Great Basin Naturalist

Volume 51 | Number 4 Article 10

12-27-1991 Oxytropis DC.—names, basionyms, types, and synonyms—Flora North America Project Stanley L. Welsh Brigham Young University

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Recommended Citation Welsh, Stanley L. (1991) "Oxytropis DC.—names, basionyms, types, and synonyms—Flora North America Project," Great Basin Naturalist: Vol. 51 : No. 4 , Article 10. Available at: https://scholarsarchive.byu.edu/gbn/vol51/iss4/10

This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. Great Basin Naturalist 51(4), 1991, pp. 377-396

OXYTROPIS DC.~NAMES, BASION)'MS, TYPES, AND SYNONYMS­ FLORA NORTH AMERICA PROJECT

Stanley L. Welshl

ABSTRACT.- All names known to apply to the genus Oxytropis de Cahdolle in NorthAmerica are listed with place of publication, type information (where known), type specimen repositories, and notations pertinent to understanding the nomenclatural aspects ofthe genus. Nomenclatural combinations proposed are: Oxytropis arctica R. Brown var. murrayi Gurtzev) Welsh and O. campestris (L.) de Cahdolle var. roaldii (Ostenfeld) Welsh.

keywords: Oxytropis, North America, names, types, synonyms.

Preparation of a revisionary summary of the'inclusion would have further burdened the genus Oxytropis de Candolle for the an already hug~ genus and overlooked the Flora North America Project necessitates that divergence ofthe oxytropes from most ofthe nomenclatural changes and type inf.ormation astragah:ts complljlxes. In North America only be presented prior to publication in that pro~ the introduced O. Tiparia and the indigenous ject. The following list consists of synonyms, o. deflexa are caulescent or have caulescent names, nomenclatural types, and new combi­ phases. The porrect beak ofthe keel is diag­ nations of names involved with this interest~ nostic for the genus, even though some spe~ ing and complex genus as it occurs in North cies ofAstragalus have extended keel apices; America. Each name involved with the genus none are truly porrect. is listed with its bibliographical citation, type Aside from the problern of nomenclatural information, places of deposit of the types, transfers, which have added to the list of syn~ and other pertinent information as necessary. onyms, the main difficulties are morphologi­ All names cited in the literature are in­ cal; there are few COnsistent mOrPhological cluded. In some few of them the place of features to serve as taxonomic criteria. The deposition of the type is unlalOwn, and the pods, with some exceptions, are mainiy alike, space for that information is left blank. especially in those taxa that are most alike Oxytropis consists of some 57 taxa in 22 spe­ otherwise. The flowers ?Ie similar through cies in its North American complement. The the genus, the main differences being in size taxonomic problems are disproportionate (and that varies greatly within SOme species) with the size ofthe genus, however. The large and color (which also varies within a species number ofsynonyms reflects the Problematic and often within the infraspecmc categories). nature of the taxa within the genus. Further The taxonomist has had to rely on features of cornplicating the number ofnames have been stipules, leaves, inflorescence, pubescence, the nomenclatural transfers to earlier pub­ and calyx Or pod inflation to arrive at a treah lished genera Aragallus Neck. and Spiesia ment that still lacks absolute consistency. Neck. The conservation of Oxytropis DC. Morphological intermediates, resulting from forestalled ultimate adoption ofeither ofthose hybridization or from overlap ofWidely vary­ generic names. Some authors, Tidestrorn for ing characteristics, tend to cloud the picture. example, discerned the close relationship of Chromosome number is helpful, to an e~tent, Oxytropis with Astragalus L., and he sub~ but plants with differing ploidy levels are sequently made wholesale transfers of the known within species and some of the vari­ names to that genus. While having consider~ eties; and chromosome number is not always able merit frorn a phylogenetic standpoint, associated with other morphological features.

lLif~ Science Museum and Department ofBotany and Range Science, Brigham Young University, Provo, Utah 8460~.

377 378 GREATBASIN NATURALIST [Volume 51

Some workers have looked at the genus as = Oxytropis eampestris (L.) de Candolle var. gracilis lithe taxa were clear and distinctive, resulting (A. Nelson) Barneby in the proliferation ofspecific and infraspecific Aragallus albiflorusA. Nelson, Erythea1: 62. lS99, nom. names. Each new morphological variant was nov. Basionym: Oxytropis lambertii Pursh var. ochroleuea stu~ considered worthy of a name by some A. Nelson dents ofthe genus. Adding to the difficulties = Oxytropis serieea Nuttall var. serieea nomenclat~ ofinterpretation oftaxonomic and var. eondensatus A. Nelson, Erythea 7: 62. lS99. ural problems was the circumboreal nature of Type: Wyoming, Bitter Creek, Red Desert, SWeet- the genus. In Alaska and other northern re­ water County, A. Nelson 4773, 16 June lS9S; gions of North America, some species were holotype RM!; isotypes NY!, US! = Oxytropis serieea Nuttall var. serieea clearly allied closely to Siberian or Eurasian taxa. A paucity of critical materials from Aragqllus alpicola Rydherg, Mem. N. Y. Bot. Gard. 1: 252,1900. Siberia for comparison with the American Type: Montana, Old Hoilowtop near Pony [South counterparts made interpretation difficult. Boulder Range, Madison County], P. A. Rydberg Specimens from Siberia in American herbaria & E. Bessey 4503, 9 July lS97; holotype NY!; are still the exception. In the campestris and isotypes GH!, NDG!, US!, NY! borealis complexes the need for such Com­ = Oxytropis campestris (L.) de Candolle var. eusickii (Greenman) Bameby parisons is critical. Both species were named prior to any of the North American counter­ Aragallus angflstatus Rydberg, Bull. Torrey Bot. Club 34: 422.1907. parts, and itwas not until the critical review of Type: Nebraska, Rush Creek, Duel County, P. A. Oxytropis by Barneby (1952) that clarification Rydberg S2c, 2 July iS91; holotype NY!; isotypes ofmany ofthe problems became a possibility, MIN!,NEBI although the equivalence was discerned ear~ = Oxytropis lambertii Pursh var. lambertii Her by Gray (1884) and others. Much addi­ Aragall'Us arctieus (R. Browil) Greene, Pittonia 3: 211. tional work and refinement ofthe infraspecific lS97. taxa in the complexes with circumboreal rep~ Basionym: Oxytropis aretiea R. Brown = Oxytropis aretiea R. BroWn resentatiOn is necess~. Aragallus argophyllus Rydb~rg, Mem. N. Y. Bot. Gard. The present Writer has examined all but a 1: 255. 1900. few ofthe taxainthefield andhas spentalmost Type: Little Blackfeet River, Montana, J. G. Cooper three decades in their pursuit. Itis hoped that S.n., lS60; holotype NY!; isotype GH! (frag), US! this summary ofnames will be helpful. Works = Oxytropis besseyi (Rydberg) Blankinship var. argo­ of special importance to this paper include phylla (Rydberg) Barneby those by Barneby (1952), Boivin (1962, 1967), Aragallus articulatus Greene, Proc. BioI. Soc. Wash. IS: Bunge (1874), Elisens and Packer (1980, 13.l905. Type: Between FortSmith and the Rio Grande [Okla­ 1982), Gray (1884), Jurtsev (1986), PorsUd and homaorwestem Texas], J. M. Bigelows.n., lS53; Cody (1980), Vasil'chenko et al. (1948), and holotype US! Welsh (1967, 1974, 1977, and 1987). = Oxytropis lambertii Pursh var. lambertii Aragallus atfopurpureus Rydberg, Bull. Torrey. Bot. Aragallus abbreviatus GrlleI!e, Proc. BioI. Soc. Wash. Club 34: 424. 1907. IS: 12. 1905. Type: Headwaters of the Tongue River, Big Hom Type: Texas, near Dallas, Limestone prairie, Dallas Mountains, Wyoming; F. Tweedy 125,126, lS9S; County, J. Reverchon 603, May lS76; holotype cotypesNY! NDG!; isotype NY!; dry calcareous soil near = Oxytropis lagopus Nuttall var. atropurpureus (Ry­ Dallas, Texas, Curtis 603, April, May; cotype dberg) Barneby GH!, NDG!, NY! = Oxytropis lambertii Pursh var. articulata (Greene) Aragall'Us aven-rl6lsonii Ltinell, Bull. Leeds Herb. 2: 6. Bameby 1905. Type: NorthDakota, Butte, Benson County, J. Lunell AragaZlus aboriginum Greene, Proc. BioI. Soc. Wash. IS: s.n., 14, 21 June, 2 July 1905; isotypes Ny!, US!, 12.1905. wrC, MIN!, NDA! Type: Oklahoma, Cimarron Valley, Cherokee Outlet, Oxytropis lambertii Pursh var. lambertii M. N. Carleton 217, June lS91; holotype US!, = photoBRY! Arqgilllus beUii (Britton) Greene, Pittonia3: 212. lS91. := Oxytropis serieea Nuttall var. serieea Basionym: Spiesia Oxytropis bellii Britton Ar.(lgaUus albertinus Greene, Proc. BioI. Soc. Wash. IS: = Oxytropis aretica R. Brown var. bellii (Britton) Boivin 15: 1905. Type: Saskatchewan, Prince Albert, Lat. 53°, J. M. Aragallus besseyi Rydberg, Mem. N. Y. Bot. Gard. 1: Macoun 12535 and 12540, July 1896; cotypes NDG! 250.1900. 1991] OXYTROPIS DC.-NAMES, BASIONYMS, 'I'¥PES, SYNONYMS 379

Type: Montana, Spanish Basin, Gallatin County, Ar4gallusfoliolosus (:aooket) Macoun, Ottawa Naturalist 6,500 ft., P. A. Rydberg & E. A. Bessey 4501, 23 13: 163. 1899. June 1897; holotype NY!; isotypes Ga:!, NpG!, Basionym: Oxytropisfoliolosa Hooker US! = Oxytropis deflexa var. foliolosa (Hooker) Barneby Oxytropis besseyi (Rydberg) Blankinship var. Aragalllis fotmosus Greene, Proc. BioI. Soc. Wash. 18: besseyi - 13.1905. AragaUus bigelovii (A. Gray) GreeQe, Pittonia 3: 212. Type: South Dakota, Fort Meade, Meade County, 1897. W. H. Forwood 95, 7 June 1887; holotype US!, Basionym: Oxytropis lambertii Pursh Var. bigelovii photoBRY! ' A. Gray . = Oxytropis lambertii Pursh vai. lambertii = Oxytropis lambertii Pursh var. lambertii AtagaUUs galioides Gre~ne, Proc. BioI. Soc. Wash. 18: MagallUs blankinshipii A. Nelson, Erythea 7: 58. 1899. 16.1905. Type: Montana, dry rocky hillsides along Middie type: Alberta, Bow River near Banff, McCalla s.n., Creek, 15 mi SW of Bozeman, Gallatin County, 10 July, 18 September 1899; holotype US! J. W. Blankinship s.n., 4 July 1898; holotype RM!; = Oxytropis splendens Douglas isotypes GH!, NY! = Oxytropis lagopus NuttBritish Columbia, Deer Park, Lower Arrow 4rag4llus inflatus (:aooket) A. Nelson, Erythea 7: 59. 1899. Lake, J. M. Macoun 5358, 8 JUlle 1890; holotype NDG! Basionym: Oxytropis arctica 8 inflata Hooker = Oxytropis campestris (1,.) de Candolle var. gracilis = Oxytropis podocarpa A. Gray (A. Nelson) Barneby 4ragaUus invenustus Greene, Proc. BioI. Soc., Wash. 18: AragaUus collinus A. Nelson, Erythea 7: 57. 1899. 12.1905. Type: Wyoming, Seminoe Mts., Carbon County, E. Type: South Dakota, about Fort Meade, Meade Nelson 4925, 21 July 1898; holotype RM!; isotypes County, W. H. Forwood 963" 96b, 3 June :(.887, GH!, NY!, US! 96b, 7 June 1887; cotypes US!, photo BRY! = Oxytropis nana Nutt. = Oxytropis sericea Nuttall var. sericea MagaUus dejlexus (Pallas) A. A. Hellet, Cat. N. Amer. AtagaUus involutus A. Nelson, Erythea 78: 64. 1899. PI., 4. 1898. Type: MiIlnesota, 1\cton, Meeker County, W. D. Basionym: Astragalus deflexus Pallas Frost s.n., June 1892; holotype EM!; isotypes = Oxytropis deflexa (Pallas) de Candolle MIN!, MO!, US! = Oxytropis lambertii Pursh var. lambertii AragaUus dispar 1\. Nelson, Erythea 7: 61. 1899. Type: North Dakota, Dickinson, Stark County, Mrs. Aragallus johannensis (Fern.) A. Hellet, Cat. N. Amer. Cook s.n., 1896; holotype RM!; photo BRY! PI., ed2: 7.1900. = Oxytropis campestris (1..) de Candolle var. dispar Basionym: Oxytropis campestris (L.) de Candolle var. (A. Nelson) Barneby johannensis Fernald AragaUusfalcutUs Greene, Proc. BioI. Soc. Wash. 18: 13. = Oxytropis campestris (L.) de Candolle var. johan­ 1905. nensis Fernald Type: Missouri" Watson, Atchison County, B. F. AragaUus knowltonii Greene, Proc. BioI. Soc. Wash. 18: Bush 204, 1 June 1894; holotype NPG!; isotypes 12.1905. GH!, NY!, US!, ISC!, MIN!, MO!, DAO! (Note: Type: Arizona, San Francisco Mountains, Coconino The same collection is also the type ofOxytropis County, F. H. Knowlton 44, 20 August 1889; holo_ bushii Gandoger.) type US! = Oxytropis lambertii Pursh var. lambertii = Oxytropis lambertii Putsh va,r. bigelovii A. Gray 380 GREAT BASIN NATURALIST [Volume5l

