A428 Black Cat to Caxton Gibbet improvements

TR010044 Volume 6 6.3 Environmental Statement Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species)

Planning Act 2008 Regulation 5(2)(a) Infrastructure Planning (Applications: Prescribed Forms and Procedure) Regulations 2009

26 February 2021

PCF XXX PRODUCT NAME | VERSION 1.0 | 25 SEPTEMBER 2013 | 5124654

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Infrastructure Planning

Planning Act 2008

The Infrastructure Planning (Applications: Prescribed Forms and Procedure) Regulations 2009

A428 Black Cat to Caxton Gibbet Improvements Development Consent Order 202[ ]

Appendix 8.16:Terrestrial Invertebrates (Including Non-Native Species)

Regulation Number Regulation 5(2)(a) Planning Inspectorate Scheme TR010044 Reference Application Document Reference TR010044/APP/6.3 Author A428 Black Cat to Caxton Gibbet improvements Project Team, Highways England

Version Date Status of Version Rev 1 26 February 2021 DCO Application

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.1

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Table of contents

Chapter Pages 1 Introduction 1 1.1 Background and scope of works 1 2 Legislation and policy 3 2.1 Legislation 3 2.2 Policy Framework 4 3 Methods 5 3.1 Survey Area 5 3.2 Desk study 5 3.3 Field survey: terrestrial invertebrates scoping survey 6 3.4 Field survey: terrestrial invertebrate survey 10 3.5 Biodiversity value 14 3.6 Competency of surveyors 17 3.7 Limitations 17 4 Results 19 4.1 Desk study results 19 4.2 Field survey results: terrestrial invertebrate survey overview 23 5 Summary and conclusions 55 6 References 56 6.1 Species list 57 Plates 96 Annex A: Figures 98

Figure 1. Invertebrate Survey Scoping Area ...... 98 Figure 2.1 Black Cat Roundabout: Areas A17, A18, G5, G6, P and Q (Part) ...... 99 Figure 2.2: Chawston (South-West Of), West of A1 Road, Area G4 (Incorporating A16 . 100 Figure 2.3. Quarry Area Q East of Black Cat Roundabout and R (Rectory Farm) ...... 101 Figure 2.4. Alington’s Land (Mainly) Near Little Barford, Areas H20, H22, H23, H24 And H25 ...... 102 Figure 2.5. Alington’s Land (Mainly) Near Little Barford, Areas H17, H19 And H20 ...... 103 [Please note there is no Figure 2.6] Figure 2.7. East of St Neots, Near Wintringham Hall And Fox Holes, Areas H16, W9 And W10 And Ash ...... 104 Figure 2.8. North of Croxton and Staggered A428 Junction, West of Minor Road, Areas H9, H10 And W7 ...... 105 Figure 2.9. North of Croxton and Staggered A428 Junction, East of Minor Road, Areas H5, H6 And H7 ...... 106 Figure 2.10. Eltisley Area G1 ...... 107 Figure 2.11. West of Caxton Gibbet Roundabout, South of A428, Areas H1 And H3 ...... 108

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.12. East of Caxton Gibbet Roundabout, South of A428, Areas A1 And W1 ...... 109

Table of Tables Table 2-1: Summary of Relevant Legislation for Terrestrial Invertebrate Species...... 3 Table 3-1: Areas included for invertebrate survey ...... 6 Table 3-2: Weather conditions during surveys ...... 10 Table 3-3: Importance of ecological features ...... 15 Table 4-1: Records returned from Cambridgeshire and Peterborough Environmental Records Centre...... 19 Table 4-2: Records returned from Bedfordshire and Luton Biodiversity Recording and Monitoring Centre...... 22 Table 4-3: Pantheon analysis of survey data organised by site codes...... 40 Table 4-4: Pantheon analysis of survey data organised by broad biotope and Habitat. .... 47 Table 6-1: Full species list for invertebrates captured during the survey...... 57

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

1 Introduction

1.1 Background and scope of works As part of the A428 Black Cat to Caxton Gibbet improvements Scheme (the Scheme), terrestrial invertebrate surveys including invasive non-natives species were carried out to inform the biodiversity assessment reported in Chapter 8, Biodiversity of the Environmental Statement [TR010044/APP/6.1]. These species could be potential constraints to the Scheme or influence the design and implementation of the Scheme. An extended Phase 1 Habitat Survey (Appendix 8.20 of the Environmental Statement [TR010044/APP/6.3]) identified and mapped the main habitats within a 1.2 kilometre-wide (0.7 mile-wide) corridor centred on the alignment of the Scheme as it was in 2016. Chapter 2, The Scheme of the Environmental Statement [TR010044/APP/6.1] provides a description of the Scheme. This appendix includes: a. Relevant legislation. b. Methods for desk and field-based assessments undertaken in 2018 to determine presence or absence of terrestrial invertebrates across the Survey Area (defined in 1.3) based on desk study and field survey visits. c. Competencies of the ecologists involved in undertaking the above surveys. d. Limitations to the surveys undertaken and any assumptions made as a result of incomplete data. e. Survey results. f. The approach for determining the nature conservation importance of terrestrial invertebrates’ populations recorded during the assessment. The information described in this appendix provides a baseline of terrestrial invertebrates’ activity recorded within the surveyed areas, including the wider area which has been used to inform the Environmental Impact Assessment (EIA) for the Scheme. Terrestrial invertebrate ecology Invertebrates comprise an enormous group of organisms, both in terms of number of species and number of individuals. There are almost thirty thousand species of macro invertebrates in Britain, excluding groups whose members are all microscopic. Invertebrates are important in both ecosystems functioning and in agricultural systems. As herbivores, predators, parasites and as a food source for other species, they are a vital element in terrestrial food chains. At the base of many food chains, arthropods are important components of the diets of invertebrates and birds and are also an integral part of the nutrient- and energy processing abilities of the soil.

Planning Inspectorate Scheme Ref: TR010044 1 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

The evaluation of habitats for overall potential for terrestrial invertebrates is a more practical method of assessing nature conservation value than individual species surveying. Ideal feeding areas include woodland, dead wood, scrub, marshy grassland, rough grassland and areas with a mosaic of habitats. Bare earth and early successional stages of vegetation can provide nesting and sunning habitats and are often of value to specialist invertebrates. Overwintering habitats are sought in terrestrial features such as field margins with long rank grassland, which tends to support more species than short swards, piles of dead wood, hedgerows, low dense scrub, below rocks and in stonewall crevices.

Planning Inspectorate Scheme Ref: TR010044 2 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

2 Legislation and policy

2.1 Legislation Whilst a large number of the UK invertebrate species are widespread and common, there are many threatened and/or vulnerable species that are afforded protection by legislation and policy. These include relevant international law, national wildlife legislation, and national and local biodiversity policies. Most of these measures aim to protect vulnerable species, but some protection against the spread of invasive species. A summary of the legislation relevant to terrestrial invertebrate species in England is presented in Table 2-1. Table 2-1: Summary of Relevant Legislation for Terrestrial Invertebrate Species.

Legislation Summary of Key Aspects

Directive 92/43/EEC on This Directive is enacted in the UK by The Conservation the Conservation of (Natural Habitats &c.) Regulations 1994 and The Natural Habitats and of Conservation of Habitats and Species Regulations 2010. The Wild Fauna and Flora annexes that include invertebrates are: (Habitats Directive) Annex IIa - designation of protected areas required within the natural range of the animal species listed Annex IVa - special protection required for the native animal species listed Annex Va - exploitation of listed animal species to be subject to management if necessary, in order to maintain their favourable conservation status. Schedule 2 of the Conservation of Habitats and Species Regulations lists those species of animals included in Annex IV(a) to the Habitats Directive that have a natural range that includes any area in Great Britain. It is illegal to trade in any of the species (or any part of the species) listed in Annex IVa of the Directive, whether or not they live in the UK.

Wildlife and Countryside Under Section 9 of this Act, wild animal species listed on Act 1981 (as amended) Schedule 5 may be protected under one, some or all of these Schedule 5, Section 9 parts: Part 1 – intentional killing, injuring, taking Part 2 - possession or control (live or dead animal, part or derivative) Part 4 (a) - intentional or reckless damage to or destruction of any structure or place used by a scheduled animal for shelter or protection Part 4 (b) - intentional disturbance of animal occupying such a structure or place

Planning Inspectorate Scheme Ref: TR010044 3 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Legislation Summary of Key Aspects Part 4 (c) - obstruction of access to any structure or place used for shelter or protection Part 5 (a) - selling, offering for sale, possessing or transporting for the purpose of sale (live or dead animal, part or derivative) Part 5 (b) - advertising for buying or selling live or dead animal, part or derivative

Wildlife and Countryside It is an offence to release in the wild any species listed on Act 1981 (as amended) Schedule 9. Schedule 9, Section 5

Natural Environment and Section 41 (S41) of the Act requires the Secretary of State to Rural Communities Act publish a list of habitats and species that are of principal 2006 importance for the conservation of biodiversity in England. Section 40 of the NERC Act places a duty on public authorities, such as ministers, local authorities and statutory undertakers, to have regard to the conservation of biodiversity in England and Wales, when carrying out their normal functions.

2.2 Policy Framework National and local planning policies relevant to biodiversity are not provided here. These are evaluated in the Environmental Statement, Volume 1 [TR010044/APP/6.1]. Overall, any development proposal should consider the identification and management of terrestrial invertebrate species to mitigate biodiversity and compensate for negative impacts like loss of important or notable habitats within or adjacent to the Scheme. The potential for adverse impacts needs to be considered during both the construction and operational stages. Habitat and Species Action Plans have been established in the different Biodiversity Action Plans covering the location of the Scheme (Cambridgeshire and Peterborough, South Cambridgeshire District Council and Bedfordshire and Luton Biodiversity Action Plans). These Habitat and Species Action Plans encourage planners and developers to consider the enhancement and management of habitats and species to ensure the recovery of some habitats to maintain their biodiversity value and protect species and sites.

Planning Inspectorate Scheme Ref: TR010044 4 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

3 Methods

3.1 Survey Area The Survey Area for the assessment of the presence or likely absence of terrestrial invertebrates was informed by published guidance (Ref 1-5) and the professional judgement of suitably qualified and experienced specialists. The Survey Area comprised the area within the Order Limits and a zone 100 metres beyond the Order Limits. Scoping was undertaken of all the habitats within the Survey Area and those that required specific assessment were surveyed using field surveys in 2018 as described below. The scoping included use of walkover surveys, consulting the Phase 1 Habitat Survey map (Appendix 8.20 of the Environmental Statement [TR010044/APP/6.3]) and analysis of aerial photography. The Survey Area for the assessment of the presence or likely absence of terrestrial invertebrates was informed by published guidance (Ref 1-5) and the professional judgement of suitably qualified and experienced specialists. The habitats which were surveyed are shown in Figure 1. Habitats within the Survey Area that would have no potential to be impacted, directly or indirectly, by the Scheme, e.g. those in the vicinity of the de-trunked section of the existing A428 for which there will be no associated works, were not surveyed. All suitable habitat within the Survey Area (Figure 1) was assessed prior to and following adoption of LA 108 (Ref 1-11). Any limitations to these surveys are acknowledged and explained in the Limitations section of this appendix with consideration given to any implications to the findings.

3.2 Desk study A desk study was undertaken based on the likely zone of influence of the Scheme on different ecological receptors and an understanding of the maximum distances typically considered by statutory consultees. Appropriate organisations were contacted to request records of notable and/or protected terrestrial invertebrate species that had been found in the Study Area, which comprised a zone within 1 kilometre (0.6 kilometres) of the Order Limits. Other resources used included aerial photography and Ordnance Survey maps which were used to evaluate habitat. The following records centres were contacted in November 2018 with a request for relevant records: a. Cambridgeshire and Peterborough Environmental Records Centre. b. Bedfordshire and Luton Biodiversity Recording and Monitoring Centre.

Planning Inspectorate Scheme Ref: TR010044 5 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

A search was undertaken to identify nature conservation designations and protected and notable habitats and species potentially relevant to the Scheme, including invasive non-native species listed in the Schedule 9 of the Wildlife and Countryside Act 1981 (as amended) (Ref 1-4). The desk study included an analysis of the information returned by the record centres and a review of previous survey reports undertaken on the study site in 2016 (i.e. the extended Phase 1 Habitat Survey report) (Appendix 8.20 of the Environmental Statement [TR010044/APP/6.3]) (Ref 1-1).

3.3 Field survey: terrestrial invertebrates scoping survey A scoping survey was undertaken on 2 and 3 May 2018 to assess habitats within the 100 meters Survey Area of the Scheme (Figure 1). This survey assessed 71 sites identified through desk study of aerial photography and habitat mapping. These sites were chosen for scoping as they held habitats with the potential to support insect assemblages or individual taxa of increased significance (local, county, regional or national significance) including hedgerows, woodland blocks, grasslands, post-industrial or recently disturbed ground, watercourse margins, inundation areas, and ponds or pools. Of the initial 71 sites identified through desk study, 31 target areas were recommended for further survey (see Figures 2 to 12). These were the areas identified as having the greatest potential to support rare or scarce invertebrates or notable invertebrate assemblages, or those protected by the Wildlife and Countryside Act 1981 (as amended) (Ref 1-4) or those that are listed on NERC S41(Ref 1-2). Table 3-1 gives brief notes of each area included in the survey: Table 3-1: Areas included for invertebrate survey

Site code and Habitat notes Grid reference figure number

A (ash) A large dead standing Ash (Fraxinus excelsior) with a TL2160 Figure 2.7 damaged trunk – notified by AECOM bat ecologists and not previously identified during the scoping survey.

A1 A linear attenuation pond running parallel to the existing TL2960, 3060 Figure 2.12 A148 carriageway, with extensive Common Reed (Phragmites australis), margin draw-down zone; no dicot emergent flora or marginals of note.

A16 Seasonal inundations within G4 grassland, including a TL1655 Figure 2.2 Reedmace (Typha latifolia) bed and also a larger permanent linear pool in post-industrial grassland; pool fringed with emergent vegetation.

A17 Seasonal inundation within G6; negligible draw-down zone TL1555 Figure 2.1 during scoping survey; pool dried out completely in summer.

Planning Inspectorate Scheme Ref: TR010044 6 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Site code and Habitat notes Grid reference figure number

A18 Scrub with Reedmace and Common Reed (Phragmites TL1555 Figure 2.1 australis) fringed pool and some willows (Salix sp.) Shaded mud margin with vegetation litter.

E A veteran English Elm (Ulmus procera) close to field TL20345880 boundary (not shown in Figures) with hollow in bole; notified late in survey.

G1 Rabbit-grazed (post-industrial) grassland with short turf and TL2759/TL2760 Figure 2.10 moss, with Nettles (Urtica dioica), Hawthorn (Crataegus Sp.) scrub and a small Ash-plantation. Also present are seasonally inundated areas of wetland/wet grassland.

G4 Large expanse of rabbit grazed grassland, some post- TL1555/TL1655 Figure 2.2 industrial, including trefoils, Bristly Ox-tongue (Helminthotheca echioides), Ground Ivy (Glechoma hederacea), Bramble (Rubus fruticosus), teasel, Medicks (Medicago lupulina), Vetch (Vicia sativa), Forget-me-not (Myosotis arvensis), Common Daisy (Bellis perennis), Dandelion (Taraxacum officinale), Ribwort Plantain (Plantago lanceolata), clovers and docks. Bordered by hedgerow with poplar saplings adjacent to and parallel to road and by the tree-lined ‘South Brook’ in the north of the site.

G5 Rabbit-grazed (post-industrial) short-turf grassland with TL1555 Figure 2.1 Chickweed (Stellaria media), Ragwort (Jacobaea vulgaris), St John’s Wort (Hypericum perforatum), cranesbills, clovers, Creeping Thistle (Cirsium arvense), Ground Ivy and Ribwort Plantain; a stand of elms present adjacent to minor road.

G6 Rabbit-grazed post-industrial grassland with large shallow TL1555 Figure 2.1 inundation and sand/gravel banks with rabbit burrows, covered in sparse prostrate plants (e.g. cudweeds), Ground Ivy, etc.

H1 Hedge with increasing frequency of tall Elms (up to 15m) TL2860/TL2960 Figure 2.11 towards and at west end, west of large Ash. Figure 2.12

H3 Hedge with many tall Elms, particularly at northern end. TL2860 Figure 2.11

H5 Hedge with deep, shaded-out ditch at base and with Ash, TL2560 Figure 2.9 Hawthorn and Elm (15m) - some dead.

H6 Hedge with Elder (Sambucus nigra), Rose (Rosa canina), TL2560 Figure 2.9 Blackthorn (Prunus spinosa) and elms (mid-way along), some 15m tall with dead examples.

Planning Inspectorate Scheme Ref: TR010044 7 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Site code and Habitat notes Grid reference figure number

H7 Hedge with Elm, Blackthorn and large Oaks with no TL2460 Figure 2.9 damage; and also, some dead Elms.

H9 Hedge along recently dredged ditch, mainly Ash and TL2360/TL2460 Figure 2.8 Hawthorn at east edge near road, but further west towards bend in ditch, with continuous stand of tall Elms, some 15m tall, some dead; more tall Elms in evidence west towards woodland plantation block W7, plus Blackthorn; grassland margin semi-improved.

H10 Hedge with Elm (tall), large Oaks (no limb damage noted) TL2360 Figure 2.8 and Rose.

H16 Hedgerow with abundant tall (15 metres) elms along road TL2159 Figure 2.7 edge of existing A428

H17 Hedge with oak, Blackthorn, Ash, Sycamore Maple (Acer TL1956 Figure 2.5 pseudoplatanus), Field Maple (Acer campestre), Norway Maple (Acer platanoides), Lime (Tilia x europaea), Rose, Hawthorn and Elm (some dead); at its eastern end, it bisects two areas of grassland (semi-improved) with vetches, Spear Thistle (Cirsium vulgare), Ribwort Plantain, Dandelion, Ragwort, Cow Parsley (Anthriscus Sylvestris), trefoil, Goat Willow (Salix caprea) saplings, Hawthorn saplings and Bramble; rabbit activity evident in northern section where there is clover, cranesbills, Field Wood-rush (Luzula campestris), and Creeping Buttercup (Ranunculus repens).

H19 Hedge with Field Maple, Hawthorn, oak, Rose, Bramble, TL1956 Figure 2.5 Dogwood (Cornus Sp.), Ash, Common Hazel (Corylus avellana), Elder, Blackthorn, and with a single Elm at east end near an oak with some limb damage (though not significant damage).

H20 Hedge with Blackthorn, Hawthorn, Elder, Ash, Sycamore, TL1956 Figure 2.4 Elm (some dead) and Bramble. Figure 2.5

H22 Hedge with Hawthorn, Elder, Bramble, Rose, Crab Apple TL1956 Figure 2.4 (Malus sylvestris), much Elm (most of hedge-length) – Plate 2 some dead.

H23 Hedge with Elder, Ash, Hawthorn and Crab Apple; some TL1955 Figure 2.4 Elms at easternmost end; potential invertebrate interest in several examples of large standing and fallen dead Ash with significant limb and bole damage, accessible heartwood etc.

H24 Hedge with Crab Apple, Elder, Ash, Hawthorn, Rose, TL1855/TL1955 Figure 2.4 Blackthorn (mainly) and Bramble – one hollow ash trunk

Planning Inspectorate Scheme Ref: TR010044 8 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Site code and Habitat notes Grid reference figure number towards western end – no Elm noted; south margin is weedy arable.

H25 Long hedge with oak, Hawthorn, Blackthorn, Rose, Elm TL1855 Figure 2.4 and Bramble – Elm mainly mid-way (4.5 to 6 metres) and towards west end (9m tall with dead examples); weedy arable margin on north edge.

P Pasture with horse dung in paddock immediately south of TL153552 Figure 2.1 A421, west of the existing Black Cat roundabout.

Q Scrub and grassland/wet grassland margin extending east TL1654/TL1655 Figure 2.1 to a linear north to south area sandwiched between active Figure 2.3 quarry works and the River Great Ouse; includes a pool with weedy draw-down zone at TL166552

R Small ditch with watercress, etc. in arable land west of TL173553 Figure 2.3 Rectory Farm.

W1 Linear woodland block with planted saplings along north- TL2960/TL3060 Figure 2.12 west edge, mainly Elder in interior with Elms (tall – 18 metres) along south edge and in east half; also, Blackthorn, Willow, Goat Willow, Ash, Crab Apple; ground flora of Nettle, Dog’s Mercury (Mercurialis perennis), Cowslip (Primula veris); some small dead Ash and larger fallen examples.

W7 Small plantation woodland containing some large Oaks and TL238604 Figure 2.8 an understorey of Nettle and Bramble situated immediately south of Gallow Brook, north of Croxton.

W9 Large damp woodland block with dense Hawthorn along TL2260 Figure 2.7 south perimeter; interior of Elms (abundant) and poplars with Elder, Hawthorn and privet understorey; ground flora of Ground Ivy, lords & ladies, garlic mustard, herb-robert on moss; some dead wood present.

W10 Woodland plantation block with cherry, crab apple, conifers, TL2259/TL2260 Figure 2.7 hazel, hawthorn and understorey of nettles; improves in west half with Elms more evident (particularly at far west end where there is a large block of tall Elms), a more open aspect and with more varied ground flora.

Survey timing Natural England report Surveying terrestrial and aquatic invertebrates for conservation evaluation (NERR005) (Drake et al., 2007) (Ref 1-5) suggests that 'a reasonable thorough survey of a terrestrial habitat can be made through seven visits at monthly intervals between April and October', but that 'four or five visits over this period will capture most species'. The timing and frequency of visits of the route survey was ideal for sampling spring and summer species.

Planning Inspectorate Scheme Ref: TR010044 9 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Terrestrial invertebrate surveys for most diurnal species groups should be undertaken during suitable weather conditions. Table 3-2 below lists the sites visited on each date and the weather conditions experienced. For site codes and descriptions see Table 3-1. Table 3-2: Weather conditions during surveys

Date Sites Visited Weather Conditions

May 23 A18, G1, G4, G5, G6, P, Q, R 12-17°C, NNE 12-15mph

June 6 A, H16, H17, H19, H20, H22, H23, 16-19°C, NNE 7-10mph H24, H25, W9, W10

June 7 G1, H1, H3, H5, H6, H7, H9, H10, 16-17°C, NE 9-11mph W1, W7

July 10 A1, G4, G5, G6, H1, H3, H16, W1 16-22°C, N 9-11mph

July 26 H17, H19, H20, H22, H23, H24, 26-33°C, mainly S 5-14mph H25

August 8 E, H6, H7, H9, H10, W9, W10 17-22°C, SW 9-12mph

No precipitation was experienced during the invertebrate survey visits detailed in Table 3-1. Areas P and R were also briefly surveyed during the scoping visits in May.

3.4 Field survey: terrestrial invertebrate survey The overall terrestrial invertebrate survey aim is to provide robust evidence for ecological quality of selected habitats or habitat features, based on their invertebrate fauna. The result of any site survey depends both on the amount of effort put into recording that location and the inherent ecological value of the location influenced by its size, geographical location, surrounding landscape and habitat biodiversity. For comparison to be most accurate, all locations within a site would have to be surveyed with the same measured effort, using standardised sampling techniques. The preferred methodology is to intuitively spend more time on areas that are obviously more diverse and that have the potential to support rare species or assemblages. An ‘exhaustive approach’ is taken, meaning that sampling is only stopped in a ‘productive’ area when new species cease to be recorded there. By using these criteria, there is a greater likelihood of finding at least some of the scarcer elements of the fauna that the site supports. A standardised methodology can miss these scarce species and produce a generalised list of nothing but common species.

