Hamburg, Dezember 2006 Mitt. hamb. zool. Mus. Inst. Band 103 S. 137-147 ISSN 0072 9612
Additions to the catalogue of the type specimens of the herpetological collection in the Zoological Museum of the University of Hamburg
JAKOB HALLERMANN
Universität Hamburg, Biozentrum Grindel und Zoologisches Museum, Martin-Luther-King-Platz 3, D-20146 Hamburg, Germany. E-mail: [email protected]
ABSTRACT. – 65 types specimens of amphibians and reptiles found in the general collection during the last ten years since the first type catalogue was published are provided together with types of recently described species. For each taxon the present name and its status are given. Taxonomic histories for some taxa are reviewed. The herpetological collection of the Zoological Museum Hamburg includes 318 type specimens, representing 137 named taxa.
KEYWORDS: Type catalogue additions, Museum Hamburg, Amphibia, Reptilia, taxonomic history
Introduction Since the first herpetological type catalogue of the Zoological Museum Hamburg was published (HALLERMANN 1998) 65 type specimens of 23 nominal type species, formerly thought to be lost, were detected by the author in the collection. The new findings are listed below. The identities of some extant type specimens were resolved previously (CADLE 1998; HALLERMANN 2000; RÖDEL & HALLERMANN 2006; ROSSMAN 2000) and many others are established in this paper. Some new species were also described from the collection (BAUER 2002; PIEH & PERÄLÄ 2002). The addition of remarkable types of historical value to the collection necessitate an update to the existing type catalogue. The herpetological collection of the ZMH contains currently about 50 000 specimens. About 17 000 specimens in 6100 lots are catalogued in a data file (FileMaker Pro 7). At present the collection contains 318 type specimens of 137 nominal species and subspecies i.e. 54 holotypes, 69 syntypes, 147 paratypes, 3 lectotypes and 45 paralectotypes. Each species or subspecies is listed in the combination as it was originally proposed. If the name has been subjected to subsequent nomenclatorial change or has been given in synonymy, the name in current use is given after an equal sign, together with a citation of the authority for the change. The arrangement in the following list is alphabetical by genus and species within each family. Localities are listed in the original spelling from the label, in English translation and if possible with coodinates and the present name of the state. Corrections of spelling mistakes and change of status of some types listed in the first type catalogue are added. Also types of questionable status are listed. For some the status could not be resolved because all documents 138 JAKOB HALLERMANN
(correspondence, catalogues, type lists) were destroyed during Word War II. (HALLERMANN in press). The family and genus concepts used follow the EMBL Reptile Data Base (http://www.reptile-database.org/) for reptiles and the amphibian species of the world data base (FROST 2006).
List of type specimens Museum acronyms BMNH The Natural History Museum, London [former British Museum (Natural History)] MCZ Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts MSNG Museo Civico di Storia Naturale “G. Doria” di Genova NMW Naturhistorisches Museum Wien SMF Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt SMNS Staatliches Museum für Naturkunde Stuttgart ZMB Museum für Naturkunde der Humboldt-Universität zu Berlin (former Zoologisches Museum Berlin) ZMH Zoologisches Museum Hamburg
Amphibia Urodela Plethodontidae
Plethodon yonahlossee DUNN, 1917 Type locality. – near the Yonahlossee Road about 1 1/2 miles from Linville, Avery County, North Carolina, altitude 4200 feet, USA, leg. E. R. DUNN, 16.VIII 1916, rec. NOBLE 2.X.1925, 1 paratype ZMH A04286 (formerly AMNH 4633). Comments: holotype is AMNH 4634. Our specimen of the type series with 117 mm total length, 16 mm head length, 48 mm body length, and 53 mm tail length corresponds exactly with DUNN’S data (DUNN 1917).
Anura Megophryidae
Leptobrachium carinense BOULENGER, 1889 [=Brachytarsophrys carinensis sensu TIAN & HU (1983)] Type locality. – Yado (Yado Taung MT 19°19’N 96°50’E), Montagni Carin (=Karen mountains), 1200-1300 m about sea level, Burma (Myanmar), L. FEA leg. 1885-89 on his travel in Burma, 3 paralectotypes ZMH A04092-4 (formerly no. 740, field no. 41, lot 1°).
