Utah Plant Types—Historical Perspective 1840 to 1981—Annotated List, and Bibliography

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Great Basin Naturalist

Volume 42 Number 2 Article 1

6-30-1982

Utah plant types—historical perspective 1840 to 1981—annotated list, and bibliography

Stanley L. Welsh Brigham Young University

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Recommended Citation Welsh, Stanley L. (1982) "Utah plant types—historical perspective 1840 to 1981—annotated list, and bibliography," Great Basin Naturalist: Vol. 42 : No. 2 , Article 1. Available at: https://scholarsarchive.byu.edu/gbn/vol42/iss2/1

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The Great Basiii Naturalist Published at Provo, Utah, by Brigham Young University

ISSN 0017-3614

Volume 42 June 30, 1982 No. 2

UTAH PLANT TYPES-HISTORICAL PERSPECTIVE 1840 TO 1981- ANNOTATED LIST, AND BIBLIOGRAPHY

St;uiley L. Welsh'

.\bstract.— Reviewed are the 144 collectors and 167 authors of 1073 Utali vascular plant types for the period from 1840 to 1981. Historical perspective yields evidence of geography of collection activities by botanists who pene- trated the boundaries of the state, and of shifting centers of emphasis in the study of Utah plants from a classical taxonomic standpoint. Philosophy and influence of contemporary authors and collectors is discussed as they affect tax- onomv of Utah plants in our first 142 years. A short biographical account of Marcus Eugene Jones and his interaction with other American botanists is included. The annotated checklist includes bibliographical citations, type locality data, collector, and place of deposition.

Acknowledgments Kaye Thorne, my colleague at the herba-

It is impossible to complete a work as rium of Brigham Young University, gave en- complex and tedious as this without the help couragement and help at all times. Mark L. of otiiers. I thank the curators of the herbaria Gabel aided me with research on the life of hospitality will- visited by me for their and Charles Christopher Parry. To him I am ingness to help in this work. They are as fol- grateful. lows: Elizabeth McClintock and John To Mr. Charles Walker, archivist for the Thomas Howell (CAS); Robert Thorne (RSA); family of Publius Virgilius and Lavinia (Bur- Ronald Hartman (RM); Patricia K. Holmgren ton) I am grateful for the access he and Rupert C. Bameby (NY); George Russell Jones, to a published family history in- (US); Lois Amow and Beverly Albee (UT); provided me cluding accounts of Marcus E. Jones and his Mary Barkworth (UTC); Theodore J. Cro- vello (NDG); and Richard Pohl and Duane wife, Anna Elizabeth Richardson. Also, he

Isely (ISC). I am indebted to George Knaphus provided me original letters written by both and Lois Tiffany who made it possible for me of them, and by a nephew of Jones, Arthur J. to visit the site of Jones Grove, where Marcus Jones, who accompanied him on attenuated E. Jones lived with his family near Grinnell, field trips by wagon. These have given me in- Iowa. Acknowledged also is the help of Mrs. sight into Jones's family life and have allowed Ralph Fleener, who resides currently on the me to present a portion of the life-style of his former Jones's property, for the checking of wife and family during his intensely goal- records and dates on markers in the cemetery oriented life. I shall be forever indebted to at Grinnell. Mr. Walker and his family for this insight.

'Life Science Museum and Department of Botany and Range Science, Brigham Young University, Provo, Utah 84602.

129 130 Great Basin Naturalist Vol. 42, No. 2

Introduction I have tried to avoid inclusion of obvious paratypes. These are specimens cited with All taxonomists, and others who work with the original description, but they are not du- plant names, have been hampered by lack of plicates of the holotype. Mostly paratypes a summary treatment of Utah plant types, do, indeed, represent the same taxon as the their collectors, authors, and places of deposi- holotype or lectotype. They are the next tion. The specimens serve to typify the most important elements in typification; but names applied to them by the authors who sometimes they represent extraneous mate- named and described them. The types are the rials, and might belong to quite different name-bearing specimens and are in- taxa. The main reason for not citing them dispensible in interpretation of taxonomic here involves the endless listing of specimens. hmits and concepts; they are necessary for They are very numerous, and will be most nomenclatural considerations. important to those undertaking revisions and There are several kinds of types, but only monographs. holotypes and their duplicates, the isotypes, Information has been gathered from exam- have been included uniformly in the follow- ination of type collections at the U.S. Nation- ing treatment. I have not tried to distinguish al (US, 500 Rocky Mountain holotypes from isotypes in the designations of Museum + );

Herbarium (RM, 150 + ); California Academy place of deposition; that is a taxonomic con- of Science (CAS, 120+ York Botanical sideration which is left to the monographer. ); New In many instances the name bearing speci- Garden (NY, 550 + ); Rancho Santa Ana mens were not designated as types by the au- (POM, 240 + ); Iowa State University (ISC, ca thors and I have followed tradition in citing 100); University of Utah (UT, 100+ ); Utah the original materials as types. Many of the State University (UTC, 150 + ) and Brigham older names have been subjected to lecto- Young University (BRY, 250 + ). Only se- typification. Where this has been obvious, lected material has been examined from Gray mainly in taxonomic revisions or as indicated Herbarium (GH), Carnegie Museum (CM), on some herbarium annotations, I have fol- University of California, Berkeley (UC), lowed that usage. Lectotypification, how- Notre Dame University (NDG), and Missouri ever, is subject to interpretation, and the Botanical Garden (MO). Information on types specimens cited might ultimately be excluded at the Greene Herbarium at Notre Dame from consideration as lectotype or isolecto- University was taken from a computer print- type. Too, some materials have been cited as out provided through the generosity of Pro- cotypes or syntypes. fessor Crovello. Literature has When a portion of the Theodore J. original collection cited is from Utah, I have been reviewed for most of the citations, but included that material as a type. It might some have been unavailable. Other citations have been designated at a lower level, i.e., as still await procedures entailed through inter- paratype, by some author or authors un- library loans. known to me. Type locality information has been abbre-

Designation of types is a function of rules viated in some instances to conform to the established mainly in the 20th-century, and computer format. Localities of deposition, or early authors were not impressed with the supposed deposition, are indicated by the necessity of typification. Mention of speci- standard abbreviations. Where I have seen mens to vouch for newly described taxa was the type, the abbreviation is followed by an often only incidental. Most of the types serv- exclamation mark (e.g., US!). Where the ab- ing as bases for taxa described in the 19th- breviation stands alone, the place of deposi- century are now interpreted through lecto- tion is taken from literature, or from infer- typification. Purists often are involved in ence with other collections or names polemical stRiggles over selection of the cor- published by the same author, but the speci- rect portion of the specimens cited (or not men has not been seen by me. cited) as the lectotype. I have not searched Although portions of more than two dec- all the literature in which lectotypes have ades have passed since this work was begun, been designated; that is the role of the the data are still incomplete for many of the monographer. plant names. I apologize for that lack of June 1982 Welsh: Utah Plant Types 131 completeness, and for the many errors that and others have undergone name changes. will become apparent as one uses the list. Present knowledge of distribution allows for Some of the problems are apparently in- accurate placement, at least to county level, soluble. Plant collectors and authors seldom of some materials which were provided with have been instilled with sufficient foresight vague or misleading locality information. to supply all the information necessary for Problems associated with collections of adequate intrepretation of specimens, local- certain individuals will be discussed in the

ities, dates, and other data. Brevity was con- historical review presented below. sidered to be a virtue by the collectors (along It is believed that the shortfalls in data do widi an almost impossible, if unstated, belief not obscure the basic picture of plant explor- that others would know what they knew), ation in Utah or of the story of publication of types. This attempt at even when that brevity left out such informa- taxa based on Utah analysis of type data in no way detracts from tion as state, locality within the state (or ter- the larger scene of plant studies which re- ritory), political subdivision of the state, and sulted from the naming of plants based on any other data. Herbarium labels are the types from other areas. The flora of Utah is sources of infonnation provided by plant col- replete with names of plants with circum- lectors, and they should have contained cer- boreal and other extra Utah representation tain basic data. Some of the deficiency, such dating to 18th- and early 19th-century collec- as lack of place names and surveys, is not the tors and authors. fault of the early workers. Boundaries of the Acknowledged also are the contributions state or territory of Utah were unknown until to botany of workers whose main thrust did long after the final reduction of the original not include classical taxonomy. Numerous Territory of Deseret to its present size in botanists have contributed in ecology, phys- 1868. When one went south from St. George, iology, anatomy, morphology, genetics, and he was never quite sure when he had entered other sophisticated fields of endeavor. Each into Arizona. There is little excuse, however, of those areas has allowed for a greater un- for the Palmer and Parry labels with only derstanding of the taxonomy of the state. "Southern Utah, Northern Arizona, &c." or Some investigators contributed to more than "Southern Utah, &c." as the only locality one of the major botanical disciplines, or to instances the localities were data. In some taxonomy of nonvascular plants, not treated to the collectors, whose passing known only herein. Their exclusion from the lists of col- of the possibility of ever has denied us lectors and authors, or their low placement in knowing. the lists does not imply that the contributions data were provided on labels, Even when are not great. The measure herein is based on published the the author who subsequently contributions in the strictly classical sense. name of the plant often failed to cite it. Neither should naming and description of publications, and some of Many of the early plants be expected to continue indefinitely. the later ones, convey little in the way of ac- But, the botany of the state is not yet eluci- ciu^ate data on collector, number, date, local- dated, and much further work is indicated. specimens. ity, or place of deposition of the This work should not be thought of as repre- taken obscure col- Some specimens were by senting the final word; certainly many cita- the locality of collection, dates, lectors, and tions of Utah types have been overlooked. and other information remains in doubt. Localities and dates for some of the plant names have been interpolated by reference Collectors of Utah Types with other collections. This is especially the case with county designations; boundaries The number of vascular plant types report- have changed and new counties have been ed for Utah between 1840 and 1981 is ap- created since the earliest collections were proximately 1073. Exact numbers are impos- made, and early collectors were not sure sible due to such equivocal label data as which county they were working in when "Southern Utah, Northern Arizona, &c." for some collections were made. Some pioneer some collections. The total number that towns and villages have passed into history. might be in error is thought to be small, and 132 Great Basin Naturalist Vol. 42, No. 2

for the purposes of this paper they seem not Table 1 continued. to affect the overall picture. The types have Number been taken by 144 collectors acting singly or Years Collectors of types active in combination with one or more collabora-

tors (Table 1). Possibly other collectors are obscured by citations where collection data are lacking. Authors include some 167 individuals acting singly or in combination, and will be discussed later. A summary of collections of Utah plant types by decade is presented in Table 2. Earliest types taken from Utah were apparently those by John Charles Fremont in 1843 [Perchiton occidentale Torr. & Frem. = Atriplex canescens (Pursh) Nutt.] on his first expedition in the American West, wherein he explored the vicinity of the Great Salt Lake. The types of Gilia stenothijrsa T. & G. and Senecio multilobatus T. & G. were collected by Fremont apparently during 1845 in the Uinta Basin. The 1850s were marked by the efforts of Captain Howard Stansbury, F. Creutzfeldt, Henry Engelmann, and Edward Griffin Beckwith. Stansbury explored the Great Salt Lake in 1850; Creutzfeldt and Beckwith were associated with the ill-fated Gunnison expedition of 1853-54. Creutzfeldt was the botanist for the expedition, whose duty was to survey a route for a railroad to the Pacific (Beckwith 1854), and he was killed along with John Williams Gminison and most of the others of the party in an am- bush by Indians near Delta on 26 October 1853. Beckwith, who was leading another portion of the expedition at the time of the disaster, was named to succeed Gunnison. Creutzfeldt had collected types of two

Table 1. Collectors of Utah plant types.

Number Years Collectors of types active

Al-Shehbaz, I.A. (b.?) June 1982 Welsh: Utah Plant Types 133

Table 1 continued. Table 1 continued.

Number Years Collectors of types active

Holmgren, N.H. (b.l937) &P.K. Holmgren (b. 1940) 4 Holmgren, N.H. (b. 1937) &J.L. Reveal (b. 1941) Huffman

Johnson, J.E. (1817-1882) Jones, M.E. (1852-1934) Kearney, T.H. (1874-1956) & H.L. Shantz (1867-1958) Klein, W. (b.?) Kuntze, C.E.O. (1843-1907) Larson Leonard, F.E. (1866-1922) Linford, J.H. (b.l863) Maguire, B. (b.l904) Maguire, B. (b.l904) & H.L. Blood (b.?) Maguire, B. (b.l904) & A.H. Holmgren (b.l912) Maguire, B. (b.l904) & R. Maguire (b.?) Maguire, B^ (b.l904) et al. Marsh Mathias, M.E. (b.l906) McKelvey, S.D. (b.l883)

Mulford,'A.L (b.?) Neese, E. (b.l934)

& S. Peterson (b.?) Neese, E.(b. 1934) & S.L. Welsh (b.l928) Neese, E.(b. 1934) &S. White (b. 1955) Nelson, A. (1859-1952)

Newberry, J.S. (1822-1892) Nord & Sargent Osterhout, G.E. (1858-1937) Ostler, W.K.(b. 1951) &D.C. Anderson (b. 1948) Ownbey, G.B. (b.l916) Palmer^ E. (1831-1911) Pammel, L.H. (1862-1931) & R.E. Blackwood (b,?) Parry, C.C. (1823-1890) Payson, E.B. (1893-1927)

Peabodv, F.J. (b.l948) Pennell', F.W. (1886-1952) Pilsbry Plummer, P. (b.l911) Porter, C.L. (b.l889) Purpus,C. A. (1851-1941) Reading Reveal,"j.L. (b.l941) Reveal, J.L.(b. 1941) & G. Davidse (b.?)

Reveal, J.L. (b.l941) & N.H. Holmgren (b.l937)

Reveal, J.L. (b. 1941) & C. Reveal (b.?) 134 Great Basin Naturalist Vol. 42, No. 2

Eriogonum taxa in the Green River vicinity. Beckwith took the type of a handsome Astra- galus, which bears his name. Several taxa were later dedicated to the memory of Gun- nison, but unfortunately none of our Utah materials bears the name of Creutzfeldt. Eng- elmann was meteorologist and botanist with the Simpson expedition (Ewan 1950) and took the type of Echinocactus whipplei var. spinosior Englemann (named by his brother George Engelmann) somewhere west of Camp Floyd in August 1859. The first woman collector of Utah types appeared on the scene in the 1850s. Jane Carrington (a Mormon lady) collected some 59 species in the basin of the Great Salt Lake in 1857 (Reveal 1972). She was apparently the first woman to collect plants in the territory since the arrival of the Mormon pio- neers, a decade earlier. Two of her collections are the basis of types. They were named by Elias Durand (1860) as Erysimum asperum var. purshii and Acerates decumhens. Perhaps this person is Jane Maria Carrington (26 Feb- ruary 1840, Wiota, Wisconsin), daughter of Albert Carrington and Rhoda Maria Woods, who married Brigham Young, Jr. in March of 1857. Albert Carrington was prominent in early day affairs of the territory of Deseret, serving as editor of the Deseret News (Cool- ey 1980). Albert Carrington accompanied Stansbury on his exploration of the Great Salt

Table 2. Decade of collection of Utah types, number of collectors,and average number of types per collector. June 1982 Welsh: Utah Plant Types 135

Parry, whose collections were taken astride City and Beaver. His host in St. George was the Utah-Arizona border, which was not sur- Joseph Ellis Johnson (Jones 1930, personal veyed until later. correspondence of Parry ISC), pioneer pub- Collectors in the 1870s include Joseph Ellis lisher, herbalist, and horticulturalist (Cham- Johnson, Edward Palmer, Mrs. Ellen Powell berlain 1950). The following year Parry Thompson (sister of John Wesley Powell), Lt. worked out of Spring Lake (Utah County), George Montague Wheeler, Captain Francis where he was the guest of Benjamin Franklin Marion Bishop, Charles Christopher Parry, Johnson, brother of Joseph Ellis Johnson. His Lester Frank Ward, and finally, in 1879, objective for that year was to obtain a collec- Marcus Eugene Jones, whose appearance on tion of the flora in the vicinity of Mt. Nebo, the scene presaged a lifetime as resident bot- whose flora he decided was depauperate 1950). Parry was an ac- anist. Jones was to dominate Utah plant tax- (Parry 1878, Ewan of Torrey, and onomy until 1923, a total of 44 years. Palmer quaintance Asa Gray, John corresponded with and Parry concentrated their efforts in south- George Engelmann. He each of them and sent his specimens to them western Utah, especially in what is now for determination and naming. His contribu- Washington County. Palmer visited the area tions to the understanding of Utah plants are twice, in 1870 and 1877, and took a total of significant. 58 specimens later designated as types. He Edward Palmer was born at Wilton, Coun- was a collector only, depending on others to ty Norfolk, England on 12 January 1831 and name the plants he sampled in Utah. Type died on 10 April 1911 in Washington, D.C. collections taken by Parry were mainly from (McVaugh 1956). He came to America in his 1874 trip. Parry was in central Utah in 1849 where he settled in Cleveland, Ohio. 1875, and again in 1877 (with Palmer) at St. After attending classes at Homeopathic Col- George, but these trips added little to his lege in Cleveland for a few months during type list for Utah. The total number of his the winter of 1856-57, Palmer moved to types is 38, and he was responsible for pub- Highland, Kansas— to practice medicine. He lishing at least 2 of those. According to Jones assumed the title of doctor, which he was to (Contr. W. Bot. 17: 3. 1930), Parry was in the carry the remainder of his life. Previously he St. George vicinity for only about a month. had been on an expedition to South America, His herbarium is at Iowa State University, and contains nvmierous collections by Palmer along with his own materials (Gabel 1981). Charles Christopher Parry was bom in Ad- mington, Gloucestershire, England, 28 Au- gust 1823, and died at Davenport, Iowa, 20 February 1890. He moved with his family to America in 1832, where they settled in Washington County, New York. He received the degree of Doctor of Medicine from Co- lumbia College in 1846, where he was influenced by Dr. John Torrey, professor at that institution (Preston 1897, Gabel 1981). Parry moved to Davenport, Iowa, in 1846. He practiced medicine for a few months only before turning to his greater interest of botany. Following a career exploring the West with various governmental surveys, the pur- suit of natural history led Parry to Utah. He came to Utah on a private trip, not associated with any of the surveys. Contrary to Jones's statement (see above). Fig. 1. Marcus Eugene Jones (1897), principal pioneer Parry was in St. George from early April to plant taxonomist in Utah (photo courtesy of the Jones June of 1874, and then moved north to Cedar family archivist, Charles Walker). 136 Great Basin Naturalist Vol. 42, No. 2

and was soon involved in other expeditions and he was badly wounded at the battle of where he collected plant, animal, and arch- Fredericksburg. He was discharged with the eological materials. In 1870 he arrived by rail rank of captain, a title which he carried for at Salt City, Lake where he heard a speech the remainder of his life. After the war he at- by Brigham Young, was introduced to him, tended Illinois Wesleyan University, where and left for southwestern Utah carrying a let- one of his instructors was Major John Wesley ter of introduction provided by Brigham Powell, whose second expedition he joined as Young. He stayed in St. George and vicinity topographer. Following his work with the for about 10 days prior to departing with a Powell survey, Bishop joined the faculty of party delivering a threshing machine to St. the University of Deseret as professor of sci- Thomas, Nevada. They left St. George on 17 ence in 1873, where he served until 1877 June 1870, and spent at least a day and a half (Chamberlain 1950). The delicately beautiful beyond that date still in Utah. He noted the Astragalus episcopus Wats, is named in his grand sight of the Joshua tree forest on the honor. Beaverdam Slope. His total collections in Ward evidently centered his efforts in the Utah for that year is thought to be very vicinity of the Aquarius Plateau and Rabbit small, for he took only about 200 plants total Valley, working out of the hamlet of Glen- for the latter part of 1869 and all of 1870 wood, Sevier County. The handsome milk- (McVaugh 1956). Only seven of the 58 Utah vetch with diaphanous inflated pods. Astra- types taken by him were collected in 1870. galus wardii, was named in his honor by Palmer was an acquaintance of C. C. Par- Gray, and the beautiful low twinpod, Les- ry, and Parry helped with distribution of the querella wardii, was named by Watson. His Palmer material. In 1876 Palmer was em- locality information is much better than pro- ployed by the Peabody to Museum do some vided by any of his contemporaries, bu t there archeological excavations in southern Utah. has been evident confusion asssociated with He was at St. George from December 1876 his labels (Pennell 1920). He was a man of to June of 1877. During June he moved his many interests, with expertise in geology, pa- base of operations to Red Creek (Paragonah) leobotany, botany, and sociology. Ward pub- where he worked in Bear Valley in early lished papers in all of those subjects. For a July, then on to Beaver, and finally to Spring time he was a clerk and librarian at the U.S. Lake. The number of plants taken in Utah in Bureau of Statistics. His collections in Utah 1877 numbered around 500 (McVaugh 1956), are from the vicinity of Glenwood, Twelve and it was from this series that the bulk of his Mile Canyon, Rabbit Valley, Dirty Devil Utah types were named. Palmer depended on River, The Button, and Aquarius Plateau others to name his plants. (Ewan 1950). Mrs. Thompson, Captain Bishop, and Les- Others who collected type materials in ter Ward were associated with the Powell Utah in the 1870s were Lt. George M. Whee- surveys of the early-day geological survey. ler, Andrew Lafayette Siler, and Townshend They worked in southwestern, south central, Stith Brandegee. The collections of Wheeler and central Utah. Mrs. Thompson lived for a are treated, at least in part, in his publication while at Kanab in 1872, and her collections in the United States Geographical Surveys are from that period. Bishop's collections are west of the 100th Meridian (Wheeler 1878). from 1872 and 1873, but with little if any Wheeler had little sense of history, as in- provenience data. Some materials credited to dicated in the total lack of geographical in- Mrs. Thompson obviously came from some formation about plants he collected. Appar- distance away from Kanab and were prob- ently he took some plants in Salt Lake ably taken by members of the Powell survey, County, as indicated in a letter by Marcus E. including her husband, Almon Harris Jones, dated "8. 1. 1904" (on file at BRY), in Thompson. which he intimated that Wheeler, too, had Bishop was born in New York. He enlisted collected CoUomia dehilis in Salt Lake Coun- in the Union army at the age of 17. He was ty, but without recording the locality. Bran- made second lieutenant at the battle of Bull degee worked with one of the government Rim first lieutenant at the battle of Antietam, surveys, mainly in present-day San Juan June 1982 Welsh: Utah Plant Types 137

County. Siler, a Mormon rancher, lived at a The greatest era for collection of Utah place called "Ranch" near Kanab and took plant types occurred in the 1890s. Marcus E. specimens in Kane and adjacent Mohave (in Jones was the main contributor of type mate- Arizona) coimties. He was much interested in rials. Of the 259 types taken during the dec- cacti and sent a collection of a low de- ade, Jones was responsible for 193 of them. pressed-hemispheric ball-cactus to Engel- Jones was employed in 1894 by the U.S. De- mann, which was later named by Britton and partment of Agriculture as "Special Field was able to spend Rose as Utahia sileri. Jones (Contr. W. Bot. Agent" (Jones 1895). He the entire field season collecting plants. Ac- 16: 46-47. 1930) says of Siler: "He was about cording to his own account (see Leafl. W. seven feet tall; and slim as a rail, and wore Bot. 10: 208-217. 1965), Jones outfitted a about a No. 14 shoe. He was awkward and team and buggy that he sent south in early uncouth, but a real man." Unfortimately, the March with a young man as driver. The driv- type of the genus commemorating the state er and outfit proceeded to Holden, where of Utah was collected in the vicinity of Pipe they were trapped by a late winter storm. Springs, in Arizona, and that generic name is Jones traveled by train to Oasis, where he now synonymized within an expanded Pedio- hired a team and reached Holden in a bliz- cactus. Siler's collections are mainly at GH zard. Jones's account is as follows: "The next (Ewan 1950). and MO morning we drove out into the snow and toil- 1880s were relatively quiescent when The ed all day in a cloudless sky with the sun daz- record- compared to the efforts and successes zling in our faces and by night our faces were ed in the statistics of the 1870s. Together, M. burnt to a blister and the serum was oozing E. Jones, Fred Eugene Leonard, Samuel Mills off our chins. That night we camped on the Tracy (alone and with someone named desert in a foot of spotless snow where there Evans), and William Trelease collected some was hardly enough brush to make a fire and 54 types. Jones worked in Washington Coun- no water but snow. A better bed I never slept ty in 1880, and widely in Utah in later years of the decade. Leonard took specimens in northern Utah, especially in Salt Lake Comi- ty. His collections were sent east, where they were named later by Rydberg. William Tre- lease stopped briefly at Helper in 1889, in transit to St. Louis, Missouri, after spending the summer with the Harriman Alaska Expe- dition. He collected the type of a yucca that he named in honor of Mrs. Harriman. Jones moved to Utah in 1880, to Salt Lake City, where he was to reside imtil 1923. At

first he taught in the old Salt Lake Academy, a congregational school. From 1881 to 1885 he conducted a private high school. He was one of the early members of the Utah Natural History Society, of which he was president in 1899, and a fellow of the AAAS. Jones was cm'ator of the museum and librarian at the University of Utah for the year 1889-90. His large herbarium of western plants was stored for a time at the University of Utah (Cham-

berlain 1950), and it was from there that the collection was retrieved by Dr. P. A. Munz Fig. 2. Alice Eastwood and John Thomas Howell and taken to Pomona College in the 1920s (1936), near Yakima Park, Mt. Ranier, contributors to (Richard Shaw, personal communication understanding of botany in Utah and the West (photo by 1982). Edith Hardin English, courtesy of John Thomas Howell). 138 Great Basin Naturalist Vol. 42, No. 2

on. The next morning it was bitterly cold to svmimer of 1895. These trips allowed her ac- get out of bed and get a fire started in that cess to some of the most remote and rugged snow, but warm tea soon thawed us out. That country in the southeastern portion of the night we slept in mud near Black Rock and state. She was able to collect numerous plants the next day were at Milford. Our faces were on both occasions, among them the types of now all scabs and sores and we looked as some of our most unusual plant taxa. Her in- though we were suffering from some loath- terest in the region was to continue until her some disease." Jones was to be in the field, death in 1953 a tthe age of 94. Her final trip except for short pauses in Salt Lake City, un- to the state (with John Thomas Howell) was til the 18th of October. During the 1894 field in 1941. Type collections taken by her in season he collected in Washington, Kane, Utah span 50 years, longer than for any other Garfield, Piute, Sevier, Emery, Carbon (then contributor. On at leas t the 1895 expedition a part of Emery County), Utah, Sanpete, and she was accompanied by B. Alfred Wetherill, Wayne counties. His collections amounted to famous for his part in the discovery of the 4060 numbers, a total of about 35,000 speci- Pueblo dwellings at Mesa Verde, and later mens. Never again would anyone have such pioneer archaeologist in the region. He was an opportunity. to send some few things to Miss Eastwood, The next most important collector of the and this accounts for his inclusion on the list decade was Alice Eastwood. Born in Canada of Utah plant type collectors. in 1859, she moved to Denver, Colorado, in Two other important figures in Utah plant 1880, where she taught high school for sever- taxomomy appear among the cast of collec- al years. She was able to devote her life to tors in the 1890s. They are Per Axel Rydberg, tlie study of plants by living initially on the a young immigrant from Westergoethland, proceeds gained through investment. In 1892 Sweden, and Aven Nelson, natural history she traveled from Thompson's Springs (pres- professor at the infant University of Wyom- ent day Tliompson) in Grand County south to ing. Rydberg had arrived in the United States San Juan County. A second trip was under- in 1882 and was about to complete his docto- taken from Mancos, Colorado, west to Wil- rate at the University of Nebraska, where he low Creek in San Juan County, Utah, in the wrote a conservative treatment of the Resales of Nebraska. Later he would contribute numerous critical publications of the Rocky Mountain flora, and would have an overwhelming role in the botany of the West. His ^:^> trip to Utah in 1895 seems not to have been long, and he picked only a few things, mainly in the vicinity of Logan. More will be said of him in later parts of this paper. Nelson had only recently arrived at Laramie (in 1887), where he had accepted a position to teach English, but upon his arrival there found that another English teacher had been hired. He was offered the chair in natural history instead, and was to contribute to knowledge of western American botany for more than five decades.

Completing the list of collectors of types for the decade were Carl Albert Purpus (a German), James Henry Linford (a Utah native), and A. Isabel Mulford, a student of Fig. 3. John Torrey (1868), author with John Charles William Trelease at the Missouri Botanical Fremont and Asa Gray of specimens taken in Utah bv Garden (Ewan 1950). Purpus collected in the Fremont and by Captain Howard Stansbury, teacher of Charles Christopher Parry, and correspondent of Parry, La Sal Mountains and in southwestern Utah. Palmer, and others (photo courtesy of Richard \V. Pohl). Linford's collections are from the northern June 1982 Welsh: Utah Plant Types 139

Wasatch Mountains, chiefly from Logan The 1920s were a time of transition. Utah Canyon, and the place of deposition is un- was involved in the consequences resulting known. Mulford traveled through the central from World War I, and the field botanists of and northern regions. the early part of the century were involved The period from 1900 to 1909 was marked in activities that did not include active field by a decline in types collected. The total investigations. Jones sold his collection to Po- number taken was 105, a large number by mona College in 1923 and left Utah for Cali- modern standards, but low when compared fornia, where he resided until his death in to the previous decade. Too, the number of 1934. Rydberg was involved in completion of contributors increased. Jones continued to be parts of the North American Flora and was for encore active, but what does one do an to pass from the scene upon his death in of the following such a performance as his 1931. The scenario shifted from these actors, 1890s? took only 17 types, and Rydberg Jones who had played such important roles in dis- (alone or with Carlton) accounted for 24 covery, to others, most of whom did not oc- types. That Jones was active is indicated by cupy the stage for any lengthy period. Only his collections, which yielded type materials 19 types were added by nine collectors. Jones for the years 1900, 1903, 1906, 1908, and took his last in 1923, in Zion Canyon, return- 1909. Rydberg's contributions were taken in ing to the region of his early pursuits in his 1905 from Salt Lake Coimty and from the old age. A. O. Garrett continued to add spec- Tushar Moimtains, astride the Piute-Beaver imens during the period. His contributions coimties boundary. The other most important have never received the meritorious notice collectors of this time, as regards historical that they deserved, although the herbarium perspective, include Ivar Tidestrom, who was at the University of Utah bears his name. Al- to publish a flora of Utah and Nevada, and though Garrett founded an herbarium at East Albert Osbun Garrett, resident Utah botanist High School in Salt Lake City in 1903, his and teacher for more than four decades. Su- private collection resides at the University of san Stokes (who later treated Eriogonum), Utah, to which it was given under terms of a Grove Karl Gilbert (?), Leslie Newton bequest following his death in 1948. It is the Goodding (student of Aven Nelson), Louis best early collection extant in Utah. Hermann Pammel and R. E. Blackwood, S. J. Walter Pace Cottam entered the botanical (?) Harkness, Frederick Vernon Coville scene in the 1920s. He graduated from Brig- (along with Tidestrom), Carleton Roy Ball (of ham Young University with a Master of Sci- willow fame), and Charles Piper Smith (pro- ence degree in 1917 (the first awarded by the ducer of infamous works in Lupinits) each university). He returned to the university as a contributed one or more types during the professor in the early part of the 1920s and decade. began a lifetime investigating the state of The 60 types taken from 1910 to 1919 rep- Utah. His primary emphasis did not M'? with resented a further decline, emphasized even plant taxonomy; he is known for his 'vork in more by the efforts of 13 collectors for this plant ecology. Plants were contributed to the period. Rydberg, alone and with Albert Os- herbarium of Brigham Young University as a bun Garrett, worked in Salt Lake, Grand, and biproduct of his investigations of the ecology San Juan counties in 1911. Their work in the of the region. inspired others to follow southeastern portion of Utah was the first He fact, has served as serious attempt at imderstanding the flora of him into the field and, in that region following the pioneer work of an inspiration to generations of scholars. In Alice Eastwood. Their collections were rich 1931 he joined the University of Utah and in type materials. C. P. Smith, A. O. Garrett, served to found that herbarium also. His Willard Webster Eggleston, E. P. Walker, early collections are at Brigham Young Uni- Thomas Henry Kearney and Homer LeRoy versity and the later ones are at the Univer- Shantz, Edwin Blake Payson, Francis Whit- sity of Utah. The herbarium at the University tier Pennell, Malmsten, Ricker, and Alice of Utah was founded in 1870 (Chamberlain

Eastwood provided type material during this 1950), prior to Cottam 's arrival there, but time. most of the collections had been destroyed. 140 Great Basin Naturalist Vol. 42, No. 2

misplaced, or pilfered by that time. He estab- basis by any of the major universities, at least lished the herbarium on a permanent basis in as far as vascular plants were concerned. He 1933. He had founded the herbarium at Brig- was an enthusiastic collector and teacher. His ham Young University in 1923. The Utah collections within Utah and the surrounding State University herbarium was formalized in states formed the basis of the herbarium at 1934. Utah State. That herbarium became the cen- Other collectors during the 1920s were E. ter of plant taxonomy in Utah in the 1930s B. Payson (botanist at the Uinversity of under Maguire's direction, and continued to Wyoming), Charles Piper Smith (professor of hold that status under his student and succes- botany at Utah State during the period 1909- sor, Arthur H. Holmgren. 12, Ewan 1950), Pilsbry, George Everett Os- The herbarium at Brigham Young Univer- terhout (amateur botanist and lumberman, sity came under the direction of Bertrand F. whose herbarium is at RM), Nord & Sargent, Harrison following Cottam's move to Salt Fallas, and Mildred Esther Mathias. Payson Lake City. Harrison was trained in plant worked on revisions of genera important in physiology, but continued to build the herba- our flora, i.e., Thelypodiian and its relatives rium throughout his tenure at Provo. This and Cryptantha (as Oreocarya). Mildred E. was accomplished during a time when he Mathias treated the complex and difficult served as chairman of the department of bot- genera in the Umbelliferae. any, taught 15-25 hours of classes per week With Rydberg and Jones both gone from (in all botanical disciplines), and functioned the scene, the 1930s became a time for re- on university committees. His favorite class evaluation of western American plant tax- was plant taxonomy, and he instilled the onomy. The divergent views of taxonomy as same kind of enthusiasm in his students that exemplified by them were put to the test by he had experienced under the direction of no less than 30 individuals who worked in Cottam, who had taught him taxonomy in Utah in the decade. Some of these, William the 1920s. Slow was the growth of the collec- D. Stanton (a student of Cottam who worked tion in this period of economic depression on tlie flora of the Henry Mountains), A. O. and difficulties of travel in the remote re- Garrett, Bassett Maguire, Edward H. Gra- gions of Utah. Nevertheless, both Harrison ham, Bertrand F. Harrison, Sevile Flowers, and Cottam were able to contribute sub- and Perry Plummer, made concerted efforts stantially to the understanding of the flora at continuing the exploration of the state. and its ecology. Others were mainly involved in summary re- The work of Maguire and his associates visions and monographs of plant genera clearly places Utah State at the center of tax- whose members included Utah as a portion of onomic study in Utah during the 1940s. Ma- their ranges. Within this category were guire, Holmgren, and C. L. Hitchcock (with George Goodman, Charles Leo Hitchcock, whom Maguire collaborated), formed a nu- Mildred Mathias, Louis Otho Williams, Fred- cleus about which Utah taxonomy func- erick Joseph Hermann, Alice Eastwood and tioned. Other workers, all from outside Utah, John Thomas Howell, Susan Stokes, E. B. collected in the state in the decade. Elzada Payson, Howard Baker, Reed Clark Rollins, Clover and Lois Jotter traversed Glen Can- Cedric Lambert Porter (curator of the her- yon in 1940. Alice Eastwood (then in her bariumn at for many years), and Susan year) T. in in RM 83rd and J. Howell were Utah Delano McKelvey. Minor contributions were 1941, allowing Miss Eastwood a final view of made by collectors named Larson, Clawson, the scenes of her earlier explorations. Dwight Carter, F. A. Barkley and M. J. Reed, and Ripley and the amazing Rupert Charles Tompkins. Barneby (already working on in his revision There occurred a shift within the state dur- of the monumental genus Astragalus) visited ing the 1930s. Bassett Maguire had arrived as the state first in 1942. Barneby, an English- plant taxonomist at Utah State University man, with field experience in Spain, where (then Utah State Agricultural College) in he first encountered the genus Astragalus, 1929. Maguire was the first professional plant was inspired to come to North America after taxonomist to be employed on a continuing seeing the name of Astragalus asclepiadoides June 1982 Welsh: Utah Plant Types 141 in a catalogue of North American plants. He During the 1970s some 47 type specimens then examined the single specimen taken by were taken. The essential group of collectors Jones on deposit at the herbarium at Kew remained largely the same as in the previous and made the decision that he must see the decade, but there was some shift in in- plant in vivo. Lyman Benson, expert on Ra- stitutional affiliation and new names ap- nunculus and cacti, was in the state in 1948 peared. Noel and Patricia Kern Holmgren and 1949. Frank Gould, specialist in grasses, became affiliated with New York Botanical collected here in 1941. A person named Garden, and James Reveal was employed by Marsh took materials in both 1945 and 1946. the University of Maryland. Leila Shultz joined Holmgren as curator of the The 1950s mark a period of quiescence in Arthur at Utah State University late in exploration and collection of Utah plant herbarium the decade. The roster of people at Brigham types. Maguire had left Utah State for the Young Universty who aided in collection of New York Botanical Garden, and the post type materials include S. L. Welsh, N. D. At- World War II era found tlie professors in- wood, Joseph Richard Murdock, Frederick volved with teaching and other pursuits, J. Peabody, Kaye Hugie Thome, Kathryn N. which did not allow them to allot much time Taylor, Elizabeth Janet Chase Neese, Susan collecting. Fewer types were taken in this to White, Kent Ostler and David Anderson, decade than any time since the 1850s. Arthur Kim Despain, Betty Jean Welsh, and S. L. T. Hermann Holmgren, B. F. Harrison, C. L. Welsh.

Porter, Rupert C. Bameby, F. J. Hermann, and others contributed only 13 types in the Counties of Collection period. The list gives Utah State and Brigham Yomig about equal, if miimpressive, billing. An examination of types taken on a per

Collectors during the 1960s were affiliated county basis is instructive as to where em- with Utah State (Arthur Holmgren, his son phasis was given in early botanical excur-

Noel Hermann Holmgren , and Garrett Da- sions. Data on types by county are presented vidse), New York Botanical Garden (Rupert in Table 3. Washington County was and is a C. Barneby and Johnnnie L. Gentry), and favorite site for investigations by those tired Brigham Yoimg University (Craig Hanson, of the cold and snow of winter, and, because Stanley Larson Welsh, Glen Moore, James of the warm desert vegetation, which reflects Reveal, Larry Charles Higgins, Stella Leim- its climate, many people traveled there to omi Tree Welsh, and Nephi Duane Atwood). collect. Palmer, Parry, and Jones were there Ishan A. Al-Shebaz (of Harvard University) was tlie only other collector of Utah plant types in the decade. Personnel from Utah State and New York Botanical Garden were engaged in a collaborative study of the intermountain flora preparatory to publication of a manual of that flora. The school that developed at Brigham Young University resulted from the serendipity that followed the acceptance of the position of plant taxonomist

(the first for the university) by Welsh in Sep- tember of 1960. Welsh, his wife, and his students began a concerted effort at exploration of the flora of Utah. The nucleus of the herbarium developed by Cottam and by Harri- son allowed for continued expansion as the flora of the state was investigated. Explor- Fig. 4. Daniel Cady Eaton (1864), author of the Com- ation and collection produced some 50 speci- positae in Watson's Botany, who is honored by the name mens that have been named as types at the Penstemon eatonii Gray (photo courtesy of Richard W. present. Pohl). 142 Great Basin Naturalist Vol. 42, No. 2 early, in the 70s and 80s, when many of the compliment of types to the early work by taxa were undescribed. Because of their at- Rydberg, and to later attempts by more re- tention to that delightful region for early cent collectors. Summit County became im- spring and autumn floras, Washington Coun- portant in types largely, 1 believe, because of ty stands in first position with 163 types. Salt the access to the westernmost portion of the Lake County is second with 102, in part be- Uinta Mountains, and especially because of cause of its unique flora that extends from the early trip to the area by Watson. moderate to very high elevations, in part and Tremendous effort and expense is neces- of its because position within the state. Early sary for exploration of the state. Most of the taxonomists visited Salt Lake City, center of remaining endemics, if past experience is cor- commerce and hub of transportation for this rect, will be found in areas that remain to be vast region. Access to the mountainous areas explored in remote parts of the state. Each around the valley of the Great Salt Lake was year a few more are discovered, and with relatively easy for those operating from this each new generation of monographers addi- base. Jones's visit to the Wasatch Mountains tional types will be designated from among in 1879 yielded many types. Watson, Jones, collections of the previous years. Rydberg, Garrett, Stokes, and many others spent time in the canyons and ranges in close proximity to Salt Lake City. The large num- Authors of Taxa Based on Utah Types ber of types from San Juan and adjacent Those who collect types may or may not Grand counties is attributable in large part to be prepared to describe taxa based on them. the early work by Eastwood and Rydberg Indeed, few are those who recognize, imme- and Garrett. Garfield County owes its large diately upon collection, that the material col-

lected is undescribed. Often, new taxa are Table 3. Utah types listed by county of collection. made to fit into concepts of previously de- County Number scribed entities, whether they fit well or not.

Beaver 25 Each worker is acquainted with only a por- Box Elder 7 tion of the var ktion of the total flora, and Cache 28 few consider themselves sufficiently prepared Carbon 21 to name as new all portions of the variation Daggett 3 encountered by them. To some investigators Davis 9 Duchesne 19 any variation is worthy of a name, regardless Emery 45 of the importance of that characteristic in Garfield 56 the total range of variation available. To Grand 43 them the value of a single character is over- Iron 20 Juab 23 riding—the character makes the species, as Kane 59 Linnaeus stated. Because of differences in Millard 19 training, philosophy, and expertise, and Morgan 1 others due to change in rmderstanding over Piute 35 more than a century of time, the lasting qual- Rich 2 Salt Lake 102 ity of names recognized as belonging to vi- San Juan 85 able entities varies from author to author, Sanpete 22 and from group to group treated by the same Sevier 48 author. There have always been more spe- Summit 53 col- Tooele 36 cialists involved with naming than with Uintah 33 lecting. Beacuse of this, many authors never Utah 30 saw the taxa that they named in vivo— they Wasatch 9 were herbarium botanists with experience in Washington 163 dried and pressed plants only. Among the Wayne 26 Weber 4 herbarium botanists quality of judgments var- Unknown 47 ied as much as between them and their field- experienced counterparts. Gray and Watson Total 1073 represent the two extremes, and both were June 1982 Welsh: Utah Plant Types 143 successful descriptive botanists whose named but where the disciplines can supplement taxa have stood the test of time. The syn- each other that course seems best to this drome of "name in print at any cost" af- worker. flicted only a few of the major authors (i.e., Some of the cast of authors is the same as Edward L. Greene and Charles Piper Smith). for the list of collectors, but many are differ-

The names of authors of Utah types are listed ent. Noted is the fact that some of the collec- in Table 4. The decade of publication of tors of many types failed to publish new taxa Utah types, number of authors, and average based on their own collections. That task fell nmnber of names per author are cited in to others, either by default or on purpose. Table 5. One will note that Jones named only 167 taxa There has always been a disparity among but collected some 305. The others were botanists regarding species concept and ap- named by some 44 botanists during and sub- plication of names. As there has always been sequent to Jones's lifetime, the latest by Re- a problem of distinguishing between no- veal (1977). This was particularly disturbing menclature and taxonomy, a problem has also to Jones, not because others purloined his existed between botanists in the recognition material, but because he did not feel that of of of species in which infraspecific categories most them were worthy being named. The 1840s demonstrate only two authors of are allowed. Some have chosen to draw spe- Utah taxa, i.e., Torrey and Charles cies concepts that are broad and allow for the John John Fremont, and John Torrey and Asa Gray. inclusion of several infraspecific taxa in a hi- Torrey was situated at Columbia College, erarchial system. Others have felt that such a New York, and Fremont was then engaged in nested system, in which species, subspecies, explorations of the West. Later, Fremont varieties, subvarieties, and forms were al- would be a candidate for the presidency of lowed, represented a return to pre-Linnaean the United States (in 1856). Together they phrase names. To them such usage was and is named Perchiton occidentale (Torrey & Fre- intolerable; the binomial system of nomencla- mont 1845). Torrey and Asa Gray collabo- ture should be adhered to at all costs. When rated in naming Gilia stenothyrsa and Se- this concept is followed, and when it is necio multilobatus (Torrey and Gray 1849). coupled with the concept that any variant is All these taxa are based on specimens taken worthy of a name, it is possible to multiply by Fremont. Asa Gray, working out of names inordinately. Names based on Utah types were supplied by workers from both of the divergent schools, and by others between. Still other workers were not bound by the sense of logic provided by either school; they were self- taught and were trapped in their preconceived notions that any variation constituted a case for a new species, and that the species were immutable. Because of the differences in approach, in imderstanding, and in training, our Utah plants were given a variety of treatments. The variation among botanists today is not as great, but there is still variation. A current move has been to recognize only one infraspecific category, the subspecies. If such a move were to succeed, then all variants now recognized at other infraspecific levels would be treated as subspecies, and levels of variation would be hid- den by the apparent uniformity of the Fig. 5. Sereno Watson (ca 1870), collector and author category. It is true, of course, that taxonomy of Utah plant types, second in importance only to Mar- should not be dependent upon nomenclature. cus E. Jones (photo courtesy of Richard W. Pohl). 144 Great Basin Naturalist Vol. 42, No. 2

Harvard University, was the principal 19th- Table 4 continued. century American plant taxonomist; his con- Number Years tributions are legion. Author of names active Five authors described taxa based on Utah types in the 1850s. These include John Tor- rey, Torrey and Gray, Elias Durand, George Bentham, and Edward Griffin Beckwith. Their published taxa are based on collections by Stansbury, Creuzfeldt, Beckwith, and Car- rington. Seven of the 12 taxa stand at some taxonomic level in contemporary treatments. Perhaps the best known of the taxa described

in the decade is Phaca mollissima var. uta- heivsis named by Torrey and now cited as As- tragalus utahensis (Torr.) T. & G. Torrey and Gray were the principal American taxonomists of the period. John Torrey (1796-1873) was a professor at Columbia College. He became associated with Asa Gray (1810-1888), his student, in writing a "Flora of North America" (1838-1840). Gray

Table 4. Authors of names of Utah plant types.

