ATOLL RLSZiRCkI BULLSTIN

Northern Expedition, 1951-1952. Land biota: Vascular plants.

by F, R. F'osberg

Issued by

THE PACIFIC SCILiVCE BOliHD

National Academy of Sciences-National Research Council

:lashington, D. C.

KW 15, 1955 Northern Marshall Islands lkpedition, 1951-1952. Land biota: Vascular plants .ly2

by F. 8. Fosberg 3

1/ Publication authorized by the Di.rector, U. S. Geological Survey

Preli!ninary results of the lkpedition to the Northern Elarshall Islands, 1951-1352, 110.2.

Botanist, U. S, Geological Survey . Northern Narshall Islands Zxpedition; 1951-1952. Land biota: Vascular plants. . . by F. 3. Fosberg , . . .

Following the nhrrative of the expedi-tion to the northern Karshalls made in 1951 and 1952, it seems north whi1.e to present the raw dsta collected in a series of: prel3minary papers, to make them available for general use and reference xitilout waiting urlbil tLey are cc:npletely studied and interpreted. Formal pubiicri.tion of a more parmanent charactsr is planned as detailed .studies are co;u$eted. This paper presents distribu- tional and ecological records, as aell as r~iitiverimes, of the vascular plants coliected on the expedition.. TQ these have beer1 added recorus from the author% 1946 and. 1950 coLlections, a few Wake &lad collections made in 1953, a~cja few miscell.acileo;ls collections by others. Other signi- ficant collections fromthe nort,%ern Marsilr:lis, not included here, are those of NU. Randolph Taylor, pubiished by hiin iri his MI'lants of Bikini," ttiose , made by Adelbert de . Ct~aniissowkii,.e on the Zotzebue ikpeditionJ and those made on several [email protected] . . .. St. John, rnostly as get unpublished...... ~he'~arsiialle~epl&t.nmeo are recorded here in tile form iri vinich they tyvece written down in .thk field? an attempt baing made to reproduce the sotlilds as hsard. Thus, they wclude, sometimes, a nwnber of variants of the same word which shply indicxte differer~cesin the way tile word was heard. 80 attgpt has been maae ,to edit these nanes. accordirig to any of the existing Mars,hallese orthographies. As this seqies of papers is intended to preserve raw data, it. :is,felt .that the naneswill he of more use exactly as recorded. i'khen a definitive flora of the hiarshalls is written, a more satisfactory orthographic form should be sought. .. Plzce names are given as they appear on the ltibels of the collec- tions, with an occasional correctio:1. giyen in,pr;l.entheses. Thus the localities for the 1946 and 1750 .collections.are given, generali?, as on the Hydrographic Office charts, &ereas those for the 1.751 and 1952 collections are given according to.the. revised list of names supplied by Mr. E. H. Bryan and Mr. Leonard Uason oi Honolulu, as a result of their work on the krsnall Islands Atlas (uqublished).

The determirititicns of the plants are all. by the author. They repre- sent the present state of the taonorny of .these species. some of them are in need of revision ar.d may later be changed. All of the &rshallese Pandanus are here referred to 2. tectorius Parkiilsoi~,.as. the nwerous "s~ecies"tha-t have. . been descri'ceci or recorded frowthese islsfdsare considered to be, actually, ndthing more than horticultur,al forms '&I genetic segreg;ites. One or tvio plants are detemir,ed to genus only, either because. . the:species hasnot yet been described, or because the m~terial . .. . , . , ...... , ...... avail.able or the state of the classii'ioatior~i~ such that convincing specific identification is at present impossible. No new taxa are described and no nominclatursl changes are made in this paper.

Collection numbers cited are from the aut.hor1s consecri-tive series except those few definitely ascribed to c'thr;r coil.ector.s, Thj.2 series applies only to plants. -- Polypodim scolopendria Bun. f . : gtirik Islet, 33674. Taka Atoll: Taka Islet, 37723. Likiep r~oll:Li.kie~ Islet, , 2701:3:. Lach Islet, 33789. i.01: Bkdwe Islet, 33913. Lae t?toXl: Lae Islet, 33994. Kmjalein iitol:?: Ihe7wetak Islet, 34135. Ujelzng toll.: Bieto Islet, 311144. i'iutho Atoll: dWotho Islet, 34237. Ujze Atoll: Bock Islet,, 34326. Inc!igenous. Common, mainiy terrestrial but fowd also on the bases of coconut trees and, more rarely, on the 1oi.eir trunks of other trees, principally in coconut plantations m5xed forest. bfarshallese: Ibkino." Panamus kctorius Park. .

Utirik Atoll: Utirik Islet,. ,~33697. ~ Taka {\toll: Taka Sslet, j3'725. : Likiep Islet, 27047; Aikini (Aekone): lsiat, 270:6; 27055; Lado Islet, 33318. : 33930, 33905, 33966. Biiuk Atoll: Ailulr Isiet, 3392'7. : Loj Islit, 3/+9140. Kwajalein: Bennett (Hikej) Islet, Isiet, 2&96; P.evistak Islet, 34.131. Ujclm~1kol.i: Ujelang I:slet, 31:212. : hotho Islet; 3/280. : SOCKIslet, 31.:?:~6. Biicsr Atoll: %.kar.Isiet, 34566; Mi~v;etoi; i,~oll.: Japtan Islet, 24317. Of aboriiinal introclxtion (?), %ar;y vli*ieties, bcth wildad planted, the latter ksed as food.ad ti~ztch.-The varieties differ mainly in fruit size, shape, juiciness, 'flavor, and other eaible cpaii-ties. Planted forms usually around v2llc~gksand formi:-ig a second story in coco- nut groves, wild ones in mixed forest, rarely in pure stands. %arshallese: "bop. " . . : . . Thalassea ---hemprichii (Wrb.) Aschers. Ujel%g ~toll:Ujelahg ISlet, 341G9. Indixenous. Form a thin sod at low tide level on 1a.roon besch on the east egd of Ujelang Islet, ; othekwise unknown in the northern Marshalls.

Cenchrus browiii R;& S. Wake !;toll: Feale Islet, 34929. Introduced weed. Common locally.

~enc11r:usechimtus L. Wake &toll: Wake Islst., -'3'362L. - . , -3LL77. . . . . Utirik Iltbll: Utirik Islet. 33665. Jemo Island: ,33868, Kxajalein xtoll: jeiIslet, 26486, 34140; Bennett (Bikej) Islet, 26504. Ujae Atoll: Ujae Isiet, 34310. Ehivietok Atoll: Eniwetok Islet, 24300; Japtan Islet, 24324;' ,hgebe Islet, 24378. Likiep Atoll: .Likiep Islet, 27636. htroduced weed. In open, s;r;dy disturbed areas, especially around villages and military establisherits, and in thin coconut plmtations. In places very abundant. iiashallese: "lek e leg. 11 Chloris inflata Link : Bennett (Bikej) Islet, 26522, 31202; Kwa jalein Islet, 31187. Eniwetok Atoll: Gngebe Islet, 24407. Wake Atoll: Peale Islet, 34928. Introduced weed. In disturb& places where there have been military operations.

Cynodon dactylon (L.) Pers. . . Wake Atoll: Make Islet, 34450; Peale Islzt, ,34931. Eniwetok Atoll: t . . Eniwetok Islet, 24292; Engebe Islet, 24379. Kwajalein Atoll: Kwajalein Islet, 261+89. .Introduced weed. Common in open places around military establishments, persisting from plantings made in 1945-46, when there was concern about revegetation of the devastated areas. . .

, ~ ~ .~ - -" "- ~ 7 .~ T-~~--~- Introduced weed. ~ccasiinalin open disturbed places around military establishments.

~ikitariamicrobachne (Presl) Henr. Likiep Atoll: Lado Islet, 33'190; Likiep Islet, 27018. Jemo,IsI&d, 33870. : Ailuk IIset, 33946. Lae Atoll: Loj Islet, 34050, Wotho ktoll: Wotho Islet, 34240. Ujae Atoll: Bock Islet, .34363. .. . Kwajalein Atoll: Kwajalein Islet, 31161, 26475; Bennett (Biltej; &let, 26498.- Eniwetok Atoll: Japtan Islet, 24340; Aomon Islet, 24355; Eniwetok Islet, 24369; Engebe Islet, 24373. . . . . Introduced ':('?) weed. Occasional to .c&non in coconut plantations. . krshallese: "ujoij,I1 %jos kulkuii, f5n,ik* pueue." . . . ,. . - Digitaria timorensis (Kunth) Bg1.1.. . . Utirilc Atoll: Ukirik Islet, 33664. aka Atoll: Taka Islet, 33751. Introduced weed. Villages and plantations. darshallese: llujoij." . . Eleusine indica (L. ) Gaertn...... :. WBke Atoll: l~Jakeislet, 33623,34482. Utirik Atoll.:. kirik Islet, . 33663. Ailuk Atoll: Aiiuk kiet, 33971.. Lae Atoll:, La? islet, 34060. Ujelang Atoll: Ujelang Islet, 34182. Wotho Atoll: '$otho.Yslet, 34235. Ujae;iitoll: Ujae islet, 34309. 5;niwetok Atoll: Eniwetok Islet, 24289; Japtan Islet; 24320; Zngebe Islet, 24384, KwajaleinAtoll: Kwajalein Islet, 26485; Bennett (siltej) Islet, 25517. Likiep Atoll: Likiep Islet, 27013. Jemo Island, 33876. Intrcduced weed. Cownon around villages and in disturbed places. Marshallese: lrujoij.ll A giant forn occasional in opRnings in coconut

,' 'plantations on Jerno kland (33876) is possibly only a response to abundant phosphate in soil.