Comments: The original type locality was given by BOULENGER (1889) as “western slope of the Karens Mountains, east of Toungoo at an attitude of 800 m and upwards”. BOULENGER noted, that FEA obtained several specimens; he gave measurements for a female (168 mm) and a male (150 mm). ZMH A04092 (107 mm): leg. FEA Xbrie (October) 1887; A04093 male (150 mm): leg. FEA Genn. (January)1889; A04094 (130.6 mm): leg FEA XII 1888. Lectotype MSNG 29689 from Karin Bia-po was designated by CAPOCACCIA 1957, 3 paralectotypes in Genova: one from Karin Bia-po 900-1200 m, one from Carin Cheba 900-1200 m, and one from Yado 1200-1400 m, 4 paralectotypes from Karin Hills in BMNH, 2 paralectotypes from West slope of Karin Mountains in NMW. Today I cannot verify which and how many specimens belonged to the type series. Additions to the herpetological type catalogue, Hamburg 139
Ranidae
Phrynobatrachus monticola FISCHER, 1884 [= Tomopterna monticola sensu RÖDEL & HALLERMANN (2006)] Type locality. – Bach (streamlet) Wasso-Njiro, „Massailand“, (Kenya, north of Lake Natron in the border region to Tanzania (approximately between 1°30' and 2°04 S / 36°12' E)), leg. G.A. FISCHER between 19 April and 1 May 1883, holotype ZMH A04402 (formerly no. 388).
Comments: det. by Fr. NIEDEN as Phrynobatrachus delalandii Bibron, was considered “incerta sedis” by FROST (2006), collection details see G. A. FISCHER (1885).
Rhacophoridae
Chirixalus doriae BOULENGER, 1893 Type locality. – Bia-po, (Bia-po Keba, District Karen-ni, 19°15' N, 97°30' E) Montagni Carin Ghecú, in 900-1000 m, Burma (Myanmar), L. Fea leg., during his travel in Burma, 1885- 1889, 1 syntype ZMH A03145 (formerly number 722, field No. 26, lat 1°). Comments: There were 5 type specimens from District of the Karin Bia-po“ and 1 specimen from „Thao“ (BOULENGER 1893). CAPOCACCIA 1957 designated a lectotype MSNG 29426A and 3 paralectotyes. All were in bad condition. The specimens are no longer present in MSNG (DORIA et al. 2001). HÄUPL & TIEDEMANN (1978), and HÄUPL et al. (1994) listed one specimen (NHW 16556) from “Karin-Gebirge” without specific locality as a syntype. It seems more probable that the ZMH specimen is a syntype, until a new lectotype designation will be made for the lost lectotype.
Ixalus carinensis BOULENGER, 1893 [= Philautus carinensis sensu SMITH (1940)] Type locality. – Montagni Carin (=Karen mountains) in 900-1000 m, Burma (Myanmar), Kayah State 19° 15’N 97° 30’E, L. FEA leg., during his travel in Burma, 1885-1889, 4 (?) paralectotypes ZMH A03138-41 (formerly number 702, field no. 23, lat 1°).
Comments: status not clear. According to BOULENGER (1893) types are from Karin Hills (1), Thao (1), and Karin Bia-po (5). There are more type specimens listed and they did not correspond to the given localities: three specimens in Genova are from Thao (CAPOCACCIA 1957), four from Karin Hills in BMNH (MCCARTHY pers. comm. 2000). BOSSUYT & DUBOIS (2001) designated BMNH 1947.2.6.24 from Karin Hills, “2900-3500 feet” as lectotype. It seems that they did not examine other probable type specimens other than the BMNH specimens. This did not clarify the situation because the specimens from Karin Bia-po could not be located with certainty and no paralectotypes were explicitly noted by BOSSUYT & DUBOIS (2001). It is not currently possible to verify which specimens belong to the type series because no measurements exist other than that of a male of 38 mm length (the lectotype ?). It may be that at least one of the specimens in Hamburg is a paralectotype.