AragaUus lagopus (Nuttall) Greene, Pittonia 3: 212. Aragallus multiceps (Torrey &: Gray) A. Heller, Cat. N. 1897. Amer. PI., 4.1898. Basionym: Oxytropis lagopus Nuttall Basionym: Oxytropis multiceps Torrey &: Gray Oxytropis multiceps Torrey &: Gray Aragallus lambettii (Pursh) Greene, Pittonia 3: 212. = 1897. var. minor (A. Gray) A. Nelson, Erythea 7: 51. 1899. Basionym: Oxytropis lambertii Pursh Basionym: Oxytropis multiceps var. minorA. Gray = Oxytropis multiceps Torrey &: Gray var. sericeus (Nuttall) A, Gray, Proc. Amer. Acad. 20: 7. i884. Aragllllus nanus (Nuttall) Greene, Pittonia3: 212. 1897. Basionym: Oxytropis sericea Nuttall Basionym: Oxytropis nana Nuttall Aragallus luteolils Greene, Proc. BioI. Soc. Wash, 18: 17. = Oxytropis nana Nuttall 1905. Aragallus oreophilus (A. Gray) A. Nelson, Erythea 7: 59. Type: Washington, Olympic Mts., Clallam County, 1899. A. D. Elmer 2532, July 1900; holotype USI; iso o Basionym: Oxytropis oreophila A. Gray types NE, NY'!, NDG!, MINl, MOl, wtc, CAS!, = Oxytropis oreophila A. Gray DS! Aragallus parryi (A. Gray) Greene, Pittonia3: 211. 1897. = Oxytropis campestris (L.) de Candolle var. gracilis Basionym: Oxytropis parryi A. Gray (A. Nelson) Bameby = Oxytropis parryi A. Gray Aragallus majusculus Greene, Proc. BioI. Soc. Wash. Aragqllus patens Rydberg, Bull. Torrey Bot. Club 34: 18:12.1905. . 421. 1907.' Type: Utah, Henry Mts., Garfield County, Utah, Type: Colorado, plains and foothills near Boulder, M. E. Jones 5674, July1894; holotype US!; isotype Boulder County, F. Tweedy 5164, 1902; holotype NY!, MO!, photo BRY! NYl = Oxytropis sericea Nuttall vat. sericea = Oxytropis lambertii Pursh var. bigelovii A. Gray Aragallus macounii Greene, Proc. BioI. Soc. Wash. 18: Aragallus pinetorum A. Heller, Bull. Tortey Bot. Club 16.1905. 26: 548. 1899. Type: Alberta, Elbow River, Rocky Mountains, Lat. Type: New Mexico, 11 mi SE of Santa Fe, Santa Fe 49°40', J. M. Macoun 18517, June-July 1897; County, A. A. &: E. G. Heller 3751, 23 June 1897; holotype NDG! holotype USI (?); isotypes MO!, lA!, MUI, GHl, = Oxytropis campestris (L.) de Candolle vat. gracilis NDG!, NY!, POM, WTC, BRY!, DS! (A. Nelson) Barneby = Oxytropis sericea Nuttall var. sericea Two specimens, J. M. Macoun 18516 and 18517, were cited as type of A,ragallus macounii Greene. vllr. veganus Cockerell, Torreya 2: 155. 1902. Batneby(1952) designated 18517 as the type ofthe Type: New Mexico, top ofLasVegas Range, on end of taxon; the other specimen belongs to O. sericea ridge, T. D. A. Cockereil 20, end of June 1901; Nuttall var. spicata (Hooker) Barneby. holotype NY!; isotype GH! = Oxytropis sericea Nuttall var. sericea Aragallus melanodontils Greene, Proc. BioI. Soc. Wash. 18: 15. 1905. Aragalltls plattensis Nuttall ex Torrey &: Gray, FI. N. . Type: Alberta, Elbow River, Rocky Mountains, Lat. Amer. 1:340. 1838 49°40', J. M. Macoun 18513, June-July 1897; holo­ Type: ? typeNDGl = Oxytropis lambertii Pursh var. lambertii Oxytropis sericea Nuttall vat. spicata (Hooker) Aragallus podocarpus (A. Gray) A. Nelson, Coulter &: Barneby Nelson, NewMan., 294.1909. Aragallus merlensianils (Ttirczaninow) Greene, Pittonia Basionym: Oxytropis podocarpa A. Gray 3: 211. 1897. = Oxytropis podocarpa A. Gray Basionym: Oxytropis mertensiana Turczaninow Aragallus richardsonii (Hooker) Greene, Pittonia 4: 69. = Oxytropis mertensiana Turczaninow 1899. Aragallus 7Tietccilfei Greene, Proc. BioI. Soc. Wash. 18: Basionym: Oxytropis splendens ~ richardsonii 12.1905. Hooker Type: New Mexico, Sawyer's Peak, Grant County, = Oxytropis splendens Douglas open glade, ca 10,000 ft., O. B. Metcalf 1079, Aragallus rigens Greene, Proc. BioI. Soc. Wash. 18: 14. 7 July 1904; holotype US!; isotypes NY!, CAS!, 1905. GHI, POM, WTC. Type: Montana, Cedar Creek, 12 mi above Glendive, = Oxytropis lambertii Pursh vat. bigelovii A. Gray Dawson County, L. F. Watd s.n., 15 July 1884; Aragallils minor (A. GraY) Cockerell.ex Daniels, Univ. holotype US!, photo BRYI MissoUri Sci. Stud. II, 2: 158. 1911. = Oxytropis lambertii Pursh var. lambertii Basionym: Oxytropis multiceps vat. minorA. Gray Atagallus saximontanus A. Nelson, Erythea 7: 190. 1899, .. = Oxytropis multiceps Torrey &: Gray nom. nov. Aragallus monticola (A. Gray) Greene, Pittonia 3: 212. Basionym: Oxytropis lambertii Pursh var. ochroleuca 1897. A. Nels. Basionym: Oxytropis monticola A. Gray = Oxytropis sericea Nuttall var. sericea = Oxytropis campestris (L.) de Candolle vat. gracilis var. condensatus (A. Nelson) A. Nelson, Erythea 7: (A. Nelson) Barneby 190.1900. 1991] OXYTROPIS DC.-NAMES, BASIONYMS, TYPES, SYNONYMS 381

Basionym: Aragallus albiflorus var. condensatus A. Astragalus albiflorus (A. Nelson) Gandoger, Bull. Soc. Nelson Bot. France 48: xiv. 1901. = Oxytropis sericeq Nuttall var. sericea Basionym: Aragallus albiflorus A. Nelson Aragallus seticeus (Nuttall) GreeIle, Pittonia 3: 212. = Oxytropis sericea Nuttall var. sericea 1897. AstragalUs alpicola (Rydberg) Tidestrom, Proc. BioI. Basionym: Oxytropis sericea Nuttall Soc. Wash. 50: 19. 1937. = Oxytropis sericea Nuttall Basionym: Aragallus alpicola Rydberg Aragallus spicatus (Hooker) Rydberg, Mem. N. Y. Bot. = Oxytropis campestris (L.) de Candolle var. cusickii Gard. 1: 251. 1900. (Greenman) Barneby Basionym: Oxytropis campestris 1) spicata Hooker Astragalus arcticus (R. Brown) Sprengel, Syst. 4: 288. = Oxytropis sericea Nuttall var. spicata (Hooker) 1827. Barneby Basionym: Oxytropis arctica R. Brown Aragallus splendens (Douglas) Greene, Pittonia 3: 211. = Oxytropis arctica R. BroWn 1897. Astragalus bellii (Britton) Tidestrom, Proc. BioI. Soc. Basionym: Oxytropis splendens Douglas Wash. 50: 18. 1931. = Oxytropis splendens Douglas Basionym: Spiesia bellii Britton Aragallusvarians Rydberg, Bull. N. Y. Bot. Card. 2: 17'6. = Oxytropis arctica R. Brown var. bellii (Britton) 1901. Boivin Type: Yukon, Lewes River, J. B. Tarleton 33b, 28 Astragalus biflorus Schweinitz ex Gray, Proc. Amer. June 1899; holotype US!; isotypes SI, NYI Acad. 6: 234. 1864, sYIlonym. = Oxytropis campestris (L.) de Candolle var. varians = Oxytropis podocarpa A. Gray (Rydberg) Barneby Astragalus bisontum Tidestrom, Proc. BioI. Soc. Wash. Aragallus veganus (Cockerell) Wooton &: Standley, 50: 18. 1937. Contr. U. S. Nat. Herb. 16: 136. 1913. = Oxytropis multiceps Torrey &: Gray Basionym: Aragallus pinetorum veganus Cockerell = Oxytropis sericea Nuttallvar. sericea var. minor (A. Gray) tidestrom, Proc. BioI. Soc. Wash. 50: 18. 1937. Aragallus ventosus Greene, Proc. BioI. Soc. Wash. 18: Basionym: Oxytropis multiceps vat. minor A. Gray 15.1905. = Oxytropis multiceps Torrey &: Gray . Type: Dry ground in the valley of the North Fork of Wind River, Wyoming, W. H. Forwood 65, Astragalus blankinshipii (A. Nelson) Tidestrom, Proc. BioI. Soc. Wash. 50: 18. 1937. 12 July 1884; holotype PSI; isotype GH! = Oxytropis besseyi (Rydberg) Blankinship var. ven­ Basionym: Aragallus blankinshipiiA. Nelson tosa (Greene) Barneby = Oxytropis lagopus Nuttall Aragallus villosus Rydberg, Bull. Torrey. Bot. Club 28: Astragalus campestris L., Sp. PI., 761. 1753. 36.1901. Type: "in Oelandia, Germania, Helvetia"; holotype Type: Montana, Craig, Lewis and Clark County, LINN 926/511 E. N. Wilcox 3'78, 20 June 1900; holotype US!; = Oxytropis campestris (L.) de Candolle isotypeNY! Astragalus coronaminis (Fernald) Tidestrom, Proc. BioI. = Oxytropis campestris (L.) de Candolle var' gracilis Soc. Wash. 50: 19. 194'7. (A. Nelson) Barneby Basionym: Oxytropis coronaminis (Fernald) Tide­ Aragallus viscidulus Rydberg, Mem. N. Y. Bot. Gard. strom 1: 253. 1900. = Oxytropis arctica R. Brown vat. arctica Type: Montana, Melrose, Silver Bow County, P. A. Astragalus deflexus Pallas, Acta Acad. Sci. Imp. Petrop. Rydberg 2716, 6 July 1895; holotype NY! 2: 268. 1179. = Oxytropis borealis de Candolle var. viscida (Nut­ Type: "ad nivalia Dauriae ... circa Balyra rivum tall) Welsh aliosque Ononem influentibus" [Siberia], P. S. var. depressus Rydberg, Mem. N. Y. Bot. Gatd. Pallas s.n.; holotype BM. 1: 253. 1900. = Oxytropis deflexa (Pallas) de Candolle var. deflexa Type: Haystack Mt., Stillwater County, Montana, var. foliolosus (Hooker) Tidestrom, Proc. BioI. Soc. F. Tweedy 120; holotype NY! Wash. 50: 18. 1937. = Oxytropis borealis de Candolle var. viscida (Nut­ Basionym: Oxytropisfoliolosa Hooker tall) Welsh = Oxytropis deflexa (Pallas) de Candollevat. foliolosa Aragqllus viscidus (Nuttllll) GreeIle, Pittonia 3: 211. (Hooker) Barneby 1897. Astragalus gaspensis (Ferilald &: Kelsey) TidestroIn; Basionym: Oxytropis viscida Nuttall Proc. BioI. Soc. Wash. 50: 19. 1937. = Oxytropis borealis de Candolle var. viscida (Nut~ Basionym: Oxytropis gaspensis Fernald &: Kelsey tall) Welsh = Oxytropis borealis de Candolle vat. viscida (Nut­ Astragalus albertinus (Greene) Tidestrom, Proc. BioI. tall) WelSh Soc. Wash. 50: 19. 1937. Astragalus grayanus Tidestrom, in Tidestrom &: Kittell, Basionym: Aragallus albertinus Greene FI. Ariz. &: New Mex., 216. 1941, nom. nov. = Oxytropis campestris (L.) de Candolle vat. gracilis =Oxytropis campestris (L.) de Candolle var. gracilis (A. Nelson) Barneby (A. Nelson) Bameby 382 GREAT BASIN NATURALIST [Volume 51