Eragrostis ar~abilis(L.) W.& A. lJtirik Atoll: Utirik Islst, 33653. Likigp Atoll: Lado Islet, 33791; Likiep Islet, 27017. Jertio sland. 33875. Ailuk Atoll:. Ailuk Islet. 33944. Lae Atoll: Lae Islet,' 34065. ujeiang !ake Atoli: hake Isl.et -~, 341,58.- Introduced weed. On bare ground near air strip. iqot prev$ously known from the Pacific Islands.

Lepturus repens. (Nrst,f. ) R.Br. : Wake Atoll: liiake islet, 33627, 311950, 34462; yeale Islet, 349Q, 34942, 349L3...... 31~9W~. . . . . Utirik Atoll: Utirik Islet,, .-33861. Taka iitoll: Taka Islet, 33732. Jemo Island: 33864. kiluk Atoll: Ail& Islet, 33923. Lae Atoil: Lae Islet, 34064. tfjelang kt.011: . Ujeiang Islet, 3LJ73. keotho i~toll:Notho Islet, 34271. Ujae Atoll: Bock Islet, 3!&359; Rua Islet, 34378. Pokak Atoll: Sibylla Islet, 34504; Kamome Islet, 34506, 34509. : Bikar Islet, 34571; Jaliitlik Islet j/.+fl'i'. Ihiwetok Atoil: Enivretok Islet, 24298; Japtari Islet, 24325; Aonion islet, 54353; jyigebe Islet, 24355. Kwajalein Atoll: Kwajalein Islet, 26477; Bennett (~~:ej)Islet, 26510. Likiep Atoll: Likiep Islet, 26995. Indigenous. A very comlon grass, found in open plsces, in p3.anta- tions and forests, but mostly confined.to places. with at least sonle- sand. or fine gravel. Extremely vsriable i;l habit and luxuriance, with greit' latitude in tolerance of different rr,oi.stureconditions. Pioneer species on newly .deposited sand and fine gravel. Marshallese: %jo j, "ujw aitok". (kiwstok), "ujuj. 11 -1' Lepturus sp. Y Pokak Atoll: Kamome Islet, ,34511. hake Atoll: ~kn3.e., Islet, 34940. Indigenous. ii form with extyemejg. coarse habit,. , 'robably a distinct '. species, fomd only verg locally on open grevel, grol;5ng with, but not intergrading with, Lepturus yewns. Apparently endemic - to qokak arid Wake Atolls, Comon,on Poicak only on a part of Kmoine Isif$,:. . +nd"on Lakeonly ...... in a small area on the west end of reale Islet. . ..

Paspalm vai.,inatun STN...... :.. .

Likieg ~$01.~:Likiep ~+t,33824, 27039. ' &,j+ein ilto~.:..Kwa.jalein Islet, 34123, 31189;. ?beye rsle.t, 31207. hake toll: Peaie Islet, 34932;, Wake. Islet,, 34949. 1 At:Aiiuk &let,, 33969a. . , , Introduced weed. In open placesand especially l~u(uriaritin taro .' pits. An aggressive weed, rkcently introauced but spreadine rapidiy, forndng adense to%h sod* Possibly. responsibie for the failure.. . of the revival of taro culture in the'riorthern &rsh&lls. . ., . . ~etkria~erti.cillata (L. ) Bern. : , . -Wake Atoll: !!take Islet, 34470. miwetok &toll: Jz~gtanIslet, 24327; Engebe Islet, 24392. Introduced weed. In disturbed places, not yet common. , Sorghumdgme yar. technicm (~oern.) ~iori,& Paoli . .. ExLvietokAtoll: En::ebe ket, 21+383. Introd-xed. .Apparently plrsisting from pia~ting'by Japanese, +are when' collected, p~obab3.y not a pi-rmane& member of the flora.. .

. . ;1/ See Fosberg, F. R., The Pacific form: of Lept.urus R. Br. (Gra.??ineae). Occ. Pap. ish hop t.fus.. 1955 (in press). -Stenotap!lrurn micranthum ( Desv. ) Hubb.' UJelarie; Atoll: U.ie:Lanpc Islet.,-. 7L200. ,.. 3L201. IntrodGced weed."^ wizespread but not coinmon Pacific species, found here for the first time in the harshtill Islands, growing along paths in coconut plantation.

Thuarea involuta orst st.) R.& S. . . Utirik. Atoll:, Utirik Islet, 33692. Likiep Atoll: Lado Islet, 33841; Likiep Islet, 26993. k:il& ikoll: niluk Islet, 33922. heAtoll: Lae .., Islet, 34020; Ujelag Atoll: Ujelang Islet, 3b74. :,jotho &toll: vcotno Islet, 34238. Ujae Atoll: Ujze Islet, 34300; bock islwi;, 31c360. Eniwetok Atoll: Eniwetok Islet, 24295; Jagtan Islet, 2431;L- fiomon Islet, 24354. Kwajalein..Atoll: Kwajalein ISlet, 261+74; Bennett tliikej) Islet, 26524. Indigenous., In sandy places, shaded 0r.h the open, 1oc~I.l~very common, forming a soft green ground cover. Marsimllese: "ujoij, 11 "u juj, I' "uios" maroro1I (Eniwet old. 2ricachne ~sul.arig(L. ) Nees . .. Wake Atoll:. hke, Islet, 34476. Eniwetol: Atoll: JAptan Islet, 24328. Kwajalein Atoll: Bennett (Bikej) Islet, 26502.

Introduced weed. 0ccasior:al in disturbed places around abandoned military installations.

Gyperus compressus i. Kwa jalein Atoll: Kwa jalein Islet, 31189. Introduced weed in dicturbed phces. &perus javariicus iioutt . Likiep Atoll: Likiep Islet, 33659, 269@. Indigenous(?). Rare around village. %rshail.ese :l~s$p&sZp,fl

Clvperus odor~tusL. Likiep Atoll: Lado 'Islet, 33845; .~ikie~:Islet, 27043. Ailuk Atoll: Ailuk Islet, 33967. Lae Atoll: Lae Islet, 34014. Notho Atoll: Uotho Islet, 34260. Kivajalein Atoll: Kwjalein Isiet, 31172, 26484. Introduced (?) weed. Comon in taro pits and othec wet places. . . Cyperus rotundus L. Kwajalein Atoll: Kwajalein Islet, 31200. keAtoll: Peale Islet, 349333 Wake Islet, 31+948. Introduced weed i.n gardens.

H.eochs?ris ~enicuiata.(L.) E.& S. Li~iepAtoll: Lado Islet, 33825. Ailuk ntol'i: ,Qil,.ukIslet.33969. Introduced (?) weed. In taro pits. . . Fimbristylis spathacea Roti Wotho Atoll: Viotho Islet, 34259, 34272. V%ke Atoll: Viake Islet, 33619, 34457, 34.478. Utirik Atoll: tiset, 33642. Taka Atoll:. Taka Islet, 33737. Likiep Atoll: Lado Isle.t, 33835. hiluk Atoll:. kiluk. Islet, 33941.. Lae Atoll: Lae Is!.et, 34063. Ujelmg ittoll: Raoj Islet, %I-54. Ujae Atoll: Bock Islet, 34327. K-wajalein atoll: Bennett (Bikej) Islet, 31204, 26518; ir~qjaleinIslet, 261;85. Likiep Atoll: Likiep Islet, 27016. Indigenous. Co~unon to sburidant in: o.&q places,, coconut plantations and ncixed woods, variable in habit, tolerant. of .aWXt' Xange o$ Goisture conditions. Piofieer in disturbed places, .Mar6,.ihalle,se: glderelij'~~ig~,lt.. .:. "drali j ',man, fl l~perelijman,I! "ujoj maroro, " Weye jisrnari.,~~, -: ... . . -Cocos nuciferz L. Jeino Island: 33899. Ehiwetok Atoll;: Igurin IsLei;, ,24216...... Introduced. Plated in large numbers qn 611 atol.l:.~xceptPOX& and wake. ~o~llin&tin the moat extensive vegetation type. M;l:rshs.liese.... 'l,ni.!t . . ,: ...... Alocasia inacrorshiza (I,. ) Seho-tt ......