Ixalus parvulus BOULENGER, 1893 [= Philautus parvulus (BOULENGER) sensu BOURRET (1942): 451] Type locality. – Montagni Carin (=Karen mountains), 900-1000 m about sea level, Burma (Myanmar), Kayah State 19° 15’N 97° 30’E, L. FEA leg. 1885-89 on his travel in Burma, 2 paralectotypes ZMH A04076-7 (formerly number 724, field no. 38 lot 1°).
Comments: according to BOULENGER (1893) there were 7 type specimens from Karin Bia- po. Today the lectotype and two paralectotypes are in MSNG, one specimen from Karin Hills 140 JAKOB HALLERMANN in NMW, and two from Karin Hills in BMNH. Including the ZMH specimens there are 8 type specimens present, one too many. Existing data do not permit me to determine which specimen does not belong to the type series.
Rhacophorus feae BOULENGER, 1893 Type locality. – Thao, (Tahò ~19°23' N 96°54' E), Montagni Carin, Carin Ghecu 1300-1400 m about sea level, Burma (Myanmar), L. Fea leg. II 1888 on his travel in Burma, 1 paralectotype ZMH A04063 (formerly no. 718).
Comments: This specimen was determined as Rhacophorus nigropalmatus feae by WOLF in 1935 and as Rhacophorus feae by N. ORLOV in 2000. Lectotype MSNG 29415 designated by CAPOCACCIA (1957). Another paralectotype is BMNH 93.10.9.24.
Diploglossus (Celestus) ohlendorffi FISCHER, 1886 [= Celestus costatus costatus (COPE, 1861) J. HALLERMANN det. 2001, HALLERMANN & BÖHME (2002)] Type locality. – Haiti, Hispaniola, Zoologische Gesellschaft ded. 1885, received from Zoological Garden Hamburg, holotype ZMH R06351 (formerly no. 855).
Comments: the specimen was determined by MEERWARTH (1901) as Diploglossus striatus. Original label is missing.
Boidae
Epicarsius cupreus FISCHER, 1856b [= Epicrates cenchia cenchria (LINNAEUS, 1758), J. HALLERMANN det. (2006)]
Type locality. – Porto Cabello, Venezuela, Dr. TAMMS leg. 1851, holotype ZMH R08256 (formerly no. 49 B 17bd 51).
Chamaeleonidae
Rhampholeo boettgeri PFEFFER, 1893 [= Rhampholeon brevicaudatus MATSCHIE, 1892 sensu WERNER (1902)].
Type locality. – Bagamoyo, Tanganyika [Tanzania], Dr. F. STUHLMANN leg. 1890, holotype ZMH R07911 (formerly no. 1287).
Comments: determined as Rhampholeon brevicaudatus by WERNER (1902). On an old label was mistakenly written: “Rhampholeo boulengeri PFEFFER = Rh. brevicaudatus”.
Colubridae
Boaedon nigrum FISCHER, 1856 [=Lamprophis virgatus (HALLOWELL, 1854) det. J. HALLERMANN (2006)] Type locality. – São Thomé Is. (West Africa) [São Tomé et Principé], leg. Chirurgius WEISS 1848, 1 syntype ZMH R08378 (formerly no. 143). Additions to the herpetological type catalogue, Hamburg 141
Comments: other syntypes formerly deposited in ZMH are lost. The taxon name Boaedon nigrum or Boodon niger was, at times, variously assigned to different dark coloured species of the genus Lamprophis: L. fuliginosus, L. capensis, L. virgatus as seen during a visit in the ZMB collection.
Coronella elegans JAN, 1863 [= Meizodon regularis FISCHER, 1856, sensu FISCHER (1883), species status] Type locality. – West Africa (no specific locality, no collector), one syntype ZMH R06837 (formerly no. 136).
Comments: G. JAN returned the specimen with a label “Ablabes elegans JAN”. Total-length: 350 mm, tail-length: 70 mm, Subcaudals 64. Other syntypes (4?) were deposited in Stuttgart and are destroyed (SCHLÜTER pers. com. 2006). The right upper jaw is missing.
