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3 (1) • September 2012: 21-29

Review Article

Checklist of the Bracon (Hymenoptera: ) in Serbia Vladimir Žikić1*, Saša S. Stanković1, Marijana Ilić1

1Faculty of Science and Mathematics, Department of Biology and Ecology, University of Niš, Višegradska 33, 18000 Niš, Serbia

* E-mail: [email protected]

Abstract: Žikić, V., Stanković, S., Ilić, M.: Checklist of the genus Bracon (Hymenoptera: Braconidae) in Serbia, Biologica Nyssana, 3 (1), September 2012: 21-29.

This is the first checklist of the genus Bracon for the territory of Serbia. Here we present 80 species of this cosmopolitan genus of the subfamily Braconinae which is one of the cyclostome braconid groups. We connected the parasitoids with their hosts where the data about them were available. There are 290 host species in total belonging to 39 families of four orders: Diptera, Coleoptera, Hymenoptera and .

Key words: Cyclostome Braconidae, Bracon, Serbia, checklist

Introduction habitats and geographical areas (S h a w & H u d d l e s t o n , 1991). Because of their important Braconinae belongs to the cyclostome group role in biological control, biology of some species is of the family Braconidae. This subfamily assembles very well studied: for B. intercessor (van ectoparasitoid wasps which lay their eggs on the A c h t e r b e r g et al., 1990; G e o r g i e v et al., host body. According to their biology, ectoparasitic 2005), for B. brevicornis (T e m e r a k , 1982), or braconid wasps belong to the group of idiobionts; many publications for B. hebetor and its host Plodia which means that they do not permit the host to interpunctella (Pyralidae) (R e i n e r t et al., 1971; continue its development after oviposition T a y l o r , 1988a, b; Yu et al., 1999; M i l o n a s , (M a t h e w s , 1974). Oviposition in this group is 2005; D w e c k et al. 2010). According to the Fauna split in two acts: paralisation of the host, than laying Europaea (v a n A c h t e r b e r g , 2012), about 250 eggs which is opposite in koinobions. As a species of the genus Bracon are recorded in . cosmopolitan group of parasitoids they represent a The present publication stemmed from the need to large and powerful biological weapon (eg. B a i r d , make the list of all registered Bracon species on the 1958; L e w i s et al., 1990) against various pests territory of Serbia after many years of investigation from the following holometabolous insect orders: and also to connect the parasitoids with their hosts. Diptera, Coleoptera, Hymenoptera and Lepidoptera. Investigation of the genus Bracon in Serbia in The species of the genus Bracon show different different aspects is presented in following degree of the host specificity, ranging from a strict publications by the authors Belokobyskij & specialization to only one host species, whereas Ž i k i ć (2009); B r a j k o v i ć (1989); B r a j k o v i ć other can be polyphagous, when parasitism occurs et al. (1991, 1994); S t a n k o v i ć et al. (2010) and in more than a hundred hosts in different types of Ž i k i ć et al. (2000, 2010, 2012).

21 BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)…

Materials and methods Curculio villosus, Mononychus punctumalbum (Curculionidae); Tetramesa brevicornis T. The majority of analyzed specimens were hyalipennis (Eurytomidae); coronillae taken from the collection of the Faculty of Biology, (); Paranthrene tabaniformis University of Belgrade and the Faculty of Sciences (). Distribution: , , Great and Mathematics, University of Niš, Serbia. A part Britain, Bulgaria, Croatia, , Finland, , of specimens is deposited at the Zoological Institute , , , Ireland, , Latvia, RAS (St. Petersburg, Russia) and The Natural Lithuania, , Russia (Central and South), History Museum of Budapest, Hungary. The data , , , Sweden, Switzerland, about braconid hosts are compiled using the Netherlands, Serbia. available literature; the list of references is bellow. Bracon (Bracon) grandiceps Thomson 1892 on The basic key for the identification of the species we scopigera (Sesiidae). Distribution: used is one by Tobias et al. (1986), updated Fauna Austria, Croatia, Turkey, Finland, France, Germany, Europaea (http://faunaeur.org) and the database Greece, Hungary, Italy, Macedonia, Poland, Russia Taxapad (Yu et al. 2012). A lot of information (South), Slovenia, Sweden, Serbia. about the hosts and the distribution of the investigated genus we collected from P a p p (1973, Bracon (Bracon) intercessor Nees 1834 on Apion 1977, 2008) and B e y a r s l a n et al. (2005). opeticum (Apionidae); Agapanthia villosoviridescens (Cerambycidae); Anthonomus Results pedicularius, A. pomorum, A. sorbi, Lixus brevirostris, L. incanescens, L. junci, L. scabricollis,