.Ailuk toll: Ail~kIslet, 33970. 'Ujel&. g-t~ll:~i&ti' rslet,; , ' , 34145. . . ~. . . . Of .aboriginal introduction. Plqted aroucd ~j.lli&s, .sg

. , L&mtosperrna chmiissonis (~cliott)Merr. Ail~~k toll: Ailuk Islet, 31968. ii'otho koll: Ivotho Islet, &61. Ujae ~to1.1: Ujae Islet, 3L+302.-Likiep xtol.1: Likiep Islet, 2701~6.. .. Of aboriginal introductio-fi., Plariteci and persisting in taro pits. MarshalLese: %.arij" or "iarej. It......

Rhoeo discolor (~"iIer.) Hance .,, , ..., . ... - ...... , Ailuk Atoll: Uiluk. Islet, 33974. , :...... arden plait...... ,

,...... , , Islet,. 27050 (?). . . Introduced. Rarely plagted......

Crinum procerum Carey; . , . . Utirik Atoll.: Utirik ~slet,33686. Likikp toll, Ledo islet, 33820. Ai..l.uk Atoll: Ailuk Islet, 33956. :.!otho A.to1.1: Wotho Islet, 34278, 34225(?). .. ,. . of aboriginal iritr&uction (?). Nid.elji planted and persisting. Marshallese: "kiep, 11 "kieb." ...... , .. Crinum rumphii Merr. -he ~ltoll:'Loj Islet, 3yJ51. Ujeiatlg.Atol1: Ujelang Islet, 34197. U jae toll: U jae Islet, 31h3l.l (?); , Bock Islet, 343ir5;; .. . . . , ,> ...... , .; :: Of aboriginal introduction (?Widely planted' &ld persisting. . . &rshailese: Ykiep, 11 l1kieb, 11 ...... -Crinm asi.aticm. L. (?) . , Likiep : AtolL: Likiep Islet, 27001. . . 1nt~oduce.d.Planted.~arshallese: "kieb.I1

. . . . Hyraenocallis littoralis (~ac~.) Salisb. Aiiuk Stol.1: Ail';; set, 33976. Lae toll: Lae Islet, '34093. Ujelang Atoll: Ujelqg Islet, 30.85. ~jaek.toli: Ujae Islet, 34284. Likiep Atoll: Likiev Islet.. . '26992.. . . . Introduced. Planted hi vill;~es. iderskiiilese: rlkiep,u "small kiep." -Zephyrathes -- rose2 (Spreng.) Lind. Likiep Atoli: Li.klep islet, 27033.

Tacca leontopetaloides (L.) 0,Ktze. -Utirik Atoll: Utirik Islet, 33695. Likiep utoll: Laao Islet, 33839; Likiep Islet, 27000. Jeido Island: 3387'7. Ailuk At,oi:L: Biluic Islet, 33938.. Lae Aioll: ~aeIslet, 3400'-). Ujelang Atoll: Ujelang Islet, 3/+206.-Yfotho Atoll: Wot!io Islet, 34242. iJjae Atoll.: Ujat! Islet, 34288; hbeju Islet, 34398. Enivvetok Atoll: Japtan Is:?&, 24333. K;laJalein Atoll: Bennett (:Bikej) Islet, 261Jj3. Of aboriginal .&troductli.on ( ?) . Aix~qcient aid spontaneous ' in coconut plantations,especialiy so on Utirik and i~iiuiKAtolls. Tdbers used for food, especially in rior.thern atolls. hkrsi:nllese: boken;ok,ll 9n0kmok.~~

ELusa-- sapientm L. ,, Ailuk Atoll: Aiiuk Islet, 31965. Lae Atoll: Lae Islet, 34021. Ujae Atoll: Ujae Islet, 34237. Kwajdeiri (not collected) : Introduced. Plated in shaoec and sheltered pl.aces, especially on Ujae. Becoming chiorotic when exposed to sdt spray. Xarshal.1ese: %anana.u

Peperomia sp. Lae Atoll: Lae Islet., -,3L027. . Indigenous. Very rare, one colony on rocks in Barriar,to& forest: first record from northern hiarsh&.is. M~~ha1lese:~~r;tikia~a.~~ , . -.-F1eur.p ruderalis (Forst. ) Gaud. ex i:edd. Utirik Atoll: Utirik Islet., --'33679. .. Taka- Atoll: Taka Islet. 337LL:. ,, Watwerok Isl.et, 33'764. 1,i.kiep Atoll: Likiep Islkt, 27004; Lado filet, ., . 33794. Jemo Island: 33880. Ail& ;!toll: Ail& Islet, 33939. Lac ,!toll: Lac Islet, 34030. Ujelmlrir: ~toi:!.: U:;e!.an: Isriet, 34175'. Ujae Atoll: Bock Islet, 34341. .Enitietok ii'ioil: ~a~t;nIslet, 21+321; kgebe Islet, 24399. Indigenous. Co~~iionlocally, espec

Introduced. ' Planted, the seeded form possibly spontaneous, arowd villages and houses and in coconut plantations oil. tile 1agoon;vai-d half or two thirds of islets, especiiU.y i'rom Ail.& swdth. Two forms, one ,with seeds, the othcr seedless. Trees were found growing within 15 rn, of passage and lagoon beaches, ~uitl~outapparent ill effect::. n'one were found close to the outer beech. &irsh~llese: llm9,15 %ei,ll hne,ll

Ximenia arnericana L. -Lae toll: Livejap Islet, 34073. Ujze toll.: Eoeju Islet, 311399, 34Lb00, 34383. lhiwetok Atoll: Jhpten Iilet, 24319. Indigenous. in thickets and in uidergrowth in phntations.

uvifera L. Wake Atoll: Wake Islet, 31t.l+'/1; pea1.e Islet, 34935. IPmjalein toll: Ebeye Islet, 31212. Introduced. 0cccisionalI.y planted around !~~erica.ninstallations.

Boerhavia diffusa L. Wake Atoll: lii'ake Islet, 33626, 34453, 31+946. Utirik Atoll: Utirik Islet, 33680. Taka Atoll.: Taka Lslet, 3374.3, 33748; i!ztwerok Islet, 33765. Ailuk Atoll: Ailuir Islet, 3331. KwaJaSein ,]toll: E?ewei;ak Islet, 34136. Pokak Atoll: Breje 1slet; 34503; ~am;me Islet, 34517, 34.54.6. Bik& Atoll: Bikar Islet, 3457%. Fhiwtok Atoll: Igurin Islet 24306; Japtan Islet, 24334; Aomon Islet, 24356. Li!

Boerhavia fon'with leaves as in ;, mutabi1is.R. Br. JVake Atoll: Wake Islet, 34947. Intro&.ced? Common 1ocal.ly on cord sand and gravel.

Boerhavia tetrads F0rst.f. Utirik Atoll: U$irik Islet, 33644. Taka Atoll: Taka Isle*, 337116; Watwerok Islet,. -33761. - . . -33762 - . fn~~brici?).. " , Jemo Island: -37881. - ii1u.k Atoll.: Aiixk Islet, 33937. Lze Atoll: l3iker;a.j Islet, 3L$?7L+, 3!+G76. Ujelang Atoll: Bokan Islet, 34163. WC+,!?J iito1.l: ;!otiio Islet, 34277. UJae Atoll: Bock Islet, 34320. Gake Atoll: Lake islet, 34454. gnixetok AtoAL: Aomon Islet, 24362; Engebe Islet, 24Yj4. Likiep Ati:ll:Aikir~i (~eicEne)Islit, 27057. Indigenous. Comaon in rocky or gravelLy, even si;n* !>laces, especiklly in the open, also 'ln o2en coconut groves. Apparently inter- grades or hybridizes with 2. --difhsa -(3171+6,31,074, :5l1076,2l+362, 2/+39&,27057 338Cl). MarshBllcse: 'tri%tt to, I) Iperial, " umaf,ok'aitokll (kn~;&,ok). b'iirabilis jaiapz L. Utirik Atoll: Utirik Islet, 33657. Ail& ~to1.l:Ailuk Islet, 33960. Eniwetok Atoll: Ihiwetok Islet, 2L1370. Likicp At0:J.l: Likiep Islet, 2.7029. Introduced. Planted in gardens adaroun2. houses and graves.