Astragalus lqgopus. (N1.ittall) Tidestrom, Proc. BioI. Soc. Astragalus septentrionalis Tidestrom, Proc. BioI. Soc. Wash. 50: 19. 1937. Wash. 50: 19.1931, substitute name. Basionym: Oxytropis lagopus Nuttall = Oxytropis podocarpa A. Gray = Oxytropis lagopus Nuttall Astragalus splendens (Douglas) Tidestrom, Proc. BioI. Astragaluslambenii(Purs!t) Sprengel, Syst. 3: 308. 1826. Soc. Wash. 50: 18. 1937. Basionym: Oxytropis lambertii Pursh Basionym: Oxytropis splendens Douglas = Oxytropis lambertii PiIrsh var. lambertii = Oxytropis splendens Douglas var. bigelovii (A. Gray) Tidestrom, Proc. Bioi. Soc. var. richardsonii (Hooker) Tidesttom, Proc. BioI. Wash. 50: 19. 1937. Soc. Wash. 50:128. 1937. Basionym: Oxytropis lambertii Pursh var. bigelovii Basionym: Oxytropis splendens 13 richardsonii Hooker A. Gray = Oxytropis splendens Douglas = Oxytropis lambertii Pursh var. bigelovii A. Gray Astragalus tomae Tidestrom, Proc. BioI. Soc. Wash. 50: Astragalus mazama (St. Johit) G. N. jones, Univ. Wash. 18. 1937, nom. nov. Pub. Bot. 7: 175. 1938. = Oxytropis nana Nuttall Basionym: Oxytropis mazama St. John. = Oxytropis campestris (L.) de Candolle var. gracilis Astragalus viscidus (Nuttall) Tidestrom, Proc. BioI. Soc. (A. Nelson) Bameby Wash. 50: 19. 1937. AstragalUs munzii Wheeler, Leafl. W. Bot. 2: 209. 1939, Basionym: Oxytropis viscida Nuttall nom. nov. = O. borealis de Candolle var. viscida (Nuttall) Welsh = Oxytropis oreophila A. Gray Oxytropis alaskana A. Nelson, Univ. Wyo. Pub. Bot. 1: Astragalus nigrescens rallas, Astragalogia, 65, tab. 53. 120.1926. 1800. Type: Kachemak Bay, Cook Inlet, M. W. Gorman Type: "inter Aldanum fl[u'm.en] et orientalem Ocean" 1560; holotype RM!; isotype WTU. urn" [between Aldan River and the Sea of = O. campestris (L.) de Candolle var. varians (Ryd" Okhotsk, Siberia], D. D. Merk; type LE. berg) Bameby = Oxytropis nigrescens (PaiIas) de Caluiolle Oxytropis albertlna (Greene) Rydberg, FI. Prair. & PI., var. arctobia (Pallas) Tidestrom, Proc. BioI. Soc. 484.1932. Wash. 50: 19. 1937. Basionym: Aragallus albertinus Greene :i3asionym: Oxytropis arctobia Pallas . = O. campestris (L.) de Candolle var. gracilis = Oxytropis nigrescens var. uniflora (Hooker) (A. Nelson) Barneby Barneby O:tytropis albiflora (A. Nelson) J(. SChumllUn, Just's Bot. Astragalus oreophilus (A. Gray) Tidestrom, Proc. BioI. Jahresb. 27: 496. 1901. Soc. Wash. 50: 19. 1937. Basionym: Aragallus albiflorus A. Nelson, nom. nov. Basionym: Oxytropis oreophila A. Gray Oxytropis lambertii Purshvar. ochroleuca A. Nel­ = Oxytropis oreophila A. Gray son Astragalus parryanus Tidestrom, Proc. BioI. Soc. Wash. = O. sericea Nuttall var. sericea 50: 19. 1941. Oxytropis alpicola (Rydberg) M. E. Jones, Mont. Bot. nom. nov. pro OxytropisparryiA. Gray Notes, 34. 1910. = Oxytropis parryi A. Gray Basionym: Aragallus alpicola Rydberg Astragalus pygmaeus Pallas, Astragalogia, 66, tab. 54. = O. campestris (L.) de Candolle var. cusickii 1800. (Greenman) Barneby Type: ? Oxytropis angustata (Rydberg) A. Nelson, Univ. Wyo. = Oxytropis nigrescens (Pallas) Fischer var. nigres" Pub. Bot. 1: 116. 1926. cens Basionyrn: Aragallus angustatus Rydberg Astragalus retro~s Pallas, Astragalogia, 33, tab. 27. = O. lambertii var. lambertii 1800. Type: ? Oxytropis arctica:J:l.. Brown, Patry's FirstVoy., Append. = Oxytropis deflexa var. deflexa 9: 278. 1824. Type: Canada, Melville Island, Parry's First Voyage, Astragalus rusbyi Green & Motris, J. Amer. Soc. Agron. Sabine, Edwards, Ross, and others, 1819-1820; 27: 546, 549, figs. 1, 2. 1935, provo nom. holotype (?) S!; isotype GHI = Oxytropis riparia Litvinov var. arctica Astragalus rydbergianus Tidestrom, Proc. BioI. Soc. Distribution: Alaska, Yukon, Canadian Arctic Archi­ Wash. 50: 19. 1937, nom. nov. pelago, N.W.T. east to north ofHudson Bay, and = Oxytropis campestfis (L.) de CandoUe var. gracilis less commonly in the interior. (A. Nelson) Barneby The var. atctica is recognizable by its racemes of Astragalus saximontanUs (A. Nelson) Tidestrom, in Tide" mainly fewer than 8 large purpie or lavender strom & Kittell, FI. Ariz. New Mex., 216.1941. flowers. on ·plants mainly less than 15 cm tall Basionym: A.ragallus saximontanus A. Nelson, nOm. and leaflets not or seldom fasciculate. Mainly they nov. Oxytropis lambenii Pursh var. ochroleuca occur in or near coastal Alaska and Canada; less A. Nelson commonly they are montane plants ofthe interior = Oxytropis sericea Nuttall var. sericea or occur in other interior situations. 1991] OXYTROPIS DC.-NAMES, BASIONYMS, TYPES, SYNONYMS 383

var. bamebyana Welsh, Great Basin Nat. 28: 15g, summits of the Rocky Mts., Drummond:' The fig. 4. 1968. plant is almost certainly typical material of vat. Type: Alaska, Kotzebue, 66°55'N, 162°40'W, S.L. inflata. Welsh 5729, 1966; holotype BRY!; isotype NYI, var. koyukukensis (Porsild) Welsh, Iowa State J. Sci. UCI 41: 280. 1967. Distribution: Coastal northwestern Alaska; endemic. Basionym: O. koyukukensis Porsild Flower size, pilose stipules, and calyx features indi­ Distribution: Umiat, Wiseman, Anaktuvuk, Koyu_ cate an alliance to the sympatric O. arctica. The . kuk, Shatolik, and Northway Vicinities, Alaska; racemes vary from subcapitate to somewhat ex­ endemic. panded. In the expanded form the plants :ite a The specimens upon which this variety are based close matchfor much ofO. arctica in a strictsense. vary considerably. They tend to be tall plants with The placement ofO. arctica var. harnebyana with severalflowered racemes, butthey approachifnot O. sordida, a taxonomic entity considered by actually pass into vat. arctica (specimens from European authors (see Flora Europaea) at infra­ Anaktuvuk). Leaflet arrangement ranges from en~ specific level within O. campestris, has consider­ tiretofasciculate to merelyscattered. Tothe south able merit. However, the treatment of var. they simulate specimens of var. murrayi, de­ barnebyana at infraspecific level within O. sor­ scribed by Jurtsev (1986) in O. sordida. dida, a highly variable taxonomic entity with both pale and colored flowers, by Jurtsev (1986), does var. murrayi (Jurtsev) Welsh comb. nov. not solve the basic problem of the similarity of Basionym: O. sotdida var. murrayi Jurtsev Arctic FI. barnebyana to O. arctica, however well that U.S.S.R. 9(2): 179.1986. placement indicates the similarity of this North Distribution: St. Elias Mts., SW Yukon, Canada; en~ American phase with theprotean campestris com­ demic. plex. The calyx teeth ofvar. barnebyana vary from This taxon is the portion ofthe arctica complex most short, as in some phases of O. campestris, to al­ similar to var. bellii, from which it is distantly most as long as in some phases ofO. arctica. The isolated, but is probably most nearly allied to the sympatry of O. arctica and the allopatry of O. nearer disjunct, var. koyukukensis. Robust mate­ campestris, while not conclUSive, weigh in the rials ofvar. murrayi closely simulate some ofvar. decision to keep the Kotzebue materials of var. koyukukensis. Thedifferences betweenvats. mur­ barnebyana with O. arctica. And, the lack of a rayi and bellii rest on such int(ingibles as the ap~ stopping pl(ice for additional inclusions of North parently larger flowers, broader calyces with American taxa within expanded versions of mixed shaggy villous vesture, and tendency to campestris orsordida is likewise a consideration. larger leaflets ofvar. murrayi. C/. subumbellata Hooker, Parry's Second Voy., Ap- ssp. bellii (Macourt) love & Love, Taxon 31: 347. pend. 4: 396. 1825. 1982. type: ? Basionym: Spiesia bellii Britton ex Macoun = O. arctica R. Brown var. arctica = O. arctica R. Brown var. bellii (Britt.) Boivin ~ uniflota Hooker, Parry's Second Voy., Append. 4: var. bellii (Britton) Boivin, Naturaliste Canad. 94: 73. 396.1825. 1967. Type: Barrow River, E coast Melville Peninsula, lat. Basionym: Spiesia bellii Britton ex Macoun 67°g1'N, on Parry's SecondVoyage; isotypes CHI, Distribution: Kewatin, vicinity of Hudson Bay, NY! Canada; endemic. O. nigrescens (Pallas) Fischer var. uniflora The simil(lrity of this large-flowered low plant of (Hooker) Barneby coastal or near coastal Hudson Bay and vicinity to specimens designated as var. mu.rrayi is readily varietas notabilis, R. Brown, ChIor. Melvill., 254. apparent. They also simulate closely plants from 1823. Armatkchene Island off the coast of Siberia. The = O. nigrescens (Pallas) Fischer var. uniflora latter are evidently included by Jurtsev (1986) as (Hooker) Barneby portions ofan expanded O. sordida. The alliance Oxytropis arctobia Bunge, Gen. Oxytropis, 114. 1874. of var. bellii to O. arctica has long been recog­ Type:? nized. Any expansion of O. sordida to include it = O. nigrescens (Pallas) Fischer var. uniflora would also, logically, include the remainderofthe (Hooker) Bameby arctica complex. Such a proposal is herein consid­ var. hyperarctica Polunin, Bot. Canad. E. Arctic, ered as both illogical and unnecessary. 293, pI. 8. 1940. 1IinflataHooker, FI. Bor.~Amer., 1: 146.1834. Type: Franklin district, Baffin Island, Arctic Bay, N. Type: "Without locality, '0. arcticacomm. Hooker'''; Polunin 2583, 8-11 Sept. 1936; holotype CAN; holotype ?; isotypes GHI, NYI, PH! isotypes GR!, BM, OXF. = O. podocarpaA. Gray O. nigrescens (Pallas) Fischer vat. uniflora The specimen at GH! is accompanied by a small label (Hooker) Barneby bearing the notation, "0. arctica 11 inflata." Below Oxytropis attoputpurea (:Jlyclberg) A. Nelson, Univ. it in Asa Gray's handwriting is the note, "0. podo­ Wyo. Pub. Bot. 1: 117.1928. carpa. Plenty and same with the rather inflated Basionym: Aragallus atropurpureus Rydberg legume in Hb. Kew." Below that, in the writing of Oxytropis lagopus Nuttall var. atropurpurea a person not identified, is the notation, "Highest (Rydberg) Barneby . 384 GREAT BASIN NATURALIST [Volume 51