Microlarinus lareynii, M. lypriformis We present 80 species of the genus Bracon (Curculionidae); Rhynchites bacchus registered on the territory of Serbia. The data (Rhynchitidae); Tetramesa hyalipennis, T. rossica regarding parasitoids are connected with the (Eurytomidae); Haplochrois theae (Agonoxenidae); adequate data about their hosts. The species of Augasma atraphaxidellum (Coleophoridae); Bracon are listed alphabetically according to their Synanthedon culiciformis, Paranthrene tabaniformis subgenera. For 25 braconid species there were no (Sesiidae); Sparganothis pilleriana (). host records for Serbia nor for other parts of Europe. Distribution: Albania, Austria, Great Britain, In addition, we give the geographical distribution of Bulgaria, Croatia, Cyprus, Turkey, France, all registered parasitoids in continental part of Germany, Greece, Hungary, Italy, Latvia, Lithuania, Europe and Russia. Macedonia, Norway, Poland, , Russia (Central, East, Northwestern and South), Slovakia, Bracon (Bracon) alutaceus Szepligeti 1901 – no Slovenia, Spain, Sweden, Switzerland, Netherlands, host data. Distribution: Austria, Great Britain, , Serbia. Bulgaria, Cyprus, , Denmark, Germany, Hungary, Lithuania, Poland, Russia Bracon (Bracon) laetus Wesmael 1838 on (Northwestern), Slovakia, Spain, Sweden, Ukraine, Anthonomus pomorum, Magdalis nitida Serbia. (Curculionidae); Amblypalpis tamaricella (); Argyresthia chrysidella, Prays citri; Bracon (Bracon) crassiceps Thomson 1892 – no Prays oleae (Yponomeutidae). Distribution: host data. Distribution: Austria, Finland, Germany, Belgium, Great Britain, Bulgaria, Croatia, Cyprus, Hungary, Russia (Northwestern), Sweden, Czech Republic, France, Germany, Greece, Switzerland, Serbia. Hungary, Spain, Serbia. Bracon (Bracon) crassungula Thomson 1892 – no Bracon (Bracon) leptus Marshall 1897 on host data. Distribution: Austria, Hungary, Italy, lapella (Gelechiidae). Distribution: Sweden, Switzerland, Serbia. Austria, Great Britain, Bulgaria, Croatia, Cyprus, Bracon (Bracon) filicornis Thomson 1892 – no Germany, Greece, Hungary, Lithuania, Macedonia, host data. Distribution: Austria, Sweden, Serbia. Romania, Russia (Central and South), Slovakia, Bracon (Bracon) flagellaris Thomson 1892 – no Slovenia, Spain, Switzerland, Netherlands, Serbia. host data. Distribution: Croatia, Germany, Hungary, Bracon (Bracon) longicollis Wesmael 1838 on Romania, Russia (South), Slovenia, Sweden, Chlorops pumilionis (Chloropidae). Distribution: Switzerland, Ukraine, Serbia. Austria, Belgium, Great Britain, Bulgaria, Croatia, Bracon (Bracon) fulvipes Nees 1834 on Apion Cyprus, Turkey, Finland, France, Germany, Greece, minimum, Cistapion cyanescens (Apionidae); Hungary, Italy, Lithuania, Macedonia, Norway, Poland, Russia (Central, Northwestern and South),

22 BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)…

Slovenia, Spain, Sweden, Switzerland, Netherlands, Distribution: Belgium, Great Britain, Finland, Serbia. France, Hungary, Ireland, Italy, Poland, Sweden, Bracon (Bracon) longulus Thomson 1892 on Netherlands, Serbia. Oscinella frit (Chloropidae). Distribution: Austria, Bracon (Bracon) speerschneideri Schmiedeknecht Bulgaria, Croatia, Cyprus, Czech Republic, 1897 on Argyresthia thuiella, Ocnerostoma sp. Denmark, Finland, France, Germany, Greece, (Yponomeutidae). Distribution: Austria, Great Hungary, Italy, Macedonia, Norway, Slovenia, Britain, France, Germany, Norway, Sweden, Spain, Sweden, Netherlands, Serbia. Switzerland, Serbia. Bracon (Bracon) luteator Spinola 1808 on Bracon (Bracon) subglaber Szepligeti 1901 on Urophora solstitialis (Tephritidae); Metzneria Noeeta pupillata, Tephritis leontodontis, Terellia aestivella, M. lappella (Gelechiidae). Distribution: serratulae, Urophora cardui, U. quadrifasciata, U Austria, Great Britain, Bulgaria, Croatia, Cyprus, solstitialis (Tephritidae); Metzneria lappella Turkey, France, Greece, Hungary, Italy, Latvia, (Gelechidae); Epichnopterix sieboldi (Psychidae); Macedonia, Russia (Central, Northwestern and Paranthrene tabaniformis, hylaeiformis South), Slovakia, Spain, Switzerland, Ukraine, (Sesiidae); Aethes williana (Tortricidae). Serbia. Distribution: Croatia; Germany; Hungary; Italy, Bracon (Bracon) mariae Dalla Torre 1898 on Kazakhstan, Moldova, Russia (Russia-Krasnodar, Chaetorellia jaceae (Tephritidae). Distribution: Yaroslavl Oblast), Slovenia, Switzerland, Ukraine, Bulgaria, Croatia, Germany, Hungary, Italy, Russia Serbia. (South), Slovakia, Spain, Sweden, Switzerland, Bracon (Bracon) trucidator Marshall 1888 on Ukraine, Serbia. Metzneria lappella (Gelechiidae). Distribution: Bracon (Bracon) mediostriatus Papp 1968 – no Albania, Austria, Belgium, Great Britain, Bulgaria, host data. Distribution: Hungary, Serbia. Croatia, Denmark, Turkey, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Poland, Bracon (Bracon) nigratus Wesmael 1838 on Romania, Russia (Central, Northwestern and South), Chaetostomella cylindrica (Tephritidae); Slovenia, Spain, Switzerland, Netherlands, Serbia. Coleophora millefolii (Coleophoridae); Grapholita compositella (Tortricidae); Zygaena loti Bracon (Bracon) variegator Spinola 1808 on (Zygaenidae). Distribution: Austria, Belgium, Great Ernobius abietis (Anobiidae); Coleophora Britain, Croatia, Czech Republic, France, Germany, iutipennella (Coleophoridae); Agonopterix Hungary, Ireland, Italy, Lithuania, Romania, Russia propinquella, (Depressariidae); Anarsia lineatella, (Central and North), Slovakia, Slovenia, Spain, Recurvaria nanella, Teleiodes luculella Sweden, Switzerland, Netherlands, Serbia. (Gelechiidae); Phyllonorycter klemannella, Ph. mespillela (Gracillariidae); Endrosis sarcitrella Bracon (Bracon) pallicarpus Thomson 1892 – no (Oecophoridae); Acrobasis obsutella (Pyralidae); host data. Distribution: Great Britain, Bulgaria, Archips rosana, Cydia strobilella, Tortrix viridana Finland, Germany, Hungary, Italy, Spain, Sweden, (Tortricidae); Yponomeuta padella Serbia. (Yponomeutidae). Distribution: Albania, Austria, Bracon (Bracon) pectoralis Wesmael 1838 on Croatia, Cyprus, France, Hungary, Italy, Macedonia, Chrysobothis affisinis, Sphenoptera gossypii, S. Russia (South), , Spain, Switzerland, lobicollis, S. montana (Buprestidae); Plagionotus Ukraine, Serbia. arcuatus, Plagionotus bobelayei (Cerambycidae). Bracon (Cyanopterobracon) sabulosus Szepligeti Distribution: Austria, Belgium, Great Britain, 1896 on Tephritidae. Distribution: Bulgaria, Croatia, Bulgaria, Croatia, Cyprus, Turkey, France, Cyprus, Greece, Hungary, Macedonia, Romania, Germany, Greece, Hungary, Italy, Macedonia, Russia (South), Serbia. Romania, Russia (Central and South), Slovakia, Spain, Switzerland, Serbia. Bracon (Cyanopterobracon) urinator (Fabricius 1798) on Larinus saussureae, L. sturnus, Bracon (Bracon) scabriusculus Dalla Torre 1898 – Rhinocyllus conicus (Curculilonidae); Tephritis no host data. Distribution: Austria, Germany, pulchra (Tephritidae). Distribution: Albania, Hungary, Italy, Romania, Russia (Central), Spain, Austria, Belgium, Bosnia and Herzegovina, Great Serbia. Britain, Bulgaria, Croatia, Cyprus, Turkey, France, Bracon (Bracon) scutellaris Wesmael 1838 on Germany, Greece, Hungary, Italy, Macedonia, Anobium abietis (Anobiidae); Euura mucronata, Portugal, Romania, Russia (Central, East, Pontania pedunculi, P. viminallis (Tenthredinidae).