Helicops infrataeniatus JAN, 1865 Type locality. – Surinam (no specific locality, no collector), lectotype ZMH RO4331 (formerly no. B88 da) was designated by ROSSMAN (2000). Comments: the other syntype from Brazil is deposited in the Essex Institute of Salem according to JAN (1865) and seems to be missing (ROSADO pers com).
Leptophis varius FISCHER, 1884 [= Liopholidophis varius (FISCHER, 1884) sensu CADLE (1998)]
Type locality. – Madagascar, LÉON HUMBLOT leg, vend.1883, lectotype ZMH R04319 (formerly no. 1175), female.
Comments: formerly 5 syntypes, destroyed, the jar label reads as collector “HUMBOLD”, perhaps a mistake by copy of the old label. FISCHER (1884) noted: “gesammelt und gekauft von Herrn Humblot in Paris” [collected and bought from Mr. Humblot in Paris]. Léon Humblot was a colonist and amateur naturalist on the Comores and he obtaind specimens from Madagascar for the Paris Museum. The present specimen was designated as lectotype by CADLE (1998). In the original description 5 specimens with the catalogue numbers 1174 and 1175 were listed. The lectotype most closely matches FISCHER’S specimen „b“ in its length and pholidosis (CADLE 1998; FISCHER 1884).
Ligonirostra stuhlmanni PFEFFER, 1893 [= Prosymna stuhlmanni, sensu SPAWLS et al. (2002)]
Type locality. – Usambáa, Usambara, Deutsch Ost-Afrika [Tanzania], F. STUHLMANN leg. Sept. 1888, one syntype ZMH R07910 (formerly no. 1646).
Comments: One of the two synytpes No. 521 or 522 of PFEFFER with 132 (133-136 of PFEFFER) ventrals, 31(31-32) subcaudals, total length 190 mm (“etwa 20 cm” after PFEFFER 1893).
Rhabdosoma univittatum JAN, 1862 [= Atractus univittatus (JAN, 1862) sensu PETERS & OREJAS MIRANDA (1970)] Type locality. – Caracas, (10°30’00"N 66°55’00"W), Venezuela, holotype ZMH R02851 (formerly no. 482).
Comments: formerly det. as Atractus guentheri. MARKEZICH (2002 in lit) determined that the specimen was the holotype. 142 JAKOB HALLERMANN
Gekkonidae
Cnemaspis indraneildasii BAUER, 2002 Type locality. – Gund (15°10’N, 74°40’E), Karnataka, North Kanara (= Uttara Kannada), India, „Hütte“ = hut, G. A. v. MAYDELL leg. 20 January 1956 (Deutsche Indien Expedition 1955/58), holotype ZMH R06125, female (formerly no. 875).
Cnemaspis heteropholis BAUER, 2002 Type locality. – Gund (15°10’N, 74°40’E), Karnataka, North Kanara (= Uttara Kannada) India, „Bambus teak, Dschungel“, G. A. v. MAYDELL leg. 20. January 1956 (Deutsche Indien Expedition 1955/58), holotype ZMH R06158, female (formerly no. 234). Comments: tail autotomized.
Gerrhosauridae
Gerrhosaurus zanzibaricus PFEFFER, 1893 [= Gerrhosaurus major major DUMÉRIL, 1851, det. J. HALLERMANN (2006)] Type locality. – Kubueni, Sansibar (=Zanzibar 6°10’00"S 39°11’00"E), Tanzania, F. STUHLMANN leg, 12.V.-9.VI 1888, 3 syntypes ZMH R07964-6 (formerly numbers 1276, 249, field no. 127). Comments: specimen R07965 with 440 mm total length leg. 12.V.1888; R07964 with 460 mm total length (no colleting date), R07966: specimen head only, without body (no collection data preserved).