Oxytropis avenmelsonii (Lunell) A. Nelson; Univ. Wyo. Oxytropis borealis de CandoIle; Prodromus 2: 275. 1825. Pub. Bot. 1: 116. 1926 Type locality: "In terra Tschuktschorum ad sinum Basionym: Aragallus aven"nelsonii Lunell Sancti~Laurentii," collector not stated. Type: "e = Oxytropis lambertii Pursh var. lambertii sinu S. Laurentii in terra Tsuchuktschorum (pays Oxytropis bellii (Britton) Palibin, Bull. Soc. Bot. Geneve des Tchouktchi) septentrionem versus a fretlls II, 2: 19. 1910. Beringii. Legumina divisa a leg. ox. montana. m. Basionym: Spiesia bellii Britton Fischer 1825"; G"DC.! = O. arctica R. BroWn var. bellii (Britton) Boivin = O. borealis de Candolle var. borealis Oxytropis besseyi (Rydberg) Blankinship, Mont. Agric. var. australis Welsh, Great Basin Nat. 50: 359. 1991. Coli. Sci. Stlld. Bot. i: 80. 1904. This southern phase ofO. borealis is mainly montane Basionym: Aragallus besseyi Rydberg in distribution, but occurs mostly on xeric sites in sagebrush, black sagebrush, grass, ponderosa pine, var. argophylla Cllydberg) Barneby, Leafl. W. Bot. and aspen parkland communities, often on ex". 5: Ill. 1951. posed ridges or outcrops. Main substrate types Basionym: Aragallus argophyllus Rydberg are of igneous origin, either granitic or basaltic Distribution: W Montana, adjacent Idaho, and NE derived soils, but limestone also serves as a sub­ Nevada. strate. Elevational range varies from 2135 to var. besseyi 3355m. Distribution: Idaho, Montana, Wyoming, and Distribution: Inyo and Mono counties, California, Saskatchewan. Nevada, and S Utah. var. fallax Barneby, Proc. Calif Acad. Sci. IV, 27: 13 Hooker & Arnott, Bot. Beechey Boy., 122. 1832. 235.1952. Type:? Type: Clay slope among boulders, 4350 ft., mouth of = Oxytropis borealis de Candolle var. borealis Shell Creek, Big Horn Mountains, Big Horn County, Wyoming, 4,350ft., H. D. Ripley&R. C. var. borealis Barneby 8010, 16 June 1946; holotype CAS!; iso" Distribution: N.W.T., Yukon, and Alaska; Chukotsk. types GH!, US!, RM! The relatively few leaflets, ample flowers, and con- Distribution: West ofthe Bighorn River, Wyoming. densed, copiously hirsute inflorescence in combi" nation allow this entity to be ratherreadily identi­ var. obnapiformis (C. t. Porter) Welsh, Great Basin fied. Itconsists, atleast in part, ofwhat has passed Nat. 38: 337. 1978. . under the name of O. glutinosa Porsild, who ex­ Basionym: O. ohnapiformis C. L. Porter cluded the type of"subsucculenta" from consider" Distribution: NW Colorado, NE Utah, and SW ation in treatment ofthe genus in"Vascular Plants Wyoming. of Continental Northwest Territories Canada" The pod ofthis variety is strictly sessile or nearly so, (Porsild & Cody 1979). Included within the con­ not stipitate as in other phases ofthe species with ceptofvar. borealis is the O. uralensis 13 subsuccu­ inflated pods. Plants from the type locality near lenta Hooker, the basis ofO. viscida var. subsuc" Mayfield, Colorado, have 13-25 leaflets and inflo" culenta (Hooker) Barneby. rescences about equaling the leaves. Those from Daggett County, Utah, have 5-9leaflets andinflo" var. kudsonica (E. Greene) Welsh, Great Basin Nat. rescences surpassing the leaves. Plants from else" 50: 357. 1991. where bridge the two phases. Basionym: Aragallus hudsonicus E. Greene, Proc. Bio!. Soc. Wash. 18: 17. 1905. var. salmonensis Barneby; Proc. Calif. Acad. Sci. IV, Distribution: Yukon east to Hudson Bay. 24: 234. 1952. This is the phase of the species that occurs in North Type: Salmon River Canyon, 12 mi below Clayton, America mainly east of the Yukon, but with some Custer County, Idaho, alt. 5,400 ft., H. D. Ripley representation in that province, where it is transi­ & R. C. Barneby 8829, 22 June 1947; holotype tional with both var. viscida and var. sulphurea. CAS!; isotypes NY!; IDS. . Distribution: Custer Co., Idaho; endemic. var. sulphutea (Porsild) Welsh, Great Basin Nat. 50: 358. 1991 var. ventosa (Greene) Barneby, Leafl. W. Bot. 5: Ill. Basionym: Oxytropis viscidula ssp. sulphurea Por­ 1951. sild, Bull. Nat. Mus. Can. 121: 247. 1951. In­ Basionym: Aragallus ventosus Greene '. cludes: O. verruculosa Porsild. Distribution: S Montana, W Wyoming, and NE Utah. Distribution: British Columbia, Yukon, and E Alaska. Oxytropis b,locularis A. Nelson, Univ. Wyo. Pub. Bot. 1: These are the pallid-flowered plants ofthe Yukon and 114. 1926 Alaska. In theirmosttypical condition theracemes Type: Arizona, high moist slopes, Miller Peak, are compactly and uniformly smallflowered. They Huachuca Mts., Cochise County, L. N. Goodding vary from that norm to elongate racemes With 2411, 22 August 1907; holotype RM!; isotypes small to large flowers. The bracts are mainly small, GH!, NY!, UC!, POM. but in some they are very long and conspicuous in = O. lambertii vat. bigelovii A. Gray the inflorescence. On the one side the plants seem Oxytropis blankinshipii (A. Nelson) K Schumann; Just's to grade with var. hudsonica and on the otherwith Bot. Jahresb. 27: 496. 1901. both var. viscida and var. borealis. Basionym: Aragallus blankinshipii A. Nelson var. viscida (Nuttall) Welsh, Great Basin Nat. 50: 359. = O. lagopus Nuttall var. lagopus 1991. 1991] OXYTROPIS DG-NAMES, BASIONYMS, TYPES, SYNONYMS 38"5

Basionym: O. viscida Nuttall, ex Torrey & Gray, Flora val'. cusickii (Gteenman) Barneby, Leafl. W. Bot. N. ArneI'. 1: 341. 1838. 5: HI. 1951. Distribution: Alaska, Yukon, N.W.T., Gaspe, British Basionym: O. cusickii Greenman Columbia, Alberta, Minnesota, Oregon, Idaho, Distribution: British Columbia, Alberta, Washington, Wyoming, Nevada, Utah, Colorado, and Califor­ Oregon, Idaho, Montana, WYOming, Utah, and nia. Colorado. This variety includes almost as much diversity as the This taxon is highly variable in flower size, especially species as a whole. The numerous subunits are where the large~flowered O. sericea var. spicata held together by tenuous characteristics that are occurs nearby. Transitionally apparent popu­ difficult to define or place in a key. Variation is lations again demonstrate the lack of consistent often great in populations from adjacent hillsides diagnostic features to separate what are otherwise or on a single gravel bar, especially in the Arctic. distinctive populations. The same problem is ap~ One is reminded ofthe conditions ofmorphologi­ parent where var. gracilis occurs nearby at lower cal variation occurring in the boreal O. nigrescens elevations than these montane phases of the var. nigrescens, as treated by this author. Unless campestris complex (see also var. varians). one is willing to support a taxonomy wherein the val'. davisii Welsh, Leafl. W. Bot. 10: 25. 1963. purported taxa are largely sympatric and consist of Type: British Columbia, mi 403.4, Alaska Hwy, morphological subunits whose genetic continuity R. J. Davis 6076, 19 July 1962; holotype BRY!; is questionable, made up of a series of similar isotype.IDS! plants held together by that similarity and not by Distribution: SW Alberta, NE British Columbia. genetic linkage, there does not seem to be a rea­ This plant is readily distinguished by its colorful flow­ sonable way to segregate the morphological varia­ ers, fasciculate leaflets or tendency to faspiculate tion as taxa. The ratherlarge numberofsynonyms, leaflets, and elongate inflorescences. Specimens often at specific or varietal level, reflects the have been known in collections from early times, attempts at segregation. but have been regarded as occasional intermedi­ Oxytropis bushii Gandoger, Bull. Soc. Bot. France 48: ates between portions of the campestris complex xvii. 1901. and o. borealis var. viscida, or, they have been Type: Missouri, Watson, Atchison County, B. F. identified, because of the fasciculate leaflets, as Bush 204, 7 June 1894; isotypes GH!, NDG, ISC! o. splendens. The plants are locally abundant on NY!, US!, MIN!, MOl stream gravels and adjacent slopes in the foothills = O. lambertii var. lambertii mainly of the Alberta Rockies and in northeast Oxytropis campestris (L.) de Candolle, Astragalogia, 59. British Columbia. The plants form apparentinter­ 1802. mediates with var. gracilis. Basionym: Astragalus campestris L. val'. dispar (A. Nelson) Barneby, Leafl. W. Bot. var. americana Brunet, Cat. PI. Canad., 39. 1865, 5: HI. 195:1.. nomen. Basionym: Aragallus dispar A. Nelson = O. campestris (L.) de Candolle var. johannensis Distribution: North Dakota and Manitoba. Fernald l'lants ofthis variety are closely allied to var. gracilis, val'. cervinus (Greene) Boivin, Naturaliste Canad. 94: from which they differ in the flowers being poly~ 75.1967. chrome in populations, and in the. somewhat Basionym: Aragallus cervinus Greene firmer texture ofthe pods. It may well be that var. = O. campestris (L.) de Candolle var. gracilis d4Paris the somewhatstabilizedproductofprevi­ (A. Nelson) Barneby ous hybridization involving the mainly disjunct val'. chartacea (Fassett) Barneby, Proc. Calif. Acad. pale-floweredvat. gracilis and thenowfar disjunct IV, 27: 269. 1952. purple-flowered vars. davisii and johannensis. Basionym: O. chartacea Fassett Purple-flowered or polychrome populations or Distribution: Known only from lake shores in central entire taxa within the campestris complex are and NWWisconsin; endemic. now known to be at least as important as are the Plants with other ofthe syndrome ofcharacteristics of pale-flowered phases. Indeed, pink-purple flow­ var. johannensis in Ontario (Farm River area) also ers show up here and there throughout North have short pods as in var. chartacea. These have America, even in othe~sewhite or ochroleucous not been examined in the field, and it seems best populations. not to relegate this otherwise Wisconsin endemic val'. EglabrataHooket, FI. Bor.-Amer. 1: 147.1834. to synonymy until more study has been com­ Type: Bear Lake to the Arctic Shores and Islands pleted. [possibly Richardson]; isotype (?) GH! vat. columbiana (St. John) Bameby, Leafl. W. Bot. = o. maydelliana Trautvetter 5: IH. 1951. ssp. gracilis (A. Neison) Boivin, Naturaliste Canad. Basionym: O. columbiana St. John 94:74.1967 Distribution: Columbia River above the mouth ofthe Basionym: AragallUS gracilis A. Nelson Spokane River, NE Washington (where possibly O. campestris de Candolle var. gracilis extirpated), andforested margin ofFlathead Lake, = (L.) (A. Nelson) Barneby Montana. This variety is characterized by its white to ochroleu­ val'. gracilis (A. Nelson) Bllrneby, Leafl. W. Bot. cous flowers with maculate keel tips and soft 5: HI. 1951. pubescence. It is still extant at Flathead Lake. Basionym: AragallUS gracilis A,. Nelson 386 GREAT BASIN NATURALIST [VolUlne 51

Distribution: Mainly south of the 65th parallel in that occur inland in northern Yukon Territory and British Columbia, Alberta, Saskatchewan, Manh west in Alaska to Prudhoe Bay and beyond. The toba, Washington, Montana, Wyoming, South variety is characterized by its small pink"purple Dakota, North Dakota, and Colorado. flowers and other features that simulate and pass This is a highly variable taxon whose morphological into the partially sympatric vat. jordalii. The var. subunits have been regarded by some previous roaldii further simulates and is probably allied to workers as belonging to several specific or infra~ the eastern var. terrae-novae. specific taxa. A portion of the problem revolves var. rydbergii {A. Nelson):I1. j. Davis, Madrono 11: around the inclusion ofmaterials with two differ­ 144.1951. entbase chromosome numbers, 3gand48. At least Basionym: O. rydbergiiA. Nelson some portion of it is the southern counterpart of = O. campestris (L.) de Candolle vat. cusickii var. varians, which is reported to have the base (Greenman) Barneby chromosome number of48. The variety is transh tional to vat. davisii in southwestern Alberta, to 13 speciosa Torrey & Gray, Fl. N. Amer. 1: 341. 1838. var. dispar in North Dakota, and more especially Type: "Dr. Hooker" NY!. See Flora Boreali-Ameri­ to the montane var. cusickii, which often occurs at cana 1: 147. 1833. "Carleton House Saskatchewan the tops ofthe same mountains whose bases bear to Rocky Mts. [T.] Drummond." The sheet at NY var. gracilis. beats the notation quoted above and the initials E. P. S[heldon]. var. johannensis Fernald, Rhodora 1: 88. 1899. = O. sericea Nuttall var. spicata (Hooker) Barneby Type: Maine, gravelly shores, valley ofSt. John River, Fort Kent, Aroostook County, M. L. Fernald II spicata Hooker, Fl. Bor.-Amer. 1: 174. 1834. 2289, 15 June 1898; holotype GH!; isotype CAN!, Type: Between Carlton House on the Saskatchewan US!,NY! . and the Rocky Mountains [Alberta], T. Drum" Distribution: Newfoundland, New Brunswick, Nova mond s.n.; holotype?;? isotypes GH, NY! Scotia, Quebec, Ontario, and Maine. = O. sericea Nuttall var. spicata (Hooker) Batneby Plants ofthis variety from the Farm River area, south vat. terrae-novae (Fernald) Barneby, Proc. Calif. of James Bay, Ontario, have fasciculate leaflets Acad. IV, 27:266. 1959. and short pods. In the latterfeature they simulate Basionym: O. terrae-novae fernald the vat. chartacea, which might best be regarded Distribution: Hudson Bay, Baffin Island, Ungava as only a disjunct phase ofthis variety. Peninsula, Labrador, and coastal Newfoundland. ssp. jordalii (Porsild) Hnlten, Ark. Bot. 7: 79.'1967. Reports of this taxon from the Mackenzie Mountains purple~flowered Basionym: O. jordalii Porsild are probably of the vat. roaldii, which is a western vicariad of var. terrae-novae = O. campestris (L.) de Candolle var. jordalii (Por­ sild) Welsh that differs in minor but consistent ways. At Churchill, Manitoba, there is a mixture ofspeci­ var. jordalii (PorsiId) Welsh, Leafl. W. Bot. 10: 25. mens variously assigned to vars. varians, terrae~ 1963. novae, orjohannensis. They should be viewed in Basionym: O. jordalii Porsild the field prior to an attempt to resolve their rela­ Distribution: N Alaska, N Yukon, and N.W.T. tionships. Mackenzie District. var. varians (Rydberg) Barneby, Proc. Calif. Acad. This is a dwarfborealvarietyofridge tops, gravel bars, IV, 27: 253. 1952. and arctic tundrawith small White to ochroleucous Basionym: Aragaztus varians Rydberg flowers. It is easily recognizable in its typical Distribution: Alaska, Yukon, N.W.T., N Manitoba, phases, butitpasses bydegree, especially in lower and N British Columbia, mainly north ofthe 65th elevation and more mesic sites into vat. varians. parallel. On some gravel bars, especially, the plants are Plants ofthis variety are highly variable, with numer­ transitional with the other dwarf boreal var. ous differing morphological phases often growing roaldii. Itis with the lattervariety, with pink"pur" together on the same gravel bar in portions of pIe flowers, thatitseems to be most closely allied. Alaska and the Yukon. Alpine portions ofthe vari~ var. ~meltLnocephalaI:looker,Fl. Bor.-Amer. 1: 147. ety, especially in SW Yukon, N British Columbia, 1834. and adjacent SE Alaska, closely simulate high"alti­ Type: Bear Lake to the Arctic Shores and Islands tude materials ofO. campestris var. cusickii at its [possibly Richardson]; isotype (?) GH. northern limits in S British Columbia andAlberta. = O. maydelliana Trautvetter Indeed, there are specimens of O. campestris var. roaldii (Ostenfeld) Welsh; comb. et stat. nov. var. gracilis, the so called "cervinus" phase in Basionym: O. roaldii Ostenfeld, Vasco pI. Arct. N. southern British Columbia, that almost match the Amer. Gjoa Exped., 54, pI. 3, fig. 16. ·1910. "alaskana" materials ofvar. varians from southern Distribution: N Alaska, N Yukon, and adjacent Alaska. The maintenance ofvar. varians as sepa­ N.W.T. ratefrom var. gracilis rests mainly on allopatry and The type ofthis variety was included in the synonymy historical perspective. Specimens ofvar. varians ofO. arctica byBarneby (1952), whose experience appear to intergrade with those ofvar. jordalii in withboreal representatives was limited by the few montane sites near Juneau, Alaska. speCimens available to him. The type specimen var. verrucosaLedebour, Fl. Ross. 1: 591. 1842. was taken on Herschell Island on the Amundsen Type: "in terra Tschuktschorum ad sinum Sancti_ Gjoa Expedition and is clearly the same as plants Laurentii," collector not stated, but probably 1991] OXYTROPIS DC.~NAMES,BASIONYMS, TYPES, SYNONYMS 387