Number Years Author of names active

Al-Shehbaz, I.A. (b.?) 1 Anderson, L.C. (13-1936) Atwood, N.D. (b.l938) Bailev, L.H. (1858-1954) Baker, M.S. (1868-1961) Baker, M.S. (1868-1961) &J.C. Clausen (b. 1891) Ball, C.R. (1873-1958) Barkley, T.M.(b. 19.34) Bameby, B.C. (b.l911) Barneby, B.C. (b. 1911) & N.H. Holmgren (b. 1937) Beal, W.J. (18a3-1924) Beaman, J.H. (b.l929) Benson, L.D. (b.l909) Bentham, G. (1800-84) Bicknell, E.P. (1859-1925) Blake, S.F. (1892-1959) Brand, A. (186.3-19.30) Brandegee, M.K. (1844-1920)

Brenckle, J.F. (b.l875) & W.P. Cottam (b.l875) Britton, N.L. (18,59-1934) Britton, N.L. (1859-1934 &J.N. Rose (1862-1928) Buchenau, F.G.P. (1831-1906) Candolle, A.L.P.P. (1806-93) Chaudliri, M.N. (b.l932) Clover, E.U.(b. 1897) & M.L. Jotter (b.l914) Cockerell, T.D.A. (1866-1948) Cottam, W.P. (b.l894) June 1982 Welsh: Utah Plant Types 145

Table 4 continued. Table 4 continued. 146 Great Basin Naturalist Vol. 42, No. 2

was to be associated with Torrey throughout his hfe, and in a biography compiled at Tor- ". rey 's death. Gray (1873) noted that . . he went again across the continent to California,

and . . . enjoyed the rare pleasure of viewing in their native soil, and plucking with his owTi hands, many a flower which he had himself named and described from dried specimens in the herbarium, and in which he felt a kind of paternal interest." The 1860s did not offer much in the way of opportunity for description of Utah plant taxa. Torrey and Gray described Navarretia setosissima, and de Candolle named Quercus stellata var. utahensis, based on a collection presumeably taken by Beckwith "between Salt Lake City and the Sierra Nevada" in 1854. Sereno Watson and Asa Gray dominated the botanical literature dealing with Utah plant types in the 1870s. They were joined by John Torrey, Daniel Cady Eaton, James Wat- son Bobbins, George Vasey, George Engel- mann, Lester Frank Ward, and Charles Christopher Parry in the publication of some 161 taxa. The most important single work of the decade is the report of the U.S. Geologi- cal Survey of the 40th Parallel, Volume 5, Botany, authored by Watson (1871), but in collaboration with D. C. Eaton (honored in the name Penstemon eotonii Gray), George in- Engelmann, and J. W. Bobbins. The work cluded 51 taxa based on Utah types (39 by

Table 5. Decade of publication of Utali types, number of authors, and average number of names per author. June 1982 Welsh: Utah Plant Types 147

Axel Rydberg, Thomas Henry Kearney, Lu- in the 1940s clearly belongs to Bassett Ma- cien Marcus Underwood, Aven Nelson, guire, who accepted a position at Utah State Thomas Albert Williams, Benjamin Lincoln Agricultural College in the 1930s. Others Robinson, Jesse Moore Greenman, Jared working in the flora of the state in the 1940s, Gage Smith, Charles Vancouver Piper, and who contributed names of new taxa, include Elias Emanuel Nelson. Alice Eastwood and Rupert Charles Barneby, Arthur John Per Axel Rydberg were both beginning their Cronquist, Alice Eastwood, C. Leo Hitch- activities within Utah; they would spend dec- cock, John Thomas Howell, Ivan M. John- ades involved, in part, with the Utah flora. ston, and David D. Keck. The decade was the golden age of descriptive The 1950s were relatively quiet. This peri- botany in Utah, and in deference to Jones's od was marked by the shift from efforts of contributions the decade should be known as World War II, and Bassett Maguire had left the "Jonesian Era." Utah State. Charles Piper Smith, in his senes- The decade from 1900 to 1909 showed cence, and suffering from a lack of under- only a slight decline in taxa named from the standing of what constituted a species within Lupinus, seven inconsequential lu- state when compared to the previous decade named pines as species, all but one of them from the (180 compared to 181), with the deficit in head of Salina Canyon, and all of those be- types collected in that period (105) being longing to Lupinus sericeus in a broad sense. made up by names being applied to speci- Barneby named five species during this mens taken previously. Some 18 authors as- period. saulted tlie collections of the state. Jones did In the decade of the 60s Barneby (with 9 not come close to matching his unparalled new taxa) was displaced by James Lauritz feats of the 1890s. He named only 21 taxa in Reveal (with 15 new taxa) as leader in au- the decade. Rydberg, an enthusiastic and thorship of new names. Reveal had begun his energetic professionally trained botanist, assault on Eriogonum, a genus rich in endem- Rished to fill whatever vacuum was left fol- ic taxa and other narrowly restrcted plants. lowing Jones's valliant efforts. Introduced to Craig Hansen, James L. Reveal, and Larry the region in the mid-90s, Rydberg had named only one taxon from Utah prior to the turn of the century. Now he assumed the leading position as author of 65 Utah types in the decade, in preparation for publication of his flora of the Rocky Mountains (1917). Prin- cipal authors besides Rydberg and Jones were Edward L. Greene (21 names) and Aven Nel- son (20 names). Rydberg had no close competition in the period from 1910 to 1919. He named another 65 taxa based on Utah materials, and Jones named only 14. The 1920s were the lowest for publication of new taxa since the 1880s. Only 59 taxa were named by 19 authors. Rydberg authored only 9, second to F. W. Pennell, then engaged in a revision of the K genus Penstemon, who named 14 taxa. Neither Jones nor Rydberg were to live beyond the early 1930s. Jones named a final 4 taxa in that decade. The period fell to Sidney

Faye Blake (4), Alice Eastwood (4), Ivan Fig. 6. Asa Gray (ca 1865), author of numerous Utah Murray Johnston (2), David Daniels Keck (4), types, based on collections corresponded by Watson, Aven Nelson (2), F. W. Pennell and Susan (3), Parry, Thompson, Siler, Bishop, Palmer, and Jones, and Gabriella Stokes (12), largely without a lead- the most important North American plant taxonomist of er. Leadership in the naming of Utah's plants the 19th century (photo courtesy of Richard VV. Pohl). .

148 Great Basin Naturalist Vol. 42, No. 2

Charles Higgins and Stanley L. Welsh, all as- Marcus Eugene Jones— His Contributions sociated with Brigham Young University, AND His Contemporaries named a third of all species described during In the summer of 1879 (Broaddus an the decade. 1935) event occurred which, because of its hap- Botanical exploration continued at an ac- penstance and because of its forthrightness, celerated pace in the 1970s. Impetus for pub- would not seem to have any lasting effect. lication of the taxa discovered in this period A young botanist, on one of his earliest trips was supplied by preparation of the grand west from the rolling farmlands of Iowa, hap- plan for publication of the "Intermountain pened to find a wallet (Munz that Flora" by members of New York Botanical 1965) be- longed to a General Wm. Palmer. The wal- Garden and associates. Included in that team J. Arthur let contained valuable papers and was are Arthur J. Cronquist, H. Holmgren, returned to its owner. Following this act of Noel H. Holmgren, Patricia K. Holmgren, an honest man. General Palmer sent the bot- and James L. Reveal. Noel Holmgren, alone anist, Marcus E. Jones, on repeated trips to or in combination with others of the group, explore regions for mining or agricultural po- published descriptions of 10 new taxa from tential. Jones was enabled to travel widely in Utah, and Reveal published 1 1 the western portions of the continent, all be- Work on exploration of floristic provinces cause of serendipity— the finding and return of Utah by botanists from Brigham Young of another's property. Because of this, west- Universtiy was expanded in the 1960s, and ern American plant taxonomy was altered for was again intensified in the 1970s. The Utah all time. He indicated his gratitude to Palmer Flora project, initiated in the 1960s, offered 1891) by naming Cleomella palmerana; stimulus. Further activity was increased by (Jones "Dedicated to General Wm. Palmer, than passage of environmental protection laws, J. there is no one more interested in sci- and especially by passage of the Endangered whom entific researches in Utah, or who has shown Species Act of 1973, as amended in 1978. interest in substantial Welsh, alone or with others, described 29 more a more way." Jones was born in Jefferson, Ashtabula taxa in the decade. That is the largest number involving any individual since the 1910-1919 County, Ohio, on 25 April 1852 and died at Bernadino, California, in his 83rd period. Atwood named 4, bringing the total San year on for those at Brigham Young University to 33, 3 June 1934. He was reared and educated or 61 percent of the total for the decade. near Grinnell, Iowa, having moved west with his family in 1865 ("on the very day that Lin- coln was shot graduating from Grinnell The Decades Beyond "), (Iowa) College in 1875 (Broaddus 1935). In

Tliat the past is prologue is illustrated by 1876 and 1877 the youthful botanist collect- the fact that several new taxa have been de- ed plants in Iowa. Early in 1878 he left for scribed from Utah since the close of the peri- Colorado, where he collected large numbers od covered by this text. Those which have of specimens that were sold as "sets." He re- been published are included in the list of turned to Colorado Springs in 1879 as "pro- type material, but they are not summarized fessor of natural science." During July he as for previous decades. Too, I recognize that traveled to Salt Lake City and collected at a decade does not begin with the zero year, Alta, City Creek Canyon, Lake Point, and but ends with it. For the sake of convenience Bingham Canyon. on my part, I have chosen to use it otherwise. Jones moved from Iowa to establish per- Those who follow can find fault, as all who manent residence in Salt Lake City in 1880. view others' work must, but it will not mate- Accompanying him was his bride (since Feb- rially affect the data presented here or my ruary 18), Anna Elizabeth Richardson view of the history of plant collection in (1853-1916), who would spend most of the Utah. Perhaps some will find this summary remainder of her life with Jones in Salt Lake iLseful, and possibly someone will choose to City. Instructive is his account (Munz 1965) keep it current for the future. of the itinerary from Salt Lake City to June 1982 Welsh: Utah Plant Types 149

Washington County and return during the His respect for women was less than over- spring of 1880. whelming. Marcus (called Mark by his family 'T left Iowa Falls, Iowa, Febmary 18th for and close friends) wrote of his great admira- Salt Lake City and got there the 24th. March tion and love for his mother (Jones and

3rd I hired a team and started to drive to St. Broaddus 1952), but he demonstrated little

George. I reached Sandy. On the 4th I re- regard for other women. His unpublished dia- turned to Salt Lake and back to Sandy (hav- ry of 1880 (POM) cites his expenses on his ing forgotten something). On the 5th reached wedding day, and then, almost as an after- Pleasant Grove, Spanish Fork on the 6th, thought, notes in the margin of the diary, Mona on the 8th, and Juab on the 9th, Warm "today I married Anna Elizabeth Springs near Gimnison on the 10th, Willow Richardson." Bend below Richfield on the 11th, Elsinore Anna Jones was a talented and educated having taught school for several on the 12th, where I stayed three days and woman, in Falls, Iowa, went 14 miles up the Clear Creek Canyon on years from the age of 16, Iowa in the surrounding area. She went to the 18th. Then went seven miles farther on and Grinell to complete one year of advanced the 19th, then to Pine Creek fifteen miles study in English, but remained to complete above Beaver, Beaver on the 20th, Buckhorn the college course. Finally, she accepted a Springs on the 22nd, Summit south of Paro- position as "Lady Principal" at Grinell wan on the 23rd, then to Fort Hamilton on (Iowa) College, a position she resigned in the 24th where I found the first spring flow- January of 1880 so that she could marry one ers. I reached Leeds on the 25th and St. of her former Latin teachers, Marcus Eugene George on the 26th. I botanized all around Jones. the city on the plains and mesas until the 9th It is evident from correspondence and of April when I went over on the Santa from the published account of her life that Clara. On the 14th I went to Washington five Anna Jones supported her husband in his life miles east. On the 20th I started for Salt Lake goal of botany. She did sewing for hire, took and reached Bellevue. On the 21st got to Ka- in individual boarders at first, and finally rarrah (Kanarrah?). On the 22nd Fort Ham- (1900-1910) established a boarding house to ilton. the 23rd Parowan. On the 24th. On provide funds for Jones to continue his work. Beaver. the 27th Creek. On the 28 On Com Jones had three main goals; to collect the Fillmore. the 29th to the ridge north of On flora of the West, to publish a treatment of Scipio. the 30th I reached Nephi. On On Astragalus, and to write a flora of the Great May 1st I reached Spanish Fork. On the 3rd,

I reached the Point of the Mountain and the 5th Salt Lake City." One cannot find in Jones's rather volumi- nous published writings whether his young and beautiful bride of a few weeks accom- y^WS fS5j panied him on this long and tiring trip. He makes no official acknowledgment that she did go with him. In an account of Anna Jones's life written by her daughter (Broaddus 1952), however, it is noted that "At first Anna went with him on wagon trips of incredible difficulty and danger, even as far as St. George in southern Utah, helping collect wild flower specimens, pressing and drying them, assembling sets for shipment to Ejro- pean and eastern herbaria, even mounting many of the most attractive flowers in little Fig. 7. George Engelmann (ca 1864), specialist in cac- booklets of 'Western Flora,' which they ti, yucca, agave, and conifers, and correspondent of Par-

" sold. Probably she was on this first trip ry, Jones, Siler, and others (photo courtesy of Richard

south; it is inconceivable to think otherwise. W. Pohl). 150 Great Basin Naturalist Vol. 42, No. 2

Basin. All funds provided by Anna and by It is not surprising that Jones was able to himself, from mining, consulting, writing, and accomplish so much, with a wife so dedicated other projects, were technically supposed to to his care and to his cause. go toward accomplishing of these goals. But His trip to St. George required 22 days, Jones was not good at managing money, ex- but he returned to Salt Lake City in just 16 pending large quantities of capital in photog- days. This compares with a drive of about raphy and unwise investments. Anna lived to seven hours each way today. The state must see practically all of the first goal and much have seemed overwhelmingly large to Jones, es- of the second one completed. Jones had as it still remains, but his youthful enthusiasm sentially completed the manuscript of Astra- seems to have overshadowed any doubts re- galus by 1910 (Bameby 1964). Neither Anna garding size or complexity of the flora. live to see final nor Marcus would the goal In the ensuing decades he researched a sur- attained. The Great Basin flora would remain prising number of places, including many of for others. the mining areas such as Frisco, Silver Reef, Broaddus (1952) notes of her mother that Detroit, and Eureka. The state, despite its "The last year of her life whe spent in Clare- great size, did not occupy all his time. He mont, California, keeping house for her traveled to California, Nevada, Montana, youngest child, Mildred, to help her finish Arizona, and finally to Mexico, but his pre- college, and earning what she could by doing occupation with Utah plants is evident from dress making." Mildred was one of three chil- his accomplishments. During his 43 years as dren; the others were Howard and Mabel resident botanist for Utah this resourceful (Broaddus 1952). person built a personal herbarium of about 100,000 unmounted sheets (Munz 1965), among them the best set of Utah plants as- sembled in all history. Included among that herbarium were 305 Utah types, almost three-tenths of all types collected in the state between 1843 and 1981. Additionally, Jones had named some 165 plant taxa based on Utah types, mainly on his own collections. If that was not enough, Jones, after becom- ing disgruntled with other publishers (his life

is filled with examples of being disgruntled), began to print his own journal "Contribu- tions to Western Botany," on a hand-oper- ated press, using badly worn type, at his home on the avenues in Salt Lake City. This remarkable pioneer undertook to complete a revision of the genus Astragalus for North America. Difficult was the task, at any time, but Jones also faced the problems of transportation, communication, and paucity of library materials; and, though largely dependent on his own collections, his meager

locality data is encyclopedic when compared to that recorded (or unrecorded) by some of his contemporaries. Despite these hurdles, the revision of As- f tragalus was essentially completed by 1910 (Barneby 1964). It was not published until Fig. 8. Albert Osbun Garrett (ca 1930), resident Utah 1923, apparently printed as were his "Contri- plant taxonomist for more than five decades, teacher, his press, with worn type. collector, and author of six editions of the Spring Flora butions" on own of the Wasatch (photo courtesy of Lois Arnow). The text was distributed on "Feb. 15, 1923" June 1982 Welsh: Utah Plant Types 151 and tlie plates on "June 20, 1923" (Jones centers of botanical investigation in the coun- 1923). try, and in part to the lack of recognition for In that same year, Jones's herbarium was his contributions, which are staggering. Nev- piu-chased by Miss Ellen B. Scripps of La Jol- ertheless, his criticisms recounted herein la, California for Pomona College, Clare- were largely unfounded. His contemporaries mont, California. That herbarium, the richest were, with some exceptions, honorable men of any in Jones's first set of Utah type mate- doing work in the best tradition open to rial, resides today at Rancho Santa Ana Bot- them. They were, in their own ways, as iso- anical Garden, having been integrated there- lated and as unrecognized as Jones. His pub- in in the 1960s. lished attacks on them yield information and The species concept of Jones was similar to insight into them and him. They are cited that of Asa Gray, the leading 19th century here for that reason. Jones thought of himself botanist with whom Jones corresponded, and as benevolent and charitable. to whom he sent early sets of his western col- The following examples are presented to lections. A tribute to Jones abilities is in- give an idea of the time in which Jones lived, dicated in the fact that more than half of the and to give a portion of the cast of characters taxa named by him from Utah are still ac- who wandered through Jones's existence as a corded taxonomic standing in contemporary botanist. treatments. In his "Contributions to Western Botany. He came to look upon western American No. 12" (1908), Jones took N. L. Britton to plant taxonomy as his own, and summarily task for provisions of the American Associ- defended his territory against all comers. His ation for Advancement of Science attempt at ego was monumental, but was occasionally botancial nomenclature. One should know tempored by flashes of knowledge that he that no standardized code was acceptable to might not be omnipotent. The field season of the botanists in the United States at that 1894 (Jones 1895) found him in the field, time, and one would not be forthcoming until hired by the U.S. Department of Agriculture publication of the International Code of Bot- as "Special Field Agent." It was to be his anical Nomenclature in 1935, following best year, with "about 50,000 specimens and Jones's death. Jones wrote of the Brittonian 1700 sets, there being 1106 species and varie- attempt: "Its chief virtue was in a rigid ad-

." " "... ties in the sets . . Or, (Munz 1965) "The to- herence to priority, and further we find tal number collected was 4060, and about that it isn't priority if it goes back farther

35,000 specimens," depending on which of than 1753, and it is not priority if it is not in the two accounts of the season one is to be- the Latin language." And, still further; "If it

Ueve. Jones (1895) opened the introductory is contended that common terms are not Lat- remarks of the report of the 1894 collections, in we can easily follow the lead of Britton

which is the most remarkable single pub- and put a Latin tail on them (a la Manihota lication in Utah's history of botanical explor- Britton for Manihot). In that case we would ation, with the following sentence: "Having have such very attractive combinations as the had an opportunity to examine the material following: Sagebrushurn tridentatum, Brit-

in the National Herbarium I have been able tonastriim tumbleweedum, Greenella slip- to correct many errors, and possibly to make penjehna, RydbergioneUa bitterroota, Covillea a few more." That publication, Jones's "Con- creosotebusha, Nelsonella greasewooda, etc."l tributions to Western Botany. No. VII.", re- Jones did not agree with acceptance of ported on some 119 new taxa for Utah, inter 1753 as the starting date for botanical no- alia. That record will stand for all time. The menclature, and neither did he believe that report was published in "Proceedings of the synonyms should stand in the way of a sub- California Academy (Second Series, Vol. 5)," sequent use of those same names. Latin as an and was one of the last of his contributions official language for publication of new spe- published elsewhere (Jones 1895). cies was unacceptable. Jones (1908) wrote: There was no one free from Jones's criti- "The purpose of science is to clarify, ... so cism, which resulted from a growing para- that those who follow us can take up work noia due in part to his isolation from the where we left off and ultimately bring out all 152 Great Basin Naturalist Vol. 42, No. 2

the underlying facts and explain causes of fit, and it is a standing joke among botanists." natural phenomena." Latin, which was in- Jones (1913) then discusses the work by Ryd- cluded in his formal education at Grinnell berg in Habenaria; "As though Rydberg had College, and which he had taught in the late done any original work in the genus." And 1870s, would not add to clarification for further, "Rydberg tries to justify this work by those who would follow him, he thought. saying that some of the changes have already Jones carried his criticism to Dr. B. T. But- been recognized by Europaean botanists.

ler who published a treatment of the western Now if there is any crazy thing that some Eu- American birches in the Bulletin of the Tor- ropean botanist has not done in the near or rey Botanical Club in August of 1909. After remote past the writer would like to have it systematically chastising Butler on the merits mentioned, and to use that as an excuse for of his work, which criticisms were merited, if doing some other crazy thing is no argument not pleasantly couched, he unfairly attacked in its favor." The last sentence is as appli- the education of Butler his and the quality of cable today as it was when Jones wrote it. teachers (Jones 1910). "The writer [Jones] Some contemporary American botanists look would be glad to subscribe several cents to- to Europe as the wellspring of all that is im- ward a fund for the education of the profes- portant or believable in 20th-century tax- sors who are responsible for this kind of work onomy. If the Europeans subscribe to splitt- so that they might go to some botanical kin- ing then we should follow. The idea is as dergarten where they could learn that a plant absurd today as it was in Jones's time; the growing in the shade or on a north slope or Europeans have mainly looked at their flora in a cold lake will differ from another from too long, if we have not looked long enough. the same parent which grows in a warmer Jones was not without humor in his criti- and drier situation, and that this condition cisms. He (1910: 34-42) wrote a rebuttal to a will be reversed when the offspring of these publication by A. A. Heller dealing with no- plants occupy reversed conditions; and where menclature. Jones wrote: "The publication of they could learn that a poodle dog running the article on nomenclature in my last Con- around a house is not necessarily two distinct tributions seems to have given Mr. Heller (in species because they see his tail only on the Muhlenbergia) a bad attack of mental colic, south side and later on his head only on the recurrent colic. His capacity to appreciate north side." the motive and scope of my criticisms re- Perhaps, but not likely, Jones would have minds one of the Englishman who after trav- been less harsh on Dr. Butler, but that stu- eling extensively in this coimtry returned dent took his Ph.D. degree at Bronx Park and home and said to his admiring friends 'The Columbia, always synonymous to Jones with Americans are a very clever people but they gross misunderstanding of "true" botany. have many uncouth expressions, for example Jones attacked members of the Brittonian they say 'Where am I at', we would say school, especially P. A. Rydberg and those as- 'Where his my at.' sociated with him. An enlightening aside on Jones (1910: 35) continued his argument the Brittonian school is found in Jones's against the principal of priority begun earlier (Jones 1913: 27) comments on Gray's Manual, (Jones 1908). After giving other examples of Seventh Edition, in which he states; "One of treating names from languages other than the most commendable things in the book is Latin, he notes, "Now that he [Heller] has the ignoring of the split genera of Britton, Hellerized all the English names . . . and got-

Rydberg, and Small. For some years the Brit- ten the Hellerian tail properly adjusted to all, tonians have raised a great hue and cry about he will find that Greek antedates the English, the new and original work done by them in then can get up another set of names from splitting up Grayan genera. A recent repeti- the Sanscrit, then the Chinese, then he might tion of this is in the Torrey Bulletin where take up the Egyptian and give still another the new Mantial was reviewed through Bronx batch of new names. By the time the bot- Park glasses. Anything published by Harvard anical public has begun to reccover from the is like a red flag before a bull to Bronx Park. last of these afflictions it is likely that the Whenever Harvard sneezes Bronx Park has a phonograph and the graphophone will b*^ '"^ "

June 1982 Welsh: Utah Plant Types 153 perfected that they can take the ripple marks Bameby (1964) has indicated that Jones made by sound waves on prehistoric mud was wrong in his malicious attacks, especially

(now turned to stone) produced by the inco- of Rydberg. There is logic in Rydberg's ap- herent babbhngs of some of the simian ances- proach, even if that approach is subject to in- tors of the Brittonians, and reproduce them terpretation. He believed in the binomial sys- so that they will be as lucid as some recent tem of nomenclature strictly. He (Rydberg descriptions of plants, and will have far 1929) believed that the variation which he stronger claim to priority of publication than observed, and he was a keen observer, should approach called to at- the Brittonian checklist had at the time when be described. Such an tention those who were his contemporaries it was said to have been published. and those who followed him. His work was Jones was wrong to oppose the principal of always limited to those few specimens he had priority, and because of his obstinacy, he cre- available. That he should see only portions of ated synonyms that were unjustified, i.e., the the entire picture is a fault not his alone; all case of Astragalus angustus (Jones) Jones, botanists are troubled with the same problem wherein he refused to use the legitimate re- in varying degree. placement "ceramicus" proposed by Sheldon But, it was on Greene and Rydberg that because earlier names were already occupied. Jones showered his continuing invective. He ultimately provides for a stability in Priority survived both of them, and heaped in- it chaos could result. nomenclature; without dignation on them in their later years, and in Jones should have understood the necessity Greene's case, even following his death. for a consistent nomenclatiire. E. L. Greene died on 10 November 1915. In the same article in which Jones (1910: Presumeably Jones wrote his scathing obitu- 35) attacked Heller, he took the opportunity ary of Greene shortly thereafter, but it was for a broadside against P. A. Rydberg and E. L. Greene. "Greene and Rydberg especially have been addicted to bolstering up fictitious species and genera by reasoning from analo- gy instead of known facts. Such arguments always get their users into difficulty. I well re- member when it was a stock argument among geologists to claim that the climate of the Arctic was Tropical in the early Qua- ternary because elephants (mastodons) lived there, animals which live only in Tropical climates. Many years afterwards one of these animals was found in a perfect state of pres- ervation imbedded in ice, and it proved to have very long hair and its stomach was full of Arctic plants which it had eaten, showing that it had its home in a frigid climate where that race of animals flourished, and not Trop- ical as has been dogmatically assumed," Jones wrote.

The mastodon case is cited by Creationists in contemporary literature as demonstrating existence of tropical climates in high lati- tudes, but the plants taken from frozen stomach contents are said to consist of tropical

plants., i.e., buttercups for example. The Fig. 9. Anna Elizabeth Richardson Jones (1913), wife rattle-headed thinking alluded to by Jones, if of Marcus Eugene Jones, who helped him to collect, and not altogether warranted in the men he at- who provided money to support him in his botanical

tacked, is not attributable to botanists alone. works (photo courtesy of Charles Walker). 154 Great Basin Naturalist Vol. 42, No. 2 not published until a decade and a half later tragedy in American botany, the death (Jones 1929). The opening paragraph of the of Rydberg before his time, the first tragedy paper (Jones 1929: 25) entitled "Greene" is as having the of been death J. N. Rose. Their follows. "There have been several notable disappointment at the treatment accorded deaths in the botanical world since my last them by the botanical public was, however, contributions. Greene, the pest of systematic partly their own fault; for no one has a right botany, has gone and relieved us from his to subordinate his own judgment to that of botanical drivel. They say that the good that another, as I think they did to Britton's, for men do lives after them, but the evil is en- the sake of financial support." Later, Jones terred with their bones. I suspect that his (1935: 142) continued, "Rydberg was inclined grave must have been a big one to hold it in his later years to consider himself per- all." Subsequently, Jones (1929) continues; secuted. I never knew that he had any per-

"Greene was first, last and all the time a bot- sonal enemies. Nor do I think any one ever anical crook, and an unmitigated liar, when it got after him for personal reasons. I certainly suited him to try to make a point against never did, though once or twice I had some someone else." reason to feel that he had not treated me fair- Rydberg survived until July of 1931, and ly in private correspondence." had published his monumental Flora of the Ivar Tidestrom and Aven Nelson were sub- Rocky Movmtains first in 1917. The work was jects of his invective too, and not without reissued in revised form in 1922 (Rydberg some justification. They were troubled by a 1917, 1922). Not until much later did Jones lack of material to make definitive taxonomic (1929: 9) take note of Rydberg's flora in decisions, and they lacked the experience and print. The opening paragraph of Jones's re- understanding that Jones had accumulated in view begins: "We are glad that at last Ryd- his decades of active field and herbarium berg has put his conception of species and studies. Despite the quality of their work, as genera in book form. His sporadic pub- is apparent in their publications (Tidestrom lications hitherto have shown botanists that 1925, Coulter & Nelson 1909), their floras he has no conception of either species or gen- were to stand for decades as the authorities era. In addition he accommodates us by on plant names for much of the Rocky Moun- saying that any plant deserving of a name tain and intermountain regions. The pub- should have a specific name. What he really lications were used by generations of botan- means is that if any fool botanist anywhere in ists who were never quite satisfied with keys, the world has applied a name to a plant as descriptions (lacking altogether in the work variety fonn or subspecies that name should by Tidestrom), or geographical data. The be raised at once to specific rank, and which shame in all of Jones's criticism is that he did he proceeds to do. At the same time he not complete a work on the flora of the Basin quibbles over priority by position as though and Range Province of his own. It, too, it was a matter of any importance. He would have been flawed, as all works are, but " 'strains at a gnat and swallows a camel.' with evidence taken over a long lifetime

"This book is what I have in the past dub- (more than four decades), it would have bed bughole botany, species depending on served well those later generations on whom the number of bugholes in the leaves. Ryd- was afflicted the work of Tidestrom, and the berg has no conception that there is any such peripheral work of Coulter & Nelson. Jones's science as ecology, or that environment has opus magnus laboris, his revision of Astra-

" any effect on species variation. galus (1923) is flawed, not by the species con- Jones (1935, published posthumously) cept or by the descriptions, but by the rattle- wrote an obituary for Rydberg also, in which headed polychotomous keys, which are diffi- he was more a gentleman, but the rancor cult if not impossible to follow, and by his comes through even then. Jones, it seems, persistence in following his own code of no- never knew when to let an argument end, menclature. The keys, of which he felt un- even when there was no chance for his oppo- justifiably proud (as noted by Bameby 1964), nent to speak. Jones (1935: 141) states: "On are based on a complex number and letter se- July 25, 1931 there occurred the second quence, which leaves the user wondering June 1982 Welsh: Utah Plant Types 155 where one goes from any particular place. can scarcely plead 'not guilty' to such a That his species concept has stood the test of charge." A portion of the problem of "over time is indicated again in the work by Barn- description" was recognized as duplication of eby (1964), who published a revision of the effort, where one taxonomist did not know ". North American species of Astragalus some that the entity was already described. . .the four decades following the appearance of more the taxonomist studies a genus or a fam- Jones (1923) work on that incredible genus. ily, the more forms he recognizes, and the Bameby was able to recognize some 60 per- better he knows the family, the finer splitter cent of the taxa named in that genus by he becomes. Then he either distinguishes Jones, but only 38 percent of those named by more restricted genera, or recognizes in- Rydberg, and most of those at varietal rank. finitesimal species; or perhaps he recognizes Jones had "seen more of the elephant," as large genera and broad species, but splits these into numerous subordinate categories: it were, than his contemporaries, and with his sections, groups, subspecies, varie- abilities of observation and his acute awar- subgenera, ties, forms, etc., without end." Thus, Rydberg ness of more of it than any other of the peri- outlined the directions which he recognized od, it is presumed that he would have de- at the ends of the spectrum of taxonomic scribed both trvmk and tail properly and have studies. known their approximate locations. His con- His case was overstated, because not all temporaries suffered, as do many of our own taxonomists are polarized to either one of the generation of taxonomists, with a myopia of two extremes. Rather, they fall remarkably understanding, and they are caught up in between, because most taxonomists let real- preconceived notions of the importance of ity, as they understand it, dictate the course plant characteristics. they should follow. It is true that reality is Rydberg let the characteristics of pods in subject to interpretation, and no one can ad- Astragalus, which he likewise monographed mit to imderstanding all of what is real. But (1929), dictate its segregation into more than few ignore the information that is available 30 genera. Carried to its logical conclusion, pod featiu-es would ultimately trap its user into absurdities. The use of pod types led Rydberg to place plants of almost identical .^v'^t.! vegetative appearance into separate genera

(i.e.. Astragalus oophorus Wats. var. caules- cens (Jones) Jones in the genus Phaca and As- tragalus beckwethii T. & G. in the genus Pliacomene). Uniformity in plant taxonomy is a trap laid by taxonomists, because the plants vary in accordance with dictates of genotype as that is influenced by habitat. Too, each taxonomic unit is the result of the genetic pathways that produced it, and each is unique both in pathway and in end result.

The system must be made to fit the plants, not the other way aroimd. Still, we allow ourselves to fall into the rut; perhaps we de- mand that we do. In a presentation to the Fourth Inter- national Botanical Congress Rydberg (1929a) outlined his approach to naming of plants.

The paper is entitled "Scylla or Charybdis," in recognition of the pitfalls of the descriptive taxonomist. Rydberg (1929a: 1539) notes, - Fig. 10. Marcus Eugene Jones (1891i, vmH ! 'Id "I will admit that phytographic work in the equipment and plant press, south of Salt Lake City (pho- last 35 vears has been overdone and I myself to Courtesy of Charles Walker). 156 Great Basin Naturalist Vol. 42, No. 2

to them; their interpretations approach real- that I am a splitter, but I have prided myself ity, more or less. on being consistent, as, for instance, in splitt- The thesis of Rydberg's speech (1929) re- ing some genera so as to make them equiva- volves aroimd the publication of the "Phy- lent to other genera in the same family or logenetic Method in Taxonomy" (Hall and tribe, but I have just discovered that in order Clements 1923). He (Rydberg 1929: 1540) to save already described species of other states, "We have seen the endless and in- botanists, even if they have a meager excuse finitesimal splitting into categories of the for their existence, I have added some of my German school and for twenty years have own of the same category and made my spe- been thoroughly disgusted with their absurd cies concept very inconsistent." nomenclature of trinomials, quadrinomials, Despite the frailties of his approach, Ryd- and polynomials. We believed that any form berg was a gentleman. He did not attack that is worth describing is worth a name and Jones in the press, as Jones attacked him. that a binomial is better than any other. But Time will serve to judge them both. we might have forgotten that there is a Edward L. Greene (Ewan 1950) was at- Scylla as well as a Charybdis." tacked more heartily than of The defence of Rydberg would have been any Jones's other contemporaries, easier except for his "leaping over the edge" and with more justification. in the direction of recognizing all variants at Of the 44 taxa named by him from species level. For the feeling expressed in his within Utah only 3 stand in modern tax- paper that he had created the species, and onomic treatments. He did not collect from had defended and upheld the creations of within the state, and relied on plants taken others, he states, "For years I have admitted by others for the basis of his taxonomic decisions. Perhaps Jones would have been less

harsh in criticism of Greene if he had chosen more substantial variants for naming. But Greene was the product of his background, as was Jones, and that background involved training in the clergy (he was a minister). He seems to have believed in the concept of fix- ity of species, and that any variant was there- fore worthy of a name. He also considered himself to be a linguist and would change names involving compounds of Latin and Greek together, without even knowing about the plants themselves. Time has served to judge him, at least as far as his Utah plant names are concerned, as incompetent. Jones judged him more harshly than that. The career of Marcus Eugene Jones ended suddenly, as noted by his daughter, Mabel Jones Broaddus. In Contributions to Western Botany. No. 18. (1935: 132) she wrote: "At this point on June 3, 1934, my father. Prof. Marcus E. Jones met with a fatal accident. He was returning alone from a day's field trip in the San Bernadino Mts. enjoyed in company with the Samuel B. Parish Bot- anical Society when, at an intersection in San Bernadino, his car was struck in the rear by Fig. 11. Charles Christopher Parry (ca 1870), student another car and overturned." of John Torrey, correspondent of Asa Gray, Sereno Wat- son, and George Engelmann, and important collector of Marcus Jones (Jones and Broaddus 1952) Utah types in 1874 (photo courtesy of Richard W. Pohl). was "... a man of unusual ability, strone^ June 1982 Welsh: Utah Plant Types 157

character, and amazing versatility. ... his Actinella biennis Gray Proc. Amer. Acad. 13: 373. central interest was always botany." 1878. Asteraceae. = Hymenoxys cooperi (Gray) Cock- erell Washington (?) Co., Arizona and Utah, Palmer 260, Jones's contributions to the understanding 1877(US!;ISC!;BRY!). are overwhelming. of Utah plant taxonomy Adiantum rimicola Slosson Bull. Torrey Bot. Club 41: He cannot be equaled or surpassed. .308. 1914. Polypodiaceae. = A. capillus-veneris var.

modestum f. rimicola (Slosson) Fern. San Juan Co, Arm- strong Canyon, Rydberg & Garrett 9423, 1913 (NY!). Utah Plant Types Agave scaphoidea Greenm. & Roush Ann. Missouri Bot. Card. 16: .391. 1929. Agavaceae. = A. utahensis Abies subalpina Engelm. ex Ward Ainer. Naturalist Engelm. var. utahensis Washington Co., St. George, Pal- 10: 555. 1876. Pinaceae. = A. lasiocarpa (Hook.) Nutt. mer sn, 1877 (MO?). Sanpete Co., Wasatch PL, above Gunnison, Ward sn, (?) Agave utahensis Engelm. in Wats. var. utahensis 1875 (US!). Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: 497. 1871. Abronia argillosa Welsh & Goodrich Great Basin Agavaceae. Washington Co., St. George, Palmer s.n. 1980. Nyctaginaceae. Co., Naturahst 40; 78. Grand 6 mi 1870 (US!). s Cisco, S, E., M. Welsh 16689, 1978 (BRY!;NY!; & Agoseris caudata Greene Leafl. Bot. Obs. & Grit. 2: US!;UT!). 124. 1911. Asteraceae. = A. glauca (Pursh) Raf. var. Abronia fallax Heimerl ex Rydb. Bull. Torrey Bot. laciniata (D.C. Eaton) Smiley Sevier Co., Salina Canyon, Club 29: 684. 1902. Nyctaginaceae. = A. elliptica A. Jones 5438, 1894 (US!;NY!). Nels. Salt Lake Co., Salt Lake City, Jones 1337, 1879 Agoseris confinis Greene Leafl. Bot. Obs. & Crit. 2: (US!;NY!;UTC!). 124. 1911. .\steraceae. = A. aurantiaca (Hook.) Greene Abronia fragrans Nutt. ex Hook. var. pterocarpa Piute Co., near Marysvale, Jones 5893z, 1894 (US!). Jones Contr. W. Bot. 11: 3. 1903. Nyctaginaceae. = A. Agoseris isomeris Greene Leafl. Bot. Obs. & Crit. 2: elliptica A. Nels. Tooele Co., Cottonwood, near Johnson 123. 1911. Asteraceae. = A. glauca (Pursh) Raf. Summit

Pass, Jones sn, 1900 (?). Co., Uinta Mts., Goodding 1397, 1902 (US!;RM!;ISC!). Abronia micrantha Torr. var. pedunculata Jones. Agoseris longirostris Greene Leafl. Bot. Obs. & Crit.

Proc. Calif. .\cad. II. 5: 716. 1895. Nyctaginaceae. = 2: 125. 191 1. Asteraceae. = A. aurantiaca (Hook.) Tripterocalyx micrantha (Torr.) Hook. Washington Co., Greene Sevier Co., Fish Lake, Jones 5743N, 1894 (US!;POM!). St. George, Jones 5101, 1894 (POM!;US!;BRY!;RM!). Agoseris taraxacoides Greene Leafl. Bot. Obs. Crit. Abronia nana Wats. Proc. Amer. Acad. 14: 294. 1879. & 2: 123. 1911. .\steraceae. = A. glauca (Pursh) Raf. Piute Nyctaginaceae. Beaver Co., near Beaver City, Palmer Co., near Marysvale, Jones 5372, 1894 (US!). 404 12, 1877 (ISC!). Aletes tenuifolia Coult. & Rose Contr. U.S. Natl. Abronia pumila Rydb. Bull. Torrey Bot. Club 29: 683. Herb. 7: 108. 1900. Apiaceae. = Musineon lineare 1902. Nyctaginaceae. = A. elliptica A. Nels. Emery Co., (Rydb.) Mathias Cache Co., Rydberg sn, 1895 Emery, Jones 554.5q, 1894 (US!;POM!;NY!). (US!;RM!;NY!). Abronia salsa Rydb. Bull. Torrey Bot. Club 29: 684. Allium biceptrum var. utahense Jones Contr. W. Bot. 1902. Nyctaginaceae. = A. elliptica A. Nels. Salt Lake 10: 33. 1902. Liliaceae. = A. biceptrum var. biceptrum Co., Salt Lake City, Watson 965, 1869 (NY!;US!). Salt Lake Co., City Creek Canyon, Jones 6647, 1898 Abronia turbinata var. marginata Eastw. Proc. Calif. (POM!). Acad. II, 6: 313. 1896. Nyctaginaceae. = A. fragrans Allium brevistylum Wats. Rep. U.S. Geol. Explor. Nutt. San Juan Co., Bartons range, Eastwood sn, 1895 40th Parallel, Bot. 5: ,350. 1871. Liliaceae. Summit Co., (CAS!). Uinta Mts., Watson 1180, 1869 (US!;NY!). Acerates var. erecta Trans. Amer. decumbens Durand Allium cristatum Wats. Proc. Amer. Acad. 14: 232. 11: = Phil. Soc. 174. 1860. .\sclepiadaceae. Asclepias as- 1879. Liliaceae. = A. nevadense Wats. Washington Co., (Decne.) Salt Salt City, perula Woodson Lake Co., Lake St. George, Palmer 454, 1877 (US!;NY!;ISC!). Carrington sn, 1857 (P). Allium pasgeyi A. & N. Holmgren Brittonia 26: 309. Acer kingii Britt. in Britt. & Shafer N. Amer. Trees 1974. Liliaceae. Box Elder Co., Howell Valley, Hol-

656. 1908. .\ceraceae. = A. negundo var. interius (Britt.) mgren et al. 13125, 1960 (NY!;BRY!;US!;UT!;UTC!). Sarg. Wasatch Mts., Watson 216, 1869 (US!). Allium reticulatum var. deserticola Jones Contr. W.