Hydrophiidae
Hydrophis pelamoides var. annulata FISCHER, 1855 [= Lapemis curtus hardwickii GRAY, 1835, det J. HALLERMANN (2006)]
Type locality. – Coast of Java, Indonesian Sea, Indonesia, MEYER and WERNER leg. 1847, ded. 1848, 2 syntypes ZMH R03353 (formerly no. 417), R03355 (formerly no. 418). Comments: The description was first issued in 1855 in the school program of the Realschule Johanneum. It was reprinted in 1856 by the „Naturwissenschaftlichen Verein zu Hamburg“, a scientific society and clearly labeled as the second edition (FISCHER 1856 a). It is ten pages longer than the original and there are three hand coloured plates added. ZMH R03353 is the smaller specimen: total-length: 337 mm, R03355 the larger one with total-length: 785 mm and both correspond with the measurements of FISCHER (1855).
Leptotyphlopidae
Glauconia unicolor WERNER, 1913 [= Leptotyphlops unicolor sensu WALLACH in lit. (2006)]
Type locality. – wahrscheinlich Brasilien (probably from Brazil), AUGUST BRÜCKMANN leg., ded. 9.XI.1910, holotype ZMH R08401 (formerly no. 4823).
Comments: The species was described by WERNER (1913) under Glauconia bilineata (SCHLEGEL). He wrote:” Eventuell würde ich, wenn es sich um eine neue Art handeln sollte, den Namen P. unicolor beantragen” [possibly, if it may be a new species I would like to propose the name P. unicolor]. “P” may be a spelling mistake and means “G.” Label reads: Glauconia bilineata var. unicolor Wern. Our collection houses other specimens of other species received Additions to the herpetological type catalogue, Hamburg 143 from BRÜCKMANN definitively from South America. VAN WALLACH (pers. comm. 2006) is currently examining the specimen and will publish its identity soon.
Scincidae
Heteropus rhomboidalis PETERS, 1869 [= Carlia rhomboidalis sensu GREER (1974)]
Type locality. – Port Mackay NE Autralia, most probably AMALIE DIETRICH leg. 1864-66, 1 syntype ZMH R06796 (formerly no. MG “Museum Godeffoy” 931).
Comments: as BAUER et al. (2003) pointed out the lectotype designation of BMNH 1946.8.16.57 by WELLS & WELLINGTON (1985) is invalid and the lectotype designation of ZMB 6509A by INGRAM & COVACEVICH (1989) is ambigous. Until clarification I regard the Berlin and Hamburg specimens as syntypes.
Lygosoma (Emoa) Kuekenthali BOETTGER, 1895 [= Emoia kuekenthali (BOETTGER, 1895) sensu BROWN (1991)] Type locality. – Halmaheira [= Halmahera], Moluccas [Maluku Propinsi], Indonesia, W. KUEKENTHAL leg., received from Senckenbergische Naturforschende Gesellschaft (no date), 2 paralectotypes ZMH R06794-5 (formerly no.1780).
Comments: SMF 15202 is the lectotype (MERTENS 1967), other paralectotypes in ZMB, BMNH, MCZ, SMF (BAUER et al. 2003). The published type locality is Soa Konorra and Tabella, North Halmaheira, up to 2500 m about sea level, W Halmaheira, and Pantani E Halmaheira. „in ziemlicher Anzahl erbeutet“ (caught in rather large quantities).
Lygosoma (Emoa) sorex BOETTGER, 1895 [ = Emoia sorex (BOETTGER, 1895) sensu BROWN (1991)] Type locality. – Halmaheira [= Halmahera], Moluccas [Maluku Propinsi], Indonesia, W. KUECKENTHAL leg., received from Senckenbergische Naturforschende Gesellschaft (no date), 2 paralectotypes ZMH R06791-2 (formerly no. 1783).
Comments: SMF 15340 was designated as lectotype (MERTENS 1967), other paralectotypes in ZMB, BMNH, SMF (BAUER et al. 2003). BROWN (1991) mistakenly referred to the lectotype, SMF 15340, as the holotype and did not mention the existence of other types.