based on the type ofO. borealis (q.v.). O:wtropis cusickii Greenman, Erythea 7: 116. 1899. = O. borealis de Candolle var. borealis Type: Oregon, alpine summits ofthe Wallowa Moun­ var. viscida (Nuttall) S. Watson, U.S. GeoI. ExpI. tains, W. C. Cusick i365, 2095, 23 August 1886; 40th Parallel, Bot. 5: 55. 1871. lectotype (Eliseris & Packer, Canad. J. Bot. 58: Basionym: O. viscida Nuttall 1827. 1980) GHI; Cusick 2095, Aug. 1898, syn­ types GHI, NYl, NDGl, UC, USl = o. borealis de Candolle var. viscida (Nuttail) Welsh O. campestris (L.) de Candolle var. cusickii var. wanapum Joyal, Great Basin Naturalist 50: 373. (Greenman) Barneby 1991. OXyttopis czukotica Jtirtsev, )3uti. Mosk. Obtz. Icp. Type: Washington, Saddle Mt. above Lower Carb Prir., Otd. BioI. 85. 6: 102.1980. Creek, E. Joyal 12643, 25 May 1987; holotype nom. nov. pro O. tschuktschorumJurtsev USI; isotypes BRY!, ISCI, OSU! = O. nigrescens (Pallas) Fischer vat. nigrescens Distribution: Known only from the type locality. This is a plant of xeric, basaltic talus; the flowers Oxytropis deflexa (Pallas) de CandoIle, Astragalogia, 33, suffused with purple are diagnostic, since no other tab. 27. 1802. phases ofthe genus in the Pacific Northwest typi­ Type: "ad nivaliajuga Dauria ... in excelsis montibus cally have colored flowers. The narrow leaflets circa balyta rivum aliosque Ononem influentibus" tend to be involute and to vary in numberfrom 20 [Siberia], S. P. Pallas s.n.; holotype BM. to 25. These vegetative features are unlike any of var. capitata BoiVin, Svensk Bot. Tidskr. 56: 499. the other several varieties of O. campestris that 1962. occur elsewhere in North America haVing laven~ type: Ontario, cordon gazonnant Ie long d'une riv­ der to purplish flowers. iere, 34 mi au sud du cap Henriette, 44°54'N, A. Dutilly & E. Lepage 34367, 17 Aout 1953; holo­ Oxytropis cascadensis St. John, Proc. BioI. Soc. Wash. typeDAO! 41: 105. 1928. = O. deflexa var. foliolosa (Hooker) Barneby Type: Washington, Grouse Creek, Mt. Baker, What­ com County, St. John 5513; holotype WTCI; photo var. culmtnis Jepson, FI. Calif. 2: 381. 1936. BRY! Type: Cottonwood Creek, White Mts., Mono Co., = O. campestris (L.) de Candolle var. gracilis (A. California, V. Duran 1650, 29 June 1926; holotype Nelson) Barneby UCI = O. deflexa var. sericea Torrey & Gray Oxytropis caudata (Greene) K. Schumann, Just's Bot. Jahresb.27:496.1901. var. de.flexa Basionyrn: Aragallus caudatus Greene Distribution: Colorado, Utah, and reported (Barneby = O. splendens Douglas 1952) less commonly elsewhere in North America; Asia. Oxytropis chanacea Fassett, Rhodora 38: 95. 1936. As interpreted by me and by Barneby(1952), plants Type: Wisconsin, sandy shore ofLake Huron, Plain­ similar to the typical Siberian material ofthe spe~ field, Waushara County, N. C. Fassett 16704, 15 cies occur disjimctly in America, mainly in the Sept. 1936; holotype WISI; isotypes GHI, ISC!, mountains of Colorado and Utah. the combina­ MOl, lAI, GHI, NY, MUI, NYI, PH!, USI, UC!, tion of racemes With 10 to 20 large flowers on DAOI plants that aremainly acaulescent or shoft caules­ = O. campestris (L.) de Candolle vat. chartacea (Fas­ cent forms a distinctive morphology. They differ sett) Barneby from var. foliolosa in about the same manner that Oxytropis collina (A. Nelson) K. Schumann, Just's Bot. that variety differs from var. sericea. Jahresb. 21: 496. 1901. var. dezhneviiJurtsev, Bot. Journ. 57: 6,64'7.1972. Basionym: Aragallus collinus A. Nelson Type: ? = O. nana Nuttall = O. deflexa vat. foliolosa (Hooker) Barneby The placementofthis plantwith var. foliolosa is based OXytropis columbiana St. John, Proc. BioI. Soc. Wash. on examination of authentic material of the pro­ 41: 100. 1928. posed entity in the herbarium at BRY. The speci­ Type: Gravelly beach ofthe Columbia River, Marcus, mens examined from western AlaSka are some­ Stevens County, H. St. John 6482,27 June 1924; what more apparently black hairy than materials holotype WTC; isotype GHI from farther east. Otherwise they seem to fit = O. campestris (L.) de Candolle var. columbiana (St. within the broad concept ofvar. foliolosa. John) Barneby ssp. dezhnevii (Jurtsev) Jurtsev, Arctic FI. S.S.S.R. Oxytropis condensata (A. Nelson) A. NelsOIl, Univ. Wyo. 9: 19.1986. Pub. Bot. 1: 115. 1926. Basionym: O. deflexa var. dezhnevii Jurtsev Basionym: Aragallus albiflorus var. condensatus A. var. foliolosa (Hooker) Barneby, Leafl. W. Bot. 6: Nelson 111. 1951. = O. sericea Nuttall var. sericea Basionyrn: Astragalus deflexus Pallas Oxytropis coronaminis Fernald, Rhodora 30: 151, pI. Distribution: Rocky Mountains, from Alaska to Ne_ 175.1928. vada and Wyoming; also on atctic and subarctic Type: Mackenzie, Arctic sea~coast, Dr. Richardson shores of Mackenzie, UngavaPeninsuIa, Baffin s.n.; holotype GHI Island, Labrador, Newfoundland, and Gaspe = Oxytropis arctica R. Brown var. arctica Peninsula. 388 GREAT BASIN NATURALIST [Volume 51

This is the most showy ofthe two common phases of Oxytropis glaberrima Hulten, Bot. Tidskr. 5: 124. 1954. the species in North America. The flowers are Type: Alaska, Kurupa Valley, 7 mi N ofKurupa Lake, typically brightly colored in compact to subcom­ A. R. Hodgdon 8060,25 June 1952; holotype S! pact racemes with up to 10 flowers. Sometimes = O. nigreseens (Pallas) Fischer var. nigreseens there are more flowers in rather lax racemes, but Oxytropisglabrata (Hooker) A. Nelson, Univ. Wyo. Pub. the racemes seldom surpass 10 em long even in Bot. 1: 117.1926 fruit. The herbage is typically sparingly pilose and Basionym: O. eampestris E glabrata Hooker. green in aspect, and both caulescent and acaules­ = O. maydelliana Trautvetter cent phases occur. Some of the variation, espe­ cially portions of that regarded as var. eapitata Oxytropis glutinosa Porsild, Bull. Nat. Mus. Canad. 121: Boivin, is geographically correlated. Further seg­ 240. 1951. regation, however, seems unnecess(lry. Transi­ Type: Yukon, Canol Rd.: Mile 132. Lower Lapie tional specimens to var. seneea are known. Crossing, dry shaly slopes of mountain west of road, A. E. Porsild &: A. J. Breitung 9730, 20 June var. parviflora Boivin, Svensk Bot. Tidskr. 56: 498. 1944; holotype CAN; isotypes GH!, ISC!, SI, NY!, 1962. US! Type: Aiberta, Ft. Smith, 1 mi SW, bl~ck sandy soil in = Q. borealis de Candollevar. viscida (Nuttall) Welsh Populus spp., Salix spp., Picea glaucawoods along trail, W. J. Cody &: C. C. Loan 4496, 17July 1950; Oxytropis gorodkovii Jurtsev, Bot. Journ. 53: 11. 1968 holotype DAOl; isotype RMl (nom. nov. pro Astragalus pygmaeus Pallas; O. pygmaea = O. deflexa (Pallas) de Candolle var. seneea Torrey [Pallas] Fernald). &:Gtay = O. nigreseens (Pallas) Fischer var. nigreseens Oxytropis gracilis (A. Nelson) K. Schumann, Just's Bot. ssp. retrorsa (Fernald) Love &: Love, Taxon 31: 347. Jahresb.24:496.1901. 1982. Basionym: Aragallus gracilis A. Nelson Basionym: O. retrorsa Fernald = O. eampestris (L.) de Candolle var. gracilis (A. = O. deflexa var. seneea Torrey &: Gray Nelson) Barneby 13 [vat.] seneea Torrey &: Gray; Fl. N. Amer. 1: 242. Oxytropis hallii Bunge, Gen. Oxytropis, 162. 1874. 1838. Type: Rocky Mts., Lat. 39°-41°, E. Hall &: H. Har­ Type: "Rocky Mountains near streams" [Wyoming or bour 143 (in part), 1862; isotypes GH!, NY (in Idaho], T. Nuttall s.n., 1834; holotype NY! packet)!, US! Distribution: Alaska, Yukon, N.W.T., British Colum~ = O. podoearpaA. Gray bia, Alberta, Manitoba, Washington, California, Idaho, Montana, New Mexico, Oregon, Utah, and Oxytropis hookeriana Nuttall ex Totrey &: Gtay, Fl. Wyoming. N. Amer., 340. 1838 This is the comIIlon phase of O. deflex(L south of the Type: "0. hooken. O. lambertii Hook. Platte Plajns. Arctic. The vllriety is highly variable, especially in T. Nuttall"; holotype BM. size and aspect ofthe flowers. The small-flowered = O.lambertiivar.lambertii phaseofthenorthernRockies was proposed as var. OXytropis huddlesonii Porsild, Bull. Nat. Mus. Canad. parviflora Boivin. Flowers ofsome ofthe variants 121: 242. pI. 11, fig. 5. 1951. appear not to open, as ifthey are cleistogamous. Type: Yukon, Golden Horn Mountain, south of Furtherworkis indicated on this group. Whitehorse, in lichen heath above timberline, elevation 5,800 feet, C. W. Huddleson s.n. 1944; Oxytropis dispar (A. Nelson) K Schumann, Just's Bot. holotype CAN! Jahresb. 27: 496. 1901. Distribution: Sand E Alaska, S Yukon, and N British Basionym: Aragallus dispar A. Nelson Columbia; endemic. = O. eampestris (L.) de Candolle var. dispar (A. The presence of unilocular, glabrous or sparingly Nelson) Bameby strigose pods distinguishes this entity from Oxytropis falcata (Greene) A. Nelson; Univ. Wyo. Pub. O. nigreseens, its apparent close ally in boreal Bot. 1: 118. 1926. northwestern North America. Basionym: Aragallusfaleatus Greene Oxytropis hudsonica (Greene) Fernald, Rhodora30: 142, = O. lambertii var. lambertii pI. 112. 1928. Oxytropisfoliolosallooket, Fl. Bor.-Amer.1: 146. 183l. Basionym: Aragallus hudsonicus Greene Type: "Carlton House to the Rocky Mountains, in lat. = O. borealis de Candolle var. hudsoniea (Greene) 54°" [Alberta], collector not noted, probably T. Welsh Drummond; holotype K; isotype NY! Oxytropis hyperborea Porsild, Sargentia4: 53. 1943. = O. deflexa var.foliolosa (Hooker) Barneby Type: N.W.T., Mackenzie R. Delta, E branch, Oxytropis g(LSpensis Fernald &: :r<:elsey, Rhodora 30: 123. 68°55'N, A. E. Porsild 1033, 21 July 1934; holo­ 1928. type CAN; isotype GHI, SI, US! Type: Quebec, Gaspe County, dry talus ofslaty cliffs, = Oxytropis eampestris (L.) de Candolle var. vanans northernface ofMt. St. Piette, atmouthofRiviere (Rydberg) Barneby a Pierre, M. L. Fernald &: L. B. Smith 25874, 14 Oxytropis inflata (Hooker) Steffen, Beih. Bot. Centralbl. August 1923; holotype GH!; isotypes MO!, CAS!, 58B: 162. 1938. NY! Basionym: O. aretiea 8 inflata Hooker = O. borealis de Candollevar. viseida (Nuttall) Welsh = O. podoearpaA. Gray 1991] OXYTROPIS DC.-NAMES, BASIONYMS, TYPES, SYNONYMS 389