Planning Inspectorate Scheme Ref: TR010044 10 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

The scoping survey identified the presence of Small-leaved Elm (Ulmus minor aggregate) throughout the site in hedgerows and woodland blocks. Elms are the foodplant of the larvae and adults of two NERC S41 Lepidoptera; White-letter Hairstreak (Satyrium w-album) and White-spotted Pinion (Cosmia diffinis). Direct searching and visual observation for the adults and larvae of these species was conducted in addition to the more general sampling. The terrestrial invertebrate surveys were conducted using best practice guidance based on Natural England Research Council Report 005 ‘Surveying Terrestrial and Aquatic Invertebrates for Conservation Evaluation’ (Ref 1-5). This guidance recommends several passive trapping, active trapping, and direct observational survey techniques that can be used and gives details on the terrestrial invertebrate taxa targeted by chosen techniques. The field techniques selected and utilised when undertaking the terrestrial invertebrate survey are detailed below: a. Sweeping. This was conducted by sweeping vegetation with a large heavy- duty net on a metal frame. This technique was favoured above all others. b. Grubbing: searching at ground level in vegetative litter, moss and under vegetative mats. c. Sieving: to sample invertebrates in vegetative litter, moss and under vegetative mats. d. Refugia search: Natural refugia such as large stones, logs and boards were lifted. e. Beating: A beating tray and pole were used to dislodge arboreal invertebrates from tree and scrub foliage. f. Close observation/spot checking: Studying small areas of exposed or sparsely vegetated ground for invertebrates or looking closely at vegetation and blossom to find invertebrates resting or feeding. g. Pitfall Trapping: installation of small pitfall traps filled with salt solution to trap nocturnally active ground roving taxa. Small vertebrates are prevented from falling by capping with a coarse mesh cap. These were left in place from May 23 to June 6 (used at Site G1 only). h. Dung search: searching in and around animal dung for invertebrate taxa, particularly the Scarabidae. (Used at Site P only). The insect groups that were targeted during the survey are detailed below: a. Coleoptera (beetles – all families). b. Dermaptera (earwigs). c. Diptera (larger Brachycera, Tephritidae and Syrphidae (hoverflies)). d. Hemiptera (true bugs, except psyllids and aphids). e. Hymenoptera (mainly sawflies and ants). f. Lepidoptera (butterflies and day-active moths).

Planning Inspectorate Scheme Ref: TR010044 11 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

g. Neuroptera (lacewings and allies). h. Mecoptera (scorpionflies). i. Odonata (dragonflies and damselflies). j. Orthoptera (bush crickets and grasshoppers). k. Trichoptera (caddisflies). Where possible, specimens were identified to species level whilst in the field, but where positive identification required the use of microscopic examination and identification literature (keys), specimens were collected. The bio-catches from each habitat and compartment were kept separate from one another inside of portable set of catches. Interpretation and analysis Methods Pantheon analysis The species lists for each survey location were processed and analysed using the Pantheon database tool. Pantheon is derived from and includes the previous Invertebrate Species-habitat Information System (ISIS) developed largely by Natural England in 2007 for the purpose of analysing species composition of a surveyed locality and interpreting this data in terms of habitat/species associations and species richness. Created by The Centre for Ecology & Hydrology in association with Natural England, Pantheon improves on the ISIS database tool by adding further analysis criteria including associated habitats and resources and habitat fidelity scores, against each taxon in a survey list. The Pantheon database tool is most effectively used where standardised sampling techniques have been employed in survey work. It enables comparison of resulting data from a fixed frequency of site visits over a fixed period and could indicate whether the ecological value of a site in terms of its invertebrate fauna, is either improving or deteriorating. This interpretation tool is less useful for surveys based on a 'snapshot' sample taken over a relatively short period and biased towards finding the scarcer invertebrates that a site supports. However, it is still a useful tool for producing a hierarchy of significance in terms of species habitat associations and assemblages at any given site and for comparing similar habitats which are surveyed at the same time of the year, using similar techniques and effort. The Pantheon database deals with around 11,000 invertebrate species, including all the most familiar and widely surveyed insect Orders. The scoring systems in Pantheon use species richness, threat status, rarity and characteristic species for each broad biotope, habitat and resource. The two Pantheon generated scores used in the interpretation of habitats and associated invertebrate assemblages are the Conservation Status and Species Quality Index (SQI).

Planning Inspectorate Scheme Ref: TR010044 12 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Each species recorded from a site list was scored according to its conservation status and the SQI was calculated by dividing this score by the number of species in the sample and multiplying by 100. SQIs for species lists with 15 or fewer species are understandably unreliable, so these index values have been removed from the resulting analyses in Section 1.4 - Results below. Conservation status Invertebrate surveys conducted between the late 1980s and 2010 relied in their interpretation of species recorded on British-specific rarity statuses for individual taxa, based on restricted distribution rather than population threat or risk. The term Nationally Scarce was applied to invertebrate species that were known to occur in 16 to 100 kilometre (9.9 to 62.1 miles) squares (hectads). The term Nationally Notable was also applied, this category was further split into Notable A (Na) for species occurring in 16 to 30 hectads of the National Grid and Notable B (Nb) for those occurring in 31 to 100 hectads. The category of ‘Red Data Book’ equating to ‘Nationally Rare’ was used for species recorded in 15 or fewer hectads in Britain and subdivided depending on the perceived or actual degree of rarity – e.g. RDB2 as Vulnerable, RDB1 as Endangered and RDBK as ‘Insufficiently Known’. Since 2010, IUCN Reviews have been produced for many invertebrate groups and are continuing to be written. In the IUCN Reviews, the restricted distribution categories have now been standardised to Nationally Rare (NR) and Nationally Scarce (NS) without further subdivision. The GB system of assessing rarity based solely on distribution is used alongside IUCN criteria which, whilst having geographical context, are primarily concerned with assessing National and International Threat. For the taxa found at the site the newly adopted GB Rarity categories 'NS' (Nationally Scarce) and ‘NR’ have been used where these appear in IUCN Reviews. Where IUCN reviews have not been written for the species recorded, the older categorisations of Nationally Scarce 'Notable Nb', 'Notable Na' and 'Notable' and Red Data Book species, 'RDB’ have been maintained. As further invertebrate groups are assessed for IUCN Reviews the current complex situation will become standardised The main categories in the IUCN Reviews for Threat status are ‘Least Concern’, ‘Near Threatened’, ‘Vulnerable’, ‘Endangered’, ‘Critically Endangered’ and ‘Extinct’. Species analysis is based on the area that it occupies and/or population statistics, and the magnitude of decline trends. BAP Priority Species (Research Only) Lepidoptera Priority Species “of principal importance for the purpose of conserving biodiversity” are covered under NERC Section 41, which requires these species to be taken into consideration by a public body when performing any of its functions with a view to conserving biodiversity.

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A number of Lepidoptera species are of National BAP Priority (Research Only) status under NERC S41. These Research Only Lepidoptera species differ from Priority Species in that they are not treated Nationally through formal individual Species Action Plans. There is concern that these Lepidoptera species have declined significantly in the UK in the last 25 to 35 years, despite still being common and widespread, they are flagged as S41 Research Only species to encourage awareness of their presence at sites and to promote recording and monitoring and are not intended to play a role in site protection.

3.5 Biodiversity value An essential prerequisite step to allow ecological impact assessment of the Scheme is an evaluation of the relative nature conservation value of the identified ecological features (encompassing nature conservation designations, ecosystems, habitat and species). This is necessary to set the terms of reference for the subsequent ecological impact assessment. The method of evaluation that has been utilised has been developed with reference to the CIEEM (Ref 1-6) and interpreted for terrestrial invertebrates by the professional judgement of Steve Lane, entomologist. These give advice on scoping and carrying out environmental assessments and place appraisal in the context of relevant policies. Data received through consultation, desk-based studies and field-based surveys are used to allow ecological features of nature conservation value or potential value to be identified, and the main factors contributing to their value described and related to available guidance. The value of terrestrial invertebrate species addressed in this appendix has been defined with reference to the geographical level at which the feature being assessed is considered to matter. Dispersal capacity and habitat connectivity has been considered to determine the ecological importance of terrestrial invertebrate species related to the Scheme. Table 3-1 describes each of the level of importance of ecological features. Guidance and criteria of potential relevance to the Habitat features and invertebrate assemblages being assessed is summarised is detailed in Table 3- 3.

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Table 3-3: Importance of ecological features

Importance of Typical Descriptors / Examples of Criteria Ecological Features

International or An internationally designated site or candidate site including; Special Protection European Areas (SPA); potential SPAs (pSPAs); Special Areas of Conservation (SAC); candidate or possible SACs (cSACs or pSACs1); and/ or Wetlands of International Importance (Ramsar sites). Biogenetic Reserves, World Heritage Sites and Biosphere Reserves. Areas which meet the published selection criteria for those sites listed above but which are not themselves designated as such. Resident or regularly occurring populations of species which may be considered at an international or European level2 where: a) The loss of these populations would adversely affect the conservation status or distribution of the species at this geographic scale; b) The population forms a critical part3 of a wider population at this scale; or, c) The species is at a critical phase4 of its life cycle at this scale.

UK or National Sites designated at national UK level e.g. Sites of Special Scientific Interest (SSSI); Marine Protection Areas (MPAs) including Marine Conservation Zones (MCZs); and National Nature Reserves (NNR). Areas which meet the published selection criteria for those sites listed above but which are not themselves designated as such. Areas of key/ priority habitats identified in the UK Post-2010 Biodiversity Framework i.e. UK Biodiversity Action Plan (BAP), including those published in accordance with Section 41 of the Natural Environment and Rural Communities Act (2006) and those considered to be of principal importance for the conservation of biodiversity. Areas of ancient woodland e.g. woodland listed within the Ancient Woodland Inventory. Resident or regularly occurring populations of species which may be considered at a national or UK level5 where: a) The loss of these populations would adversely affect the conservation status or distribution of the species at this geographic scale; b) The population forms a critical part of a wider population at this scale; or, c) The species is at a critical phase of its life cycle at this scale.

Regional Habitats or populations of species of value at a regional level (i.e. East Anglia). Areas of key/ priority habitat identified as being of Regional value in the appropriate Natural Area profile (or equivalent) now referred to as the National

Character Area (NCA).

Key/ priority habitat or species listed within the Highways England (HE) / Highways Agency (HA) BAP.

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Importance of Typical Descriptors / Examples of Criteria Ecological Features Resident or regularly occurring populations of species which may be considered at a regional level6 where:

a) The loss of these populations would adversely affect the conservation status or

distribution of the species at this geographic scale; b) The population forms a critical part of a wider population at this scale; or, c) The species is at a critical phase of its life cycle at this scale.

County or Unitary Habitats or populations of species of value at a County (i.e. Cambridgeshire) level Authority or or District (i.e. South Cambridgeshire). District Designated sites, such as County Wildlife Sites (CWS), Local Wildlife Sites (LWS) or Sites of Importance for Nature Conservation (SINCs), and Local Nature Reserves (LNRs) designated in the county or unitary authority area i.e. District context. Areas which meet the published selection criteria for those sites listed above but which are not themselves designated as such. Areas of key/ priority habitats identified in the Local Biodiversity Action Plan (LBAP). Resident or regularly occurring populations of species which may be considered at a County (or District) level7 where: a) The loss of these populations would adversely affect the conservation status or distribution of the species at this geographic scale; b) The population forms a critical part of a wider population at this scale; or, c) The species is at a critical phase of its life cycle at this scale.

Local Habitats or species populations of value in a local (i.e. within ~ 5km of the site) context. Designated sites include LNRs designated in the local context. Trees that are protected by Tree Preservation Orders (TPOs). Areas of habitat, or populations/ communities of species considered to appreciably enrich the habitat resource within the local context (such as veteran trees), including features of value for migration, dispersal or genetic exchange.

Site Habitat that is of value in the context of the site only. Populations of common and widespread species. A degraded/ impoverished example of a common or widespread habitat in the local area.

*As well as assigning importance there is also a need to identify all legally protected species that could be affected by the Scheme in order that measures can be taken to ensure that adherence to the relevant legislation is observed. This may include the adoption of mitigation and appropriate licensing which is acceptable to Natural England. 1 pSACs are sites which have been formally advised by to UK Government but have not yet been submitted to the European Commission. These sites should be valued at an international (European)

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Importance of Typical Descriptors / Examples of Criteria Ecological Features level on the basis that they meet the relevant selection criteria for a SAC but are not yet designated as such. 2 Such species include those listed within the Directive 2009/147/EC on the Conservation of wild birds (i.e. EC Birds Directive) (codified version of Council Directive 79/409/EEC as amended) or animal/ plant species listed within Council Directive 92/43/EEC on the Conservation of natural habitats and of wild flora and fauna (i.e. Habitats Directive). 3 Such populations include sub-populations that are essential to maintenance of metapopulation dynamics e.g. critical emigration/ immigration links between otherwise discrete populations. 4 Seasonal activity or behaviour upon which survival or reproduction depends. 5 Species which may be considered at the UK or national level means; birds, other animals and plants which receive legal protection in the basis of their conservation interest (those listed within the Wildlife and Countryside Act 1981 (as amended) Schedule 1, 5 and 8); species listed for their principal importance for biodiversity (in accordance with the Natural Environment and Communities Act 2006 Section 41 England) (Ref 1-2); priority species listed within the UK Post 2010 Biodiversity Framework (i.e. UKBAP); or species listed within the Red Data Book. 6 Such species include those listed in the appropriate National Character Area and key/ priority species. 7 Such species include those at county level (i.e. Bedford or Cambridge) including unitary authority area i.e. District level (e.g. Bedford Borough Council); as listed on the LBAPs; and listed as a county designated site.

3.6 Competency of surveyors The surveys were undertaken in line with AECOM’s Integrated Management System (IMS). Our IMS places great emphasis on professionalism, technical excellence, quality, environmental and Health and Safety management. All staff members are committed to establishing and maintaining our certification to the international standards BS EN ISO 9001:2015 and 14001:2015 and BS OHSAS 18001:2007. In addition, our IMS requires careful selection and monitoring of the performance of all sub-consultants and contractors. All AECOM Ecologists who worked on this project are members of (at the appropriate level) the Chartered Institute of Ecology and Environmental Management (CIEEM) and follow their code of professional conduct (Ref 1-3) when undertaking ecological work.

3.7 Limitations The aim of a desk study is to help characterise the baseline context of a scheme and provide valuable background information that would not be captured by a single site survey alone. Information obtained during the course of a Desk Study is dependent upon people and organisations having made and submitted records for the area of interest. As such, a lack of records for a particular habitats or species does not necessarily mean that the habitats or species do not occur in the study area. Likewise, the presence of records for particular habitats and

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species does not automatically mean that these still occur within the area of interest or are relevant in the context of the Scheme. Resolution of grid references varies between records. Some records have an accurate grid reference, meanwhile others were provided as a 1, 2 or 10- kilometre (0.6, 1.2 or 6.2 miles) squares. A survey can only assess the site as it was found at the time of the survey and if the habitats found on the site are subject to change, the results of this survey may no longer be representative of the site. At the time of the scoping and further terrestrial invertebrate surveys, access to some of the lands in the Survey Area was not possible due to landowner permission not being agreed (Figure 1). Wet weather in late May and subsequent warm conditions in early June impacted the survey opportunity for late instar larvae of White Spotted Pinion. This presents a minor constraint to the results regarding this species. A nesting lapwing discovered at Roxton (TL15505523) in May, prevented access to survey post-industrial grassland in this location. It is not considered that the access limitations to this location has caused any significant impact on the data obtained. An area of grassland with pools, situated immediately west of the A1 and south of Chawston (site G4/A16 Figure 2.2) had been altered significantly between the May and July survey visits. Habitats identified as suitable for survey were adversely impacted by industrial developments undertaken by a third party. However, this is not considered to have a major constraint to the results of the terrestrial invertebrate survey. Route re-alignments impacted upon a mature Elm at the corner of arable land at TL20345880 (site E) late in the survey programme. Survey of this tree conducted on August 8th, 2018 was unsatisfactory as notable saproxylic taxa would not be active as adults. This is not considered a constraint as further ecological and arboricultural surveys have influenced route re-alignments that removed direct impacts from this mature Elm.

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4 Results

4.1 Desk study results Two biological record centres were consulted for terrestrial invertebrates species within 1 kilometre (0.6 miles) of the Scheme, Cambridgeshire and Peterborough Environmental Records Centre (PERC) and Bedfordshire and Luton Biodiversity Recording and Monitoring Centre (BRMC). Due to the typically low levels of terrestrial invertebrate recording effort, records presented have not been limited to within last ten years as usually specified with vertebrate taxa. Data provided by PERC comprised 117 records representing 46 species. These records included 29 species that are included on Section 41 of the Natural Environment and Rural Communities Act 2006, and 17 species that have National significance designations. These records are presented in Table 4-1 below. Table 4-1: Records returned from Cambridgeshire and Peterborough Environmental Records Centre.

Common name Scientific Location Grid Precision Date Designation name reference (metres)

Aleochara Aleochara Croxton Park TL2559 1000 22/07/1998 Notable (N) (Xenochara) (Xenochara) CWS discipennis discipennis

Bloody Cranesbill Zacladus Railway TL193592 100 20/07/2010 Notable b (Nb) Weevil exiguus Meadow, St Neots

Cobweb Beetle Ctesias serra Croxton Park TL2559 1000 22/07/1998 Nb CWS

Cramp-Ball Platyrhinus Croxton Park TL2559 1000 22/07/1998 Nb Fungus Weevil resinosus CWS

Euglenes Euglenes Croxton Park TL2559 1000 22/07/1998 Nb oculatus oculatus CWS

Orthoperus Orthoperus Croxton Park TL2559 1000 22/07/1998 Nb nigrescens nigrescens CWS

Phytoecia Phytoecia Cambourne TL3159 1000 20/05/2004 Nb cylindrica cylindrica

Plegaderus Plegaderus Croxton Park TL2559 1000 22/07/1998 Nb dissectus dissectus CWS

Prionychus ater Prionychus ater Croxton Park TL2559 1000 22/07/1998 Nb CWS

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Common name Scientific Location Grid Precision Date Designation name reference (metres)

Scaphisoma Scaphisoma Croxton Park TL2559 1000 22/07/1998 Nb boleti boleti CWS

Sepedophilus Sepedophilus Croxton Park TL2559 1000 22/07/1998 N testaceus testaceus CWS

Tychius pusillus Tychius pusillus Railway TL193592 100 20/07/2010 N Meadow, St Neots

Small Heath Coenonympha Cambourne TL311598 100 05/09/2013 NERC S41 pamphilus

Wall Lasiommata St Neots TL16 1000 1985 NERC S41 megera

White-letter Satyrium w- Railway TL193592 100 24/07/2010 NERC S41 Hairstreak album Meadow, St Neots

Alder Signal Stathmopoda The Gorse, TL247614 100 21/07/2005 Nb pedella Yelling

Beaded Chestnut Agrochola Tithe Farm, St TL210605 100 27/09/2011 NERC S41 lychnidis Neots

Blood-Vein Timandra The Gorse, TL247614 100 26/08/2003 NERC S41 comae Yelling

Buff Ermine Spilosoma lutea Eynesbury, St TL188586 100 20/06/2003 NERC S41 Neots

Bulrush Veneer Calamotropha Yelling TL260614 100 25/07/2009 Nb paludella

Centre-barred Atethmia Tithe Farm, St TL210605 100 27/09/2011 NERC S41 Sallow centrago Neots

Cinnabar Tyria jacobaeae Crane Hill, TL1955 1000 01/07/2013 NERC S41 Waresley

Dark-barred Xanthorhoe The Gorse, TL247614 100 24/06/2003 NERC S41 Twin-spot Carpet ferrugata Yelling

Dot Moth Melanchra The Gorse, TL247614 100 17/07/2006 NERC S41 persicariae Yelling

Double Dart Graphiphora The Gorse, TL247614 100 10/07/2003 NERC S41 augur Yelling

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Common name Scientific Location Grid Precision Date Designation name reference (metres)

Dusky-lemon Cirrhia gilvago Tithe Farm, St TL210605 100 27/09/2011 NERC S41 Sallow Neots

Ghost Moth Hepialus humuli St Neots TL1960 1000 01/07/1992 NERC S41

Green-brindled Allophyes The Gorse, TL247614 100 14/10/2006 NERC S41 Crescent oxyacanthae Yelling

Grey Dagger Acronicta psi Eynesbury, St TL1858 1000 16/09/2013 NERC S41 Neots

Knot Grass Acronicta Tithe Farm, St TL210605 100 05/08/2011 NERC S41 rumicis Neots

Lackey Malacosoma Railway TL1959 1000 11/06/2013 NERC S41 neustria Meadow, St Neots

Large Nutmeg Apamea Tithe Farm, St TL210605 100 19/06/2012 NERC S41 anceps Neots

Long-legged Synaphe Eynesbury, St TL195579 100 15/07/2005 NERC S41 Tabby punctalis Neots

Mottled Rustic Caradrina Eynesbury, St TL188586 100 10/07/2010 NERC S41 morpheus Neots

Mouse Moth Amphipyra Eynesbury, St TL186585 100 03/09/2011 NERC S41 tragopoginis Neots

Oak Hook-tip Watsonalla Yelling TL260614 100 25/07/2009 NERC S41 binaria

Pale Eggar Trichiura Sir John's TL198552 100 25/08/2007 NERC S41 crataegi Wood, Crane Hill

Rustic Hoplodrina Tithe Farm, St TL210605 100 13/08/2013 NERC S41 blanda Neots

Sallow Cirrhia icteritia The Gorse, TL247614 100 25/09/2003 NERC S41 Yelling

Small Phoenix Ecliptopera The Gorse, TL247614 100 21/07/2005 NERC S41 silaceata Yelling

Small Square- Diarsia rubi Sir John's TL198552 100 25/08/2007 NERC S41 spot Wood, Crane Hill

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Common name Scientific Location Grid Precision Date Designation name reference (metres)

Sprawler Asteroscopus The Gorse, TL247614 100 05/11/2003 NERC S41 sphinx Yelling

White-spotted Cosmia diffinis Tithe Farm, St TL210605 100 20/08/2012 NERC S41 Pinion Neots

Chlorops racilis Chlorops Railway TL193592 100 13/06/2011 N gracilis Meadow, St Neots

Stratiomys Stratiomys Railway TL193592 100 13/06/2011 N potamida potamida Meadow, St Neots

Data provided by BRMC comprised 137 records representing 14 species. These records included seven species that are included on Section 41 of the Natural Environment and Rural Communities Act 2006, and five species that have National significance designations and two species with local interest group notations. These records are presented in Table 4-2 below. Table 4-2: Records returned from Bedfordshire and Luton Biodiversity Recording and Monitoring Centre.

