Coleoptera of the Idaho National Engineering Laboratory: an Annotated Checklist

Great Basin Naturalist

Volume 46 Number 2 Article 18

4-30-1986

Coleoptera of the Idaho National Engineering Laboratory: an annotated checklist

Michael P. Stafford University of Idaho, Moscow, Idaho

William F. Barr University of Idaho, Moscow, Idaho

James B. Johnson University of Idaho Moscow, Idaho

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Recommended Citation Stafford, Michael P.; Barr, William F.; and Johnson, James B. (1986) "Coleoptera of the Idaho National Engineering Laboratory: an annotated checklist," Great Basin Naturalist: Vol. 46 : No. 2 , Article 18. Available at: https://scholarsarchive.byu.edu/gbn/vol46/iss2/18

This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. COLEOPTERA OF THE IDAHO NATIONAL ENGINEERING LABORATORY: AN ANNOTATED CHECKLIST

Michael P. Stafford", William F. Barr" and James B. Johnson"

AbstR'VCT. —An insect survey was conducted on the Idaho National Engineering Laboratory during the summers of Vi 1981-1983. This site is on the Snake River Plains in southeastern Idaho. Presented here is an annotated checkUst of the Coleoptera collected. Successful collecting methods, dates of adult occurrence, and relative abundance are given for

each species. Relevant biological information is also presented for some species.

The Idaho National Engineering Labora- tory (INEL) was designated an Environmen- tal Research Park in 1975. Because of the increased number of ecological studies on the INEL since this time, there is a need for an entomofaunal list for this area. This information will be valuable to many biologists since insects interact with many other organisms, e.g., plants as pollinators and herbivores or vertebrates as food and parasites. Therefore, an insect survey was conducted on the INEL during the summers of 1981-1983. The objec- tives of this survey were to document the taxonomic composition of the insect fauna and the seasonal occurrence, relative abundance, and host information for the insect species present. Several arthropod surveys were conducted on the INEL, formerly known as the National Reactor Testing Station, under the auspices of Brigham Young University. Allred (1968) re- Eig. 1. Location and detail of the Idaho National Engi- ported seven species of ticks from a variety of neering Laboratory. hosts. Forty-two species of spiders were collected in a pitfall trap survey (Allred 1969). Allred and Cole (1971) published a checklist of Mountain Ranges in southeast Idaho (Fig. I). 22 species of ants, and Allred and Muma The topography is flat to gently rolling; mean (1971) listed the solpugid species found on the elevation is 1,490 m. Soils are generally aeo- INEL. Allred (1973) reported the occurrence lian sandy loams and loess and classified as of scorpions. Although these studies provide Aridisols (McBride et al. 1978), with scattered valuable information on the ant and arachnid lava outcroppings. The vegetation is domi- faunae, information on other insect ta.\a is nated by big sagebrush {Artemisia tridentata lacking. Nutt.) Other conspicuous shrubs include green rabbitbrush (Chrijsothamnus viscidi- Study Area floris [Hook.] Nutt.), gray rabbitbrush (C. The INEL is on the Upper Snake River nauseosus [Pall.] Britt.), gray horsebrush {Te- Plains at the base of the Lost River and Lemhi tradymia canescens D.C.), and winter fat

Published with the approval of the director of the Idaho .Agriculture Station as Research Paper 85724. ^Department of Plant, Soil and Entomological Sciences, University of Idaho, Moscow, Idaho 83843.

287 288 Great Basin Natur.\list Vol. 46, No. 2

= = (Ceratoides lanata [Piirsh] J. T. Howell). The SW Sweep Net; PE Plant Examination; dominant grasses are bluebunch wheatgrass PF = Pitfall Trap; WP = Windowpane Trap. (Agropyron spicatum [Pursh] S. & S.), Indian Relative Abundances ricegrass {Orijzopsis hijmenoides [R. & S.] Ricker), bottlebrush squirreltail {Sitanion U = Uncommon, 1-15 specimens; C = Com- mon, 16-50 specimens; A = Abundant, 51 + hystrix [Nutt.] J. B. Smith), and needle-and- thread {Stipa comata Trin. & Rupr.). The specimens. more alkaline areas are dominated by shad- The assignment of the abundance rating is scale {Atriplex confertifola [Torr. & Frem.] subjective. It is based upon both the number Wats.) and saltsage (A. nuttalii Wats.). In de- of specimens collected and on personal obser- pressions prone to flooding are stands of Great vations. Because of population fluctuations we Basin wildrye (Elyjyms cinereus Scribn. & have observed over the years, these ratings Merrifl). E.xtensive stands of Utah juniper may not always be indicative of the abundance {Juniperus osteospenna [Ton.] Little) occur of a species in that particular year. Also, the on the foothills of the surrounding mountains. collecting methods employed may not always Harniss and West (1973) described the vege- reflect the relative abundance of a species. tation patterns and presented a detailed map Coleoptera Checklist of the vegetation found on the INEL. Jeppson and Holte (1978) listed the plant species found Carabidae in the area. Agonum balesi Gray: PF; May, Jun; U. Agonum placidiim (Say): LT; Jul; U. Results Amara apricaria Paykull: LT; Jul; U.