Oxyttopis involuta (A. Nelson) K SChUmlmn, Just's Bot. rather than the merely purplish suffused ones o Jahresb. '/,7: 496. 1901. ofO. kobukensis. Basionym: Aragallus involutus A. Nelson = Oxytropis lambertii Pursh var. lambertii Oxytropis kokrinensis Porsild, Rhodora 41: 251, tab. 553. i939. Oxytropis ixodes Butters & Abbe, Rhodora 45: 2, tab. Type: Kokrines Mountains, north side of divide 745, figs. 1-6. 1943. towards Melozitna River, 65°20'N, 154°30'W, Type: Minnesota, slate cliffs on northside ofa high hill alpine slopes 2000-4000 ft, A. E. &c R. T. Porsild 1/2 mi. west of the outlet of South Fowl Lake, 76'/" 1926; holotype CAN!, fragment S! Cook County, F. K. Butters, E. C. Abbe, &G. W. Distribution: Ridge tops in the Kokrines Mountains Bums 611, 27 June 1940; holotype MIN; isotype and western Brooks Range, Alaska; endemic. GH!, NY!, PH!, US!, UC!, DAO! This species is evidently allied to O. podocarpa by its = O. borealisdeCandollevar. viscida (Nuttall) Welsh inflated, stipitate pods. It is easily distinguished from that entityby its purplish stipules. var. ecaudata Butters & Abbe, Rhodora 45: 4. 1943. Type: Ontario, slate cliffs east of North Fowl Lake, Oxytropis koyukukensis Porsild, Canad. Field~Nat. 65: Thunder Bay District, F. K. Butters, E. C. Abbe, 78.1951. & G. W. Bums 682, 1 July 1940; holotype MIN; Type: Alaska, Wiseman, L. H. Jordal1763, 15 June isotype GH! 1944; holotype CAN!; isotype US!, ISC! = O. borealis de Candolle var. viscida (Nuttall) Welsh = O. (Lrctica R. Brown var. koyukukensis (Porsild) OX!ftropis johannensis (Fernald) Fernald, Rhodora 30: Welsh 1435, pI. 113. 1928. OX!ftropis lagopus Nuttllll, J. Acad. Philad. 7: 17.1834. Basionym: O. campestris (L.) de Candolle var.johan­ Type: Sources of the Missouri, N. B. Wyeth s.n., nensis Fernald 1833; holotype PH!; isotype NY! f. bicensis Victorin llr. Rousseau, Cont. Inst. Bot. var. attopurputea (Rydberg) Barneby, Leafl. W. Univ. Montreal 36: 32. 1940. Bot. 5: 111. 1951. Type": Quebec, Rimoufski Co., St. Fabien, Montagne Basionym: Aragallus atropurpureus Rydberg, Bull. du Bic, J. Rousseau 50297, 15 July 1986; holotype Torrey Bot. Club 34: 4'/,4. 1907. ?; isotype DAO! Distribution: SE Montana, Wyoming, and South = O. campestris (L.) de Candolle var. johannensis Dakota. Fernald var. conjugans Barneby, Proc. Cali£ Acad. IV, 27: Oxytropis jonesii Barneby, Proc. Calif. Acad. IV, 27: ~7.1952. 215.1952. Type: Vicinity of Helena, Montana, E. O. Wooton Type: Utah, pink calcareous gravel benches, Red s.n., 19'/,1; holotype US!; isotype NY! Canyon, Garfield County, alt. 7150 ft., 1947, Distribution: W Montana and S Alberta; endemic. H. D. Ripley & R. C. Bameby 8550, 7 June 1947; holotype CAS!, isotypes GH!, NY!, RM! Vlir. iagopus =0. oreophila var.jonesii (Barneby) Bameby Distribution: Nand E Idaho, W Montana, and N Wyoming. Oxyttopis jotdallii Porsild, Canad. Field"Nat. 65: 11, tab.!. 1951. Oxyttopis lambertii Pursh, FI. Amer. Sept., 740. 1814. Type: Alaska, OldJohn Lake, L. H. Jordal3580, 1950; "Type: "On the Missouri, on the bluffs from the Maha holotype CAN; isotypes US!, S! village to the Ponc!tts, Louisillna [NE Nebraska or = O. campestris (L.) de Candolle var. jordalii (Por" adjacent South Dakota or Iowa], Bradbury s.n., sild) Welsh 1811; holotype PH! ssp. davisii (Welsh) Elisells llr. Plicker, Canad. J. Bot. [3 Hooker, FI. Bor.-Amer. 1: 107.1834. 58: 1830. 1980. Type: Oxytropis lambertii [3. Dr. Hooker; isotype NY! Basionym: O. campestris (L.) de Candolle var. davisii = O. campestris (L.) de Candolle vat. davisii Welsh Welsh "I Torrey llr. Gray, FI. N. Amer. :I.: 339. 1838. = O. campestris (L.) de Candolle var. davisii Welsh Type: ? Oxytropis kobukensis Welsh, Iowa State J. Sci. 41: 286. = O.lambertiivar.lambertii 1967. 8 ? Torrey & Gray, FI. N. Amer. 1: 338. 1838. Type: Alaska, sand dunes, Kobuk River, across from Type: Quebec, nellr Quebec, Mrs. Percival s.n.; holo­ themouth ofthe HuntRiver, R. D. Hamiltons.n., type NY! 1938; holotype BRY! = O. campestris (L.) de Candolle var. johannensis Distribution: Sand dunes, Kobuk River, W Alaska; Fernald endemic. The enlarged, purplish stipules on elongllte caudex var. articultit(L (Greene) Bllrneby, Leafl. W. Bot. 5: branches are characteristic of this pink-purple 111. 1951. flowered plant of dunes in western Alaska. The Basionym: Aragallus articulatus Greene relationship ofthis entity likely lies with Siberian Distribution: Kansas (Meade Co.), Oklahoma, and species. Jurtsev points to O. ochotensis Bunge as Texas. having possible relationship. The few specimens This plant ofthe southern prairies 3.I1d plains is distill" of that taxon eXllmined seem to be smaller in guished by its short calyxteeth, large flowers, and all parts and to have chestnut colored stipules shortly exserted or included pods. 390 GREAT BASIN NATURALIST [Volume 51

ssp. bigelovii (A. Gray) W. A. Weber, Phytologia 51: Oxytropis leucantha (Pallas) Persoon, Syn. 2: 331. 1807. 374.1982. Basionym: Astragalus lellcanthlls Pallas B:iSionym: O. lambertii var. bigelovii A. Gray Note: All combinations cited are synonyms ofphases = O. lambertii var. bigelovii A. Gray ofO. borealis de Candolle-O. lellcantha (Pallas) Persoon belongs to o. carnpestris sens. lat., but var. bigeloviiA. Gray, Proc. Amer. Acad. 20:7. 1884. Juttsev (1986) regards itas a separate species allied Type: Rocky hillsides, upper Canadian [Guadalupe or to O. sordida (Willdenow) Persoon. San Miguel County, New Mexico], J. M. Bigelow s.n., Sept. 21,1853; holotype GH!; isotype US! Var. depressa (Rydberg) Boivin, Naturaliste Canad. Distribution: SE Wyoming, Colorado, New Mexico, 94: 77. 1967. Arizona, and Utah. Basionym: Aragallus viscidulus var. depressus Ryd~ This plant typically has large flowers and short to berg elongate calyx teeth and pods. Itis mainly a mon­ = O. borealis de Candolle var. viscida (Nuttall) Welsh tane or intermontane variety, which abuts with var. hudsonica f. galactantha Boivin, Natmaliste var. lambertii in Colorado and Wyoming. MateJi­ Canad. 94: 76. 1967. als ofvar. bigelovii from the canyonlands of Utah Type: Canada: Franklin District, Melville Peninsula, and adjacent Arizona have malpighian hairs with Repulse Bay, along NaujaRiver, P. F. Bruggeman very short attachment. Furthermore, the plants 52,21July 1950; holotype DAO! tend to be tall, with features of scape, inflores­ = o. borealis de Candolle var. hudsonica (Greene) cence, and leaflets attep.uated. Theleaflets further Welsh tend to disarticulate from the rachis readily. var. gaspensis (Fernald &: Kelsey) Boivin, Naturaliste Northward from the canyonlands ofKane and San Canad.94:76.1967 Juan counties, Utah, the attenuation ofparts fails, Basionym: Oxytropis gaspensis Fernald &: Kelsey even though the short branch of the malpighian O. borealisde Candolle var. viscida (Nuttall) Welsh hair persists. There does not seem to be sufficient = correlation of morphological features to warrant var. hudsonica (Greene) Boivlp., Naturaliste Canad. taxonomic status for these plants thatare obviously 94: 76. 1967. in transition. Basionym: Aragallus hudsoniclls Greene = O. borealis de Candolle var. hudsonica (Greene) f. canadensis Gandoger, BulL Soc. Bot. France 48: Welsh xvi. 1901. Type: Southern Manitoba in the valley of the Souris var. ixodes (Butters &: Abbe) Boivin, Naturaliste River, Portage"la~Prairie, MacMorine s.n., Aug. Canad. 94: 76. 1967. 1897; syntype DAO! Basionym: Oxytropis ixodes Butters &: Abbe = O. campestris (J.... ) de Candolle var. gracilis = O. borealis de Candolle var. viscida (Nuttall) Welsh var. ieuchippiana Boivin, Naturaliste Canad. 94: 76. var. lambertii 1967. DIstribution: British Columbia, Saskatchewan, Mani­ Type: Yukon: Whitehorse, airport area, steep slope, toba, Montana, North Dakota, Minnesota, South flowers Varying in colour from yellow to purple, Dakota, Wyoming, Nebraska, Iowa, Kansas, Mis­ abundant, J. M. Gillett&:J. A. Calder3181, 4 June souri, and Wyoming. 1949; lectotype here selected bAO!; isolectotype The coincidence oflongcalyx teeth, smallflowers, and CAS! elongate pods is diagnostic for this Variety of the = O. borealis de Candolle var. hudsonica (Greene) central and northern prairies and plains. Welsh var. lilacina Cockerell, W. Amer. Sci. 5: ll. 1888, var. magnifica Boivin, Naturaliste Canad. 94: 77. nomen. 1967. = O. sericea Nuttall vfir. sericea Type: Alberta, High River, J. Fletcher 457 1/2, 27 f. mixta Gandoger, Bull. Soc. Bot. France 48: xvii. June 1902; holotype DAO! 1901 = O. borealis de Candolle var. viscida (Nuttall) Welsh Type: Minnesota, Montevideo, L. Moyer s.n., 1909; Vln:. viscida (Nuttall) Boivin, Naturaliste Canad. 94: Wyoming, head of Pole Creek, A. Nelson 1320, '77.1961. 27 June 1895; syntypes NY! Basionym: O. viscida Nuttall = O. lambertii var. lambertii = O. borealis de Candolle var. viscida (Nuttall) Welsh var, ochroleucaA. Nelson, Bull. Wyo. Exper. Sta. 28: OXytropis luneZliana A. Nelson, Univ. Wyo. Pub. Bot. 1: 98.1896. 117. 1926, nom. nov. Type: Wyoming, Pole Creek, Albany County, A. Nel­ Basionym: Aragallus collinus A. Nelson. son 119, 2 June 1894; holotype RM!; isotype NY!, = Oxytropis sericea Nuttall var. spicata (Hooker) photo BRY! Nelson named this plant three times, Barneby in 1896 as O. lambertii var. ochroleuca, in 1899 as O:tytropis luteola (Greene) Piper &: Beattie, Fl. N.-W. Aragallus albiflorus, and in 1900 as Aragallus saxi­ Coast, 334. 1915. montanus. All are typified on the same collection. Basionym: Aragallus luteolus Greene = O. sericea Nuttall var. sericea = O. campestris (L.) de Candolle var. gracilis (A. vllr. sericea (Nuttall) A. Nelson, Erythea1: 62. 1899. Nelson) Bameby Basionym: O. sericea Nuttall Oxyttbpis macounii (Greene) Paytoll, Proc. BioI. Soc. = O. sericea Nuttall var. sericea Wash. 40: 120.1927. 1991] OXYTROPIS DC.-NAMES, BASIONYMS, TYPES, SYNONYMS 391