Acer sacharum Marsh, var. trilobum E. Murray Kal- Bot. 10: .30. 1902. Liliaceae. = A. macropetalum Rydb. mia 10: 2. 1980. Aceraceae. = A. grandidentatum Nutt. Grand Co., Cisco, Jones sn, 1890 (POM!;US!). Sanpete Co., Ephraim Canyon, Tidestrom 1003, 1908 Allium tribracteatum Torr. var. diehlii Jones Contr. (US!). W. Bot. 10: 18. 1902. Liliaceae. = A. brandegei Wats. Aconitum divaricatum Rydb. Fl. Rocky Mts. 314, Summit Co., Parleys Park, Jones 6590, 1901 1062. 1917. Ranunculaceae. = A. columbianum Nutt. in (US!;CAS!POM!). T. & G. Salt Lake Co., City Creek Canyon, Leonard Allocarya cognata Greene Pittonia 4: 235. 1901. Bo- 204182, 1884 (NY!). raginaceae. = Plagiobothrys scouleri (H. & A.) Johnst. Aconitum glaberrimum Rydb. Bull. Torrey Bot. Club Cache Co., Cache Valley, Mulford sn, 1888 (US!;NDG!). 29: 151. 1902. Ranunculaceae. = A. columbianum Nutt. Allocarya orthocarpa Greene Pittonia 3: 109. 1896.

in T. & G. Washington (?) Co., (S. Utah) Palmer 11, 1877 Boraginaceae. = Plagiobothrys leptocladus (Greene) (NY!). Johnst. Cache Co., Mulford sn, 1898 (US!;NDG!). 158 Great Basin Naturalist Vol. 42, No. 2

Ahine pabneri Rydb. Bull. Torrey Bot. Club 39: 315. Antirrhinum kingii Wats. Rep. U.S. Geol. Explor. 1912. Caryophyllaceae. = Stelkiria obtiisa Engelni. Bea- 40th Parallel, Bot. 5: 215. 1871. Scrophulariaceae. ver Co., Beaver Valley, Palmer 54, 1877 (NY!). Tooele Co., Stansbury Island, Watson 767, 1869 (US!). Amelanchier ittahensis Koehne Wiss. Bei. Progr. Aquilegia caerulea James var. calcarea Jones Proc. Falk- Realgyni. Berl. 95: 25. 1890. Rosaceae. Washing- Calif. Acad. II 5: 619. 1895. Ranunculaceae. = A. sco- ton Co., Bellevlew, Jones 1716, 1880 (US!;POM!;UTC!). pulonim var. calcarea (Jones) Munz Garfield Co., above Amelanchier titahensis Koehne var. cinerea Goodd- Cannonville, Jones 5312a, 1894 (POM!;US!). ing Bot. Gaz. 37: 55. 1904. Rosaceae. Washington Co., Aquilegia depauperata Jones Contr. W. Bot. 8: 1. St. George, Goodding 780, 1902 (RM!). 1898. Ranunculaceae. = A. flavescens Wats. Utah Co., Amsinckia eatotiii Suksd. Werdenda 1: 64. 1931. Bo- Provo Canyon, Jones sn, 1896 (POM!;US!). raginaceae. = A. retrorsa Suksd. Utali, Eaton 251, 1869 Aquilegia flavescens Wats. Rep. U.S. Geol. Explor. (CAS!). 40th Parallel, Bot. 5: 10. 1871. Ranunculaceae. Syn: A. Amsinckia utahensis Suksd. Werdenda 1: 106. 1931. depauperata Jones Salt Lake (?) Co., Wasatch Mts., Boraginaceae. = A. tesseUcita Gray Salt Lake Co., Salt Watson 35, 1869 (US!;NY!). Lake City, Vicker sn, 1901 (UC!). Aquilegia flavescens Wats. f. minor Tidestr. Amer. Amsonia brevifolia Gray Proc. Amer. Acad. 12: 64. Midi. Naturalist 1: 171. 1910. Ranunculaceae. Sanpete 1876. Apocynaceae. = A. jonesii Woodson Washington Co., Wasatch Plateau, Tidestrom sn (?). (?) Co., Palmer 302, 1877 (NY!). Aquilegia micrantha Eastw. Proc. Calif. Acad. II, 4: Amsonia eastwoodiana Rydb. Bull. Torrey Bot. Club 559. 1895. Ranunculaceae. Syn: A. pallens Payson; A. 40: 465. 1913. Apocynaceae. Grand Co., Moab vicinity, Payson; Rydberg & Garrett 8468, 1911 (NY!;RM!;US!). Uthophila A. rubicunda Tidestr. San Juan Co., near Bluff City, Wetherill sn, 1894 (CAS!;POM!). Amsonia latifolia Jones Contr. W. Bot. 12: 50. 1908. Payson Bot. 60: Apocynaceae. = A. jonesii Woodson Sevier Co., Monroe Aquilegia pallens Gaz. 375. 1915. Ra- Jones 6446, 1899 (CAS!;US!;RM!;POM!;BRY!;NY!). nunculaceae. = A. micrantha Eastw. San Juan Co., LaS- Androstephium breviflorum Wats. Amer. Naturalist al Creek, Payson 443, 1914 (?).

7: 303. 1873. Liliaceae. Syn: Brodiaea paysonii A. Nels. Aquilegia rubicunda Tidestr. Amer. Midi. Naturalist Kane Co., near Kanab, Thompson 59, 1872 (US!). 1: 168. 1910. Ranunculaceae. = A. micrantha Eastw. Anemone styhsa A. Nels. Bot. Gaz. 42: 52. 1906. Ra- Emery Co., near Emery, Tidestrom 1418, 1908 (US!). nunculaceae. = A. multifida Poir. Sevier Co., Fish Aquilegia scopulorum Tidestr. Amer. Midi. Naturalist Lake, Jones 5763, 1894 (US!";BRY!;RM!;POM!;NY!). 1: 167. 1910. Ranunculaceae. Sanpete Co., Wasatch Angelica dilatata A. Nels. in Coult. & Rose Contr. Peak, Tidestrom 1788, 1908 (US!). U.S. Natl. Herb. 12: 446. 1909. Apiaceae. = A. wheeleri Arabis demissa Greene var. russeola Rollins Rhodora Wats. Salt Lake Co., City Creek Canyon, Garrett 2127, 43: 387. 1941. Brassicaceae. Uintah Co., 18 mi N Vernal, 1907 (US!;RM!). Rollins 1757, 1937 (NY!;US!;CAS!;RM!;BRY!;UTC!). Angelica leporina Wats. Proc. Amer. Acad. 12: 252. Arabis diehlii Jones Contr. W. Bot. 14: 38. 1912. 1871. Apiaceae. = A. pinnata Wats. Wayne Co., Rabbit Brassicaceae. = A. pendulina Greene Beaver Co., Mt. Valley, Ward 612, 1875 (US!;NY!). Belknap, Jones sn, 1899 (POM!). Angelica pinnata Wats. Rep. U.S. Geol. Explor. 40th Arabis glabra (L.) Bernh. var. furcatipilis Hopkins Parallel, Bot. 5: 126. 1871. Apiaceae. Syn: A. leporina Rhodora 39: 109. 1937. Brassicaceae. Cache Co., Logan Wats. Summit Co., Uinta Mts. Watson 458, 1869 Canyon, Maguire .3437, 1932 (RM!;UTC!). (US!;NY!). Arabis hirsuta (L.) Scop. var. laevis Tuzon Bericht

Angelica wheeleri Wats. Amer. Naturalist 7: 301. Frein Verein Pflanzengeog. Syst. Bot. 1919: 43. 1921. 1873. Apiaceae. Svn: A. dilatata A. Nels. Northern and Brassicaceae. = A. hirsuta (L.) Scop. Utah Co., Ameri- Central Utah, Wheeler 1872 (US!). can Fork Canyon, Jones sn, 1880 (UTC!).

Anotites jonesii Greene Leafl. Bot. Obs. & Grit. 1: Arabis lasiocarpa Rollins Systematic Botany 6: 58. 102. 1905. Caryophyllaceae. = Silene menziesii Hook. 1981. Brassicaceae. Rich Co., 6 mi w Garden City, R.C. Utah Co., American Fork Canyon, Jones 1372, 1880 & K.W. Rollins 79307, 1979. (GH). (US!;RM!;POM!;UTC!). Arabis longirostris Wats. Rep. U.S. Geol. Explor. 40th Antennaria austromontana E. Nels. Proc. U.S. Natl. Parallel, Bot. 5: 73. 1871. Brassicaceae. = Streptanthella Mus. 23: 703. 1901. Asteraceae. = A. alpina var. media longirostris (Wats.) Rydb. Tooele Co., Stansbury Island, (Greene) Jeps. Piute Co., Marysvale, Jones 5522, 1894 Watson 72, 1869 (NY!). (US!). Arabis oreophila Rydb. Bull. Torrey Bot. Club 34: Antennaria dimorpha (Nutt.) T. & G. var. macroce- 437. 1907. Brassicaceae. = A. hjaUii Wats. Salt Lake phala D.C. Eaton Rep. U.S. Geol. Explor. 40th Parallel, Co., Rydberg & Carlton 6678, 1905 (NY!). Bot. 5: 186. 1871. Asteraceae. = A. dimorpha (Nutt.) T. Arabis pulchra Jones var. duchesnensis Rollins Sys- & G. Salt Lake Co., Salt Lake City, Watson 654, 1869 tematic Botany 6: 59. 1981. Brassicaceae. Duchesne Co., (US!). 3.8 mie Duchesne, R.C & K.W. Rollins 79113, 1979

Antennaria obtusata Greene Feddes Repert. 5: 241. (GH). 1908. Asteraceae. = A. parvifolia Nutt. Uintah Co., Arabis pulchra Jones var. pallens Jones Contr. W. Uinta Mts., Goodding 1209, 1902 (US!). Bot. 14: 42. 1912. Brassicaceae. Grand Co., Westwater, Anticlea vaginata Rydb. Bull. Torrey Bot. Club 39: Jones sn, 1891 (US!;NY!). 108. 1912. Liliaceae. = Zigadenus vaginatus (Rydb.) Arabis setulosa Greene Leafl. Bot. Obs. & Grit. 2: 81. Macbr. San Juan Co., Natural Bridges, Rydberg & Gar- 1910. Brassicaceae. = A. pendulina Greene Piute Co., rett 9407, 1911 (NY!;RM!;US!;BRY!;UT!). Marysvale, Jones 5.330, 1894 (US!;CAS!;POM!;NY!). June 1982 Welsh: Utah Plant Types 159

Arabis stokesiae Rydb. Fl. Rocky Mts. 361, 1062. Asclepias Jeucophylla Engelm. in Parry Amer. Natu- 1917. Brassicaceae. Syn: A. divaricarpa A. Nels.; = A. ralist 9: .348. 1875. Asclepiadaceae. = A. erosa Torr. confinis Wats. Salt Lake Co., Parleys Canvon, Stokes sn, Washington Co., near St. George, Parry 207, 1874 (MO). 1901 (US!;NY!). Asclepias ruthiae Maguire & Woodson Ann. Missouri Aragallus majusculus Greene Proc. Biol. Soc. Wash. Bot. Card. 28: 245. 1941. Asclepiadaceae. Emery Co., 18: 12. 1905. Fabaceae. = Oxtitwpis sericea Nutt. Gar- Calf Spring Canvon, Maguire 18310, 1940 (US!;Ufc!). field Co., Henry Mts., Jones 5674, 1894 (US!;PO.\l!;NY!). Asclepias welshii N. & P. Holmgren Brittonia 31: 110. Arctoniecon humilis Gov. Proc. Biol. Soc. Washington 1979. .\sclepiadaceae. Kane Co., Coral Pink Sand Dunes, 7: 67. 1892. Papaveraceae. Washington Co., St. George, N. & P. Holmgren 9009. 1978 (NY!;BRY!;UTC!). Parry sn, 1874 iUS!:NY:;ISG!;NDG!). Astephanus utahensis Ejiglem. in Parry Amer. Natu- Arenaria fendJeri (Rydb.) Fern. ssp. brevifolia Ma- ralist 9: 349. 1875. Asclepiadaceae. = Cynanchum uta- guire Madrono 6; 2.3. 1941. Carvophvllaceae. = A. fend- hense (Engelm.) Woodson Washington Co., near St. leri var. brevifolia (Magiiire) Magiiire Grand Co., LaSal George, Parry 209, 1874 (ISC!). Mts., Magiiire 17972, 1933 (UTC!). Aster canescens Pursh var. aristatus Eastw. Proc. Ca- Bull. Arermria filiortim Maguire Torrey Bot. Club 73: lif. Acad. II. 6: 296. 1896. Asteraceae. = Machaeran- 326. 1946. Carvophvllaceae. Iron Co., beach, Navajo thera canescens (Pursh) Grav San Juan Co., Willow Lake, Magiiire 19472, 1940 (\Y!;BRY!). Creek, Eastwood 45, 1895 (CAS!). Arenaria hookeri Nutt. in T. & G. var. desertorum Aster glancodes Blake ssp. pulcher Blake Proc. Biol. Maguire Amer. Midi. Naturalist 46: 506. 1951. Carvo- Soc. W'ashington 35: 174. 1922. Asteraceae. = A. glanc- phyllaceae. Duchesne Co, near Duchesne, Riplev & odes Blake var. pulcher (Blake) Kearnev & Peebles Kane Barneby8701, 1947 (US!;NY!). Co., Elk Ranch, Jones 6037, 1894 (US!;RM!;POM!). Arenaria kingii (Wats.) Jones ssp. plateauensis Ma- Aster glaucus var. wasatchensis Jones Proc. Calif. guire Bull. Torrey Bot. Club 74: 54. 1947. Carvophvl- .\cad. II. 5: 694. 1895. Asteraceae. = A. wasatchensis laceae. = A. kingii var. plateauensis (Maguire) Reveal Jones Blake Piute Co., near Marvsvale, Jones 5861, 1894 Iron Co., Cedar Breaks rim, Maguire 19024, 1940 (US!;RM!:BRY!;POM!;NY!). (US!;NY!;UTC!). Aster halophilus Greene Leafl. Bot. Obs. & Grit. 2: 8. Arenaria niittallii Pa.\ var. gracilipes Jones Proc. Ca- 1909. Asteraceae. = A. chiletisis ssp. adscendens (Lindl.) hf. Acad. II, 5: 626. 1895. Caryophyllaceae. = A. niit- Cronq. Salt Lake Co., Beck's Hot Springs, Garrett 2057, 1906 (UT!;NdDG!). tallii Pax Piute Co., Brigham ' Peak, Jones .5951, 1894 (US!). Aster kingii D.C. Eaton in Wats. Rep. U.S. Geol. Ex- = Argemone corymbosa Greene ssp. arenicola G. B. plor. 40th Parallel, Bot. 5: 141. 1871. Asteraceae. Ownbey Mem. Torrey Bot. Club 21(1): 118. 1958. Pa- Machaeranthera kingii (D.C. Eaton) Cronq. & Keck. Salt paveraceae. Emerv Co., 37.9 mi s\v Green River, Own- Lake Co., Cottonwood Canvon, Watson 507, 1869 bey 2146, 1954 (US!:CAS!;RM!:BRY!). (US!;NY!). Arnica arachtwidea Rydb. N. Amer. Fl. 34: 353. Aster leucopsis Greene Leafl. Bot. Obs. & Crit. 2: 8. 1927. .\steraceae. = A. mollis Hook. Salt Lake Co., Big 1909. Asteraceae. = A. chilensis ssp. adscendens (Lindl.) Cottonwood Canvon, Rvdberg & Carlton 6609, 1905 Cronq. Salt Lake Co., Salt Lake Citv, Garrett 1694, 1905 (NY!). (UT!;NDG!). Arnica catidata Rydb. Bull. Torrev Bot. Club 37: 463. Aster thermalis Jones Proc. Calif. Acad. II, 5: 694. 1910. .\steraceae. = A. mollis Hook. Salt Lake Co., Big 1895. Asteraceae. = A. pauciflorus Nutt. Sevier Co., Cottonwood Canyon. Garrett 1547, 1894 (RM!;NY!). Monroe, Jones 5410, 1894 (POM!;US!). = Arnica chamissonis Less. var. longinodosa A. Nels. Aster venustus Jones Zoe 2: 247. 1891. .\steraceae. Bot. Gaz. 30: 199. 1900. Asteraceae. = A. mollis Hook. Machaeranthera venusta (Jones) Cronq. & Keck. Grand Piute Co., Marvsvale, Jones 5883, 1894 (RM!;NY!l Co., Cisco, Jones sn, 1890 (POM!:US!;C.\S!; Arnica jonesii Rydb. Fl. Rockv Mts. 979. 1917. .\ste- RM!;BRY!;NY!). var. raceae. = A. latifolia Bong. Salt Lake Co., Alta, Jones Astragalus amphioxys Gray cymbellus Jones 119, 1879 (NYI;POM!;UTCl;NDG!). Rev. Astragalus 215. 1923. Fabaceae. = A. cymboides Arnica longifolia D.G. Eaton in Wats. Rep. U.S. Jones Emery Co., San Rafael Swell, Jones sn, 1914 Geol. E.xplor. 40th Parallel, Bot. 5: 186. 1871. .Aste- (POM!). raceae. Summit Co., Uinta Mts., Watson 655, 1869 Astragalus ampullarius Wats. Amer. Naturalist 7: (NY!). 300. 1873. Fabaceae. Kane Co., Kanab, Thompson sn, Arnica ovata Greene Pittonia 4: 161. 1900. Aste- 1872 (GH;US!). raceae. = A. mollis Hook. Salt Lake Co., Alta, Jones Astragalus araneosus Sheld. Bull. Geol. & Nat. Hist. 1128, 1879 (POM!:NY!;BRY!;UTC!;NDG!). Surv. Minnesota 9: 170. 1894. Fabaceae. = A. lentigi- Artemisia norvegica Fries var. piceetorum Welsh & nosus Dougl. var. araneosus (Sheld.) Barneby Beaver Frisco, Goodrich in Welsh Brittonia .33: 295. 1981. Asteraceae. Co., Jones 1807, 1880 (US!;RM!;BRY!; Duchesne Co., Garfield Basin, Welsh et al 18970, 1979 POM!:NY!;UT!;UTC!). (BRY!:NY!). Astragalus argillosus Jones Zoe 2: 241. 1891. Fa- Asclepias itivohicrata Engelm. var. tomentosa Eastw. baceae. = A. flavus Nutt. var. argillosus (Jones) Barneby Zoe 4: 120. 1893. Asclepiadaceae. = A. macrospenua Emerv Co., Greene River, Jones sn, 1890 (POM!; Eastw. Grand Co., Courthouse Wash, Eastwood sn, 1892 US!;B'rY!). (CAS!;NY!). Astragalus argophyllus Nutt. var. cnicensis Jones Rev. Astragalus 207. 1923. Fabaceae. = A. argophyllus Asclepias labriformis Jones Proc. Calif. Acad. II, 5: 708. 1895. Asclepiadaceae. Wavne Co., Capitol Wash, var. martinii Jones Utah Co., Thistle, Jones sn, 1898 Jones .5650, 1894 (P0M!;US!;RM'!;BRY!;NY!). (POM!). 160 Great Basin Naturalist Vol. 42, No. 2

Astragalus arietinus Jones Proc. Calif. Acad. II. 5: Astragalus detritalis Jones Contr. W. Bot. 13: 9. 653. 1895. Fabaceae. = A. cibarius Sheld. Sanpete Co., 1910. Fabaceae. Syn: A. spectabilis C.L. Porter Duch- Fairview, Jones 5544o, 1894 (POM!). esne Co., near Theodore, Jones sn, 1908 Astragalus asclepiadoides Jones Zoe 2: 238. 1891. Fa- (POM!;BRY!;NY!). baceae. Grand Co., Cisco, Jones sn, 1889 (POM!). Astragalus diehlii Jones Rev. Astragalus 194. 1923. Astragalus beckwithii T. & G. var. beckwithii Rep. Fabaceae. = A. flexuosus var. diehlii (Jones) Barneby U.S. Explor. & Surv. R.R. Pacific 2: 120. 1855. Fabaceae. Carbon Co., Farnham, Jones sn, 1898 (POM!;RM!; Tooele Co.?, west of Lone Rock, Beckwith, 1854 (GH). NY!;BRY!;UTC!). Astragalus beckwithii T. & G. var. purpureus Jones Astragalus dodgeanus Jones Zoe 3: 289. 1893. Fa- Zoe 3: 288. 1893. Fabaceae. Juab Co., Deep Creek Mts., baceae. = A. wingatanus Wats. Grand Co., Thompsons Jones sn, 1891 (US!;POM!;BRY!;NY!). Springs, Jones sn, 1891 (POM!;BRY!;NY!). Astragalus castaneiformis var. consobrinus Barneby Astragalus duchesnensis Jones Contr. W. Bot. 13: 6. Amer. Midi. Naturalist 41: 496. 1949. Fabaceae. = A. 1910. Fabaceae. Duchesne Co., Theodore to Myton, consobrinus (Barneby) Welsh Wayne Co., Bicknell, Rip- Jones sn, 1908 (POM!;CAS!;RM!;BRY!;US!;NY!). ley & Barneby 8605, 1947 (CAS!;RM!;NY!;UTC!). Astragalus ensiformis Jones var. gracilior Barneby Astragalus chamaeleuce Gray in Ives var. pan- Proc. Calif. Acad. IV. 25: 158. 1944. Fabaceae. = A. en- guicensis Jones Proc. Calif. Acad. II. 5: 671. 1895. Fa- siformis Jones Washington Co., s Veyo, Ripley & Barn- baceae. = A. argophyllus var. panguicensis (Jones) Jones eby 4951, 1942 (CAS!;NY!). Garfield Co., Pangiiitch Lake, Jones 6023f, 1894 (POM!). Astragalus episcopus Wats. Proc. Amer. Acad. 10: Astragalus chloodes Barneby Leafl. W. Bot. 5: 6. 345. 1875. Fabaceae. Kane (?) Co., Bishop sn, 1872 1947. Fabaceae. Uintah Co., 6 mi se Jensen, Ripley & (GH;US!;NY!). Barneby 7797, 1946 (CAS!;RM!;NY!;UTC!). Astragalus equisolensis Neese & Welsh Rhodora 83: Astragalus cibarius Sheld. Bull. Geol. & Nat. Hist. 457. 1981. Fabaceae. Uintah Co, s Jensen, Neese & Surv. Minnesota 9: 149. 1894. Fabaceae. Syn: A. ariet- Welsh 7380, 1979 (BRY!;NY!;US!;POM!;UTC!). inus Jones Utah Co., Utah Valley, Jones 1679, 1880 Astragalus eremiticus Sheld. Bull. Geol. & Nat. Hist. (MINN;POM!;US!;NY!). Surv. Minnesota 9: 161. 1894. Fabaceae. Washington Astragalus cicadae var. laccoliticus Jones Proc. Calif. Co., Beaver Dam Mountains, Parry 45, 1874 (MINN;

Acad. II. 5; 672. 1895. Fabaceae. = A. chamaeleuce US!;ISC!;NY!). Gray Garfield Co., Henry Mts., Jones 5658q, 1894 Astragalus eurekensis Jones Contr. W. Bot. 8: 12. (POM!). 1898. Fabaceae. Syn: Xylophacos medius Rydb. Juab Astragalus coltonii Jones var. aphyllus Jones Rev. As- Co., Eureka, Jones sn, 1891 (POM!;BRY!). tragalus 71. 1923. Fabaceae. Syn: A. coltonii var. coltonii Astragalus flavus Nutt. in T. & G. var. candicans Sevier Co., near Richfield, Jones sn, 1898 (POM!). Gray Proc. Amer. Acad. 12: 54. 1876. Fabaceae. Syn: A. Astragalus coltonii Jones var. coltonii Zoe 2: 237. confertiflorus Gray Sevier Co., near Richfield, Ward 1891. Fabaceae. SynAstragalus coltonii var. aphyllus 246, 1875 (GH;NY!). Astragalus hamiltonii C.L. Porter 54: 159. Jones Carbon Co., Castle Gate, Jones sn, 1889 Rhodora (POM!;US!;NY!). 1952. Fabaceae. Uintah Co., 5 mi sw Vernal, Hamilton & Beathsn, 1952 (RM!). Astragalus coltonii Jones var. moabensis Jones Contr. Astragalus harrisonii Barneby Mem. New York Bot. W. Bot. 8: 11. 1898. Fabaceae. Syn: Homalobus canovi- Card. 13: 271. 1964. Fabaceae. Wayne Co., near Fruita, rens Rydb. San Juan Co., Monticello, Eastwood 9, 1892 Barneby 13131, 1961 (CAS!;US!;BRY!;NY!;UTC!). (POM!). Astragalus haydenianus Gray ex Brandegee var. ma- Astragalus convallarius Greene var. finitimus Barn- jor Jones Zoe 2: 240. 1891. Fabaceae. = A. bisulcatus eby Leafl. W. Bot. 7: 192. 1954. Fabaceae. Washington var. major (Jones) Welsh Kane Co., Johnson, Jones sn, Co., 3 mi s Enterprise, Ripley & Barneby 4967, 1942 1890(POM!;BRY!;NY!). (CAS!). Astragalus ibapensis Jones Zoe 3: 290. 1893. Fa- Astragalus cottamii Welsh Rhodora 72: 189. 1970. Fa- baceae. = A. diversifolius Gray Juab Co., Deep Creek baceae. San Juan Co., e Clay Hills Divide, Welsh 5207, Mts., Jones sn, 1891 (POM!). 1966 (BRY!;NY!;ISC!). Astragalus iselyi Welsh Great Basin Naturalist 34: Astragalus cronquistii Barneby York Bot. Mem. New 305. 1974. Fabaceae. San Juan Co., Brumley Ridge, Card. 13: 258. 1964. Fabaceae. San Juan Co., w side Welsh 10970, 1971 (BRY!;UT!;POM!;UTC!;ISC!). Comb Wash, Cronquist 9123, 1961 (NY!;UTC!). Astragalus jejunus Wats. Rep. U.S. Geol. Explor. 40th Astragalus Proc. Calif. Acad. II, 5: cymboides Jones Parallel, Bot. 5: 73. 1871. Fabaceae. Summit (?) Co., 650. 1895. Fabaceae. Syn: A. amphyoxys var. cymbellus Bear River Valley, Watson 279, 1869 (NY!). Jones Emery Co., Huntington, Jones 5464j, 1894 Astragalus junceus var. attenuatus Jones Rev. Astra- (POM!;US!;N'y!). galus 76. 1923. Fabaceae. = A. convallarius Greene Astragalus desereticus Barneby Mem. New York Bot. Carbon Co., Price, Jones sn, 1898 (POM!;BRY!;NY!). Card. 13: 635. 1964. Fabaceae. Sanpete Co., Indianola, Astragalus lentiginosus Dougl. ex Hook. var. char-

Tidestrom 2249, 1909 (GH). taceus Jones Proc. Calif. Acad. II. 5: 673. 1895. Fa- Astragalus desperatus Jones var. desperatus Zoe 2: baceae. = A. lentiginous var. araneosus (Sheld.) Barn- 243. 1891. Fabaceae. Grand Co., near Cisco, Jones sn, eby Sanpete Co., Ephriam, Jones 5627m, 1894 (POM!). 1890(US!;RM!;BRY!). Astragalus lentiginosus Dougl. ex Hook. var. pohlii Astragalus desperatus Jones var. petrophilus Jones Welsh & Barneby Isleya 2: 1. 1981. Fabaceae. Tooele Rev. Astragalus 204. 1923. Fabaceae. Emery Co., San Co., 4.5 mi N Vernon, Welsh et al. 16743, 1978 Rafael Swell, Jones sn, 1914 (POM!;RM!;BRY!;NY!). (BRY!;NY!;UTC!). June 1982 Welsh: Utah Plant Types 161

var. vitreus Astragalus lentiginosus Dougl. ex Hook. Astragalus stocksii Welsh Great Basin Naturalist 34: Wash- Bameby Leafl. W. Bot. 4: 119. 1945. Fabaceae. 307. 1974. Fabaceae. = A. henriinontanensis Welsh ington Co., 5 mi w Leeds, Maguire & Blood 4413, 1933 Garfield Co., Henry Mts., Welsh 11740, 1972 (UTC!). (BRY!;ISC!). Astragalus lentiginosus Dougl. ex Hook. var. wah- Astragalus straturensis Jones Contr. W. Bot. 8: 19. weapensis Welsh Great Basin Naturahst 38: 286. 1978. 1898. Fabaceae. Washington Co., Silver Reef, Jones Fabaceae. Kane Co., Four Mile Bench, Welsh 12426, 5175, 1894 (POM!;US!;RM!;NY!;BRY!). 1974 (BRY!;ISC!). Astragalus striatiflorus Jones Proc. Calif. Acad. II. 7: Astragalus limnocharis Bameby Leafl. W. Bot. 4: 643. 1895. Fabaceae. Washington Co., Springdale, Jones 236. 1946. Fabaceae. Kane Co., Navajo Lake, Maguire 6080k, 1894 (POM). 19474, 1940(NY!;RM!;UTC!). Astragalus subcinereus Gray var. basalticus Welsh Astragalus nidularius Bameby Leafl. W. Bot. 8: 16. Great Basin Naturalist 38: 302. 1978. Fabaceae. Sevier Co., White Canyon, Barneby 1956. Fabaceae. San Juan Co., 10 mi s Fremont Jet., Welsh et al. 6447, 1967 12778, 1955 (CAS!;US!;RM!;NY!;UT!;UTC!)'. (BRY!;ISC!). Astragalus palans Jones Zoe 4: 37. 1893. Fabaceae. Astragalus subcinereus Gray var. subcinereus Proc. Co., = A. lentiginosus var. palans (Jones) Jones San Juan Amer. Acad. 13: .366. 1878. Fabaceae. Syn: A. sileranus (RM!;NY!). Montezuma Canyon, Eastwood sn, 1892 Jones; A. sileranus var. caraicus Jones Southern Utah, peabodianus Jones Zoe 3; 295. 1893. Fa- Astragalus (Northern Arizona ?), Palmer 117 1877 (US!;ISC!;NY!). baceae. = A. pubentissinuis var. peabodianus (Jones) Astragalus tegetarius Wats. var. rotundus Jones Proc. Welsh Grand Co., Thompsons Springs, Jones sn, 1891 Calif. Acad. II. 5; 650. 1895. Fabaceae. = A. kent- (POM!). rophyta var. implexus (Canby) Barneby Wayne Co., near Astragalus perianus Barneby Mem. New York Bot. Loa, Jones 5649b, 1894 (POM!;UC!). Card. 13(2): 973. 1964. Fabaceae. Piute Co, n Bullion Astragalus tetrapterus Gray Proc. Amer. Acad. 13: Creek, Rydberg & Carlton 7104, 1905 (NY!;RM!;US!). .369. 1878. Fabaceae. Washington Co., 25 mi n St. Astragalus pictus var. angustus Jones Zoe 4: 37. 1893. George, Palmer 111, 1877 (NY!;ISC!). Fabaceae. = A. ceramicus Sheld. San Juan Co., Mon- Astragalus thompsonae Wats. Proc. Amer. Acad. 10; tezuma Canyon, Eastwood 7, 1892 (POM!;US!). .344. 1875. Fabaceae. = A. moUissimus var. thompsonae Astragalus pictus var. magnus Jones Rev. Astragalus (Wats.) Barnebv Kane Co., Kanab, Thompson sn, 1872 109. 1923. Fabaceae. = A. ceramicus Sheld. Washington (GH;US!). Acad. 13: 367. Co., Silver Reef, Jones 5160, 1894 (POM!;RM!). Astragalus ursinus Gray Proc. Amer. = var. ursinus (Gray) Astragalus pinonis Jones Contr. W. Bot. 8: 14. 1898. 1878. Fabaceae. A. lentiginosus Valley, Palmer sn, 1877 Fabaceae. Beaver Co., Frisco, Jones sn, 1880 (POM!). Barneby Iron (?) Co., Bear Astragalus preussii Gray var. latus Jones Zoe 4: 36. (GH!;NY!;ISC!). Astragalus wardii Gray Proc. Amer. Acad. 12: 55. 1893. Fabaceae. = A. preussii Gray var. preussii Emery 1876. Fabaceae. Garfield Co., Aquarius PI., Ward Co., Green River, Jones sn, 1891 (NY). (?) 424, 1875 (GH;US!;NY!). Astragalus preussii Gray var. sulcatus Jones Zoe 4: Astragalus woodruffii Jones Rev. Astragalus 77. 1923. 37. 1893. Fabaceae. = A. eastwoodiae ' ]ones Southeast Fabaceae. Emery Co., San Rafael Swell, Jones sn, 1914 Utah, Cane Spring, Eastwood sn, 1892 (CAS!;BRY!; (POM!;US!;CAS!';RM!;BRY!;NY!;UTC!). NY!;POM!). Astragalus zionis Jones Proc. Calif. Acad. II, 5: 652. Astragalus rafaelensis Jones Rev. Astragalus 146. 1895. Fabaceae. Washington Co., Springdale, Jones 1923. Fabaceae. Emery Co., Cedar Mt., Jones sn, 1915 5261w, 1894 (POM!;US!;RM!;BRY!). (POM!;CAS!;RM!;BRY!;NY!;UTC!). Atriplex bonnevillensis C.A. Hanson Stud. Syst. Bot. Astragalus sabulosus Jones Zoe 2: 239. 1891. Fa- (BRY) 1: 2. 1962. Chenopodiaceae. Millard Co., Pine baceae. Grand Co., Cisco, Jones sn, 1890 (POM!;BRY!). Valley playa, Hanson 354, 1960 (BRY!;UTC!). Astragalus saurinus Barneby Leafl. W. Bot. 8: 17. Atriplex caput-medusae Eastw. Proc. Calif. Acad. II. 1956. Fabaceae. Uintah Co, 6 mi N Jensen, Holmgren & 6: 316. 1896. Chenopodiaceae. = A. argentea var. caput- Tillett 9527, 1953 (NY!;CAS!;RM!). medusae (Eastw.) Fosberg San Juan Co., Recapture Astragalus serpens Jones Proc. Calif. Acad. II, 5: 641. Creek, Eastwood 116, 1895 (US!;CAS!). 1895. Fabaceae. Wayne Co., Loa Pass, Jones 56.39i, 1894 Atriplex cornuta Jones Proc. Calif. Acad. II. 5; 718. (POM!;US!;NY!). 1895. Chenopodiaceae. = A. saccaria Wats. Emery Co., Astragalus sesquiflorus Wats. Proc. Amer. Acad. 10: Green River, Jones 5481, 1894 (POM!;US!). 345. 1875. Fabaceae. Kane Co., near Kanab?, Bishop sn, Atriplex cuneata A. Nels. Bot. Gaz. 34: 357. 1902. 1873 (GH;US!). Chenopodiaceae. Emery Co., Emery, Jones 5443, 1894 Astragalus sileranus Jones Zoe 2: 242. 1891. Fa- (US!;RM!). baceae. = A. subcinereus Gray Kane Co., Sink Valley, Atriplex cuneata A. Nels. ssp. introgressa C.A. Han- Jones sn, 1890 (POM!;CAS!;US!;BRY!;NY!). son Stud. Syst. Bot. (BRY) 1: 4. 1962. Chenopodiaceae. Astragalus sileranus Jones var. caraicus Jones Proc. Carbon Co., Wellington, Hanson 346, 1961

Calif. Acad. II. 5: 642. 1895. Fabaceae. = A. sub- (BRY!;POM!). Rydb. Bull. Torrey Bot. Club 39; cinereus Gray Kane (?) Co., Elk Ranch, Jones 6036, 1894 Atriplex garrettii (POM!;BRY!). 312. 1912. Chenopodiaceae. Grand Co., Moab vicinity, Astragalus spectabilis C.L. Porter Rhodora 54; 160. Rydberg & Garrett 8465, 1911 (NY!;US!;UT!). Proc. Calif. Acad. II, 5: 1952. Fabaceae. = A. detritahs Jones Uintah Co., 5 mi Atriplex graciliflora Jones Co., Blue Valley, sw Vernal, Porter 5309, 1950 (RM!;CAS!;BRY!; 717. 1895. Chenopodiaceae. Wayne (POM!;US!;RM!;BRY!). POM!;NY!). Jones 5697, 1894 162 Great Basin Naturalist Vol. 42, No. 2

Atriplex nuttallii Wats. var. utahensis Jones Contr. Bigelovia leiosperma Gray var. abbreviata Jones W. Bot. 11: 19. 1903. Chenopodiaceae. = A. tridentata Proc. Calif. Acad. II, 5: 693. 1895. Asteraceae. = Kuntze Salt Lake Co., Salt Lake City, Jones 1760, 1879? Chrysothamnus nauseosus (Pallas) Britt. Sevier Co., (UC). Clear Creek Canyon, Jones 6105, 1894 (POM!;US!;NY!). Atriplex rydbergii Standi. N. Amer. Fl. 21: 47. 1916. Bigelovia menziesii var. scopuhrum Jones Proc. Calif. Chenopodiaceae. = A. argentea var. argentea San Juan Acad. II, 5: 692. 1895. Asteraceae. = Haplopappus sco- (?) Co., s Moab, Rydberg & Garrett 9110, 1911 pulorum (Jones) Blake Iron Co., near Cedar City, Jones (US!;NY!). 5204v, 1894 (POM!;US!). Atriplex subdecumbens Jones Proe. Calif. Acad. II. 5: Bigelovia turbinata Jones Proc. Calif. Acad. II, 5: 716. 1895. Chenopodiaceae. = A. truncata (Torr.) Gray 691. 1895. Asteraceae. = Chrysothamnus nauseosus var. Sevier Co., Fish Lake, Jones 5745, 1894 (POM!;US!). turbinatus (Jones) Blake Garfield (?) Co., Canaan Ranch, Atriplex tenuissima A. Nels. Bot. Gaz. 34: 359. 1902. Jones 6066c, 1894 (POM!;US!). Chenopodiaceae. = A. wolfii Wats. Sanpete Co., Gun- BrickelUa atractyloides Gray Proc. Amer. Acad. 8: nison, Jones 6.525, 1900 (US!;RM!;POM!;NY!). 290. 1870. Asteraceae. Utah?, Colorado River, Palmer Atriplex tridentata Kuntze Rev. Gen. 2: 546. 1891. sn, 1870 (US!). Chenopodiaceae. Syn: A. nuttallii var. utahensis Jones BrickelUa linifolia D.C. Eaton Rep. U.S. Geol. Ex- Box Elder Co., Corinne, Kuntze .3084, 1874 (NY!). plor. 40th Parallel, Bot. 5: 137. 1871. Asteraceae. = B. Atriplex welshii C. A. Hanson Stud. Syst. Bot. (BRY) oblongifolia Wats. var. linifolia (D.C. Eaton) Robins. 1: 1. 1962. Chenopodiaceae. Grand Co., 4 mi s Cisco, Utah Co., Jordan Valley, American Fork, Watson 493, Hanson ,322, 1961 (BRY!;ISC!). 1869 (US!;CAS!). Atropis laevis var. rigida Beal Grasses N. Amer. 2: BrickelUa watsonii Robins. Mem. Gray Herb. 1: 42. 578. 1896. Poaceae. = Poa canbyi (Scribn.) Howell 1917. Asteraceae. = B. microphylla var. watsonii (Rob- Tooele Co., Lake Point, Jones 1021, 1879(US!). ins.) Welsh Utah Co., American Fork Canyon, Watson Aulospermum minimum Mathias Ann. Missouri Bot. 494, 1869 (US!;NY!). Card. 17: 353. 1930. Apiaceae. = Cijmopterus minimus Buddleja utahensis Gov. Proc. Biol. Soc. Washington (Mathias) Mathias Iron Co., Cedar Breaks, Mathias 723, 7: 69. 1892. Loganiaceae. Washington Co., St. George, 1929 (MO;CAS!;BRY!). Palmer 400, 1877 (US!;NY!;BRY!;ISC!). Aulospermum rosei Jones in Coult. & Rose Contr. Caesalpinia repens Eastw. Zoe 4: 116. 1893. Fa- U.S. Natl. Herb. 7: 179. 1900. Apiaceae. = Cijmopterus baceae. = Hoffmanseggia repens (Eastw.) Cockerell rosei (Jones) Jones Sevier Co., Richfield, Jones 30, 1899 Grand Co., Courthouse Wash, Eastwood sn, 1892 (US!;POM!). (CAS!;US!). Bahia desertorum Jones Zoe 2: 249. 1891. Asteraceae. Calamagrostis scopulorum Jones Proc. Calif. Acad. II, = Platyschkuhria integrifolia var. deserertorum (Jones) 5: 722. 1895. Poaceae. Syn; C. scopulorum var. lucidu- Ellison Grand Co., Cisco, Jones sn, 1890 (US!;POM!). lum Kearney Washington Co., Springdale, Jones 6075, Bahia ourolepis Blake Proc. Biol. Soc. Washington 35: 1894(POM!;NY!). 175. 1922. Asteraceae. = Platyschkuhria oblongifolia Calamagrostis scopulorum Jones var. lucidula (Gray) Rydb. Emery Co., Green River, Jones 5482#1 Kearney USDA Agrostol. Bull. = 1894 (US!;POM!). 11: 33. 1898. Poaceae. C. scopulorum Jones Salt Lake Co., Alta, Jones 1145, Bahamorhiza hispidula Sharp Ann. Missouri Bot. 1879 (POM!;NY!;BRY!;UTC!). Card. 22: 137. 1935. Asteraceae. = B. hookeri var. hispi- Calochortus aureus Wats. Amer. Naturalist 7: 303. dula (Sharp) Cronq. Tooele Co., Lake Point, Jones 1727, 1880 (UT!;UTC!). 1873. Liliaceae. Syn: C. nuttallii var. aureus (Wats.) Ownbey Kane (?) Co., "S. Utah" Thompson, Batidophaca sabinarum Rydb. N. Amer. Fl. 24: 320. 1872 (US!). Calochortus flexuosus Wats. 1929. Fabaceae. = Astragalus argophyllus var. pan- Amer. Naturalist 7: 303. 1873. guicensis (Jones) Jones Iron Co., Cedar Canyon, Garrett Liliaceae. Kane (?) Co., "S. Utah or N. Arizona", 2660, 1920 (NY!). Thompson sn, 1872 (GH).