23 BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)…

Northwestern and South), Slovakia, Spain, Sweden, Bracon (Glabrobracon) coniferarum Fahringer Switzerland, Netherlands, Ukraine, Serbia. 1928 on Millieria dolosalis (Choreutidae); Bracon (Cyanopterobracon) uromelas Costa 1888 Coleophora gallipennella, C. serratella – no host data. Distribution: Albania, Austria, (Coleophoridae). Distribution: Great Britain, Croatia, Cyprus, France, Hungary, Italy, Macedonia, Bulgaria, Czech Republic, Germany, Italy, Poland, Russia (South), Sardinia, Spain, Switzerland, Switzerland, Netherlands, Serbia. Ukraina, Serbia. Bracon (Glabrobracon) delibator Haliday 1833 on Bracon (Glabrobracon) abbreviator Nees 1834 on Gymnetron campanulae, Hylobius piceus Anthonomus pomorum (Curculionidae); Lipara (Curculionidae); Olibrus aeneus (Phalacridae); lucens (Chloropidae); Cephus pygmaeus Urophora cuspidata (Tephritidae); Cydia strobilella (Cephidae); Coleophora trogldytella (Tortricidae). Distribution: Austria, Belgium, (Coleophoridae); Mesapamea secalis, Oria Bosnia and Herzegovina, Great Britain, Bulgaria, musculosa, Sesamia nonagrioides (Noctuidae); Croatia, Cyprus, Czech Republic, Turkey, Finland, Pennisetia hylaeiformis (Sesiidae); Cochylis France, Germany, Greece, Hungary, Ireland, Italy, posterna, Pandemis heparana, Rhyacionia buoliana Macedonia, Poland, Russia (Central and South), (Tortricidae). Distribution: Austria, Great Britain, Slovakia, Slovenia, Spain, Sweden, Switzerland, Croatia, Cyprus, Czech Republic, Finland, Netherlands, Ukraine, Serbia. Germany, Hungary, Poland, Romania, Slovakia, Bracon (Glabrobracon) delusor Spinola 1808 – no Spain, Sweden, Switzerland, Netherlands, Serbia. host data. Distribution: Austria, Croatia, France, Bracon (Glabrobracon) arcuatus Thomson 1892 – Germany, Italy, Latvia, Poland, Russia no host data. Distribution: Austria, Great Britain, (Northwestern), Serbia. Bulgaria, Croatia, Cyprus, Czech Republic, Bracon (Glabrobracon) dichromus Wesmael 1838 Germany, Greece, Hungary, Slovenia, Sweden, on Gymnetron asellus (Curculionidae); Platyedra Netherlands, Serbia. subcinerea (Gelechidae); Grapholita funebrana Bracon (Glabrobracon) atrator Nees 1834 on (Tortricidae). Distribution: Austria, Belgium, Omphalapion buddebergi (Apionidae); Gymnetron Bulgaria, Croatia, Cyprus, France, Germany, villosulum (Curculionidae); Amauromyza flavifrons Hungary, Italy, Poland, Romania, Slovakia, Spain, (Agromyzidae); Tephritis conura, T. separata Serbia. (Tephritidae); Coleophora coronillae Bracon (Glabrobracon) discoideus Wesmael 1838 (Coleophoridae). Distribution: Austria, Belgium, on Saperda populnea (Cerambycidae); Anthonomus Great Britain, Bulgaria, Croatia, Cyprus, Turkey, pomorum, A. spilotus, Ceutorhynchus assimilis, C. Finland, France, Germany, Greece, Hungary, quadridens, Curculio crux, C. nucum, C. Ireland, Italy, Latvia, Lithuania, Macedonia, Poland, pyrrhoceras, C. salicivorus, Pissodes notatus Russia (Central, Northwestern and South), Slovakia, (Curculionidae); Byctiscus betulae, B. populi, Spain, Sweden, Switzerland, Netherlands, Serbia. Rhynchites pauxillus (Rhynchitidae); Chlorops Bracon (Glabrobracon) caudatus Ratzeburg 1848 pumilionis (Chloropidae); Biorhiza pallida on Hylesinus fraxini (Curculionidae). Distribution: (Cynipidae); Euura mucronata, Pontania Austria, Great Britain, Bulgaria, Czech Republic, bridgmanii, P. collactanea, P. foetidae, P. Germany, Greece, Hungary, Poland, Serbia. pedunculi, P. proxima, P. vesicator, P. viminalis (Tenthredinidae); Anticlea badiata (Geometridae); Bracon (Glabrobracon) caudiger Nees 1834 on Adoxophyes orana, Rhyacionia buoliana Coleophora obscenella (Coleophoridae); Aphelia (Tortricidae). Distribution: Austria, Belgium, Great amplana, Cydia splendana, C. strobilella Britain, Bulgaria, Czech Republic, France, (Tortricidae). Distribution: Austria, Belgium, Czech Germany, Hungary, Ireland, Italy, Poland, Romania, Republic, Turkey, France, Germany, Hungary, Italy, Russia (Northwestern), Slovakia, Slovenia, Sweden, Poland, Romania, Russia (Central, North, Switzerland, Netherlands, Serbia. Northwestern and South), Sweden, The Netherland, Serbia. Bracon (Glabrobracon) fumatus Szepligeti 1901 – no host data. Distribution: Austria, Croatia, Cyprus, Bracon (Glabrobracon) colpophorus Wesmael Turkey, France, Greece, Hungary, Russia (South), 1838 on Coleophora sp. (Coleophoridae). Spain, Switzerland, Serbia. Distribution: Austria, Belgium, Great Britain, Czech Republic, France, Germany, Hungary, Italy, Bracon (Glabrobracon) fuscipennis Wesmael 1838 Moldova, Poland, Romania, Russia (Northwestern), on Gastrophysa polygoni, G. viridula, Plagiodera Netherlands, Ukraine, Serbia. versicolora (Chrysomelidae). Distribution: Austria,