Presented here is an annotated checklist of Amara impuncticollis Say: PF; Jul; U. the Coleoptera we collected in approximately Amara laticollis LeConte: LT; Jul; U. 800 man-hours of collecting during this survey Amara miisculus Say: PF; Jun, Jul; U. and previous collecting efforts on the INEL Amara sp. (poss. quenseli Schonherr): PF; Jul; over the last 25 years by one of us (WFB). The U. list includes 214 species in 29 families, making Axinopalpus hiplagiatus (Dejean): PF, WP; U. it the most extensive faunal list yet published Jul; for the INEL. This information should be use- Bembidion immaculosum Hatch: LT, PF; Jul, ful for those undertaking entomological and Aug; U. ecological studies in semiarid regions of the Bembidion nebraskense LeConte: LT; Aug; West, particularly in the Great Basin and U. Snake River Plains. The placement of family Bembidion obscurelhim Motschulsky: LT; names follows A Catalog of the Coleoptera of Jul, Aug; A. America North of Mexico (Kingsolver 1979). Bembidion rupicola Kirby: LT; Aug; U. Genus and species names are arranged in al- Bembidion timidum LeConte: LT; Aug; U. phabetical order. Determinations, except as Calosoma luxatmn Say: PF; May-Jul; U. noted in the acknowledgments, were made by Cicindela decemnotata Sav: PF, GC; Mav- Aug; U. the authors and Paul J. Johnson (also at the Universitv of Idaho) using published kevs, Cymindis planipcnnis LeConte: PF; Jun- e.g. Arnett (1963) and Hatch (1953-1973), and Aug; U. the University of Idaho insect collection as Harpalus amptitatu.s Si\\: PF; May; U. reference material. Following each species Harpalus basilaris Kirby: PF; Jun; U. are the collecting methods, dates of occur- Harpalus fraternus LeConte: LT, PF; Jun, rence, and abundance. Biological information Jul; U. is presented for some species. Voucher speci- Lebia vittata (Fabricius): WP; May-Jul; U. mens are housed in the William F. Barr Ento- Microlestes nigrinus (Mannerheim): PF; Jun, mological Museum at the University of Idaho. Jul; U. Philophuga viridis Dejean: PF, PE; May-Jul; U. Gollecting Methods Larva obserxed feeding on larva of the BF = Berlese Funnel; GC ^ General Collect- sagebrush defoliator, Aroga websteri Clark, ing; LT = Light Trap; MT = Malaise Trap; (Lepidoptera: Gelechidae), on A. tridentata. April 1986 Stafford ET AL. : CoLEOPTERA OF Idaho 289

Piosoma setosa LeConte: PF, WP; Jun, Jul; Serica barri Dawson: LT, PF; Jul; A. U. New state record. Trox sp.: LT; Jun; U. Pseudomorpha behrensi Horn: PF; Jul, U. Leiodidae Dytiscidae Hydnobius sp.: LT, PF; Jun, Jul; U. Laccophilus decipiens LeConte: LT; Aug; U. Ptomophagus californicus (LeConte): PF; Jul; U. Hydrophilidae Silphidae Berosus fraternus LeConte: LT; Jul; U. Nicrophorus hecate Bland; PF, WP; May- Berosus stylifertis Horn: LT; Jul, Aug; U. Aug; C. Cercyon quisquiliiis (Linnaeus): LT; Jul; U. Heloplionis sp.: LT; Jul, Aug; U. Staphylinidae Sphaehdium scarabaeoides (Linnaeus): PF; Anotylus sp.: LT; Aug; U. Jun-Aug; U. Bledius strenuus Casey: LT; Jul; U. Histeridae Philonthus concinnus (Grcivenhorst): PF; May; U. Psiloscelis corrosa Casey: WP; U. New Jun; Philonthus cruentatus (Gmelin): PF; May; U. state record. Platystethus americanus (Erichson): LT; Aug; Saprinus lugens Erichson: GC; May; U. U. Saprinus oregonensis LeConte: PF; Jun, Jul; Tachinus angustatus Horn: WP; Jun; U. U. Xerosaprinus acilinea (Marseul): PF; Jun, Jul; Pselaphidae U. Pilopius sp.: GC; May; U. Xerosaprinus lubricus (LeConte): PF; Jun- Aug; C. Heteroceridae Lanternarius brunneus (Melsheimer): LT; Scarabaeidae Jun; U. Aphodius denticidatus Haldeman: LT, PF; Buprestidae May-Jul; U. Aphodius distinctus Miiller: PF; May; U. Acmaeodera itnmaculata Horn: GC, PE; Jul;