Basionym: Aragallus macounii Greene ssp. dispar (A. Nelson) Elisens & Packer, Canad. J. =0. campestris (L.) de Candolle var. gracilis (A. Bot. 58: 18g6. 1980. Nelson) Bameby Basionym: Aragallus dispar A. Nelson O. campestris (L.) de Candolle var. dispar (A. Oxytropis maydelliana Trautvetier, Act. Hort. Petrop. = .Nelson) Barneby 6: 16.1879. Type: "... in tractu fljlvium Anadyr inter et latus Oxytropis multicepsTorrey& Gray, FI. N. Amer. 1: 341. meridionale montium jugi a fluvio hoc septentri" 1838. onem versus siti" [Chukchi, from the basin ofthe Type: "Summit of lofty hills in the Rocky Mountain Anadyrandits northern tributaries], Baron G. von range, towards Lewis's River [S Wyoming], Rocky Maydell s.n., 1869; holotype LE? Mts.," T. Nuttall s.n., 1834; holotype GH!; iso­ Distribution: Alaska, Yukon, N British Columbia, types NY!, PH! N.W.T., E to Hudson Bay and Baffin Island; Distribution: Colorado, NE Utah, S Wyoming, andW Chukotsk and Kamchatica. Nebraska. The yellowish flowers and reddish brown stipules The accrescent calyces, broad bracts, and few flowers easily characterize this arctic species. are characteristic for the species, which stands alone in the genus in North America in its mor­ ssp. melanocephala (Hooker) Porsild, Nat. Mus. phology. Canada Bull. 216: 47. 1966. Basionym: O. campestris (L.) de Candolle var. vat. minor A. Gray, Proc. Amer. Acad. 20: g. 1884. melanocephala Hooker Type: Clear Creek County, Colorado, C. C. Parry = O. maydelliana Trautvetter 991, 1861; lectotype (here designated by Welsh, cf. Barneby, Proc. Calif. Acad. Sci. IV, 27: 220. Oxytropis mazama St. John, Proc. BioI. Soc. Wash. 41: 1952) GH!; isolectotype NY! 101.1928. = O. multiceps Torrey & Gray Type: Washington, Goat Mts., Pierce County, O. D. OXljtropis nana NuUall ex Torrey & Gray, FI. N. Amer. Allen 6 July, 30 September 1896; holotype 245, 1: 340. 1838. WTC; isotype NY!, US!, CAS!, UC! Type: Plains of the Platte in the Rocky Mountain O. campestris (L.) de Candolle var. gracilis (A. Range [Wyoming], T. Nuttall s.n. 1834; holotype Nelson) Bameby PH!; isotype NY! Oxytropis mertensiana Turczaninow, Bull. Soc. Nat. Distribution: Drainage of the North Platte and Mosc., 1840:68. 1840. Cheyenne rivers, westward to the Wind River Type: "ad sinum Sancti Laurentii," Dr. Mertens; Mountains, Albany, Carbon, Converse, Fremont, holotype LE. Natrona, Platte, and Sweetwater counties, Distribution: Disjunct in E, N, W Alaska, and N Wyoming; endemic. Yukon; arctic Siberia and west to ca 60° E longi" This is a beautiful species ofclays, shales, and gravelly tude (Jurtsev 1986). bluffs and ridge tops endemic to Wyoming. This plantis easily distinguishedby its simple primary Barneby (1952)' postulated that it might have and trifoliolate secondary leaves in conjunction arisen through hybridization ofO. sencea and O. with few-flowered, densely black-villous inflores­ multiceps, a very likely supposition. Flowercolors cences. are variable in a given population from pale pinks through lavender and purpie, and white-flowered Oxytropis minor (A. Gray) Cockerell, Torteya 18: 180. 1918. . populations are known. A contribution £rpm O. lambertii is also suggested by the presence of Basionym: Oxytropis multiceps var. minorA. Gray incipiently malpighian hairs in some specimens. O. multiceps Torrey & Gray = The relationship to segregates ofO. besseyi seems Oxytropismollis Nuttall exA. Gray, Proc. Amer. Acad. 6: tenuous at best. The relegation ofO. nalla to that 235. 1864, synonym. species might require a realignment ofothet taxa = O. borealis de Candolle var. viscida (Nuttall) Welsh as well, including combination of lambertii, Oxytropis monticola A. Gray, Proc. ArneI'. Acad. 20: 6. sencea, campestris, and even multiceps. Such a 1884. proposal is, ofcourse, absurd. Taxonomy must be Type: Montana, Valley o£Iocko River, Lake County, both practical and reflect biological reality. W. M. Canby 91, 18 July' 1883; lectotype var. atgophylla (Rydbt1rg) Isely, Syst. Bot. 8: 425. (Bameby, Proc. Calif. Acad. Sci. IV, 27: 257. 1952) 1983. GH! Basionym: Aragallus argophyllus Rydberg = O. campestris (L.) de Candolle var. gracilis (A. = O. besseyi (Rydberg) Blankinship var. argophylla Nelson) Bameby (Rydberg) Isely This name is lectotyp$ed by Bameby (195g), who vat. besseyi (Rydberg) Isely, Syst. Bot. 8: 425. 1983. selected the type from among a Series of speci" Basionym: Aragallus besseyi Rydberg mens cited by Dr. Gray. The syntypes were a = O. besseyi (Rydberg) Blankinship var. besseyi mixed lot, with" C. C. Parry 87 being a putative hybrid between O. campestris var. gracilis and O. var. fallax (Barneby) Isely, Syst. Bot. 8: 425. 1983. borealis var. viscida, Jenney s.n. 1875 belonging BasionYm: O. besseyi (Rydberg) Blankinship var. to O. campestris var. dispar, and Lyall s.n. 186g fallax Bameby belonging to O. campestris var. cusickii. The lec­ var. obnapifotmis (C. L. Porter) Isely, Syst. Bot. 8: totypification was reviewed by Elisens and Packer 425.1983. (1982). Basionym: O. obnapiformis C. L. Porter 392 GREAT BASIN NATURALIST [Volume 51

var. salmonensis (Batneby) Isely, Syst. Bot. 8: 425. glabrous plants growing intermingled. And, it is 1983. with the often mixed variants that infraspecific Basionym: O. besseyi (Rydberg) Blankinship var. segregation has been attempted. The synonymy salmonensis Batneby is long, indicating the diversity of form but not var. ventosa (Greene) Isely, Syst. Bot. 8: 425. 1983. necessarily the presence ofseveral taxa within the Basionym: Aragallus ventosa Greene complex morphology ofthe species. ssp. pygrnaea (Pallas) Htilten, FI. Alaska & Yukon, O:tytTopis nigrescens (fallas) Fischer ex de CandoiIe, n02.1947 Prodr.2:278.1825. Basionym: Astragalus pygmaeus Pallas ssp. arctobia (Bunge) Hulten, Ark. Bot. (n.s.) 7: 80. = O. nigrescens (Pallas) Fischer V(lr. nigrescens 1968. var. pygmaea (Pallas) Chamisso, Linnaea 6: 546. Basionym: O. arctobia Bunge 183!. O. nigrescens (Pallas) Fischer val'. uniflora = Basionym: Astragalus pygmaeus Pallas (Hooker) Batneby = O. nigrescens (Pallas) Fischervar. nigrescens var. arctobia (Bunge) A. Gray, Proc. Amer. Acad. var. unijiora (Hooker) Barneby, Proc. Calif. Acad. 20:3.1884 IV, 27: 209. 1952. Basionym: O. arctobia Bunge Basionym: O. arctica 13 uniflora Hooker = O. nigrescens (Pallas) Fischer vat. uniflora Distribution: Islands of the Canadian Arctic Archi­ (Hooker) Bameby pelago and along the arctic coast from Mackenzie ssp. bryophita (Greene) HtIlteIi, FI. Alaska & Yukon, Delta E to Baffin Island and Hudson Bay, less nog, map 833. 1947. commonly in interior sites in Yukon. Basionym: Aragallus bryophilus Greene, Ptoc. BioI. This plant, as interpreted by me, is an American Soc. VVash. 18:17. 1905. (more precisely a Canadian) endemic, but there = O. nigrescens (Pallas) Fischer var. nigrescens are plants from the Chukotsk that match them val'. btyophila (Greeile) Lepllge, Amer. MidI. Nat. closely (see O. gorodkovii Jurtsev, a newname for 46:758.1952 O. pygmaea, a phase of var. nigrescens). The Basionym: Aragallus bryophilus Greene Siberian materials seem to representmere ecolog­ = O. nigrescens (Pallas) Fischervar. nigrescens ical variants within an expanded vat. nigrescens, however. var. bryophilaf. albida Lepage, Amer. MidI. Nat. 46: 758.1952. Oxytropis obnapiformis C. L. Porter, Madrono 9: 133, Type: Alaska, Kodiak, Old VVoman Mt., Lepage fig. 1947. 25089, 24 June 1949; holotype ?; isotype DAO!, Type: Colorado, in the sand hills 8-9 mi VV of May­ RIM!, fragment BRY! . bell, MOffat County, 5,900 ft., C. L. Porter 3864, = O. nigrescens (Pallas) Fischer vat. nigrescens 19 June 1946; holotype RM!; isotypes CHI, RSA, US! v~r. genuina 1<:jellnwn, Vega-Exped. Vetensk. Arb., O. besseyi (Rydberg) Blankinship var. obnapi­ 16. 1883, nomen. formis (C. L. Porter) VVelsh = O. nigrescens (Pallas) Fischer var. nigrescens Oxytropis okanoganea St. John, PrOc. BioI. Soc. VVash. Vllr. lonchopoda Barneby, Leaf!. VV. Bot. 10: 23. 41: 109. 1928. 1963. Type: VVashington, n.w. of Riverside, Okanogan Type: Yukon, Catheclral Rocks, Ogilvie llange, J. A. County, H. St. John 7728; holotype VVTC. Cll1der & J. M. Gillett 26013A, 29 June 1960; = O. campestris (L.) de Candolle var. gracilis holotype DAO!; isotype NY! (A. Nelson) Bameby Distribution: Ogilvie Mts., Yukon; endemic. The variety rests almost solely upon the pods being Oxytropis olympica St. John, Proc. BioI. Soc. VVash. 41: cIefinitely stipitate. Thereis considerable variation 103.1928. in stipe length through the range ofO. nigrescens Type: VVashington, Olympic Nits., Jefferson County, apart from the phase segregated as var. loncho­ B. Flett 134, 24 July 1897; holotype VVTC; isotype poda. The localization of an eiollgated stipe in US!, photo BRY! plants from the OgilVie Mts. might prove ulti­ O. campestris (L.) de Candolle var. gracilis mately to represent a mere continuum and lack (A. Nelson) Bameby value in designation ofa taxon. Plants from there OXytropis oreophila A. Gray, Proc. Amer. Acad. 20: 3. in vegetative condition are essentially like those of 1884. phases ofO. nigrescensfrom elsewhere. The elon­ Type: Utah, Aquarius Plateau, 10,500 ft., Garfield gate stipe and slightly inflated pods are remines­ County, L. F. VVard 541, 9 August 1875; lectotype centofO. podocarpa, but the leaflets are as in var. (here designated by VVelsh, cf. Barneby, Proc. nigrescens. Ca!i£ Acad. Sci. IV, 27: 214. 1952) GH!, isolecto­ val'. nigrescens type NY! Distribution: Amphiberingian,. coastal Yukon to var. jonesii C!3arneby) Barneby, Intermountain FI. Bering Strait, south to Kodiak, and in interior 3B: 183. 1989. Yukon and Alaska, N. VV. T. (Mackenzie Mts.), and Basionym: O. jonesii Bameby British Columbia; Siberia and Kamchatka. Distribution: Uinta Basin, VVasatch Plateau, and This is a highly variable taxon with green and silvery, Paunsagnnt and Markagunt plateaus, Utah; en~ loose and compact, villous to strigose or even demic. 1991] OXYTROPIS DC.-NAMES, BASIONYMS, TYPES, SYNONYMS 393

This is a caespitose, mat- or mound-forming plant, of folded, falcate leaflets are useful in distinguishing moderate elevations in mountain brush or pon­ this from closely related mat" or mound"forming derosapine zones. Its compact growth, large flow­ species, such as O. nigrescens, in vegetative con­ ers, and short peduncles are characteristic for the dition. variety. Itis more orless transitional with the next v!1r. in}lata (Hooker) Boivin, Naturaliste Canad. 94: variety. 78.1967. var. juniperina Welsh, Great Basin Nat. 38: 339. Basionym: O. arctica 1) inflata Hooker 1978. = O. podocarpaA. Gray Type: Utah, ca 1 mi E of Bicknell, Wayne County, Oxytropis pygmaea (:I:'aIlas) Fel"Ilald, Rhodora 30: 153. S. L. Welsh & G. Moore 13828, 1976; holotype 19~8 .. BRY!; isotype NY! Basionym: Astragalus pygmaeus Pallas Distribution: Nevada and Utah; endemic. = Oxytropis nigrescens (Pallas) Fischer var. nigres­ This ia a caespitose, mat- or mound-forming piant, of cens low-elevation, xeric sites. The compact growth, smallish flowers, and short peduncles are charac­ Oxytropis rettorsa fernald, Rhodora 30: 140. 1928. teristic for the variety. Type: Colorado, vicinity of Como [Park Co.], South Park, C. S. Crandall & J. S. Cowen 152, August 3, vat. oreophiltL 1895; holotype GH!; fsotype NY! Distribution: Arizona, California, Nevada, and Utah. = O. deflexa var. sericea Torr. &: Gray . This is the taller, more robust phase of the species with elongate scapes bearingclustered, bladdety­ vat. sericea (Torrey & Gray) Fernald, Rhodora 30: inflated pods, more typical of high elevations 140. 1998. ' where it often grows intermingled with O. parnJi. O. retrorsa ssp. sericea (Torrey & Gray) Love & Love, Taxon 31: 341. 1982. OXytropis parnJi A. Gray, Proc. Amer. Acad. 20: 4. 1884. = O. deflexa var. sericea Torrey & Gray Type: "Rocky Mountains ofnorthem New Mexico and OxytfOpis richardsonii (Hooker) K. Schumann, Just's Colorado, near the limit of trees" [Sangre de Bot. Jahresb. g1: 496. 1901. Cristo Pass, Taos Co., N.M.], C. C. Patry 41, Basionym: O. splendens 13 richardsonii Hooker 1861; lectotype (here designated by Welsh, cr. = O. splendens Douglas Barneby, Proc. CaIi£ Acad. Sci. IV, 27: 218. 1959) GHI. Rocky Mountains, E. Hall & H. Harbour; Oxytropis riplZria titvinov, Sched. Herb. FI. Ross. 6: 98. cotypes GR!, NY! 1908. . DistribUtion: Montana (?), Idaho, Wyoming, Califor­ Type: Russian Turkestan, from Farab (on the Amu nia, Nevada, Utah, Colorado, and New Mexico. Darya); holotype LE. Distribution: Idaho, Montana, North Dakota, and Oxytropis patens (Rydberg) A. Nelson, Univ. Wyo. Pub. Wyoming; introduced from central Asia. Bot. 1: 114.1926. This is a coarse perennial herb with the general aspect Basionym: Aragallus patens Rydberg of O. deflexa (Pallas) de Candolle. It grows in = Oxytropis lambertii Pursh var. bigelovii A. Gray riparian habitats where it is grazed by wildlife, Oxytfopis paysoniana A. Nelson, Univ. Wyo. Pub. Bot. especially by sage hens. The species is expected 1: 119. 1996. to spread in SUitable habitats through much of Type: Wyoming, Piney Mt., 25 miles west of Big the American West. Piney, Sublette County, E. & L. Payson g100, Oxytropis roaldii Ostenfeld, Vase. PI. Arctic N. Amer. July 12, 199Z; holotype RM!; isotypes NY!, US! GjoaExped., 54, pI. 3, fig. 16. 1910. = O. campestris (L.) de Candolle var. cusickii Type: Iter arcticus Roald Amundsen (Gjoa-Expedi" (Greenman) Barneby tion) 1903-1906. America Arct., Herschell Island, Oxytropis pinetorum (A. Heller) K. SchuIllann, Just's Lat N 69°35' Long. W 138°50', 13 July 1906, A. H. Bot. Jahtesb. 21: 496. 1901. Lindstrom s.n.; isotype NY! Basionym: Aragallus pinetorum A. Heller = O. campestris (L.) de Candolle var. roaldii (Osten­ = O. sericea Nuttall var. sericea feld) Welsh OXytropis plattensis Nuttall ex Torrey & Gray, FI. N. Oxytropis rydbetgii A. Nelson, Univ. Wyo. Pub. Bot. 1: Amer. 1: 340. 1838. 117.1926. ~om. Type: Platte Plains, Nuttall s.n.; isotype NY! nov. pro Aragallus alpicola Rydberg = O.lambertiivar.lambertii = O. campestris (L.) de Candolle var. cusickii (Greenman) Barneby OxytropispodocqrpaA. Gray, Proc. Amer. Acad. 6: 234. 1864. Oxytfopis saximontana (A. Nelson) A. Nelson, Univ. Type: Labrador (Schweinitz) and Alberta (Bourgeau); Wyo. Pub. Bot. 1: 113. 1926. cotypes GH! The specimens are cotypical, both Basionym: Aragallus saximontanus A. Nelson, nom. having been used by Gray in characterization of nov. Oxytropis lambertii Pursh var. ochroleuca the species. A. Nelson Distribution: Rocky alpine ridges and coastal shores, = Oxytropis sericeq Nuttall var. sericea in Colorado, Wyoming, Montana, Alberta, N.WT., Oxytropis scammaniana :Hulten, Ark. Bot. 33B: 4, figs. g, Ungava Peninsula, Labrador, and Baffin Island. 3 (map). 1947. The bladdery~inflatedstipitate pods are characteristic Type: Alaska, Eagle Summit, near Steese Highway of this and few other species of oxytrope. The 109 miles north ofFairbanks, elevation 3880 feet, 394 GREAT BASIN NATURALIST [Volume 51