Lygosoma zebratum BOULENGER, 1887 [= Sphenomorphus indicus (GRAY, 1853) sensu SMITH (1935)] Type locality. – Montagni Mooleyit in 1100-1200 m about sea level (= Muleyit mountains, Dawna Chain (Mula-a Taung 16° 07' N 98° 39' E), Tenasserim, Burma (= Myanmar ), L. FEA leg., during his travel in Burma, 1885-1889, 1 syntype ZMH R06566 (formerly no. 1444, collecting no.: 71 lat 1°).
Comments: the published type locality is Pla-poo, Mt. Mooleyt. BOULENGER (1887) noted several specimens as types but giving measurement for only one. Later BOULENGER regarded L. zebratum as a variety of L. indicum (BOULENGER 1893: 319) and TAYLOR as a subspecies of Sphenomorphus indicus (TAYLOR 1963: 1020). The ZMH syntype and BMNH 1889.3.25.23-24 specimens are smaller than the specimen for which BOULENGER (1887) provided measurements. 5 MNSG specimens also belong to the type series. G. DORIA (pers com. 2006) provided measuremens and photos of all specimens of the type series. MNSG 27883 -1 matches the data of BOULENGER (1887). 144 JAKOB HALLERMANN
Mabuya quinquetaeniata scharica STERNFELD, 1917 [= Trachylepis quinquetaeniata (LICHTENSTEIN 1823) sensu BROADLY & BAUER (1998) species status and BAUER (2003) genus status]
Type locality. – Fort Archambault, [Sarh, Chad], H. SCHUBOTZ leg. 2. Deutsche Zentral-Afrika-Expedition des Herzogs ADOLF FRIEDRICH ZU MECKLENBURG 1910-1911, 10 paralectotypes ZMH R06771-80 males (formerly no. 3478), 10 paralectotypes ZMH R06781-90 (females, formerly no. 3479), 5 paralectotypes ZMH R06764-68 (formerly no. 3480); Route Fort Crampel-Archambault [Road from Kaga Bandoro, Central African Republic to Sarh, Chad], HABERER leg. 2. Deutsche Zentral-Afrika-Expedition des Herzogs ADOLF FRIEDRICH ZU MECKLENBURG 1910-1911, 4 paralectotypes ZMH R06760-3, males (formerly no. 3481); Route Fort Possel-Fort Crampel [road between Possel and Kaga Bandoro, Central African Republic], H. SCHUBOTZ leg. 2. Deutsche Zentral-Afrika-Expedition des Herzogs ADOLF FRIEDRICH ZU MECKLENBURG 1910-1911, 2 paralectotypes ZMH R06769-70, females, gravid (formerly no. 3477).
Comments: SMF 14271 designated by MERTENS (1967) as lectotye, specimens exchanged in 1921 by RICHARD STERNFELD.
Testudines Testudinidae
Testudo graeca marokkensis PIEH & PERÄLÄ, 2004
Type locality. – near Casablanca (Morocco), H. WITTKE leg.1974, S. SCHIELZETH ded. VI 1976, 1 paratype ZMH R01092, female.
Corrections of the type catalogue of 1998 page 200, line 9 from below: Acanthosaura fruhstorferi ZMH R04357 no longer a syntype (see HALLERMANN 2000). Comments: the lectotype is NMW 1141, paralectotype SMNS 4155. page 202, line 9:“Cantonophis praefrontalis WERNER, 1909 a, b [ = Opisthropis latouchii“ not Opistrophis. page 208 line 12 from below ; Hydrophis (Pelamis) bicolor var. alternans FISCHER, 1855 not 1856 a. page 209, line 30: Lacerta cappadocica muhtari EISELT, 1979. „R00782, adult“, not „juvenile“ page 221, line 12: „OTA, H. & HIKIDA, T.“, not „OTA & ZSUTOMU, H.“
ACKNOWLEDGEMENTS. I thank COLIN MCCARTHY (BMNH) and GIULIANO DORIA for information on specimens in their care, AARON M. BAUER (Pennsylvania) and RAINER GÜNTHER (ZMB) for helpful comments on the manuscript.
References
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Received: 30 June 2006; accepted: 27 September 2006.