Betula utahensis Britt. Bull. Torrey Bot. Club 31: Camassia quamash (Pursh) Greene ssp. utahensis 165. 1904. Betulaceae. = B. occidentalis x B. papyrifera Gould Amer. Midi. Naturalist 28: 740. 1942. Liliaceae. Salt Lake Co., City Creek Canyon, Stokes sn, 1900 = C. quamash (Pursh) Greene Cache Co., Blacksmith

(NY!). Canyon, Maguire & Maguire 3265, 1932 (?). Bigelovia douglasii var. spathulata Jones Proc. Calif. Camissonia gouldii Raven Contr. U.S. Natl. Herb. 37: Acad. II, 5: 690. 1895. Asteraceae. = Chrysothamnus 368. 1969. Onagraceae. Washington Co., N of St. viscidiflorus var. pumilus (Nutt.) Jeps. Sevier Co., Fish George, Gould 1423, 1941 (US!;CAS!). Lake, Jones 5758m, 1894 (POM;US!). Capnodes brachycarpum Rydb. Bull. Torrey Bot. Bigelovia glareosa Jones Zoe 2: 247. 1891. Asteraceae. Club 34: 426. 1907. Fumariaceae. = Corydalis caseana = C. nauseosus (Pallas) Britt. var. glareosa (Jones) ssp. brachycarpa (Rydb.) M. Ownbey Salt Lake Co., Welsh Piute Co., Marysvale, Jones sn, 1890 (POM). Alta, Jones 1197, 1879 (NY!;US!;RM!;POM!;BRY!;UTC!). Bigelovia howardii var. attenuata Jones Proc. Calif. Cardamine cordifoUa Gray var. pubescens Gray ex Acad. II, 5: 691. 1895. Asteraceae. = Chrysothamnus Schuiz Engl. Bot. Jahrb. 32: 439. 1903. Brassicaceae. = parryi var. attenuatus (Jones) Kittell Piute Co., Mary- C. cordifoUa Gray Garfield (?) Co., Thousand Lake Mtn., svale, Jones 5912, 1894 (US!;RM!;NY!). Ward .396, 1875 (US!). Bigelovia leiosperma Gray Syn. Fl. 1(2): 129. 1884. As- Cardamine palustris var. jonesii Kuntze Rev. Gen. 1: teraceae. = Chrysothamnus nauseosus var. leiosperma 125. 1891. Brassicaceae. = Rorippa curvipes var. cur- (Gray) Hall Washingto n Co., St. George, Palmer sn, vipes Salt Lake Co., City Creek Canyon, Jones 1352, 1875 (NY!;BRY!). 1879(POM!;NY!). '

June 1982 Welsh: Utah Plant Types 163

Cardamine uinlahensis F.J. Hermann Rhodora 36: Castilleja viscida Rydb. Bull. Torrey Bot. Club 34: 410. 1934. Brassicaceae. = C. cordifolia Gray Summit 32. 1907. Scrophulariaceae. Syn: C. applegatei var. vis- Co., Mt. Elizabeth Ridge, Hermann 5894, 1933 (GH). cida (Rydb.) Ownbey Salt Lake Co., Big Cottonwood Carduus lacerus Rydb. Bull. Torrey Bot. Club 37: Canyon, Rydberg & Carlton 6593, 1905 (NY!). 543. 1910. .A.steraceae. = Cirsium scariosum Nutt. Castilleja zionis Eastw. Leafl. W. Bot. 3: 91. 1941. Wasatch Co., near Midway, Carlton & Garrett 6732, Scrophulariaceae. Washington Co., Clear Creek, East- 1905(RM!;NY!). wood & Howell 9227, 1941 (CAS!). Carduus olivescens Rydb. Bull. Torrey Bot. Club 37: Caulanthus crassicaulis (Torr.) Wats. var. glaber 544. 1910. .\steraceae. = C. scariosum Nutt. Garfield Jones Zoe 4: 266. 1893. Brassicaceae. Kane Co., Sink Co., .\quarius Plateau, Rydberg & Carlton 7450, 1905 Valley, Jones sn, 1890 (POM!). (NY!). Caulanthus crassicaulis (Torr.) Wats. var. major €arex camptjlocarpa T.H. Holm ssp. affinis Maguire Jones Proc. Calif. Acad. II, 5: 623. 1895. Brassicaceae. Leafl. W. Bot. 4: 262. 1946. Cyperaceae. Garfield Co., Henry Mts., Jones 5685, 1894 & Holmgren ' = C. scopulorum T.H. Holm Juab Co., Indian Farm (POM!;US!;BRY!;NY!). Creek, Maguire & Holmgren 21947. 1943 Caulanthus divaricatus Rollins Contr. Gray Herb. II, (NY!;US!;CAS!;UTC!). 201: 8. 1971. Brassicaceae. = Thelypodiopsis divaricata Carex canescens L. var. dubia Bailey Bot. Gaz. 9: (Rollins) Welsh & Reveal San Juan Co., 10 mi E of Hite,

119. 1884. Cvperaceae. = C. cmiesceiis L. Summit (?) Cronquist 90.33, 1961 (GH;NY!). Co., Bear River Canyon, Watson 1231A, 1869 (NY!). Caulanthus hastatus Wats. Rep. U.S. Geol. Explor. Carex epapillosa Mack, in Rydb. Fl. Rocky Mts. 138. 40th Parallel, Bot. 5: 28. 1871. Brassicaceae. = Chloro- 1060. 1917. Cvperaceae. = C. atrata L. Piute Co., Mary- crambe hastatus (Wats.) Rydb. Salt Lake (?) Co., svale, Jones 5.345, 1894 (NY!;US!;POM!;BRY!). Wasatch Mts., Watson 114, 1869 (US!;NY!). Carex interimus Maguire Brittonia 5: 200. 1944. Ceanothus fendleri Gray var. viridis Jones Proc. Ca- = C. fendleri Cvperaceae. = C. aquatilis Wahl. Cache Co, Tony lif. Acad. II, 5: 629. 1895. Rhamnaceae. 60.32n, 1894 Grove Lake, Maguire 16098, 19.38 (NY!;C.\S!;US!;UTC!)'. Gray Kane (?) Co., Elk Ranch, Jones Carex pelocarpa F. Hermann Rhodora 39: 492. 1937. (POM!). Ceanothus martinii Jones Contr. W. Bot. 8: 41. 1898. Cyperaceae. = C. nova Bailey Summit Co., Lamotte Rhamnaceae. Syn: C. utahensis Eastw. Sanpete Co., Peak, Hermann 5983, 19,33 (NY!';C.\S!). Manti Canyon', Jones sn, 1895 (POM!;RM!;BRY!; Carex rachillis Maguire Brittonia 5: 199. 1944. Cype- ' NY!;UTC!). raceae. = C. sidmigricans Stacev Summit Co, Gilbert Ceanothus utahensis Eastw. Proc. Calif. Acad. IV. 16: Peak. Maguire et al 14668, 1936 (NY!;US!;CAS!;UTC!). 363. 1927. Rhamnaceae. = C. martinii Jones Wasatch Carex vernacula Bailey var. hobsonii Maguire Brit- Co., Soldier Summit, Eastwood 7689, 1918 (CAS!). tonia 5: 199. 1944. Cvperaceae. = C. neurophora Mack. Celtis villosula Rydb. Fl. Rocky Mts. ed 2. 1116. 1922. Cache Co., Bear River Range, Maguire et al 14013, 1936 Ulmaceae. = C. reticulata Torr. Utah, without definite (NY!;C.\S!;US!;UTC!). locality, Jones sn, 1894 (NY!). Carum garrettii A. Nels. in Coult. & Rose Contr. U.S. Cerastium variable Goodding Bot. Gaz. 37: 54. 1904. Natl. Herb. 12: 443. 1909. Apiaceae. = Perideridia Carvophvllaceae. = C. beeringianum C. & S. Uintah gairdneri (H. & A.) Mathias Salt Lake (?) Co., Wasatch Co.; Dyer Mine, Goodding 1246, 1902 (US!;RM!; Mts.. Garrett 2053, 1906 (US!;NY!). BRY!;NY!;UT!;ISC!). Castilleja aquariensis N. Holmgren Bull. Torrey Bot. Cercis orbiculata Greene Feddes Repert. 11: 111. Club 100: 87. 1973. Scrophulariaceae. Garfield Co., 1912. Fabaceae. = C. occidentalis var. orbicidata Aquarius Plateau, N. & P. Holmgren 4726, 1970 (Greene) Tidestr. Washington Co., Diamond Valley, (NY!;US!;BRY!;UTC!). Goodding 899, 1902 (US!;NY!). Castilleja arcuata Rydb. Bull. Torrey Bot. Club 34: Cercocarpus flabellifolius Rydb. N. Amer. Fl. 22: 422. 35. 1907. Scrophulariaceae. = C. linariifolia Benth. Se- 1913. Rosaceae. = C. montanus Raf. Sevier Co., near vier Co., Fish Lake, Rydberg & Carlton 7-508, 1905 Glenwood, Ward 122, 1875 (US!;NY!). (NY!:RM!:US!). Cercocarpus intricatus Wats. Proc. .\mer. .\cad. 10: Castilleja flava Wats. Rep. U.S. Geol. Explor. 40th 346. 1875. Rosaceae. Syn: C. intricatus var. villosus Parallel, Bot. 5: 2.30. 1871. Scrophulariaceae. Summit (?) Schneid. Utah Co., American Fork Canyon, Watson 314, Co., Upper Bear River Valley, Watson 813, 1869 1869 (US!;NY!). (US!;NY!). Cercocarpus intricatus Wats. var. villosus Schneid. Castilleja konardii Rydb. Bull. Torrey Bot. Club 34: = Mitt. Deutsch. Dendr. Ges. 14: 129. 1905. Rosaceae. .36. 1907. Scrophulariaceae. Utah Co., American Fork C. intricatus Wats. Tooele (?) Co., Deep Creek, Jones sn, Canyon, Leonard 151, 1885 (NY!). 1891 (US!). Castilleja parvula Rydb. Bull. Torrey Bot. Club 34: Cercocarpus ledifolius Nutt. var. intercedens Schneid. 40. 1907. Scrophulariaceae. Piute Co., Bullion Creek, f. hirsutus Schneid. Mitt. Deutsch. Dendrol. Ges. 14: Rydberg & Carlton 71.58, 1905 (NY!;US!). 129. 1905. Rosaceae. = C. ledifolius Nutt. Weber Co., Castilleja revealii N. Holmgren Bull. Torrey Bot. Ogden Canyon, Pammel & Blackwood 3726, 1902 Club 100: 87. 1973. Scrophulariaceae. Garfield Co., (ISC!;BRY!). Bryce Canyon, Holmgren & Reveal 2017, 1965 Cercocarpus ledifolius Nutt. var. intercedens f. sub- (NY!;US!;BRY!;UT!:UTC'). glaber Schneid. Mitt. Deutsch. Dendr. Ges. 14: 128. Castilleja variabilis Rydb. Bull. Torrey Bot. Club 34: 1905. Rosaceae. = C. ledifolius Nutt. Utah Co., Slate ..37. 1907. Scrophulariaceae. = C. miniata Dougl. Salt Canyon, Jones 5613b, 1894 (MO;NYl;US!;RM!; Lake Co., Big Cottonwood Canyon, Rydberg 6773, 1905 BRY!;POM!). (NY!;RM!). 164 Great Basin Naturalist Vol. 42, No. 2

Cercocarpus parvifolius Wool. var. minimus Schneid. Cirsium lactucinum Rydb. Fl. Rocky Mts. 1010, 1068. 111. Handb. Laubholzk. 1: 532. 1905. Rosaceae. = C. 1917. Asteraceae. = C. rijdbergii Petrak San Juan Co., montuntis Raf.? Utah ?. Bluff, Rydberg & Garrett 10001, 1911 (NY!;US!;UT!). Chaenactis brachiata Greene Leafl. Bot. Obs. & Crit. Cirsium pulchellum (Greene) Woot. & Standi, var. 2: 224. 1912. Asteraceae. = C. douglasii (Hook.) H. & A. glabrescens Petrak Beih. Bot. Centr. 3.5(2): 511. 1917. Washington Co., Springdale, Jones 5261q, 1894 A.steraceae. = C. bipinnatum (Eastw.) Petrak San Juan (US!;POM!). Co., Elk Mts., Rydberg & Garrett 9335, 1911 (US!). Chaenactis brachiata Greene var. stansburiana Cirsium rydbergii Petrak Beih. Bot. Centralbl. 35(2): Stockwell Contr. Dudley Herb. .3: 111. 1940. Asteraceae. 315. 1917. Asteraceae. Syn: C. lactucinum Rydb. San = C. douglasii (Hook.) H. & A. Tooele Co., Stansbury Juan Co., Bluff, Rydberg & Garrett 10001, 1911 Island, Stansbury sn, 1850 (US;NY!). (US!;UT!;NY!). Chaenactis douglasii (Hook.) H. & A. var. alpina Cirsium undulatum (Nutt.) Spreng. var. albescens Gray Syn. Fl. N. Amer. = 1(2): 341. 1884. C. alpina D.C. Eaton Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: (Gray) Jones Salt Lake Co., Alta, Jones 1232, 1879 194. 1871. Asteraceae. = C. undulatum (Nutt.) Spreng. (NY!;UTC!). Tooele Co., Stansbury Island, Wat.son 687, 1869 (US!). Cheirinia brachycarpa Rydb. Bull. Torrey Bot. Club Cirsium utahense Petrak Beih. Bot. Centr. 35(2); 470. 39: .325. 1912. Brassicaceae. = Erijsimum asperum Nutt. 1917. Asteraceae. Washington Co., Silver Reef, Jones San Juan Co., Abajo Mts., Rydberg & Garrett 9713, 1911 516.3q, 1894 (POM !;US!). (NY!;US!;UT!). Clematis alpina ssp. occidentalis var. repens Kuntze Chrysopsis caespitosa Jones, not Nutt. Proc. Calif. Verb. Bot. Ver. Brandenb. 26: 161. 1885. Ranuncu- Acad. II, 5: 694. 1895. Asteraceae. = Heterotheca pnesii laceae. = C. columbiana (Nutt.) T. & G. Utah Co., (Blake) Welsh & Atwood Washington Co., Springdale, American Fork, Jones 1887, 1880 (NY!;POM!). Jones .5249u, 1894 (POM!;US!). Clematis douglassii ssp. jonesii Kuntze Verb. Bot. Ver. Chrysopsis villosa Nutt. var. cinerascens Blake Proc. Brandenb. 25: 180. 1886. Ranunculaceae. = C. hir- sutissima Pursh Biol. Soc. Washington .35: 173. 1922. Asteraceae. = Het- Utah Co., American Fork Canvon, Jones 1351, erotheca villosa (Nutt.) Shinners Beaver Co., Beaver 1880 (US!;POM!;NY!). Canyon, Tidestrom 2873, 1901 (US!). Clematis pseudotragene ssp. tvenderothioides Kuntze Verb. Bot. Ver. Brandenb. Chrysopsis villosa Nutt. var. scabra Eastw. Proc. Ca- 26: 160. 1885. Ranuncu- laceae. lif. = C. columbiana var. columbiana Kane Co., Acad. II, 6: 294. 1896. Asteraceae. = Heterotheca vil- Ka- nab, Siler sn, 1873 (K). losa (Nutt.) Shinners San Juan Co., Willow Creek, East- Cleome integrifolia var. wood .38, 1895 (CAS!). angusta Jones Proc. Calif. Acad. II, 5; 625. 1895. Capparidaceae. = C. serrulata Chrysothamnus marianus Rydb. Bull. Torrey Bot. var. angusta (Jones) Tidestr. Piute Co., Marysvale, Club 37; 131. 1910. Asteraceae. = C. viscidiflorus var. Jones 6057, 1894 (US!). puhendus (D.C. Eaton) Jeps. Piute Co., near Marysvale, Cleomella cornuta Rydb. Bull. Torrey Bot. Rydberg & Carlton 6993, 1905 (US;R]V1!;NY!). Club .30; 249. 1903. Capparidaceae. = C. palmeriana Jones Chrysothamnus nauseosus (Pallas) Britt. ssp. iridis Wayne Co., Caineville, Jones 5656, 1894 L.C. Anderson Great Basin Naturalist 41: 311. 1981. As- (US!;BRY!;RM!;NY!). teraceae. Sevier Co., Rainbow Hills, Welsh 19258, 1979 Cleomella nana Eastw. Bull. Torrey (BRY!;FSU!). Bot. Club 30: 490. 1903. Capparidaceae. = C. palmeriana Jones Chrysothamnus nauseosus (Pallas) Britt. var. psilo- Grand Co., S of Thompsons Spr., Eastwood sn, 1892 carpus Blake Washington Acad. Sci. 27: 376. 1937. As- J. (CAS!;RM!). teraceae. Syn: C. nauseosus ssp. psilocarpus (Blake) L.C. Cleomella palmerana Jones Zoe 2: 236. 1891. Cap- Anderson Emery Co., Huntington Canyon, Garrett 7021 paridaceae. Emery Co., Green River, sn, 1935 (US). Jones 1890 (POM;US!). Chrysothamnus oliganthus A. Nels. Univ. Wyoming Cnicus calcareus Jones Proc. Calif. Acad. II, 5: 704. Publ. Bot. 1: 65. 1924. Asteraceae. = C. nauseosus var. 1895. Asteraceae. = Cirsium calcareum (Jones) Woot. & leiospermus (Gray) Hall Washington Co., Zion National Standi. Garfield Co., Bromide Pass, Jones 569.5bh, 1894 Park, Nelson 9975, 1922 (RM!). (POM!;BRY!;US!;NY!). Chrysothamnus salicifolius Rydb. Bull. Torrey Bot. Cnicus clavatus Jones Proc. Calif Acad. II, 5; 704. Club 37: 1,30. 1910. Asteraceae. = C. nauseosus (Pallas) 1895. Asteraceae. = Cirsium clavatum (Jones) Petrak Se- Britt. var. salicifolius (Rydb.) Hall Wasatch Co., Straw- vier Co., Fish Lake, Jones 5715, 1894 berry Valley, Leonard 288, 1883 (NY!;BRY!). (POM!;BRY!;US!;NY!). Chrysothamnus zionis A. Nels. Univ. Wyoming Publ. Cnicus nidulus Jones Proc Calif. Acad. II, 5: 705. Bot. 1: 66. 1924. Asteraceae. = C. nauseosus var. 1895. Asteraceae. = Cirsium nidulum (Jones) Petrak gnaphaloides (Greene) Hall Washington Co., 20 mi N St. Garfield Co., Paria, Jones 5290a, 1894 (POM!;US!;NY!). George, Nelson 9980, 1922 (RM!). Cnicus rothrockii var. diffusus Eastw. Proc. Calif. Cirsium barnebyi Welsh & Neese in Welsh Brittonia Acad. II, 6; 303. 1896. Asteraceae. = C. rothrockii 33: 296. 1981. Asteraceae. Uintah Co., 1.5 mi e Ignacio, (Gray) Petrak San Juan Co., Willow Creek, Eastwood Welsh 19606, 1980 (BRY!;NY!;US!;MINN!). 65, 1895 (US!;CAS!). Cirsium eHocephalum var. leiocephalum D.C. Eaton Cogswellia cottamii Jones Contr. W. Bot. 16: .36. Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: 196. 1871. 1930. Apiaceae. = Lomatium parriji (Wats.) Macbr. Asteraceae. = C. eatonii Robins. var. eatonii Summit? Washington Co., Beaverdam Mts., Cottam et al. 4098, Co., Uinta Mts., Watson 691, 1869 (US!;NY!). 1929 (BRY!;UT!). June 1982 Welsh: Utah Plant Types 165

Cogswellia millefolia var. depauperata Jones Contr. Cryptantha leptophylla Rydb. Bull. Torrey Bot. Club W. Bot. 12: 38. 1908. Apiaceae. = Lomatium graiji 36: 679. 1909. Boraginaceae. = C. nevadensis Nels. & Coiilt. & Rose Tooele Co., Dugway, Jones sn, 1891 Kennedy Washington Co., St. George, Palmer 350, 1877 (US!;POM!;NY!). (NY!). Cogswellia minima Mathias Ann. Missouri Bot. Card. Cryptantha ochroleuca Higgins Great Basin Natural-

19: 497. 1932. Apiaceae. = Lomatium rninimtan (Math- ist 28: 197. 1968. Boraginaceae. Garfield Co., Red Can- ias) Mathias Garfield Co., Bryce Canyon, Mathias 670, yon, Higgins 1788, 1968 (BRY!;US!). 1929 (MO;CAS!;NY!). rollinsii Arb. 20: 391. Cryptantha Johnst. J. Arnold Coleosanthes garretti A. Nels. Proc. Biol. Soc. Wash- 1939. Boraginaceae. Uintah Co., 22 mi s Ouray, Rollins ington 20: 38. 1907. Asteraceae. = Brickellia grandiflora 1715, 1937 (GH;US!;CAS!;RM!;UTC!). (Hook.) Niitt. Salt Lake Co., City Creek Canyon, Gar- Cuscuta denticulata Engelm. in Parry Amer. Natural- rett 1061, 1904 (RM!;UT!). ist 9: 348. 1875. Cuscutaceae. Washington Co., near St. Collomia tenella Gray Proc. Amer. Acad. 8: 259. George, Parry 205, 1874 (US!;NY!;ISC!;BRY!). 1870. Polemoniaceae. Summit Co., Parleys Park, Watson Cuscuta warneri Yuncker Brittonia 12: 38. 1960. Cus-

900, 1869 (NY!). cutaceae. Millard (?) Co., Flowell, Warner sn, 1957 Coloptera jonesii Coult. & Rose Rev. N. Amer. Um- (US!;CAS!;BRY!;UT!;NY!;UTC!;ISC!). bell. 49. 1888. Apiaceae. = Cymopterus newberriji Cycladenia jonesii Eastw. Leafl. W. Bot. 3: 159. 1942. (Wats.) Jones Beaver Co., Milford, Jones 1792, 1880 Apocynaceae. = C. humilis var. jonesii (Eastw.) Welsh (POM!;UT!). & Atwood Emery Co., San Rafael Swell, Jones sn, 1914 Comandra linearis Rydb. Fl. Rocky Mts. 818. 1066. (CAS!;US!;BRY!). 1917. Santalaceae. = C. uynbellata var. pallida (DC.) Cymopterus basalticus Jones Contr W. Bot. 12: 16.

Jones Emery Co., Green River, Tracy 716, 1887 (NY!). 1908. Apiaceae. Beaver (?) Co., Wah Wah, Jones sn, Cordylanthus parryi Wats, in Parry Amer. Naturalist 1906 (POM !;BRY!).

9; 346. 1875. Scrophulariaceae. = C. maritimus ssp. ca- Cymopterus beckii Welsh & Goodrich in Welsh Brit- nescens (Gray) Chuang & Heckard Washington Co., tonia 33: 297. 1981. Apiaceae. Wayne Co., Fruita, Beck Parry 155, 1874 (US!;NY!;ISC!). sn, 1938 (BRY!). Corydalis engelmannii var. exaltata Fedde Feddes Cymopterus corrugatus var. scopulicola Jones Contr. = coulteri Repert. 11: 497. 1913. Fumariaceae. = C. aurea Willd. W. Bot. 14: 39. 1912. Apiaceae. C. (Jones) Mathias Juab Co., Sevier Bridge, Jones sn, 1910 (POM!). Grand (?) Co., La Sal Mts., Purpus 6550, 1897 (US!). decipiens Zoe 2: 247. 1891. Coryphantha marstonii Clover Bull. Torrey Bot. Cymopterus Jones Apiaceae. = C. fendleri Gray Grand Co., Cisco, Jones Club 65: 412. 1938. Cactaceae. = C. Tuissouriensis var. sn, 1891 (POM!;US!;NY!). marstonii (Clover) L. Benson Garfield Co., Hells Back- Cymopterus duchesnensis Jones Contr. W. Bot. 13: bone, Clover 1909, 1937 (?) (MICH). 12. 1910. Apiaceae. Duchesne Co., Myton, Jones sn, Cowania stansburiana Torr. in Stansb. Explor. Great 1908 (CAS!;US!;RM!;POM!;BRY!). Salt Lake 386. 1852. Rosaceae. = C. mexicana D. Don Cyrhopterus higginsii Welsh Great Basin Naturalist Davis (?) Co., Great Salt Lake, Stansbury sn, 1850 (NY!). 1976. Apiaceae. Kane Co., 17 mi e Glen Canyon Bot. 18: 685. 1931. 35: 377. Crassipes annuus Swallen Amer. J. City, Welsh 12740, 1975 (BRY!;NY!). Poaceae. = Sclerochloa dura (L.) Beauv. Davis (?) Co., Cymopterus ibapensis Jones Zoe 3: 302. 1893. between Salt Lake City & Ogden, Fallas sn, 1928 (US!). Apiaceae. Tooele Co., Deep Creek Valley, Jones sn, Crepis aculeolata Greene Leafl. Bot. Obs. & Grit. 2: (?) 1891 (POM!;US!;RM!). 86. 1910. Asteraceae. = C. runcinata var. runcinata jonesii Coult. Rose Rev. N. Amer. Um- Utah, Ward 606, 1875 (US!). Cymopterus & 80. 1888. Apiaceae. Syn: Aulospermum jonesii Crepis occidentalis Nutt. var. costatus Gray Geol. bell. (Coult. & Rose) Coult. & Rose Beaver Co., Frisco, Jones Surv. Calif. Bot. 1: 435. 1880. Asteraceae. = C. occiden- 1808, 1880 (US!;CAS!;POM!;RM!;NY!;UT!;UTC!). talis Nutt. Tooele Co., Stansbury Island, Watson 715, longipes Wats. Rep. U.S. Geol. Explor. 1869 (US!). Cymopterus Parallel, Bot. 5: 124. 1871. Apiaceae. Syn: Peuceda- Cressa erecta Rydb. Bull. Torrey Bot. Club 40: 466. 40th S.alt Lake Co., near Salt Lake 1913. Convolvulaceae. = C. truxillensis H.B.K. Salt num lapidosum Jones 1869 (US!;NY!). Lake Co., Becks Hot Springs, Garrett 870f, 1905 (NY!). City, Watson 451, Cynomarathrum latilobum Rydb. Bull. Torrey Bot. Croton longipes Jones Proc. Calif. Acad. II, 5: 721. 1913. Apiaceae. = Lomatium latilobum 1895. Euphorbiacea. Washington Co., 2 mi e Leeds, Club 40: 73. (Rydb.) Mathias Grand Co., Wilson Mesa, Rydberg & Jones 5213, 1894 (POM!;US!;CAS!;RM!;NY!). (NY!;US!;UT!). Arnold Arb. 29: 240. Garrett 8371, 1911 Cryptantha bamebyi Johnst. J. Cynomarathrum scabrum Coult. & Rose Contr. U.S. 1948. Boraginaceae. Uintah Co., 30 mi s Ouray, Ripley Natl. Herb. 7: 247. 1900. Apiaceae. = Lomatium sca- & Barneby 8748, 1947 (GH;NY!). Const. Beaver Co., Cryptantha compacta Higgins Great Basin Naturalist brum (Coult. & Rose) Math. & Frisco, 1864, 1880 (US!;CAS!;POM!;BRY!; 28: 196. 1968. Boraginaceae. Millard Co., 8 mi w Desert Jones Experimental Range, Higgins 1613, 1963 NY!;UT!;UTC!). 409. (BRY!;US!;POM!). Cystium stramineum Rydb. N. Amer. Fl. 24: = Astragalus lentiginosus var, stra- Arb. 18: 23. 1929. Fabaceae. Cryptantha grahamii Johnst. J. Arnold Washington Co., "S. Utah" Pal- 1937. Boraginaceae. Uintah Co., mouth Sand Wash, mineus (Rydb.) Barneby Graham 7924, 1933 (GH). mer sn, 1870 (NY!;US!). Great Basin Naturalist 31: 90. Cryptantha johnstonii Higgins Great Basin Naturalist Dalea epica Welsh 1971. Fabaceae. San Co., 10 mi e Halls Crossing, 28: 195. 1968. Boraginaceae. Emery Co., 15 mi w Hwy Juan (BRY!;NY!;ISC!). 50-6, Higgins 1310,1968 (BRY!;POM!;US!;NY!). Welsh 5205, 1966 166 Great Basin Naturalist Vol. 42, No. 2

Dalea johnsonii Wats. Rep. U.S. Geol. Explor. 40th Draba spectabilis Greene var. glabrescens Schulz Parallel, Bot. 5: 64. 1871. Fabaceae. = Psorotharnnus Pflanzenr. 4. Fam. 105 (Heft 89) : 84. 1927. Brassicaceae. fremontii (Torr.) Barneby Washington Co., near St. = D. spectabilis Greene San Juan Co., LaSal Mts., George, Johnson sn, 1870 (GH). Walker 275, 1912 (NY!;US!). Dalea nummularia Jones Contr. W. Bot. 18: 41. 1933. Draba subalpina Goodm. & Hitchc. Ann. Missouri Fabaceae. = Psorotharnnus polyadenius var. pnesii Bot. Card. 19: 77. 19.32. Brassicaceae. Iron Co., Cedar Barneby Emery Co., Green River, Jones sn, 1914 Breaks, Goodman & Hitchcock 1622, 1930 (NY!;RM!). (US!;CAS!;NY!).' Draba uncinalis Rydb. Bull. Torrey Bot. Club 30: Daucophyllum lineare Rydb. Bull. Torrey Bot. Club 251. 1903. Brassicaceae. = D. sobolifera Rydb. Piute 40: 69. 1913. Apiaceae. = Musineon lineare (Rydb.) Co., Tate Mine, Jones 5940am, 1894 (US!;POM!). Mathias Cache Co., near Logan, Rydberg sn, 1895 (?). Draba valida Goodding Bot. Gaz. 37: 55. 1904. Delphinium ahietorum Tidestr. Proc. Biol. Soc. Brassicaceae. = D. lanceolata Royle Uintah Co., Dyer Washington 27: 61. 1914. Ranunculaceae. = D. occiden- Mine, Goodding 1402, 1902 (US!;RM!). tale Wats. Sanpete (?) Co., Wasatch Plateau, Coville & Draba zionensis C.L. Hitchc. Univ. Washington Tidestrom 19, 1908 (US!). Publ. Biol. 11: 49. 1941. Brassicaceae. = D. asprella var. Delphinium coelestinum Rydb. Bull. Torrey Bot. zionensis (C.L. Hitchc.) Welsh & Reveal Washington Club .39: 320. 1912. Ranunculaceae. Syn: D. ajnabile Co., Zion Canyon, Jones sn, 1923 (POM!;CAS!;BRY!). Tidestr. (new name) = D. scaposum Greene Washing- Drymocallis micropetala Rydb. N. Amer. Fl. 22: 375. ton Co., St. George, Palmer 10, 1877 (NY!;US!). 1908. Rosaceae. = Potentilla glandulosa var. micropetala Delphinium leonardii Rydb. Bull. Torrey Bot. Club (Rydb.) Welsh Johnst. Salt Lake Co., City Creek Can- 39: .320. 1912 Ranunculaceae. = D. andersonii Gray Salt & yon, 61,53, 1905 Lake Co., Garfield, Leonard 20.5, 1884 (NY!). Rydberg (US!;NY!). Delphituum scoptilorum var. attenuafum Jones Proc. Echinocactus johnsonii Parry ex Engelm. Rep. U.S. Parallel, Bot. 5: 117. 1871. Calif. Acad. II, 5: 617. 1895. Ranunculaceae. = D. bor- Geol. Explor 40th Cactaceae. = heifi Huth. Piute Co., head Bullion Creek, Jones 5893d, NeoUoydia johnsonii (Parrv) L. Benson Washington 1894(POM!:US!). Co., near St. George, Johnson sn, 1870? (GH?). Descurainia richardsonii (Sweet) Schulz var. macros- Echinocactus whipplei Engelm. & Bigel. var. spin- perma Schulz Pflanzenr. 4. Fam. 105(Heft 86): 319. osior Engelm. Trans. Acad. Sci. St. Louis 2: 199. 1863. 1924. Brassicaceae. = D. richardsonii var. brevipes Cactaceae. = Sclerocactus pubispinus {Enge]m.) L. Ben- (Nutt.) Welsh & Reveal Salt Lake Co., Alta, Jones 1117, son Utah, desert valley west of Camp Floyd, H. Engel- 1879 (UTC!). mann sn, 18.59 (MO).

Dicoria paniculata Eastw. Proc. Calif. Acad. II, 6: Echinocereus engelmannii (Parry) Rumpler var. purpureus L. Benson 41: 127. 298. 1896. Asteraceae. = D. brandegei Gray San Juan Cact. & Succ. J. (Los Angeles) Co., McElmo & Recapture, Eastwood 51, 1895 (CAS!). 1969. Cactaceae. Washington Co., near St. George, Ben-

Dicoria wetherillii Eastw. Proc. Calif. Acad. II, 6: son 1.3637, 1949 (POM). 299. 1896. Asteraceae. = D. brandegei Gray (monstrous Edwinia macrocalyx Small N. Amer. Fl. 22: 176. form?) San Juan Co., San Juan River, Wetherill sn, 1895 1905. Saxifragaceae. = Jamesia americana T. & G. Utah (CAS!). Co., American Fork Canyon, Watson 371, 1869 (US!).

Distichlis maritima var. laxa Holm Bot. Gaz. 16: 277. Elymus salina Jones Proc. Calif. Acad. II, 5: 725. 1891. Poaceae. = D. stricta (Torr.) Rvdb. Utali, Lake 1895. Poaceae. Sevier Co., Salina Pass, Jones 5447, 1894

Park, Tracy sn, 1887 (?). (RM!;BRY!;US!;NY!). Dodecatheon zionense Eastw. Leafl. W. Bot. 2: 37. Emmenanthe foliosa Jones Zoe 4: 278. 1893. Hydro- 1937. Primulaceae. = D. pulchcUitm (Raf.) Merr. Wash- phyllaceae. = PhaceUa salina (A. Nels.) J.T. Howell ington Co., Zion Canyon, Eastwood & Howell 1144, Tooele (?) Co.. Deep Creek, Jones sn, 1891 (POM!;US!). 19.33 (CAS!). Encelia microcephala Gray Proc. Amer. Acad. 8: 657. Draha apiculata C.L. Hitchc. Univ. Washington 1873. Asteraceae. = Helianthella microcephala (Gray) Publ. Biol. 11: 72. 1941. Bra.ssicaceae. = D. densifolia Gray San Juan Co., Sierra .\bajo, Newberry sn, 1859 var. davisiae (C.L. Hitchc.) Welsh & Reveal Summit (NY!). Co., Uinta Mts, Payson & Pavson 5048, 1926 Encelia nudicaulis Gray Proc. Amer. Acad. 8: 656. (NY!;US!:RM!). 1873. Asteraceae. = Enceliopsis nudicaulis (Gray) A. Draha brachystylis Rydb. Bull. Torrey Bot. Club 29: Nels. Utah, Bishop sn, 1872 (US!). 240. 1902. Brassicaceae. Salt Lake Co., Wasatch Mts., Epilobium palmeri Rydb. Bull. Torrey Bot. Club 31: Alta, Jones 1.357, 1879 (NY!;US!;CAS!;RM!;POM!). 569. 1904. Onagraceae. = E. saximontanum Hausskn. Draha maguirei C.L. Hitchc. var. burkei C.L. Washington Co., (s. Utah), Palmer 156, 1877 (NY!). Hitchc. Univ. Washington Publ. Biol. 11: 70. 1941 Epilobium tracyi Rydb. Bull. Torrey Bot. Club 40: 63. Brassicaceae. Box Elder Co., Cottonwood Canyon, 1913. Onagraceae. = E. hrachycarpiim Presl Weber Burke 2968, 19.32 (UTC!). Co., Ogden, Tracy & Evans 547, 1887 (NY!). Draba maguirei C.L. Hitchc. var. maguirei Univ. Eremocarya muricata Rydb. Bull. Torrey Bot. Club Washington Publ. Biol. 11: 70. 1941. Brassicaceae. 36: 677. 1909. Boraginaceae. = Cryptantha micrantha

Cache Co., Bear River Range, Maguire et al. 14161, (Torr.) Johnst. Southern Utah, Parry 164, 1874 19.36 (WTU!;UTC!;NY!). (NY!;1SC!). Draba sobolifera Rydb. Bull. Torrey Bot. Club 30: Erigeron abajoensis Cronq. Brittonia 6: 168. 1947. As- 251. 1903. Brassicaceae. Syn: D. uncinalis Rydb. Piute teraceae. San Juan Co., Abajo Mts., Rydberg & Garrett Co., Tate Mine, Jones .59.36, 1894 (NY!;US!;POM!). 97.55, 1911 (NY!;UT!). June 1982 Welsh: Utah Plant Types 167

Erigeron caespitosus Nutt. var. laccoliticus Jones Erigeron pulvinatus Rydb. non Wedd. Fl. Rocky Mts. Proc. Calif. Acad. II, 5: 696. 1895. Asteraceae. = E. 911, 1067. 1917. Asteraceae. = £. compactus Blake Juab caespitosus Nutt. Garfield Co., Henry Mts., Jones 5661, Co., Deep Creek, Jones sn, 1891 (US!;POM!;BRY!;NY!). 1894 {POM!:US!). Erigeron pumilus Nutt. ssp. concinnoides Cronq. var. Erigeron caespitosus Nutt. var. laccoliticus Jones subglaber Cronq. Brittonia 6: 183. 1947. Asteraceae. San Proc. Calif. Acad. II, 5: 696. 1895. Asteraceae. = £. Juan Co., Monticello, Rvdberg & Garrett 9141, 1911 caespitosus Nutt. Garfield Co., Henry Mts., Jones 5661, (US!;RM!;UT!). 1894 (POM!;US!). Erigeron religiosus Cronq. Brittonia 6: 258. 1947. .As- Erigeron caespitosus Nutt. var. nauseosus Jones Proc. teraceae. Washington Co., Clear Creek, Eastwood &

Calif. .\cad. II, 5: 696. 1895. Asteraceae. = E. nauseosus Howell 6.3.39, 1938 (CAS!;US!). (Jones) Cronq. Piute Co., Marysvale, Jones 5386, 1894 Erigeron sionis Cronq. Brittonia 6: 258. 1947. Aste- (POM!;US!;NY!). raceae. Washington Co., Zion National Park, Pilsbry .sn, Erigeron cinereus var. aridus Jones Proc. Calif. Acad. 1925 (PHI).

II, 5: 695. 1895. Asteraceae. = E. divergens T. & G. Erigeron sparsifolius Eastw. Proc. Calif. Acad. II, 6: Washington Co., Silver Reef, Jones 5149v, 1894 297. 1896. Asteraceae. = £. utahensis Gray var. spar- (POM!;US!). sifolius (Ea.stw.) Cronq. San Juan Co., Willow Creek, Erigeron controversus Greene Leafl. Bot. Obs. & Grit. Eastwood 48, 1895 (CAS!).