Ap/iot/ius/ossor (Linnaeus): GC; Jul; U. C. Larvae bore roots of C. lanata , adults on Aphodius hirsutus Brown: PF; May; U. flowers oiOpuntia polyacantha Haw. Aphodius granarius (Linnaeus): PF; Jun; U. Agrilus politus (Say): GC, PE; Jul-Aug; U. Aphodius vittatus Say: PF; Jul; U. Larvae bore Salix spp. Aphodius sp. (poss. militaris LeConte): PF, Agrilus pubifrons Fisher: GC, PE, SW; Jul, WP; May; U. Aug; C. Larvae bore roots and adults feed in Boreocanthon simplex (LeConte): PF; May; the flowers of C. viscidifloris. U. Agrilus walsinghami Crotch: GC, PE; Aug; C remastocheihis crinitus bifoviatus Van U. Larvae bore roots of C. nauseosus. Dyke:PF, GC; Jun, Jul; U. Anthaxia retifer LeConte: GC, PE; May; U. Dichelonyx truncata LeConte: GC; Jun, Jul; Adults on flowers of Balsamorhiza sagittata C. (Pursh) Nutt. Diplotaxis haydenii LeConte: GC, LT, WP; Chrysobothris deleta LeConte: GC, PE; Aug; Jun, Jul; U. U. Larvae bore roots of A. tridentata. Diplotaxis obscura LeConte: PF; May-Aug; Chrysobothris horyiingi Barr: GC, PE; May; C. U. Larvae bore roots o{ Eriogonum spp. Diplotaxis subangulata LeConte: LT, WP; Chrysobothris idahoensis Barr: GC, PE; Jun- Jul; U. Aug; U. Larvae bore roots of Eriogonum spp. Glaresis canadensis Brown: PF; Jun-Aug; C. Chrysobothris texana LeConte: GC, PE; Ochodaeus simplex LeConte: LT, WP; Jul, U. Jun-Aug; C. Larvae bore branches of/, os- Paracotalpa granicollis (Haldeman): GC; Jul; teosperma. U. Nanntdaria brunneata (Knull): GC, PE; Jul; U. Serica anthracina LeConte: GC, PF; Jun; U. Larvae bore roots of Eriogonum sp. 290 Great Basin Naturalist Vol. 46, No. 2

Elateridae Nitidulidae

Ainpedus ursinus (Van Dyke): PF; Jun; U. Brachypterolous pulicahus (Linnaeus): WP; Anchastus cinereipennis (Ecshscholtz): PF; Juh'u. May-Jul; C. Adults associated with E. Carpophilus pallipennis (Say): GC, PE, WP; cinereus. Jun-Aug; C. Adults feed on pollen in flow- Cardiophorus sp. 1: BF, PF; Mar, May, Jun; ers of O. polyacantha. U. Phalacridae Cardiophorus sp. 2: PF; May, Jun; U. Cardiophorus sp. 3: GC, PF; May, Jun; U. Olibrus rufipes LeConte: GC, PE; Aug; C. Cardiophorus sp. 4: WP; Jun; U. Adults feed on pollen in flowers of C. visci- Cardiophorus sp. 5: BF, PF, WP; Mar, Mav; difloris. U. Phalacrus penicillatus Say: GC, PE, SW; U. Ctenicera noxia (Hyslop); PF; May, Jun; U. Adults associated with E. cinereus. Ctenicera semivittata (Say): GC; May; U. Coccinellidae Hypolithus bicolor (Eschschohz): PF; Jul; U. Brachyacantha dentipcs socialis Casev: GC; Cantharidae Aug;U. Brachyacantha ursina utcella Casev: GC, Malthodes sp.: GC; May, U. SW;Jun, Jul;U. Brumus septcntrionus Weise: GC; Jun-Aug; Dermestidae U.