Common name Scientific Location Grid Precision Date Designation name Reference (metres)

Ceutorhynchus Ceutorhynchus Begwary Brook TL1656 1000 16/05/1987 Nb constrictus constrictus Pits CWS

Small Heath Coenonympha Chawston Lake TL1556 1000 07/06/2016 GB- pamphilus RL2001:NT, NERC

Wall Lasiommata Tempsford TL15R 2000 12/06/1905 GB- megera RL2001:NT, NERC

White Admiral Limenitis Tempsford TL1653 1000 27/09/2014 GB- camilla RL2001:VU, NERC

Emerald Lestes sponsa Bedfordshire TL15 1000 25/04/1905 BedsDragon Damselfly

Scarce Chaser Libellula fulva River Great TL160540 100 30/05/2016 BedsDragon, Ouse CWS GB-RL2001:NT

Ectemnius Ectemnius Bedfordshire TL15 1000 01/01/1930 Nb (Clytochrysus) (Clytochrysus) - ruficornis ruficornis 31/12/1960

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Common name Scientific Location Grid Precision Date Designation name Reference (metres)

Small Spurred Nysson Bedfordshire TL15 1000 01/01/1930 Nb Digger Wasp dimidiatus - 31/12/1960

Little Black Wasp Pemphredon Bedfordshire TL15 1000 01/01/1930 GB-RLpre94:R (Cemonus) - lethifera 31/12/1960

Mouse Moth Amphipyra Wyboston TL15T 2000 15/09/1989 NERC tragopoginis

Small Square- Diarsia rubi Wyboston TL15T 2000 01/09/1989 NERC spot

Rosy Rustic Hydraecia Wyboston TL15T 2000 15/09/1989 NERC micacea

Blood-Vein Timandra Wyboston: TL1656 1000 06/06/2010 NERC comae Begwary Brook NR

Palliduphantes Palliduphantes Bedfordshire TL25 1000 01/06/1905 BedsSpider Insignis insignis

4.2 Field survey results: terrestrial invertebrate survey overview The terrestrial invertebrate survey was carried out in May, June, July and August 2018 (refer to Table 3-2, Section 1.3 for survey details). The surveyor specialises in British beetles and true bugs so these groups are particularly well-detailed in the resulting samples. A total of 661 invertebrate species (including aggregates of species, species complexes and unresolved species pairs) were recorded during six full days of field survey and one day of the scoping survey. This total includes 353 Coleoptera (beetles), 156 Hemiptera (true bugs), 41 Lepidoptera (butterflies and moths) and 36 Diptera (flies). No species that are afforded full protection under UK or International legislation were recorded during the survey. Three NERC Act 2006, Section 41 Lepidoptera species of BAP Priority status were recorded; the Cinnabar Moth (Tyria jacobaeae), the Small Heath Butterfly (Coenonympha pamphilus) and the White-spotted Pinion (Cosmia diffinis). No White-letter Hairstreak (Satyrium w-album) was recorded during the survey. No invasive non-native invertebrate species listed in Schedule 9 of the Wildlife and Countryside Act 1981 (Ref 1-4) were recorded during the terrestrial invertebrate survey.

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The non-native Elm Zig-zag Sawfly (Aproceros leucopoda) was recorded at 13 of the survey sites (E, G5, H1, H3, H5, H6, H7, H9, H10, H16, H20, H25, and W1) in June, July and August, typically identified from the characteristic zigzag larval feeding damage on elm leaves (Plate 3). First recorded in the UK in 2017, this species has the potential to adversely impact upon native elm species. The non-native Harlequin Ladybird (Harmonia axyridis) was recorded at 15 of the survey sites. This species was first introduced to the UK in 2004 and has since spread considerably. Monitoring of this species and potential negative impacts is ongoing. Two NERC S41 Research Only species were recorded, the Cinnabar Moth and the Small Heath Butterfly. Nationally Rare and Nationally Scarce species Two species recorded by the survey, the Small Heath butterfly and the Scarce Chaser (Libellula fulva) are designated as Near Threatened in recent IUCN Reviews. Thirty-eight species of Nationally Scarce (Notable, NS) status were recorded during the survey. See Table 6-1 below. Seven species of category Red Data Book status considered Nationally Rare were recorded; a. A flea weevil (Isochnus sequensi). b. Rusty Red Click Beetle (Elater ferrugineus). c. A rove beetle (Tachinus flavolimbatus). d. A tachinid fly (Cistogaster globose). e. A true bug (Anthocoris amplicollis). f. A leafhopper (Ribautodelphax imitans). g. White-spotted Pinion Moth (Cosmia diffinis). Brief summaries for the Nationally Rare, Nationally Scarce and IUCN-designated species found during the survey follow, ordered by Habitat-type. Statuses in square brackets are considered out-of-date and inaccurate based on current species distributions: Grassland and disturbed ground specialists a. A longhorn beetle (Phytoecia cylindrica) – Nationally Scarce (Nb Notable). Identification: A moderately large longhorn beetle, ash-grey with pale legs. Requirements: Grassland at the base of hedgerows, in field margins, on road verges and woodland edges where the food-plants, cow parsley, hogweed, wild carrot and rough chervil grow. Larvae bore into the stems and pupate at the stem bases, killing the plant in the process. Adults visit a variety of flowers.

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British distribution: Widely distributed south of a line from the Humber to the Bristol Channel with scattered records beyond this from Wales and England as far north as Yorkshire. Survey distribution: Adults were swept in areas G4, H9, H24 and Q in May and June. b. A flea beetle (Longitarsus lycopi) – Nationally Scarce NS. Identification: A small leaf beetle with modified hind femora for leaping. Very similar to a number of other species. Requirements: Short sward grassland with food-plants such as Ground Ivy (Glechoma hederacea), Calamint (Calamintha nepeta), Self-heal (Prunella vulgaris) and other labiates. British distribution: Widespread but local in southern England, becoming more local in northern England, Scotland and Wales. Survey distribution: 14 adults were recovered from pitfall traps in rabbit- grazed short sward grassland at Eltisley G1 in June. At this site, the food- plant is probably ground ivy c. A weevil (Calosirus terminates) – Nationally Scarce (Nb Notable) Identification: A small rounded weevil with distinctive white scale patch at the front of the wing cases and reddish colouration to the apices of the wing cases and the legs. Requirements: Grassland, including disturbed ground, where the Food-plant Wild Carrot (Daucus carota) occurs. Larvae feed in the stems and petioles of the food-plant. British distribution: Widespread but locally distributed throughout, with many records occurring close to coastal areas. Survey distribution: An adult was swept from post-industrial grassland next to the existing Black Cat roundabout at G6 in July. d. A weevil (Hypera meles) – [Nationally Scarce (Na Notable)]. Identification: A small non-descript brownish weevil Requirements: Grassland, roadside verges and field margins where it feeds on clovers, particularly White Clover (Trifolium repens). The larvae probably feed externally on the food-plant. British distribution: England as far north as Yorkshire and Wales. Most records appear to come from the Midlands and eastern England. This species is undoubtedly spreading and probably no longer merits its early 1990’s designation as Nationally Scarce. Survey distribution: Adults were swept from grassland at G4 and G6 in May and July.

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e. A weevil (Larinus planus) – Nationally Scarce (Nb Notable) Identification: A moderately large, cryptically coloured elongate weevil. Requirements: Feeds on a variety of thistles (Cardus spp. and Cirsium spp.) in grassland. British distribution: Primarily found in south-west England and Wales and particularly well-represented in coastal grassland localities. Possibly increasing in range in a similar pattern of population expansion as found recently in the closely related Thistle-feeder (Rhinocyllus conicus). Few records currently from eastern England. Survey distribution: A single adult was swept from spear thistle in disturbed rabbit-grazed grassland near the existing Black Cat roundabout, area G5 in May. f. A weevil (Rhinocyllus conicus) – [Nationally Scarce (Na Notable)] Identification: – A moderately large grey weevil, similar in appearance to Larinus. Requirements: Found in grassland where it is phytophagous on spear thistle and Musk Thistle (Cirsium natans). The adults are recorded from April to September and are known to hibernate in the turf mat. British distribution: Until recently, more-or-less confined geographically to the south coast of England, but it has very recently colonised inland counties in England and is certainly increasing, so its status of Na Notable is likely to be already obsolete. Its presence at the survey site is unexceptional. Survey distribution: Adult swept from the quarry edge at Site Q in May. g. A weevil (Zacladus exiguous) – Nationally Scarce (Nb Notable) Identification: A small black weevil with a strongly raised rim at the front of the pronotum Requirements: Phytophagous on several small-flowered Crane’s-bill (Geranium species). in grassland, typically verge habitats. The larvae feed at the roots of the food-plant. British distribution: Widespread but locally distributed in England and Wales; probably more frequent in the southern part of its range. Survey distribution: Adults were swept in grassland at G4, along an arable field margin at H19 and again on the margin of an arable field at R, all in May. h. A weevil (Catapion pubescens) – Nationally Scarce (Nb Notable) Identification: A small apionid weevil, grey-black in colour Requirements: Phytophagous on clovers (Trifolium spp.) in grassland habitats. The larva feeds in stem galls produced on the food-plant. British distribution: Mainly found in central and eastern England; predominantly coastal in the west, including Wales, as far north as Cumbria

Planning Inspectorate Scheme Ref: TR010044 26 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Survey distribution: An adult swept in grassland in a broad woodland ride at W9 in August. i. A rove beetle (Aleochara brevipennis) – Nationally Scarce (Notable) Identification: A small and nondescript brown rove beetle Requirements: Associated most commonly with the turf mat in grassland, but also found in other situations. Adults and larvae are probably both predatory on smaller invertebrates. British distribution: It is widely distributed but local in Britain. Some sources suggest that there have been recent declines, however this is disputable. Survey distribution: Adults sieved from moss etc in grassland at Eltisley area G1 in May. j. A tachinid fly (Cistogaster globosa) – Nationally Rare, Red Data Book (pRDB2, Vulnerable) Identification: A diminutive but distinctive fly, with a more-or-less shiny globose abdomen and the wings partly splayed when at rest. Requirements: The species is a parasite of heteropteran bugs, mainly the Bishop’s Mitre shieldbug (Aelia acuminate). The fly lays an egg on the dorsal surface of the host's abdomen and the larva when fully fed within the host, leaves the bug and pupates in the ground. The fly is typically found in dry grassland habitats. Adults have been observed to seek nectar on wild carrot. British distribution: Mainly found in southern England with a concentration of records in the south-east into East Anglia. Survey distribution: An adult was observed and captured in arable grassland at the elm site E in August. k. A lacehopper (Reptalus quinquecostatus)– Nationally Scarce (Nb Notable) Identification: A small hopper with transparent wings, similar to a number of closely-related species in Britain. Requirements: ‘The ecology of this species remains obscure, although it is often associated with grasslands in which the ground tends to crack during the summer. Since the nymphs are root-feeders, this perhaps allows the adults to lay eggs below ground’ (Ref 1-9). British distribution: Geographically restricted to southern England Survey distribution: An adult was swept at the weedy arable field margin at H25 in July. The ground here was cracked throughout the site, due no doubt in part to the drought conditions that had prevailed up to and around this time. l. A planthopper (Asiraca clavicornis) – Nationally Scarce (Nb Notable) Identification: Unmistakeable large leafhopper with flattened front legs. Requirements: Rough grassland and scrub.

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

British distribution: Previously more widespread in southern England, but now has a very restricted distribution: Thames estuary, a few areas in East Kent, the East Anglian Brecks and increasingly found in East Anglia away from this apparent stronghold. Survey distribution: Adults were swept on the margins of arable land at E and H9 in August. m. A planthopper (Ribautodelphax imitans) – Nationally Rare (Red Data Book RDBK) Identification: A small non-descript pale brown planthopper. Requirements: The hopper’s food-plant is Tall Fescue in grassland habitats. British distribution: Formerly rare in Britain but has either been overlooked or appears to be increasing its range. Old records are all in the Lyme Regis area, but ‘recent’ (post 1990) ones are from: Canvey Island, Essex, Portland, Dorset, Bincombe, Dorset, Axmouth, Devon, Cambridge, Cambs, Kingston, Sussex and Yarmouth, East Norfolk. There are further unconfirmed records from Northamptonshire, Huntingdonshire, South Lincolnshire and South Essex. Previously a BAP Priority Species and NERC S41 species, this status is likely to be removed in upcoming IUCN Review (pers. comm. Dr Alan Stewart, National Recorder, 2017). Survey distribution: Adults were swept from the grassland area adjacent to hedgerow H17, in July. n. A ground bug (Graptopletus lynceus) - Nationally Scarce (Nb Notable) Identification: A large and distinctively patterned ground bug Requirements: Dry sandy soils such as those found in dunes, the Brecks and in sand and chalk pits. The food-plant is Viper's bugloss (Echium vulgare). British distribution: Scattered records across southern England into Wales and some outliers in north-west England. Mainly found in the south-east. Survey distribution: Single adults were swept at the edge of the active sand quarry at Q and within the grassland sward at G4 in May. o. A ground bug (Megalonotus antennatus) - Nationally Scarce (Nb Notable) Identification: A brownish, ground-dwelling true bug with yellow legs and antennae. Requirements: The bug is believed to feed on seeds, although no specific plant species-associations are listed in the JNCC review. It is recorded from diverse habitat-types and substrates which include woodland rides, dry and wet grassland, disused clay workings and limestone quarries, most frequently found by grubbing, sieving and pitfall-trapping as it rarely strays from the ground. British distribution: Found sparsely in southern England but probably increasing.

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Survey distribution: An adult was sieved from moss in the grassland sward at G4 in May. p. Six-belted Clearwing (Bembecia ichneumoniformis) – Nationally Scarce (Nb Notable) Identification: A distinctive clearwing moth with yellow banding on the abdomen. Requirements: This moth inhabits calcareous soils including chalk downland, quarries and sea-cliffs. The foodplants are Bird's-foot Trefoil (Lotus corniculatus) or Kidney Vetch (Anthyllis vulneraria). The day-flying adults are on the wing between June and August. The larvae feed in the roots of the host plants. British distribution: Locally distributed in the southern half of Britain Survey distribution: An adult was swept from disturbed, rabbit-grazed short sward grassland at Black Cat junction, area G6, in July (see Plate 4). q. Small Heath (Coenonymphus pamphilus) – NERC S41 Priority Species, IUCN Near Threatened, LBAP Identification: A small orange-brown butterfly. Requirements: This moth inhabits well-drained, short-sward grassland sites where the larva utilises grasses of the genera Festuca, Poa and Agrostis. The flight period is from late April through to September with peak numbers usually in July and August. British distribution: Locally distributed throughout Britain. Population decline is ongoing. Estimated 27% decline since 1970’s and recent, due to habitat loss and fragmentation. Considered to be close to Vulnerable IUCN status (Ref 1-10). Survey distribution: An adult was observed in flight in grassland close to the hedge at H17 in July. Wetland specialists r. A weevil (Drupenatus nasturtii) – Nationally Scarce (Nb Notable) Identification: A small greenish-grey weevil. Requirements: Associated with the Food-plant Watercress (Nasturtium officinale) in ditches and streams. The species is not truly aquatic but the food-plant is always water-associated. Adults are usually recorded between April and October. British distribution: Widespread but locally distributed in England and Wales. Survey distribution: Adults were swept off the food-plant in the small open ditch site R on the approach to Rectory Farm in May. s. A weevil (Dorytomus filirostris) – Nationally Scarce (Nb Notable) Identification: An ash to light tan-coloured elongate weevil with a long rostrum and diagnostically long and thin antennal segments.

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Requirements: Found in wetland and fen habitats usually, where it is associated with Black and Italian Poplars (Populus nigra and P. nigra italica) which it feeds on. The larvae develop in the catkins. Adults have been recorded from May to October. British distribution: The species is widespread but local in central and eastern England. Survey distribution: Adults were beaten off poplars growing along the stream edge just east of the woodland block at W9, in June. t. A flea weevil (Isochnus sequensi) – [Nationally Rare, Red Data Book RDBK = insufficiently known] Identification: A small brown weevil with partly pale legs. Requirements: Found in wetland and at the drier edges of wetland features such as ditches and rides. As a larva it creates blotch mines in the leaves of Salix spp. pupating within the leaf mine. British distribution: Until recently, a rare species in Britain. First recorded in 1952 in Canterbury, Kent, it has since spread, being found in Sussex, Surrey, Essex, Middlesex, Norfolk, Cambridge and Huntingdonshire by 2012 and since then, appearing across most south and Midland counties of England. This is a species which is rapidly colonising northwards and consequently, its ‘official’ status of RDBK is no longer justified. Survey distribution: Found at G4 on Salix species. u. A rove beetle (Philonthus fumarius) – Nationally Scarce (Notable) Identification: A medium-sized black rove beetle with red legs. Requirements: Primarily a fen and marsh species, mostly associated with reed or sedge debris where it predates other invertebrates. British distribution: Distributed throughout England, but with an apparent bias for eastern England, particularly East Anglia and the south-east. There are scattered records north into Scotland. Survey distribution: Adults were sieved from wet grassland habitat at Eltisley G1 in May. v. A rove beetle (Platystethus nodifrons) – Nationally Scarce (Notable) Identification: A small, elongate black rove beetle with proportionately deep and distinct punctures on the wing cases. Requirements: Undisturbed pond margins and marshes with fluctuating water levels. Often found along river flood-plains. British distribution: Southern England north to Yorkshire, north Wales and north Scotland with most of the records in the Midlands and eastern England. Survey distribution: Recovered from pitfall traps in wet grassland at Eltisley, G1, in June.

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

w. Scarce Chaser (Libellula fulva) – Nationally Scarce, IUCN Near Threatened Identification: An unmistakeable libellulid dragonfly with a combination of dark patches at the base of the wings and either a bright orange abdomen with a central longitudinal dark streak (females and immatures) or a blue abdomen with terminal segments black (adult males). Requirements: Primarily found in wet meadows and in and at the edges of floodplains close to sluggish flowing water courses with substantial marginal vegetation. The larvae undertake an aquatic two-year development. Both adults and larvae are predatory on other invertebrates. British distribution: Southern and eastern England, with a much-fragmented distribution within its main range, although localised population expansions and colonisation have been noted in recent years. Survey distribution: Adult females or immatures were observed at close range at G4 (presumably associating either with the water bodies there or the South Brook (the latter heavily shaded though and therefore unlikely to support the species) and also at the edge of the River Great Ouse at Q in amongst willow carr/nettle beds etc, both sightings in May. x. A capsid bug (Agnocoris reclairei) – Nationally Scarce (Nb Notable) Identification: A medium-sized reddish-brown capsid bug. Requirements: Associated with its food-plant white willow (Salix alba). British distribution: It was first recorded in Britain in 1913 and has extended its range during the last thirty or forty years. It was previously a great rarity, restricted to fenland districts in Cambridgeshire, Huntingdonshire and Bedfordshire but is now more widely distributed in England although still relatively scarce, with recent records from Warwickshire, Kent, Northamptonshire and Derbyshire amongst other counties. Survey distribution: Adults were beaten off Salix along the south edge of South Brook at site G4 in July. y. Arboreal specialists including dead wood specialists (saproxylic and related species indicated in the following listings by SAP) An anthribid beetle (Anthribus nebulosus) – Nationally Scarce (Nb Notable) Identification: An ash-black convex beetle of non-descript appearance. Requirements: Found in mixed woodland and hedgerow habitats where it is associated with a variety of trees and shrubs. The larvae predate scale insects and the adults are most often encountered by beating trees and shrubs between April and July. British distribution: The species is widespread but local in England and has also been recorded from Wales and Scotland. Survey distribution: Adults were beaten from foliage along H1 in June and July.

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

z. An anthribid beetle (Platystomos albinus) - Nationally Scarce (Nb Notable) SAP Identification: A large, cryptically coloured black and white weevil which despite its appearance as a bird-dropping, is spectacularly marked and very conspicuous when active. Requirements: Associated with dead wood (hornbeam, oak, hazel, hawthorn) during its larval development. Recorded primarily from deciduous and pasture woodland. British distribution: Southern England and north to Westmorland. Survey distribution: An adult was beaten from dead boughs at H6 in June. aa. A soldier beetle (Malthodes pumilus) – Nationally Scarce (NS) SAP? Identification: A diminutive grey soldier beetle easily overlooked due to its small size and appearance. Requirements: “A poorly known species ecologically due to the wide range of situations in which it is to be found. It has been thought to develop in decaying heartwood of large old trees because many localities are ancient wood-pastures and well-wooded riverbanks. It can be swept in large numbers, for instance, beneath the canopy of old oaks in Moccas Park (J. Cooter, pers. comm.). However, it also occurs quite regularly on calcareous grassland sites in southern England, albeit usually those with some scrub. It may be that it is associated with epiphytes and saxicolous lichens, etc, rather than decaying wood, the common factor being bare surfaces which have been colonized by lichens. The larval habits remain very unclear however” (Ref 1-7). British distribution: Recorded from only around 80 hectads between 1980 and 2012. Widespread throughout Britain, but apparently more sparsely distributed in Scotland and Wales. Survey distribution: Widely distributed across the survey area with adults beaten from dead boughs of small dead-standing ash and elms in hedgerows at H5, H6, H10, H16, H19, H20, H23 and H25, all in June. bb. A soldier beetle (Rhagonycha lutea) – Nationally Scarce (NS) Identification: A moderately small, elongate soldier beetle with soft wing cases that are ochreous with black tips. Requirements: Usually associated with open woodland or scrubby calcareous grassland. Predatory, but also feeding on honeydew, nectar etc Larvae are thought to inhabit leaf litter on the ground. British distribution: Locally distributed in England and Wales, northwards up into Yorkshire. Survey distribution: Quite widely distributed across the survey site with adults beaten from various trees and shrubs at H1, H9, H10, H23 and W1, all in June.

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

cc. A longhorn beetle (Glaphyra umbellatarum) – Nationally Scarce (Nb Notable) SAP Identification: A distinctive elongate reddish-brown species with long antennae and short elytra leaving the wings are partly exposed. Requirements: Found in broad-leaved woodland, scrub and hedgerows. The larvae develop in dead wood of Rosa spp. and adults have been observed at flowers. British distribution: Southern England predominantly, but with records north to mid-west Yorkshire. Survey distribution: Adults were beaten from Rosaceous shrubs including Crab Apple (Malus sylvestris), at H7, H22, W1 and W10 in June. dd. A Clerid beetle (Opilo mollis) – Nationally Scarce (NS) SAP Identification: A distinctively marked, elongate, moderately-sized clerid beetle. Requirements: Associated with ancient broad-leaved woodland, pasture woodland and isolated trees. It is predatory on wood-boring beetles in the family Ptinidae (which include the woodworm beetle Anobium punctatum). The larvae live and develop inside the dead wood where they prey on the ptinid beetles, but they also eat dead wood and the contents of the hosts burrows. Adults have been noted under the bark of elm and Norway spruce as well as willow, poplar, beech and sycamore. British distribution: The species is widespread but locally distributed in south and central England with outlying records from north-east England and Wales. Survey distribution: Adults were beaten mainly off the boughs of small standing dead elms in hedgerows at H16, H22 and H25. ee. A clerid beetle (Tillus elongatus) – Nationally Scarce (NS) SAP Identification: An elongate beetle with serrate antennae, a usually red pronotum and blue-black vaguely metallic elytra. Requirements: Ancient broad-leaved and pasture woodland habitat where it is a predator on beetles of the family Anobiidae (Ptilinus, Pectinicornis and Anobium in particular), which it seeks out in larval galleries under the bark of dead standing trees. Adults can be found by beating flowering shrubs and have been recorded between April and September. British distribution: It is widely distributed throughout England and Wales, except for northern England where it is particularly scarce. Survey distribution: Adults were beaten from boughs of standing dead wood in hedges at H6, H9 and H25 in June. ff. A weevil (Anthonomus ulmi) – Nationally Scarce (Nb Notable) Identification: A small reddish-brown weevil with indistinct paler markings on the wing cases.

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Requirements: Broad-leaved woodland, parkland and hedgerows. Phytophagous on English and Wych Elms. Larval development occurs within flower buds. The adults having emerged, then aestivate during summer. British distribution: Distributed sparsely throughout Britain. Survey distribution: Adults were beaten from foliage at H9 and W9 in June. gg. A weevil (Polydrusus formosus) – [Nationally Scarce (Na Notable)] A medium-sized weevil covered in bright green scales, arboreal in habits and recorded from a wide variety of deciduous shrubs and trees. Until around twenty years ago, the species was relatively scarce in Britain, but it has since undergone a meteoric population expansion and is now frequent in south, central and eastern England at least. It is no longer Nationally Scarce despite its official status. Adults beaten from hedgerow at H17 in June. hh. Rusty Red Click Beetle (Elater ferrugineus) – Nationally Rare (Red Data Book RDB1 – Endangered) SAP Identification: A very distinctive and very large click beetle, mainly rust-red in colour, but with a black head, black hind edges to the pronotum and black legs. Requirements: Woodland, parkland and isolated trees. The larvae develop in black wood mould and red-rotten heartwood of deciduous trees, mainly in elm, beech and ash British distribution: The strongholds are the Thames Basin and East Anglia (Breckland is a main area for the beetle). Windsor Great Park was long known as a classic site for the species and it wasn’t until pheromone lures were produced that the species was found to be much more widespread. Adults are active in July and August and are rarely seen without the use of pheremone lures. Survey distribution: Two adults were attracted to a pheromone lure in the author’s shirt pocket which had survived the washing machine and was still surprisingly effective. The first adult alighted on Colin Smith’s back on the driveway at Top Farm, Alington’s Land (TL1905 5640) at 9.30 am, closely followed by another individual. That day was particularly hot and still. The origin of the beetles is unknown as there are no obvious breeding sites in the immediate vicinity. However parallels might be drawn with a report of female pregnant Barbastelles observed nearby at this site in the absence of any immediate large standing dead trees, which they too would require for habitation (Mike Padfield pers comm). Due to the location of origin not being known, the species has not been included in the data analyses at this site. ii. A melandryid beetle (Anisoxya fuscula) – Nationally Scarce (NS) SAP Identification: A small elongate convex, brown Melandryid beetle.