2: 206. 1912. Asteraceae. = E. garrctti A. Nels. Salt Lake Erigeron stenophyllus D.C. Eaton Rep. U.S. Geol. Ex- Co., Alta, Jones 1207, 1879 (US!;POM!). plor. 40th Parallel, Bot 5: 152. 1871. Asteraceae. = £. Erigeron cronquistii Maguire Brittonia 5: 201. 1944. arenarioides (D.C. Eaton) Rydb. Salt Lake Co., Cotton- Asteraceae. Cache Co., Logan Canyon, Maguire 16681, wood Canyon, Watson 547, 1869 (US!;NY!). 1939 (NY!;US!;UTC!). Erigeron stenophyllus var. tetrapleuris Gray Proc. Erigeron eatonii Gray Proc. Amer. Acad. 16: 91. 1880. Amer. Acad. 8: 650. 1873. .Asteraceae. = £. utahensis Asteraceae. Summit Co., Uinta Mts., Watson 546, 1869 Gray Kane Co., Kanab, Thompson sn, 1870 (GH). (US!:NY!). Erigeron uintahensis Cronq. Bull. Torrey Bot. Club Mill Creek, Fay- Erigeron eatonii Gray ssp. typicus Cronq. var. mo- 70: 270. 1943. Asteraceae. Summit Co., 19.36 (RM!;US!). lestus Cronq. Brittonia 6: 172. 1947. .Asteraceae. = £. son 4894. Erigeron ursinus D.C. Eaton Rep. U.S. Geol. Explor. eatonii Grav Tooele Co., Mt. Deseret Peak, Stansbury 148. 1871. Asteraceae. Summit Co., Range. Maguire and Holmgren 21773, 1943 (NY!;UTC!).' 40th Parallel, Bot. 5; Bear River Canyon, Watson 5.34, 1869 (US!;NY!). Erigeron flagellaris Gray var. trilobatus Maguire & Erigeron vagus Payson Univ. Wyoming Publ. Bot. 1: Cronq. Brittonia 6; 258. 1947. Asteraceae. = £. prose- 179. 1926. Asteraceae. Grand Co., LaSal Mts., Payson & hjticus Nesom Iron Co., Cedar Breaks, Maguire 14947, Payson 40.33, 1924 (US!;BRY!;RM!:NY!). 19.34 (NY!;UTC!). Eriogonum ammophilum Reveal Phytologia 23: 163. Erigeron fruticetorum Rydb. Fl. Rocky Mts. 906, 1972. Polvgonaceae. Millard Co., Ibex Warm Point, N. 1067. 1917. Asteraceae. = E. formosissimus Greene San & P. Holmgren 46.50, 1970 (US!;NY!;BRY!;UTC!). Juan Co., La Sal Mts., Rvdberg & Garrett 8912, 1911 Eriogonum aretioides Barneby Leafl. W. Bot. 5: 154. (NY!). 1949. Polvgonaceae. Garfield Co., Widtsoe, Ripley & Erigeron garrettii A. Nels. in Coult. & Nels. Man. Barneby 8.570, 1947 (CAS!;NY!;BRY!). Bot. Centr. Rocky Mts. 526. 1909. Asteraceae. Syn: E. Eriogonum aureum Jones Proc. Calif. Acad. II, 5: 718. controversus Greene Salt Lake Co., Big Cottonwood 1895. Polygonaceae. = £. corymbosum var. glutinosusm Canyon, Garrett 1310, 1905 (RM!;UT!). Washington Co., St. George, Jones 6091, Erigeron kachinensis Welsh & Moore Proc. Utah (Jones) Jones 1894(POM!;US!;NY!).' Acad. 45: 2.31. 1968. Asteraceae. San Juan Co, Natural Eriogonum batemanii Jones Contr. W. Bot. 11: 11. Bridges, Welsh & Moore 2.398, 1963 (BRY!;NY!). 1903. Polvgonaceae. Carbon Co., Price Valley, Jones sn, Erigeron leiomerus Gray Syn. Fl. N. Amer. II. 1: 211. 1903(US!';POM!). 1884. .\steraceae. Svn: E. minuscuhis Greene Summit Eriogonum bicolor Jones Zoe 4: 281. 1893. Polyg- Co., Uintas, Watson'504, 1869 (NY!). onaceae. Grand Co., Thompsons Spring, Jones sn, 1891 Erigeron leiophyllus Greene Leafl. Bot. Obs. & Grit. (US!;POM!:NY!). 2: 218. 1912. Asteraceae. = E. speciosus var. macr- Eriogonum biumbellatum Rydb. Bull. Torrey Bot. anthus (Nutt.) Cronq. Salt Lake Co., Fort Douglas, Jones Club .39: .306. 1912. Polvgonaceae. = £. umbellatum sn, 1880 (US!;POM!). var. subaridum Stokes Sevier Co., Fish Lake, Rydberg & Erigeron maguirei Cronq. Brittonia 6: 165. 1947. .As- Carlton 7409. 1905 (US!;RM!:NY!). teraceae. Emerv Co., Calf Spring Wa.sh, Maguire 18459, brevicaule Nutt. var. pumilum Stokes in 1940 (NY!:UTC!). Eriogonum Jones Contr. W. Bot. 11: 10. 1903. Polygonaceae. = £. Erigeron mancus Rydb. Fl. Rocky Mts. 902, 1067. brevicaule var. laxifolium (T. & G.) Reveal Carbon Co., 1917. Asteraceae. Syn: £. pinnatisectus var. insoJens between Colton and Kyune, Jones 5603, 1894 (POM?). Macbr. & Pavson Grand Co., La Sal Mts., Rvdberg & Eriogonum cernuum Nutt. var. tenue T. & G. Proc. Garrett 8671, 1911 (NY!;US!;UT!;RM:). Amer. Acad. 8: 182. 1870. Polygonaceae. = £. cernuum Erigeron minusculus Greene Leafl. Bot. Obs. & Grit. var. cernuum Weber Co., Weber Valley, Watson 10.36, 2: 8. 1909. .Asteraceae. = £. leiomcris Gray Salt Lake 1869 (US!). Co., Big Cottonwood Canyon, Garrett, 1906 (UT?). Eriogonum cernuum Nutt. var. umbraticum Eastw. Erigeron pinnatisectus A. Nels. var. insolens Macbr. Proc. Calif. Acad. II, 6:: 319. 1896. Polygonaceae. = £. & Payson Contr. Gray Herb. II, 49: 79. 1917. Aste- cen^uum var. cernuum San Juan Co., McElmo Creek, raceae. = £. mancus Rvdb. Grand Co., La Sal Mts., Eastwood sn, 1892 (CAS?). Walker 271, 1912 (GH;MiNN;RM!). 168 Great Basin Naturalist Vol. 42, No. 2

Eriogonum chrysocephalum Gray ssp. alpestre Stokes Eriogonum effusum Nutt. ssp. orbiculatum Stokes Genus Eriogonum 93. 1936. Polygonaceae. = E. pan- Genus Eriogonum 79. 1936. Polygonaceae. = E. co- guicense var. alpestre (Stokes) Reveal Iron Co., Cedar rymbosum var. orbiculatum (Stokes) Reveal & Broth- Breaks, Goodman & Hitchcock 1601, 1930 (CAS!; erson Emery Co., Green River, Jones sn, 1915 NY!;UTC!). (POM!;BRY!). clavellatum Small Bull. Eriogonum Torrey Bot. Club Eriogonum effusum Nutt. ssp. pallidum var. shandsii 25: 48. 1898. Polygonaceae. San Juan Co., Bartons range, Stokes Genus Eriogonum 81. 1936. Polygonaceae. = E. Eastwood 1,32, 1895 (US!;NY!). leptocladon var. leptocladon San Juan Co., 3 mi s Indian Eriogonum confertiflorum var. stansburyi Benth. in Creek, Shands 1,30, 1931 (UT!). DC Prodr. 14: 17. 1856. Polygonaceae. = E. brevicaiile Eriogonum ephedroides Reveal Madrono 19: 295. var. brevicaule Utah (label data illegible), Stansbury sn, 1969. Polygonaceae. Uintah Co., 10 mi s Bonanza, Hol- 1850 (NY!). mgren et al. 2265, 1965 (UTC!;BRY!;UT!;RM!;ISC!). Eriogonum corymbosum Benth. in DC. var. albogil- Eriogonum eremicum Reveal Phytologia 23: 165. vum Reveal Great Basin Naturalist 27: 218. 1967. Polyg- 1972. Polygonaceae. Millard Co., se Garrison, Holmgren onaceae. = E. X duchesnense Reveal Duchesne Co., In- et al. 2247, 1965 (UTC!;BRY!;UT!;RM!;ISC!). Eriogonum filicaule Stokes Genus 32. dian Canyon, J & C Reveal 726, 1967 (US!;BRY!; Eriogonum NY!;UTC!). 1936. Polygonaceae. = £. subreniforme Reveal Wash- Eriogonum corymbosum Benth. in DC. var. davidsei ington Co., Springdale, Eastwood & Howell 1171, 1933 Reveal Great Basin Naturalist 27: 216. 1967. Polyg- (CAS!;US!;NY!). onaceae. Carbon Co., Wellington, Reveal & Davidse Eriogonum filiforme L.O. Williams Bull. Torrey Bot. 956, 1967 (US!;BRY!;UT!;RM!;NY!;UTC!;ISC!). Club 59: 428. 1932. Polygonaceae. = £. wetherillii Eastw. Wayne Co., w Hanksville, Garrett 5975, 1931 Eriogonum corymbosum Benth. in DC. var. divarica- (UT!;RM!). tum T. & G. in Beckwith Rep. U.S. Explor. & Surv. R.R. Eriogonum flexum Jones var. ferronis Jones Contr. Pacific 2: 29. 1857. Polygonaceae. Emery Co., Green W. Bot. 11: 15. 1903. Polygonaceae. = £. flexum Jones River, Creutzfeldt sn, 1853 (NY!). Emery Co., w Perron, Jones 5454, 1894 (US!;POM!). Eriogonum corymbosum Benth. in DC. var. erectum Eriogonum friscanum Jones Contr. W. Bot. 11: 14. Reveal & Brotherson in Reveal Great Basin Naturalist 1903. Polygonaceae. = £. microthecum var. foliosum (T. 27: 213. 1968. Polygonaceae. Wasatch Co., W Duch- & G.) Reveal Beaver Co., Frisco, Jones (POM?). esne, Holmgren & Reveal ,3022, 1966 (UTC!;US!; Eriogonum grayii Reveal Phytologia 25: 193. 1973. BRY!;UT!;NY!;ISC!). Polygonaceae. Salt Lake Co., Lake Blanche, Holmgren Eriogonum corymbosum Benth. in DC. var. mat- et al. 7121, 1947 (UTC!;BRY!;ISC!). thewsiae Reveal Phytologia .35: 441. 1976. Polygonaceae. Eriogonum heracleoides Nutt. var. utahense Gandg. Washington Co., Springdale, al. Welsh et 9509, 1969 Bull. Soc. Bot. Belgium 42: 190. 1906. Polygonaceae. = (BRY!;UT!). £. heracleoides Nutt. var. heracleoides Cache Co., Lin- Eriogonum crispum L.O. Williams Bull. Torrey Bot. ford sn, 1897 (US!). Club 59: 427. 19,32. Polygonaceae. = E. corymbosum Eriogonum hookeri Wats. Proc. Amer. Acad. 14: 295. var. ghitinosum (Jones) Jones Iron Co., Cedar Canyon, 1879. Polygonaceae. Syn: £. deflexum ssp. hookeri var. Garrett 6027, 1931 (RM!;UT!). gilvum Stokes Utah Co., American Fork Canyon, Wat- Eriogonum cronquistii Reveal Madrono 19: 289. 1969. son 10,33, 1869 (US!;NY!). Polygonaceae. Garfield Co., Henry Mts., Holmgren & Eriogonum howellianum Reveal Phytologia 25: 204. Reveal .3010, 1966 (BRY!;RM!;UT!;NY!;US!;UTC!;ISC!). 1973. Polygonaceae. Syn: £. glandidosum authors, not (Nutt.) Nutt. Millard se Garrison, al Eriogonum deflexum Terr, in Ives ssp. hookeri var. Co, Holmgren et 2248, 1965 (US!;BRY!;UT!;RM!;UTC!). gilvum Stokes Genus Eriogonum 45. 1936. Polygonaceae. Eriogonum humivagans Reveal Madrono 19: 219. = E. hookeri Wats. Davis Co., Antelope Island, Cottam 1969. Polygonaceae. San Juan Co., 13.5 miles e Mon- 6407, 19,35 (UT!;UTC!). ticello, Holmgren & Reveal 3001, 1969 (UTCi; Eriogonum deflexum Torr. in Ives ssp. ultrum Stokes US!;RM!;BRY!;UT!;NY!;ISC!). Genus Eriogonum 45. 1936. Polygonaceae. = £. nutans Eriogonum hylophilum Reveal & Brotherson Great var. nutans Sevier Co., Sevier Valley, Eastwood & How- Basin Naturalist 27: 190. 1968. Polygonaceae. Duchesne ell 623, 1933 (CAS!). Co, Gate Canyon, Holmgren & Reveal 3017, 1966 Eriogonum duchesnense Reveal (hybrid) Great Basin (UTC!;US!;RM!;BRY!;UT!;NY!;ISC!). Naturalist 27: 194. 1968. Polygonaceae. Syn: E. corymbo- Eriogonum insigne Wats. Proc. Amer. Acad. 14: 295. sum var. albogilvum Reveal Duchesne Co., Indian Can- 1879. Polygonaceae. Iron Co., Red Creek, Palmer 431, yon, Reveal 678, 1964 (UTC;US!;BRY!;NY!;UTC!). 1877 (NY!;BRY!;ISC!). Eriogonum dudleyanum Stokes Genus Eriogonum 90. Eriogonum intermontanum Reveal Madroiio 19: 293. 1936. Polygonaceae. = £. kearneyi Tidestr. var. 1969. Polygonaceae. Grand Co., Middle Canyon, Roan kearneyi Tooele Co., Skull Valley, Jones sn, 1896 Cliffs, Holmgren et al. 2278, 1965 (UTC!;BRY!; (CAS!;UT!). UT!;RM!;ISC!). Eriogonum effusum Nutt. var. durum Stokes Genus Eriogonum jamesii Benth in DC. var. rupicola Reveal Eriogonum 80. 1936. Polygonaceae. = E. corymbosum Phytologia 25: 202. 1973. Polygonaceae. Washington var. corymbosum Carbon Co., Sunnyside, Jones 1113a, Co., Zion National Park, Reveal & Reveal 2874, 1972 1901 (POM!;NY!). (US!;BRY!;NY!;UTC!). June 1982 Welsh: Utah Plant Types 169

Eriogonum kearneyi Tidestr. Proc. Biol. Soc. Wash- Eriogonum palmeri Wats. Proc. Amer. Acad. 14: 267. ington 26: 122. 1913. Polygonaceae. Syn: E. dtidleyanum 1879. Polygonaceae. = £. plumatella Dur. & Hilg.

Stokes Tooele Co., w Tooele, Kearney & Shantz 3218 Washington (?) Co., Palmer sn, 1870 (US!). (US!). Eriogonum parryi Gray Proc. Amer. Acad. 10: 77. Eriogonum kingii var. laxifolitim T. & G. Proc. Anier. 1874. Polygonaceae. = £. brachypodum T. & G. Wash- Acad. 8: 165. 1870. Polygonaceae. = £. brevicaule Nutt. ington Co., St. George, Parry 239, 1874 (US!;NY!;BRY!). var. kixifoUum (T. & G.) Reveal Salt Lake Co., Parleys Eriogonum pauciflorum var. panguicense Jones Peak, Watson 1021, 1869 (US!). Contr. W. Bot. 11: 9. 1903. Polygonaceae. = £. pan- Eriogonum lancifolium Reveal & Brotherson in Re- guicense (Jones) Reveal Garfield Co., Panguitch, Jones veal Great Basin Naturalist 27: 188. 1967. Polygonaceae. sn, 1890 (POM!). Carbon Co., e Wellington, Reveal & Davidse 955, 1967 Eriogonum pharnaceoides Torr. in Sitgr. var. cervi- (UTC!;US!;RM!;BRY!;UT!;NY!;ISC!). num Reveal Great Basin Naturalist 34: 245. 1974. Polyg- Eriogonum leptocladon T. & G. in Beckwith var. lep- onaceae. Washington Co., Pine Valley Mts., Atwood & tocladon Rep. U.S. Explor. & Surv. R.R. Pacific 2: 129. Higgins 5895, 1973 (US!;BRY!;RM!;UTC!). 1857. Polygonaceae. Syn: E. effusum spp. pcdlidum var. Eriogonum porteri Small Bull. Torrey Bot. Club 25: shandsii Stokes Einerv Co., Green River, Creuzfeldt sn, 41. 1898. Polygonaceae. = £. umbellatum Torr. var. 1853 (NY!). porteri (Small) Stokes Summit Co., Bear River Canyon, Eriogonum hganum A. Nels. Bot. Gaz. 54: 149. 1912. Watson 1014, 1869 (US!;NY!). Polygonaceae. Cache Co., Logan, Smith 1704, 1909 Eriogonum puberulum Wats. Proc. Amer. Acad. 14: (RM!;BRY!). 295. 1879. Polygonaceae. Iron Co., Red Creek, Palmer Eriogonum longilobum Jones Proc. Calif. Acad. II, 5: 429, 1877 (US!;NY!;BRY!;ISC!). 720. 1895. Polygonaceae. = E. shockleyi var. longilobum Eriogonum pulvinatum Small Bull. Torrey Bot. Club (Jones) Reveal Carbon Co., near Price, Jones 5590, 1894 25: 44. 1898. Polygonaceae. = £. shockleyi Wats. var. (POM!;US!). shockleyi Beaver Co., Milford, Jones 1775, 1880 Eriogonum medium Rydb. Fl. Rocky Mts. 220, 1061. (US!;UT!;POM!;NY!;UTC!). 1917. Polvgonaceae. = E. brevicaule Nutt. var. laxifo- Eriogonum ramosissimum Eastw. Proc. Calif. Acad. = lium (T. & G.) Reveal Juab or Utah Co., Mt. Nebo, Ryd- II, 6: 322. 1896. Polygonaceae. £. leptocladon T. & G. (Eastw.) Co., Butter berg sn, 1905 (US!;RM!;NY!). var. ramosissiinum Reveal San Juan Eriogonum nanum Reveal Phytologia 25: 194. 1973. Spring, Eastwood 134, 1895 (US!;CAS!). Welsh Great Basin Naturalist Polygonaceae. Box Elder Co., Willard Peak, Reveal & Eriogonum revealianum 30: 17. 1970. Polygonaceae. = £. var. re- Holmgren 665, 1964 (UT!;BRY!;RM!). corymbosum (Welsh) Reveal Garfield Co., s Antimony, Eriogonum natum Reveal in Welsh, Atwood, & Re- vealianum 1969 (BRY!;US!;UT!;RM!;NY!). veal Great Basin Naturalist 35: 363. 1975. Polygonaceae. Welsh & Welsh 9389, Zoe 4: 281. 1893. Polyg- Millard Co, 43 mi w Delta, Reveal & Reveal 3924, 1975 Eriogonum rubiflorum Jones = (BRY!;NY!;UTC!). onaceae. E. nutans T. & G. var. nutans Tooele Co., sn, 1891 (POM!;US!;NY!). Eriogonum nehonii L.O. Williams Bull. Torrey Bot. Dugway, Jones saurinum Reveal Great Basin Naturalist Club 59: 428. 19'37. Polygonaceae. = £. microtheciim Eriogonum 27: 197. 1967. Polygonaceae. Uintah Co., 10 mi e Vernal, Nutt. var. foliosum (Torr.) Reveal San Juan Co., Geyser Holmgren Reveal 3019, 1966 Basin, Walker .368, 1912 (RM!;UT!). & (UTC!;UT!;BRY!;US!;RM!;NY!;ISC!). Eriogonum nudicaule Small ssp. garrettii Stokes Eriogonum scabrellum Reveal Ann. Missouri Bot. Genus Eriogonum 8.3. 19.36. Polygonaceae. = E. brevi- 55: 74. 1968. Polygonaceae. Grand Co, West- caule Nutt. var. brevicaule Summit Co., Echo Reservoir, Card. water, Reveal Davidse 949, 1967 Garrett 7068, 19.35 (US!;UT!;UTC!). & (UTC!;US!;BRY!;UT!;RM!;NY!;ISC!). Eriogonum nudicaule Small ssp. ochroflorum Stokes smithii Reveal Great Basin Naturalist 24: Genus Eriogonum 83. 1936. Polygonaceae. = E. spa- Eriogonum 1967. Polygonaceae. Emery Co., San Rafael Desert, thulatum Gray Sevier Co., Clear Creek Canyon, Jones 202. &' .30i2, 1966 (UTC!:US!; 6104, 1894 (p6m!;BRY!). Holmgren Reveal BRY!;UT!;RM!;ISC!). Eriogonujn nudicaule Small ssp. parleyense Stokes Eriogonum soredium Reveal Great Basin Nat. 41: 229. Genus Eriogonum 84. 19.36. Polygonaceae. = E. brevi- 1981. Polygonaceae. Beaver Co., Grampian Hill, near caule Nutt. var. brevicaule Salt Lake Co., Parleys Can- al. (BRY!). yon, Stokes 213, 1934 (CAS!;NY!;UTC!;BRY!). Frisco, Welsh et 20192,1980 Eriogonum spathuliforme Rydb. Bull. Torrey Bot. Eriogonum nummulare Jones Contr. W. Bot. 11: 13. 1912. Polygonaceae. = E. ostlundii Jones 1903. Polvgonaceae. Tooele Co., Dutch Mt., Jones sn Club 39: 307. (POM?). Piute Co., near Mt.Belknap, Stokes sn, 1900 (US!). Gray Proc. Amer. .\cad. 10: Eriogonum ochrocephalum var. angustum Jones Eriogonum spathulatum 76. 1874. Polygonaceae. Syn: £. nudicaule Small ssp. Contr. W. Bot. 11: 9. 1903. Polygonaceae. = £. brevi- ochroflorum Stokes Sevier Co., Sevier River Valley, caule Nutt. var. laxifolium (T. & G.) Reveal Tooele Co., (?) Parry 245, 1874 (ISC!). Johnsons Pass, Jones sn (POM?). Eriogonum subreniforrne Wats. Proc. Amer. Acad. 12: Eriogonum ostlundii Jones Contr. W. Bot. 11: 12. 1877. Polygonaceae. Syn: £. Stokes Wash- 1903. Polvgonaceae. Sevier Co., near Elsinore, Jones sn 260. filicaide ' (POM?). ington Co., Parry 237, 1874 (ISC!). sulcatum Wats. Proc. Amer. Acad. 14: Eriogonum ovalifolium Nutt. var. utahense Gandg. Eriogonum 296. 1879. Polygonaceae. = £. heennanii var. sulcatum Bull. Soc. Bot. Belgium 42: 191. 1906. Polygonaceae. = (Wats.) Reveal Washington Co., near St. £. ovalifolium Nutt. var. ovalifolium Cache Co., Linford Munz & George, Palmer 432, 1877 (US!;NY!;BRY!). sn, 1897 (US!). 170 Great Basin Naturalist Vol. 42, No. 2

Eriogonum sulcatum Wats. var. argense Jones Contr. Eritrichium pterocaryum Torr. var. pectinatum Gray W. Bot. 11: 15. 1903. Polygonaceae. = E. heermannii Proc. Amer. Acad. 10: 61. 1874. Boraginaceae. = Cryp- Dur. & Hilg. var. argense (Jones) Munz Utah, Jones. tantha pterocarya var. pterocarya Southern Utali, Parry Eriogonum tenellum ssp. cottamii Stokes Genus sn, 1874 (NY!).

Eriogonum 70. 1936. Polygonaceae. = E. brevicaiile var. Eritrichium setosissimum Gray Proc. Amer. Acad. 12: cottamii (Stokes) Reveal Utah Co., West Mountain, Cot- 80. 1876. Boraginaceae. = Cryptantha setosissima tarn 411, 1925(BRY!;UT!). (Gray) Payson Sevier Co., Fish Lake, Ward sn, 1875 Eriogonum tenellum var. grandiflorum Gandg. Bull. (GH). Soc. Bot. Belgium 42: 197. 1906. Polygonaceae. = E. mi- Erysimum asperum (Nutt.) DC. var. purshii Durand crothecum var. laxiflorum Hook. Rich Co., Linford sn, Trans. Amer. Phil. Soc. II. 159. 1860. Brassicaceae. Syn:

1897 (?). E. capitatum (Dougl.) Greene Salt Lake Co., near Great Eriogonum thompsonae Wats. var. albiflorum Reveal Salt Lake, Salt Lake Valley, Carrington sn, 1857 (P). Madroiio 19: 299. 1969. Polygonaceae. Washington Co., Erythrocoma brevifolia Greene Leafl. Bot. Obs. & w Virgin, Holmgren & Reveal 2991, 1966 (UTC!; Grit. 1: 176. 1906. Rosaceae. = Geum triflorum Pursh US!;BRY!;UT!;RM!;NY!;ISC!). Garfield Co., Panguitch Lake, Jones 6002q, 1894 (US!). Eriogonum thompsonae Wats. var. thompsonae Amer. Erythronium utahense Rydb. Fl. Rocky Mts. 165, Naturalist 7: .302. 1873. Polygonaceae. Kane Co., Kanab, 1061. 1917. Liliaceae. = E. grandiflorum Pursh Salt Thompson sn, 1872 (GH;US!;BRY!;NY!). Lake Co., Salt Lake, Stansbury sn, 1850 (NY!). Eriogonum triste Wats. Proc. Amer. Acad. 10: 347. Eschscholzia ludens Greene Pittonia 5: 272. 1905. Pa- pa veraceae. = E. minutiflora Wats. Washington Co., St. 1875. Polygonaceae. = E. alatum Torr. in Sitgr. Kane George, Jones 5110a, 1894 (US!;POM!). Co., Siler sn, 1873 (US!;NY!;BRY!;ISC!). Euphorbia nephradenia Barneby Leafl. W. Bot. 10: Eriogonum umbellatum Torr. var. desereticum Reveal 314. 1966. Euphorbiaceae. Kane Co., Cottonwood Can- in Welsh, Atwood, & Reveal Great Basin Naturalist 35: yon, Barneby 14434, 1966 (NY!;US!;BRY!;UTC!). .365. 1975. Polygonaceae. Utah Co., Mt. Timpanogos, Euphorbia parryi Engelm. in Parry Amer. Naturalist near Timpooneke Campground, Reveal 3702. 1974 9: .350. 1875. Euphorbiaceae. Wa.shington Co., near St. (US!;BRY!;NY!;UTC!). George, Parry s47, 1874 (MO). Eriogonum umbellatum Torr. var. glabratum Stokes Euphorbia podagrica I.M. Johnston Univ. Calif. Publ. Genus Eriogonum 109. 1936. Polygonaceae. = E. um- Bot. 7: 440. 1922. Euphorbiaceae. Utah, Purpus 6432, bellatum Torr. var. aureum ( Gandg.) Reveal Emery Co., 1898 (US!). Huntington Canyon, Garrett 7036, 1935 (UT!). Euphorbia robusta (Engelm.) Small ex Britt. & Eriogonum villiflorum Gray var. tumulosum Barneby Brown var. interioris J.B.S. Norton N. Amer. Euphorbia Leafl. W. Bot. 5: 153. 1949. Polygonaceae. = £. tumulo- 49. 1899. Euphorbiaceae. = E. robusta (Engelm.) Small sum (Barneby) Reveal Emery Co., sw Woodside, Ripley Wasatch Mts., Watson 1081, 1869 (US!). & Barneby 8678, 1947 (CAS!;NY!;BRY!;UTC!). Eurotia subspinosa Rydb. Bull. Torrey Bot. Club 39. Eriogonum viridulum Reveal Proc. Utah Acad. 42: 313. 1912. Chenopodiaceae. = Ceratoides subspinosa 287. 1966. Polygonaceae. Duchesne Co., 8 mi e Duch- (Rydb.) J.T. Howell Washington Co., St. George, Goodd- esne, Reveal 675, 1964 (UTC!;BRY!;UT!;RM!;NY!;ISC!). ing 810, 1903 (US!). Festuca brevifolia R. Br. var. utahensis St-Yves Can- Eriogonum wasatchense Jones Contr. W. Bot. 11: 11. dollea 2: 257. 1925. Poaceae. = F. ovina L. Salt Lake 1903. Polygonaceae. = £. brevicaule var. wasatchense Co., Alta, M.E.D. (possibly M.E.Jones) sn, 1895 (NY!). (Jones) Reveal Utah Co., American Fork Canyon, Jones Festuca dasyclada Hack, ex Beal Grasses N. Amer. 2: 1877, 1880(US!;POM!;UTC!). 602. 1896. Poaceae. Emery (?) Co., Parry 93, 1874 Eriogonum wetherillii Eastw. Proc. Calif. Acad. II, 6: (US!;ISC!;NY!). 319. 1896. Polygonaceae. Syn: E. filiforme L.O. Williams Festuca jonesii Vasey Contr. U.S. Natl. Herb. 1: 278. San Juan Co., San Juan River, Eastwood 124, 1895 1893. Poaceae. = F. subukita Trin. Salt Lake Co., City (CAS!). Creek Canyon, Jones sn, 1880 (US!;POM!). Eriogonum zionis T. Howell Leafl. W. Bot. 2: 253. J. Fragaria virginiana Duchesne var. glauca Wats. Rep. 1940. Polygonaceae. Washington Co, Zion National U.S. Geol. Explor. 40th Parallel, Bot. 5: 280. 1871. Ro- Park, Eastwood & Howell 6344, 1938 (CAS!). saceae. = F. virginiana Duchesne Summit Co., Parleys Eritrichium barbigerum Gray Synop. Fl. N. Amer. Park, Watson .322, 1869 (US!;NY!). 2(1): 194. 1878. Boraginaceae. = Cryptantha barbigera Frasera albomarginata Wats. Rep. U.S. Geol. Explor. (Gray) Greene Washington Co., St. George, Parry 171, 40th Parallel, Bot. 5: 220. 1871. Gentianaceae. = 1874 (GH;ISC!). Sivertia albomarginata (Wats.) Kuntze Washington Co., Eritrichium elongatum Wight var. paysonii Johnst. J. near St. George, Palmer, 1879 (US!;NY!). Arnold Arb. 33: 67. 1952. Boraginaceae. = £. nanum Frasera utahensis Jones Zoe 2: 14. 1891. Gen- var. elongatum (Wight) Cronq. Summit Co., Uinta Mts, tianaceae. = Swertia paniculata (Torr.) St. John Kane (?) Maguire et al 14385, 19.36 (RM!). Co., (possibly House Rock Valley), Buckskin Mts., Jones Eritrichium holopterum Gray Proc. Amer. Acad. 12: sn, 1890 (US!;POM!). 81. 1876. Boraginaceae. = ? Southern Utah, Bishop sn, Fraxintts anomala Torr. in Wats. Rep. U.S. Geol. Ex- 1873 (GH?). plor. 40th Parallel, Bot. 5: 283. 1871. Oleaceae. San Juan

Eritrichium holopterum Gray var. submolle Gray (?) Co., Labrynth Canyon, Newberry sn, 1859 (NY!). Proc. Amer. Acad. 13: 374. 1878. Boraginaceae. = Cryp- Fritillaria dichroa Gandg. Bull. Soc. Bot. France 66: tantha utahensis (Gray) Greene Washington Co., St. 291. 1920. Liliaceae. = F. pudica (Pursh) Spreng. Cache George, Palmer sn, 1870 (NY!). Co., Linford sn, 1897 (?). June 1982 Welsh: Utah Plant Types 171

Gaiidg. Bull. Soc. Bot. France 66: Fritillaria leucella Geranium marginale Rydb. ex Hanks & Small N. 291. 1920. Liliaceae. = F. pudicci (Pursh) Spreng. Cache Amer. Fl. 25: 16. 1907. Geraniaceae. Garfield Co., Co., Linford sii. 1897 (?). Aquarius Plateau, Rydberg & Carlton 7401, 1905 (NY!). Fritillaria utahensis Gandg. Bull. Soc. Bot. France Gilia aggregata (Pursh) Spreng. ssp. aggregata var. 66: 291. 1920. Liliaceae. = F. pticUca (Pursh) Spreng. attenuata f. utahensis Brand Pflanzenreich 4. Fani. 250: Cache Co., Linford sn, 1897 (?). 116. 1907. Polemoniaceae. = G. aggregata var. riiacrosi- Gaillardia acaulis Gray Proc. Anier. Acad. 10: 73. phon Kearney & Peebles Salt Lake Co., Alta, Jones 1122, 1874. Asteraceae. = G. parnji Greene Washington Co., 1879 (POM !;UTC!). St. George, Parrv 120, 1874 (NY!;NDG!). Gilia arenaria Benth. var. rubella Brand in Engler Gaillardia crassifolia Nels. & Macbr. Bot. Gaz. 61: Pflanzenreich 4. Fam. 250: 103. 1907. Polemoniaceae. = 46. 1916. Asteraceae. = G. pinnatifida Torr.? Washing- G. hutchinsifolia Rydb. Washington Co., St. George, ton Co., LaVerkin, Jones 5177, 1894 (US!;NY!;BRY!). Jones 1651, 1880 (Us'!;POM!;NY!). Gaillardia flava Rydb. N. Amer. Fl. .34: 139. 1915. Gilia caespitosa Gray Proc. Amer. Acad 12; 80. 1876. -Asteraceae. Emerv Co., Lower Crossing, Jones 6412, Polemoniaceae. Wayne Co., Rabbit Valley, Ward 575, 1898(US!;BRY!;POM!). 1875 (US!;NY!;BRY!). Gaillardia gracilis A. Nels. Bot. Gaz. 37: 276. 1904. Gilia congesta Hook. var. nuda Eastw. Proc. Calif. Asteraceae. = G. pinnatifida Torr. Washington Co., Di- Acad. II, 6: 308. 1896. Polemoniaceae. = G. roseata amond Valley, Goodding 894, 1902 (US!;RM!). Rvdb. Syn: G. nuda (Eastw.) Rydb. San Juan Co., Wil- Gaillardia spathulata Gray Proc. Amer. Acad. 12: 59. low Creek, Eastwood 80, 1895 (US!;CAS!). 1876. Asteraceae. Wavne Co.', Rabbit Valley, Ward 401, 1875(US!;NY!). Gilia congesta Hook. var. paniculata Jones Proc. Ca- = Gaillardia straminea A. Nels. Univ. Wyoming Publ. lif. Acad. II, 5: 712. 1895. Polemoniaceae. G. congesta var. congesta Emery Co., Huntington, Jones 5464m, Bot. 1: 137. 1926. Asteraceae. = G. pinnatifida Torr. Washington Co., LaVerkin, Jones 5177, 1894 1894 (POM!;US!). (RM!;\Y!:BRY!). Gilia debilis Wats. Amer. Naturalist 7: .302. 1873. Po- Galium bifolium Wats. Rep. U.S. Geol. E.\plor. 40th lemoniaceae. = Collomia debilis (Wats.) Greene Salt Mts., Salt City, Wheeler Parallel, Bot. 5: 134. 1871. Rubiaceae. Salt Lake Co. (?), Lake Co., Wasatch above Lake Wasatch Mts., Watson 480, 1869 (US!;NY!). sn, 1872 (US!). Galium desereticum Dempst. & Ehrend. Brittonia 17: Gilia depressa Jones in Gray Proc. Amer. Acad. 16: 314. 1965. Rubiaceae. = G. mtiltiflonnn var. multi- 106. 1880. Polemoniaceae. Syn: Ipomopsis depressa florum Juab Co., Fish Springs, Jones sn, 1891 (Jones) V. Grant Millard Co., Deseret, Jones 1772, 1880 (US!;POM!). (GH;US!:CAS!;POM!;BRY!;NY!;UTC!). Galium filipes Rydb. Fl. Rocky Mts. 809, 1066. 1917. Gilia filiformis Parry ex Gray Proc. Amer. Acad. 10: Rubiaceae. = G. mexicanum var. asperuhim Gray Salt 75. 1874. Polemoniaceae. Washington Co., near St. Lake Co., Citv Creek Canyon, Leonard 16665, 1883 George, Parry 187, 1874 (NY!;ISC!;BRY!). (NY!). Gilia floribunda var. arida Jones Proc. Calif. Acad.

Galium hypotrichium Gray ssp. scabriusculum Eh- II, 5: 713. 1895. Polemoniaceae. = Leptodactylon wat- rend. Contr. Dudley Herb. 5: 13. 1956. Rubiaceae. = G. sonii (Gray) Rydb. Wayne Co., Capitol Wash, Jones multiflorum var. multiflorum Emery Co., Calf Spring 5701a, 1894(POM!;US!).' Wash, Maguire 18437, 1940 (CAS!;Utc!). Gilia frutescens Rydb. Bull. Torrey Bot. Club 40: 471. Galium hypotrichium ssp. utahense Ehrend. Contr. 1913. Polemoniaceae. = G. congesta var. frutescens Dudley Herb. 5: 12. 1956. Rubiaceae. = G. multiflorum (Rvdb.) Cronq. Washington Co., Springdale, Jones 5247, var. multiflorum Utah Co., Provo, Goodding 1116, 1902 1894 (NY!;US!). (ISC!). Gilia gracilis Hook. ssp. spirillifera var. nana Brand var. Gray Syn. Galium multiflorum Kellogg watsonii in Engler Pflanzenr. 4. Fam. 250: 92. 1907. Polemo- 1884. = multi- Fl. N. Amer. 1(2): 40. Rubiaceae. G. niaceae. = Microsteris gracilis (Hook.) Greene Millard var. Syn: G. ivatsonii (Grav) Heller florum multiflorum Co., Fillmore, Jones 1684 (POM!;UT!). Wasatch Mts., Watson 484, 1869 (US!;NY!). Gilia inconspicua (J.E. Sm.) Sweet ssp. eu- Galium scabriusculum (Ehrend.) Dempst. & Ehrend. incotispicua var. variegata Brand in Engler Pflanzenr. ssp. protoscabriusculum Dempst. & Ehrend. Brittonia 4. Fam. 250: 105. 1907. Polemoniaceae. = G. inconspic- 17: 312. 1965. Rubiaceae. = G. multiflorum var. multi- ua (J.E. Sm.) Sweet Kane Co., Kanab, Jones 5280, 1894 florum Carbon Co.. Castle Gate, Ehrendorfer & Stutz 5954. 1959(RM!:UTC!). Gilia latifolia Wats, in Parry Amer. Naturalist 9: 347. Galium utahense Eastw. Leafl. W. Bot. 1: 55. 1933. 1875. Polemoniaceae. Washington Co., near St. George, Rubiaceae. = G. horeale L. Wasatch Co., Soldier Sum- Parry 188, 1874 (GH;US!;CAS!;NY!;BRY!;ISC!). mit, Eastwood 7668. 1918 (CAS!). Gilia leptomeria Gray Proc. Amer. .-Kcad. 8: 278. Gentiana calycosa Griseb. ssp. apetala Maguire 1870. Polemoniaceae. Syn: G. leptomeria var. tridentata Madrono 6: 151. 1942. Gentianaceae. = G. calycosa Jones; G. triodon Eastw. Tooele Co., Stansbury Island, Griseb. Duchesne Co., w Mt. .\gassiz, Maguire et al. Watson 27, 1869 (NY!). 4225. 19.33 (UTC!). Gilia leptomeria Gray var. tridentata Jones Proc. Ca- Gentiana tortuosa Jones Proc. Calif. .\cad. II, 5: 707. lif. Acad. II, 5: 713. 1895. Polemoniaceae. = G. lepto- 1895. Gentianaceae. = Gentianella tortuosa (Jones) Gil- meria Gray Emery Co., Emery, Jones 5445n, 1894 lett Garfield Co., Panguitch Lake, Jones 6008, 1894 (POM;US!).' (POM!:US!). 172 Great Basin Naturalist Vol. 42, No. 2

Gilia incvickerae Jones Proc. Calif. Acad. II, 5: 712. Cymnolomia his])ida Robins. & Greenm. var. ciliata 1895. Polemoniaceae. = G. pinnatifida Nutt. Piute Co., Robins. & Greenm. Proc. Boston Soc. Nat. Hist. 29: 93. Marysvale, Jones 5-378, 1894 (POi\1!;US!;CAS!; 1899. .Asteraceae. = Heliomeris hispida (Grav) Cockerell BRY!:RM!;NY!). Syn: Viguiera ciliata (Robins. & Greenm.) Blake South- Gilia scoptilontm Jones Bull. Torrey Bot. Club 8; 70. ern Utah, Palmer 245, 1877 (US!). 1881. Polemoniaceae. Syn: G. scopulorum var. deformis Gymnolomia linearis Rydb. Bull. Torrey Bot. Club Brand Washington Co., St. George, Jones 1659, 1880 37: 327. 1910. Asteraceae. = Heliomeris multiflora var.

(US!;POM!;NY!;UTC!). nevadensis (A. Nels.) Yates Washington Co., St. George, Gilia scopulorum Jones var. deformis Brand Pflan- Palmer 241, 1877 (NY). zenr. 4. Fani. 2.50; 109. 1907. Polemoniaceae. = G. sco- Gymnolomia multiflora var. annua Jones Proc. Calif. ,\cad. II, 5: 698. 1895. = pulonnn Jones? .Southern Utiili, Parry 198, 1874 (?). Asteraceae. Heliomeris long- var. Gilia stenothrysa Gray Proc. Amer. Acad. 8: 276. ifolia annua (Jones) Yates Utah?, Jones ?, (?). Hackelia ibapensis Shultz Shultz Brittonia 1870. Polemoniaceae. Duchesne (?) Co., "Among cedars & 33: 157. 1981. Boraginaceae. Co., between Duchesne and Lake Fork," Fremont sn, 1845 Juab Deep Creek Range, L. & Shultz 4:^50, 1980 (NY!;UTC!). (NY!). J. Hackelia patens (Nutt.) Johnst. var. harrisonii Gentry Gilia straminea Rydb. Bull. Torrev Bot. Club 40: 472. Southw. Naturalist 19: 140. 1974. Boraginaceae. Wash- 1913. Polemoniaceae. = G. inconspicita var. sinuata ington Co., Pine Valley, Gentry 2002, 1968 (NY!; (Hook.) Gray Washington Co., St. George, Palmer 325, ' CAS!;BRY!:UTC!). 1877 (NY!). Halostachys occidentalis\ Wats. Rep. U.S. Geol. Ex- Gilia superba Eastw. Zoe 4: 123. 1893. Polemo- plor. 40th Parallel, Bot. 5: 293. 1871. Chenopodiaceae. niaceae. = G. siihnuda Torr. San Juan Co., Hatches = AUcnrolfia occidentalis (Wats.) Kuntze Box Elder Wash, Eastwood sn, 1892 (CAS!;POM!). Co., Raft River Valley, Watson 995, 1869 (US!). Gilia tenerrima Gray Proc. Amer. Acad. 8: 277. 1870. Hamosa atratiformis Rydb. Bull. Torrey Bot. Polemoniaceae. Summit Co., Bear River Vallev, Wat.son Club 34: 48. 1907. Fabaceae. = Astragalus 922, 1869 (NY!). straturensis Jones Wahington Co., Parry 47, 1874 (NY!;ISC!). Gilia tenuituba Rydb. Bull. Torrey Bot. Club 40: 472. Haplopappus cervinus Wats. Amer. Naturalist 7: 301. 1913. Polemoniaceae. = G. aogregata var. macwsiphon 1873. Asteraceae. Utah, Antelope Canvon, Wheeler sn, Kearney & Peebles Beaver (?) Co., Beaver City, Palmer 1872 (US!). 320, 1877 (NY!;US!). Haplopappus nuttallii T. & G. var. depressus Ma- Gilia tridactyla Rydb. Fl. Rocky Mts. 692,1065. 1917. guire .Amer. Midi. Naturalist 37: 144. 1947. Asteraceae. Polemoniaceae. Piute Co., Brigham Peak, Jones .5949, = M. griudelioides var. depressa (Maguire) Cronq. 1894(US!;NY!;POM!). & Keck Millard Co., Desert Experimental Range, .Maguire Gilia triodon Eastw. Zoe 4: 121. 1893. Polemo- 20859, 1941 (US!;NY!;UTC!). niaceae. = G. leptomeria Gray San Juan Co., Ruin Can- Haplopappus parryi Gray var. minor Gray Svn. Fl. N. yon. Eastwood sn, 1892 (CAS!). Amer. 1(2): 131. 1884. Asteraceae. = Solidagv parryi Gilia watsonii Gray Proc. Amer. Acad. 8: 267. 1870. (Gray) Greene Salt Lake Co., Alta, Bald Mtn., Jones sn?, Polemoniaceae. = Leptodactyhn watsonii (Gray) Rydb. 1879(POM!:BRY!;NDG!). Salt Lake Co., Cottonwood Canyon, Watson sn, 1869 Haplopappus scopulorum (Jones) Blake in Tidestr. (NY!). var. hirtellus Blake Proc. Biol. Soc. Washington 48: 170. Glycosma maxima Rydb. Bull. Torrev Bot. Club 40: 1935. .Asteraceae. Iron Co., Cedar Canyon, Garrett 6051, 67. 1913. Apiaceae. = Osmorhiza occidentalis (Nutt.) 1931 (US!;UT!). Torr. Juab Co., Mt. Nebo, Rydberg & Carlton 7585, Hedysarum gremiale Rollins Rhodora 42: 230. 1940. 1905 (NY!). Fabaceae. = H. boreale var. gremiale (Rollins) North- Glyptopleura setulosa Gray Proc. Amer. .Acad. 9: 211. strom & Welsh Uintah Co., 14 mi n Vernal, Rollins 1874. Asteraceae. Washington Co., St. George, Palmer 6, 17.33, 19.37 (US!;CAS!;RM!;UTC!). 1870 (US!;NY!;BRY!). Hedysarum occidentale Greene var. canone Welsh Grayia brandegei Proc. Gray Amer. Acad. 11: 101. Great Basin Naturalist 38: 314. 1978. Fabaceae. Carbon 1876. Chenopodiaceae. San Co., R., Juan (?) San Juan Co., Soldier Creek, Welsh & Tavlor. 1.52.56, 1977 Brandegee sn, 1875 (GH), (BRY!;NY!). Grindelia laciniata Rydb. Fl. Rocky Mts. 848, 1066. Hedysarum utahense Rydb. Bull. Torrey Bot. Club = 1917. Asteraceae. G. fastigiata Greene San Juan Co., .34: 425. 1907. Fabaceae. = H. boreale var. boreale Salt

Montezuma Canyon, Rydberg & Garrett 9692, Lake Co., Salt Lake City, Leonard .55, 1883 (NY!). 1911(NY!;USI). Helianthella multicaulis D.C. Eaton in Wats. Rep.

Grindelia stylosa Eastw. Proc. Calif. Acad. II, 6: 293. U.S. Geol. Explor. 40th Parallel, Bot. 5: 170. 1871. Aste- 1896. Asteraceae. = Vanclevea stylosa (Eastw.) Greene raceae. = H. uniflora (Nutt.) T. & G. Summit Co., Par- San Juan Co., Bartons range, Eastwood 713, 1895 leys Park, Watson 605, 1869 (US!;NY!). (US!;CAS!;NY!). Helianthus anomalus Blake Washington Sci. J. Acad. Gutierrezia sarothrae (Pursh) Britt. & Rusby var. 21: .3.33. 1931. Asteraceae. Wayne Co.^ s Hanksville, pomariensis Welsh Great Basin Naturalist .30: 19. 1970. Stanton 4806, 19,30 (US!;UT!). Asteraceae. = Xantliocephahim sarothrae var. poma- Helianthus bracteatus E.E. Watson Papers .Michigan riensc (Welsh) Welsh Uintah Co., Orchard Creek Draw, Acad. Sci. 9: .393. 1929. Asteraceae. = H. nuttallii T. & Welsh etal. 9471, 1969 (BRY!). G. Cache Co., Logan, Mulford 177, 1898 (MO). June 1982 Welsh: Utah Plant Types 173

Helianthus deserticolus Heiser Proc. Indiana Acad. Hymenopappus nudipes Maguire Amer. Midi. Natu- Sci. 70: 209. 1961. Asteraceae. Washington Co., w Hur- ralist 37: 143. 1947. Asteraceae. = H. filifolius var. al- ricane, Stoiitamire 2574, 1957 (IND). pestris (Maguire) Shinners Kane Co., 15 mi n Orderville, Helianthus giganteus var. utahensis D.C. Eaton in Maguire 18740, 1940 (NY!;US!). Wats. Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: 169. Hymenopappus nudipes Maguire var. alpestris Ma- = 1871. Asteraceae. H. nuttallii T. & G. Wasatch Mts., guire Amer. Mid!. Naturalist 37: 144. 1947. Asteraceae. Watson 603, 1869 (US!). = H. filifolius var. alpestris (Maguire) Shinners Iron alipes Wats. var. pallidum C.L. Porter Hennidium Co., Cedar Breaks, Maguire 19023, 1940 (US!; = Rhodora 54: 158. 1952. Nyctaginaceae. Mirabilis NY!;UTC!). alipes (Wats.) Pi!z Uintah Co., 5 mi sw Vernal, Porter Hymenopappus pauciflorus Johnst. Contr. Gray 5308, 1950(RM!;CAS!). Herb. II, 68: 97. 1923. Asteraceae. = H. filifolius var. Hesperanthes alhomarginata Jones Zoe 2: 251. 1891. pauciflorus (Johnst.) Turner San Juan Co., near Bluff, Liliaceae. = Eremocrinum albomarginatum (Jones) Rydberg & Garrett 9951, I91I (UT!;NY!). Jones Emery Co., Green River, Jones sn, 1890 (?). Hymenopappus tomentosus Rydb. Bull. Torrey Bot. Heuchera rubescens Torr. in Stansb. Explor. Great Club 27: 633. 1900. Asteraceae. = H. filifolius var. to- Salt Lake 388. 1852. Saxifragaceae. Syn: H. versicolor mentosus (Rydb.) Turner Washington Co., St. George, Greene; H. versicolor i. puinila Rosend., Butters, & La- Palmer 270, 1877 (NY!;ISC!;BRY!). kela Davis (?) Co., Great Salt Lake, Stansbury sn, 1850 Hymenoxys lemmonii (Greene) Cockerell ssp. greenei (NY!). Cockerell Bull. Torrey Bot. Club 31: 479. 1904. Aste- Heuchera utahensis Rydb. N. Amer. Fl. 22: 114. 1905. raceae. = H. lemmonii (Greene) Cockerell Washington Saxifragaceae. = H. parvifolia Nutt. Salt Lake Co., City Co., Rock Creek, Palmer 261, 1877 (US!;NY!). Creek Canyon, Jones 1458, 1880 (US!;CAS!; (?) Hymenoxys richardsonii (Hook.) Cockerell var. uta- RM!;POM!;UT!;BRY!;UTC!). hensis Cockerell Bull. Torrey Bot. Club 31: 477. 1904. Heuchera versicolor Greene f. pumila Rosend., flut- Asteraceae. = H. richardsonii (Hook.) Cockerell Emery ters, & Lakela Minnesota Studies PI. Sci. 2: 85. 1936. Sa- Co., Emery, Jones 5442, 1894 (US!;POM!). xifragaceae. = H. rubescens Torr. Utali? (?). Ivesia utahensis Wats. Proc. Amer. Acad. 10: 71. Hieracium utahense Gandg. Bull. Soc. Bot. France 1874. Rosaceae. Salt Lake Co., Bald Mt., 1231, 65: 49. 1918. Asteraceae. = H. gracile Hook. Cache Co., Jones 1879(US!;NY!;UTC!)). Linford sn, 1897 (?). var. Bot. 12: Holodiscus microphyllus Rydb. Bull. Torrey Bot. Juncus canadensis kuntzei Buch. Jahrb. = Club 31: 559. 1904. Rosaceae. = H. dumosus (Nutt.) 272. 1890. Juncaceae. /. tweedyi Rydb. Box Elder Heller Salt Lake Co., Alta, Jones 1142, 1879 Co., near Corinne, Kuntze 3133, 1874 (NY!). (P0M!;BRY!;NY!;UTC!). Juncus jonesii Rydb. Fl. Rocky Mts. 153, 1061. 1917.