Dermestes marmoratus Say: GC, LT; Jul, Coccinella difficilis Crotch: WP; May, Jun; U. Aug; C. Coccinella novemnotata degener Casey: SW; Aug; U. Cleridae Coccinella prolongata Crotch: WP; Jun; U. Coccinella transversoguttata richardsoni Enoclerus acerhus Wolcott: GC, PF, WP; Brown: GC, LT; May-Aug; U. May-Jul; C. Hippodamia apicalis Casey: GC, SW, WP Enoclerus barri Knull: GC; Jul, Aug; U. Jun-Aug; A. PhyUobaenus sp.: GC, PE, SW, WP; Jun- Hippodamia convergens Guerin: SW Aug; C. Adults often found on A. tridentata May-Aug; U. preying on small Lepidoptera larvae. Hippodamia glacialis lecontci Mulsant: WP Trichodes ornatus Say: WP; Jul; U. Aug; U. Hippodamia quinquesignata Kirbv: GC; May Melyridae U. Amecoceris spp.: BF, PF, WP; Mav-Jul; A. Hippodamia tredecimpunctata tibialis (Say) Attahis glab reikis Fall: GC, WP; May-Jul; C. LT; Aug; A. Attalus morulus smithi Hopping: GC, SW; Hyperaspidius hercules Belicek: PF, WP May, Jun; U. Adults associated with E. Jun-Aug; U. cinereus. Hyperaspidius vittigera LeConte: WP; Aug Attalus oregonensis Horn: GC, PE, PF; Jun, U. Jul; C. Adults feed on pollen in flowers of O. Hyperaspis lateralis montanica Casev: MT, polyacantha. SW, WP; May-Aug; U. Collops bipunctatus (Say): PE, SW; Jul; U. Nephus sordidus (Horn): PF; Jun; U. Adults feed on pollen of £. cinereus. Scymnus caurinus Horn: SW; Jun; U. Collops bridgeri Tanner: GC, PF; Mav-Jul; Scymnus marginicollis Maniierheini: PF, U. WP; Jun-Aug; U. Collops hirtellus LeConte: SW; Jun; U. Scynmus })ostj)ictus (]asey: WP; Jul; U. Collops punctidatus LeConte: PF, WP; U. Lathridiidae Dasytellus sp.: PF, WP; Jul; C. Hoppingiana nitida Hatch: WP; Jul; U. Melanoptlialma americana Mannerheim: BF; Trichochrous spp.: GC, PE, WP; C. Adults May; U. feed on pollen in flowers of O. polyacantha. Melanopthalma sp.: SW; Jul; U. . . .