24 BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)…

Belgium, Croatia, France, Germany, Hungary, Italy, Cyprus, Czech Republic, Denmark, Turkey, Poland, Slovenia, Serbia. Finland, France, Germany, Greece, Hungary, Italy, Bracon (Glabrobracon) hyalinipennis Szepligeti Macedonia, Poland, Romania, Russia (Central, 1901 – no host data. Distribution: Hungary, Serbia. Northwestern and South), Slovakia, Slovenia, Spain, Sweden, Switzerland, Netherlands, Serbia. Bracon (Glabrobracon) immutator Nees 1834 on Anthonomus rubi, Ceutorhynchus pleurostigma, Bracon (Glabrobracon) osculator Nees 1811 on Cryptorhynchus lapathi, Curculio villosus, Pissodes Millieria dolosalis (Choreutidae); Coleophora notatus (Curculionidae); Urophora stylata albicostella, C. alticolella, C. caespititiella, C. (Tephritidae); Biorhiza pallida, (Cynipidae). colutella, C. discordella, C. flaviella, C. frischella, Distribution: Austria, Belgium, Bosnia and C. gallipennella, C. lutipennella, C. milvipennis, C. Herzegovina, Great Britain, Croatia, Cyprus, Czech obscenella, C. ochrea, C. saponariella, C. serenella, Republic, Finland, France, Germany, Hungary, C. serratella, C. solitariella, C. spinella, Co. Italy, Macedonia, Poland, Romania, Russia (Central, spiraeella, C. therinella, C. trifariella, C. vibicella, North, and Northwestern), Slovenia, Sweden, C. vicinella, C. vulpecula, Coleophora sp.1, Switzerland, Netherlands, Ukraine, Serbia. Coleophora sp.2 (Coleophoridae); Cosmopterix eximia (Cosmopterigidae); Elachista Bracon (Glabrobracon) larvicida Wesmael 1838 – contaminatella, Elachista sp. (Elachistidae); no host data. Distribution: Austria, Belgium, Great Phyllonorycter sp. (Gracillariidae); Mompha Britain, Bulgaria, Croatia, Turkey, France, conturbatella (Momphidae); Ectoedemia Germany, Greece, Hungary, Italy, Poland, Romania, agrimoniae (Nepticulidae); Retinia perangustana Russia (Central, Northwestern and South), Slovakia, (Tortricidae). Distribution: Austria, Belgium, Slovenia, Serbia. Bosnia and Herzegovina, Great Britain, Bulgaria, Bracon (Glabrobracon) marshalli Szepligeti 1901 Croatia, Cyprus, Czech Republic, Denmark, Turkey, on Trachys sp. (Buprestidae); Myopites Finland, France, Germany, Greece, Hungary, inulaedyssentericae (Tephritidae); Cydia nigricana Ireland, Italy, Latvia, Lithuania, Macedonia, Poland, (Tortricidae). Distribution: Great Britain, Bulgaria, Romania, Russia (Central, East, Northwestern and Croatia, Cyprus, Denmark, Finland, Germany, South), Slovakia, Slovenia, Spain, Sweden, Greece, Hungary, Italy, Norway, Portugal, Switzerland, Netherlands, Serbia. Romania, Slovakia, Slovenia, Spain, Sweden, Bracon (Glabrobracon) peroculatus Wesmael 1838 Switzerland, Serbia. on chrysantheana (Tortricidae). Bracon (Glabrobracon) minutator (Fabricius 1798) Distribution: Belgium, Finland, France, Germany, on Anthonomus pomorum (Curculionidae); Lipara Hungary, Italy, Russia (Central), Serbia. lucens (Chloropidae); Trypeta jaceae (Tephritidae); Bracon (Glabrobracon) picticornis Wesmael 1838 Cephus pygmaeus (Cephidae); Caloptilia syringella on Clytus sp., Plagionotus arcuatus (Cerambicidae); (Gracillariidae); Mesapamea secalis, Oria Nematus bipartitus, N. salicis, Pontania pedunculi, musculosa, Sesamia nonagrioides (Noctuidae); P. proxima, P. vesicator, P. viminallis Pennisetia hylaeiformis (Sesiidae); Cochylis (Tenthredinidae); Cochylis pallidana (Tortricidae). posterana, Leguminivora glycinivorella, Rhyacionia Distribution: Austria, Belgium, Great Britain, buoliana (Tortricidae). Distribution: Albania, Bulgaria, Turkey, Finland, France, Germany, Austria, Belgium, Great Britain, Bulgaria, Cyprus, Greece, Hungary, Ireland, Italy, Lithuania, Poland, Czech Republic, Denmark, Turkey, Finland, France, Russia (Central, Northwestern and South), Slovakia, Germany, Greece, Hungary, Ireland, Italy, Latvia, Spain, Sweden, Switzerland, Netherlands, Serbia. Lithuania, Macedonia, Norway, Poland, Russia (Central and South), Slovakia, Slovenia, Spain, Bracon (Glabrobracon) pineti Thomson 1892 on Sweden, Switzerland, Netherlands, Serbia. Ernobius abietis, E. longicornis (Anobiidae); Cydia strobilella (Tortricidae). Distribution: Austria, Bracon (Glabrobracon) obscurator Nees 1811 on Bulgaria, Croatia, Czech Republic, Finland, Trachys pumilus, T. troglodytes (Buprestidae); Germany, Hungary, Lithuania, Moldova, Poland, Hylesinus fraxini, Ips typographus, Orthotomicus Russia (Central, North, Northwestern and South), suturalis, Trypodendron domesticum, T. signatum Slovakia, Sweden, Serbia. (Curculionidae); Coleophora caespititiella (Colephoridae); Epermenia fulviguttella Bracon (Glabrobracon) ratzeburgii Dalla Torre (Epermeniidae); Homoeosoma sinuella (Pyralidae). 1898 on Ceutorhynchus maculaalba, Hylesinus Distribution: Austria, Belgium, Bosnia and fraxini, Scolytus rugulosus (Curculionidae). Herzegovina, Great Britain, Bulgaria, Croatia, Distribution: Great Britain, Germany, Italy, Serbia.