Homalobus canovirens Rydb. Fl. Rocky Mts. ed. 2. Juncaceae. = /. regelii Buch. Salt Lake Co., Alta, Jones 1126. 1922. Fabaceae. = Astragalus coltonii var. moa- 119, 1879(NY!;POM!). bensis Jones Grand Co., La Sal Mts, Rydberg & Garrett Juncus tracyi Rydb. Fl. Rocky Mts. 155, 1061. 1917.

85.36, 1913 (NY!). Juncaceae. = /. ensifolius Wikstr. Weber Co., Ogden, Homalobus humilis Rydb. Bull. Torrey Bot. Club 34: Tracy 389, 1887 (US!;NY!). 417. 1907. Fabaceae. = Astragalus miser var. oblongi- Juncus utahensis Martin Rhodora 40: 69. 1938. Jun- folius (Rydb.) Cronq. Piute Co., n Bullion Creek, Ryd- caceae. = /. ensifolius var. brunnescens (Rydb.) Cronq.

berg & Carlton 7147, 1905 (NY!;US!;RM!;BRY!). Summit (?) Co., Ashley National Forest, Nord & Sargent

Homalobus paucijugus Rydb. Bull. Torrey Bot. Club 1, 1927 (US!). 34: 418. 1907. Fabaceae. = Astragalus miser var. tenui- Juniperus californica var. utahensis Vasey Cat. For- (Nutt.) folius Barneby Syn: A. garrettii Macbr. Salt Lake est Trees U.S. 37. 1876. Cuprcssaceae. = /. osteosperma Co., Big Cottonwood Canyon, Garrett 1580, 1905 (Torr.) Little Wasatch Mts., Ward sn, 1875 (ISC!). (NY!;US!;UT!). Juniperus californica var. utahensis Engelm. Trans. Hordeum pusillum Nutt. var. pubens A.S. Hitchc. J. Acad. Sci. St. Louis 3: 588. 1878. Cupressaceae. = /. os- Washington Acad. Sci. 23: 453. 1933. Poaceae. = H. pu- teosperma (Torr.) Little Syn: /. utahensis (Engelm.) Lem- sillum Nutt. Washington Co., LaVerkin, Jones 5196ro, mon Washington Co., St. George, Palmer? sn, 1877 (?). 1894 (POM!). Kochia americana Wats. var. vestita Wats. Proc. Hosackia rigida Benth. var. nummularia Jones Proc. Amer. Acad. 9: 93. 1874. Chenopodiaceae. = K. ameri- Calif. Acad. II, 5: 633. 1895. Fabaceae. = Lotus long- cana Wats. Tooele Co., "Tuilla Valley" Watson 991, ebracteatus Rydb. Washington Co., Rockville, Jones 1869 (US!;NY!). 5224, 1894 (US!;RM!). Krynitzkia echinoides Jones Proc. Calif. Acad. II, 5: Houstonia saxicola Eastw. Proc. Calif. Acad. II, 6: 709. 1895. Boraginaceae. = Cryptantha ftdvocanescens 291. 1896. Rubiaceae. = H. rubra Cav. San Juan Co., var. echinoides (Jones) Higgins Garfield (?) Co., Pahria Butler Spring, Eastwood 33, 1895 (CAS!). Canyon, Jones 5297p, 1894 (US!;POM!). Hymenopappus eriopodus A. Nels. Bot. Gaz. 37: 274. Krynitzkia glomerata var. acuta Jones Zoe 2: 250. 1904. Asteraceae. = H. filifolius var. eriopodus (A. 1891. Boraginaceae. = Cryptantha wetherillii (Eastw.) Nels.) Turner Washington Co., Diamond Valley, Goodd- ing 880, 1902 (RM!;US!;NY!). Payson Grand Co., Cisco, Jones sn, 1890 (?). Krynitzkia glomerata var. virginensis Contr. Hymenopappus niveus Rydb. N. Amer. Fl. .34: 52. Jones 13: 5. = virgi- 1914. Asteraceae. = H. filifolius var. tomentosus (Rydb.) W. Bot. 1910. Boraginaceae. Cryptantha Turner Washington Co., Springdale, Jones 5261, 1894 nensis (Jones) Payson Washington Co., LaVerkin, Jones (US!;NY!). 5195a, 1894 (US!;RM!;POM!). 174 Great Basin Naturalist Vol. 42, No. 2

Krynitzkia leucophaea var. alata Jones Proc. Calif. Lepidium montanum Nutt. in T. & G. var. alpinum

Acad. II, 5; 710. 1895. Boraginaceae. = Cryptuntha con- Wats. Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: 29. fertiflora (Greene) Payson Kane Co., Johnson, Jones 1871. Brassicaceae. Salt Lake Co., Cottonwood Canyon, 5289f, 1894 (POM!;US!;'bRY!;NY!). Watson 122, 1869 (US!;NY!). Krynitzkia mensana Jones Contr. W. Hot. 13: 4. Lepidium montanum Nutt. in T. & G. var. demissum 1910. Boraginaceae. = Cryptuntha mensana (Jones) Pay- C.L. Hitchc. Madrofio 10: 157. 1950. Brassicaceae. = L. son Emery Co., Emery, Jones 5445p, 1894 (POM!; barnebyanum Reveal Duchesne Co., sw Duchesne, Rip- ' US!;RM!). ley & Barneby 8699, 1947 (US!;CAS!;NY!). Krynitzkia multicaulis var. setosa Jones Cotr. W. Lepidium montanum Nutt. in T. & G. var. neeseae Hot. 13: 4. 1910. Boraginaceae. = Cryptantha cinerea Welsh & Reveal Great Basin Naturalist 34: 334. 1977. (Greene) Cronq. Syn: C. jainexii var. setosa (Jones) Brassicaceae. Garfield Co., Hells Backbone, Neese & Johnst. Millard Co., near Cove Fort, Jones sn, 1901 White 3332, 1977 (BRY!;NY!;UT!). (POM!). Lepidium montanum Nutt. in T. & G. var. stellae Krynitzkia titahensis Gray Syn Fl. N. Anier. ed.2. Welsh & Reveal Great Basin Naturalist 34: .3.34. 1977.

2(1): 427. 1886. Boraginaceae. = Cryptantha iitahensis Brassicaceae. Kane Co., se Cannonville, Welsh & Welsh (Gray) Greene Washington Co., St. George, Palmer 352, 12841, 1975 (BRY!). 1877 (US!). Lepidium ostler Welsh & Goodrich Great Basin Natu-

Krynitzkia watsonii Gray Proc. .Amer. Acad. 20: 271. ralist 40; 80. 1980. Brassicaceae. Beaver Co., Frisco, Os- 1885. Boraginaceae. = Cryptantlia watsonii (Gray) tler & Anderson 1258, 1978 (BRY!). Greene Wasatch Mts., Watson 858, 1869 (US!;NY!). Lepidium utahense Jones Zoe 4: 266. 1893. Brassi- Langloisia setosissima (T. & G.) Greene var. camp- caceae. = L. integrifolium Nutt. Beaver Co., Milford, yloclados Brand Pflanzenreich 4. Fam. 250: 171. 1907. Jones 1821, 1880 (POM!;NY!;BRY!;UT!;UTC!;ISC!). Polemoniaceae. = L. setosissima (T. & G.) Greene Lepidium zionis A. Nels. Bot. Gaz. 42: 50. 1906. Washington Co., near St. George, Parry 190, 1874 Brassicaceae. = L. integrifolium Nutt. Sevier Co., Rich- (US!;ISC!). field, Jones 5411, 1894 (RM!;BRY!;POM!;NY!). Laphamia palmeri Gray Proc. Amer. Acad. 13: 372. Leptodactylon brevifolium Rydb. Bull Torrey Bot. = pungens 1878. Asteraceae. = Perityle tenella (Jones) Macbr. Club. 40: 474. 1913. Polemoniaceae. L. Co., Range, Purpus 6306, 1898 Washington Co., (Note: At Beaverdam in Ariz.) Palmer (Torr.) Nutt. Iron Juniper (US!). 199, 1877 (US!;NY!). Leptotaenia eatonii Coult. & Rose Rev. N. Amer. Laphamia palmeri Gray var. tenella Jones Proc. Ca- Umbell. 52. 1888. Apiaceae. = Lomatium disseetum var. lif. Acad. II, 5: 703. 1895. Asteraceae. = Perityle tenella eatonii (Coult. & Rose) Cronq. Utah, Eaton 147, 1869 (?). (Jones) Macbr. Washington Co., Springdale, Jones Lesquerella garrettii Payson Ann. Missouri Bot. Card. 52491, 1894 (US!;POM!;NY!;BRY!). 8; 213. 1921. Brassicaceae. Salt Lake Co., Big Cotton- Laphamia stansburyi Gray PI. Wright. 1: 101. 1852. wood Canyon, Garrett 1.344, 1908 (MO!;RM!). Asteraceae. = Perityle stansburyi (Gray) Macbr. Tooele Lesquerella gordonii var. sessilis Wats. Proc. Amer. Co., Stansbury Island, Stansbury sn, 1850 (NY!;BRY!). Acad. 23; 253. 1888. Brassicaceae. = L. tenella A. Nels. Lappula collina Greene Pittonia 4: 96. 1899. Boragi- Washington Co., near St. George, Parry sn, 1874 (GH). naceae. = L. oeeidentalis (Wats.) Greene sens. lat. Piute Lesquerella hemiphysaria Maguire var. hemiphysaria Co., Kingston, Jones sn, 1894 (:'). .\mer. Midi. Naturalist 27: 456. 1942. Brassicaceae. San- Lathyrus brachycalyx Rydb. var. brachycalyx Bull. pete Co., Wasatch Plateau, Maguire 20053, 1940 Torrev Bot. Club 34: 425. 1907. Fabaceae. Salt Lake (UTC!). Co., City Creek Canyon, Leonard 101, 1883 (NY!). Lesquerella hemiphysaria Maguire var. lucens Welsh Lathyrus coriaceus White Bull. Torrey Bot. Club 21: Reveal Great Basin Naturalist 37: .338. 1977. Bra.ssi- 452. 1894. Fabaceae. = L. lanzwertii var. lanzwertii & caceae. Carbon Co., Range Creek, Welsh & Taylor Wasatch Mts., Watson 297, 1869 (US!). 151.39, 1977 (BRY!). Lathyrus utahensis Jones Proc. Calif. Acad. II, 5: Lesquerella hitchcockii ssp. tumulosa Barneby Leafl. 678. 1895. Fabaceae. = L. puuciflorus var. utahensis W. Bot. 10: 313. 1966. Brassicaceae. = L. tumulosa (Jones) Peck Sevier Co., Irelands Ranch, Jones 54411, (Barnebv) Reveal Kane Co., se Cannonville, Barneby 1894 (US!;NY!). 14424, 1966(US!;BRY!;UTC!;NY!). Lathyrus zionis C.L. Hitchc. Univ. Washington Publ. Lesquerella multiceps Maguire Amer. Midi. Natural- Biol. 15: 36. 1952. Fabaceae. = L. braehyealyx var. zi- ist 27: 465. 1942. Brassicaceae. Cache Co., Tony Grove onii (C.L. Hitchc.) Welsh Kane Co., 10 mi e Zion N.P., Lake, Maguire 16030, 1938 (UTC!;RM!). Hitchcock 19013, 1949(CAS!,RM!;UTC!;NY!). Lesquerella rubicundula Rollins Contr. Dudley Herb. Lepidium brachybotryum Rydb. Bull. Torrey 3: 178. 1941. Brassicaceae. Garfield Co., Red Canyon, Bot.club 34: 427. 1907. Brassicaceae. = L. montanum Egglcston 8198, 1912 (US!). var. montanum Juab Co., Juab, Goodding 1075, 1902 Lesquerella subumbellata Rollins Rhodora 57: 255. (NY!;US!;RM!). 1955. Brassicaceae. Uintah Co., n Vernal, Rollins 17,5, Lepidium georginum Rydb. Bull. Torrey Bot. Club 30; 19.37 (US!;RM!;NY!;UTC!). 253. 1903. Brassicaceae. = L. lasiocarpum var. georgi- Lesquerella utahensis Rydb. Bull. Torrey Bot. Club num (Rydb.) C.L. Hitchc. Washington Co., Southern .30: 252. 1903. Brassicaceae. Utah Co., American Fork Utah, Parry 19, 1874 (NY!;ISC!). Canyon, Jones 1.354, 1880 (NY!;US!;CAS!). Lepidium jonesii Rydb. Bull. Torrey Bot. Club 29; Lesquerella wardii Wats. Proc. Amer. Acad. 23; 252. 233. 1902. Brassicaceae. = L. montanum var. jonesii 1888. Brassicaceae. Garfield Co., Aquarius Plateau, (Rydb.) C.L. Hitchc. Washington Co., St. George, Jones Ward 589, 1875 (GH;US!;RM!). 1636, 1880 (US!;CAS!;RM!;POM!;NY!). June 1982 Welsh: Utah Plant Types 175

Ligusticum brevilobum Rydb. Fl. Rocky Mts. 613, Lupinus huffmanii C.P. Sm. Sp. Lupinorum 693. 1064. 1917. Apiaceae. = L. porteri Coult. & Rose Gar- 1951. Fabaceae. = L. sericeus var. sericeus sens. lat. Se- field Co., Aquarius Plateau, Rydberg & Carlton 7473, vier Co., Salina Experiment Station, Huffman 47-7A, 1905 (NY!;US!). 1949 (CAS!). Ligusticum filicimim Wats. Proc. Anier. Acad. 11: Lupinus jonesii Rydb. Bull. Torrey Bot. Club 30: 256. 140. 1876. Apiaceae. Summit (?) Co., Uinta Mts., Wat- 1903. Fabaceae. Syn: L. leucanthus Rydb. Washington son 454, 1869 (US!;NY!). Co., Silver Reef, Jones 514.3, 1894 (US!;RM!;BRY!;NY!). Limnia utahensis Rydb. Bull. Torrey Bot. Club 39: Lupinus larsonanus C.P. Sm. Sp. Lupinorum 696. 314. 1912. Portulacaceae. = Montia perfoliata (Donn) 1951. Fabaceae. = L. sericeus var. sericeus sens. lat. Se- Howell Washington Co., St. George, Palmer 56, 1877 vier Co., inter Browns Hole & Hoodoo, Larson sn, 1934 (NY!). (CAS!). Linosyris serrulata Torr. in Stansb. Explor. Great Salt Lupinus leucanthus Rydb. Bull. Torrey Bot. Club .30; Lake 389. 1852. Asteraceae. = Chrijsothamnus vis- 259. 1903. Fabaceae. = L. jonesii Rydb. Washington cidiflorus var. viscidiflorus Salt Lake Co., Salt Lake Val- Co., Springdale, Jones .5249e, 1894 (US!;RM!; ley," Stansbury sn. 1850 (NY!). BRY!;POM!;NY!). Linum aristatum Engelm. in Wisliz. var. subteres Lupinus leucophyllus Dougl. ex Lindl. var. lupinus Trel. in Eastwood Proc. Calif. Acad. II, 6: 285. 1896. Li- Rydb. Bull. Torrey Bot. Club 40: 44. 1913. Fabaceae. naceae. = L. subteres (Trel.) Winkler San Juan Co., Syn: L. utahensis Moldenke = L. caudatus var. caud- Willow Creek, Eastwood sn, 1895 (?). atus San Juan Co., Bears Ears, Rydberg &c Garrett 9363, Linum kingii Wats. Rep. U.S. Geol. Explor. 40th Par- 1911 (NY!). allel, Bot. 5: 49. 1871. Linaceae. Summit Co., Uintas, Lupinus maculatus Rydb. Bull. Torrey Bot. Club 30: Watson 203, 1869 (US!). 257. 1903. Fabaceae. Utah Co., Pleasant Valley (P.V.) Linum kingii Wats. var. pinetorum Jones Proc. Calif. Jet., Jones sn, 1883 (NY!;US!;POM!;UTC!). Acad. II, 5: 628. 1895. Linaceae. = L. kingii Wats. Gar- Lupinus marianus Rydb. Bull. Torrey Bot. Club 34: field Co., canyon above Tropic, Jones 5306, 1894 41. 1907. Fabaceae. = L. sericeus var. marianus (Rydb.) (US!;F0M!;NY!;'BRY!). Welsh Piute Co., Bullion Creek, Rydberg & Carlon Lomatium jonesii Coult. & Rose Contr. U.S. Natl. 7024, 1905(US!;RM!;NY!). Herb. 7: 233. 1900. Apiaceae. = L. foeniculaceum var. Lupinus prunophilus Jones Contr. W. Bot. 13: 7. macdougalii (Coult. & Rose) Cronq. Sevier Co., Irelands 1910. Fabaceae. Juab Co., Robinson, Jones sn, 1909 Ranch, Jones 5435, 1894 (US!). (CAS!;POM!;BRY!;NY!). Lomatium junceum Barneby & Holmgren Brittonia Lupinus pulcher Eastw. Leafl. W. Bot. 3: 173. 1942. 31: 96. 1979. Apiaceae. Emery Co., San Rafael Swell, Fabaceae. = L. hillii Greene Iron Co., 18 mi s Cedar Holmgren et al. 8778, 1978 (NY!;BRY!;UTC!). City, Barkley & Reed 4069, 1939 (CAS!). Lonicera utahensis Wats. Rep. U.S. Geol. Explor. Lupinus puroviridis C.P. Sm. Sp. Lupinorum 694. 40th Parallel, Bot. 5: 133. 1871. Caprifoliaceae. Salt 1951. Fabaceae. = L. sericeus var. sericeus sens. lat. Se- Lake Co., Cottonwood Canyon, Watson 477, 1869 vier Co., Salina Experiment Station, Huffman 49-7B, (US!;NY!). 1949 (CAS!). Lotus longebracteatus Rydb. Bull. Torrey Bot. Club Lupinus quercus-jugi C.P. Sm. Sp. Lupinorum 696. .30: 254. 1903. Fabaceae. Syn: L. oroboides var. niimmii- 1951. Fabaceae. = L. sericeus var. sericeus sens. lat. Se- Jarius (Jones) Isely Washington Co., "South Utah" ("Mo- vier Co., Browns Hole, Clawson C241, 1935 (CAS!). kaik Pass"), Palmer 94, 1877 (US!;NY!;BRY!). Lupinus rickeri C.P. Sm. Sp. Lupinorum 695. 1951. Lotus utahensis Ottley Brittonia 5: 108. 1944. Fa- Fabaceae. = L. sericeus var. sericeus sens. lat. Sevier baceae. Kane Co., w Carmel, Ferguson & Ottley 5613 Co., Salina Experiment Station, Ricker 14080, 1917 (?) (CAS!). Lupinus aegraovium C.P. Sm. Sp. Lupinorum 694. Lupinus rubens Rydb. Bull. Torrey Bot. Club 34: 45. 1951. Fabaceae. = L. sericeus var. sericeus sens. lat. Se- 1907. Fabaceae. = L. pusiUus var. rubens (Rydb.) vier Co., Salina Experiment Station, Huffman 49-9, 1949 Welsh Washington Co., St. George, Parry 41, 1874 (CAS!). (NY!;ISC!). Lupinus aridus var. utahensis Wats. Proc. Amer. Lupinus salinensis C.P. Sm. Sp. Lupinorum 695. Acad. 8: 5.34. 1873. Fabaceae. = L. caespitosus Nutt.; 1951. Fabaceae. = L. sericeus var. sericeus sens. lat. Se- Svn; L. watsonii Heller Summit Co., Parleys Park, Wat- vier Co., Salina Experiment Station, Clawson C.202, son 2.34, 1869 (US!). 1935 (CAS!). Lupinus barbiger Wats. Proc. Amer. Acad. 8: .528. Lupinus sileri Wats. Proc. Amer. Acad. 10: 345. 1875. 1873. Fabaceae. = L. sericeus var. barbiger (Wats.) Fabaceae. = L. kingii Wats. Southern Utah, Siler sn Welsh Kane Co., S. Utah, Siler sn, 1872? (GH). (GH). Lupinus eatonanus C.P. Sm. Sp. Lupinorum 671. Lupinus spatulatus Rydb. Bull. Torrey Bot. Club 29: 1949. Fabaceae. = L. leucophyllus Dougl. Wasatch Co., 244. 1902. Fabaceae. = L. argenteus var. boreus (C.P. Hailstone, Smith .3995, 1925 (CAS!). Sm.) Welsh Wasatch Mts., Watson 225, 1869 (NY!). Lupinus garrettianus C.P. Sm. Sp. Lupinorum 672. Lupinus tooelensis C.P. Sm. Sp. Lupinorum 640. 1949. Fabaceae. = L. argenteus var. argenteus Duch- 1948. Fabaceae. = L. prunophilus Jones Tooele Co., esne Co., 1 mi w Duchesne, Garrett 8.303, 1940 (UT!). Deep Creek Mts., Cottam 7203, 1937 (UT!). Lupinus holosericeus var. utahensis Wats. Proc. Lupinus watsonii Heller Muhlenbergia 1: 114. 1905. Amer. Acad. 8: 5.33. 1873. Fabaceae. = L. caudatus var. Fabaceae. = L. caespitosus Nutt. Summit Co., Parleys caudatus Salt Lake Co. ?, Watson sn?, 1869 (US!). Park, Watson 234, 1869 (US!). 176 Great Basin Naturalist Vol. 42, No. 2

12: 247. 1877. Lychnis kingii Wats. Proc. Amer. Acad. Mammilluria chlorantha Engelm. in Wheeler Rep. Co., head Bear River, Watson Caryophyllaceae. Summit U.S. Geogr. Surv. W. 100th Meridian 6: 127. 1878. Cac- 153,1869 (US!;NY!). taceae. = Coryphantha vivipara var. deserti (Engelm.)

Lygodesmia entrada Welsh & Goodrich Great Basin W.T. Marshall Washington Co., e St. George, Parry sn, NaturaHst 40: 83. 1980. Asteraceae. Grand Co., Tusher 1874 (ISC!;BRY!). Canyon, Welsh & Welsh 16725, 1978 (BRY!;UT!;NY!). Mentzelia albicaulis Dougl. in Hook. var. integrifolia Lygodesmia grandiflora (Nutt.) T. & G. var. stricta Wats. Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: 114. Maguire Amer. Midi. Naturalist 37: 145. 1947. Aste- 1871. Loasaceae. = M. dispersa Wats. Davis Co., Ante- raceae. Carbon Co., 1 mi s Price, Maguire 18417, 1940 lope Island, Watson 4.30, 1869 (US!;NY!). (NY!;US!;UTC!). Mentzelia argillosa Darlington Ann. Missouri Bot. Lygodesmia juncea (Pursh) D. Don var. dianthopsis Card. 21: 153. 1934. Loasaceae. Sevier Co., Vermillion, D.C. Eaton Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: Jones 5631, 1894 (MO;POM!;US!). 200. 1871. Asteraceae. = L. dianthopsis (D.C. Eaton) Mentzelia multiflora (Nutt.) Gray var. integra Jones

Proc. Calif. Acad. Sci 11, 5: 689. 1895. Loasaceae. = Tomb Davis (?) Co., Great Salt Lake Islands, Watson M. 707, 1869 (US!). integra (Jones) Tidestr. Washington Co., Rockville, Jones Machaeranthera commixta Greene Pittonia 4: 71. 6082C, 1894 (US!). 1899. Asteraceae. Syn: M. leptophylla Rydb.? Garfield Mentzelia pterosperma Eastw. Proc. Calif. Acad. 11, 6: 290. 1896. Loasaceae. San Co., Willow Creek, East- Co., Bromide Pass, Jones 5695v, 1894 (US!). Juan 1895 (CAS!). Machaeranthera glabriuscula (Nutt.) Cronq. & Keck wood 31, Mertensia brevistyla Wats. Rep. U.S. Geol. Explor. var. confertiflora Cronq. Leafl. W. Bot. 10: 11. 1963. 40th Parallel, Bot. 5: 239. 1871. Boraginaceae. Summit Asteraceae. = Xylorhiza confertiflora (Cronq.) T.J. Wat- (?) Co., Wasatch Mts., Watson 845, 1869 (NY!). son Garfield Co., 11 mi ne Henrieville, Cronquist 9164, Mertensia leonardii Rydb. Bull. Torrey Bot. Club 36: 1961 (NY!;UTC!). 681. 1909. Boraginaceae. = M. arizonica Greene var. Machaeranthera kingii (D.C. Eaton) Cronq. & Keck leonardii (Rydb.) Johnst. Salt Lake Co., Mill Creek Can- var. bamebyana Welsh & Goodrich in Welsh Brittonia yon, Leonard sn, 1884 (NY!). .33: 299. 1981. Asteraceae. Millard Co., Canyon Mts., Mertensia paniculata var. nivalis Wats. Rep. U.S. Goodrich 14929, 1980 (BRY!;GH!;NY!;US!;RM!; Geol. Explor. 40th Parallel, Bot. 5: 239. 1871. Boragi- UC!;USFS!;UT!;UTC!). naceae. = M. viridis A. Nels. var. cana (Rydb.) L.O. Machaeranthera latifolia A. Nels. Proc. Biol. Soc. Williams Summit Co., Bear River Canon, Watson 844, .38. Asteraceae. Syn: rubr- Washington 20: 1907. M. 1869 (CAS!;US!;NY!). = icaulis Rydb.; Aster nihrotinctiis Blake M. bigelovii Mertensia praecox Smiley in Macbr. Contr. Gray ? Salt Canyon, Garrett Gray Lake Co., Big Cottonwood Herb. 11, 48: 10. 1916. Boraginaceae. = M. oblongifolia 1933, 1906(US!;RM!;UT!). var. nevadensis (A. Nels.) L.O. Williams Cache Co., Lo- Machaeranthera leptophylla Rydb. Bull. Torrey Bot. gan Canyon, Smith 2160, 1910 (UTC!). Club 37: 147. 1910 Asteraceae. = M. commixta Greene? Mertensia sampsonii Tidestr. Proc. Biol Soc. Wash- Cache Co., Logan, Rydberg sn, 1895 (NY!;BRY!). ington 26: 122. 1913. Boraginaceae. = M. arizonica var. Machaeranthera paniculata A. Nels. Proc. Biol. Soc. leonardii (Rydb.) Johnst. Sanpete Co., e of Ephraim, Washington 20: 38. 1907. Asteraceae. = M. bigelovii Sampson 677, (US!). Contr. Gray ? Salt Lake Co., Parleys Canyon, Garrett 2083, Mertensia toyabensis var. subnuda Macbr. 1906(US!;RM!;UT!). Gray Herb. II, 48: 7. 1916. Boraginaceae. = M. arizona var. (Macbr.) L.O. Williams Wayne Co. Machaeranthera pulverulenta var. vacans A. Nels. subnuda (?), Fish Lake Mountain. Ward 329, 1875 (GH). Bot. Gaz. 56: 70. 1913. Asteraceae. = M. canescens Micropuntia barkleyana Daston Amer. Midi. Natu- (Pursh) Gray? San Juan Co., Walker 360, 1912 ralist 36: 662. 1946. Cactaceae. = Opuntia pulchella (RM!;NY!). Engelm. Millard Co., Desert Experimental Range, Machaeranthera tortifolia (T. & G.) Cronq. & Keck Marsh sn, 1945 (F?). var. imberbis Cronq. Leafl. W. Bot. 10: 12. 1963. Aste- Micropuntia brachyrohopalica Daston Amer. Midi. raceae. = Xylorhiza tortifolia var. iinberbis (Cronq.) T.J. Naturalist 36: 661. 1946. Cactaceae. = Opuntia pul- Watson Grand Co., w Moab, Cronquist 8994, 1961 chella Engelm. Millard Co., Desert Experimental Range, (NY!;UTC!). Marsh sn, 1945 (F?). Macronema obovatum Rydb. Bull. Torrey Bot. Club Micropuntia spectatissima Daston Amer. Midi. Natu-

27: 618. 1900. Asteraceae. = Haplopappus rydbergii ralist 36: 661. 1946. Cactaceae. = Opuntia pulchella Blake Salt Lake Co., City Creek Canyon, Jones 1081, Engelm. Millard Co., Desert Experimental Range, 1895(US!;CAS!;RM!;POM!;NY!;UTC!). Marsh sn, 1946 (F?). Madronella oblongifolia Rydb. Bull. Torrey Bot. Club Mimulus eastwoodiae Rydb. Bull. Torrey Bot. Club 36: 686. 1909. Lamiaceae. = Monardella odoratissima 40: 483. 1913. Scrophulariaceae. San Juan Co., San Juan Benth. Juab Co., Mt. Nebo, Rydberg & Carlton 7706, River, near Bluff, Rydberg 9883, 1911 (NY!; 1905(NY!;US!;RM!). US!;RM!;UT!). Madronella sessilifolia Rydb. Bull. Torrey Bot. Club Mimulus glabratus H.B.K. ssp. utahensis Pennell 36: 685. 1909. Lamiaceae. = Monardella odoratissima Acad. Nat. Sci. Philadelphia Monogr. 1: 123. 1935. Scro-

Benth. Washington Co., St. George, Palmer .393, 1877 phulariaceae. = M. glabratus H.B.K. Millard Co., Preuss (NY!). Lake, Tidestrom 11180, 1919 (PH). June 1982 Welsh: Utah Plant Types 177

Mimulus parryi Gray Proc. Amer. Acad. 11: 97. 1876. Oenothera californica ssp. avita W. Klein Aliso 5: Scrophulariaceae. Washington Co., near St. George, Par- 179. 1962. Onagraceae. = O. avita (W. Klein) W. Klein ry 147, 1874 (NY!;US!;ISChNDG!). Washington Co., 0.5 ne of Leeds, Klein 1049, 1959 Mirabilis gltitinosa A. Nels. Proc. Biol. Soc. Washing- (RSA). ton 17: 92. 1904. Nyctaginaceae. = M. bigelovii Gray Oenothera gauraeflora var. hitchcockii H. Lev. Mon- var. retrorsa (Heller) Munz Washington Co., near St. ogr. Oenothera 226. 1905. Onagraceae. = Camissonia George, Goodding 778. 1902 (RM!). boothii ssp. ah/ssoides (H. & A.) Raven Simpsons Park, Mitella stenopetala Piper Erythea 7: 161. 1899. Sa- without coll., 1859 (MO). xifragaceae. = M. stauropetala Piper Salt Lake (?) Co., Oenothera hookeri T. & G. var. angustifolia Gates

Wasatch Mts., Watson 365, 1869 (?). Mut. Factors Evol. 10: 30. 1915. Onagraceae. Utah Co., Muhlenbergia ctirtifolia Scribn. Bull. Torrey Bot. Asphalt (s Thistle), Jones 5624, 1894 (US!;NY!). Club 38: 328. 1911. Poaceae. = M. thurberi Rydb. Kane Oenothera johnsonii Parry Amer. Nat. 9: 270. 1875. Co., inter Kanab & Carmel, Jones 6077j, 1894 (US!). Onagraceae. = O. primiveris Gray Washington Co., Najas flexilis ssp. caespitosus Maguire in Maguire & near St. George, Parry 64, 1874 (ISC!).

Jensen Rhodora 44: 7. 1942. Najadaceae. = N. caespi- Oenothera hngissima Rydb. Bull. Torrey Bot. Club tosus (Maguire) Reveal Sevier Co., Pelican Point, Fish 40: 65. 1913. Onagraceae. San Juan Co., Natural Bridges Lake, Maguire 19888, 1940 (UTC!;BRY!). Rydberg & Garrett 9410, 1911 (NY!;US!). Nasturtium obtusum Nutt. var. alpinum Wats. Rep. Oenothera multijuga Wats. Amer. Naturalist 7: 300. 1873. Onagraceae. = Camissonia multijuga (Wats.) Rav- U.S. Geol. Explor. 40th Parallel, Bot. 5: 15. 1871. Brassi- near sn, 1872 caceae. = Rorippa curvipes var. alpina (Wats.) Stuckey en Kane Co., Kanab, Thompson (NY?). Oenothera multijuga Wats. var. orientalis Summit Co., head of Bear River Canyon, Uinta Mts., Munz Amer. Bot. 15: 232. 1928. Onagraceae. = Camissonia Watson 60, 1869 (US!). J. walkeri ssp. walkeri Grand Co., Moab, Jones sn, 1913 Navarretia setosissitna T. & G. in Ives Rep. Colorado (POM!). Riv. W. 4: 22. 1860. Polemoniaceae. = Langloisia seto- Oenothera parryi Wats, ex Parry Amer. Naturalist 9: sissima (T. & G.) Greene Washington Co., Virgin River, 270. 1875. Onagraceae. = Camissonia parryi (Wats.) Fremont 414, (NY!). Raven Washington Co., near St. George, Parry 72, 1874 Nemophila breviflora Gray Proc. Amer. Acad. 10: (US!;NY!;BRY!;ISC!;NDG!). 315. 1875. Hydrophvllaceae. Summit Co., Parleys Park, Oenothera scapoidea Nutt. in T. & G. ssp. utahensis Watson 169, 1869 (NY!;US!). Raven Univ. Calif. Publ. Bot. 34: 96. 1962. Onagraceae. NothoJaena parryi D.C. Eaton Amer. Naturalist 9: = Camissonia scapoidea ssp. utahensis (Raven) Raven 351. 1875. Polvpodiaceae. Washington Co., St. George, Salt Lake Co., Black Rock, Watson 414, 1869 (US!). Parry 263, 1874 (US!;ISC!;BRY!). Oenothera tenuissima Jones Proc. Calif. Acad. II, 5: Nuttallia lobata Rydb. Bull. Torrey Bot. Club 40: 61. 683. 1895. Onagraceae. = Camissonia parryi (Wats.) 1913. Loasaceae. = Mentzelia Integra (Jones) Tidestr. Raven Washington Co., Rockville, Jones 6083, 1894 Washington Co., near St. George, Palmer 172, 1877 (POM!;US!;RM!;NY!). (NY!). Oenothera triloba var. ecristata Jones Proc. Calif. Oenothera acutissima W.L. Wagner Systematic Bot- Acad. II, 5: 681. 1895. Onagraceae. = O. flava (A. Nels.) any 6: 153. 1981. Onagraceae. Daggett Co., Greendale Garrett Garfield Co., Panguitch Lake, Jones 60155, 1894 Peterson 1978 (BRY!). Campground, Neese & 5428, (POM!). Oenothera albicaulis Pursh var. decumbens Wats, ex Opuntia barhata M. Brandegee ex J. A. Purpus Mon- Parry Amer. Naturalist 9: 270. 1875. Onagraceae. = O. ats. Kakteenk. 10: 97. 1900. Cactaceae. = O. poly- deltoides ssp. ambigua (Wats.) W. Klein Washington acantha (?) Haw. San Juan (?) Co., La Sal Mts., Purpus Co., near St. George, Parry 63, 1874 (GH?). sn, 1897 (?). Oenothera alyssoides H. & A. var. minutiflora Wats. Opuntia barbata M. Brandegee ex J.A. Purpus var. Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: 111. 1871. gracillima M. Brandegee ex J.A. Purpus Monats. Kak- Onagraceae. = Camissonia minor (A. Nels.) Raven teenk. 10: 110, 120. 1900. Cactaceae. = O. polyacantha Tooele Co.. Stansbury Island, Watson 421, 1869 (US!). (?) Haw. San Juan (?) Co., La Sal Mts., Purpus sn, 1897 Oenothera alyssoides Pursh var. villosa Wats. Proc. (?)• Amer. Acad. 8: 591. 1873. Onagraceae. = Camissonia Opuntia basilaris Engelm. & BigeL var. woodburyi boothii ssp. alyssoides (H. & A.) Raven Salt Lake Co., (?) Earl Sahuaroland Bull. 34: 15. 1980. Cactaceae. Wash- near Salt Lake, Stansbury sn, 1850 (NY!). ington Co., Fort Pierce Wash, Woodbury 2060a, 1977 Oenothera Wats. Proc. Acad. 14: 293. ambigua Amer. (BRY!). 1879. Onagraceae. = O. deltoides var. ambigua (Wats.) Opuntia palmeri Engelm. Contr. U.S. Natl. Herb. 3: Munz Washington Co., near St. George, Palmer 162, 423. 1896. Cactaceae. = ? O. phaeacantha var. discata 1877 (US!;ISC!). (Griffiths) Benson & Walkington Washington Co., near var. in Oenothera brevipes parviflora Wats, Parry St. George, Palmer sn, 1877 (MO). Amer. Naturalist 9: 271. 1875. Onagraceae. = Cam- Opuntia rubrifolia Engelm. Contr. U.S. Natl. Herb. issonia multijuga (Wats.) Raven Washington Co., near 3: 424. 1896. Cactaceae. = O. erinacea var. ursina Par- St. George, Parry 74, 1874 (GH;MO;F). ish Washington Co., near St. George, Palmer 3, 1877 Oenothera caespitosa Nutt. var. jonesii Munz Amer. (MO). 18: J. Bot. 731. 1931. Onagraceae. Syn: O. caespitosa ssp. Opuntia utahensis J.A. Purpus Monats. Kakteenk. 19: jonesii (Munz) Munz Tooele Co., Fish Springs, Jones sn, 133. 1909. Cactaceae. = O. macrorhiza Engelm. San

1891 (POM!). Juan (?) Co., La Sal Mts., Purpus sn, 1897 (US!). 178 Great Basin Naturalist Vol. 42, No. 2

Oreocarya breviflora Osterh. in Payson Univ. Wyom- Oxytropis oreophila Gray var. juniperina Welsh ing Publ. Bot. 1: 169. 1926. Boraginaceae. = Crijptantha Great Basin Naturalist 38: 339. 1978. Fabaceae. Wayne breviflora (Osterh.) Payson Uintah Co., n Jensen, Os- Co., 1 mi e Bicknell, Welsh & Moore 13828, 1976 terhout 6414, 1925 (RM!;BRY!). (BRY!;UT!;ISC!). Oreocarya Macbr. commixta Contr. Gray Herb. II, Pachylophus crinitis Rydb. Fl. Rocky Mts. 598, 1064. 48: 33. = 1916. Boraginaceae. Cryptantha humilis var. 1917. Onagraceae. = Oenothera caespitosa var. crinita commixta (Macbr.) Higgins Juab Co., Juab, Goodding (Rydb.) Munz Wayne Co., Rabbit Valley, Ward 526, 1074, 1902 (GH;RM!). 1875 (US!). Oreocarya disticha Eastw. Bull. Torrey Bot. Club 30: Paronychia pulvinata Gray var. longiaristata 258. 1903. Boraginaceae. Syn: Cryptantha jamesii var. Chaudri Revis. Paronychia 194. 1968. Caryophyllaceae. disticha (Eastw.) Payson = C. cinerea (Greene) Cronq. Duchesne Co., Mt. Emmons, Hermann 5041, 1933 San Juan Co., Bartons range, Eastwood sn, 1895 (BRY!). (CAS!;NY!). Parrya platycarpa Rydb. Bull. Torrey Bot. Club 39:

Oreocarya dolosa Macbr. Contr. Gray Herb. II, 48: 326. 1912. Brassicaceae. = P. rydbergii Botsch. Summit

32. 1916. Boraginaceae. = Cryptantha humilis var. (?) Co., Uinta Mts, Watson 54, 1869 (US!;NY!). shantzii (Tidestr.) Higgins Cache Co., Logan, Smith Parthenium alpinum (Nutt.) T. & G. var. ligulatum 1605, 1909 (RM!). Jones Contr. W. Bot. 13: 16. 1910. Asteraceae. Syn: Oreocarya jonesiana Payson Univ. Wyoming Publ. Bolophyta ligulata (Jones) W.A. Weber = P. ligulatum

Bot. 1: 168. 1926. Boraginaceae. = Cryptantha jone- (Jones) Barneby Duchesne Co., Theodore, Jones sn, 1908 siana (Payson) Payson Emery Co., San Rafael Swell, (POM!;US!;CAS!;BRY!). Pediocactus Jones sn, 1914 (RM!;POM!;BRY!). despainii Welsh & Goodrich Great Basin Naturalist 40: 83. 1980. Cactaceae. Emery Co., Ra- Oreocarya pustulosa Rydb. Bull. Torrev Bot. Club San fael Swell, Despain 266a, 1978 (BRY!;NY!)' 40: 480. 1913. Boraginaceae. = Cryptantha pustulosa Pediocactus hermannii W.T. Marshall Saguaroland (Rydb.) Payson Syn: C. jamesii var. pustulosa (Rydb.) Bull. 8: 78. 1954. Cactaceae. = P. simpsonii (Engelm.) Harrington San Juan Co., Elk Mts., Rydberg & Garrett Britt. & Rose Garfield Co., Hatch, Hermann & Hermann 9320, 1911 (NY!). sn, 1953 (BRY!). Oreocarya rugulosa Payson Univ. Wyoming Publ. Pediocactus winkleri Heil Cact. &: Succ. (U.S.) J. 51: Bot. 1: 166. 1926. Boraginaceae. = Cryptantha rugulosa 28. 1979. Cactaceae. Wayne Co., Winkler sn, (UNM). (Payson) Payson Tooele Co., Fish Springs, Jones sn, 1891 Penstemon abietinus Pennell Contr. U.S. Natl. Herb. (GH;RM!;POM!). 20: 376. 1920. Scrophulariaceae. Sevier Co., Irelands Oreocarya shantzii Tidestr. Proc. Biol. Soc. Washing- Ranch, Jones 5440, 1894 (US!;RM!;BRY!;NY!). ton 26: 122. 1913. Boraginaceae. = Cryptantha humilis Penstemon acuminatus Dougl. var. congestus Jones var. shantzii (Tidestr.) Higgins Salt Lake Co., s Great Proc. Calif. Acad. II, 5: 714. 1895. Scrophulariaceae. Salt Lake, Kearney & Shantz 3098, 1912 (US!). Syn: P. pachyphyllus ssp. congestus (Jones) Keck; do var. Oreocarya tenuis Eastw. Bull. Torrey Bot. Club 30: congestus (Jones) N. Holmgr. = P. congestus (Jones) 244. 1903. Boraginaceae. = Cryptantha tenuis (Eastw.) Pennell Washington Co., near Cannan Ranch, Rockwell, Payson Grand Co., Courthouse Wash, Eastwood sn, Jones 5262, 1894 (POM!;US!). 1892(CAS!;RM!;NY!). Penstemon angustifolius Nutt. var. vernalensis N. Holmgren Brittonia 31: 229. Oreocarya torva A. Nels. Amer. Bot. 23: 1978. Scrophulariaceae. J. 269. 1936. Uintah Co., n Maeser, Holmgren et al. 8748, 1979 Boraginaceae. = Cryptantha flava (A. Nels.) Payson (NY!;BRY!;UTC!). Carbon Co., Price, Flowers 6395, 1933 (UTl). Penstemon atwoodii Welsh Great Basin Naturalist 35: Oreocarya wetherillii Eastw. Bull. Torrey Bot. Club 378. 1976. Scrophulariaceae. Kane Co., sse Canaan Peak, 30: 242. 1903. Boraginaceae. = Cryptantha wetherillii Welsh & Welsh 12821, 1975 (BRY!;UT!;NY!;UTC!). (Eastw.) Payson Grand Co., Courthouse Wash, East- Penstemon azureus var. ambiguus Gray Syn. Fl. N. wood sn, 1892 (CAS!;RM;NY!). Amer. 2: 272. 1886. Scrophulariaceae. = P. sepalulus A. Oreocarya iLnlliamsii A. Nels. Amer. Bot. 21: 578. J. Nels. Utah Co., near Provo, Watson 786, 1869 (US!;NY!). 1934. Boraginaceae. = Cryptantha stricta (Osterh.) Pay- Penstemon bracteatus Keck Leafl. W. Bot. 1: 82. son Daggett Co., Flaming Gorge, Williams 489, 1932 1934. Scrophulariaceae. Garfield Co., Red Canyon, East- (RM!;NY!). wood & Howell 783, 1933 (CAS!). Orogenia linearifolia Wats. Rep. U.S. Geol. Explor. Penstemon caespitosus Nutt. e.\ Gray ssp. perbrevis 40th Parallel, Bot. 5: 120. 1871. Apiaceae. Summit Co., Pennell Contr. U.S. Natl. Herb. 20: 375. 1920. Scro- Wasatch Mts., north of Parleys Park, Watson 440, 1869 phulariaceae. Carbon Co., near Castle Gate, Pennell (US!;NY!). 6138, 1915 (NY!:US!). Oxybaphus angustifolia var. viscidus Eastw. Proc. Penstemon caespitosus Nutt. ex Gray var. suffruti- Calif. Acad. II, 6: 313. 1896. Nyctaginaceae. = O. lin- cosus Gray Syn. Fl. N. Amer. 2: 270. 1878. Scrophula- earis (Pursh) Robins. San Juan Co., Butler Spring, East- riaceae. = P. tiisharensis N. Holmgren Beaver Co., near wood 101, 1895 (CAS!). Beaver, Palmer sn, 1877 (ISC!). Oxybaphus glaber Wats. Amer. Naturalist 7: 301. Penstemon carnosus Pennell in Graham Ann. Car- 1873. Nyctaginaceae. Kane Co., Kanab, Thompson sn, negie Mus. 25: 329. 1937. Scrophulariaceae. Emery Co., 1872 (US!). San Rafael Swell, Cottam 5229, 1935 (UT!;NY!;UTC!). Oxytropis jonesii Barneby Proc. Calif. Acad. IV, 27: Penstemon comarrhenus Gray Proc. Amer. Acad. 12:

215. 1952. Fabaceae. Garfield Co., Red Canyon, Ripley 81. 1876. Scrophulariaceae. Garfield (?) Co., Aquarius & Barneby 8550, 1947 (CAS!). Plateau, Ward 462, 1875 (NY!;US!). June 1982 Welsh: Utah Plant Types 179

Penstemon coticinnus Keck Amer. Midi. Naturalist Penstemon laevis Pennell Contr. U.S. Natl. Herb. 20: 23: 608. 1940. Scrophulariaceae. Millard Co., Tunnel .347. 1920. Scrophulariaceae. Washington Co., Spring- Springs Range, Cottam 5634, 1933 (CAS!;US!;UT!;NY!). dale, Jones 5250, 1894 (US!;POM!). Penstemon confertus var. aberrans Jones Proc. Calif. Penstemon leiophyllus Pennell Contr. U.S. Natl. Acad. II, 5: 715. 1895. Scrophulariaceae. = P. procerus Herb. 20: 346. 1920. Scrophulariaceae. Garfield Co., var. aberrans (Jones) A. Nels. Wasatch Co., Soldier Sum- Mammoth Creek, Jones 6026b, 1894 (US!). mit, Jones 5601i, 1894 (POM!;US!). Penstemon lentus Pennell ssp. albiflorus Keck Amer. Penstemon confusus Jones Zoe 4: 280. 1893. Scro- Midi. Naturalist 23: 616. 1940. Scrophulariaceae. San phulariaceae. Juab Co., Detroit, Jones sn, 1891 Juan Co., 8 mi w Blanding, Porter 1801, 19.39 (POM!;CAS!;BRY!). (RM!;NY!;UTC!). Penstemon crandallii A. Nels. ssp. atratus Keck Bull. Penstemon leonardii Rydb. Bull. Torrey Bot. Club 40: Torrey Bot. Club 64: 370. 1937. Scrophulariaceae. = F. 483. 1913. Scrophulariaceae. Wasatch Mts., Leonard sn, crandallii var. atratus (Keck) N. Holmgren San Juan (?) 1884 (NY!). Co., La Sal Mts., Jones sn, 1914 (CAS!;US!; Penstemon leptanthus Pennell Contr. U.S. Natl. POM!;NY!;BRY!). Herb. 20: 339. 1920. Scrophulariaceae. Sanpete Co., Penstemon cyananthus Hook. ssp. compactus Keck Twelve Mile Creek, Ward 280, 1875 (US!). Amer. Midi. Naturalist 23: 615. 1940. Scrophulariaceae. Penstemon linarioides Gray in Torr. var. sileri Gray = P. compactus (Keck) Crosswhite Cache Co., Mt. Syn. Fl. N. Amer. 2(1): 270. 1878. Scrophulariaceae.