April 1986 Stafford ETAL.:C()LEOPTERA OF Idaho 291

Tenebrionidae Anthicidae

Alaudes sin^iilaris Horn: BF; Mar; U. Anthicus cervinus LaFerte: LT; Aug; U. Araeoschizus airmeti Tanner: GC, PF; May- Anthicus fonnicarius LaFerte: PF; Jul; U. Aug; U. Inquiline in the nests of the harvester Anthicus hastatus Casey: LT; Aug; U. ant, Pogotuwiyrmex owhyheei Cole Notoxns serratiis LeConte: GC, PE, PF, WP; Blapstimishani Boddy: PF, WP; May, Jun; U. May-Jul; C. Adults feed on pollen and de- Blapstiniis substnatiis Champion: PF, WP; veloping seeds of £. cinereus. May-Jul; C. Cerambycidae Coclocnemis punctatus LeConte: PF; Aug; U. Coniontisobesa LeConte: PF; May, Jun; U. Centrodera nevadica nevodico LeConte: LT, Coniontissetosa Casey: BF, PF; Mar, Mav, Jun; PF, WP; Jul, Aug; C. U. Cortodera barri Linsley & Chemsak: GC, Eleodes cordata Eschscholtz: PF; May-Aug; U. MT, PE, PF, WP; Jun;'u. Adults associated Eleodes extricato cognata Haldemann: GC, with E. cinereus PF; May-Aug; A. Crossidius (iter LeConte: GC, PE; Aug, Sep; Eleodes hispilahhs connexa LeConte: PF; U. Larvae bore roots of A. tridentata. May-Jul; U. Crossidius coralinus LeConte: GC, PE; Aug; Eleodes nigrina LeConte: GC; Jul, Aug; U. U. Larvae bore roots and adults feed on Eleodes ohsciira Say: PF; May-Aug; U. pollen of C. nauseosus. Eleodes pilosa Horn: GC, PF; May-Aug; A. Crossidius hirtipes allgeicahri LeConte: GC, Emhaphion elongatum Horn: PF; Mav-Jul; PE; Aug; U. Larvae bore roots and adults U. feed on pollen of C. viscidifloris Helops californicus Mannerheim: PF; Mav, C rossidius punctatus LeConte: GC, PE; Aug, Jun; U. Sep; U. Larvae bore roots and adults feed Helops convexiilus LeConte: BF; Mar; U. on pollen of C. nauseosus Helops opacits LeConte: PF; May-Jul; U. Judolia gaurotoides Casey: GC, PF; May; U. Melanastus ate r (LeConte): BF, PF, Mar, Jul; Mecas bicallosa Martin: PE; Jun; U. Larvae U. bore roots of sagebrush. Oxygonodera hispidiila (Horn): PF; Mav- Megacheuma brevipennis (LeConte): GC, Aug; C. PE; Aug; C. Larvae bore roots ofA. confer- Sphaerkmtis muricata (Casey): PF; Jun; U. tifolia, A. nuttali, and Sacrobatus vermicu- latus (Hook.) Torr. Alleculidae Megasemum asperum (LeConte): LT; Jul; U. Mycetochara procera Casey: GC, WP; May, Prionus californicus Motschulsky: GC; Aug; U. Jun;U. Bruchidae Oedemeridae Acanthoscelides pauperculus (LeConte): GC, Oxacis hicolor (LeConte): WP; Jul; U. WP;Jun-Aug;U.

Mordellidae Chrysomelidae

Mordellistena idahoensis Ray: PF; Jun; U. Altica sp. : GC; Jun; U. Adults associated with Mordellistena sericans Fall; MT, WP; Jun, E. cinereus. Jul; U. Anisostena californica Van Dyke: GC; Aug; LI. Adults associated with E. cinereus. Meloidae Brachycoryna montana (Horn): GC; Jul; U. Epicauta immerita Walker: WP; Jun; U. Chaetocnenia sp.: PF; Jun, Jul; U. Epicauta piceiventris Maydell: WP; Jul; U. Crepidodera nana (Say): GC; Jun; U. Gnathium eremicola Macswain: GC, PE; Cryptocephalus spurcus LeConte: GC, SW; Aug; A. Adults feed on neetar in flowers of Jul; U. Adults and larvae feed on foliage of C. viscidifloris. C. viscidifloris. Lytta viilnerata cooperi LeConte: GC, PE; Disonycha latifrons Schaeffer: PF, SW; May- Aug; U. Aug; C. Larvae and adults feed on foliage of Nemognatha lutea LeConte: GC; Jun; U. C. viscidifloris. . . —

292 Great Basin Naturalist Vol. 46, No. 2

Exenia conspersa (Mannerheim): SW; Jul, Mynnex vittatus (Horn): GG; Jul, aug; U. Aug; U. Ophryastes latirostris LeGonte: GG, PE; Jun; Glyptoscelis sp. (poss. arteniisiae Blake): PF, U. Larvae and adults feed on foliage of C. SW, WP; May, Jun; U. lanata. Monoxia consputa LeConte: PF, SW, WP; Sitona hispidula (Fabricius): LT; Aug; U.