Pisonia --.-grandis R.Br. Utirik :1i,oll: Utirik Islet, 33676. :'aka ~toll:Taka Islet, 33735. Likier, Atoll: Lado Islet, 33852; Aikirii (nek~ne)Islet, 27052. Jeruo 1slard: 33895, 33898. ~il&~tcli: Akdwe &let, 33YlO:~ae Atoll.: he Islet, 34026, Ujehng ALO~~:Ujelan,: Islet, 31,208. ~otho toll: '8otho Islet, 34263, 6;Oixi~clln~ Islet, 31,430, Ujae iito!.L: Bock Islet, 34336, 34337. Pokak Atoll: Krsnonoiiie Is.Lct, ji+50r7. Bikap Atoll: Bikar Isl.et, 34569. Wake Atoll: bliake Islet, 34472. biwetok Atoll: Igurin islet, 24304, 24309; Japtar, Islet, .?4332, 24337; Aomoil Islet, 2425';; mgebe Islet, 241dX2. Kwjalein toll: Kwajalein Islet, 26512. Indigenous. Frequent tree in nixed forests, usuall! away from direct exposure to wind. Impcr'mnt in fornk-ig pure ~%~ild.forests. ~6vo'rc;dby bircis for roosting and nesting. M~rsiialLese: "kangl., I! Itkangae" (Eniwve-tok). Ac!lyranthes aspercr L. Ehiwetok Atoll: Aamon Islet, 2411r8. Introrhced weed. &re in disturbed pl.aces. Karshallese: "kale:clekilf

Acivranthes canescens R. Ba. Utirik Atoll: Bekrak Islet, 33714. Taka i~toll:Taka Islet, 33729. era 1:3389s. 1ndigenous.Very local in scrub on drier islets.

Amaranthus dubius &tart. -7~-flajamtoll: !Wajalein Islet, 31178, 31185; Bennett( ~ikej). Islet, 26503. Wake toll: Wake Isiet, 3l+L+52;Pealo Islet, 311938. Eniwetok toll: Emebe: Islet, 24390. Introduced we&. Cormon arour~dr&I.itary installations.

Amaranthus ~J~~~QSUSL. --- Kwa jalein Atoll: Kwa jalein Islet, 31174. ~ntroducedweed. Infreciuent around military installations.

Amaranthus viridis L. !7ake Atoll: Wake Islet, 3/+/+1+6.Eni->.etok Atoll: Ihiwetok Islet, 21+372. Introduced weed. Infrequent around military installations.

Gornphrena nlobosa L. Ujae toll: Ujae Islet, 34291,. Introduced. Rare in gardens. Portulaca -lutea Sol. Utirik Atoll: auk Islet., ?37CI. Poksir Atoll: iiamorfie Islet. 54510. Wotho Atoll: Biken Islet, 34434. Bikhr ~ttill:Bikar hlet, 34567; 34570; Jaliklik Islet, 34578. Wake Atoll: !:ake .r.siet,7. j4468. Ehivretok Atoll: Igurin Islet, 21,307; Japttn Islet, 24326; Aoinon Islet, 21,358. Indigenous. Abmdant in the nwthern atcLls, less so southward, in open sandy or gravelljj places and uncer trees vrhere other ground cover is '. not thick. Pioneer on new sand and gravel bars, beaches, and typhoon- stripped areas. Marshallese: "puryn," 1Ikiran.lt Portulaca oleracea Id...... ,, Nake ~$011:V?&e Islet, 33620, 34481. ~RkaAtoll: Taka 'lsl&t) 337Lc2. Ujae toll: Ujne Islet, 34308. Fhivietok /ii;oll: Biiwetok Islet, 24296;. &gebe Islet, 24408. Kwa.ja.lein .~toL1:Kviajalein Islet,' 26491; Bennett .

7. htroauced weed. Coninon in sudy placesaromd villagesand in ... disturbed. . places. p~rt,&acasambensis v. ~oo~n...... Ut'irik Atoll: Utirik Islet, 33675. ~ikiepAtoll: Lado Islet, 33848;. Likiep' Islet,, 2700 5. Ailuk fitoll: Aii-k Is:Let, 33940. UJeiing Ato&t: . . 7 Ujelang Islet,, 34216. ~rkeiitoll: qiake hie:, 34.46G. Wiwetok. .Atoll: .. ' . Ehiwetok Islet, 24297; J~aptanAs:Let,. 24.32; Engebe hlet, 24406.. Kwajalein Atoll: ~wajalein'Ts1et;:26469, 26513. : 'Introduced ('1. weed. Gommori in sandy open or distwbed places. :.. . . barshallese: %at 'io," "'uujang. 11 . .

Sesuvium portulacastrw: L...... , . .. . -IVake Atoll: VWe Isletj 34461, . . . . Indigenous. Common on open saline flats. ....

. . ... Cassytha filifoi-mis L...... Utirik Atoll: Utirik Islet, 33694. $&$ Atol.1:' TakaIslet., 33724. no1: 33889. ~il.ukAtoll.: Ailuk Islet,.- 33936. L'ae~kboll: Lae Islet. 34080. UJelang Atoll: Ujelang Islet, $4172. lvotho toil: Wotho Islet, 34266. Ujae Atoll: @ck Islet, 34310. hiwetok Atoll: B~iwetokIslet, 24288; Igurin IsTel;, 2t53i3; Japta;?..Isl.ei;, -24338; iiomoi? &let, 24365. Kwajaiein %olli Hennett ,(~ikej)~slet,, 264%. . .. ~iitii?~. Atoll: Likiep Is:iet, ... 27009...... Indigen6us. (;&non 5enerally in,openin&, :&open..... coconut groves; and in loose scrub, pcirasitizing grasses, sirrubs anileveri small. tree*; forming loose mtits covering vegetation, J-arshailese: "kaanin," Vkmi ... a "kenen, ltk&inun. 11 ... .: ...... " . . .,, .. :. ... . ~ . . . , ...... ---Hernmdia sonora L. ... Lae -Lae Islet, 33977; Loj Islet, 3L1.0Lrl. Ujelmg Atoll: U~elaiigIslet, 34211. -Likiep litol.1: .!iikj,ili (Aek5ne~ IsLet, 27060. Indle,&nous. Rare ill.mixed foresi;..eu:d in coconut groves. Seedli;~~ found in gerxinated beach drift ai; to{> 01 heir& (33997). HarstiKllese: "pingiping, !thing bing. " ...... ,. . . . Brassica sp. . . . . -Xniwetok litoll: beebe Isle-t, 24388. Elare,: persisting a:'cer cultivetion, probably liiis now disappeared. . -...... pibidi& .'.&ii5.~iens& Chiin.. -& ~chlecht..'...... Wake Atoll: .Peal.e Islet, 34930;. .. , . , . ~di&rlous.. Very common locally on steep banks of 1kke. . Peale," Caesalpinia ~ulcherrima(L. ) Sw. Likiep Atoll: Likie~Islet. , 27031..- ~ntroduced. Rare. pianted in gardens.

, . Caesalr~iniabonduc (L.) Roxb. (?) Ujae Atoll:- Nle Islet, 3W6. Indigenous. Rarely seen as seedling from drift seeds; seem's not to . . . . p.ersist:in horthern darshails.

Caesalpinia (Medic.) Dandy & Bell (?) Lae Atoll: Lae Islet. -.3L031r. -. Rarely seen as seedlbg from drift seeds; seems not to persist in northern krshalls.

Canavalia microcarpa ~Dc.) Piper Likiep Atoll: Liklep Islet, 33857. Jew Island: 33888. Lae Atoll: Lae Islet, 34012, 34031. Ujelang Atoll: Bieto Islet, 34146; Ujelang Islet, 342-04. Notho Atoll: Wotho Islot, 3l+227; Kabben Islet, 34431. Ujae Atoll: Bock Islet, 34319. Biwetok Atoll: Igurin Islet, 24309. Kwajalein Atoll: Bennett (~ikej)Islet, 2651.5. Indigenous. ~oinhonbut loca,l in coconut plantations and in mixed forests. Roots of seedlings,at least,.bear bacterial nodules. Idarshallese: "manlap, "marlap......

cassia occidentalis L, . . Ujelang Atoll: Ujelang Islet, 3/+l84. Introduced. Persisting after planting by Japanese; said to have been 'used by them as coffee ,substitute...... ,. Desmadiun canurn (Gmel. ) Schinz & Thell. Kivajalein Atoll: Kwajalein Islet. 3fi25. Introduced weed. Conion around 116lltGestablishment on Kwajalein. When clipped makes fairly attractive lawn. . . , . @ythrina variegata var. orientalis! (k.')' Merr. . . Likiep Atoll: Likiep -I.slet, ' 270k5. . . Introduced. Rarely planted: ' " , ...... Intsia (Cblebr. ) Kt& -Lae toll: Lae Islet, 33995; Bikenaj Islet., '34077; Bikelabet Islet, 34079; aemanman Jsl~t,3~09~.-3~oth8'~toll: Wotho Islet, 34246. Ujae