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Requirements: Associated during its larval development with dead wood, particularly the twigs of ash, willow, beech, field maple and lilac. It is found primarily in deciduous ancient woodland and pasture woodland but has been noted to occur in more open sites also. Adult beetles generally occur in the tree canopy. British distribution: Distributed mainly south of an imaginary line drawn from the Humber estuary in the east to the Severn estuary in the south-west. Not recorded in Scotland, but present in Wales. The species is sparsely distributed across eastern and central England with additional clusters of records around the Thames region and the Bristol Channel and outliers in northern England and Wales. Survey distribution: Adults were beaten from dead boughs, principally of ash, in hedges at H1, H16, H17 and H25 in July. jj. A melandryid beetle (Osphya bipunctata) – Nationally Scarce (NS) SAP Identification: A very distinctive beetle which occurs in orange and grey colour forms and with much variation in extent and patterning of colour on the elytra and pronotum. Males often have very well-developed, thick hind femora. Requirements: Ancient broad-leaved woodland primarily. The adults are usually observed on scrub blossom, most often flowering hawthorn, in May and June. The larvae probably develop in dead wood. British distribution: A scattered distribution from Somerset through the Midlands counties into East Anglia and down to Kent and Essex. There is also an old record for Yorkshire. Survey distribution: An adult was beaten off foliage in the hedgerow at H5 in June. kk. A mordellid beetle (Mordellistena neuwaldeggiana) – [Nationally Scarce (NS)] SAP A small orange-brown beetle that is characteristically convex with a distinctive elongated terminal abdominal segment and a jumping habit when captured. Designated as a Red Data Book species in the 1990s due to distribution based on conclusive records, was restricted to a handful of southern and south Midland counties. It has increasingly frequent in the last three decades and is now widely distributed across Midland and eastern England with records as far as Yorkshire in the north and into Devon and Wales in the west. The larvae develop in woody stems in the early stages of decay and the adults are often recorded visiting flowers or by general sweeping. Its most recent designation (Ref 1-8) is as Nationally Scarce (NS), but any British Rarity designation is currently disputable. ll. A mordellid beetle (Mordellistena variegate) – Nationally Scarce (NS) SAP Identification: A small tumbling flower beetle that has an elongated terminal abdominal segment and a jumping habit when captured or disturbed. Fuscous brown, patterned with darker markings.

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Requirements: The larvae develop in decaying wood. Adults are most often encountered visiting flowers such as flowering hawthorn and umbellifers in and at the edge of ‘wooded’ habitats. British distribution: Primarily distributed in Midland, south-east and eastern England. Very scarce elsewhere in England. Survey distribution: Adults were swept at G4 and H1, in June and July. mm. A salpingid beetle (Lissodema denticolle) – Nationally Scarce (NS) SAP Identification: A small brown beetle with pale reddish-yellow markings. Requirements: Found in woodland, pasture woodland, hedgerow and isolated trees, usually in dead wood or under bark, but also by sweeping under and around trees. Found on a variety of tree species, but with a probable preference for ash. The larvae probably develop in the dead wood. British distribution: Widespread but local in England Survey distribution: Adults were widespread throughout the survey area and were beaten from dead boughs of small standing dead ash trees in hedgerows at H1, H3, H6, H9, H16, H17, H22, H25 and also at W1, in June, July and August. nn. A Darkling beetle (Pseudocistela ceramboides) – Nationally Scarce (NS) SAP Identification: A distinctive and relatively large beetle with yellow-brown elytra, a black head and pronotum and long serrate antennae. Requirements: Larvae inhabit wood-mould of hollow decayed oaks, but also beech, generally beneath bird nests and most often in red-rot. Adults can be found at blossom, particularly at flowering hawthorn. British distribution: Locally distributed throughout southern and Midland England, scarce elsewhere in England. Survey distribution: Adults were beaten off foliage at H23 and W1 in June. oo. A tue bug (Anthocoris amplicollis) – Nationally Rare (Red Data Book RDB3, Rare) Identification: A small, relatively dark Anthocorid bug identified by its relatively large size for that genus, the location of matt versus glossy sheen on the wings and the shape of the pronotum. Requirements: Predatory on invertebrates on ash trees. Little is known of its ecology. British distribution: First recorded in Britain in 1982 from specimens taken between 1978 and 1980 and until relatively recently, known only from North- east Yorkshire (hence the National status designation of RDB). Now also known from East Suffolk (Breckland district) and other sites in England (personal communication M. Telfer).

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Survey distribution: Adults were beaten off ash trees at H17, H23 and H24 in July. pp. A leafhopper (Iassus scutellaris) – [Nationally Scarce (Na Notable)] Identification: A relatively large plant-hopper, more-or-less green in colour and can sometimes appear similar to the relative Iassus lanio found on oak. Requirements: Feeds on elm foliage. British distribution: Found in south-east England up through the Midlands and as far north as Lincolnshire. Expanding in distribution and the Nationally Scarce designation for this species is disputable. Survey distribution: Widely distributed across the survey site on elms. Adults beaten off the host tree at E, G5, H1, H3, H6, H7, H9, H10, H16, H20, H25, W1 and W10, in July and August. qq. Brown Ant (Lasius brunneus) – Nationally Scarce (Na Notable) Identification: A small ant identifiable by the pale brown head and trunk contrasting with the darker black-brown gaster (abdomen). Requirements: This is a species that nests in old mature trees and also stumps etc in hedgerows. It is perhaps mainly associated with oak. The adults feed on honeydew collected from large tree aphids although they may supplement this with small invertebrates. British distribution: The ant has only been recorded from southern and central English counties, from Essex to Shropshire. Survey distribution: Colonies were recorded at H5, H6, H9, H23 and H25 in May, June and August.

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

rr. White-Spotted Pinion (Cosmia diffinis) - Nationally Rare (Red Data Book, pRDB), NERC S41 Identification: Adults are distinctive by way of the large pure white marks on the leading edge of the forewing. These are more defined and larger than in the other superficially similar Pinion Moths. The larva is particularly diagnostic, being green with a series of parallel longitudinal white stripes and having a black or dark brown head capsule and black pro-legs. Requirements: Found in woodland, copses, shelterbelts, hedgerows etc. The larvae feed on English elm, small-leaved elm and apparently also wych elm. They are purported to prefer epicornic growth (suckers growing directly from the tree trunk). The adult flight period is from late July to early September. The overwintering stage is an egg. Larvae are present on the food-plant from April through to mid-June. British distribution: The species experienced a dramatic decline because of Dutch elm disease during the 1970s and it is now sparsely distributed in parts of southern England with its stronghold in the Huntingdonshire, Cambridgeshire, Bedfordshire region. Survey distribution: Larvae were located at H20 and W10. At H20, a single fully-grown larva was beaten off tall (20’) elms in a hedgerow at Top Farm, Little Barford (TL192564) on June 6th (see Plates. 4, 5 and 6). On the same date, a final instar larva was beaten off elms at the extreme north-west corner of the plantation block at W10. Here the elms were very tall, approximately 30-40’ high and forming a dense stand along the edge of the woodland. Other Species (non-specific Habitat) ss. A flea beetle (Psylliodes luteolaIa) – Nationally Scarce (NS) Identification: A small orange beetle with enlarged hind femora adapted for leaping. There are several similar species. Requirements: Various habitats including woodland, hedgerows, grassland. Feeds primarily on cultivated grass crops (wheat, barley etc) and wild grasses, but also known to skeletonise oak leaves. British distribution: Distribution is centralised on south and southern central England as far north as Derbyshire, with the strongholds perhaps in the southern Midlands (Oxfordshire and south Warwickshire). Survey distribution: A single adult was beaten off oak on the southern edge of H19 in July.

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

tt. A rove beetle (Tachinus flavolimbatus) – [Nationally Rare (Red Data Book RDBK)] This small blackish rove beetle with tapered abdomen and yellow margins to the fore-body, is associated with open ground including gardens, fields, coastal habitats and the foreshore where it is found in a variety of decaying organic matter, including dung. There are an increasing number of British records (well over 50), from West Norfolk, West Kent and Cambridgeshire. Its main centre of distribution appears to be East Anglia, and in Norfolk alone, it has been recorded from over 15 hectads. In the forthcoming IUCN Review for this group, the British Rarity status is to be downgraded so it no longer merits RDB status. On the survey, an adult was swept at G4 in May. uu. ThElm Zig-zag Sawfly (Aproceros leucopoda) - non-native Several adults were beaten off elms during a survey visit on June 6, 2018. Once identification was confirmed, prompt communication with AECOM and with Highways Agency followed, as it was thought to be the first occurrence in Britain of a notifiable and invasive species. However, it transpired that the species had already been recorded in Britain; in Surrey in autumn 2017. The species appears to have been found quite widely throughout southern and eastern England in 2018. This species was recorded from areas E, G5, H1, H3, H5, H6, H7, H9, H10, H16, H20, H25, W1, in June, July and August. The June records were of adults, but the characteristic zigzag larval feeding damage on elm leaves was noted widely on visits in July and August (e.g. Plate 3). The concern is that this species could potentially cause wholesale defoliation of elms if the population is large enough. Elm-feeding species of conservation importance in Britain such as White-letter Hairstreak and White- spotted Pinion have the potential to be adversely affected by loss of feeding sites. Interpretation and analysis Pantheon analysis The results for the Pantheon data analysis for grouped habitats in the survey area are shown in Table 4-3 and Table 4-4.

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Table 4-3: Pantheon analysis of survey data organised by site codes (see Table 3-1 and Figs. 2-12 for locations and descriptions of sites).

Sites Broad Habitat No. of SQI Species with Conservation Favourable (Codes) biotope species Score conservation status status

A, H16 wetland marshland 1 N

A, H16 tree- shaded 4 N associated woodland floor

A, H16 open short sward 9 N habitats & bare ground

A, H16 tree- decaying 15 180 4 NS|NS|NS|NS N associated wood

A, H16 tree- arboreal 16 123 1 NA N associated

A, H16 open tall sward & 51 100 1 Section 41 N habitats scrub Priority Species - research only

A1, W1 open short sward 6 N habitats & bare ground

A1, W1 wetland marshland 12 N

A1, W1 tree- arboreal 17 138 2 NA|NS N associated

A1, W1 tree- decaying 27 146 5 NS|DD N associated wood (European)|NA| Na|NS|NS

A1, W1 open tall sward & 33 100 1 Section 41 N habitats scrub Priority Species - research only

A18, P tree- arboreal 1 N associated

A18, P wetland marshland 1 N

A18, P open short sward 1 N habitats & bare ground

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Sites Broad Habitat No. of SQI Species with Conservation Favourable (Codes) biotope species Score conservation status status

A18, P tree- decaying 6 N associated wood

A18, P open tall sward & 33 100 N habitats scrub

E open short sward 1 1 Nb N habitats & bare ground

E tree- decaying 2 N associated wood

E tree- arboreal 6 1 NA N associated

E open tall sward & 16 100 N habitats scrub

G1 open upland 1 N habitats

G1 wetland wet 3 1 Notable N woodland

G1 tree- arboreal 3 N associated

G1 tree- wet 3 1 Notable N associated woodland

G1 tree- shaded 5 1 Notable N associated woodland floor

G1 tree- decaying 7 N associated wood

G1 wetland marshland 10 1 Notable N

G1 open short sward 12 1 NS N habitats & bare ground

G1 open tall sward & 39 100 N habitats scrub

G4 tree- wet 2 N associated woodland

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Sites Broad Habitat No. of SQI Species with Conservation Favourable (Codes) biotope species Score conservation status status

G4 wetland wet 2 N woodland

G4 wetland running 4 1 NT N water

G4 tree- shaded 5 N associated woodland floor

G4 tree- decaying 7 N associated wood

G4 wetland marshland 8 N

G4 open short sward 24 113 1 Nb N habitats & bare ground

G4 tree- arboreal 25 125 2 [RDB K]|RDB N associated K|Nb

G4 open tall sward & 76 116 5 Nb|[Nb]|Section N habitats scrub 41 Priority Species - research only|Nb|Nb

G5, G6 tree- decaying 1 N associated wood

G5, G6 wetland lake 1 N

G5, G6 wetland running 1 N water

G5, G6 wetland marshland 2 N

G5, G6 tree- shaded 2 N associated woodland floor

G5, G6 tree- arboreal 8 1 NA N associated

G5, G6 open short sward 23 152 3 [Nb]|NS|Nb N habitats & bare ground

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Sites Broad Habitat No. of SQI Species with Conservation Favourable (Codes) biotope species Score conservation status status

G5, G6 open tall sward & 69 104 2 [Nb]|Section 41 N habitats scrub Priority Species - research only

H1, H3 wetland marshland 1 N

H1, H3 open short sward 2 N habitats & bare ground

H1, H3 tree- shaded 3 N associated woodland floor

H1, H3 tree- decaying 12 2 NS|NS N associated wood

H1, H3 tree- arboreal 23 141 3 NS|Nb|NA N associated

H1, H3 open tall sward & 46 100 N habitats scrub

H17, tree- shaded 1 N H19, H20 associated woodland floor

H17, wetland marshland 1 N H19, H20

H17, open short sward 8 1 Section 41 N H19, H20 habitats & bare Priority ground Species|NT

H17, tree- decaying 19 150 4 |NS|NS|NS N H19, H20 associated wood

H17, tree- arboreal 38 143 4 NA|[Na]|RDB N H19, H20 associated 3|Section 41 Priority Species

H17, open tall sward & 63 114 4 Nb|NS|RDB N H19, H20 habitats scrub K|Section 41 Priority Species|Sectio n 41 Priority Species - research only

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Sites Broad Habitat No. of SQI Species with Conservation Favourable (Codes) biotope species Score conservation status status

H22, wetland marshland 2 N H23, H24, H25

H22, tree- shaded 3 N H23, associated woodland H24, H25 floor

H22, open short sward 7 1 Nb N H23, habitats & bare H24, H25 ground

H22, tree- arboreal 33 139 3 NS|NA|RDB 3 N H23, associated H24, H25

H22, tree- decaying 37 192 10 NS|NS|NS|NS| YES H23, associated wood NS|NS|NA|Nb| H24, H25 NS|NA|Na

H22, open tall sward & 70 104 1 Nb N H23, habitats scrub H24, H25

H5, H6, wetland wet 1 N H7 woodland

H5, H6, tree- wet 1 N H7 associated woodland

H5, H6, open short sward 3 N H7 habitats & bare ground

H5, H6, wetland marshland 3 N H7

H5, H6, tree- shaded 4 N H7 associated woodland floor

H5, H6, tree- arboreal 27 112 1 NA N H7 associated

H5, H6, tree- decaying 36 167 8 [Nb]|NA|Na|NS| YES H7 associated wood NS|NS|NA|Nb| NS

H5, H6, open tall sward & 42 100 N H7 habitats scrub

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Sites Broad Habitat No. of SQI Species with Conservation Favourable (Codes) biotope species Score conservation status status

H9, H10, wetland marshland 3 N W7

H9, H10, tree- shaded 5 N W7 associated woodland floor

H9, H10, open short sward 5 1 Nb N W7 habitats & bare ground

H9, H10, tree- decaying 27 158 4 NS|NS|NS|NA N W7 associated wood

H9, H10, tree- arboreal 28 136 3 NA|[Nb]|NS N W7 associated

H9, H10, open tall sward & 46 107 2 Section 41 N W7 habitats scrub Priority Species - research only|Nb

Q tree- shaded 1 N associated woodland floor

Q wetland running 3 1 NT N water

Q tree- decaying 3 N associated wood

Q wetland marshland 4 N

Q tree- arboreal 6 N associated

Q open short sward 7 3 NS|[Nb]|Nb N habitats & bare ground

Q open tall sward & 34 109 1 Nb N habitats scrub

R wetland lake 1 1 [Nb] N

R open short sward 1 N habitats & bare ground

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Sites Broad Habitat No. of SQI Species with Conservation Favourable (Codes) biotope species Score conservation status status

R wetland marshland 2 N

R wetland running 3 1 [Nb] N water

R open tall sward & 9 1 Nb N habitats scrub

W10 wetland marshland 1 N

W10 open short sward 2 N habitats & bare ground

W10 tree- decaying 7 1 Na|NA N associated wood

W10 tree- arboreal 11 2 Section 41 N associated Priority Species|NA

W10 open tall sward & 35 100 N habitats scrub

W9 wetland wet 1 N woodland

W9 tree- wet 1 N associated woodland

W9 tree- shaded 2 N associated woodland floor

W9 open short sward 3 N habitats & bare ground

W9 tree- arboreal 9 2 Nb|[Nb] N associated

W9 tree- decaying 10 1 Nb N associated wood

W9 open tall sward & 33 109 1 [Nb] N habitats scrub

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Table 4-4: Pantheon analysis of survey data organised by broad biotope and Habitat. (for Site Codes, locations and site descriptions, see Table 3-1 and Figs. 2- 12).

Sites Broad Habitat Species SQI Species Conservation Favourable (Codes) biotope Score with status (Y/N) conservatio n status

G4 open short sward 24 113 1 Nb N habitats & bare ground

G5, G6 open short sward 23 152 3 [Nb]|NS|Nb N habitats & bare ground

A, H16 open short sward 9 N habitats & bare ground

A1, W1 open short sward 6 N habitats & bare ground

A18, P open short sward 1 N habitats & bare ground

E open short sward 1 1 Nb N habitats & bare ground

G1 open short sward 12 1 NS N habitats & bare ground

H1, H3 open short sward 2 N habitats & bare ground

H17, H19, open short sward 8 1 Section 41 N H20 habitats & bare Priority ground Species|NT

H22, H23, open short sward 7 1 Nb N H24, H25 habitats & bare ground

H5, H6, H7 open short sward 3 N habitats & bare ground

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Sites Broad Habitat Species SQI Species Conservation Favourable (Codes) biotope Score with status (Y/N) conservatio n status

H9, H10, open short sward 5 1 Nb N W7 habitats & bare ground

Q open short sward 7 3 NS|[Nb]|Nb N habitats & bare ground

R open short sward 1 N habitats & bare ground

W10 open short sward 2 N habitats & bare ground

W9 open short sward 3 N habitats & bare ground

A, H16 open tall sward & 51 100 1 Section 41 N habitats scrub Priority Species - research only

A1, W1 open tall sward & 33 100 1 Section 41 N habitats scrub Priority Species - research only

A18, P open tall sward & 33 100 N habitats scrub

E open tall sward & 16 100 N habitats scrub

G1 open tall sward & 39 100 N habitats scrub

H1, H3 open tall sward & 46 100 N habitats scrub

H5, H6, H7 open tall sward & 42 100 N habitats scrub

W10 open tall sward & 35 100 N habitats scrub

G5, G6 open tall sward & 69 104 2 [Nb]|Section 41 N habitats scrub Priority Species - research only

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Sites Broad Habitat Species SQI Species Conservation Favourable (Codes) biotope Score with status (Y/N) conservatio n status

H22, H23, open tall sward & 70 104 1 Nb N H24, H25 habitats scrub

H9, H10, open tall sward & 46 107 2 Section 41 N W7 habitats scrub Priority Species - research only|Nb

Q open tall sward & 34 109 1 Nb N habitats scrub

W9 open tall sward & 33 109 1 [Nb] N habitats scrub

H17, H19, open tall sward & 63 114 4 Nb|NS|RDB N H20 habitats scrub K|Section 41 Priority Species|Section 41 Priority Species - research only

G4 open tall sward & 76 116 5 Nb|[Nb]|Section N habitats scrub 41 Priority Species - research only|Nb|Nb

R open tall sward & 9 1 Nb N habitats scrub

H5, H6, H7 tree- arboreal 27 112 1 NA N associated

A, H16 tree- arboreal 16 123 1 NA N associated

G4 tree- arboreal 25 125 2 [RDB K]|RDB N associated K|Nb

H9, H10, tree- arboreal 28 136 3 NA|[Nb]|NS N W7 associated

A1, W1 tree- arboreal 17 138 2 NA|NS N associated

H22, H23, tree- arboreal 33 139 3 NS|NA|RDB 3 N H24, H25 associated

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Sites Broad Habitat Species SQI Species Conservation Favourable (Codes) biotope Score with status (Y/N) conservatio n status

H1, H3 tree- arboreal 23 141 3 NS|Nb|NA N associated

H17, H19, tree- arboreal 38 143 4 NA|[Na]|RDB N H20 associated 3|Section 41 Priority Species

A18, P tree- arboreal 1 N associated

E tree- arboreal 6 1 NA N associated

G1 tree- arboreal 3 N associated

G5, G6 tree- arboreal 8 1 NA N associated

Q tree- arboreal 6 N associated

W10 tree- arboreal 11 2 Section 41 N associated Priority Species|NA

W9 tree- arboreal 9 2 Nb|[Nb] N associated

A1, W1 tree- decaying 27 146 5 NS|DD N associated wood (European)|NA| Na|NS|NS

H9, H10, tree- decaying 27 158 4 NS|NS|NS|NA N W7 associated wood

H5, H6, H7 tree- decaying 36 167 8 [Nb]|NA|Na|NS| YES associated wood NS|NS|NA|Nb|N S

A, H16 tree- decaying 15 180 4 NS|NS|NS|NS N associated wood

H22, H23, tree- decaying 37 192 10 NS|NS|NS|NS|N YES H24, H25 associated wood S|NS|NA|Nb|NS| NA|Na

H17, H19, tree- decaying 19 150 4 NS|NS|NS N H20 associated wood

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Sites Broad Habitat Species SQI Species Conservation Favourable (Codes) biotope Score with status (Y/N) conservatio n status

A18, P tree- decaying 6 N associated wood

E tree- decaying 2 N associated wood

G1 tree- decaying 7 N associated wood

G4 tree- decaying 7 N associated wood

G5, G6 tree- decaying 1 N associated wood

H1, H3 tree- decaying 12 2 NS|NS N associated wood

Q tree- decaying 3 N associated wood

W10 tree- decaying 7 1 Na|NA N associated wood

W9 tree- decaying 10 1 Nb N associated wood

A, H16 tree- shaded 4 N associated woodland floor

G1 tree- shaded 5 1 Notable N associated woodland floor

G4 tree- shaded 5 N associated woodland floor

G5, G6 tree- shaded 2 N associated woodland floor

H1, H3 tree- shaded 3 N associated woodland floor

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Sites Broad Habitat Species SQI Species Conservation Favourable (Codes) biotope Score with status (Y/N) conservatio n status

H17, H19, tree- shaded 1 N H20 associated woodland floor

H22, H23, tree- shaded 3 N H24, H25 associated woodland floor

H5, H6, H7 tree- shaded 4 N associated woodland floor

H9, H10, tree- shaded 5 N W7 associated woodland floor

Q tree- shaded 1 N associated woodland floor

W9 tree- shaded 2 N associated woodland floor

G1 tree- wet 3 1 Notable N associated woodland

G4 tree- wet 2 N associated woodland

H5, H6, H7 tree- wet 1 N associated woodland

W9 tree- wet 1 N associated woodland

G5, G6 wetland lake 1 N

R wetland lake 1 1 [Nb] N

A, H16 wetland marshland 1 N

A1, W1 wetland marshland 12 N

A18, P wetland marshland 1 N

G1 wetland marshland 10 1 Notable N

G4 wetland marshland 8 N

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Sites Broad Habitat Species SQI Species Conservation Favourable (Codes) biotope Score with status (Y/N) conservatio n status

G5, G6 wetland marshland 2 N

H1, H3 wetland marshland 1 N

H17, H19, wetland marshland 1 N H20

H22, H23, wetland marshland 2 N H24, H25

H5, H6, H7 wetland marshland 3 N

H9, H10, wetland marshland 3 N W7

Q wetland marshland 4 N

R wetland marshland 2 N

W10 wetland marshland 1 N

G4 wetland running 4 1 NT N water

G5, G6 wetland running 1 N water

Q wetland running 3 1 NT N water

R wetland running 3 1 [Nb] N water

G1 wetland wet 3 1 Notable N woodland

G4 wetland wet 2 N woodland

H5, H6, H7 wetland wet 1 N woodland

W9 wetland wet 1 N woodland

Yellow-highlighted analysis results are those for which the SQI equals or exceeds 150 (Table 6-1). This figure is chosen as it flags up the highest scoring sites in the survey and indicates the location of notable invertebrate assemblages.