E. Scamman 806, 7-11 July 1937; holotype S!; ssp. murrayi Jurtsev, Arctic FI. U.S.S.R. 9(2): 179. isotype GH!; photo BRYI 1986. Distribution: Alaska, Yukon, andN.W.T. (Mackenzie Type: Yukon, St. Elias Mts., Observation Mt. and Mts.); endemic. vicinity; at terminus ofJOxytropis sordida (Willdenow) Petsoon, Syn. PI. 2: 332. (Fernald) Barneby 1807. Oxyttopis tschuktschorum Jurtsev, Bot. Journ. 53: 11, Basionym: Astragalus sordidus Willdenow 1538. Hl68. ssp. batnebyana (Welsh)Jurtsev, Arctic Fl. U. S. S.R. = O. nigrescens (Pallas) Fischer var. nigrescens 9(2)= 109. 1986. Oxyttopis uralensis S arctica (R. Brown) Ledebour, FI. . Basionym: O. arctica R. Brown var. bamebyana Ross. 1: 594. 1842. Welsh Basionym: Oxytropis arctica R. Brown = O. arcticaR. Brown vat. bamebyana Welsh = O. arctica R. Brown var. arctica 1991] OXYTROPIS DC:-NAMES, BASIONYMS, TYPES, SYNONYMS 395

'Y minorHooker, FI. Bor.~Amer. 1: 146. 1834. Type: Yukon, Canol Rd.: Rose-Lapie R. Pass, mile Note: Barneby cites this name as problematical, to 105; schist mountain eastoflake. Elev. 4000, shaly be excluded because of inadequate typification. cliffs by waterfall, A. E. Porsild & A. l Breitung The Labrador specimens cited with the original 10918, July 19, 1944; holotype CAN; isotype ISCI, description are likely what now passes under O. GHI, Sf, US!, NY! campestris var. terrae-novae; the remainder is = O. borealis de Candolle vat. sulphurea (Porsild) citedfrom dryhills and prairies in the Rock Moun­ Welsh tains, and remains obscure. Physocalyxmulticeps Nuttallex Gray, Proc. Amer. Acad. f3 subsucculenta Hooker, FI. Bor.~Amer. 1: 146. 6: 234. 1864. 183l. Type: Rocky Mountains, T. Nuttall s.n., 1834; holo­ Type: "Arctic seashore, to the east of the Mackenzie type PHI; isotype NY! Riv.er," Dr. Richardson s.n.; holotype K. = Oxytropis multiceps Torrey & Gray = O. borealis de Candolle var. borealis Spiesia arctica (ll. Brown) Kuntze, Rev. Gen. 205. 189l. Oxytropis vamns (Rydberg) K. Schumann, Just's Bot. Basionym: Oxytropis arctica R. Brown var. arctica Jahresb. 29: 543.1903. = Oxytropis arctica R. Brown vat. arctica Basionym: Aragallus varians Rydberg = O. campestris (L.) de Candolle vat. varians (Ryd­ Spiesia arctobia (Bunge) Kuntze, Rev. Gen. 205. 1891. berg) Barneby Basionym: Oxytropis arctobia Bunge = O. nigrescens (Pallas) Fischer var. uniflora Oxytropis vegana (Cockerell) Wooton & Standley, Contt. (Hooker) Barneby U. S. Nat. Herb. 16: 136. 1913. Basionym: Aragallus pinetorumveganus Cockerell Spiesia bellii Britton ex Macotm, Can. Rec. Sc. 6: 148. = O. sericea Nuttall var. sericea 1894. Type: Canada, Digges Island, R. Bell s.n., 15 Sept. Oxytropis verruculosa Porsild, Bull. Nat. Mus. Canad. 1884; holotype CAN!; isotype NY!; US! 121: 246. 1951. = Oxytropis arctica R. Brown var. bellii (Britton) Type: Yukon: Canol Rd.: Rose-Lapie Pass, southwest Boivin ofgranite mountain west ofmile 116, alpine slopes from road to below summit. Elev. 5000 ft. rocky Spiesia inflata (Hooker) Britton, Mem. Torrey Bot. Club ledges, A. E. Porsild & A. J. Breitung 100'72, 5: 20l. 1894. 1 July 1944; holotype CANI; isotype GHI, SI, US! Basionym: Oxytropis arctica 1\ inflata Hooker = O. borealis de Candolle var. sulphurea (Porsild) = O. podocarpaA. Gray Welsh Spiesia lagopus (Nuttall) Kuntze, Rev. Gen., 206. 189l. Oxytropis vilZosa (Rydberg) :K. Schumann, Just's Bot. Basionym: Oxytropis lagopus Nuttall Jahresb. 29: 543. 1903. = Oxytropis lagopus Nuttall Basionym: Aragallus villosus Rydberg Spiesia lambertii (Pursh) Kuntze, Rev. Gen., 206. 189l. = Oxytropis campestris (L.) de Candolle var. gracilis Basionym: Oxytropis lambertii Pursh (A. Nelson) Batneby = Oxytropis lambertii Pursh var. lambertii Oxytropis viscida Nuttall ex Torrey & Gray, Flora N. var. sericea (Nuttall) Rydberg, FI. Neb. 21: 43, pI. Amer. 1: 34l. 1838. XI, figs. 107-109. 1895. Type: Rocky Mountains, near the sources ofthe Ore­ Basionym: Oxytropis sericea Nuttall gon [SW Wyoming], T. Nuttall s.n., 1834; syn­ = Oxytropis sericea Nuttall types NYI, PH; and "R. Mts. Oxyttopis mollis," GH! Spiesia mertensiana (Turczaninow) Kuntze, Rev. Gen., = O. borealis de Candolle var. viscida (Nuttall) Welsh 206. 189l. Basionym: Oxytropis mertensiana Turczaninow var. hudsonica (Greene) Barneby, Proc. Calif. Acad. = Oxytropis mertensiana Turczaninow IV, 2,7: 245. 1952. Basionym: Aragallus hudsonicus Greene Spiesia monticola (A. Gray) Kuntze, Rev. Gen., 206. 189l. ssp. hudsonica (Greene) Love & Love, Taxon 31: 347. Basionym: Oxytropis monticola A. Gray 1982. = Oxytropis campestris (L.) de Candolle var. gracilis BasionYm: Aragallus hudsonicus Greene (A. Nelson) Barneby var. subsucculenta (Hooker) Barneby,Proc. Calif. Spiesia multiceps (Torrey & Gray) Kuntze, Rev. Gen., Acad. IV, 27: 246. 1952. 206. 189l. Basionym: Oxytropis uralensis f3 subsucculenta Basionym: Oxytropis multiceps Torrey & Gray Hooker = Oxytropis multiceps Torrey & Gray var. viscida Spiesia nana (Nuttall) Kuntze, Rev. Gen., 206. 189l. Oxytropis viscidula (Rydberg) Tidesttom, Contt. U.S. Basionym: Oxytropis nana Nuttall Nat. Herb. 25: 332.1925. = Oxytropis nana Nuttall Basionym: Aragallus viscidulus Rydberg Spiesia nigrescens (Pallas) Kuntze, Rev. Gen., 206. 189l. = O. borealis de Candolle var. viscida (Nuttall) Welsh Basionym: Astragalus nigrescens Pallas ssp. sulphurea l'orsild, Bull. Nat. Mus. Canad. 121: = Oxytropis nigrescens (Pallas) Fischer var. nigres­ 2,47. 1951. cens 396 GREAT BASIN NATURALIST [VolumeSl

Spiesia oreojJhila (A. Gray) I<:tintze, Rev. Gen., 206. ELISENS. W. J., AND J. G. PACKER. 1980. A contribution'to 1891. the taxonomy ofthe Oxytropis campestris complex Basionym: Oxytropis oreophila A. Gray innorthwestern North America. Canadian Journal = Oxytropis oreophila A. Gray ofBotany 58: 1820-183l. Cana~ Spiesia parryi (A. Gray) KUIitze, R.ev. Gen., 206. 189l. =. 1982. Typification of Oxytropis monticola. dian Journal ofBotany 60: 1580-1582. Basionym: Oxytropis parryi A. Gray GRAY, A. 1884. A revision ofthe North American species = Oxytropis parryi A. Gray of Oxytropis DC. Proceedings of the American Spiesia podocarpa (A. Gray) Kuntze, Rev. Gen., 206. Academy gO: 1-7. 189l. JURTSEV, B. A. 1986. Oxytropis DC. Pages 61-146 in Basionym: Oxytropis podocarpa A. Gray A. A. Korobkov, M. V. Sokolova, N. N. Taraskina, = Oxytropis podocarpa A. Gray and B. A. Jurtsev, Arctic flora U.S.S.R 9(2). Spiesia splendens (Douglas) Kuntze, Rev. Gen., 207. 187pp. 189l. PORSILD, A. E., AND W. J. CODY. 1980. Oxytropis DC. Basionym: Oxytropis splendens Douglas Pages 438-442 in Vascular plants ofthe continen" = Oxytropis splendens Douglas tal Northwest Territories, Canada. National Museum ofNatural Sciences, National Museums Spiesia viscida (Nuttall) Kuntze, Rev. Gen., 206. 189l. ofCanada. 667 pp. Basionym: Oxytropis viscida Nuttall VASIL'CHENKO, 1. T., B. A. FEDCHENKO. AND B. K. = Oxytropis borealis de Candolle var. viscida (Nut" SHISHKIN. 1948. Oxytropis DC. in Flora U.S.S.R tall) Welsh 13: 1-229. WELSH, S. L. 1967. Legumes ofAlaska II: Oxytropis DC. LITERATURE CITED Iowa State Journal ofScience 41: 277-303. ~. 1974. Oxytropis DC. Pages 215-282inAnderson's BARNEBY, R. C. 1952. A revision of the North American flora of Alaska and adjacent parts of Canada. species ofOxytropis DC. Proceedings ofthe Cali" Brigham Young Press, Provo, Utah. 724 pp. forniaAcademy6fScience IV, 21: 177-312. ~. 1977. On the typification of Oxytropis leucantha BONIN, B. 1962. Etudes sur les Oxytropis DC. I. (Pallas) Pers. Taxon 21: 155-157. Oxytropis dejlexa (Pallas) DC. Svensk Botaniska ~. 1981. Oxytropis DC. Pages .396-398 in S. L. Tidskrift 56: 496~500. Welsh, N. D. Atwood, S. Goodrich, and L. C. __. 1967. Etudes surles Oxytropis DC. II. Naturaliste Higgins, A Utah flora. Great Basin Naturalist Canadiene 94: 73-78. Memoirs 9. 984 pp. BUNGE, A. 1874. Species Generis Oxytropis, DC. Mem" oires l'Academie Imperiale des Sciences de St. Received 19 August1991 PetersbourgVII, 22(1): 1-166. Accepted15 October1991