. . , . Atoll: Bock Islet, 34317; Wo jia islet, 34387. Indigci~ous.Compcnent of mixed forest.on southem atolls of northern

Marshalis. Marshallese: Itkubuk." , , ~. ,.. . ,: : . . . Leucaena a(L'.) Bentfi;" . .. '" Utirik Atoll: Utirik Islet, 33669. Ailuk Atoll: Ailuk Islet, 33972. Kwajalein Atoll: Kwajalein Islet,, 31176. . Introduced. Persisting:'aftkr~~cultivation.Rare. around former Japanese establishments. %ts seed .but seems not to 'establish itself in northern &rshalls. Hawaiian: nkoa haole. . . Sophora tomeritosa L. Ujelang Atoll, Ujelang Islet, 34210. Ujae Atoll: Zbeju Islet, 34385. , ... Liltiep Atoll: Aikizi (Ael;?he) Islet, 27065...... , Indigenous (?). Aiong lagoon sides of islets, just bsck of beaches in edges of cocon3.rt plmtations, not com;i.or;. iarshallese: "kileJtt nkuli.lt

Vima marina(Burn1. ) I$err. Lae Atoll: Loi Islet., -.3LOL75L: ,., Lae Islet.. -.3lr036. - Likieu Atoll: Aikbii (~ekzne)Islet, 27082; Lado Islet, 33822. Ujelan:: htbll: ~j.elangI:siet, 34179. Wotho Atoll: VJotho 1;le.L 34234. Ujde Atcil.: Ujae Islet, 3b285. Kwajalein Atoll: Bennett (Bikejj Islet, 26521. Indigenous. Comon ground cover in coconut groves, climi?ing in bushes in mixed foi-es~,2ioneer on cleared. . areas arid ,on nev: sad and gravel bars. Roots bear hacterid nodules. rm-shaliese: "marukurien jo jo, "marlapt! (Uje- lang).

,. . ... Tribulus cistoides L...... hiwetok toll: Fngebe Islet, 2&03. . . InLroduced. (?) weed. Rare on open sand; probably intmduced acciden- :., tally with military equipment. . ~ . . .. Citrus aurantifolia- (Christm,) 3v;ingle Ujae A.boll: Ujae Islet, 31;.303 (:') . Likiep &toll: Aikini (~ek~ne) Islet, 27051. Introduced. Rarely planted in viiiages.

Soulamea amara L. .. . . --Ailuk Atoll: Akulvre Islet, 33912. Lae toll: &ananman Islet, 34lCO. Wotho Atoll: Wotho Islet, 34269. a Atoll: Eoeju Islet, 34384. Indigenous. Locally common in mixed forest. ~arsI~allese:~kil'li,f~ tkabi jiling. II . . Suriana rnaritima L. , Utirik &~oll:Utirik Islet.. 31668:-- auk Islet.,. --53699. .. Taka Atoll: Taka Islet, 33738. Likiep Atoll.: Lsdo -!%let, 33788; Likiep Islet, 26996; Aikini (kek8ne) Islet, 27066. .Ailuk Atoll: Akulwe Islet, 33909; Ailuk Islet, 33947. Lae Atoll: L~Ekkt., 34033. Uje1ar.g Atoll: Bokan Islet, 34166. Lotho Atoll: iuotho Islet, 34273; Eokanaetok Islet, 3444.l. Ujae Atoll: Bock Islet, 34315. Jkiwetok Atoll: Igurin Islet, 24310; Puraai Islet, Hosaka 2691. Indigenous. Comon in sandy places in scrub, especially along outer beaches, occasion.ally. cio~ninmt. &arshallbse:. "ngienge, It "kaianpi, ," "ngiangi. t~ Acalypha zodseffima ihast. Likiep litoll: Likiep Islet, 27022. ~ntro~uced.~a~ely planted; aoes not. appear hedthy.

Acalyphs hispida hrm. f . ~ Likiep iitoll: Likiep Islet,.27027., ~ntroduced. Rbrely planted;. does not appear healthy.

Acalypha :?ilkesiana X. -k. AiPi Atoll: Ailuii Islet, 33977. Lae Atoll: Lae Islet, 34097. Introduced. Unco:iunon in villages, does not appear heaithy. --Codiaeum variei;atun- (L. ) 01. Liki.ep Atoll: Lilciep Islet, 570214. Introduced. Xarelyplanted;

Euunhorbia chauissonis Boiss. Utirik Atoll: Utirik Islet, 33662. Likiep Atol!.: Lado Islet, 33'793; Likiep Islet, 27012. Biluk Atoll: liiluli islet, 33921. Lae Atoll: Enemanman Islet, 31c10.2. Ujelang Kcoll: U.jelan# Islet, 311.207. Notho Atoll: lsotho

1slet; 34232. ~jae~icll: ~ock-~s~et, 3~8325. ~ Ehiiiietok Atoll: hivcetok Islet, 21+286; Fhgebe Islet, 24335. Kviajalein ktol'l: Kwajalein Islet, 26470. Indinegous. In cocvflut groves, domi!i+t in ground cover ifi may places. HarshaTlese: "ma1 dok, 11 %arm, 11 ilpuripur, $1 "perul. " --bia - het,erophyll~L. Utirik Atoll: kirik Islet, 33672. Likiep Atoll: La& Islet, .3.38&?. Likiep Islet, 27311, li2ake Atoll: Kake islet, 34474. liiiuk (not collected$.

Kwajalein Atoll: Kwajalein Islet, jll&, ' 2648'2. Introduced weed. Cormon in uisturbed places, abundant locally, especially so on :eke Atoll. Marshhllese: flnukuni;" Euphorbia hirta -. -. L. v-'A k e &toll: Kake Islet, 3362tr, 34465. Jtlno Islad: 338'73. Kwajalein Atoll: Bennett (Eikejj Islet, 265iil. mivie-tok Atoll: &.iwietok Islet,

24367.. Likiep Atoll: Likiep Islet, 2699;. . ' . . , . Introduced. . weed. Corn&& in dis%urb&. . yxices...... Euphorbia @ericifolia,---- L. Wake ~t0.11:Lake Islet, 33618, 33622. Rviajalein Atoll: Kwajalein Islet, 31166. Introduced weed.' In. ,~ tiisturbed . . places,.'local' but common where found.

Euphorbia prostrzta kit. '. ' Likiep Atoll: Lbclo Islet,, 3%29. ,Jemo Ialand: 33874. K3;jajalein Atoll: Ki~ajaLeinIslet, 342.22, 26473 (1. a 1 Ujae Islet, 34307. Wake Atoll: Wake Islet,- '34455;~. ~.. peale Islet,- '34939. Introduced weed. Comnon in disturbed plzces and aroma villzqps.

. . -Euphorbia thymifolis L. -. Kviajalein liviajalein Islet, ' 343.3'7, Ujelang Atoll:. U jelang toll: ...... IsI&t;, . 34181.. Introduced weed. Very loic;liy cbmmoh indisturbed places. I\farshaJ1- me: "xiat lto.11 Phyll~ntnusrii~uri-- L.

Utirik Atoli: Utirik Islet,. 33688.~.~ Likiep Atoll: Lzdo Islet, 33832;- - Likiep Islet, 27015. Jemq Islaria: 333'69. Lae Atoll: Lae Isiet, 34692. Ujelang Atoll: U jeiang Islet, 3i~~l.83.U jae Atoll: Ujae Islet,. 34306.. Kwajalein Atoll: KwajaLein Islet, F173, 26487. Introduced weed. Cor~nonin disturbed places and around villages. .. . Ricinus colmunis L. Ehii!etok ktoll: Ehgebe Isict, 24377. Introduced. Occasionally plantea armd military instdlations, probably by Japsnese. 33710. Lae Atol.1: Lae Islet, 33991. Kwajalein iitoll: Ehewetak Islet, 34134; Bennett (Bikej) Islet, 31201, 26511. Ujelan~Atoll: U.jei.c!.ng Is!.&, 3I+199. Wotho Atoll: Ilotho Islet,, 34226. ujae iiioll: ~ock-~sl.e<,3!;j~j; Ujae Islet, 34419. Indigenous. Cormon in mixed forest in southern atolls; in iitirik in scrub. Marsiiallese: tlk$tak, 11 I~lmi;alc, or t'keda.