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Biodiversity evaluation The principal findings of the invertebrate survey are as follows. The confirmed presence of White-spotted Pinion moth at H20 and W10 is significant because the two locations for this BAP Priority NERC S41 species have possible implications for the Scheme of the new dual carriageway. The presence of the BAP Priority NERC S41, IUCN Near Threatened Small Heath butterfly in the survey area should be a consideration in any mitigation strategy. The survey also evidences wood fauna of between County and Regional Importance, which is utilising standing dead ash and elm in many of the hedgerows surveyed. This is particularly significant at the H5, H6 and H7 grouping as the habitat here scored as ‘Favourable’ in Pantheon analysis; a score which could be construed as placing the importance of the assemblages here at or close to SSSI standard and therefore of National or near-National importance. However, due to the exhaustive and non-standardised sampling methodology, it is sensible to consider these clusters as above County and below Regional Level significance. The same is true of hedgerows H22, H23, H24 and H25. The highest SQI scores (scores of 150 or more) are achieved in analysis for dead wood resource and its associated invertebrates at H5, H6, H7, H10, H16 (and A), H17, H19, H20, H22, H23, H24, H25. All of the hedgerows in the target survey area with the exception of H1 and H3, are shown to be important for their saproxylic fauna. Due to the high scoring SQI values, hedgerows H10, H16 (and A), H17, H19 and H20 are evaluated as of County Level Importance. The hedgerows selected for full survey were the best examples identified in the scoping survey. It is likely that the saproxylic fauna will be of some importance, even if only at a Local level, in any other hedgerows and woodland blocks that contain standing dead elm and ash elsewhere within the Survey Area. Depending on the precise land-take required by the Scheme, some of the hedgerow areas within the 100 meters Survey Area of the Scheme are expected to be exposed to major adverse impacts as a results of habitat loss. Due to their composition which incorporates a significant resource of dead standing trees, these habitats have inherent inability to absorb change and it is unlikely that it would be possible to re-create these habitats in the short term. The grassland areas close to the existing Black Cat roundabout (G5 and G6) score highly with an SQI equal to or exceeding a value of 150 in the analysis. On the basis of the species assemblages in this combined small area, these locations would be assessed as between Local and County Level importance. The invertebrate assemblages associated with the other sites surveyed are unremarkable based on the findings of this survey and so these are evaluated as only being of Site Importance. These are sites A1, A18, E (see ‘Limitations’ in Section 3.7 of this Appendix), G1, G4, H1, H3, P, Q, R, W1 and W9.

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5 Summary and conclusions

A desk study, an extended Phase 1 Habitat Survey and a terrestrial invertebrate survey were carried out for the Scheme to look for, among other ecological receptors, protected and/or notable invertebrate species or assemblages. The results of a two-day scoping survey in May 2018 indicated that further survey work for invertebrates should be commissioned along the Scheme. This further work was undertaken over the course of six visits during May, June, July and August 2018. There were 661 species recorded from the survey visits. Thirty-eight of these species are designated as Nationally Scarce (Notable, NS) in terms of their British rarity. Seven species of category Red Data Book status (considered to be Nationally Rare) were recorded; the flea weevil (Isochnus sequensi), the Rusty Red Click Beetle (Elater ferrugineus), the rove beetle (Tachinus flavolimbatus), the tachinid fly (Cistogaster globose), the true bug (Anthocoris amplicollis), the leafhopper (Ribautodelphax imitans) and the White-spotted Pinion moth (Cosmia diffinis). Three NERC Section 41 species observed were the White-spotted Pinion, Cinnabar Moth (Tyria jacobaeae) and Small Heath butterfly (Coenonympha pamphilus). The White-spotted Pinion moth is of BAP Priority conservation significance, but the latter two species are of BAP Priority (Research Only) status. Of these, Small Heath is Near Threatened IUCN status but has no British rarity status and Cinnabar has no IUCN or British Rarity status. In practice, the NERC legislation requires all local authorities to consider biodiversity when undertaking their public duty. No fully legally-protected species were found during the survey. In terms of conservation value, hedgerows in two areas of the survey (hedgerow group H5, H6 and H7 and group H22, H23, H24 and H25) were found by analysis to be of above County importance but below Regional importance in terms of the dead wood habitat and associated invertebrate assemblages present. The majority of the other hedgerows surveyed were evaluated as being of County importance for their dead wood habitat and associated saproxylic invertebrate fauna. Two closely-associated small areas of rabbit-grazed short sward grassland were evaluated as being of above Local to County importance for invertebrates of open habitats.

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6 References

Ref 1-1. Jacobs UK LTD (2016a). A428 Black Cat to Caxton Gibbet Stage 1 Environmental Assessment Report.

Ref 1-2. Natural Environment and Rural Communities Act 2006. HMSO (2006). https://www.legislation.gov.uk/ukpga/2006/16/contents

Ref 1-3. Professional Code of Conduct. Chartered Institute of Ecology and Environmental Management (CIEEM), January 2019.

Ref 1-4. Wildlife & Countryside Act 1981. HMSO (1981). https://www.legislation.gov.uk/ukpga/1981/69

Ref 1-5. Drake, C. M, Lott, D. A, Alexander, K. N. A. and Webb, J. (2007). Surveying terrestrial and freshwater invertebrates for conservation evaluation. Natural England Research Report NERR005.

Ref 1-6. Chartered Institute of Ecology and Environmental Management (CIEEM) (2018). Guidelines for Ecological Impact Assessment in the UK and Ireland: Terrestrial, Freshwater, Coastal and Marine, September 2018.

Ref 1-7. Alexander, K.N.A. (2014). A review of the beetles of Great Britain: The Soldier Beetles and their Allies. Species Status No.16. Natural England Commissioned Reports, Number 134.

Ref 1-8. Alexander, K.N.A, Dodd, S. and Denton, J.S. (2014). A review of the beetles of Great Britain: The darkling beetles and their allies. Species Status No. 18. Natural England Commissioned Reports, Number 148.

Ref 1-9. Bantock, T. and Botting, J. (2018). An online identification guide to British Hemiptera. Available on: http://www.britishbugs.org.uk

Ref 1-10. Fox, R., Warren, M., Brereton, T.M., Roy, D.B. and Robinson, A. (2010). A new Red List of British Butterflies. Insect Conservation and Diversity doi: 10.1111/j.1752-4598.2010.00117.x

Ref 1-11. Design Manual for Roads and Bridges (DMRB) LA 108 Biodiversity (2020). Biodiversity.

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6.1 Species list Table 6-1: Full species list for invertebrates captured during the survey.

(*Yellow-highlighted analysis results are those for which the SQI equals or exceeds 150)

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Araneae – Araneidae – Araneus diadematus Garden G6, H17 7 Spiders Orb Web Spider Spiders

Araneidae Araniella cucurbitina H7, W1 6 sens. str.

Araneidae Araniella opisthographa H1, H5, H6, 6 H16, H17, H23

Araneidae Gibbaranea gibbosa H25 6

Araneidae Nuctenea umbratica H5, H23, W1 6,7

Clubionidae Clubiona brevipes H17 7

Clubionidae Clubiona pallidula H1 6

Gnaphosidae Drassyllus pusillus G1 6

Hahniidae Hahnia nava G1 6

Lycosidae – Alopecosa pulverulenta G1 6 Wolf Spiders

Lycosidae Arctosa leopardus G1 6

Lycosidae Pardosa prativaga G1 6

Lycosidae Pirata latitans G1 6

Lycosidae Trochosa ruricola G1 6

Philodromidae Philodromus aureolus H16, H17 6

Philodromidae Tibellus oblongus H10, H17 7,8

Pisauridae Pisaura mirabilis H17, W9 5,7,8

Tetragnathidae Pachygnatha degeeri G1 6

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Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Araneae – Theridiidae Phylloneta impressa formerly H17 6 Spiders Theridion impressum

Thomisidae Ozyptila simplex G1 6

Thomisidae Xysticus cristatus G1 6

Coleoptera Anthribidae Anthribus nebulosus H1 6,7 NS (Nb – Beetles Notable)

Anthribidae Platystomos albinus H6 6 NS (Nb Notable)

Apionidae – Apion frumentarium G4, G5, G6, R 5,7 Weevils (part)

Apionidae Apion haematodes E 8

Apionidae Aspidapion aeneum G5, G6 7

Apionidae Aspidapion radiolus G5, G6 5,7

Apionidae Catapion pubescens W9 8 NS (Nb Notable)

Apionidae Ceratapion carduorum G4, G5 7

Apionidae Ceratapion gibbirostre H16, H25 7

Apionidae Ceratapion onopordi Q, W1 5,6

Apionidae Holotrichapion pisi G4, G5 5

Apionidae Ischnopterapion loti G6 5,7

Apionidae Malvapion malvae G5 5,7

Apionidae Perapion hydrolapathi G5, G6 7

Apionidae Perapion violaceum H10, H24, R, 5,6,7 W9, W10

Apionidae Protapion apricans G4, H17 5,6

Apionidae Protapion assimile G4, W10 5,6,7

Apionidae Protapion fulvipes White Clover G5, H16, H23, 6,7 Seed Weevil H24, H25, W1

Planning Inspectorate Scheme Ref: TR010044 58 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Apionidae Protapion nigritarse E, H9, H17, 6,7,8 H19, H25, W1

Apionidae Protapion trifolii G5, W10 7,8

Coleoptera Apionidae Pseudapion rufirostre G5, G6 5,7 – Beetles Apionidae Taeniapion urticarium G1, Q 5

Attelabidae Attelabus nitens Oak Leaf- G4 7 roller

Byrrhidae – Pill Byrrhus pilula Pill Beetle G1 6 Beetles

Byrrhidae Cytilus sericeus G1 6

Byturidae Byturus ochraceus Q, W1, W9 5,6

Byturidae Byturus tomentosus Raspberry G1, G4, H9, 5,6,7 Beetle W9

Cantharidae – Cantharis cryptica A18, H1, H5, 5,6 Soldier beetles H7, H9, H10, H16, H25, W1

Cantharidae Cantharis decipiens H1, H16, H22 5,6

Cantharidae Cantharis lateralis A, H3, H6, H9, 6 H17, W10

Cantharidae Cantharis livida G4, Q 5

Cantharidae Cantharis nigricans H1, H6, H9, Q 5,6

Cantharidae Cantharis rufa H19 6

Cantharidae Cantharis rustica G4, H19, Q 5,6

Cantharidae Malthinus flaveolus H6, H10 6

Cantharidae Malthinus H10, H16, W9 6,7 seriepunctatus

Cantharidae Malthodes marginatus W1 6

Cantharidae Malthodes minimus H1, H6, H7, 6 H9, H19, H20, H25, W1, W9, W10

Planning Inspectorate Scheme Ref: TR010044 59 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Cantharidae Malthodes pumilus H5, H6, H10, 6 NS H16, H19, H20, H23, H25

Cantharidae Rhagonycha fulva G4, G6, H1, 7 H16, W1

Cantharidae Rhagonycha lignosa W9 6

Cantharidae Rhagonycha limbata H5, H6, H7, 5,6 H9, H22, H24, Q

Coleoptera Cantharidae Rhagonycha lutea H1, H9, H10, 6 NS – Beetles H23, W1

Carabidae – Acupalpus dubius G1, G4 5,6 Ground Beetles

Carabidae Agonum emarginatum G1, G4 5,6

Carabidae Agonum muelleri G4 7

Carabidae Amara aenea G6 5

Carabidae Amara communis G1 6

Carabidae Amara convexior G4 5

Carabidae Amara ovata R 5

Carabidae Amara plebeja H24 6

Carabidae Amara tibialis G4 5

Carabidae Anisodactylus binotatus G1 6

Carabidae Bembidion articulatum A1, Q 5,7

Carabidae Bembidion biguttatum G1 5,6

Carabidae Bembidion clarkii G1 6

Carabidae Bembidion illigeri A1, G4, Q 5,7

Carabidae Bembidion lunulatum A1, G1, Q 5,6,7

Carabidae Bembidion obtusum G1, H25 6

Planning Inspectorate Scheme Ref: TR010044 60 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Carabidae Bembidion properans G1, G4 5,6

Carabidae Bembidion A1, G1 6,7 quadrimaculatum

Carabidae Calathus fuscipes G6 7

Carabidae Calodromius spilotus H6, H17, H19, 6,7 H23, H25, W1

Carabidae Chlaenius vestitus G4 5

Carabidae Clivina fossor G1 6

Coleoptera Carabidae Demetrias atricapillus E, H1, H7, H9, 5,7,8 – Beetles H10, H16, H17, H19, H20, H23, H25, R, W10

Carabidae Demetrias imperialis A18 5 one swept from Phragmites at edge of pool

Carabidae Dromius meridionalis H7, H9, H23, 6,7 W1

Carabidae Harpalus affinis G4 5

Carabidae Harpalus rufipes H9, H22 6

Carabidae Harpalus tardus general site 5 in heap of old grass

Carabidae Leistus spinibarbis W1, W7, W10 5,6

Carabidae Nebria brevicollis G1, G4 5,6

Carabidae Notiophilus biguttatus H25 6

Carabidae Paradromius linearis E, G4, H1, 6,7,8 H7, H9, H10, H16, H17, H20, H22, H24, H25

Carabidae Paranchus albipes G4 7

Planning Inspectorate Scheme Ref: TR010044 61 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Carabidae Philorhizus H3, H7, H9, 6,8 melanocephalus H20, H22, W1, W10

Carabidae Poecilus cupreus G1 6

Carabidae Pterostichus madidus G1, G4 6,7

Carabidae Pterostichus minor A18 5

Carabidae Pterostichus niger G4 7

Carabidae Pterostichus nigrita G1, G4 5

Carabidae Pterostichus strenuus G1 6

Carabidae Pterostichus vernalis G1 6

Carabidae Syntomus foveatus G4 5

Carabidae Trechus obtusus G1 6

Cerambycidae Anaglyptus mysticus H5, H7, H25, 6 – Longhorn W9 Beetles

Coleoptera Cerambycidae Clytus arietis Wasp Beetle H7, H9, H16 6 – Beetles Cerambycidae Glaphyra umbellatarum H7, H22, W1, 6 NS (Na W10 Notable)

Cerambycidae Grammoptera ruficornis A, A18, G1, 5,6 G4, H1, H5, H6, H7, H9, H10, H16, H17, H20, H23, W1, W7, W9, W10

Cerambycidae Phytoecia cylindrica G4, H9, H24, 5,6 NS (Nb Q Notable)

Cerambycidae Pogonocherus hispidus G1 5

Cerambycidae Stenocorus meridianus G4, H9 6,7

Cerambycidae Tetrops praeustus H6, H22 6

Chrysomelidae Altica lythri G4 7 – Leaf Beetles

Planning Inspectorate Scheme Ref: TR010044 62 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Chrysomelidae Altica palustris G1, H24 6,7

Chrysomelidae Aphthona euphorbiae G4, G5, G6, 6,7,8 H1, H6, H7, H16, H17, H19, H23, W1, W9

Chrysomelidae Bruchidius varius H17, H24 7

Chrysomelidae Bruchus loti A1, G4, W1 5,6,7

Chrysomelidae Bruchus rufimanus Bean Beetle E, G4, G6, 5,7,8 H1, H6, H9, H17

Chrysomelidae Bruchus rufipes G4, Q 5

Chrysomelidae Cassida rubiginosa Thistle G4, G5, G6, 5,6,7 Tortoise H20, W10 Beetle

Chrysomelidae Cassida vibex Q, W1, W10 5,6,7

Chrysomelidae Chaetocnema concinna E, H7, H24, 7,8 H25

Chrysomelidae Chaetocnema hortensis G1 6

Chrysomelidae Chaetocnema picipes H1 7

Chrysomelidae Crepidodera aurata H7, Q 5,6,8

Coleoptera Chrysomelidae Crepidodera aurea G4, H19, Q 5,6,7 – Beetles Chrysomelidae Cryptocephalus fulvus G6 7

Chrysomelidae Cryptocephalus labiatus H9 6

Chrysomelidae Cryptocephalus moraei G5 7

Chrysomelidae Cryptocephalus pusillus G4, H1, H9, 7,8 H19, H20, H24

Chrysomelidae Galerucella lineola Brown G4, Q 5,7 Willow Beetle

Chrysomelidae Longitarsus dorsalis G1, G4, G5, Q 5,6

Planning Inspectorate Scheme Ref: TR010044 63 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Chrysomelidae Longitarsus exoletus H9 6

Chrysomelidae Longitarsus flavicornis G4 7

Chrysomelidae Longitarsus luridus G1 6

Chrysomelidae Longitarsus lycopi G1, Q 5,6 NS

Chrysomelidae Longitarsus parvulus A18, H1, H6, 5,6,7, H9, H16, Q, 8 W10

Chrysomelidae Longitarsus pellucidus E, H6 8 swept off field bindweed at edge of arable (E) and on field bindweed at edge of track - arable (H6)

Chrysomelidae Longitarsus rubiginosus E 8

Chrysomelidae Longitarsus succineus G5, G6 7

Chrysomelidae Longitarsus suturellus G1, H17 6

Chrysomelidae Oulema melanopus H6 8

Chrysomelidae Oulema obscura H16 7

Chrysomelidae Oulema rufocyanea R 5

Chrysomelidae Phaedon cochleariae R 5

Coleoptera Chrysomelidae Phaedon tumidulus Celery Leaf general site 5 – Beetles Beetle

Chrysomelidae Phratora vitellinae Brassy general site 5 in profusion Willow on willows Beetle next to River Ouse

Chrysomelidae Phyllotreta nigripes E, H9, H17, 5,7,8 H24, H25

Chrysomelidae Phyllotreta ochripes H10 5,6

Planning Inspectorate Scheme Ref: TR010044 64 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Chrysomelidae Phyllotreta undulata H9, H24, Q 5,7,8

Chrysomelidae Phyllotreta vittula H24, W1 7

Chrysomelidae Psylliodes affinis W9 8

Chrysomelidae Psylliodes E, G4, G5, 6,7,8 chrysocephala G6, H1, H7, H9, H10, H16, H17, H19, H20, H22, H23, H24, H25, W1, W9, W10

Chrysomelidae Psylliodes laticollis R 5

Chrysomelidae Psylliodes luteola H19 7 NS – one beaten off oak

Chrysomelidae Psylliodes napi Q 5

Chrysomelidae Pyrrhalta viburni Guelder- G5 5,7 includes rose Leaf larvae Beetle

Chrysomelidae Sphaeroderma rubidum H16 7

Chrysomelidae Sphaeroderma G4, G6, H9, 7,8 testaceum H16

Ciidae Cis pygmaeus H19 6

Ciidae Ennearthron cornutum H10 6

Ciidae Orthocis alni W1 6

Cleridae Opilo mollis H16, H22, 6 NS – off elm H25

Cleridae Tillus elongatus H6, H9, H25 6 NS

Coccinellidae – Adalia decempunctata 10-spot H7, H10, H17, 6,7,8 Ladybirds Ladybird H19, H25, W10

Coleoptera Coccinellidae Anisosticta Water G4 7 – Beetles novemdecimpunctata Ladybird

Planning Inspectorate Scheme Ref: TR010044 65 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Coccinellidae Coccinella 7-spot E, G4, G5, 5,6,7, septempunctata Ladybird G6, H1, H3, 8 H9, H16, H17, H19, H20, H22, H23, H24, H25, Q, W1, W9, W10

Coccinellidae Exochomus Pine H1 6 quadripustulatus Ladybird

Coccinellidae Halyzia sedecimguttata Orange A18 5 Ladybird

Coccinellidae Harmonia axyridis Harlequin A, G4, G5, 5,6,7, Ladybird G6, H1, H3, 8 H6, H9, H16, H17, H19, H23, H25, Q, W1

Coccinellidae Propylea 14-spot G4, G5, H5, 5,6,7 quattuordecimpunctata Ladybird H9, H17, H20, H24

Coccinellidae Psyllobora 22-spot G6 7 vigintiduopunctata Ladybird

Coccinellidae Rhyzobius litura G6, H17, R 5,6,7

Coccinellidae Rhyzobius lophanthae W10 8

Coccinellidae Scymnus auritus H19 7

Coccinellidae Scymnus frontalis G5, H25 7

Coccinellidae Scymnus G1 6 haemorrhoidalis

Coccinellidae Stethorus punctillum H1, H16, H19, 6,7 H23, W1

Coccinellidae Subcoccinella 24-spot E, G4, G6, 5,7,8 vigintiquattuorpunctata Ladybird H7, H17

Coccinellidae Tytthaspis 16-spot G6, H17, H22 7 sedecimpunctata Ladybird

Cryptophagidae Atomaria linearis Pygmy W1 6 Beetle

Planning Inspectorate Scheme Ref: TR010044 66 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Cryptophagidae Atomaria testacea G4, P 5

Cryptophagidae Telmatophilus typhae A18 5

Coleoptera Curculionidae – Acalles misellus H6 ,H9, W1 6 – Beetles Weevils (part)

Curculionidae Anthonomus rubi Strawberry A, G5, G6, 5,6,7, Blossom H23, H25, W9 8 Weevil

Curculionidae Anthonomus ulmi H9, W9 6 NS (Nb Notable)

Curculionidae Archarius pyrrhoceras W10 6

Curculionidae Archarius salicivorus Willow Gall G4 5,7 Weevil

Curculionidae Barypeithes pellucidus G4, G5, H6, 5,6 H9, H10, H16, W1, W7, W9, W10

Curculionidae Calosirus terminatus G6 7 NS (Nb Notable)

Curculionidae Ceutorhynchus H1, H16, H19, 7 obstrictus W1

Curculionidae Ceutorhynchus Cabbage G1, G6, H24, 5,6,7 pallidactylus Stem Weevil H25, W1

Curculionidae Ceutorhynchus typhae H9, R 5,6

Curculionidae Cionus scrophulariae Figwort Q 5 Weevil

Curculionidae Curculio glandium Acorn H19 6 Weevil

Curculionidae Curculio venosus H7 6

Curculionidae Dorytomus dejeani H19 6 off aspen

Curculionidae Dorytomus filirostris W9 6 NS (Nb Notable) – from stream edge near wood

Planning Inspectorate Scheme Ref: TR010044 67 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Curculionidae Dorytomus longimanus W9 6 from stream edge near wood

Curculionidae Dorytomus taeniatus H7, H25 6

Curculionidae Drupenatus nasturtii R 5 NS (Nb Notable) - on watercress in stream bed