Naomi, Maguire 16148, 1938 (UTC!). Kane (?) Co., S. Utali, Siler sn, 1873 (NY!). Penstemon cyananthus Hook. ssp. longiflorus Pennell Penstemon mucronatus N. Holmgren Brittonia 31:

Contr. U.S. Natl. Herb. 20: 353. 1920. Scrophulariaceae. 234. 1979. Scrophulariaceae. Daggett Co., s. Manila,

= P. longiflorus (Pennell) S. Clark Beaver Co., near Bea- Holmgren et al. 8447, 1978 (NY!;BRY!;UTC!). ver, Palmer 376, 1877 (US!;NY!). Penstemon nanus Keck Amer. Midi. Naturalist 23: Penstemon dolius Jones ex Pennell Contr. U.S. Natl. 607. 1940. Scrophulariaceae. Millard Co., Desert Experi- Herb. 20: 341. 1920. Scrophulariaceae. Tooele Co., Deep mental Range, Plummer 7313, 1939 (CAS!;BRY!; Creek Mts., Jones sn, 1891 (POM!;BRY!). UT!;NY!;UTC!). Penstemon dolius Jones var. duchesnensis N. Hol- Penstemon navajoa N. Holmgren Brittonia 30: 419. mgren Brittonia 31: 219. 1979. Scrophulariaceae. Duch- 1978. Scrophulariaceae. San Juan Co., Navajo Mt., N. & esne Co., e Duchesne, Holmgren et al, 8762, 1978 P Holmgren 8587, 1977 (NY!;BRY!;UTC!). (NY!;BRY!;UTC!). Penstemon obtusifolius Pennell Contr. U.S. Natl. Penstemon eatonii Gray Proc. Amer. Acad. 8: 395. Herb. 20: 370. 1920. Scrophulariaceae. = P. humilis var. 1872. Scrophulariaceae. Utah Co., Provo Canyon, Wat- obtusifolius (Pennell) Reveal Washington Co., Spring- son 776, 1869 (US!;NY!). dale, Jones 5249am, 1894 (US!;POM!;NY!). Penstemon eatonii Gray var. undosus Jones Proc. Ca- Penstemon ophianthus Pennell Contr. U.S. Natl. lif. Acad. II, 5: 715. 1895. Scrophulariaceae. Syn: P. ea- Herb. 20: 343. 1920. Scrophulariaceae. Syn: P. jamesii tonii ssp. undosus (Jones) Keck Washington Co., St. ssp. ophianthus (Pennell) Keck Wayne Co., Thurber, George, Jones 51101h, 1894 (POM!;US!;NY!). Jones 5708, 1894 (US!;POM!;NY!). Penstemon garrettii Pennell Contr. U.S. Natl. Herb. Penstemon palmeri Gray ssp. eglandulosus Keck

20: .3.53. 1920. Scrophulariaceae. = P. scariosus Pennell Amer. Midi. Naturalist 18: 797. 19.37. Scrophulariaceae. = sens. lat. Wasatch Co., near Midway, Carlton & Garrett P. palmeri var. eglandulosus (Keck) N. Holmgren 6697, 1905 (NY!;US!). Kane Co., 2.5 mi n Kanab, Maguire et al. 12279, 1935 Penstemon glaber var. utahensis Wats. Rep. U.S. (RM!;NY!;UTC!). Geol. Explor. 40th Parallel, Bot. 5: 217. 1871. Scrophula- Penstemon parvus Pennell Contr. U.S. Natl. Herb. 20:

riaceae. = P. subglaber Rydb. Summit Co., Uinta Mts, 345. 1920. Scrophulariaceae. Garfield Co., Aquarius Watson 771, 1869 (US!;NY!). Plateau, Ward 546, 1875 (US!). Penstemon goodrichii N. Holmgren Brittonia 30: 416. Penstemon patricus N. Holmgren Brittonia 31: 238. 1978. Scrophulariaceae. Uintah Co., e Lapoint, Hol- 1979. Scrophulariaceae. Juab Co., Thorns Creek Canyon,

mgren et al 8760, 1978 (NY!;BRY!;UTC!). Holmgren et al. 9018, 1978 (NY!;BRY!;UT!;UTC!). Penstemon grahamii Keck in Graham Ann. Carnegie Penstemon phlogifolius Greene Leaf!. Bot. Obs. & Mus. 26: 331. 1937. Scrophulariaceae. Uintah Co., Sand Grit. 1: 164. 1906. Scrophulariaceae. = P. watsonii Gray Wash, Graham 7883, 1933 (CM!). Carbon Co., Castle Gate, Jones 5486s, 1894 (US!;POM!).' Penstemon heterophyllus var. latifolius Wats. Rep. Penstemon platijphyllus Rydb. Bull. Torrey Bot. Club U.S. Geol. Explor. 40th Parallel, Bot. 5: 222. 1871. Scro- 36: 690. 1909. Scrophulariaceae. Syn: P. heterophyllus

phulariaceae. = P. platijphyllus Rydb. Salt Lake Co., var. latifolius Wats. Salt Lake Co., Cottonwood Canyon, Cottonwood Canyon, Watson 787, 1869 (US!). Watson 787, 1869 (US!). Penstemon humilis Nutt. in Gray var. brevifolius Penstemon pseudohumilis Jones Contr. W. Bot. 12: Gray Syn. Fl. N. Amer. 2(1): 267. 1878. Scrophula- 65. 1908. Scrophulariaceae. = P. marcusii (Keck) N. riaceae. Salt Lake Co., Cottonwood Canyon, Watson Holmgren? Carbon Co., Price, Jones sn, 1898 (POM!). 781, 1869 (US!;NY!). Penstemon pumilus var. thompsoniae Gray Syn. Fl. Penstemon jonesii Pennell (hybrid) Contr. U.S. Natl. N. Amer. 2(1): 269. 1878. Scrophulariaceae. Syn: P. Herb. 20: 338. 1920. Scrophulariaceae. = P. eatonii x P. caespitosus var. thompsoniae (Gray) A. Nels. = P. leiophyllus Washington Co., Springdale, Jones 5250, thompsoniae (Gray) Rydb. Kane Co., Kanab, Thompson 1894 (US!). sn, 1872 (GH). 180 Great Basin Naturalist Vol. 42, No. 2

Penstemon scariosus Pennell Contr. U.S. Natl. Herb. Phacelia demissa Gray var. heterotricha J.T. Howell 20: 353. 1920. Scrophulariaceae. Syn: P. garrettii Pennell Amer. Midi. Naturalist 29(1): 8. 1943. Hydrophyllaceae. Sanpete Co., Musinia Peak, Tidestrom (US!). 568, 1907 Piute Co., Marysvale, Jones 5388o, 1894 (POM!;US!). Penstemon tidestromii Pennell Contr. U.S. Natl. Phacelia foetida Goodding Bot. Gaz. 37: 58. 1904. Herb. 20: 379. 1920. Scrophulariaceae. Sanpete Co., San Hydrophyllaceae. = P. palmeri Torr. Washington Co., Pitch Mts., Tidestrom 1296, 1908 (US!). Diamond Valley, Goodding 833, 1902 (RM!). Penstemon uintahensis Pennell Contr. U.S. Natl. Phacelia howelliana Atwood Rhodora 74: 456. 1972. Herb. 20: ,350. 1920. Scrophulariaceae. Uintah Co., Dyer Hydrophyllaceae. San Juan Co., Bluff, Atwood 2454, Mine, Goodding 1221, 1902 (US!;RM!;BRY!;NY!). 1970 (BRY!;RM!;NY!;US!;UTC!). Penstemon utahensis Eastw. Zoe 4: 124. 1893. Scro- Phacelia incana Brand Beit. z. Kenntnis der Hydro- phulariaceae. San Juan Co., n Monticello, Eastwood sn, phyll. 8. 1911. Hydrophyllaceae. Tooele Co., Dugway, 1892 (CAS!;US!). Jones sn, 1891 (POM!;RM!;NY!). Penstemon wardii Gray Proc. Amer. Acad. 12: 82. Phacelia indecora J.T. Howell Amer. Midi. Naturalist 1876. Scrophulariaceae. Sevier Co., near Glenwood, 29: 12. 1943. Hydrophyllaceae. San Juan Co., Bluff, Ward 162, 1875 (US!;NY!). Jones sn, 1919 (CAS!). Petalonyx parryi Gray Proc. Amer. Acad. 10: 77. Phacelia mammillariensis Atwood Phytologia 26: 437. 1874. Loasaceae. Washington Co., St. George ("within a 1973. Hydrophyllaceae. Kane Co., e Glen Canyon City, stones throw of the great Mormon temple"). Parry 75, Welsh & Atwood 9809, 1970 (BRY!). 1874 (NY!;BRY!;ISC!). Phacelia nudicaulis Eastw. Zoe 4: 123. 1893. Hydro- Petalostemon flavescens Wats. Amer. Naturalist 7: phyllaceae. = P. demissa Gray var. demissa Grand Co., 299. 1873. Fabaceae. = Dalea flavescens (Wats.) Welsh near Moab, Eastwood sn, 1892 (CAS!;POM!;US!). Kane Co., "Kanab," Thompson sn, 1871 (US!). Phacelia orbicularis Rydb. Bull. Torrey Bot. Club 40: Peteria thompsonae Wats. Amer. Naturalist 7: 300. 479. 1913. Hydrophyllaceae. = P. corrtigata A. Nels. 1873. Fabaceae. Kane Co., Kanab, Thompson sn, 1872 (GH;US!;NY!;BRY!). Wayne Co., Marvine Laccolite, Jones 5663, 1894 (US!;NY!). Peucedanum bicolor Wats. Rep. U.S. Geol. Explor. Phacelia palmeri Torr. in Wats. 40th Parallel, Bot. 5: 129. 1871. Apiaceae. = Lornatium Rep. U.S. Geol. Ex- bicolor (Wats.) Coult. & Rose Summit Co., Parleys Park, plor. 40th Parallel, Bot. 5: 251. 1871. Hydrophyllaceae. Watson 467, 1869 (US!;NY!). Syn: P. foetida Goodding Washington Co., near St. Peucedanum graveolens Wats. Rep. U.S. Geol. Ex- George, Palmer 4, 1870 (NY!;US!). plor. 40th Parallel, Bot. 5: 128. 1871. Apiaceae. = Lo- Phacelia pinetorum Jones Zoe 4: 279. 1893. Hydro- rnatium nuttallii (Gray) Macbr. Wasatch Mts., Watson phyllaceae. = Eucrypta micrantha Torr. Tooele or Juab 463, 1869 (US!;NY!). Co., Deep Creek Mts., Jones sn, 1891 (POM!;CAS!).

Peucedanum juniperinum Jones Contr. W. Bot. 8: 29. Phacelia pulchella Gray Proc. Amer. Acad. 10: 326. 1898. Apiaceae. = Lomatiiim juniperinum (Jones) Coult. 1875. Hydrophyllaceae. Washington Co., near St. & Rose Summit Co., Coalville, Jones sn, 1889 George, Parry 182, 1874 (GH;POM!;NY!;US!;BRY!). (POM!;US!;BRY!). Phacelia pulchella Gray var. sabulonum J.T.Howell Peucedanum lapidosum Jones Zoe 2: 246. 1891. Amer. Midi. Naturalist 29: 12. 1943. Hydrophyllaceae. Apiaceae. = Cymopterus longipes Wats. Summit Co., = P. pulchella Gray Kane Co., Kaiparowits Plateau, Echo, Jones sn, 1890 (US!;RM!;POM!). Tompkins sn, 1939 (CAS!). Peucedanum millefolium Wats. Rep. U.S. Explor. Phacelia rafaelensis Atwood Rhodora 74: 454. 1972. 40th Parallel, Bot. 5: 129. 1871. Apiaceae. = Lornatium Hydrophyllaceae. Wayne Co., Capitol Reef National grayi Coult. & Rose Davis Co., Antelope Island, Watson Monument, Atwood & Higgins 1834, 1969 (BRY!; 466, 1869 (NY!;US!). CAS!;RM!;NY!;US!;UTC!). Peucedanum parryi Wats. Proc. Amer. Acad. 11: 143. Phacelia rotundifolia Torr. ex Wats. Rep. U.S. Geol. 1876. Apiaceae. = Lomatium parryi (Wats.) Macbr. Explor. 40th Parallel, Bot. 5: 253. 1871. Hydro- Washington (?) Co., Southern Utah, Parrv 85, 1874 phyllaceae. Southern Utah, Palmer sn, 1870 (NY!;US!). (NY!). Phacelia utahensis Voss Bull. Torrey Bot. Club 64: Peucedanum triternatum var. platycarpum Torr. in 135. 1937. Hydrophyllaceae. Sanpete Co., Gunnison, Standb. Explor. Great Salt Lake 389. 1852. Apiaceae. = Jones sn, 1910 (POM!). Lomatium triternatum var. platycarpum (Torr.) Cronq. Phlox austromontana Gov. Contr. U.S. Natl. Herb. 4: Great Salt Lake, Stansbury sn, 18.50 (NY!). 151. 1893. Polemoniaceae. Washington Co., Beaverdam Phaca mollissima var. utahensis Torr. in Stansb. Ex- Mts., Bailey 1944, 1891 (US!). plor. Great Salt Lake 385. 1852. Fabaceae. = Astragalus Phlox austromontana Gov. var. prostrata E. Nels. utahensis (Torr.) T. & G. Tooele Co., w shore Stansbury Rev. W. N. Amer. Phloxes 19. 1899. Polemoniaceae. Island, Great Salt Lake, Stansury sn, 1850 (NY!). Syn: P. jonesii Wherry Washington Co., Silver Reef, Phaca pardalina Rydb. N. Amer. Fl. 24: 352. 1929. Jones 5163y-z, 1894 (NY!;US!;RM!;POM!;BRY!). Fabaceae. = Astragalus pardalinus (Rydb.) Barneby Phlox caesia Eastw. Leafl. Bot. 2: 54. 1937. Emery Co., Cedar Mt., Jones sn, 1915 (NY!;POM!). W. Po- lemoniaceae. = Phacelia argillacea Atwood Phytologia 26: 437. 1973. P. gladiformis (Jones) E. Nels. Garfield Hydrophyllaceae. Utah Co., Spanish Fork Canyon, At- Co., Red Canyon, Eastwood & Howell 752, 1933 (CAS!). wood et al. 3091, 1971 (BRY!;UT!;NY!;US!;UTC!). Phlox canescens T. & G. in Beckwith Rep. U.S. Ex- Phacelia cephalotes Gray Proc. Amer. Acad. 10: 325. plor. & Surv. R.R. Pacific 2: 122. 1857. Polemoniaceae. 1875. Hydrophyllaceae. Washington Co., near St. = P. hoodii var. canescens (T. & G.) Peck Tooele Co., s George, Parry 179, 1874 (NY!;US!;BRY!;ISC!). Great Salt Lake, Beckwith 4, 1854 (NY!). June 1982 Welsh: Utah Plant Types 181

Phlox cluteana A. Nels. Amer. Botanist 28: 24. 1922. Poa festucoides Jones Proc. Calif. Acad. II, 5: 723. San Co., Navajo Mt., Clute 18, Polemoniaceae. Juan 1895. Poaceae. = Festuca thurberi Vasey Garfield Co., 1919(?). Mt. Ellen, Jones 5671, 1894 (POM!;US!). Phhx densa Brand Pflanzenr. 4. Fam. 250. 83. 1907. Poa longiligula Scribn. & Will. USDA Div. Agrostol. Polemoniaceae. = P. austromontana Gov. Beaver Co., Circ. 9: 3. 1899. Poaceae. = P. fendleriana (Steud.) Frisco, Jones 2021, 1880 (US!;POM!;NY!;UTC!). Vasey sens. lat. Washington Co., Silver Reef, Jones Phlox grahamii Wherry Brittonia 5: 63. 1943. Polemo- 5149at, 1894 (RM!;NY!;US!). niaceae. = P. longifolia Nutt. Uintah Co., Sand Wash, Poa scabriuscula Williams USDA Div. Agrostol. Circ. Graham 7884, 1933 (CM!). 10: 4. 1899. Poaceae. = P. fendleriana (Steud.) Vasey Phlox jonesii Wherry Notul. Nat. Acad. Nat. Sci. Sevier Co., s Glenwood, Ward 136, 1895 (US!). Philadelphia 146: 8. 1944. Polemoniaceae. = P. austro- Polemonium albiflorum Eastw. Bot. Gaz. 37: 437. montana var. prostrata E. Nels. Washington Co., Zion 1904. Polemoniaceae. = P. foliosissimum var. alpinum Canyon, Jones sn, 1923 (US!;CAS!;POM!). Brand. Carbon Co., Scofield, Harkness sn, 1902 Phlox longifolia Nutt. var. gladiformis Jones Proc. (CAS!;RM!). Calif. Acad. II, 5: 711. 1895. Polemoniaceae. = P. gladi- Polemonium foliosissimum Gray ssp. albiflorum var. formis (Jones) E. Nels. Iron Co., Cedar Canyon, Jones alpinuum Brand Pflanzenr. 4. Fam. 250: 34. 1907. Po- 5208c, 1894 (POM!;US!). lemoniaceae. Syn: P. albiflorum Eastw.; P. foliosissimum Physaria acutifolia Rydb. var. purpurea Welsh & Re- ssp. albiflorum (Eastw.) Brand Salt Lake Co., Alta, Jones veal Great Basin Naturalist 37: 345. 1977. Brassicaceae. 1114, 1879 (NY!;POM!;UT!). Grand Co., near Sego, Welsh 6902, 1968 (BRY!). Gray Proc. Amer. Acad. 11: Physaria chambersii Rollins var. membranacea Roll- Polygala acanthoclada 73. 1876. Polygalaceae. Syn: P. acanthoclada var. in- ins Rhodora 41: 404. 1939. Brassicaceae. = F. lepidota R., Brandegee sn, var. membranacea (Rollins) Rollins Garfield Co., Red tricata Eastw. San Juan Co., San Juan Canyon, Rollins & Chambers 2448, 1938 (GH; 1873 (NY!). RM!;US!;NY!). Polygala acanthoclada Gray var. intricata Eastw. Physaria grahamii Morton in Graham Ann. Carnegie Proc. Calif. Acad. II, 6: 283. 1896. Polygalaceae. = P. Mus. 26: 220. 1937. Brassicaceae. Uintah Co., Chandler acanthoclada Gray San Juan Co., Willow Creek, East- Canyon, Graham 9976, 1935 (US!;CM!). wood 10, 1895 (CAS!). Physaria lepidota Rollins Brittonia 33: 335. 1981. Polygonum minimum Wats. Rep. U.S. Geol. Explor. Brassicaceae. Kane Co., between Kanab and Zion, Roll- 40th Parallel, Bot. 5: 315. 1871. Polygonaceae. Summit ins & Rollins 79198, 1979 (GH). Co., Uinta Mts., Watson 1058, 1869 (US!). Physaria newberryi Gray var. racemosa Rollins Brit- Polygonum utahense Brenckle & Cottam Bull. Univ. tonia 33: 339. 1981. Brassicaceae. Washington Co., 13.5 Utah 30: 3. 1940. Polygonaceae. Garfield Co., 6 m n Es-

mi s St. George, Holmgren et al. 9183, 1979 (GH). calante, Cottam 6507, 1935 (UT!;US!;BRY!;CAS!; Pinus contorta Dougl. ex Loud. var. latifolia Engelm. RM!;UTC!). ex Wats. Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: Potamogeton pusillus L. var. vulgaris subvar. inter- 331. 1871. Pinaceae. = P. contorta Dougl. Summit Co., ruptus J.W. Robbins Rep. U.S. Geol. Explor. 40th Paral-

Uinta Mts., Watson 1110, 1869 (?). lel, Bot. 5: 338. 1871. Potamogetonaceae. = P. pusillus Plantago major L. var. pachyphylla Pilger Feddes var. pusillus Summit Co., Parleys Park, Silver Cr., Wat- Repert. 18: 277. 1922. Plant aginaceae. = P. major L. son 1137, 1869 (NY!). Salt Lake Co., Salt Lake City, Jones 1030, 1879 Potentilla dissecta var. decurrens Wats. Proc. Amer. (POM!;UT!). Acad. 8: 557. 1873. Rosaceae. Syn: P. decurrens (Wats.)

Plantago myosuroides Rydb. Mem. New York Bot. Rydb. = P. ovina var. decurrens (Wats.) Welsh & John- Card. 1: 369. 1900. Plantaginaceae. = P. elongata Pursh ston Summit Co.?, Uinta Mts., Watson 329, 1869 (US!). Salt Lake Co., Salt Lake Valley, near mouth of Jordan Potentilla diversifolia var. pinnatisecta Wats. Rep. River, (US!). Watson 749, 1869 U.S. Geol. Explor. 40th Parallel, Bot. 5: 87. 1871. Ro- Platystemon remotus Greene Pittonia 5: 190. 1903. saceae. = P. ovina Macoun Summit Co., Uinta Mts., Papaveraceae. = P. californicus Benth. Washington Watson 331, 1869 (NY!). Co., near St. George, Parry 8, 1874 (GAS!;1SC!). Potentilla modesta Rydb. N. Amer. Fl. 22: 331. 1908. Platystemon rigidulus Greene Pittonia 5: 167. 1903. Rosaceae. = P. concinna var. modesta (Rydb.) Welsh & Papaveraceae. = P. californicus Benth. Washington Johnston Piute Co., Mt. Barrette, near Marysvale, Ryd- Co., near St. George, Parry 8a, 1874 (CAS!). berg & Carlton 7261, 1905 (NY!). Platystemon terminii Fedde Pflanzenr. 4. Fam. 104: Potentilla paucijuga Rydb. N. Amer. Fl. 22: 348. 128. 1909. Papaveraceae. = F. californicus Benth. 1908. Rosaceae. = P. pensylvanica var.- paucijuga Washington Co., Diamond Valley, Jones 5121, 1894 (Rydb.) Welsh & Johnston Grand (?) Co., La Sal Mts., (POM!;US!). Purpus 251, 1899 (NY!). Pleurophragma platypodum Rydb. Bull. Torrey Bot. Potentilla pectinisecta Rydb. Bull. Torrey Bot. Club Club 34: 4.34. 1907. Brassicaceae. = Thelypodium integ- 24: 7. 1897. Rosaceae. = P. gracilis Dougl. Salt Lake rifolium var. gracilipes Robins. Grand Co., Moab, Jones Co., Salt Lake City, Jones 1765, 1880 (RM!;NY!;BRY!). sn, 1891 (NY!;CAS!;RM!;US!;BRY!). Potentilla proxima Rydb. N. Amer. Fl. 22(4): 339. Poa eatonii Wats. Rep. U.S. Geol. Explor. 40th Paral- 1908. Rosaceae. = F. concinna var. proxima (Rydb.) lel, Bot. 5: 386. 1871. Poaceae. = F. fendleriana (Steud.) Welsh & Johnston Piute Co, s Belknap Peak, Rydberg & Vasey Salt Lake Co., Cottonwood Canyon, Eaton 1891, Carlton 7369, 1905 (NY!). 1869 (NY!). 182 Great Basin Naturalist Vol. 42, No. 2

Potentilla sabulosa Jones Proc. Calif. Acad. Sci. II, 5: Psoralea stenophylla Rydb. Bull. Torrey Bot. Club 680. 1895. Rosaceae. = Ivesia sabulosa (Jones) Keck 40: 46. 1913. Fabaceae. = F. lanceolata var. stenophylla Garfield Co., Sevier River, Jones 6032, 1894 (US!; (Rydb.) Toft & Welsh Grand Co., Wilson Mesa, Rydberg POM!;CAS!;RM!;NY!). & Garrett 8367, 1911 (NY!).

Potentilla wardii Greene Leafl. Bot. Obs. & Crit. 2: Psoralea stenostachys Rydb. Bull. Torrey Bot. Club 138. 1911. Rosaceae. = P. hippiana Lehm. Wayne Co., 40: 46. 1913. Fabaceae. = P. lanceolata var. stenos- Thousand Lake Mt., Ward sn, 1875 (US!). tachys (Rydb.) Welsh Tooele Co., Government Well,

Primula incana Jones Proc. Calif. Acad. II, 5: 705. Jones 6221, 1900 (NY!;US!;CAS!;POM!;RM!;BRY!). 1895. Primulaceae. Garfield Co., Sevier River, Jones Ptelea neglecta Greene Contr. U.S. Natl. Herb. 10: 5312av, 1894 (POM!). 71. 1906. Rutaceae. = P. trifoliata ssp. pallida var. lutes- cens (Greene) V.K. Bailey Kane Co., near Kanab, Primula maguirei L.O. Williams Amer. Midi. Natu- Weth- erill sn, 1897 (CAS!). ralist 7: 747. 1936. Primulaceae. Cache Co., Logan Can- Pterchiton occidentale Torr. & Frem. in Frem. Rep. yon, Maguire & Maguire 3650, 1932 (M0;UT!;UTC!). Exped. Rocky Mts. 318. 1845. Chenopodiaceae. = Atri- Primula specuicola Rydb. Bull. Torrey Bot. Club 40: plex canescens (Pursh) Nutt. Davis (?) Co., Great Salt 461. 1913. Primulaceae. San Juan Co., near Bluff, Ryd- Lake, Fremont sn, 1843 (NY!). berg9882, 1911 (NY!;US!;UT!). Ptilocalais macrolepis Rydb. Bull. Torrey Bot. Club Prosartes trachycarpa Wats. Rep. U.S. Geol. Explor. 38: 11. 1911. Asteraceae. = Microseris nutans (Geyer) 40th Parallel, Bot. 5: 344. 1871. Liliaceae. = Disponim Schulz-bip Salt Lake Co., Salt Lake City, Garrett 182, trachycarpum Wats. Summit Co., Parleys Park, Watson 1903 (US!;NY!;BRY!). 1166,1869 (NY!). Pyrrocoma cheiranthifolia Greene Leafl. Bot. Obs. & Psathyrotes pilifera Gray Proc. Amer. Acad. 19: 50. Crit. 2: 47. 1910. Asteraceae. = Haplopappus dementis 1883. Asteraceae. Kane Co., Kanab, Thompson sn, 1872 (Rydb.) Blake Sanpete Co., w. Ephriam, Tidestrom 534, (GH). 1907 (US!). Pseudocymoptems tidestromii Coult. & Rose Contr. Pyrrocoma lapathifolia Greene Leafl. Bot. Obs. & U.S. Natl. Herb. 12: 447. 1909. Apiaceae. = Cymopterus Crit. 2: 13. 1909. Asteraceae. = Haplopappus integri- lemmonii (Coult. & Rose) Dorn Sevier Co., Mt. Terrell, folius Gray? Utah, Ward 596, 1875 (US!). Tidestrom 1811, 1908 (US!). Pyrrocoma subcaesia Greene Leafl. Bot. Obs. & Crit. 2: 12. 1909. Pseudocymoptems versicolor Rydb. Fl. Rocky Mts. Asteraceae. = Haplopappus dementis 623, 1064. 1917. Apiaceae. = Cymopterus lemmonii (Rydb.) Blake Aarfield Co., Panguitch Lake, Jones 6005, 1894 (US!;RM!;POM!;NY!;BRY!). (Coult. & Rose) Dorn Garfield Co., Aquarius Plateau, Quercus eastwoodiae Rydb. Bull. New York Bot. Rydberg & Carlton 7426, 1905 (NY!). Card. 2: 210. 1901. Fagaceae. = Q. gambelii Nutt. San Pseudopteryxia longiloba Rydb. Bull. Torrey Bot. Juan Co., Butler Wash, Eastwood 141, 1895 (NY!;US!). Club 40: 72. 1913. Apiaceae. = Cymopterus hendersonii Quercus stellata var. utahensis A. DC. Prodr. 16(2): (Coult. & Rose) Cronq. San Juan Co., Abajo Mts., Ryd- 22. 1864. . = Q. gambdii Nutt. Utah? (between Salt berg & Garrett 9761, 1911 (NY!;UT!). Lake and Sierras) Beckwith (?) sn, 1854?(G). Pseudotsuga globosa Flous Bull. Soc. Hist. Nat. Tou- Ranunculus acriformis Gray var. aestivalis L. Benson louse 66: 334. 1934. Pinaceae. = P. menziesii (Mirb.) Amer. Midi. Naturalist 40: 43. 1948. Ranunculaceae. Franco Salt Lake Co., Big Cottonwood Canyon, Ryd- Garfield Co., 8.3 mi n Panguitch, Benson 13421, 1948 berg & Carlton 6587, 1905 (?). (POM;RM;BRY!;UT!;UTC!). Psoralea castorea Wats. Proc. Amer. Acad. 14: 291. Ranunculus alismaefolius Geyer ex Benth. var. mon- 1879. Fabaceae. Beaver Co. (more likely Beaverdam, tanus Wats. Rep. U.S. Geol. Explor. 40th Parallel, Bot.

Arizona), near Beaver City, Palmer 96, 1877 5: 7. 1871. Ranunculaceae. Summit (?) Co., Uinta Mts., (US!;NY!;BRY!;ISC!). Watson 18, 1969 (US!).

Psoralea epipsila Bameby Leafl. W. Bot. 3: 193. 1943. Ranunculus andersonii Gray var. tenellus Wats. Rep. Fabaceae. Kane Co., 17 mi e Kanab, Ripley & Barneby U.S. Geol. Explor. 40th Parallel, Bot. 5: 7. 1871. Ra- 4832, 1942 (CAS!). nunculaceae. = R. juniperinus Jones Salt Lake (?) Co.,

Pilot Rock Point, Salt Lake, Watson 17, 1869 (?). Psoralea juncea Eastw. Proc. Calif. Acad. 11, 6: 286. Ranunculus juniperinus Jones Proc. Calif. Acad. II, 1897. Fabaceae. San Juan Co., Epsom Creek, Eastwood 5: 616. 1895. Ranunculaceae. Syn: R. andersonii var. te- 21, 1895 (CAS!;US!;NY!). nellus Wats. Washington Co., 18 mi w St. George, Jones Psoralea mephitica Wats. Proc. Amer. Acad. 14: 291. 5011, 1894 (US!). 1879. Fabaceae. Washington Co., near St. George, Pal- Ranunculus multifidus Pursh var. repens Hook, in mer 97, 1877 (US!;NY!;BRY!;1SC!). Wats. Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: 8. Psoralea pariensis Welsh & Atwood in Welsh, At- 1871. . = R. gmdinii var. hookeri (D. Don) L. Benson wood, & Reveal Great Basin Naturalist 35: 353. 1975. Summit Co., Weber Valley, Watson 24, 1869 (US!). Fabaceae. Garfield Co., Bryce Canyon, Welsh & Mur- Ranunculus orthrorhynchus Hook. var. alpinus Wats. dock 12859, 1975 (BRY!;UT!;NY!;US!;UTC!;ISC!). Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: 9. 1871. Psoralea rdfaelensis Jones Contr. W. Bot. 18: 41. Ranunculaceae. = R. adoneus var. alpinus (Wats.) L. 1933. Fabaceae. = P. aromatica Payson Grand Co., LaS- Benson Salt Lake (?) Co., Wasatch Mts., Watson 30, al Mts., Jones sn, 1914 (US!;CAS!). 1869 (US!). Psoralea rafaelensis Jones var. magna Jones Contr. Ranunculus orthrorhynchus Hook. var. platyphyllus W. Bot. 18: 41. 1933. Fabaceae. = P. aromatica Payson Gray Proc. Amer. Acad. 21: 377. 1886. Ranunculaceae. Emery Co., San Rafael Swell, Jones sn, 1914 (CAS!). Salt Lake (?) Co., Wasatch Mts., Watson 28, 1869 (US!). June 1982 Welsh: Utah Plant Types 183

Bull. Torrey Bot. Club Ranuticulus utahensis Rydb. Sclerocactus parviflorus Clover & Jotter Bull. Torrey 29: 158. 1902. Ranunculaceae. = R. inamoenus var. al- Bot. Club 68: 419. 1941. Cactaceae. Syn: S. whipplei au- L. Salt Lake Co., Alta, peaphiltis (A. Nels.) Benson Jones thors, not (Engelm. & Bigel) Britt. San Juan Co., Glen 1130, 1879 (NY!;POM!;UTC!). Canyon, Clover & Jotter 2398, 1940 (US!). Rhus nitens Greene Proc. Washington Acad. Sci. 8: Sclerocactus terrae-canyonae Heil Cact. Succ. J. Anacardiaceae. = R. glabra L. Utah Co., Pro- 190. 1906. (U.S.) 51: 26. 1979. Cactaceae. San Juan Co., without lo- vo, Jones 5612, 1894 (US!;RM!;NY!). cality, Heil sn, (UNM). Rhus utahensis Goodding Bot. Gaz. 37: 57. 1904. Sclerocactus wrightiae L. Benson Cact. & Succ. J. = var. (Greene) .\nacardiaceae. R. trilobata simplicifolia (U.S.) 38: 55. 1966. Cactaceae. Emery Co., San Rafael Barklev Washington Co., Diamond Valley, Goodding Ridge, Benson & Benson 16595 (POM). 832, 1902 (US!;RM!). Scrophularia utahensis Gandg. Bull. Soc. Bot. France jonesii Coult. Rose Contr. U.S. Natl. Rhysopterus & 66: 219. 1919. Scrophulariaceae. = S. lanceolata Pursh? Herb. 7: 186. 1900. Apiaceae. = Cymopterus coulteri Cache Co., Linford sn, 1897 (?). 1880 (Jones) Mathias Juab Co., Juab, Jones 1691, Sedum meehanii Gray Proc. Amer. Acad. 16: 105.