Dodonea viscosa (L. ) Jacc,. Likiep Atoll: Likiep Islet, 26998. Indigenous. Rare in openings in coconut plantatioris. Marshaliese: ~tk*&~,$1 ---Trimi'etta ~cumbensForst. Utirik ~toll:Utirik Islet, 3:369:3. Taka Atoll.: Taka Islet, 33721. Jemo Island: $3863. Aiiuk iitoll: .Aildc islet, 33932.. Lae ktoll: -lac Islet, 34107. Ujelag Atoll: Uj"lang Islet, 31,193. Xotho Atoll: liotho Islet, 34270. Ujae Atoll: Bock Islet, 31r-322. Biktr Atoii: Biker Islet, 34575. Fniwetok Atoll: Eriiwetok Isle'c, 24299; Igurin Islet, 21;311; Japtan Islet, 24322; Aomon Islet, 24366; Dgebe Islet, 2440i.. Kwajtilein Atoll: Kwajalein Islet, 26478; Bennett (Bikej) Islet, 2651./+. Likiep li.tol1: Likiep Isl.et, 27014. Indigenous. Very general in disturbr2d places, coconut groves, mixed forest, at top of bezches, and other pioneer habitxte, sometiws fomir.g pure ground cover. Flowers open in inorriing and afternoon, rAosi?d in middle of day. iYiarshtillese: "atlatJ tta~;-t.It

Abutilon iridicum (L. j Sweet ' Nake Atoll: hake Islet, 34151; Pcale Islei;, 3,4937. . Indigenous (?Conunm loc~llyin open scrub.

Gossypim barbadense L...... Ririk Atoll: Utirik Islet, 33656. KwjaLein (not collected).. Introduced. Rare, planbed in vilLage. Seen bui not coliected in nursery on Kwajaiein.

Gossypium hirs~tumvsr. is(L. j Watt Irake Atoll: V:&e 1sietm9. Introduced. Persisting after cuitivatioh. Cor~non, especially arounu old Japanese installations. This plant is identical with a specknerl collec- ted in 1923, Pollock & Fryan -.33, when it was reported to be rare, on:!y one plant seen. . ,

Hibiscus tilihceus L. Likiop Atoll: Lido Islet, 33823. Lae Atoll: E0.i Islet. 3L@Li2.

Hibiscus (hybrid) Likieu Atoll: Likieu Islet, 2'7026. Introducud. %.rely plated. Sida acuta Burm.f. . .. . %. --Ujelmg Atoll: U jelmg Islet, :j&18O. Introduced weed. r$cry - local, only seen in Ujelag village.

--Sida -- fa1ll.a bialp. . . Utiriic Atol.1: ~tirikIslet, 33687.. Taka litoil: Taka, TSlet, 33726. Likiei, +toll: iado Islet, 8.Ailuk xtoll: Ailuic Isget, ,33962, Lae Atoll: Lae Islet, 34996. U.jelan~Atoll: Nelle Isli?t, 4 Bokan Islet, 34164. hotho toil: iiotko islet,; 34215. Ujae ~,toll:~UjakIslet,-34304. Pokak Atoll: Kanome Islet, 34,535, 34518. Enivi'etok Atoll: Aomon Islet, 24359; Eniwetok islet, 2437:l.j En;;&e Islet, 2l+389, %14393,.ajae Atoll: Kvva jalein Islet, 26481. Wake Atol:i.: Peaie Islet, 31445. '' Indigenous. Very abmdsnt on -L'okak in open arezs; local in openings on drier ormore saline islets on other &t,olls; planted and prsisting.. after plating on more favorable islets 1,rhei.e not found naturally. wr- . . shallese:. .~~kio.fl -l'hespesia pwpulnea (L. ) Sol. ... . ~wajalexotcollect,ed). . . Introduced. 131gtud' in militbry esthblislment . &xaiian: "milo, I' Calophyllum inw,>hyllum L. s : Utirilc p.toll: UCirik Islet. ..33678. . Jemo Island: 33871. Ail& Atoll: Aiiuk Islet, 33926. uje.1arig Ato!.:.: Enellap Islet, 34139. Wotho Atoll: Wotho islet, 34228. Uj+e Ltoll: Yock Islet, 34343. Likiep Atol.1: Likieg . .. Islet., 2'7020. Of aboriginal introduction. planted in villages, especially along lagoon behcnes: occasionally established in mixed forest :lea,r lagoon side . of islets. On Medjurwon Isiet., k;otno, this species forms corispicuous . . thickets. Marshallese: %i$j," "luge j," "luech. " ......

wi2.acid& Forst. ... ' ' Utirik Atoll: ~tirikIslet, 33682. TaiiaAtoll: Taka Islet, 33'71~7. Likiep Atoll: Lado Islet, 3381+7.7kj.luk Jitoli: Ulika Islet, 33~34.iae Atoll: Lae Islet, 3LI.03?. .Kw+jalein iitoll': Jinebuoj Islet, 34l27; Bennett (Bikej) Islet, , %6&99. Ujel-ang Atoll: Ujelang Islet, 34188. Ujae Atoll: . . Bock Islet, 34334. liiotho .iitoll: Yokanuetok Islet, 341+42,. . Enioetok Atoll: Puiraai Islet, no.2690. Indigenous. Comon, grovtng especially on denuded rock flats; rarely on sand. Forming dense pure s-Lands. ~arshall.ese:!'ng,ier~gie,~~"ker~gi,~~ or ...... "kungi . .. . . ," . . :, ,; . .. Citrullw vulgaris Schrad.

Ehiffetok. Atoll: Wgebe Islet, ,24375.. , . . , Introduced. Rare3: persisting after irilitary. occupation. - ...... --Cucurf~s melo- Id. Eniwetok Atoll: En~ebe- Islet. < 21~499... . Introduced. Rare, persisting after military occup&tion. . , , . . , . . --Cucurbita maxima Duch, Ujelang ~toll:Yieto Islet, 3iJ47. Eniwetok Atoll: Ehijebe Islet, 243'/6, 24380 (?) (no gray spots on leaves). Introduced. Persisting after ~nilitaryoccupation; also planted by Marshallese on Ujelang.

Caricapapaya L. Utirik Atoll: Utirik Islet, 3960. Taka Atoll: Taka Islet, 33736. Jeino Islad: 33882. Ailtoll: Ailuir Islet, 33961. Lae Atoll: Lae Islet, 34059. Kwa jalein ~tol!.: Ehebuo j Islet, 341281 LT jelang Atoll: U jelang Islet, 34213. Ujae kLo1.i: Bock Islet, 31344. Introduced. 'Conkonly plaited in interiors of islets in coconut groves, sometjih& persisting tiiere. Fruit &ppreci;.ted by :iiarshalLese. Harshallese: "papaya. 1'

Bru~uieraconjup:,ata (L.) hierr. Utirik Atoll: Bekrak Islet, 33712. Aj.l.uk Atol.1: Ailuii ~sl'et, 33964. Lae Atoll: Lae Islet, 33993; Lo> Islet, 34043. Indigenous (?). Rare, in wet depressions. %rshallese: "chor:g, fl jong. It Terminalia catappa L. Likiep Atoll: Likiep Islet, 27019. Kwajalein Atoll: (no.: col.lected). Introduced. Rarely plaited in villages. Garshdlese :butil.

Terrnina1i.a samoe~sisRech. Ujelang Atoll: Bi&o Islet, ,?1+143; Enellap-Islet, 3$Lw. Utirik Atoll: Bekrak Islet, j3713. Taka iitoll: Taka Islet, 33739. Likiep Atoll: kdo Islet, 33783; Likiep Islet, 2699'7; Aiitini (~ek~rie). Islet, 27059.

Jemo Island: 33891.~~ ~ Ailuk ~tol!.: Aku'xe Islet, 3391.1. Lae Atoll: Ehenlanman Islet, 34037. I

Barrin,:tor;ia asiatica (L.) Xwrz Likiep Atoll: Lado Isle-t, 31860. Lac Atoll: Lae Islet, -,?k:)28. Indigenous,' Forming pu~eforest i+one area on Lae, seen as drift seed1i.n~on Likieu, not seen otheruise except as drift.fruits on beaches.

--Jussiaea -suffruticosa L. Likiep Atoll: Likiep Islet, 33853, 27044. Introtiuccd (?) weed. In ttaro pits.