25 BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)…

Bracon (Glabrobracon) terebella Wesmael 1838 on Bracon (Habrobracon) hebetor Say 1836 on Miarus campanulae (Curculionidae); Cephus Ostrinia nubilalis (Crambidae); Pexicopia malvella pygmaeus, Trachelus tabidus (Cephidae). (Gelechiidae); Helicoverpa armigera, Heliothis Distribution: Austria, Belgium, Great Britain, peltigera, Spodoptera littoralis (Noctuidae); Denmark, Finland, Germany, Hungary, Italy, Ephestia elutella, E. kuehniella, Etiella zinckenella, Lithuania, Poland, Romania, Russia Galleria mellonella, Plodia interpunctella (Central,Northwestern and South), Slovakia, (Pyralidae); Adoxophyes orana, Lobesia botrana Sweden, Switzerland, Serbia. (Tortricidae). Distribution: Austria, Belgium, Great Bracon (Glabrobracon) titubans Wesmael 1838 – Britain, Bulgaria, Croatia, Cyprus, Czech Republic, no host data. Distribution: Austria, Belgium, Turkey, Finland, France, Germany, Greece, Bulgaria, Finland, France, Germany, Hungary, Italy, Hungary, Italy, Lithuania, Macedonia, Poland, Poland, Russia (Northwestern), Slovakia, Sweden, Russia (Central and South), Slovakia, Slovenia, Switzerland, Netherlands, Serbia. Spain, Netherlands, Serbia. Bracon (Glabrobracon) tornator Marshall 1885 – Bracon (Lucobracon) apricus Schmiedeknecht no host data. Distribution: Austria, Great Britain, 1897 – no host data. Distribution: Germany, Greece, Bulgaria, Croatia, Finland, Germany, Greece, Serbia. Hungary, Italy, Sweden, Switzerland, Serbia. Bracon (Lucobracon) exarator Marshall 1885 – no Bracon (Glabrobracon) variator Nees 1811 on host data. Distribution: Great Britain, Germany, Bruchidius lividimanus, Bruchus atomarius, B. Serbia. laticollis, B. lentis, B. viciae (Bruchidae); Bracon (Lucobracon) femoralis Brullé, – no host Anthonomus pomorum, Baris chlorizans, B. data. Distribution: Cyprus, Greece, Hungary, Spain, cuprirostris, B. laticollis, Ceutorhynchus punctiger, Switzerland, Serbia. Gymnetron asellus, G. tetrum, Larinus jaceae, L. Bracon (Lucobracon) fortipes Wesmael 1838 on turbinatus, Magdalis rufa, Miarus campanulae, Plagionotus floralis (Cerambycidae); Lixus Pissodes validirostris, Sibinia viscariae, Sitona anguinus (Curculionidae); Gortyna xanthenes longulus (Curculionidae); Chaetostomella (Noctuidae); Bembecia scopigera (Sesiidae). cylindrica, Noeeta pupillata, Sphenella marginata, Distribution: Belgium, Bulgaria, Croatia, Cyprus, Tephritis leontodontis (Tephritidae); Hoplocampa France, Germany, Greece, Hungary, Italy, Poland, brevis, H. flava (Tenthrenidae); Colephora Slovenia, Switzerland, Serbia. coronillae, C. medelichenis (Coleophoridae); Mesophleps corsicella, Pexicopia malvella, Bracon (Lucobracon) fumarius Szepligeti 1901 – Platyedra subcinerea, (Gelechiidae); Phyllonorycter no host data. Distribution: Austria, Croatia, Cyprus, mespilella (Gracillariidae); Hadena bicruris Hungary, Romania, Serbia. (Noctuidae); Dioryctria abietella, Etiella Bracon (Lucobracon) hungaricus (Szepligeti 1896) zinckenella, Myelois circumvoluta (Pyralidae); on Clytus arietis (Cerambycidae). Distribution: Synathedon andrenaeformis (Sesiidae); Barbara Bulgaria, Croatia, Turkey, Hungary, Russia (Central herrichiana, Cydia strobilella, Eucosma cana, and South), Slovakia, Ukraine, Serbia. Grapholita funebrana, G. lunulana, Pandemis cerasana, Retinia resinella (Tortricidae). Bracon (Lucobracon) hylobii Ratzeburg 1848 on Distribution: Austria, Belgium, Bosnia and Hylobius abietis, H. piceus, Pissodes harcyniae, P. Herzegovina, Great Britain, Bulgaria, Canary Is., notatus, P. piceae, P. pini, Polygraphus Croatia, Cyprus, Czech Republic, Turkey, Finland, polygraphus (Curculionidae). Distribution: Austria, France, Germany, Greece, Hungary, Ireland, Italy, Great Britain, Bulgaria, Czech Republic, Denmark, Latvia, Lithuania, Macedonia, Moldova, Poland, Germany, Hungary, Italy, Lithuania, Poland, Russia Romania, Russia (Central, East, North, (Central and Northwestern), Sweden, Switzerland, Northwestern and South), Slovakia, Slovenia, Spain, Netherlands, Serbia. Sweden, Switzerland, Netherlands, Ukraine, Serbia. Bracon (Lucobracon) megapterus Wesmael 1838 Bracon (Habrobracon) brevicornis Wesmael 1838 on Glaphyra umbellatarum (Cerambycidae); on Sesamia cretica (Noctuidae). Distribution: Chaetorellia jaceae (Tephritidae). Distribution: Belgium, Great Britain, Bulgaria, Czech Republic, Belgium, Bulgaria, Czech Republic, Finland, France, Germany, Hungary, Italy, Poland, Portugal, France, Germany, Sweden, Serbia. Romania, Russia (Central and South), Slovakia, Bracon (Lucobracon) nigriventris Wesmael 1838 Spain, Switzerland, Netherlands, Serbia. on Plagionotus floralis (Cerambycidae); Hylobius piceus, Pissodes pini (Curculionidae). Distribution:

26 BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)…

Albania, Austria, Belgium, Bulgaria, Croatia, (Scatophagidae); Acleris rhombana (Tortricidae). Cyprus, Turkey, Germany, Hungary, Italy, Poland, Distribution: Belgium, Great Britain, Bulgaria, Romania, Russia (South), Slovakia, Switzerland, Cyprus, Turkey, Finland, France, Germany, Ukraine, Serbia. Hungary, Italy, Latvia, Lithuania, Norway, Poland, Bracon (Lucobracon) ochraceus Szepligeti 1896 – Russia (Central, North, Northwestern and South), no host data. Distribution: Croatia, Hungary, Russia Slovakia, Sweden, Switzerland, Netherlands, Serbia. (South), Spain, Switzerland, Serbia. Bracon (Orthobracon) laevigatissimus Dalla Torre Bracon (Lucobracon) planiceps Telenga 1936 – no 1898 on Plagionotus arcuatus (Cerambycidae); host data. Distribution: Russia (North), Serbia. Pontania pedunculi, P. proxima, P. vesicator, P. viminalis (Tenthredinidae). Distribution: Austria, Bracon (Lucobracon) romani Fahringer 1927 on Great Britain, Finland, France, Germany, Italy, Chamaesphecia bibioniformis, Paranthrene Poland, Romania, Switzerland, Netherlands, Serbia. tabaniformis, Pennisetia hylaeiformis, Sesia apiformis, S. bembeciformis, Synanthedon Bracon (Orthobracon) ochropus Nees 1834 – no culiciformis, S. spheciformis (Sesiidae). host data. Distribution: Austria, France, Germany, Distribution: Greece, Hungary, Slovakia, Slovenia, Italy, Serbia. Sweden, Netherlands, Serbia. Bracon (Orthobracon) roberti Wesmael 1838 on Bracon (Lucobracon) thuringiacus Synanthedon andrenaeformis (Sesiidae); Cydia Schmiedeknecht 1896 – no host data. Distribution: strobilella (Tortricidae). Distribution: Austria, Great Britain, Germany, Greece, Italy, Serbia. Belgium, Great Britain, Turkey, France, Germany, Ireland, Russia (Northwestern), Netherlands, Serbia. Bracon (Lucobracon) triangularis Nees 1834 on Synanthedon tipuliformis (Sesiidae). Distribution: Bracon (Orthobracon) subcylindricus Wesmael Great Britain, Bulgaria, Czech Republic, Finland, 1838 – no host data. Distribution: Belgium, Great Germany, Hungary, Ireland, Italy, Poland, Slovakia, Britain, France, Poland, Russia (Northwestern), Switzerland, Netherlands, Serbia. Serbia. Bracon (Orthobracon) epitriptus Marshall 1885 on Bracon (Orthobracon) tenuicornis Wesmael 1838 Cryptorhynchus lapathi, Hylobius on Phloetribus scarabaeoides (Curculionidae). transversovittatus, Pissodes spp. (Curculionidae); Distribution: Belgium, Great Britain, France, Agromyza flaviceps (Agromyzidae); Iteomyia Hungary, Russia (South), Sweden, Netherlands, capreae (Cecidomyiidae); Fenusa pumila Serbia. (Tenthredinidae). Distribution: Austria, Great Bracon (Striobracon) idrianus Fahringer 1934 on Britain, Bulgaria, Germany, Greece, Hungary, Italy, Hylesinus oleiperda (Curculionidae). Distribution: Lithuania, Poland, Russia (Central, Northwestern Serbia. and South), Slovakia, Slovenia, Switzerland, Netherlands, Serbia. Discussion Bracon (Orthobracon) erraticus Wesmael 1838 on Gastrophysa viridula, Phaedon cochleariae, The present account is the first summary of Phyllotreta nemorum (Chrysomelidae); all available data about the genus Bracon and its Chaetostomella cylindrca, Urophora eriolepidis, hosts until now on the territory of Serbia. Out of Terellia serratulae (Tephritidae); Coleophora total number 250 Bracon species in Europe, 80 laricella, C. lutipennella (Coleophoridae); species are present in Serbian fauna. We have Metzneria lappella (Gelechiidae); Pennisetia registered 290 hosts for 55 parasitoid species hylaeiformis (Sesiidae); Acleris rhombana connected in 409 trophic associations. The hosts (Tortricidae). Distribution: Austria, Belgium, come from four insect orders. The most frequent Bosnia and Herzegovina, Great Britain, Bulgaria, hosts belong to Lepidoptera and Coleoptera, Croatia, Cyprus, Turkey, Finland, France, Germany, whereas Diptera and Hymenoptera are less Greece, Hungary, Ireland, Italy, Lithuania, parasitized. Numbers in brackets refer to the number Macedonia, Poland, Russia (Central, East, North, of registered host species. Northwestern, and South), Slovakia, Slovenia, 1. Diptera (28) with five families: Agromyzidae Spain, Sweden, Switzerland, Netherlands, Ukraine, (2); Chloropidae (3); Cecidomyiidae (1) Serbia. Scatophagidae (1) and Tephritidae (21). Bracon (Orthobracon) exhilarator Nees 1834 on 2. Coleoptera (109) with nine families: Anobiidae Omphalapion hookerorum (Apionidae); Nanna sp. (3); Apionidae (5); Bruchidae (5); Buprestidae