Curculionidae Glocianus distinctus G4, G6 5,7

Coleoptera Curculionidae Hylesinus toranio A, H17, H23, 6,7 – Beetles H24

Curculionidae Hypera meles G4, G6 5,7 NS (Na Notable)

Curculionidae Hypera nigrirostris G4 5

Curculionidae Hypera plantaginis G6 7

Curculionidae Hypera pollux R 5

Curculionidae Hypera postica Clover Leaf G4 5 Weevil

Curculionidae Hypera rumicis Q 5

Curculionidae Isochnus sequensi G4 7 [Red Data Book, RDBK]

Curculionidae Larinus planus G5 5 NS (Nb Notable) – one swept off spear thistle on short turf rabbit grazed post- industrial ground near roundabout

Curculionidae Liophloeus tessulatus H3, H9 6

Planning Inspectorate Scheme Ref: TR010044 68 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Curculionidae Magdalis armigera A, H1, H3, H6, 6,7 H7, H9, H16, H17, H22, H25, W1, W10

Curculionidae Magdalis ruficornis H6 6

Curculionidae Mecinus pascuorum G4, G6, H7, Q 5,6,7

Curculionidae Mecinus pyraster G6 5

Curculionidae Microplontus rugulosus H25 7

Curculionidae Mogulones G4 5 asperifoliarum

Curculionidae Nedyus Small Nettle G5, H3, H6, 5,6 quadrimaculatus Weevil H23, R, Q, W9

Curculionidae Orchestes alni E, H6, H9, 5,6,7, H10, H20, 8 H22, H25, W1, W10

Coleoptera Curculionidae Pachyrhinus lethierryi A 6 – Beetles Curculionidae Parethelcus pollinarius H1, H3, W7 6,7

Curculionidae Phyllobius maculicornis Green Leaf Q 5 Weevil

Curculionidae Phyllobius oblongus Brown Leaf H1, H9, H20 6 Weevil

Curculionidae Phyllobius pomaceus H1, H3, H5, 5,6 H6, Q, W10

Curculionidae Phyllobius pyri Common G4, G6, H1, 5,6 Leaf Weevil H9, H16, H23, W1

Curculionidae Phyllobius roboretanus Small Green A, H1, H3, H5, 5,6 Nettle H6, H7, H9, Weevil H10, H16, H17, H20, H22, H23, Q

Curculionidae Polydrusus cervinus H10, W10 6

Planning Inspectorate Scheme Ref: TR010044 69 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Curculionidae Polydrusus formosus H17 6 [NS (Na Notable)]

Curculionidae Polydrusus H7, H9, H10, 6 pterygomalis W9

Curculionidae Rhamphus H6, H16 6,8 oxyacanthae

Curculionidae Rhinocyllus conicus Q 5 [NS (Na Notable)]

Curculionidae Rhinoncus pericarpius G1, G5, G6, 5.6.7 H24, Q, W10

Curculionidae Scolytus multistriatus Small E Bark H6, W1 6,7 Beetle

Curculionidae Scolytus pygmaeus H9, W1 6,7

Curculionidae Scolytus rugulosus Fruit Bark H24 7 Beetle

Curculionidae Scolytus scolytus Large E H6, H22 6 Bark Beetle

Curculionidae Sitona humeralis G4 5

Curculionidae Sitona lineatus E, G1, G4, 5,6,7, G6, H1, H6, 8 H7, H9, H10, H17, H19, H20, H22, H23, H24, H25, Q, W1, W9, W10

Curculionidae Trichosirocalus G4, G5, G6, Q 5 troglodytes

Coleoptera Curculionidae Tychius picirostris G4 5 – Beetles Curculionidae Zacladus exiguus a cranesbill G4, H19, R 5 NS (Nb weevil Notable)

Dasytidae Dasytes aeratus G1, G4, G5, 5,6 H6, H17

Dermestidae Anthrenus fuscus G4 7

Planning Inspectorate Scheme Ref: TR010044 70 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Dermestidae Anthrenus verbasci G1 5

Dermestidae Ctesias serra Cobweb E, H23 6,8 Beetle

Elateridae – Agriotes acuminatus G1, H1, H23, 5,6 Click Beetles W1, W9, W10

Elateridae Agriotes lineatus G1 5,6

Elateridae Agriotes obscurus G1, Q 5,6

Elateridae Agriotes sputator G1, G4, H5, 5,6 H20, H22, Q, W10

Elateridae Athous haemorrhoidalis A18, G4, H1, 5,6 H3, H6, H9, H10, H17, H22, H23, H25, Q, W1, W7, W9, W10

Elateridae Denticollis linearis W9 6

Elateridae Elater ferrugineus H20 7 Red Data Book (RDB1 – Endangered ) – 2 females attracted to pheromone lure on main farm track nr buildings at c. 9.30am!

Elateridae Hemicrepidius hirtus H1, H3 6,7

Elateridae Kibunea minuta G1. G4, G5, 5,6 H9, H17, H24

Erotylidae Dacne rufifrons W1 6

Erotylidae Triplax aenea W1 6

Coleoptera Histeridae – Atholus P 5 – Beetles Clown Beetles duodecimstriatus

Planning Inspectorate Scheme Ref: TR010044 71 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Histeridae Hister unicolor G1, G4 5

Histeridae Margarinotus ventralis P 5

Histeridae Onthophilus striatus P 5

Hydrophilidae Cercyon lateralis G1 5

Hydrophilidae Cercyon unipunctatus H22 6

Hydrophilidae Cryptopleurum minutum P 5

Hydrophilidae Sphaeridium P 5 bipustulatum

Kateretidae Brachypterus glaber G1, G5, H3, 5,6,7 W1

Kateretidae Brachypterus urticae Nettle Pollen G1, G5, H25, 5,6 Beetle Q

Latridiidae Cartodere bifasciata H16, W9 6

Latridiidae Cortinicara gibbosa H7, H9, H17, 6,7 H20, H23, H25, W1

Latridiidae Enicmus transversus H22, W9 6

Malachiidae Cordylepherus viridis G4, G6, H5, 5,6,7 H9, H16, H17, H24, W9

Malachiidae Malachius bipustulatus Malachite A, A18, G1, 5,6 Beetle H1, H5, H24, W9, W10

Melandryidae Anisoxya fuscula H1, H16, H17, 7 NS H25

Melandryidae Osphya bipunctata H5 6 NS

Monotomidae Monotoma brevicollis P 5

Mordellidae Mordellistena H23, W1 7 [NS] neuwaldeggiana

Mordellidae Mordellistena pumila G4 5

Mordellidae Mordellistena variegata G4, H1 6,7 NS

Planning Inspectorate Scheme Ref: TR010044 72 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Coleoptera Mordellidae Mordellochroa A18, H10 5,6 – Beetles abdominalis

Mycetophagida Mycetophagus W1 6 e quadripustulatus

Nitidulidae – Meligethes aeneus Common A, G1, G5, 5,6,7 Pollen Beetles Pollen G6, H1, H3, Beetle H5, H7, H9, H10, H16, H17, H20, H22, H24, H25, R, W1, W7, W10

Nitidulidae Meligethes flavimanus G5 5

Oedemeridae Ischnomera cyanea A, H7 6

Oedemeridae Oedemera lurida A18, G4, G5, 5,6,7 G6, H7, H16, H17, H20, Q, W1

Oedemeridae Oedemera nobilis Swollen- A, G4, G5, 5,6,7 thighed G6, H1, H3, Beetle H5, H7, H9, H16, H17, H20, H23, Q, W1, W10

Phalacridae Olibrus aeneus E 8

Phalacridae Olibrus affinis H17 6,7

Phalacridae Olibrus corticalis H24, H25 7

Phalacridae Olibrus liquidus G4, G5, G6 5,7

Phalacridae Phalacrus fimetarius H6, H19, H19, 6,7 H22, H24, H25

Ptinidae Anobium fulvicorne H6, H9, H10, 6 H22, H23, H25, W1, W9, W10

Ptinidae Anobium punctatum Woodworm G4, H16, W1 7

Planning Inspectorate Scheme Ref: TR010044 73 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Ptinidae Grynobius planus H9 6

Ptinidae Hedobia imperialis H10 6

Ptinidae Ochina ptinoides Ivy Boring G4, H7, H9, 5,6 Beetle W1

Ptinidae Ptilinus pectinicornis Fan-bearing H1, H9, H17, 6 Wood-borer W1

Pyrochroidae – Pyrochroa serraticornis Common H23 6 Cardinal Cardinal Beetles Beetle

Coleoptera Rhynchitidae Neocoenorrhinus Strawberry H7, W9 6 – Beetles germanicus Rhynchites

Rhynchitidae Tatianaerhynchites Apple Fruit G1, G4, G5, 5,6 aequatus Rhynchites H6, H16

Salpingidae Lissodema denticolle H1, H3, H6, 6,7,8 NS H9, H16, H17, H22, H25, W1

Salpingidae Salpingus planirostris H1, H6, H17, 6,7 H22, H23, H25, W1

Salpingidae Vincenzellus ruficollis H22 6

Scarabaeidae – Aphodius fimetarius G1 5 Dung Beetles and Chafers

Scarabaeidae Aphodius granarius H22 6

Scarabaeidae Aphodius P 5 horse dung haemorrhoidalis in paddock

Scarabaeidae Aphodius prodromus P 5

Scarabaeidae Aphodius sphacelatus P 5

Scarabaeidae Oxyomus sylvestris P 5 horse dung in paddock

Scirtidae Cyphon coarctatus H1, H9, H16, 6,7 male W1

Scirtidae Cyphon ochraceus H7, H10, W1 6

Planning Inspectorate Scheme Ref: TR010044 74 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Scirtidae Microcara testacea H1, H5, H7, 6 W1, W7

Scraptiidae Anaspis fasciata H3, H7 5,6

Scraptiidae Anaspis garneysi A18, H6, H7, 5,6 H19, H22, H23, W9

Scraptiidae Anaspis maculata A, A18, G1, 5,6 G4, H1, H3, H5, H6, H7, H9, H10, H16, H17, H22, H23, H25, Q, W1, W7, W9, W10

Coleoptera Scraptiidae Anaspis pulicaria H1, H3, H5, 6 – Beetles H6, H7, H9, H16, H20, H22, H23, H25, W1, W9

Scraptiidae Anaspis regimbarti A, A18, H1, 5,6 H19, H22, H23, Q, W1

Scydmaenidae Scydmaenus tarsatus G1 5

Silvanidae Psammoecus A18 5 bipunctatus

Staphylinidae – Alaobia pallidicornis H1 6 Rove Beetles

Staphylinidae Aleochara brevipennis G1 5 [NS (Notable)]

Staphylinidae Aleochara intricata P 5

Staphylinidae Aleochara tristis P 5

Staphylinidae Aloconota gregaria A1 7

Staphylinidae Anotylus rugosus G1 6

Staphylinidae Anotylus tetracarinatus P 5

Staphylinidae Autalia rivularis P 5

Planning Inspectorate Scheme Ref: TR010044 75 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Staphylinidae Bisnius fimetarius G1 5

Staphylinidae Cypha longicornis H5, H9, H16, 6,7 H19, H25

Staphylinidae Cypha seminulum H6 6 male

Staphylinidae Datomicra sordidula P 5

Staphylinidae Dimetrota atramentaria P 5

Staphylinidae Dimetrota setigera P 5

Staphylinidae Dropephylla vilis G1 5 male

Staphylinidae Gabrius breviventer A1, Q 5,7

Staphylinidae Gabrius piliger G1 5

Staphylinidae Gyrohypnus fracticornis P 5

Coleoptera Staphylinidae Habrocerus G1 5 – Beetles capillaricornis

Staphylinidae Lathrobium brunnipes G1 6

Staphylinidae Lathrobium fulvipenne G1 6

Staphylinidae Lathrobium geminum G1 6

Staphylinidae Ocypus brunnipes W9 6

Staphylinidae Ocypus olens Devil's G4 7 Coach-horse

Staphylinidae Oxytelus laqueatus P, W1 5,6

Staphylinidae Philonthus albipes P 5

Staphylinidae Philonthus cognatus R 5

Staphylinidae Philonthus fumarius G1 5 [NS (Nb Notable)]

Staphylinidae Philonthus P 5 sanguinolentus

Staphylinidae Philonthus splendens P 5

Staphylinidae Philonthus tenuicornis G1, P 5

Planning Inspectorate Scheme Ref: TR010044 76 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Staphylinidae Philonthus varians P 5

Staphylinidae Platystethus arenarius H22, H23 6,7

Staphylinidae Platystethus nitens G1 6

Staphylinidae Platystethus nodifrons G1 6 NS (Notable)

Staphylinidae Quedius mesomelinus H22 6

Staphylinidae Quedius semiobscurus G4 5

Staphylinidae Rugilus orbiculatus P 5

Staphylinidae Stenus canaliculatus A1 7

Staphylinidae Stenus clavicornis P 5

Staphylinidae Stenus flavipes W9 6

Staphylinidae Stenus impressus W10 6

Coleoptera Staphylinidae Stenus picipes W9 6 – Beetles Staphylinidae Tachinus flavolimbatus G4 5 [NR (RDBK)]

Staphylinidae Tachyporus atriceps W10 6

Staphylinidae Tachyporus hypnorum G4, G5, G6 5,7

Staphylinidae Tachyporus nitidulus G1, H24, W10 6

Staphylinidae Tachyporus solutus H10, Q, W1, 5,6 W10

Staphylinidae Tinotus morion P 5

Staphylinidae Xantholinus linearis G4 5

Staphylinidae Xantholinus longiventris G1 6

Tenebrionidae Lagria hirta H1, H16, W1 7

Tenebrionidae Pseudocistela H23, W1 6 NS ceramboides

Dermaptera Forficulidae Forficula auricularia Common A, A18, G1, 5,6,7, – Earwigs Earwig G4, H1, H3, 8

Planning Inspectorate Scheme Ref: TR010044 77 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1 H5, H6, H7, H9, H10, H16, H17, H19, H20, H22, H23, H24, H25, W1, W7, W9, W10

Diptera – Asilidae – Dioctria atricapilla G4 5 Flies Robberflies

Asilidae Dioctria linearis H9, H10, W9 6

Asilidae Dioctria rufipes Q 5

Asilidae Leptogaster cylindrica G6, H9, H16, 6,7,8 H17, W9

Bibionidae Bibio marci St Mark’s Fly general site 5

Bombyliidae – Bombylius major Bee-fly general site 5 Beeflies

Dolichopodidae Dolichopus festivus H17 6

Empididae Empis livida H17 6

Diptera – Limoniidae Euphylidorea dispar H5 6 Flies Platystomatidae Platystoma A, H3, H5, Q 5,6 seminationis

Rhagionidae – Chrysopilus asiliformis G4 7 Snipe Flies

Rhagionidae Chrysopilus cristatus G4 7

Stratiomyidae – Beris chalybata A 6 Soldierflies

Stratiomyidae Chloromyia formosa H7, H16 6,7

Stratiomyidae Chorisops tibialis G4, G5, H1, 7,8 H25

Stratiomyidae Microchrysa flavicornis A 6

Stratiomyidae Oxycera trilineata G5 7

Planning Inspectorate Scheme Ref: TR010044 78 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Stratiomyidae Pachygaster atra G4, G5, H1, 6,7 H9, H22

Stratiomyidae Pachygaster leachii G5, H7 7,8

Syrphidae – Epistrophe eligans general site 5 Hoverflies

Syrphidae Episyrphus balteatus G4, H1, H16, 6,7,8 W9, W10

Syrphidae Myathropa florea Q 5

Syrphidae Rhingia campestris general site 5

Syrphidae Sphaerophoria scripta G5, G6, H16 7

Syrphidae Syritta pipiens H16 7

Tabanidae – Chrysops relictus Q 5 Horseflies and Clegs

Tabanidae Haematopota pluvialis H23 6

Tachinidae Cistogaster globosa E 8 [Red Data Book (pRDB2 – Vulnerable)]

Tachinidae Gymnochaeta viridis general site 5

Diptera – Tachinidae Phasia hemiptera G4 7 Flies Tachinidae Tachina fera H6 6

Tephritidae – Chaetorellia jaceae G4, G6 5,7 Picture-winged Flies

Tephritidae Tephritis ruralis G4 5

Tephritidae Terellia ruficauda H16 6

Tephritidae Urophora stylata H16, W1 6

Tipulidae Tipula maxima G4 5

Planning Inspectorate Scheme Ref: TR010044 79 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Ephemeropt Ephemeridae Ephemera vulgata Q 5 era – Mayflies

Hemiptera – Anthocoridae Anthocoris amplicollis H17, H23, 7 Red Data True Bugs H24 Book (RDB3)

Anthocoridae Anthocoris confusus G4, H7, H10, 5,7,8 H17, H19, H25, Q

Anthocoridae Anthocoris gallarum- A, H22, H23, 6,7 ulmi H25, W1

Anthocoridae Anthocoris nemoralis H1 6

Anthocoridae Anthocoris nemorum G1, H1, H3, 5,6,7, H6, H7, H25, 8 W9

Anthocoridae Anthocoris simulans H6, H9, H17, 7,8 H23, H24

Anthocoridae Brachysteles A 6 parvicornis

Anthocoridae Cardiastethus H7, H19, H24 7,8 fasciiventris

Anthocoridae Temnostethus gracilis H16, H17, 7 H24, W1

Anthocoridae Temnostethus pusillus H1 7

Aphrophoridae Aphrophora alni E, G4, G5, 6,7,8 – Froghoppers G6, H1, H6, H7, H9, H16, H17, H19, H20, H23, H24, H25, W1, W9, W10

Aphrophoridae Aphrophora salicina G4, G6 7

Hemiptera – Aphrophoridae Neophilaenus G6 7 True Bugs campestris

Aphrophoridae Neophilaenus lineatus H1, H9, H17, 7,8 H22

Planning Inspectorate Scheme Ref: TR010044 80 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Aphrophoridae Philaenus spumarius E, G4, G5, 7,8 G6, H1, H6, H7, H9, H10, H16, H17, H19, H20, H22, H23, H24, H25, W1, W9, W10

Aradidae – Aneurus avenius H6, H9, H25, 6,7 Flatbugs W1

Cercopidae Cercopis vulnerata Q 5

Cicadellidae – Agallia consobrina H17, H20, 7 Leafhoppers H24, H25

Cicadellidae Allygus mixtus H7, H10 8

Cicadellidae Aphrodes makarovi H22, H24 7 male

Cicadellidae Aphrodes sp (makarovi G4, G5, G6 7 complex females)

Cicadellidae Athysanus argentarius H17 7

Cicadellidae Doratura stylata H17 7

Cicadellidae Edwardsiana lethierryi H1, H17 6 male

Cicadellidae Eupteryx aurata Q 5

Cicadellidae Eupteryx vittata W9 6

Cicadellidae Iassus lanio H19 7

Cicadellidae Iassus scutellaris E, G5, H1, 7,8 [NS (Na H3, H6, H7, Notable)] H9, H10, H16, H20, H25, W1, W10

Cicadellidae Idiocerus herrichi G4 7

Cicadellidae Idiocerus lituratus G5 7

Cicadellidae Ledra aurita H23, H25 7 nymph

Cicadellidae Liguropia juniperi E 8

Planning Inspectorate Scheme Ref: TR010044 81 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Cicadellidae Macropsis prasina G4 7

Hemiptera – Cicadellidae Mocydia crocea H25, W10 6,7 True Bugs Cicadellidae Populicerus confusus H17 7

Cicadellidae Populicerus laminatus G4 7

Cicadellidae Populicerus populi H19 7

Cicadellidae Viridicerus ustulatus G4 5

Cixiidae Cixius nervosus H1, H9 6,7

Cixiidae Reptalus H25 7 NS (Nb quinquecostatus Notable)

Cixiidae Tachycixius pilosus H9, H17, W9, 6 W10

Coreidae – Coreus marginatus Dock Bug E, G4, G5, 5,7,8 Squashbugs H17

Coreidae Coriomeris denticulatus Denticulate G4, G6, H17 5,7 Leatherbug

Coreidae Gonocerus Box Bug G5, H1, H7, 7,8 Nymphs acuteangulatus H24 and adults (8)

Coreidae Syromastus rhombeus Rhombic G4 5,7 Leatherbug

Cydnidae – Sehirus luctuosus Forget-me- G1, H25 6,7 Shieldbugs not (part) Shieldbug

Cydnidae Tritomegas bicolor Pied G4 5 Shieldbug

Delphacidae Asiraca clavicornis E, H9 8 NS (Nb Notable)

Delphacidae Criomorphus H19 6 male albomarginatus

Delphacidae Hyledelphax H25 7 elegantulus

Delphacidae Javesella pellucida Q 5

Planning Inspectorate Scheme Ref: TR010044 82 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Delphacidae Kosswigianella exigua G6, H17 5,7

Delphacidae Ribautodelphax imitans H17 7 [Red Data Book (RDBK)]

Hemiptera – Delphacidae Stenocranus minutus H17, Q 5,7 True Bugs Delphacidae Stiroma affinis W9 6

Lygaeidae – Chilacis typhae A1, G4 7 Groundbugs

Lygaeidae Cymus glandicolor A1 7

Lygaeidae Cymus melanocephalus G5, Q 5,7

Lygaeidae Drymus sylvaticus H10, H17 6

Lygaeidae Graptopeltus lynceus G4, Q 5 NS (Nb Notable)

Lygaeidae Heterogaster urticae G1, H3, H7, 5,6,7, H23, H24, 8 H25

Lygaeidae Kleidocerys resedae W10 8

Lygaeidae Megalonotus G4 5 NS (Nb antennatus Notable)

Lygaeidae Nysius huttoni G4 5

Lygaeidae Scolopostethus affinis H25 6

Lygaeidae Scolopostethus W1, W9, W10 6 thomsoni

Microphysidae Loricula elegantula H1, W1 7

Microphysidae Loricula pselaphiformis G1, H6 5,6

Miridae – Adelphocoris lineolatus G4, G5 7 Capsid Bugs

Miridae Agnocoris reclairei G4 7 NS (Nb Notable)

Miridae Amblytylus nasutus G5, G6, H1, 6,7 H19

Planning Inspectorate Scheme Ref: TR010044 83 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Miridae Apolygus spinolae H1, H3, H24, 7 H25

Miridae Atractotomus mali G5, H1, H16, 7 W1

Miridae Blepharidopterus E, H6, H7, H9, 7,8 angulatus H10, H23, H25

Miridae Campyloneura virgula G4, H1, H20, 7 W1

Hemiptera – Miridae Capsus ater H5, H9, H19, 6 True Bugs W9, W10

Miridae Closterotomus G4, G6, H1, 7 norwegicus H3, H16, H22, W1

Miridae Cyllecoris histrionius G4 5

Miridae Deraeocoris flavilinea G4, G5, H1, 6,7 H3, H16, H24, H25, W1

Miridae Deraeocoris lutescens G4, H7, H10, 7,8 H17, H19, H20, H23, H24

Miridae Deraeocoris ruber G5 7

Miridae Dicyphus epilobii G4 7

Miridae Dicyphus globulifer G4, G5 7

Miridae Dicyphus stachydis W10 6

Miridae Dryophilocoris H7, H19, W9 6 flavoquadrimaculatus

Miridae Grypocoris stysi H25, W10 6

Miridae Heterotoma planicornis E, G4, G5, 7,8 H1, H6, H9, H16, H23, H25, W1

Planning Inspectorate Scheme Ref: TR010044 84 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Miridae Leptopterna dolabrata H1, H5, H6, 6,7 H7, H16, H17, H24

Miridae Leptopterna ferrugata G6 7

Miridae Liocoris tripustulatus G4, G5, H1, 5,6,7, H3, H6, H16, 8 H23, H25, Q, W9, W10

Miridae Lygocoris pabulinus H3, W1, W9, 6 W10

Miridae Lygus sp (pratensis G5, G6, H1, 7 complex) H16

Miridae Macrotylus horvathi G6 7

Miridae Megacoelum infusum H19 7

Miridae Megaloceroea G4 7 recticornis

Hemiptera – Miridae Miris striatus G1, G4, H1, 5,6 includes True Bugs H6, H20 nymphs