Polyscias gellaria (Burm.f.) ~osb. filuk Atoll: kiluir Islet, 3395'). Lae iitoll,: Lae Islet, 34.066. Ujae Atoll: Ujae ~slet,14289. Kwajalein %ol'l: Ekeye Islet, 31211. ~nt~oduced.Occasionaliy planted around &l.ages;

Polysci.es sp. Likiep Atoll: Likiep Islet, 27034. Introduced. Rarely planteu. Centella asiatica (L.) Urb. Likiep Atoll: Lado Islet, 338.30; ~ikini(Aek-dne) Islet, 27064. Ailuk Atoll: Ailuk Islet, 33943. Ujae boll: Ujae Islet, 311298. Kwajalein Atoll: Kwaialeiz." Islet. 26L90... Local introduced weed in villages, plantztions, and around military establis!ments. Xarshallese : lllnsrukko, " ''margo, 'lmnariko .'1

, . -Catharanthus =us 1 ) .. Likiep Atoli: Lado: Islet, 33843; Likiep Isiet, 27037. Ailuk Atoll: /ii.l.uk Islet, 33.978. ,Wake Atoll: hake Islet 34~~56. . . , . . . Introduced. Planted around houses and graves, sohetiines. . persisting. <. . . . Nerj.im indicumUil1. L_ --, . . Ujae. Atol.1: Ujae Islet, 3429.i.' Likiep ktoll: Aikini (Aekuie) Islet, 27063...... -mtraduced. :Plaited ar.oune viliages.

Ne't-ium oleander L. Lae Atoli: Lae Islet, 34000.. U jao Atoll: Ujae Islet, 3!+293. Introduced. P1u;ted arowd vLlages. fjeriuni- sp. . . litirik toll: utirik 1:jlet j 336285. Introduced. Planted arounci .house.

Ochrosia oppositifolia (Lam. ) K. Schfim. Likiep Atoll: Likiep Islet,. 2 Ailuk Atoll: .~k.ulwe~slet, 33914. Lac Atoll: Enemannan.Islet, 34028. htja2.ein ikoll: Ehew?t& Islet, 34133; , Bennett (Bikej) Islet, 26492. Ujelsng Atoli: Ujelcng Uslet, ,34215. Tiotho Atoll: Viotho Islet, 34262, 34279. Ujae Atoll: Koeju Islet, 34377. Eniwetok ~Itoll:Japt= Islet, 24329. Indigenous. In mixed forest and, forning pure for-st$. ~iarshzllose: 'Jkabuijiling, 11 !'kijbar, '1. %e jebar, " "Ice j'~ar.~' : . .

Plumeria rubra L......

Utirik ni;oll: utirik :Islet, .31689.' J&O Island: ' 33865. 'iiikuk Atoll: Ailuk Islet, 33966. Laeiitoll: Lae islet, 0 Ujae Atoll: Ujae Islet, 34296. Likie~Atoll: Likiea Islet., 27021, Introduceti. Phi-Led around villages, i~&rshallese: "meria. 1' ...

&clepias. curassavica. L. . . Utiri~Atoll: Utiriii Islet, 33658. Ailuk Atoll: Aiiuk Islet, 33975. Lae p.tolL:. Lae %let.. 3~068.-. ... Introduced weed. Planted around houses rid paves, occa~io.g&~ : escaped. ldarshallsse: nyelo. " . . ,, , . .. umoea- --batatas L...... !5&e toll: !.jake Islet, 34449. Likiep iitol.l:l;ikiep Islet, 27038. Introduced. Rarely, plaited; not doing 1i;el.l; on hake persisting aromd Japnese garden site.

Ipomoea littoralis %.I. Lae Atoll: Loj Islet, 34053. Ujclang 1,toll.: Ujelafig Islet, 34202. Introducec: (?) veed. Occasional around viliaLes, hbuses and paths. --Ipomoea pes-caprae (L.) Roth. Lae Atoll: Lae Islet, 33399; Loj Islet, 34044. Wake Atoll: V!ake Is!.et, 34473. Enivretok H.toll: ai\;etok Islet, 24287; Eri~ebeIsl.et, 24404. Kwaja'bir~toll:Kwajalein Islet, 26480; ~er,nett(~fkej) Islet, 26506. Introduced (?) weed. Very abundant around military installiltions, especially abandoned ones; sezn eisawhere only as drift seedling. Ipomoea tuba (~chlecht.) G.Uon ka!ie Atoli: i,ake Islet., --'$36111, . utiril; Atcll: Utirik Islet,' 33661, 33677; Eluk Islet, 33700. Jemo lslund: 33890. Ailuk Ato1.l: Ai:i.u:c Lsl.et; 33942. Lae Atoll: Loj Islet, 3Lt056; Eiierlbao 1sle-c 31108:. Iiwajalein Atoll: Lo j jairong Islet, 34119; Kwajal.ein Islet, 2647%; benriett (~ikej) Islet, 26519. Ujelmg Atoll: Ujelang Is!.et, Sil78. YGoti-ii; Akli: Wotho Islet, 34239, Ujae toll: Bock Islet,. 34362, .Pokhk Atoll: Sibylla Islet, 34526. Ehiwetok toll: Eniwetok Islet, 24285; Igurin Islet, 2!+305; Japtan Islet, 24342; Aomon Islet, 24349. Idciep ktoll: Aikini (iieitijne) Islet, 27053. Indigenous. Very common to rSoundant in coconut plmtcstions, mixed Corest and scrub: also on open rock flats or1 Pokalc Atoll. Ih,rshallese: %mrbele, It Ifbeleu (Ujelang, ~niwetok). Cordia subcordata Lam. -utirik Atoll: Utirik Islet, 33671. Liiiiep Atoll: Lado Islet, 33784. Jemo Island: 33883. Lae toil: Lze Islet, 3j035; Ehenbso Islet,, 34082. Kwajalein Atoll: Enenetak Islet, 3i11.32. ~jelan~Atoli.: Ujelang Isl.et, 34265. Ujae ii.toll: Bock Islet, 31+3?2. wootho Atoll: :D1ea:5k Islet, 34427; Kab'oen Islet, 34435. i'take A';oll: yzake Isiet,, 34480. Fhiwetok Atoll.: Iguriii Islet, 24303; Japtm Islet, 24'38; Aonon Is:Let, 24351. Indigenous. Rare to copmori in mixeci. forest arid scrub, locally foriidng pure staqds. Ymrshallese: llk&o, 11 "kho. " lieliotropfum_ anomalum H. Bi A. Wake, Atoli: Uake Isl.et, 33625. Indigenous. Cornn~on on open ground between trees ard arou~dbuildings.

Eeliotropim ovalifoli.j;n var. (Cn:m.) ?+!err. I(waja1ein &dl:Kivajale.izl 'islet, jit124. introduced weed. 0cc~~sior:al;long air sstrip.

&jesserschinidiz 5rgentea (L.f .) Johnst. Wake ~tcll.:Wake Islet, 33615. Utirik ;

Cestrum r:octurnw~ L. Likie:p Atoll: Likiep Islet, 27035. Introduced. Rarely planted.

---fiicotiana -.--tabacum L. Nake ~toll:';.&ice islet, 144L18. Introduced. Persisting i..ft..r cultivation. riare, ;round old Japanese garden site. Physaiis --ang-ulata. I,. Jerno Island: 33872. USae ,@.toil: Ujae Islet, 31ii86. Kvmjalein Atoll.: IkiajaLein Islet, 34381. Iiltroduced weed. Occasional in disturbed places. Solanum lycopersicon-- L,. -Eniwetok Atoll: E~gebeIslet, 24374. Introuuced. Persisting after military occupa-tion.

Sol.am~mnigrm L. -Kwajalein toll: ltwajalein Islet, 31177, 311130. Introduced weed. Occ~sionalLi disturbed places around niiiitaly installations. Clerodendrum-- inerme (L. ) Gaerti:. Vcirik Atoil: Utirik Islet, 33673. Likiep ~toll:iado Islet, :38116; Likiep Islet, 27908. Jeno island: 331367. Ailuk ~toli:i let33Yl+5. Lac Atoll: Lae Islet, 34O67. Ujelcuig toll: Ujelang Islet, 34186. li~otho Atoll: ?fotho Islet, 34229. 1:wa.iaiei.i- iitoli: Zbeye Islet, 31213;-, Kwa.ialein-

Islet, 26468 (ultirited).< . -Indipnous. Cormion as uncie:c&ro!-th in coconut pla~italions. Also filling abimionea taro pits. Pianted near house o~~Jemo."l:;rshallese: glulij,lI Ilulej.11 -- --Premna -obtusifolia R.Br. Utirik .4toli: Utirik Islet.,--., 33696: Uekrak Islet., 33711. Likieu Atoll: Lado Islet, 3331, 3381+$1; Aiiiini (iieltiiile) Islet, 27061. Ail& ktoll: Ail& Islet, 33963. Lae ~tdl:Lne Islet, 3lrDil. Ujelang Atoll: Uje1ru:g Islet, 34203. ?;otho Atcil: Wkio Islet, 34250. Ujx Atoll.: Ujae Islet, 34418. Indiganous. .Occasional in mixed forest and arowd villtiges and houses. Uarshallese: "kaar J " llkar.fl Stachytarpheta jmaicei~,sis--- (L.) V~hl Likiep iitoll: Likiep IsLet, 2'1028. Introduced. Planted. in gsr3en.