27 BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)…

(7); Cerambycidae (12); Chrysomelidae (6); Achterberg, C. van, Hemminga, M.A., Soelen, J. Curculionidae (66); Phalacridae (1) and van 1990: New host record of Bracon intercessor Rhynchitidae (4). Nees f. megasomides Strand (Hymenoptera: 3. Hymenoptera (26) with four families: Cephidae Braconidae), a parasite of Agapanthia (2); Cynipidae (1); Eurytomidae (3) and villosoviridescens DeGeer (Coleoptera: Tenthredinidae (20). Cerambycidae) in salt marshes. Zoologische 4. Lepidoptera (128) with 21 families: Mededelingen, 64 (3): 25-29. Agonoxenidae (1); Choreutidae (1); Achterberg, C. van. 1985: Notes On Braconidae I- Coleophoridae (37); Cosmopterigidae (1); IV. Zoologische Mededelingen, 59 (15): 163- Crambidae (1); Depressariidae (1); Elachistidae 167. (1); Epermeniidae (1); Gelechiidae (11); Baird, A.B. 1958: Biological control of and Geometridae (1); Gracillariidae (4); Momphidae plant pests in Canada. In: Baecker, E.C., (ed.), (1); Nepticulidae (1); Noctuidae (5); Proceedings of 10th International Congress of Oecophoridae (1); Psychidae (1); Pyralidae (9); Entomologist, Ottawa, 4: 483-485. Sesiidae (14); Tortricidae (24); Yponomeutidae Belokobylskij, S.A., Žikić, V. 2009: New data on (6) and Zygaenidae (1). cyclostome braconid subfamilies Doryctinae, Exothecinae, Rogadinae and Braconinae The most suitable host species in these (Braconidae: Hymenoptera) of Serbia and trophic relations parasitoid-host, belongs to the neighbouring territories. Acta Entomologica coleopteran family Curculionidae and lepidopteran Serbica, 14 (1): 65-71. family Coleophoridae especially the genus Beyarslan, A., Çetin Erdoğan, Ö., Aydoğdu, M. Coleophora which has 11 parasitoids registered in 2005: A survey of Braconinae (Hymenoptera, this work. The torticid Cydia strobilella has Braconidae) of Turkish Western Black Sea remarkably wide range of parasitoids, six Bracon Region. Linzer Biologische Beiträge, 37 (1): species were registered here as parasitoids of this 195–213. moth (B. (Bracon) variegator, B. (Glabrobracon) Brajković, M., 1989: Knowledge of the Braconidae caudiger, B. (Glabrobracon) delibator, B. (Hymenoptera) fauna in Yugoslavia. Glasnik (Glabrobracon) pineti, B. (Glabrobracon) variator Prirodnjačkog muzeja, B (43/44): 127-138. and B. (Orthobracon) roberti). We noted that Brajković, M., Krunić, M., Tomanović, Ž., 1991: A Bracon idrianus is present only on the territory of Contribution to the braconid Fauna (Braconidae: Serbia. This basic research shows better insight into Hymenoptera) of Yugoslavia. In: Hawksworth, the trophic interactions about each parasitoid species D.L. (ed.): Proceedings of the Fourth European which can help in further biological control Congress of Entomology and the XIII programs. Nonetheless, it can ease the identification Internationale Symposium für Entomofaunistik of some parasitoid species according to their hosts. Mitteleuropas, Hungarian Natural History Museum, Budapest, (2): 424-428. Acknowledgements. The authors wish to express their Brajković, M., Krunić, M., Tomanović, Ž., 1994: deepest gratitude to Dr. Sergey A. Belokobylskij from the Research of braconids (Braconidae: Zoological Institute RAS (St. Petersburg, Russia) and Dr. Hymenoptera) of Deliblatska peščara. In: Jenő Papp from The Natural History Museum of Marinković, P. (ed.): Proceedings of Deliblatski Budapest, Hungary for the identification of parasitoid pesak, Srbijašume, 6: 197-502 specimens and to Dr. Aleksandar Ćetković from the Faculty of Biology, University of Belgrade for the loan of Dweck, H.K., Svensson, G.P., Gündüz, E.A., some specimens from his collection. For the present work Anderbrant, O. 2010: Kairomonal response of the authors were supported by the Ministry of Science the parasitoid, Bracon hebetor Say, to the male- and Environment Protection of the Republic of Serbia produced sex pheromone of its host, the greater (Grant No. III43001). waxmoth, Galleria mellonella (L.). Journal of Chemical Ecology, 36: 171-178. References Fischer, M. 1959: Neue und wenig bekannte Braconiden aus Jugoslawien (Hymenoptera). Achterberg, C. van 2012: Fauna Europaea: Acta Musei Macedonici Scientiarum Naturalium, Braconidae. In: van Achterberg (ed.): Fauna 6 (1): 53: 1-25. Europaea: Hymenoptera: Symphyta + Georgiev, G. 2005: Bioecological characteristics of Ichneumonoidea. Fauna Europaea, Version 2.5. Bracon intercessor Nees (Hymenoptera: http://www.faunaeur.org Braconidae) as a parasitoid of the poplar clearwing moth, Paranthrene tabaniformis

