritus ponds and bog pools. 107 (= Copelatus ) 108 109 DYTISCINAE 111 Acilius sulcatus (Linnaeus) ● ● ● ● Local in a wide variety of still waters, but intolerant of fish stocking. 117 Dytiscus circumflexus Fabricius ● ● ● Scarce, increasing? In ponds with open silty substrates. 120 Dytiscus marginalis Linnaeus ● ● ● ● Ubiquitous in standing water of all kinds, especially where amphibians are breeding, as larvae specialise in tadpoles. 121 Dytiscus semisulcatus Müller, O.F. ● ● ●2011● Rare, most records from stagnant water in heath ponds and peat pools. 124 Hydaticus seminiger (De Geer) ●2009●2011● recnt colonist increasingly found n rich pools in shade and open situations 126 127 HYDROPORINAE 130 Hydroglyphus geminus (Fabricius) ● ● ● ● Rare, most records from stagnant water in heath ponds and peat pools. 135 Graptodytes granularis (Linnaeus) ● ● ● ● Rare. Richly weeded ponds 136 Graptodytes pictus (Fabricius) ● ● ● ● very locally abundant in ponds and ditches. 137 Hydroporus angustatus Sturm ● ● ● Widespread, and common in ponds, canals, and lakes, avoiding acidic waters 138 Hydroporus discretus Fairmaire & Brisout● ● ● ● Very Local in spring fed puddles and pools. 140 Hydroporus erythrocephalus (Linnaeus)● ● ● ● Widespread and common in ponds and lakes, especially in acidic heath ponds. Subfossil Vc22 VCH Vc23 1939-2012 VC22 modern Oxon ecology 141 Hydroporus ferrugineus Stephens ● ● rare in Subterranean spring heads 143 Hydroporus gyllenhalii Schiødte ● ● RARE in acidic heath ponds. 145 Hydroporus incognitus Sharp ● ● ● ● Local in shaded pools and marshy pools beside streams. 148 Hydroporus marginatus (Duftschmid)● ● ● Semi subterranean in winterbourne streams. 150 Hydroporus memnonius Nicolai ● ● ● ● Widespread in woodland pools in flooded leaf litter. Hydroporus neglectus Schaum Nb ● ● Shaded acis waters and seepages 156 Hydroporus nigrita (Fabricius) ●1996●1998● Local in a variety of small water bodies including small seasonal pools. Hydroporus obscurus Sturm ●1906● [ in Sphagnum pools] 159 Hydroporus palustris (Linnaeus) ● ● ● ● Widespread in detritus ponds and marshy areas by lakes and flowing water. 160 Hydroporus planus (Fabricius) ● ● ● ● Widespread rapid coloniser of new and recently cleared ponds also in water troughs. 161 Hydroporus pubescens (Gyllenhal) ● ● ● ● Widespread in acid standing water. 164 Hydroporus striola (Gyllenhal) ● ● ● ● Local in standing water. 165 Hydroporus tessellatus (Drapiez) ● ● ● ● Widespread. Rapid coloniser of new and recently cleared ponds also in water troughs, isolated puddles. Hydroporus umbrosus (Gyllenhal) ● ● rare in mossy fen pools 170 Nebrioporus elegans (Panzer) ● ● ● ● Local in rivers and permanent streams. Also occasionally in deep ponds in pits etc. 175 Oreodytes sanmarkii (Sahlberg, C.R.)● ● Rare. Gravelly streams in root tangles, shade tolerant. 178 Porhydrus lineatus (Fabricius) ● ●1998 ● Rare. Ponds, ditches etc.] 179 Scarodytes halensis (Fabricius) ● ● Local, increasing 180 Stictonectes lepidus (Olivier) ● ● Extinct?? 181 Stictotarsus duodecimpustulatus E.Ho(Fabricius)● ● ● ● Local in rivers and streams especially in bare sections under submerged ledges etc. 184 Suphrodytes dorsalis sensu lato ● ● ● ● recently split status under review 187 Hygrotus (Coelambus) confluens (Fabricius)● ● ● ● Widespread a rapid coloniser of new ponds with silty substrates. 188 Hygrotus (Coelambus) impressopunctatus● (Schaller)● ● ● Widespread in field ponds, Hygrotus (Coelambus) nigrolineatus (Steven)● ● Recent immigrant increasing in new ponds. 194 Hygrotus (s.str.) inaequalis (Fabricius)H/Rom● ● ● ● Widespread and common in ponds, canals, and lakes. 196 Hygrotus (s.str.) versicolor (Schaller) ● ● ● ● Local in gravel ponds, canal margins. 197 Hyphydrus ovatus (Linnaeus) ● ● ● ● Widespread and common in ponds, canals, and lakes. 199 200 LACCOPHILINAE Subfossil Vc22 VCH Vc23 1939-2012 VC22 modern Oxon ecology 201 Laccophilus hyalinus (De Geer) ● ● ● ● Local, in slow flowing water, occasionally in ponds, 202 Laccophilus minutus (Linnaeus) ● ● ● ● Widespread and common in ponds, canals, and lakes. 205 206 CARABIDAE 207 CICINDELINAE 209 Cicindela campestris Linnaeus ● ● ● ● Local on open dry soils, scattered in woodlands and disturbed ground. 215 216 BRACHININAE 217 Brachinus crepitans (Linnaeus) ● ●2003 ● Rare on chalky soils, no post 1980 records, possibly extinct? 219 223 CARABINAE 224 CARABINI 225 Calosoma inquisitor (Linnaeus) ● ●1953● ● Extinct? [Arboreal on oaks etc.] 229 Carabus arvensis Herbst ? ? extinct Specimen labelled Oxford in JJW coll. 230 Carabus granulatus Linnaeus E.Hol● ● ● ● Local in damp meadows, 10 post 1980 records. 231 Carabus monilis Fabricius ● ● ● ● Widespread in open habitats. 232 Carabus nemoralis Müller, O.F. Rom● ● ● ● local in dry areas 236 Carabus problematicus Herbst ● ● ● ● scarce primarily on calcareous grassland in Oxon. 238 Carabus violaceus Linnaeus ● ● ● ● Widespread and common occurring in woods, parks, gardens, and more open habitats. 239 CYCHRINI 240 Cychrus caraboides (Linnaeus) ● ● ● ● Widespread in wooded districts. A flightless snail predator. 241 NEBRIINI 243 Leistus rufomarginatus (Duftschmid) B215● ● ● 20th century colonist (first noted in the 1940s), now widespread especially in woods