Stacliytarpheta inuica Vahl Kwaja~zn-: hajalein Islet, 3w1, 31182. Introdwed. v;eed around xilitary insi;alLtions.. -Oci~nw.sanctum L. Utirilc Atoll: Utirik Islet, 33659. Ailuk Atoll: Biluk Islet, 33979. Lae Atoll: Loj Islet, 34052. Ujae Atoll: Ujae Islet, 31~292. ' . Lrtroduced. Planted in gordens. &rshsllese: I1katarin. --Russelia -etiformis Schl.echt. & Cham. Likiep ~toll:?dkiep Islet, 33858, 27046. ~ntroduced. F;arely planted-in gardens.

S~p.ariadulcis L. Kvrajaieiii Atoli: Kvrajalein Islet, 31184. Introduced weed. Rare around milihsry establishir,erits,

Graptophyllu;li pictwo (L. ) Griff, . . Likiep .&toll: Likiep Islet, 27023. . . Introduced. Rarely planted...... Pseuderanthem atropurpure~m(Eull.) Bailey ---.-.- - - Jeino Island: 33866. 1 0 Ailuk Islet, 23973. LaeAtoll: Lae .... Islet, 34063. Ujae toll: Ujae Islet, 34290, 34/+21.. ~wajaleinAtoLi:Ebeye ... Islet, 31206. Likiep Atoll: Likiep Islet, 27025...... -Introduced. Conmonly planted around vili.ai;es.

...... , . Pseuderanthe~nwn- carruthersii.Sebm. .: . . Utirik Atoll: Uti-rik.IsLet, 33690. Lne Htol.1: Lse Islet, 34098.. .. ~otho toll: ~othoislet, 34236. Ujae ki;oli: Ujae Islet, 34295. Kwaj~lein Atoll: Ebege Islet, 31296. Likiep J:to:LL: L-ikiep Isl.et, 27330. ~ntroduced. ~ommonly~lanted aroixid vilizges. "iie form ;:ith entire.1~ . - ... '..' green leaves aid white flowers riiti: crimson dots, in the center. hfars&dle.se:' nulij, 18 ltviut .tiroj I1 , . . . . .

-.- Guettarda speciosa L. ' . . Utirik i:toll: Utirik Islet, 33667. Taks Atoll: Takt; Jslet, 337/+0. Jemo Island: 33897. -Ailuk Atoll: Aihk Islet, 33934.. Lae Atoll: Lae islet, 34106. ~jel&gj\toll: U jelang Islet, 34196. j;otho ~tdl:Notho Islet, 34276. Ujae toll: Bock Islet, 31+316. Eniwetok Atoll: Aoinon Islet, 24364; Fniwetok Islet, 24294; Igurin Islet, 24312; Japtan Islet, 24318; &&be -.Islet, 24398. Kwjalein Atoll: Berinett (hej) Islet, 26~94...Lilciep Atoll: nkiep Islet, 27002. Indigenous. Common in mixed forest and scrub,..'also .in coconut groves. Yershallese: l%ut, It but i lormP.

Hedyotis biflora (L.) Laii. Liltiey, Atoll: Lsdo Islet, 33792; Likiep Islet, 27041; Aikini (~ekzne)Islet, 27049. Introduced (?) weed. Rare in coconut ~l~tations. --Ixora cmei [lance Likien P.toll: Likieu Islet, 27032. Introhuced. Rarely planted; . . Uorinda citriiolia L. ~tirikAtoll: Utirik Idet, 33684. Toka :,toll: Tlika Islet, 33734, 33745. Je:m Island: 33~7%.~iluk~~to~l: iiil~islet, 33955. Lac-iitoli:. Lae Islet, 34008. Ujeiang toll: Ujelmg Met, 34177. kotho iitoll: Wotho Islet, 34231. Ujtie Atoll: Bock Islet, 3vjl5. E:iweiok Atoll: Igurin Islet, 24302; Japtan Islet, 24335; Aomon Islet, 2435J; Engebe Islet; 24381. .. Kwajalein toll: Bennett (Bikej) Isict; 26537. Liltlep ktoll: Aikini (~ekzne)Islet, 27056. Of aboriginal introductiori. Coriunon in coconut and breadfruit groves and in mixed forest. uarshallese: Iben," "nin.11 --Scaevola -frutescens (hIill. ) Xrauise hake Atoll: kake Islet, 33616. Utirik toll:, Utirik Isletj 3915. Taka Atoll: Taka Islet, 337'20. Jemo Island: 33887. "~iluk'Atoll.: Aiiuk Islet, 33933. Lae Atoll: Lae Islet, 34090. Ujelang Atoll: Ujelang Islet, 34194. Wotho Atoll: V/otho Islet, 34275. Ujae Atoll: Eock Islet, 34318. Pokak Atoll: Pokak Islet, 3452/+. Bikair htoll: Bikar ,islet, 311.573. kiwetoic .,' .,' ,/' Atoll: Fniwetok Islet, 24291; Igurin Islet, 243il;;';Japt,m Islet, 21+3?6; & Aomon Islet, h4361; Engebe Islet, 24397. kajdeia Atoll: Kwajaiein Islet, 26471; Bennett (Bikej) Islet, 26516. Likiep Atoll: Likiep Islet, 26999. Indigenous. Coiimon in scrub and as under{;;r.owth in cyconut plantations, 'usually forms a fringe aloilg top of beach. Marshalle'se:"~annat,~~flmartt ' (~jelan~),lrmar kinat" (~nir~etok),rtkoimat.lt. Hippobroma longiflora (L.) G.Con Likiep ~tol1:i;ikieu Islet., 2'70L2... IntroGed weed. ~l&earound taro pits on.Likiep, now probably gone, asnatives were advised in 1.946 to destroy it; not seen in 1951. . . Adenoste!ma lavenia (L. ) O,.Xtze. ---I__.- Lae ~toll:Lae Islet, ?l+Wg. 1 . ...' . . . Introduced (?) weed. In;coconut and breadfruit groves, but only . . seen on Lac. i~larshallese: ubulibulgas. " . . . . Conyza bonariensis (L. ) Crony. Tj!ake ii.toll: Xake Islet, 34463. Kmjalein Atoll: Kwajalein Islet, 31168,.- 311'71. . Introduced weed. Common around air strips.

Conyza canaden& (L.) Cronq. KwajaZLein toll: Kwajalein Islet, 31169. Introduced weed. Around hirstrip. Ecliuta -alba (L. ) Hassk. Kwajalein toll: Kwajalein Islet, 31163. Introduced weed. Occasional around airstrip.

pmi1ia jwanica (Burm.f. ) Rob. Kwa jalein &toll: Kwa.jalein Islet, 31i86. ~ntroducedweed. Coin?lon around i&itar.y installations. Pluchea --indica (I,.) Less. Kwajalein Atoll: Kwajalein Islet, 26523; Ebeye Islet, 31210. Introduced weed. Abundant around-military irii-tallat;i&s.

Pluchea odorata (L.) Cass. hake toll: W;ke Islet, 341+47. Kwajalein Atoll: Kwajalein Islet, 31170; Ebeye Islet, 31208. Eniwetok Atoll: mgebe Islet, 2/.+405. Introdwed weed. Abundant arowd military insth11Gtions...... ,. ,Sonchus oleracea L. . . . , Wake toil: $Jake Islet, 3446&, 34483. Introduced weed. Iri disturbed web.

*medrella nodiflbra (L.) Gaertn. Kviajaiein ktoll: Kwajalein Islet, '3a20. , :' ~nt&uced weed. ho&d @litary instsllations, . -. : Tridak procumbens L. -Kwajalein Atoll: Kwajalein Iblkt, 31165. Introduced i~eed.~ro&d airstx+ii>...... Vernonia cineres. (L.) Less. . . Kwajalein Atoll: Ebeye Islet, 31209. Eniwetok titoll: '~2liv:etok Islet, 24368; beebe Islet, 24382. Likiep jitoll: .Likiep Islet, 269.70. Introduced weed. Common around ailitapy in;rjtailations; in Likiep village.

Wedelia biflora (L.) DC. ~iEie~Atoll: Lado Islet, 33819; Likiep Islet, 27003. Ail& Atoll: Harib Islet, 33983. Lse Atoll: Lae Islet, 34061. UJae Atoll: Ujae Islet, 34.301. Ehiwetok Atoll: Elleebe Islet, 24391. %vaj&ein toil:. . hvajalein Islet, 26476; Bennett (~ikej)islet, 26508. . . Indigenous, Common as ground cover and undergrowth in plantations, climbing in trees in mixed fore^.';, 'and forming derise thick bl~ketover abandoned military installations. ~arshull.e~e:~rriargueue,"llmamjej.u . . . .