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(Rott.) (Lepidoptera: Sesiidae) in Bulgaria. hehetor. Journal of Ecology, 57: 163- Journal of Pest Science, 78: 161-165. 172. Lewis, W.J., Vet, L.E.M., Tumlinson, J.H., van Taylor, A.D. 1988b: Host effects on functional and Lenteren, J.C., Papaj, D.R. 1990: Variations in ovipositional responses of Bracon hebetor. Journal parasitoid foraging behavior: essential element of of Animal Ecology 57: 173-184. a sound biological control theory. Environmental Temerak, A.S. 1982: Laboruntersuchungen zur Entomology, 19: 1183-1193. Wirtshabitat-Wahl von Bracon brevicornis Mathews, R.W. 1974: Biology of Braconidae. Wesm. (Hym., Braconidae), einem Parasitoiden Annual Review of Entomology, 19: 15-32. der Raupen von Sesamia cretica Led. (Lep., Milonas, P.G. 2005: Influence of initial egg density Noctuidae) in Sorghum-Stengeln. Anzeiger für and host size on the development of the Schädlingskunde, Berlin und Hamburg, 55: 152- gregarious parasitoid Bracon hebetor on three 154. different host species. Biocontrol, 50: 415-428. Tobias, V.I., Belokobylskij, S.A., Kotenko, A.G. Papp, J. 1973: Contributions to the braconid fauna 1986: Family Braconidae. - In: Medvedev G.S. of Yugoslavia (Hymenoptera, Braconidae) I. (ed.). A Key to Insects of the European Part of Acta Musei Macedonici Scientarium Naturalium, the USSR. Hymenoptera 3: 4. Leningrad: Nauka. Skopje, 14 (1: 119): 1-23. 500 p. [In Russian]. Papp, J. 1977: Contribution to the Braconid fauna of Yu, D.S., Achterberg, C. van, Horstmann, K. 2012: Yugoslavia. (Hymenoptera: Braconidae) III. World Ichneumonoidea 2011. , Folia Entomologica Hungarica (Series nova), biology, Morphology and distribution 30: 105-117. (Braconidae). Taxapad (Scientific names for Papp, J. 2008: A revision of the Bracon (subgenera information management) Interactive catalogue Bracon s. str., Cyanopterobracon, on DVD/CDROM. Vancouver. Glabrobracon, Lucobracon, Osculobracon, Yu, S.-H., Ryoo, M.II., Na, J.H. 1999: Life History subgen. n., Pigeria) species described by of Bracon hebetor (Hymenoptera: Braconidae) Szépligeti from the western Palaearctic region. on Plodia interpunctella (Lepidoptera: Pyralidae) Biologia, 65 (1): 110-112. on a Dried Vegetable Commodity. Journal of Reinert, J.A., King, E.W. 1971. Action of Bracon Asia-Pacific Entomology, 2 (2): 149-152. hehetor Say as a parasite of Plodia interpunctella Žikić, V., Achterberg, C. van, Stanković, S. 2010: A at controlled densities. Annals of the contribution to Braconidae, Hybrizontidae Entomological Society of America, 64: 1335- (Ichneumonoidea: Hymenoptera) and Stephanidae 1340. (Stephanoidea: Hymenoptera) from the south- Shaw, M.R. and Huddleston, T. 1991: Classification West . Acta Entomologica Serbica, 15 and biology of braconid wasps (Hymenoptera: (2): 227-235. Braconidae). Royal Entomological Society of Žikić, V., Brajković, M., Tomanović, Ž. 2000: London, London. 126p. Preliminary results of Braconid fauna research Stanković, S., Žikić, V., Ilić, M. 2010: Betula (Braconidae: Hymenoptera). Acta Entomologica species as host plants for various insects Serbica, 5 (1/2): 95-111. parasitized by braconids (Hymenoptera: Žikić, V., Stanković, S.S., Ilić, M., Kavallieratos, Braconidae) in Serbia. Biologica Nyssana, 1 (1- N.G. 2012: Braconid parasitoids (Hymenoptera: 2): 117-122. Braconidae) on poplars and aspen (Populus spp.) Taylor, A.D. 1988a: Host effects on larval in Serbia and Montenegro. North-Western competition in the gregarious parasitoid Bracon Journal of Zoology [in press].

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