Miridae Neolygus contaminatus H17 6

Miridae Neolygus populi G4 7

Miridae Neolygus viridis H24, H25 7

Miridae Oncotylus viridiflavus G6, H3, H7, 7,8 H16, H17, H22, W10

Miridae Orthocephalus saltator G4, G5, G6 7

Miridae Orthonotus rufifrons G4 7

Miridae Orthops basalis E, G4 7,8

Miridae Orthops campestris G4 7

Miridae Orthops kalmii G6 7

Miridae Orthotylus marginalis G4 7

Miridae Orthotylus ochrotrichus G4 7

Planning Inspectorate Scheme Ref: TR010044 85 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Miridae Orthotylus tenellus H19 6

Miridae Orthotylus viridinervis H16, W1 7

Miridae Phylus melanocephalus W7 6

Miridae Phytocoris tiliae H25 7

Miridae Phytocoris ulmi H24 7

Miridae Phytocoris varipes E, G5, G6, 7,8 H10, H17, H22

Miridae Pilophorus clavatus H9 8

Miridae Pilophorus perplexus H25 7

Miridae Pinalitus cervinus H19, H22 6

Miridae Plagiognathus G4, G5, H1, 7,8 arbustorum H3, H9, H16, H17, H22, H23, H24, H25, W1

Miridae Plagiognathus G4, G5, G6, 7 chrysanthemi H1, H16, W1

Hemiptera – Miridae Polymerus unifasciatus G6 7 True Bugs Miridae Psallus lepidus H1, H23, W1 6,7

Miridae Psallus perrisi H19, Q, W10 5,6

Miridae Psallus variabilis H19, W1 6

Miridae Psallus varians H19 6

Miridae Pseudoloxops G4 7 coccineus

Miridae Rhabdomiris striatellus H10, Q, W9 5,6

Miridae Stenodema calcarata H25 7

Miridae Stenodema laevigata H17, H24, W9 6,7,8

Miridae Stenotus binotatus G4, H16 7

Planning Inspectorate Scheme Ref: TR010044 86 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Miridae Sthenarus rotermundi G4 7

Nabidae – Himacerus apterus H3, H6, H23, 7,8 Damselbugs H24, H25, W9, W10

Nabidae Himacerus major G6 7

Nabidae Himacerus mirmicoides G6, H6 7,8

Nabidae Nabis ferus E, H7, H17, 7,8 H19, H22, H24, H25

Nabidae Nabis flavomarginatus H17 7

Nabidae Nabis limbatus H17, H22, 7 H23, H24, H25

Nabidae Nabis rugosus G4 5

Pentatomidae – Acanthosoma Hawthorn A, H20, H23, 6,7 Shieldbugs haemorrhoidale Shieldbug H24 (part)

Pentatomidae Aelia acuminata Bishop's G4, H10, H17, 5,7,8 Mitre H22, W9, Shieldbug W10

Pentatomidae Dolycoris baccarum Hairy G4, G5, G6, 5,7,8 Shieldbug H3, H7, H17, H24, H25, Q

Hemiptera – Pentatomidae Eysarcoris Woundwort W10 6 True Bugs venustissimus Shieldbug

Pentatomidae Palomena prasina Common E, G4, H6, 6,7,8 Green H9, H10, H17, Shieldbug H19, H20, H22, H24, W10

Pentatomidae Pentatoma rufipes Red-legged E, G4, H1, 7,8 Shieldbug H9, H16, H23, H24, , W1, W10

Pentatomidae Podops inuncta Turtle G1 6 Shieldbug

Planning Inspectorate Scheme Ref: TR010044 87 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Pentatomidae Zicrona caerulea Blue G1 6 Shieldbug

Psyllidae Cacopsylla ulmi H6 8

Reduviidae – Empicoris culiciformis H24 7 Assassin Bugs

Rhopalidae Myrmus miriformis G5, G6 7

Rhopalidae Rhopalus G5 7 parumpunctatus

Rhopalidae Rhopalus subrufus H6, H17, W9 7,8

Rhopalidae Stictopleurus abutilon G4 7

Rhopalidae Stictopleurus G4 5 punctatonervosus

Saldidae – Saldula saltatoria G1 6 Shore Bugs

Scutelleridae – Eurygaster testudinaria Tortoise G4 7 Shieldbugs Shieldbug (part)

Tingidae – Physatocheila E, H6, H7, H9, 6,7,8 Lacebugs dumetorum H17, H20, H22, H23, H24, H25

Tingidae Tingis ampliata H17, Q 5,7

Tingidae Tingis cardui H7, H16, H25 6,7

Hymenopter Apidae – Bees Apis mellifera Honey Bee H1, H16, H20 6,7 a – Bees, Ants, Wasps, Sawflies

Hymenopter Apidae Bombus hypnorum Tree H16 7 a – Bees, Bumblebee Ants, Wasps, Apidae Bombus lapidarius Red-tailed G6, H16 5,7 Sawflies Bumblebee

Apidae Bombus lucorum agg. White-tailed H16 5,7 Bumblebee

Planning Inspectorate Scheme Ref: TR010044 88 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Apidae Bombus pascuorum Common general site 5 Carder Bee

Apidae Bombus terrestris Buff-tailed A18 5 Bumblebee

Apidae Nomada flava Flavous A18 5 Nomad

Argidae – Aproceros leucopoda Zigzag E, G5, H1, 6,7,8 very recent Sawflies (part) Sawfly H3, H5, H6, arrival in H7, H9, H10, Britain H16, H20, H25, W1

Formicidae – Formica fusca Negro Ant G4, G5, H16 7 Ants

Formicidae Lasius brunneus Brown Ant H5, H6, H9, 5,6,8 NS (Na H23, H25 Notable)

Formicidae Lasius fuliginosus Jet Ant H24, H25 6,8

Formicidae Lasius niger agg. Black A18, G4, G5, 5,6,7 Garden Ant G6, H1, H16, W1

Formicidae Myrmica rubra Red Ant P 5

Tenthredinidae Athalia rosae Turnip H17 6 – Sawflies Sawfly (part)

Tenthredinidae Cladius rufipes E, H1 7,8

Vespidae – Vespa crabro Hornet H9, H22 6 Wasps

Isopoda – Armadillidiidae Armadillidium vulgare Common Pill G1, G4, W10 5,6 Woodlice Woodlouse

Philosciidae Philoscia muscorum Common A18, G1 5,6 Striped Woodlouse

Julida – Julidae Cylindroiulus punctatus Blunt-tailed G1 6 Millipedes Snake Millipede

Planning Inspectorate Scheme Ref: TR010044 89 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Lepidoptera Adelidae – Adela croesella Small Barred H16 6 – Moths and Longhorn Longhorn Butterflies Moths

Adelidae Nemophora degeerella Yellow- W7 6 barred Longhorn

Arctiidae Tyria jacobaeae Cinnabar G4, G5, G6, 5,6,7 includes H9, H16, H17, larvae – UK W1 BAP Priority Species (Research Only) S. 41

Argyresthiidae Argyresthia H6, H7 8 bonnatella/spinosella

Bucculatricidae Bucculatrix albedinella Elm Bent- H6 8 wing

Geometridae Opisthograptis luteolata Brimstone E 8 Moth

Gracillariidae Phyllonorycter Small Elm H16 7 schreberella Midget

Hesperiidae Thymelicus lineola Essex G4, G6 7 Skipper

Hesperiidae Thymelicus sylvestris Small H3, H16 7 Skipper

Lycaenidae Aricia agestis Brown Argus G5, G6, H7, 5,7,8 H16, H17, W9

Lycaenidae Celastrina argiolus Holly Blue W1 7

Lycaenidae Polyommatus icarus Common E, G1, G4, 5,7,8 Blue G5, H16, H17

Momphidae Mompha raschkiella Little G4 7 Mompha

Noctuidae Autographa gamma Silver Y H25 6

Noctuidae Cosmia diffinis White- H20, W10 6 Larva - UK spotted BAP Priority Pinion Species, NERC S.41,

Planning Inspectorate Scheme Ref: TR010044 90 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1 provisionally Red Data Book

Noctuidae Orthosia cerasi Common A, H1, H9, 6 larva Quaker H25

Noctuidae Orthosia gothica Hebrew H5 6 larva Character

Lepidoptera Notodontidae Clostera curtula Chocolate- G4 7 – Moths and tip Butterflies Nymphalidae Aglais io Peacock H16, H25 5,7

Nymphalidae Aglais urticae Small H1, H3 5,7 Tortoiseshell

Nymphalidae Aphantopus hyperantus Ringlet G4, H1, H16, 7,8 H19, H24

Nymphalidae Coenonympha Small Heath H17 7 UK BAP pamphilus Priority Species (Research Only), NERC S.41, IUCN Near Threatened

Nymphalidae Maniola jurtina Meadow E, G4, G5, 6,7,8 Brown G6, H5, H9, H10, H16, H19, W1, W9

Nymphalidae Melanargia galathea Marbled H16 7 White

Nymphalidae Pararge aegeria Speckled H23, W9, 6,8 Wood W10

Nymphalidae Polygonia c-album Comma H1, H9 5,7,8 includes larva

Nymphalidae Pyronia tithonus Gatekeeper G4, G6, H16, 7,8 H17, H24, H25

Nymphalidae Vanessa atalanta Red Admiral H1 7

Planning Inspectorate Scheme Ref: TR010044 91 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Oecophoridae Crassa unitella Golden- H23 8 brown Tubic

Pieridae Anthocharis Orange Tip G4, H24 5,6 cardamines

Pieridae Gonepteryx rhamni Brimstone G1 5

Pieridae Pieris brassicae Large White G5, G6, H1, 6,7,8 H3, H16, H24, H25

Pieridae Pieris napi Green- E, H1, H16, 5,7,8 veined White H19, W1, W9

Pieridae Pieris rapae Small White E, H1, H16, 6,7,8 H19, H23, H24

Praydidae Prays fraxinella Ash Bud H6, H9 6 Moth

Sesiidae Bembecia Six-belted G6 7 NS (Nb ichneumoniformis Clearwing Notable)

Lepidoptera Tortricidae Epinotia abbreviana Brown Elm H1, H5, H6, 6 – Moths and Bell H9, H16, H17, Butterflies W1

Tortricidae Hedya pruniana Plum Tortrix W1 6

Tortricidae Pseudargyrotoza Yellow-spot A, H9 6 conwagana Twist

Tortricidae Ptycholoma leacheana Brindled general site 6 Twist

Tortricidae Tortrix viridana Green Oak H10 6 Tortrix

Lithobiomor Lithobiidae Lithobius forficatus G1 6 pha – Centipedes

Mecoptera Panorpidae Panorpa cognata H1, H7, W1 6 – Scorpionflie s

Planning Inspectorate Scheme Ref: TR010044 92 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Megalopter Sialidae Sialis lutaria general site 5 a – Alder Flies

Neuroptera Hemerobiidae Hemerobius lutescens G4 5 – Lacewings Hemerobiidae Micromus variegatus W9 8

Hemerobiidae Sympherobius elegans H1, H9 6,7

Sisyridae Sisyra fuscata Q 5

Odonata – Aeshnidae Aeshna cyanea Southern H9, W10 8 Dragonflies Hawker and Damselflies Aeshnidae Brachytron pratense Hairy G4 5 Dragonfly

Calopterygidae Calopteryx splendens Banded Q 5 Demoiselle

Coenagrionidae Coenagrion puella Azure G4, H22, W1 5,6,7 Damselfly

Coenagrionidae Enallagma cyathigerum Common H5 6 Blue Damselfly

Coenagrionidae Ischnura elegans Blue-tailed A1, A18, G4, 5,6,7 Damselfly H22

Odonata – Coenagrionidae Pyrrhosoma nymphula Large Red G4, Q 5 Dragonflies Damselfly and Damselflies Libellulidae Libellula fulva Scarce G4, Q 5 NS Chaser (Category 3) – IUCN Near Threatened

Libellulidae Libellula Four-spotted G4 5 quadrimaculata Chaser

Libellulidae Orthetrum cancellatum Black-tailed A1 7 Skimmer

Libellulidae Sympetrum striolatum Common H16, W1 7 Darter

Planning Inspectorate Scheme Ref: TR010044 93 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Orthoptera Acrididae Chorthippus Lesser G4, G5, G6, 7,8 – Bush- albomarginatus Marsh H9, H17, H24 crickets, Grasshopper Grasshoppe rs etc Acrididae Chorthippus brunneus Common G4, G6, H16 7 Field Grasshopper

Acrididae Chorthippus parallelus Meadow H17 7 Grasshopper

Conocephalida Conocephalus dorsalis Short- G4 7 e winged Conehead

Meconematidae Meconema thalassinum Oak Bush A, E, G4, H1, 6,7,8 includes Cricket H5, H6, H7, nymphs H9, H10, H16, H17, H22, W1, W9, W10

Phaneropterida Leptophyes Speckled A, G1, G4, 5,6,7, includes e punctatissima Bush Cricket G5, H1, H3, 8 nymphs H6, H7, H9, H10, H25, W1, W7, W9, W10,

Tetrigidae Tetrix subulata Slender G1 6 Ground Hopper

Tettigoniidae Metrioptera roeselii Roesel's G4, G6, H1, 5,6,7 Bush Cricket H9, H17, W1

Tettigoniidae Pholidoptera Dark Bush A, H6, H24, 5,6,8 includes griseoaptera Cricket Q, W9, W10 nymphs

Psocoptera Psocidae Loensia fasciata H1, W1 6 – Bark Lice

Pulmonata Cochlicopidae Cochlicopa cf lubrica G1 6 – Snails Patulidae Discus rotundatus A18 5

Raphidopter Rhaphidiidae Subilla confinis H6, H10 6 a – Snakeflies Rhaphidiidae Xanthostigma G4 5 xanthostigma

Planning Inspectorate Scheme Ref: TR010044 94 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Order Family Taxon Vernacular Site (Codes) Date Comment/ – refer to Status Table 3-1

Trichoptera Limnephilidae Limnephilus lunatus G5 7 – Caddisflies

Planning Inspectorate Scheme Ref: TR010044 95 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Plates

Plate 1 - Brown Argus (Aricia agestis) found Plate 2 - H22 showing standing dead elms widely across the target area; G5, G6, H7, which support a range of saproxylic H16, H17, W9. invertebrates.

Plate 3 – The diagnostic larval feeding Plate 4 – Six-belted Clearwing (Bembecia damage on elm, of Zigzag Sawfly ichneumoniformis) at Black Cat junction G6. (Aproceros leucopoda).

Planning Inspectorate Scheme Ref: TR010044 96 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Plate 5 - White-spotted Pinion (Cosmia Plate 6 - White-spotted Pinion larva at H20, diffinis) larva at H20, Little Barford, June Little Barford, June 6th, 2018. 6th, 2018.

Planning Inspectorate Scheme Ref: TR010044 97 Application Document Ref: TR010044/APP/6.3

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Annex A: Figures

Figure 1. Invertebrate Survey Scoping Area

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

NOTES N

LEGEND Order Limits The Scheme Scoping Area (100m) Access not granted for invertebrate surveys

Fig 2.8 Fig 2.11 Fig 2.10 Fig 2.7 Fig 2.9 Fig 2.6 Fig 2.12

Fig 2.2 Fig 2.5 d x m . a e r A

Fig 2.4 g n i p o c Fig 2.3 S

e BF 26/02/21 P01 t FIRST ISSUE a JG r b

e By t

r Revision Details Date Suffix e Fig 2.1 Check v n

I Purpose of Issue l a i r t s

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T Client

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- Highways England

6 Woodlands 1

_ Manton Lane 0 Manton Industrial Estate 3

0 Bedford 0 -

I MK41 7LW G -

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_ TR010044 Z _

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- TO CAXTON GIBBET M C

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1 Drawing Title 5 5 E H \ y

g FIGURE 1 o l o c OVERVIEW OF INVERTEBRATE E \ s p SCOPING AREA a M _ 2 0

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8 Internal Project No Suitability 2

4 60541541 D7 A \ s Scale @ A3 Zone t c e

j 1:60,000 o r THIS DOCUMENT HAS BEEN PREPARED PURSUANT TO AND SUBJECT TO THE P

TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO e LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL v i

t CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND c ONLY FOR THE PURPOSES FOR WHICH IT WAS PREPARED AND PROVIDED. A \ : Drawing Number Rev W

: Highways England Pin | Originator | Volume e m a HE551495 -ACM -EBD- n e l P01 i 0 1 2 3 4 5 Km F LGocaEtioNn _Z_Z_ZZ -| GTypSe -| G RoIle- 0 | N0um3b0er This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office © Crown copyright. Unauthorised reproduction infringes Crown copyright and may lead to prosecution or civil proceedings. Highways England 100030649 2020

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.1. Black Cat Roundabout: Areas A17, A18, G5, G6, P and Q (Part)

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

Riverside NOTES G4/A16 Farm N Nurseries

Track LEGEND Order Limits

19.8m Motor Sports Circuit

P

a

t h The Scheme

(

u

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) El Sub Sta

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X

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R 1

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Issues

A 421

Black Cat Roundabout

Balancing Pond Pa th ( um) R ock ham G5 D itch G6,A17

Balancing Pond

21.9m

T r a c k

AD RO 21.6m D OR DF BE

Roxton Garden P Centre Balancing Pond 21.0m

A18 The Bungalow

R o c k h a m D i tc h T rack 21.0m 19.5m Green Acres

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TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO S a 0 G r Cattle Grid 4 l D e r E Tank LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL e v 13 v i i

t CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND

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A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.2: Chawston (South-West Of), West of A1 Road, Area G4 (Incorporating A16

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

e

s

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c MORR 3 k IS WALK 5 55 18.9m

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5 4 B e Water g 25.0m 27 w a Invertebrate Survey Area r y 3 Dove House B 3 L r

T n E i Farm o a o 2 r k D 54 Water

3 k 1a c a r 0 1 2 T LB Waddon Lodge Pond k Scmallholdings Lyn a da r le T K imme ridge 5

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Nursery Sewage Works 26.2m Thatch Cottage

1 Pond

k

c

a r n i T a r

D Laburnham Works Cottage

11 South Well Roxmead Haven 2

The Old Foundry Nursery Brooklands Spinney Chawston El Sub Sta 21.3m View TON LANE CHAWS Farm

Box Cottage Chawston Pump FB Lodge College 20.4m Windrush Moat Farm Clare 24.7m Claygates House w Cottage o The Pump Hol lly Cottage Cottage e a

h Pump g Lilacs T n

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1 Bridge Client - 17 2

9 - Highways England 7a Brook Cottage 6 1 OAD Y R Woodlands 1 NNE SPI _ Manton Lane 1 Manton Industrial Estate 3 18

0 Bedford 0 -

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S Development Consent Order Number

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T _ A ack G4/A16 E Tr Z Nurseries R Project Title Tank G So 1 u _ 0 th Brook N

0 1 E G - Riverside A428 BLACK CAT D Farm B E

- TO CAXTON GIBBET M C

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1 Drawing Title ack 5 Tr 5 E H

\ FIGURE 2.2 y g o l CHAWSTON (SOUTH-WEST OF), WEST o c E \

s OF A1 ROAD, AREA G4 p a 19.8m Motor Sports Circuit

M (INCORPORATING A16) _ 2 0

\ Designed Drawn Checked Approved Date S I El Sub Sta SR BF LH JG 26/02/21 G \

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e Hotel j 1:5,000 o r THIS DOCUMENT HAS BEEN PREPARED PURSUANT TO AND SUBJECT TO THE P

TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO e LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL v i

t CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND c ONLY FOR THE PURPOSES FOR WHICH IT WAS PREPARED AND PROVIDED. A \

: Issues Drawing Number Rev W

: Highways England Pin | Originator | Volume e m a HE551495 -ACM -EBD- n Q e l P01 i 0 250 500 m A 421 F G5 LGocaEtioNn _Z_Z_ZZ -| GTypSe -| G RoIle- 0 | N0um3b1er This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office © Crown copyright. Unauthorised reproduction infringes Crown copyright and may lead to prosecution or civil proceedings. Highways England 100030649 2020

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.3. Quarry Area Q East of Black Cat Roundabout and R (Rectory Farm)

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

Glebe Farm

Track NOTES N L T E Nursery

Water 22.3m

Bean Wood LEGEND Order Limits e

s

u

O

t

a

e r

G Water r The Scheme e

v i

R Scoping Area (100m) 24.1m Invertebrate Survey Area

NVC Woodland

26.5m

26.2m

MP .5

k Trac

1

2

The Barns

Pond

21.0m

D rain H25

L ET

8 9 11 12 e 10 s 7 u 6 O t SL a e r 22.9m G r e iv R MP .25 5 d x 2 m 4 . 3 y

e 1 v r u S e t a

r R BF 26/02/21 P01 b FIRST ISSUE e JG t n r i a r e D By v Revision Details Date Suffix n

I Check l a i 27.7m Purpose of Issue r t s e r r e DCO APPLICATION T

3 Q

1 Client - 2

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6 Woodlands 1

_ Manton Lane 1 Manton Industrial Estate 3

0 Bedford 0 -

I MK41 7LW

G Silos -

S FB Development Consent Order Number G - Z

n

Z i a r _ TR010044 D Z _

Z Project Title _ N E G -

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D a r Rectory Farm D B E

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M L ET C Dra

A IMPROVEMENTS in - 5 9 4

1 Drawing Title 5 5 E H

\ 19.2m FIGURE 2.3 y g o l QUARRY AREA Q EAST OF BLACK CAT o c E

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8 Internal Project No Suitability 2

4 60541541 D7 A \

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c 21.0m e E j 1:5,000 T L o r THIS DOCUMENT HAS BEEN PREPARED PURSUANT TO AND SUBJECT TO THE in a P r

TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO Po D e p lar LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL v S i pi t n CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND ne c y ONLY FOR THE PURPOSES FOR WHICH IT WAS PREPARED AND PROVIDED. A \ : Drawing Number Rev W

: Highways England Pin | Originator | Volume e m a HE551495 -ACM -EBD- n e l P01 i 0 250 500 m F LGocaEtioNn _Z_Z_ZZ -| GTypSe -| G RoIle- 0 | N0um3b1er This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office © Crown copyright. Unauthorised reproduction infringes Crown copyright and may lead to prosecution or civil proceedings. Highways England 100030649 2020

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.4. Alington’s Land (Mainly) Near Little Barford, Areas H20, H22, H23, H24 And H25

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

NOTES 29.3m N

Silos 32.9m H20

Top Farm LEGEND Track Order Limits

44.8m The Scheme

Pond Issues Scoping Area (100m)

49.7m NVC Woodland Pond

52.1m

MP .75 Tr Boys Wood ack Issues

Bean Wood

53.0m

Track H22

Highbarns

Alington Hill H23

MP .5

SP 50.0m

T rack 11 12 7 8 9 10 55.8m 6

5 d x 2 m H24 4 . 37.2m 3 y

e 1 v r u S e t a r BF 26/02/21 P01 b FIRST ISSUE e JG t r e By v Revision Details Date Suffix n

I Check

32.3m l a i Purpose of Issue r t s e r r e DCO APPLICATIO N T

3

1 Client - 2

- Highways England

6 Woodlands 1

_ Manton Lane 1

n Manton Industrial Estate i 3 a r 0 D Bedford 0

- 28.3m I H25 MK41 7LW G -

S Development Consent Order Number G -

Z Sir John's Wood Z

_ TR010044 Water Z _ Tra Z ck Project Title _ N

E SL G - A428 BLACK CAT D B

E MP .25

- TO CAXTO N GIBBET M C

A IMPRO VEMENTS - 5 9 4

1 Drawing Title 5 5 E H

\ FIGURE 2.4 y

g 25.0m o l ALINGTO N’S LAND (MAINLY ) NEAR o c E \

s LITTLE BARFO RD, AREAS H20, H22, p a

M H23, H24 AND H25 _ 2 0

\ Designed Drawn Checked Approved Date S I SR BF LH JG 26/02/21 G \

8 Internal Project No Suitability 2

4 60541541 D7 A \

s Scale @ A3 Zone t c e j T 1:5,000 rack o in r THIS DOCUMENT HAS BEEN PREPARED PURSUANT TO AND SUBJECT TO THE a r D P

TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO e LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL v i Dr t ain CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND c ONLY FOR THE PURPOSES FOR WHICH IT WAS PREPARED AND PROVIDED.