May, Jun; U. Smicronyx sp. : PF; SW; Jun-Aug; U. Monoxia pallida Blake: WP; Jul; U. Sphenophorus gentilis LeGonte: PE, PF, Monoxia puberula Blake: GC; Jul; U. SW; May-Jul; A. Adults feed on developing Pachybrachys caelatus LeGonte; PE, SW; seed heads of£. cinereus Jun-Aug; G. Feeds on foliage of C. viscidi- Trachyphloeni sp. (undescribed genus): PF; floris. Jul, Aug; U. Pachybrachys jacobyi Bowditch: GG; Jun- Tychius tectus LeGonte: PF; June; U. Aug; U. Phyllotreta albionica LeGonte: PF, SW; Acknowledgments May-Aug; G. Adults commonly found on wild mustards, Descurania spp. and Appreciation is extended to the following Schoenocrambe linifolia (Nutt.) Greene. persons for the identification of specimens: R. Pyrrhalta luteola (Miiller): GG; Jul; U. S. Anderson, University of Alberta (Gur- Pyrrhalta nymphaeae (Linnaeus): GG; Jun; culionidae: C/eonws); A. F. Newton, Museum U. ofGomparitive Zoology— Harvard University Saxinis saucia LeGonte: GG; Jul; U. (Staphylinidae); R. L. Penrose, Galifornia De- Scelolyperus nigrovirescens (Fall): GG; Jun; partment of Agriculture (Gerambycidae: Cen- U. trodera); G. A. Triplehorn, Ohio State Uni- Stenopodius vandiizeei Blaisdell: GG; Jul; U. versity (Tenebrionidae: Coniontis); R. L. Systena blanda (Melsheimer): GG; Jul; U. Wenzel, Field Museum of Natural History Trirhabda nitidicollis LeGonte: GG, PE, Jul; Ghicago (Histeridae); and D. R. Whitehead, G. Adults and larvae feed on C. nauseosus. Systematics Entomology Laboratory, USDA (Gurculionidae: Trachyphloenini). We thank Anthribidae O. D. Markham, U.S. Department of En- Trigonorhiniis sp. (near annulatus Garr): WP; ergy, and Frank Merickel, University of Jun, Jul; U. Idaho, for their help and support. This work Gurculionidae was conducted for the IN EL Radioecology and Ecology Programs sponsored by the Of- Acmaegenius granicollis Van Dyke: GG; Jun; fice of Health and Environmental Research, U. United States Department of Energy. Anthonomtissp. I: GG, PE, SW; Jun-Aug; U. Adults feed on foliage of C. viscidifloris. Anthonomus sp. 2: GG, PE, SW; Jun-Aug; G. Literature Gited Adults feed on foliage of C. viscidifloris Allred. D M 1968. Ticks of the National Reactor Test- Apion sordidum Smith: PE, PF, SW, WP; ing Station. Brigham Young Univ. Sci. Bull. Biol. May-Jul; U. Inquiline in tephritid and ce- Ser., 10(1): 1-29. cidomyiid galls on A. tridentata. 1969. Spiders of the National Reactor Testing Sta- tion. Great Basin Nat. 29: 105-108. Brachyrhinus ovatus (Linnaeus): GG; Jun- 1973. Scorpions of the National Reactor Testing Aug; U. Station. Great Basin Nat. 33: 251-254.

Cleonus kirbyi Gasey: PF; May, Jun; U. Allred. D M , and A C Cole. 1971. Ants of the Na- Cleonus quadrilineatus (Chevrolcii): PF; May, tional Reactor Testing Station. Great Basin Nat. 31:237-242. Jun; U. Allred, D. M., and M M Muma. 1971. Solpugicls of the Cosmobaris americana Gasey: MT; U. Jun; National Reactor Testing Station, Idaho. Great Dinocleus denticollis Gasey: PF; May, Jun; U. Basin Nat. 31: 164-168. Arneti-. Dyslobus alternatus Horn: PF; May-Aug; G. R H , JR 1963. The beetles of the United States. Catholic University Press, Wa.shington, D.C. Gyrotus sinuatus Hatch: PE; Jul; U. Larvae 1,112 pp. bore in Arte7nisia stems. Harni.ss.R () ,andN E. West 1973. Vegetative patterns Miloderoides maculatus VanDyke: PF; Jun, of the National Reactor Testing Station, South- Jul; U. eastern Idaho. Northw. Sci. 47: 30-43. 293 April 1986 Stafford etal.:Coleoptera of Idaho

Kingsolver. M., ed. 1979. A catalog of the Coleoptera Hatch, M. H. 1953-1973. The beetles of the Pacific North- J. west, Parts I-V. University of Washington Press. of America north of Mexico. USDA Agric. Hand- K. HOLTE 1978. Flora of the Idaho book 529. JEPPSON, R. J., AND E National Engineering Laboratory Sites^ Pages y^^^ r N R. FRENCH, A. H. Dahl, AND E, Det- studies on J. 129-143 in O. D. Markhamed., Ecological ,- ^ ^- ^ j _f MER. 1978. Vegetation types and surface soilsifof 1 . 1 1 Ki . 1 r- LaboratoryI k t^ ,cit^iQ78Site 1978 the Idaho National Engineering .^ • , ,, , x, . i i u . c-^ the Idaho National Engineering Laboratory Site. Progress Report IDO-12087. U.S. Department of IDO-12084. National Technical Information Ser- Energy. National Technical Information Service, Virginia. Sprin^ield, Virginia, 371 pp. vice, Springfield, 29 pp.