A 24.4m \ : Pond Drawing Number Rev W

: Highways England Pin | Originator | Volume e m a HE551495 -ACM -EBD- n e l P01 i 0 250 500 m F LGocaEtioNn _Z _Z _Z Z -| GTypSe -| G RoIle- 0 | N0um3b1er This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office © Crown copyright. Unauthorised reproduction infringes Crown copyright and may lead to prosecution or civil proceedings. Highways England 100030649 2020

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.5. Alington’s Land (Mainly) Near Little Barford, Areas H17, H19 And H20

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

NOTES N 31.1m 28.0m B 1046 29.3m Drain 25.3m

36.9m

D ra in

Pond LEGEND D ra in Rectory Farm Cottage Order Limits Rectory Farm

2 Rectory Farm The Scheme Cottage

n i

a r

D Scoping Area (100m) 38.4m

Silos

ETL

P O T T O N R O A D

T r a c k

T ra c k

Parker's Farm

Mast Parkers Farmhouse Eynesbury The Bungalow Warehousing Gallow Hill Drain Pond Reservoir (covered)

Pond

42.1m Pond

in ra D Bramley Bungalow

Glen Eden

n i n

i

a

r a

r

D

D

43.0m

P O T T O N

R O A 12 D 11 7 8 9 10 Abbotsley Golf Course 6 and Hotel 5 45.7m d x 2 m 4 . 3 y

e 1 Post v

r Ponds u

S H17 and Pond e t a r BF 26/02/21 P01 b grassland FIRST ISSUE e JG t r e Issues By v Revision Details Date Suffix n

I Check l a i Purpose of Issue r t s e r r e DCO APPLICATIO N T

3

1 Client - 2

- Highways England

6 Woodlands 1

_ Manton Lane 1 Pond Manton Industrial Estate 3 47.5m

0 H19 Bedford 0 -

I MK41 7LW Pond G - Golf Driving Range S Development Consent Order Number G - Z Z

_ P TR010044 O T Z T O _ N

R Z O Project Title A _ D 22 N E

G 14 - A428 BLACK CAT D

1 9 10

B 1

5

E k

1 c - 1 TO CAXTO N GIBBET a

r

T M C 1

A IMPRO VEMENTS - 5

9

9 6 Abbotsley Country Homes 4

1 Drawing Title 5 El Sub Sta 5 48.8m Pond E k c a r H T Jenny Wisson Wood \ FIGURE 2.5 y 47.2m g H20 o l

5 ALINGTO N’S LAND (MAINLY ) NEAR o

c 2

E n Pond i \ a r

s LITTLE BARFO RD, AREAS H17, H19 D p

a Pond

M AND H20 _ 2 0 D 46.6m \ R Designed Drawn Checked Approved Date

E

S W I E 48.8m L SR BF LH JG 26/02/21

S G

\ L

A

8 N Internal Project No Suitability

E 2

4 60541541 D7 A \

s Scale @ A3 Zone t

c ) f

o e e j 1:5,000 s r

u o o r c THIS DOCUMENT HAS BEEN PREPARED PURSUANT TO AND SUBJECT TO THE (

D P

A

TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO

O e R LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL N v i A

t CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND M

c O R ONLY FOR THE PURPOSES FOR WHICH IT WAS PREPARED AND PROVIDED. A \ : 50.6m Drawing Number Rev W Silos

: Highways England Pin | Originator | Volume e

m 32.9m a HE551495 -ACM -EBD- n e l P01 i 0 250 Top Farm 500 m F LGocaEtioNn _Z _Z _Z Z -| GTypSe -| G RoIle- 0 | N0um3b1er This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office © Crown copyright. Unauthorised reproduction infringes Crown copyright and may lead to prosecution or civil proceedings. Highways England 100030649 2020

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.7. East of St Neots, Near Wintringham Hall And Fox Holes, Areas H16, W9 And W10 And Ash

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

NOTES N

Tra ck

Sheep Walk Plantation

Fox Holes LEGEND Order Limits

k c a r T The Scheme

Scoping Area (100m)

T ra ck Invertebrate Survey Area

k c a r T NVC Woodland

42.2m

D

A ) f O o R

e N s r A u o M c O (

R

Fox Holes

W9

46.4m

ETL

D rain

n i

a r

D

k

c

a

r

T ETL

in a r D

11 12 7 8 9 10 6

Lay-by 42.5m A E CAMBRROIDAGD 5 d ash x 2 m 4 . 3 y

e 1 v A r 428

u H16 A S 42

8 e t 1 Wintringham W10 2 a Cottages r BF Lay 26/02/21 P01 b -b FIRST ISSUE y Tank e JG t A 428 r 43.2m Track 54.1m e in By

v a r Suffix D Revision Details Date n Pond n i I Check

a

r

l MP

D a i Wintringham Purpose of Issue r CAM t B RID GE s R OAD

e T rack r Toll Gate Cottage r La y-by e 52.3m DCO APPLICATION

T Pond

Eltisley Manor 3

1 Client - A 4 3 28 4 2 Wintringham Cottages A - 428 Highways England

6 Woodlands 1

_ L Manton Lane ay-by 1 AD Manton Industrial Estate 3 O E R Moat DG

0 I BR AM Bedford

0 C

- k I rac MK41 7LW T Medieval Moat G

- Village of 51.1m Wintringham S n Wintringham Farm Drai Development Consent Order Number

G (site of) - Wintringham Hall Z Z

_ TR010044 FB

Z Cottage _

Z Project Title _

N Moat E G - k A428 BLACK CAT c

D a Tr Pond B

E Pond

- TO CAXTON GIBBET Weald M

C Pond

A IMPROVEMENTS Pond - 45.4m Pond 5 9 4

1 Drawing Title

5 51.8m Weald Farm 5 Pump k

E c a r T Ponds

H Pond

\ Pond FIGURE 2.7 y g o l EAST OF ST NEOTS, NEAR o Pond c Pond E \

s WINTRINGHAM HALL & FOX HOLES, p a Pond

M H16, W9, W10, HEDGEROW A AND ASH _ Pond 2 0

\ Designed Drawn Checked Approved Date S I Wintringham Lodge Pond SR BF LH JG 26/02/21 G \

8 Pond Internal Project No Suitability 2

4 60541541 D7 A \

s Scale @ A3 Zone t c Medieval Village of Weald e (site of) Pond j 1:5,000 Medieval Village of Weald o r 43.9m THIS DOCUMENT HAS BEEN PREPARED PURSUANT TO AND SUBJECT TO THE

P Pond

TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO e LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL v i

t CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND c ONLY FOR THE PURPOSES FOR WHICH IT WAS PREPARED AND PROVIDED. 47.5m A \ : Drawing Number Rev W Wintringham

: Highways England Pin | Originator | Volume

e k

c

a

r m ) T k c a a r T HE551495 -ACM -EBD- n ( E e N l P01 A i 0 250 500 m L

L

F I A H LGocaEtioNn _Z_Z_ZZ -| GTypSe -| G RoIle- 0 | N0um3b1er This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office © Crown copyright. Unauthorised reproduction infringes Crown copyright and may lead to prosecution or civil proceedings. Highways England 100030649 2020

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.8. North of Croxton and Staggered A428 Junction, West of Minor Road, Areas H9, H10 And W7

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

NOTES N

57.3m

LEGEND Order Limits Pond The Scheme

57.5m Scoping Area (100m)

Invertebrate Survey Area

NVC Woodland

56.8m

Pond

ETL

n

i

a

r

D

n

i

a

r

D

Pond

52.4m

G a Brook l w l allo o G w

B r o o k

New Gorse

k ac Tr

T ra c k

54.0m

ETL H9

n i

a

r W7 D 11 12 7 8 9 10 H10 6

k

c

a

r T 5 d x

n 2 i m a 4 r .

D 3 y

e 1 v r u S e t a r BF 26/02/21 P01 b FIRST ISSUE e JG t r

n e i By

a v r Revision Details Date Suffix D n

I 53.2m Check l a i Purpose of Issue r t s e r r e DCO APPLICATION T

3

1 Client - 2

- Highways England

6 Woodlands 1

_ Manton Lane 1 Manton Industrial Estate 3

0 Bedford 0 -

I MK41 7LW G -

S Development Consent Order Number G - 2 to 5

Z 6 7 Z

_ TR010044

Z El Sub Sta _

Z Drain Industrial Estate Project Title

1 _ N

E Pond 55.0m

G WB - A428 BLACK CAT D A 4 28 55.5m B

E AD - O TO CAXTON GIBBET R GE RID M MB CA C

A IMPROVEMENTS - Drain 5

9 Mast (Telecommunications) White Hall 4 55.8m Drain

1 Drawing Title

5 Pond

5 k c a r E T d n North Farm H MP o \ P FIGURE 2.8 y g o l NORTH OF CROXTON & STAGGERED o

c 56.5m

E Pond \

s A428 JUNCTION, WEST OF MINOR p Pump a House

M ROAD, AREAS H9, H10 AND W7

_ LB

2 El Sub Sta 428 TCB

0 A

\ Designed Drawn Checked Approved Date S I Weald 26/02/21 4 Cottages Pond SR BF LH JG

1

G 2 \

2

8 55.7m Internal Project No Suitability

2 D GE ROA 4 ID A 428 CAMBR 60541541 D7

A 54.1m \

s Scale @ A3 Zone t

c n i

a

r e

j 1:5,000 D o r THIS DOCUMENT HAS BEEN PREPARED PURSUANT TO AND SUBJECT TO THE 57.9m P

TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO e LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL v i

t CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND c Eltisley Manor ONLY FOR THE PURPOSES FOR WHICH IT WAS PREPARED AND PROVIDED. A \ :

6 1 Drawing Number Rev W

: Highways England Pin | Originator | Volume e m a TEexlcehpahnogne HE551495 -ACM -EBD- n e l P01 i 0 250 500 m F LGocaEtioNn _Z_Z_ZZ -| GTypSe -| G RoIle- 0 | N0um3b1er Pond This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office © Crown copyright. Unauthorised reproduction infringes Crown copyright and may lead to prosecution or civil proceedings. Highways England 100030649 2020

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.9. North of Croxton and Staggered A428 Junction, East of Minor Road, Areas H5, H6 And H7

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

NOTES

Pond N

57.5m LEGEND

ETL Order Limits

Track Pond

Tra The Scheme ck

Scoping Area (100m)

NVC Woodland

56.8m

n i Tra a ck

r

D

)

m

u (

h t

n a i

P a r

D

n

i

a

r

D

H7 H27 H6 H5

52.4m

Brook Gallow

D

r a

i H9 n

Pond

Trac Ingles k Spinney

54.0m

k

c

a

r

T

Ga llow Brook

n i

a

r

D

Tr ack

D rain

n i

a r

D

11 12 7 8 9 10 6

Pond 53.2m 5 d x 2

m 4 . 3 y

e 1

v k

c r

a

r u

T S

e t a r BF 26/02/21 P01

b FIRST ISSUE

e JG t r

e By v Revision Details Date Suffix n

I Check

l a i Purpose of Issue r t s e r 2 to r 5 6 e 7 DCO APPLICATION T

Issues 3

1 El Sub Sta Client - Industrial Estate 2 - 1 Highways England

6 Woodlands 1

_ Manton Lane 1 WB Manton Industrial Estate 3

0 Bedford

0 Pond -

I MK41 7LW G -

S North Development Consent Order Number

G Lodge -

Z Plantation Z Mast (Telecommunications) White Hall _ TR010044 Z

_ Pond

Z Project Title _

N Track Pivet and Gorse Plantation E G - El Sub Sta A428 BLACK CAT D 58.7m B E

- Pond TO CAXTON GIBBET T

r

a M c Pump A 428 k C House

A IMPROVEMENTS 59.7m -

5 LB

9 A TCB 428 4

1 Drawing Title

1 5

5 The Downs E Hall H

\ FIGURE 2.9 Downs Barn North y North Lodge g Plantation

6 o l Downs Barn East NORTH OF CROXTON & STAGGERED o c E \

s A428 JUNC, EAST OF MINOR ROAD,

p Downs Barn South Village Oak Plantation k a c a 9 Plantation r T M Cattle AREAS H5, H6, H7 AND H27 Grid _ 2 0

\ Designed Drawn Checked e Approved Date North Lodge s

u S

o I

H

l SR BF LH JG 26/02/21 o G

o \ h

c

3

1 Old School 8 S Internal Project No Suitability Sawmill 2 Pond Cottage 4 Pond Drain 60541541 D7 A 60.9m \ Elmwood House

s Scale @ A3 Zone

t 5

1

c Drain e 1

2 j Ponds Pond Pond 1:5,000

o Pond r THIS DOCUMENT HAS BEEN PREPARED PURSUANT TO AND SUBJECT TO THE 7 1 63.2m P

62.4m TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO T E e

E Lay-by LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL R v i T

t CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND S

c H Mill Plantation

G ONLY FOR THE PURPOSES FOR WHICH IT WAS PREPARED AND PROVIDED. 2 I 1 A H \ Broad Moor Spinney : Drawing Number Rev W Croxton

: Pond Highways England Pin | Originator | Volume e m a HE551495 -ACM -EBD- n

e 60.7m

l P01 i 0 250 500 m

F Pond LGocaEtioNn _Z_Z_ZZ -| GTypSe -| G RoIle- 0 | N0um3b1er This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office © Crown copyright. Unauthorised reproduction infringes Crown copyright and may lead to prosecution or civil proceedings. Highways England 100030649 2020

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.10. Eltisley Area G1

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

61.6m NOTES N Poultry Houses

61.9m

LEGEND

n

i

a

r

D Order Limits

The Scheme

Pond Track Scoping Area (100m) D

r a

i H27 Fairview n Farm Invertebrate Survey Area

62.8m NVC Woodland

Pond

H27

63.5m

D

A

O

R

S

E

V I

T W11 S

Track

Track

63.8m W12

Pillar Plantation

AD O R E 64.0m G ID BR AM n C ai in Dr ra Pond D

Track 11 12 7 8 9 10 6

5 d x 2 m 4 . 3 y

e 1 v r u S

H30 e t a r BF 26/02/21 P01 b FIRST ISSUE e JG t r e By v 64.2m Revision Details Date Suffix n

I Check l a i Purpose of Issue r MP t s

e in

r a r r D e DCO APPLICATION T

3 MS 64.1m

1 Client - 2

- Highways England

6 Woodlands 1

_ Manton Lane 1 Manton Industrial Estate 3 8 42 A 0 Bedford 0 -

I MK41 7LW G -

S Development Consent Order Number G -

D Z r a

Z i n G1 in _

T a s i TR010044 r

D t

r r Pond n

a Z u

c n o

k e

_ T C

Z D Project Title A _ O R

N E k c G Pond a ID E r El Sub Sta T R B

G M S A - H28 T C Lion House A428 BLACK CAT 28 RIV D 4 A OE SA D B E

- TO CAXTON GIBBET M C

A Pond IMPROVEMENTS

- 1 5

9 Croxton Old Rectory The S 4 Eltisley T T HO 1 7 M (PH) AS Drawing Title 5 5 rain E D 63.2m k

c 5 a H

r \ T FIGURE 2.10 y 4 4 g

o 7 l Mill Plantation 63.9m ELTISLEY AREA G1 AND H30 AND o c 18

1 1 E

\ 62.1m 22 5

s AREAS NORTH OF A428 - H27, H28, Pumping 26 House 36 Pond p 46 26 Pond a

14 30 M W11 AND W12 34 _ 61.6m in ra 7 D 1 2

0 38 TCB \ Designed Drawn Checked Approved Date ST N 1 EOTS RO r 2 AD 0 e LB S 4 lt

I e h S SR BF LH JG 26/02/21

G 0 6

\ 62.0m

8 8 39 Internal Project No Suitability 5 3 1 52 2 The 64.2m 54 4 Green

2 3 Moat 60541541 D7 A \

s Scale @ A3 Zone t 62.7m c Lych e Gate j The Green 1:5,000 Hall o Pond 81 r St John the Baptist's THIS DOCUMENT HAS BEEN PREPARED PURSUANT TO AND SUBJECT TO THE Church 83 20 P

TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO Cricket Ground 31 e rain k D LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL

v c

i a r t T El CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND 6 c 7 Pavilion 6 Sub Church 5 21 ONLY FOR THE PURPOSES FOR WHICH IT WAS PREPARED AND PROVIDED. Sta A The

\ Old House

: 5 9 9 9 2

3 55 G Drawing Number Rev R W 63.6m E

E

: The Rectory S N Highways England Pin | Originator | Volume FIELD Ponds 1 N F 2

e E GRE 1 I E

1 L 51 5 D m Church End

S a 17 19 HE551495 -ACM -EBD- n

e 6 47 l 1 P01 i 0 250 500 m

F 63 GEN_Z_Z_ZZ -| GTypSe -| G RoIle- 0 | N0um3b1er 6 Location Drain 7 This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office © Crown copyright. Unauthorised reproduction infringes Crown copyright and may lead to prosecution or civil proceedings. Highways England 100030649 2020

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.11. West of Caxton Gibbet Roundabout, South of A428, Areas H1 And H3

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

NOTES

Papley Grove Farm N

LEGEND

k c a r T Order Limits

The Scheme

Scoping Area (100m)

Pond Drain

n i

a r

D

Reservoir

n i

a

r

D

Water Pond

Pembroke Farm Solar Panels

Pond

North East Farm

n i

a r

D W13 Ponds

ROAD TCP RIDGE (AA) CAMB

Lay-by

428 H1 64.8m A 11 12 7 8 9 10 Mile Post 64.5m 6 Drain CAMBRIDGE ROAD

Drain 64.2m

63.7m 5 d x 2

m 4 . 3 y A 428

e 1 v r u

S 63.5m

e H3 t a r BF 26/02/21 P01

b FIRST ISSUE

e JG t r

e By v Revision Details Date Suffix n

I Check

l a i Purpose of Issue r t s e

r 63.8m r

e DCO APPLICATION T

3

1 Client - 2

- Highways England D 6 r a Woodlands

1 in

_ Manton Lane

1 T r a Manton Industrial Estate 3 c k 0 Bedford 0 -

I MK41 7LW G - D S ra Development Consent Order Number in G - Z n Z ai Dr _ TR010044 Z _

Z Project Title _ N E

k

G c

a -

r

T A428 BLACK CAT D B E

- TO CAXTON GIBBET M C

A IMPROVEMENTS - 5 9 4

1 ck Tra Drawing Title 5 5 E H

\ FIGURE 2.11 y

g M oa o t l WEST OF CAXTON GIBBET o c E \

s ROUNDABOUT, SOUTH OF A428, p a

M AREAS H1 AND H3 _ 2 0

\ Designed Drawn Checked Approved Date S I SR BF LH JG 26/02/21 G \

8 Internal Project No Suitability

2 Pastures Farm 4 60541541 D7 A \

s Scale @ A3 Zone t c Bungalow e The

j 1:5,000 o

r The Dovecote THIS DOCUMENT HAS BEEN PREPARED PURSUANT TO AND SUBJECT TO THE P

TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO

e LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL v i

t CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND

c ONLY FOR THE PURPOSES FOR WHICH IT WAS PREPARED AND PROVIDED. Pond A \ : Drawing Number Rev W Moat

: Pond Highways England Pin | Originator | Volume e Moat m a HE551495 -ACM -EBD- n e

l P01 i 0 250 500 m F LGocaEtioNn _Z_Z_ZZ -| GTypSe -| G RoIle- 0 | N0um3b1er This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office © Crown copyright. Unauthorised reproduction infringes Crown copyright and may lead to prosecution or civil proceedings. Highways England 100030649 2020

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Figure 2.12. East of Caxton Gibbet Roundabout, South of A428, Areas A1 And W1

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3

D ra in 52.7m NOTES N

E r m i n R e O S M t r A e e N t R O A D k c ra T LEGEND

D r Order Limits a in

64.6m The Scheme

Scoping Area (100m)

Track T rack Invertebrate Survey Area

Drain Tra ck NVC Woodland

Drain

4

1

Common Farm Cottages

62.2m 65.8m

Pond

A

1 1 9 8 Papworth Hotel

D r a in

H29

65.2m

n i a r

D 66.4m L a y - b y

D r a in

Cycle Way

A 12

1 11 1 8 9 9 10 8 7 A 428 A1 6 A 428

Drain Balancing Pond 5 d in 66.8m x 8 Dra A 42 65.9m 66.5m 2 m 4 . 3 y 28 e A 4 1 v MP r u Pond S

ESS e W13 W1 67.2m t

2 Oak Tree Cottage A 428 a

1 r n BF ing Statio 26/02/21 P01 b ll FIRST ISSUE Fi A 1

e 1 9 JG t 8

r New Bungalow ROAD TCP RIDGE ay-by e L 65.2m (AA) CAMB By v Revision Details Date Suffix 4 n A I E Check in 42 -by R 8 ay a D l L M r

Caxton Gibbet D r IN a a

i i E n Purpose of Issue r S A t T 42 R in 8 s Dra E Drain E

e T r r e DCO APPLICATION

T H1

3

1 Client - 65.8m 2

- Highways England

6 Woodlands 1

_ Manton Lane R 1 O Manton Industrial Estate M 3 E A r N m 0 i Bedford R n e

0 O

A S - t D r

I MK41 7LW e e t G -

S Development Consent Order Number G - Z Z

_ TR010044 Z _

Z Project Title _ N E G - A428 BLACK CAT D B E

- TO CAXTON GIBBET n M ai Dr C 66.1m

A IMPROVEMENTS - 5 9 4

1 Drawing Title 5 5 E H

\ FIGURE 2.12 y D

r g a i n o l IN THE VICINITY OF CAXTON GIBBET o c E \

s ROUNDABOUT, AREAS A1, H1, H29, W1 p a

M AND W13 _ in 2 a Dr 0

\ Designed Drawn Checked Approved Date S I MS SR BF LH JG 26/02/21 G

\ Swansley

8 Internal Project No 66.0m Wood Suitability 2

4 in Dra 60541541 D7 A \

s Scale @ A3 Zone t c Farm e j Pond 1:5,000 o r THIS DOCUMENT HAS BEEN PREPARED PURSUANT TO AND SUBJECT TO THE P

TERMS OF AECOM'S APPOINTMENT BY ITS CLIENT. AECOM ACCEPTS NO e LIABILITY FOR ANY USE OF THIS DOCUMENT OTHER THAN BY ITS ORIGINAL v i

t CLIENT OR FOLLOWING AECOM'S EXPRESS AGREEMENT TO SUCH USE, AND c ONLY FOR THE PURPOSES FOR WHICH IT WAS PREPARED AND PROVIDED. A \ : Drawing Number Rev W

: Highways England Pin | Originator | Volume e m a HE551495 -ACM -EBD- n e l P01 i 0 250 500 m F LGocaEtioNn _Z_Z_ZZ -| GTypSe -| G RoIle- 0 | N0um3b1er This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office © Crown copyright. Unauthorised reproduction infringes Crown copyright and may lead to prosecution or civil proceedings. Highways England 100030649 2020

A428 Black Cat to Caxton Gibbet improvements Environmental Statement – Appendix 8.16: Terrestrial Invertebrates (Including Non-Native Species

Planning Inspectorate Scheme Ref: TR010044 Application Document Ref: TR010044/APP/6.3