(US!;POM!;BRY!). 1880. Crassulaceae. = ? S. lanceolatum Torr. Salt Lake Rites lacustre (Pers.) Poir. in Lam. var. lentutn Jones Co., City Creek Canyon, Reading (GH). Proc. Calif. Acad. II, 5: 681. 1895. Saxifragaceae. = R. 39: 83. Selaginella utahensis Flowers Amer. Fern J. Garfield Co., Henry Mts., montigenum McClatchie 1949. Selaginellaceae. Washington Co., s St. George, Jones 56950, 1894 (POM!). Cottam 5644, 1931 (UT!;BRY!). Riddellia tagetina var. sparsiflora Gray Syn. Fl. N. Selaginella watsonii Underwood Bull. Torrey Bot. = Psilostrophe spar- Amer. 1(2): 318. 1884. Asteraceae. Club 25: 127. 1898. Selaginellaceae. Salt Lake Co., Cot- Utah), Bish- siflora (Gray) A. Nels. Kane Co., (Southern tonwood Canyon, Watson 2370, 1869 (US!). op, Mrs. Thompson sn, 1872 (GH?). Senecio aquariensis Greenm. Ann. Missouri Bot. integra Rydb. Bull. Torrey Bot. Club 29: 236. Rorippa Card. 3: 144. 1916. Asteraceae. = S. streptanthifolius = (Rydb.) 1902. Brassicaceae. R. curvipes var. integra Greene Garfield Co., Aquarius Plateau, Ward 505, 1875 Stuckey Wasatch Mts., Watson 64, 1869 (US!;NY!). (GH;US!). Rosa chrysocarpa Rydb. Bull. Torrey Bot. Club 44: Senecio convallium Greenm. Ann. Missouri Bot. 74. 1917. = R. woodsii Lindl. San Co., Rosaceae. Juan Card. 1: 266. 1914. Asteraceae. = S. canus Hooker Garrett Allen Canyon, sw of Abajo Mts., Rydberg & Wavne Co., Rabbit Valley, Ward 704, 1875 (US!). 9312, 1913 (NY!). Senecio dimorphophyllus Greene var. intermedius Rosa puberulenta Rydb. Fl. Rocky Mts. 443. 1917. T.M. Barkley Trans. Kansas Acad. 65: 362. 1963. Aste- = Rosaceae. R. woodsii Lindl. San Juan Co., Mon- raceae. San Juan Co., LaSal Mts., Payson & Payson 4097, tezimia Canyon, e of Monticello, Rydberg & Garrett 1924 (MO;GH;RM!;UC). 9705, 1913 (NY!). Senecio incurvus A. Nels. Univ. Wyoming Publ. Bot. maritimus L. var. athrix St.John Rhodora 17: Rumex 1: 141. 1926. Asteraceae. = S. spartioides T. & G. Wash- 79. 1915. Polvgonaceae. = R. maritimus L. Sevier Co., ington Co., Zion National Park, Nelson 9988, 1922 Vemiillion, Jones 58.39, 1894 (MO;GH;POM!). (RM!). II, 720. Rumex subalpina Jones Proc. Calif. Acad. 5: Senecio jonesii Rydb. Bull. Torrey Bot. Club 27: 179. 1895. = R. pycnanthus Rech. f. Piute Polvgonaceae. 1900. Asteraceae. = S. streptanthifolius Greene Salt Co., Brigham Peak, Jones 5957, 1894 (POM!; Lake Co., Alta, Jones 1125, 1879 (NY!;POM!;UTC!). US!;RM!;BRY!;NY!). Senecio kingii Rydb. Bull. Torrey Bot. Club 37: 468. Rech. f. Repert. Sp. Nov. 40: 298. Rumex utahensis 1910. Asteraceae. = S. eremophilus var. kingii (Rydb.) = Wein. Carbon 19.36. Polvgonaceae. R. salicifolius Greenm. Salt Lake Co., Cottowood Canyon, Watson 676 Co., Kyime, Jones 5603, 1894 (US!). (NY!;US!). Salicornia utahensis Tidestr. Proc. Biol. Soc. Wash- Senecio lapidum Greenm. Ann. Missouri Bot. Gard. 4: ington 26: 13. 1913. Chenopodiaceae. = S. pacifica var. 18. 1917. Asteraceae. = S. multilobatus T. & G. ex Gray utahensis (Tidestr.) Munz Utah (?). Washington Co., Silver Reef, Jones 5163v, 1894 Salix lutea Nutt. var. platyphylla Ball Bot. Gaz. 71: (NY!;BRY!). = 4.30. 1921. Salicaceae. S. rigida Muhl. Salt Lake Co., Senecio leonardii Rydb. Bull. Torrey Bot. Club 37: City Ball 1.336, 1908 (US!). Creek Canyon, 468. 1910. Asteraceae. = S. streptanthifolius Greene var. purpusii Saxifraga caespitosa L. ssp. exaratoides Utah Co., American Fork Canyon, Leonard 143, 1884 4. 377. 1916. Engler & Irmsch. Pflanzenr. Fam. 117(1): (NY!). = Saxifragaceae. S. caespitosa var. minima Blankinship Senecio lugens Richards var. hookeri D.C. Eaton in (RM!). Grand (?) Co., LaSal Mts., Purpus 6642, 1899 Wats. Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: 188. Leafl. Bot. Obs. Crit. 1: Schmaltzia affinis Greene & 1871. Asteraceae. = S. sphaerocephalus Greene Summit 135. 1905. .\nacardiaceae. = Rhus trilobala var. sim- Co., Salt Lake City to Uintas, Watson 661, 1869 plicifolia (Greene) Barkeley Kane Co., Kanab, Jones (US!;NY!). 5286e, 1894 (POM!;US!). Senecio malmstenii Blake in Tidestr. Proc. Biol. Soc. Schoencrambe pinnata Greene Pittonia 3: 127. 1896. Washington 36: 183. 1923. Asteraceae. = S. streptanthi- Brassicaceae. = S. linifolia (Nutt.) Greene Utah, Ward folius Greene Kane Co., Little Podunk Creek, Malmsten 1875 (US!). 131, 1916 (US!). Heil Cact. Succ. (U.S.) 51: 25. multilobatus T. G. ex Gray Mem. Amer. Sclerocactus contorus J. Senecio & 1979. Cactaceae. San Juan Co., Canyonlands National Acad. II, 4: 109. 1849. Asteraceae. Uintah Co., Uinta Park, Heil sn, (UNM). River, Fremont 549, 1845 (NY!). 184 Great Basin Naturalist Vol. 42, No. 2

Senecio pammelii Greenm. Ann. Missouri Bot. Card. Sphaeralcea psoraloides Welsh Great Basin Naturalist 3: 118. 1916. Asteraceae. = S. streptanthifalius Greene 40: 36. 1980. Malvaceae. Wayne Co., Salt Wash, Welsh Morgan Co., Peterson, Pammel & Blackwood .3870, 1902 13348, 1977 (BRY!). (MO;GH). Sphaeralcea subrhomhoidea Rydb. Bull. Torrey Bot. Senecio platylohus Rydb. Bull. Torrey Bot. Club 27: Club 40: .59. 1913. Malvaceae. = S. munroana (Dougl.) 181. 1900. Asteraceae. = S. streptanthifolius Greene Spach Wasatch Co., Midway, Carlton & Garrett 6691, Wasatch Mountains, Watson 671, 1869 (NY!). 1905 (NY!;US!). Senecio rubricaulis var. aphanactis Greenm. Ann. Sphaeromeria ruthiae Holmgren, Schultz, & Lowrey Missouri Bot. Card. 3: 174. 1916. Asteraceae. = S. strep- Brittonia 28: 257. 1976. Asteraceae. Washington Co., tanthifolius Greene Cache Co., Logan, Smith 2208, 1910 Zion Canyon, Holmgren et al. 16603, 1974 (UTC!; (UTC!). BRY!;UT!;US!;NY!). Senecio wardii Greene Pittonia 4: 116. 1900. Aste- Sphaerostigma utahense Small Bull. Torrey Bot. Club raceae. = S. streptanthifolius Greene Sevier Co., Fish 23: 191. 1896. Onagraceae. = Camissonia boothii ssp. Lake Mt., Ward 332, 1875 (GH;US!). alyssoides (H. & A.) Raven Beaver Co., Milford, Jones Shepherdia rotundifolia Parry Amer. Naturalist 9: 1773, 1880 (NY!;POM!). 350. 1875. Elaeagnaceae. Kane Co., Valley of the Virgin, Spiraea caespitosa Nutt. in T. & G. var. elatior Wats. Siler sn, 1875 (US!;NDG!). Rep. U.S. Geol. Explor. 40th Parallel, Bot. 5: 81. 1871. SidaJcea crenulata A. Nels. Proc. Biol. Soc. Washing- Rosaceae. = Petrophytum caespitosum (Nutt.) Rydb.. ton 17: 93. 1904. Malvaceae. = var. S. neomexicana cre- Box Elder Co., Raft R. Mts., Watson .307, 1869 (NY!). nulata (A. Nels.) C.L. Hitchc. Juab Co., Juab, Goodding Stachys asperrima Rydb. Bull. Torrey Bot. Club 36: 1091, 1902 (RM!;BRY!;NY!;ISC!). 682. 1909. Lamiaceae. = S. palustris L. Salt Lake Co., Sieversia scapoidea A. Nels. in Coult. & Nels. New Jordan, Leonard 138, 1884 (NY!). Man. Bot. = Rocky Mts. 263. 1909. Rosaceae. Geum Stanleya canescens Rydb. Bull. Torrey Bot. Club rossii (R. Br.) Ser. Piute Co., of Marysvale, w Jones 5871, 29(1): 232. 1902. Brassicaceae. = S. pinnata var. pinnata 1894 (RM!;POM!). Beaver Co., Frisco, Jones 1809, 1880 (NY!;POM!). Silene petersonii Maguire var minor Hitchc. & Ma- Stenotus falcatus Rydb. Bull. Torrey Bot. Club 27: guire Univ. Washington Publ. Biol. 13: 38. 1947. Caryo- 616. 1900. Asteraceae. = Haplopappus acaulis var. gla- phyllaceae. Garfield Co., Red Canyon, Maguire 19550, bratus (D.C. Eaton) Hall Washington Co. (s. Utah), Red 1940 (UTC!;NY!). Creek, Palmer 202, 1877 (US!;NY!;ISC!). Silene petersonii var. Maguire petersonii Madrono 6: Stenotus latifolius A. Nels. Bot. Gaz. 37: 266. 1904. 24. 1941. Caryophyllaceae. Sanpete Co., Wasatch Asteraceae. = Haplopappus acaulis var. glabratus (D.C. Plateau, Maguire 20000, 1940 (UTC!;NY!;BRY!). Eaton) H.M. Hall Utah Co., near Provo, Goodding 1111, Sisyrinchium radicatum Bickn. Bull. Torrey Bot. 1902 (US!;NY!;BRY!;1SC!). Club. 28: 576. 1901. Iridaceae. = S. demissiim Greene Stipa arida Jones Proc. Calif. Acad. 11, 5: 725. 1895. Washington Co., St. George, Palmer 456, 1877 Poaceae. Syn: S. mormonum Mez Piute Co., near Mary- (NY!;ISC!). svale, Jones 5377, 1894 (POM!;RM!;US!;NY!). Sitanion insulare J.G. Sm. USDA Div. Agrostol. Bull. Stipa mormonum Mez. Feddes Repert. 17: 209. 1921. 18: 14. 1899. Poaceae. = Agrositanion saxicola (Scribn. Poaceae. = S. arida Jones Beaver Co., Milford, Jones & Sm.)Bowden Tooele Co., Carrington Island, Watson 2106, 1880(US!frag;UT!;UTC!). 1338, 1869 (US!). Stipa parishii var. depauperata Jones Contr. W. Bot. Solidago garrettii Rydb. Bull. Torrey Bot. Club 37: 14: 11. 1912. Poaceae. = S. coronata var. depauperata 134. 1910. Asteraceae. = S. sparsi flora Gray Salt Lake (Jones) A.S. Hitchc. Juab Co., Detroit, Jones, 1891 (US!). Co., Big Cottonwood Canyon, Garrett 2041, 1906 (NY!;UT!). Stipa pinetoTum Jones Proc. Calif. Acad. II, 5: 724. 1895. Poaceae. Garfield Co., Panguitch Lake, Solidago radulina Rydb. Bull. Torrey Bot. Club 31: Jones 6023, 1894 (POM!;US!). 650. 1904. Asteraceae. = S. nana Nutt. Salt Lake Co., Streptanthus crassicaulis Torr. in Cottonwood Canyon, Watson .558, 1869 (NY). Stansbury Expl. Sonchus asper (L.) All. var. glanduliferus Garrett Spr. Great Salt Lake, Append. D. Botany 383. 1852. Brassi- = Fl. Wasatch ed. 3. 119. 1917. Asteraceae. = S. asper (L.) caceae. Caulanthus crassicaulis (Torr.) Wats. Tooele

Hill Utah (?). Co., w shore of Great Salt Lake, Stansbury sn, 1850

Sophia leptostylis Rydb. Bull. Torrey Bot. Club .39: (NY!). 325. 1912. Brassicaceae. = Descurainia californica Suaeda intermedia Wats. Proc. Amer. Acad. 14: 296. (Gray) Schulz Salt Lake Co., Big Cottonwood Canyon, 1879. Chenopodiaceae. Sevier Co., Glenwood, Ward Rydberg & Carlton 6629, 1905 (NY!;US!;RM!). 718, 1875 (US!). Sphaeralcea caespitosa Jones Contr. W. Bot. 12: 4. Swertia fritillaria Rydb. Bull. Torrey Bot. Club 40:

1908. Malvaceae. Beaver (?) Co., Wah Wah, Jones sn, 465. 1913. Gentianaceae. = S. perennis L. Salt Lake Co, 1906 (POM!;CAS!;US!;BRY!;NY!). Big Cottonwood Canyon, Garrett 1566, 1905 (NY!;UT!). Sphaeralcea grossulariifolia (H. & A.) Rydb. var. Synthris laciniata (Gray) Rydb. ssp. ibahpensis Pen- moorei Welsh Great Basin Naturalist 40: 35. 1980. Mal- nell Proc. Acad. Nat. Sci. Philadelphia 85: 92. 1933. vaceae. Kane Co., Lake Powell, Welsh & Atwood 11597, Scrophulariaceae. = S. laciniata (Gray) Rydb. Juab Co., 1972 (BRY!). Mt. Ibapah, Stanton 1000, 1932 (PH). Sphaeralcea leptophylla (Gray) Rydb. var. janeae Synthris pinnatifida Wats. Rep. U.S. Geol. Explor. Welsh Great Basin Naturalist 40: 36. 1980. Malvaceae. 40th Parallel, Bot. 5: 227. 1871. Scrophulariaceae. Utah San Juan Co., White Rim road, Welsh 7085, 1968 Co., American Fork Canyon, Watson 802, 1869 (BRY!). (US!;NY!). June 1982 Welsh: Utah Pla>jt Types 185

Synthris pinnatifida Wats. var. laciniata Gray Syn. Thlaspi fendleri Gray var. tenuipes Maguire Amer. Fl. N. Amer. 2(1): 286. 1878. Scrophulariaceae. = S. Midi. Naturalist 24: 469. 1942. Brassicaceae. = T. mon- laciniata (Gray) Sevier Co., Fish Lake Mts., Ward 327, tanum var. montanum Sanpete Co., Mayfield Canyon, 1875 (US!). Maguire 19998, 1940 (UTC!). Tanacetum diversifolium D.C. Eaton Rep. U.S. Geol. Thlaspi prolixum A. Nels. Amer. Bot. 32: 287. 1945. = J. Explor. 40th Parallel, Bot. 5: 180. 1871. Asteraceae. Brassicaceae. = T. montanum var. montanum Piute Co., Sphaeromeria diversifolia Rydb. Utah Co., American Marysvale, Jones 5374, 1894 (RM!;POM!). Fork Canyon, Watson 632, 1869 (US!). Thysanocarpus trichocarpus Rydb. Bull. Torrey Bot.

Tetradymia linearis Rydb. Bull. Torrey Bot. Club 32: Club 30: 253. 1903. Brassicaceae. = T. curvipes Hook. 130. 1905. Asteraceae. = T. canescens DC. Iron Co., Washington Co., Silver Reef, Jones 5163b, 1894 (US!). Rock Creek, Palmer 264, 1877 (NY!). Tithonia argophylla D.C. Eaton Rep. U.S. Geol. Ex- Tetradymia spinosa H. & A. var. longispina Jones plor. 40th Parallel, Bot. 5: 423. 1871. Asteraceae. = En- Proc. Calif. Acad. II, 5: 698. 1895. Asteraceae. = T. ax- celiopsis argophylla (D.C. Eaton) A. Nels. Washington illaris var. longispina Jones Washington Co., St. George, Co., St. George, Palmer sn, 1877 (US!).

Jones 5110, 1894 (POM!;US!;NY!;BRY!). Townsendia annua Beaman Contr. Gray Herb. II, Tetraneuris epunctata A. Nels. Bot. Gaz. 37: 275. 183. 132. 1957. Asteraceae. San Juan Co., 1.5 mi n. Bluff, 1904. Asteraceae. = Hymenoxys acaulis var. caespitosus Maguire 13509, 1936 (UTC!). (A. Nels.) Parker Uintah Co., Dyer Mine, Goodding Townsendia aprica Welsh & Reveal Brittonia 20: 375.

1236, 1902 (RM!;US!;UT!;NY!;BRY!;ISC!). 1968. Asteraceae. Sevier Co., s Fremont Jet., Reveal & Thalictrum duriusculum Greene Leafl. Bot. Obs. & Welsh 721, 1966 (BRY!;CAS!). Crit. 2: 92. 1910. Ranunculaceae. = T. alpinum L. Se- Townsendia dejecta A. Nels. Bot. Gaz. 37: 267. 1904. vier Co., near Fish Lake, Jones 5826a, 1894 (US!;POM!). Asteraceae. = T. montana var. montana Uintah Co., Thelesperma subnudum Gray Proc. Amer. Acad. 10: Dyer Mine, Goodding 1283, 1902 (US!;RM!). 72. 1874. Asteraceae. Washington Co., near St. George, Townsendia florifer (Hook.) Gray var. communis Parry 109, 1874 (US INY!;ISC!;BRY!). Jones Proc. Calif. Acad. II, 5: 697. 1895. Asteraceae. = Thelypodiopsis argillacea Welsh & Atwood Great Ba- r. florifer (Hook.) Gray Piute Co., Kingston, Jones 5322f, sin Naturalist 37: 95. 1977 Brassicaceae. Uintah Co., Big 1894 (POM!;US!;BRY!). Pack Mountain, Atwood 6627, 1976 (BRY!). Townsendia incana Nutt. var. prolixa Jones Contr. Thelypodiopsis bamebyi Welsh & Atwood in Welsh W. Bot. 13: 15. 1910. Asteraceae. = T. strigosa Nutt. Brittonia 33: 300. 1981. Brassicaceae. Emery Co., San Duchesne Co., Duchesne Valley, Jones 5323, 1908 Rafael Swell, Welsh 20345, 1981 (BRY!;NY!;GH!;UTC!). (POM!;BRY!). Thelypodiutn elegans Jones Zoe 4: 265. 1893. Brassi- Townsendia mensana Jones Contr. W. Bot. 13: 15. caceae. = Tlielypodiopsis elegans (Jones) Rydb. Grand 1910. Asteraceae. Duchesne Co., near Theodore, Jones Westwater, sn, 1891 Co., Jones (POM!;RM!;BRY!;NY!). sn, 1908 (POM!;BRY!). Thelypodium macropetalum Rydb. Bull. Torrey Bot. Townsendia mensana Jones var. jonesii Beaman Club 29: 233. 1902. Brassicaceae. = T. sagittatiim var. Contr. Gray Herb. II, 183: 88. 1957. Asteraceae. = T. jo- sagittatum Davis Co., Farmington, Jones 1841, 1881 nesii (Beaman) Reveal Juab Co., Mammoth, Jones sn, (NY!;US!;CAS!;POM!;BRY!;UTC!). 1910(POM!;BRY!). Thelypodium ovalifolium Rydb. Bull. Torrey Bot. Townsendia minima Eastw. Leafl. W. Bot. 1: 206. Club 30: 253. 1903. Brassicaceae. = T. sagittatum var. 1936. Asteraceae. Garfield Co., Bryce Cyn., Eastwood & ovalifolium (Rydb.) Welsh & Reveal Garfield Co., Pan- Howell 727, 1933 (CAS!). guitch Lake, Jones 6015e, 1894 (US!;POM!;NY!). Townsendia montana Jones Zoe 4: 262. 1893. Aste- Thelypodium palmeri Rydb. Bull. Torrey Bot. Club raceae. Syn: T. dejecta A. Nels. Salt Lake Co., Alta, 34: 433. 1907. Brassicaceae. = T. sagittatum var. ovalifo- above Flagstaff Mine, Jones sn, 1879 (POM!). lium (Rydb.) Welsh & Reveal Washington Co., (s. Utah), Townsendia scapigera var. ambigua Gray Proc. Palmer 25, 1877 (NY!;US!;BRY!;ISC!). Amer. Acad. 16: 84. 1880. Asteraceae. = T. florifer Thelypodium rollinsii Al-Shehbaz Contr. Gray Herb. (Hook.) Gray Wayne Co., Rabbit Valley, Ward 523, II, 204: 97. 1973. Brassicaceae. Juab Co., 12 mi n Scipio, 1875 (US!). I. & N. Al- Shehbaz 6913, 1969 (GH). Thelypodium sagittatum (Nutt.) Endl. var. vermicu- Townsendia watsonii Gray Proc. Amer. Acad. 16: 84. = laris Welsh & Reveal Great Basin Naturalist 37: 358. 1880. Asteraceae. T. florifer (Hook.) Gray Tooele Co., 1978. Brassicaceae. Sevier Co., 4 mi se Sigurd, Welsh & Stansbury Island, Watson 520, 1869 (US!). Atwood 11718, 1972 (BRY!). Toxicodendron longipes Greene Leafl. Bot. Obs. & Thelypodium suffrutescens Rollins in Graham Ann. Crit. 1: 118. 1905. Anacardiaceae. = T. rydbergii (Small) Carnegie Mus. 26: 224. 1937. Brassicaceae. = Glauco- Greene Sevier Co., s Glenwood, Ward 212, 1875 (US!). carpum suffrutescens (Rollins) Rollins Uintah Co., Big Trifolium andersonii Gray var. friscanum Welsh Pack Mtn, Graham 8959, 1935 (GH;CM!;NY!). Great Basin Naturalist 38: .355. 1978. Fabaceae. Beaver Thelypodium utahense Rydb. Bull. Torrey Bot. Club Co., Grampian Hill, Peabody et al 406, 1976 (BRY!;NY!). 29: 233. 1902. Brassicaceae. = Caulanthus lasiophyllus Trifolium confusum Rydb. Bull. Torrey Bot. Club 34: var. utahensis (Rydb.) Jeps. Washington Co., St. George, 46. 1907. Fabaceae. = T. longipes var. pygmaeutn Gray Jones 1648, 188(T(US!;UT!;POM!;NY!). Iron Co., near Cedar City, Parry 35, 1874 (NY!). Thelypodium wrightii Gray var. tenellum Jones Proc. Trifolium inequale Rydb. Bull. Torrey Bot. Club 34: Calif. Acad. II, 5: 622. 1895. Brassicaceae. = T. laxiflo- 47. 1907. Fabaceae. = T. parryi var. montanense (Rydb.) rum Al-Shehbaz Utah Co., Slate Canyon, Jones 5559, Welsh Summit Co., Bear River Canyon, Watson 243, 1894 (POM!;US!;CAS!;RM!;NY!). 1869 (NY!). 186 Great Basin Naturalist Vol. 42, No. 2

Trifolium kingii Wats. Rep. U.S. Geol. Explor. 40th Viola mamillata Greene Leafl. Bot. Obs. & Crit. 2: Parallel, Bot. 5; 59. 1871. Fabaceae. Summit Co., Parleys 33. 1910. Violaceae. = V. adunca J.E. Sm. Uintah Co., Park, Watson 239, 1869 (US!;NY!). Dyer Mine, Goodding 1202, 1902 (US!;RM!;UT!;BRY!). Trifolium longipes Nutt. var. brachypus Wats. Biblio. Viola oxysepala Greene Leafl. Bot. Obs. & Crit. 2: Index N. Amer. Bot. 264. 1878. Fabaceae. = T. longipes 34. 1910. Violaceae. = V. adunca J.E. Sm. Sanpete Co., var. pijgmaeum Gray Washington Co., near St. George, inter Willow & Ephriam creeks, Tidestrom 2476, 1909 Palmer sn, 1870 (GH). (NDG!).

Trifolium macilentum Greene Pittonia 3: 223. 1897. Viola tidestromii Greene Leafl. Bot. Obs. & Crit. 2:

Fabaceae. Washington Co., (s. Utah), Palmer sn, 1877 234. 1910. Violaceae. = V. adunca J.E. Sm. Sanpete (US!;NY!). Co., Ephriam Creek, Tidestrom 1143, 1908 (NDG!).

Trifolium uintense Rydb. Bull. Torrey Bot. Club 34: Viola utahensis Baker & Clausen Leafl. W. Bot. 5:

47. 1907. Fabaceae. = T. dasyphyUtim var. uintense 145. 1949. Violaceae. = V. purpurea Kellogg Cache Co., (Rydb.) Welsh Summit Co., Uintas, Watson 241, 1869 Providence Canyon, Maguire 16026, 19.37 (UTC; (NY!;US!). US!;NY!;UTC!). Trifolium villiferum House Bot. Gaz. 41: 335. 1906. Whipplea utahensis Wats. Amer. Naturalist 7: 300.

Fabaceae. = T. eriocephahim var. villiferum (House) 1873. Saxifragaceae. = Fendlerella utahensis (Wats.)

Martin Washington Co., (s. Utah), Palmer 91, 1877 Heller Kane Co., Kanab, Thompson 243, 1872 (US!). (US!;NY!;BRY!). Wyethia scabra Hook. var. attenuata W.A. Weber Midi. Naturalist 35: 425. 1946. Asteraceae. Triglochin maritima L. var. dehile Jones Proc. Calif. Amer. Kane Co., 10.5 mi n Kanab, Carter 1424, 1938 (WS). Acad. II, 5: 722. 1895. Juncaginaceae. = T. concinna Wyethia scabra Hook. var. canescens W.A. Weber var. debile (Jones) Howell Kane Co., Johnson, Jones Amer. Midi. Naturalist .35: 425. 1946. Asteraceae. San 5289, 1894 (US!;POM!). Juan Co., s Mexican Hat, Goodman & Hitchcock 1352, Valeriana puberulenta Rydb. Bull. Torrey Bot. Club 19.30 (CAS!;RM!;NY!). .36: 697. 1909. Valerianaceae. = V. acutiloba var. pub- Wyomingia vivax A. Nels. Bot. Gaz. 56: 70. 1913. As- icarpa (Rydb.) Cronq. Piute Co., Bullion Creek, Rydberg teraceae. = Erigeron utahensis var. sparsifolius (Eastw.) & Carlton 7065, 1905 (NY!;US!;BRY!). Cronq. San Juan Co., Geyser Canyon, Walker 355, 1912 Valeriana pubicarpa Rydb. Bull. Torrey Bot. Club (US!). 36: 697. 1909. Valerianaceae. = V. acutiloba var. pub- Xanthocephalum petradoria Welsh & Goodrich in icarpa (Rydb.) Cronq. Juab Co., Mt. Nebo, Rydberg & Welsh Brittonia 33: 301. 1981. Asteraceae. Millard Co., Carlton 7717, 1905 (NY!;US!;RM!). Canyon Mts., Goodrich 15240, 1980 (BRY!;ASU!; Valeriana utahensis Gandg. Bull. Soc. Bot. France GH!;MO!;NY!;RM!;TEX!;UC!;US!;UT!). 65: 37. 1908. Valerianaceae. = V. acutiloba var. pub- Xylophacos aragaloides Rydb. Bull. Torrey Bot. Club icarpa (Rydb.) Cronq. Cache (?) Co., Linford sn, 1907 (?). 34: 48. 1907. Fabaceae. = Astragalus amphioxys var. Verbesina scaposa Jones Zoe 2: 248. 1891. Aste- amphioxys Washington Co., St. George, Jones 1633, raceae. = Enceliopsis nutans (Eastw.) A. Nels. Grand 1880 (NY!;POM!;ISC!). Co., Cisco, Jones sn, 1890 (POM!;BRY!). Xylophacos marianus Rydb. Bull. Torrey Bot. Club Viguiera soliceps Barneby Leafl. W. Bot. 10: 316. 52: 233. 1925. Fabaceae. = Astragalus marianus (Rydb.) 1966. Asteraceae. = Heliomeris soliceps (Barneby) Yates Barneby Piute Co., Marysvale, Jones 5355, 1894 Kane Co., Cottonwood Canyon, Barneby 14435, 1966 (NY!;US!;POM!). (NY!;US!;UTC!). Xylophacos medius Rydb. Bull. Torrey Bot. Club 52: = Vilfa depauperata Torr. exHook. var. filiformis 232. 1925. Fabaceae. Astragalus eurekensis Jones Tooele Co., Lake Point, 1743, 1880 (US!; Thurb. in Wats. Rep. U.S. Geol. Explor. 40th Parallel, Jones CAS!;POM!;BRY!;NY!;UT!). Bot. 5: 376. 1871. Poaceae. = Muhlenbergia filiformis Xylophacos melanocalyx Rydb. Bull. Torrey Bot. (Thurb.) Rydb. Summit Co., Uinta Mts., Watson 1281, Club 52; 149. 1925. Fabaceae. = Astragalus amphioxys 1869 (US?).' var. amphioxys Washington Co., Beaverdam Mts., Jones Viola beckivithii T. & G. in Beckwith Rep. U.S. Ex- 5009, 1894 (NY!). plor. & Surv. R.R. Pacific 2: 119. 1855. Violaceae. Utah, Xylophacos uintensis Rydb. Bull. Torrey Bot. Club between Salt Lake and Sierra Nevada, Beckwith sn, .32: 662. 1905. Fabaceae. = Astragalus argophyllus var. 1854(NY!). argophyUus Salt Lake Co., Salt Lake V., Jones 1633b, Viola beckunthii T. & G. var. cachensis C.P. Sm. 1880 (POM!). Muhlenbergia 7: 136. = 1912. Violaceae. V. beckivithii Xylorhiza cronquistii Welsh & Atwood in Welsh Brit- T. & G. Utali. tonia .33: .302. 1981. Asteraceae. Kane Co., Horse Mt., Viola bellidifolia Greene ssp. valida Baker Madrofio Welsh & Welsh 12819, 1975 (BRY!). 5: 223. 1940. Violaceae. = V. adunca J.E. Sm. Salt Lake Xylorhiza glabriuscula Nutt. var. linearifolia T.J. Co., Brighton, Baker 8519, 19.36 (UTC!). Watson Brittonia 29: 215. 1977. Asteraceae. Grand Co., Viola bonnevillensis Cottam Bull. Univ. Utah Biol. 6 mi nw Moab, Watson 679, 1971 (TEX; Ser. 3: 3. 19.39. Violaceae. = V. beckivithii T. & G. Salt COLO;GH;MP;MONTU;NY!;UC). Lake Co., Salt Lake City, Cottam 7067, 1937 (UT!). Xylorhiza lanceolata Rydb. Bull. Torrey Bot. Club

Viola clauseniana Baker Madrofio 4: 194. 1938. Vio- 37: 146. 1910. Asteraceae. = X. tortifolia var. tortifolia laceae. = V. nephrophylla Greene Washington Co., Washington Co., St. George, Palmer 208, 1877 Zion National Park, Baker 84348, 1936 (UC). (NY!;US!). June 1982 Welsh: Utah Plant Types 187

Yucca angustissima Engelm. var. avia Reveal Inter- 1945. Pugillus Astragalorum— IV: The section mountain Fl. 6; 534. 1977. Liliaceae. Piute (?) Co., Loa Diplocystium. Leafl. W. Bot. 4: 65-147. Pass, Jones 5639a, 1898 (US!). 1946. Pugillus Astragalorum-V. Leafl. W. Bot. 4: Yucca harrimaniae Trel. Rep. Missouri Bot. Card. 13: 228-238. 59. 1902. Agavaceae. Svn: Y. harrimaniae var. gilher- 1947. Pugillus Astragalorum— VI: Notes on sec- tiana Trel. Carbon Co., Helper, Trelease 3233, 1889 tion Drabellae. Leafl. W. Bot. 5: 1-9. (MO!). 1947. Pugillus Astragalorum— VII: A revision of Yucca harrimaniae Trel. var. gilbertiana Trel. Rep. the Argophvlli. .4mer. Midi. Naturalist 37: Missouri Bot. Card. 18: 225. 1907. Agavaceae. = Y. har- 421-451. rimaniae Trel. Juab Co., Fish Springs or House Range, 1949. Pugillus .'Astragalorum— X: New species, varieties combinations. Amer. Midi. Naturalist Gilbert sn. 1901 (US!). and Yucca kanabensis McKelvey Yuccas S.W. U.S. 2: 122. 41: 496. 1949. villiflorum and its near relatives 1947. Agavaceae. Syn: Y. angustissima var. kanabensis Eriogonum Basin. Leafl. W. Bot. 5: 151-154. (McKelvey) Reveal Kane Co., between Mt. Carmel & in the Great 1952. A revision of the North American species of Kanab, Mckelvey 4347a, 1934 (A). Oxijtropis. DC. Proc. Calif. Acad. IV, 27: Yucca toftiae Welsh Great Basin Naturalist 34: 308. 177-312. 1974. .\gavaceae. Syn: Y. angustissima var. toftiae 1953. Pugillus Astragalorum—XV: Four new spe- (Welsh) Reveal San Juan Co., Lake Powell, Welsh cies. Leafl. W. Bot. 7: 31-37. 11935a, 1973 (BRY!;NY!). 1954. Pugillus Astragalorum—XVI: Minor novel- Yucca utahensis McKelvey Yuccas S.W. U.S. 2: 94. ties from Nevada. Leafl. W. Bot. 7: 192-196. 1947. Agavaceae. Syn: Y. elata var. utahensis 1956. Pugillus Astragalorum—XVII: Four new (McKelvev) Reveal Washington Co., Santa Clara V., species and one variety. Leafl. W. Bot. 8: 14-23. McKelvey 4167, 1934 (A). 1964. Atlas of North American Astragalus. Mem. Zauschneria garrettii A. Nels. Proc. Biol. Soc. Wash- N.Y. Bot. Garden 13: 1-1188. ington 20: 36. 1907. Onagraceae. = Epilobium canum 1966. New sorts of Lesquerella, Euphorbia, and (Greene) Raven Salt Lake Co., Big Cottonwood Canyon, Viguiera from Kane County, Utah. Leafl. W. Bot. Garrett 2031. 1906 (UT!). 10: 31.3-317. Barneby, R. C, and N. H. Holmgren. 1979. A new spe- from Utali. Brittonia Bibliography cies of Lomatium (.\piaceae) 31: 96-100. Beal, W. 1896. Grasses of North America. Vol. 2. .\l-shehbaz, I. A. 1973. The biosystematics of the genus J. New York, Henry Holt and Company. nehjpodiiim (Cmciferae). Contr. Gray Herb. II, Beaman, H. 1957. The systematics and evolution of 204: 3-148. J. Townsendia (Compositae). Contr. Gray Herb. II, .\.NDERSON, L. C. 1981. Chrysothamnus nauseosus ssp. 183: 1-151. iridis (.\steraceae): a new endemic from Utah. Beckwith, E. G. 1855. Reports of explorations and sur- Great Basin Naturalist 41(3): 311-313. vevs, to ascertain the most practicable and eco- .Atwood, N. D. 1972. New species in the Phacelia crenu- nomical route for a railroad from the Mississippi lata group (Hvdrophyllaceae). Rhodora 74: River to the Pacific Ocean. House Rep. E.xec. 451-463. Doc, Vol 2, No. 91. 1973. Two new species of Phacelia (Hydro- Benson, L. 1948. A treatise on the North American Ra- phyllaceae). Phytologia 26: 437-438. nunculi. Amer. Midi. Naturalist 40: 1-264. Bailey, L. H. 1884. Notes on Carex— I. Bot. Gaz. 1966. A revision of Sclerocactus. Cact. & Succ. J. 9:117-122. (Los Angeles) 38: 50-57, 100-106. Baker, S. 1938. An undescribed species of Viola from M. 1969. The cacti of the United States and Can- Utah. Madrono 4: 194-196. ada—new names and nomenclatural com- 1940. Studies in western violets— III. Madrono 5: (Los .\ngeles) 41: binations— 1. Cact. & Succ. J. 218-231. 124-28. S. 1949. Studies in western Baker, M. and J. Clausen. 1969. The cacti of Arizona. Ed. 3, Tucson. Uni- Violets-IV. Leafl. W. Bot. 5: 141-156. versity of Arizona Press. 218 pp. Ball, C. R. 1921. Undescribed willows of the section Bentham, G. 1856. Eriogonum confertiflorum var. stans- Cordatae. Bot. Gaz. 71: 426-4.37. bunji. Page 17 in A. P. de Candolle. Prodromus

Barkley, T. M. 1962. .\ revision of Senecio aureus Linn. Svstematis Naturalis Regni Vegetabilis. Vol. 14. and allied species. Trans. Kansas Acad. Sci. 65: Bicknell, E. P. 1901. Studies in Sisyrinchium— IX: the 318-408. species of Texas and the Southwest. Bull. Torrey Barneby, R. C. 1943. .Miscellaneous diagnoses. Leafl. W. Bot. Club 28: 570-592.

Bot. 3: 193-199. Blake, S. F. 1922. New Asteraceae from Utah and Ne- 1944. Alice Eastwood semi-centennial pub- vada. Proc. Biol. Soc. Washington 35: 173-177. Asteraceae. Wash- lications No. 3. Pugillus Astragalorum Alter. 1931. Nine new American J. Proc. Calif. Acad. IV, 25:147-169. ington Acad. Sci. 21: 325-336. 188 Great Basin Naturalist Vol. 42, No. 2

1935. Five new plants of the genus Aplopappus. 1909. Supplement to the monograph of the North Proc. Biol. Soc. Washington 48: 169-174. American Umbelliferae. Contr. U.S. Natl. Herb. 1937. Eleven new Asteraceae from North and 12: 441-451. Sci. South America. J. Washington Acad. 27: Coville, F. V. 1892. Descriptions of new plants from ,374-391. southern California, Utah and Arizona. Proc.

Blauer, H. C. 1965. Helianthus (Compositae) in Utali. Biol. Soc. Wa.shington 7: 65-80. Proc. Utah Acad. 42: 241-251. 1893. Botany of the Death Valley Expedition. Brand, A. 1907. Polemoniaceae. Pflanzenr. 4, Fam. 2.50 Contr. U.S.Natl. Herb. 4: 1-363. (Heft 27): 1-203. Cronquist, a. 1943. The North American species of 1911. Beitrag zur Kenntnis der Hydrophyllaceae. Erigeron centering about £. speciosus (Lindl.) Brandegee, M. K. 1900. Opitntias. Pages 97,110,120 in DC. and £. glabellus Nutt. Bull. Torrev Bot. A. Purpus, Club 70: 265-274. J. Die Succulenten der La Sal Moun- tains in Utah. Monats. Kakteenk. 10: 95-120. - 1947. Revision of the North American species of F., P. Erigeron, north of Mexico. Brittonia 6: 121-302. Brenckle, J. AND W. Cottam. 1940. A new Polyg- onum from Garfield County, Utah. Bull. Univ. 1963. Two new varieties in Machaeranthera. Utah 30: 2-7. Leafl. W. Bot. 10: llinl3. Cronquist, A., A. Holmgren, N. H. Britton, N. L. 1904. Four new North American birches. Holmgren, J. L. Bull. Torrey Bot. Club 31: 165-166. Reveal, and P. K. Holmgren. 1977. Inter- Britton, N. L., N. Rose. 1920. mountain Flora. Vol. 6. New York, Columbia and J. Cactaceae. Vol. 1 Univ. Press. 584 & 4, Carnegie Inst. Washington Publ. 248. 80 pp. pp. Darlington, 19.34. A monograph of the genus Britton, N. L., A. J. Men- and J. Shafer. 1908. North American tzelia. Ann. Missouri Bot. Garden 21: 103in226. Trees. New York. Henry Holt and Company. 894 Daston, S. 1946. Three noteworthy cacti of south- pp. J. western Utah. Amer. Midi. Naturalist 36: Broaddus, M. 1935. Biographical sketch of Marcus E. J. 661 111662. Jones. Contr. W. Bot. 18: 132. Dempster, L. T., and F. Ehre.vdorfer. 1965. Evolution 1952?. VI. Anna Elizabeth (Richardson) Jones, of the Galium miiltiflorum complex in western 1853-1916 (Anna). Pages 48-51 in A. Jones, J. North America— II. Critical taxonomic revision. Publius Virgilius and Lavinia (Burton) Jones and Brittonia 17: 289in334. their family- 1800- 1950. Published by the family; Durand, E. M. 1860. A sketch of the botanv of the basin without locality indicated. of the Great Salt Lake of Utah. Trans. Amer. Buchenau, F. G. P. 1890. Juncus canadensis var. ktin- Phil. Soc. 11: 155m 180. tzei. Bot. Jahrb. 12: 272. Earle, H. W. 1980. A new Opuntia identified in south-

Candolle, a. P. DE 1864. Quercus stellata var. ttta- western Utali. Saguaroland Bull. 34: 15. hensis. Prodromus Systematis naturalis Regni Eastwood, A. 1893. List of plants collected in south- vegetabilis. Vol. 16. Parisiis. 160 pp. eastern Utah, with notes and descriptions of new Chamberlain, R. V. 1950. Life Sciences at the Univer- species. Zoe 4: 113-127. sity of Utah. Background and History. Deseret 1895. Two species of Aquilegia from the upper Sonoran zone of Colorado and Utah. Proc. Calif. News Press; Salt Lake City. 417 pp. Acad. II, 4: 559-562 Chaudhri, M. N. 1968. Revision oi Paronychia. 194. 1896. Report on a collection of plants from San Chuang, T. I., and L. R. Heckard. 1973. Taxonomy of Juan County, in southeastern Utali. Proc. Calif. Cordylanthus subgenus Hemistegia (Scrophula- Acad. II, 6: 270-329. riaceae). Brittonia 25: 135-158. 1903. New species of Oreocarya. Bull. Torrey Clover, E. U. 1938. The Cactaceae of southern Utah. Bot. Club 30: 238-246. Bull. Torrey Bot. Club 65: 397-412, 1903. New species of western plants. Bull. Torrev Clover, E. U., and L. Jotter. 1941. Cacti of the canyon Bot. Club 30: 483-502. of the Colorado River and tributaries. Bull. Tor- 1904. Some new species of western Polemo- rey Bot. Club 68: 409-419. niaceae. Bot. Gaz. 37: 437-447. Cockerell, T. D. 1904. A North American species of 1927. New species of Ceanothus. Proc. Calif. Hymenoxys. Bull. Torrey Bot. Club 31: 461-509. Acad. 16: 361-363. CooLEY, E. L. 1980. Diary of Brigham Young 1857. Uni- 1933. Two new species of western Galium. Leafl. versity of Utah Library. 105 pp. W. Bot. 1: 55-56. Cottam, W. P. 1939. A new violet from Utah. Bull. 1936. A new Townsendia from Utah. Leafl. W. Univ. Utah Biol. Ser. 4 (3). 8 pp. Bot. 1: 206-207. Coulter, 1896. J. M. Preliminary revision of the North 1937. New species of Dodecatheon. Leafl. W. American species of Echinocactits, Cereiis and Bot. 2: 36-38. Opuntia. Contr. U.S. Natl. Herb. 3: 355-462. 1937. New species of western plants. Leafl. W.

Coulter, J. M., and A. Nelson. 1909. New manual of Bot. 2: 54-,56. botany of the central Rocky Mountains. New 1941. New species of CastiUeja. Leafl. W. Bot. 3: York. American Book Co. 646 pp. 87-92. Coulter, M., N. 1942. New western plants IV. Leafl. W. Bot. 3: J. and J. Rose 1888. Revi.sion of North American Umbelliferae. Crawfordsville. 144 pp. 157-160. 1900. Monograph of the North American Um- 1942. New species and varieties of Lupinus. belliferae. Contr. U.S. Natl. Herb. 7: 7-256. Leafl. W. Bot. 3: 169-192. June 1982 Welsh: Utah Plant Types 189

D. C. 1871. Compositae. Pages 136-208 in S. Eaton, Gould, F. W. 1942. A systematic treatment of the genus Botany. Report of U.S. Geological Ex- Watson, Camassia Lindl. Amer. Midi. Naturalist 28: ploration of the Fortieth Parallel. 5: Xiii-525. 712-742. Government Printing Office. Washington: Graham, E. H. 1937. Botanical studies in the Uinta Ba- 1875. parriji. Amer. Naturalist 9: 351. Notholeana sin of Utah and Colorado. Ann. Carnegie Mus. of the multi- Ehrendorfer, F. 1956. Survey Galium 26: 1-432. in western North America. florum complex Gray, A. 1849. Plantae Fendlerianae Novi-Mexicanae. Dudley Herb. 5: .3-21. Contr. Mem. Amer Acad. 4: 1-116. Engelmann, G. 1863. x\dditions to the cactus-flora of 1852. Plantae Wrightianae Texano-Neo-Mexi- the territory of the United States. Trans. Acad. canae. Part 1. Smithsonian Contr. Knowl. 3: Sci. St. Louis 2: 199. 5-146. Agave. Page 497 in S. Watson, 1871. Yucca and 1870. Revision of the North American Polemo- Botany. Report of U.S. Geological Exploration of niaceae. Proc. Amer. Acad. 8: 247-282. Parallel. Bot. 5: xiii-525. Washing- the Fortieth 1870. Miscellaneous botanical notes and charac- ton: Government Printing Office.. ters. Proc. Amer. Acad. 8: 282-296. Sabina. Trans. 1878. American junipers of section 1872. Botanical contributions. Proc. Amer. Acad. St. Louis Acad. Science 3: 588-589. 8: 365-412. 1896. Opuntia. Pages 42.3-24 in M. Coulter, J. 1873. Characters of new genera and species of revision of the North spe- Preliminary American plants. Proc. Amer. Acad. 8: 620-661. cies of Echinocactits, Cereits, and Opuntia. Contr. 1874. Notes on Compositae and characters of cer- U.S. Natl. Herb. 3: .357-462. tain genera and species. Proc. Amer. Acad. 9: Engler, H. G., and E. Irmscher. 1916. Saxifragaceae— 187-218. Saxifraga. Pflanzenr. 4. Fam. 117 (Heft 67 & 1874. Contributions to the botany of North 69); 1-707. America. Proc. Amer. Acad. 10: 39-78. 1950. Naturalists. The Uni- EwAN, J. Rocky Mountain 1875. A conspectus of the North American Hy- versitv of Denver Press. 358 pp. drophyllaceae. Proc. Amer. Acad. 10: 3I2-.332. Fedde, F. K. G. 1909. Platystemon. Pflanzenr. 4. Fam. 1876. Miscellaneous botanical contributions. 104. (Heft 40): 106-131. Proc. Amer. Acad. 11: 71-104. Fedde, F. K. 1913. Neue Arten aus der Verwandtschaft 1876. Contributions to the botany of North der Con/dalis aurea Willd. von Nord Amerika - America— V. Proc. Amer. Acad. 12: 51-84. VIII. Feddes Repert. 11: 497. 1876. Geological survey of California. Botany. Flous, F. 19.34. Diagnoses d' especes et varietes nou- Vol. I. ed. I. - velles de Pseudotsuga americains. Bull. Soc. Hist. 1878. Synoptical flora of North America. Vol. II Nat. Toulouse 66: 329-346. Part 1. New York: Ivison, Blakeman, Taylor and Co. 402 Flowers, S. 1949. A new Selaginella from southwestern pp. 1878. Contributions to the botany of North Utah. 39: 83-86. Amer. Fern J. America. Proc. Amer. Acad. 13: 361-374. 1845. of the exploring expedition Fre.mont, J. C. Report 1880. Geological .survey of California. Botany. to the Rocky Mountains in the year 1842, and to Vol. I. 628 pp. Oregon and north California in the years 1880. Contributions to North American botany. 184.3-1844. House Rep. Doc. 199. Proc. Amer. Acad. 16: 78-108. Gabel, M. L. 1981. The Parry herbarium. Proc. Iowa 1883. Contributions to North American botany. Acad. Sci. 88: 179. Proc. Amer. Acad. 19: 1-96. Gandoger, M. 1906. Le genre Eriogonum (Polyg- 1884. Synoptical flora of North America. Vol. I. onaceae). Bull. Soc. Bot. Belgium 42: 183-200. Part 2. New York; Ivison, Blakeman, Taylor, and 1918. Sertum plantarum novarum. Pars prima. Co. 473 pp. Bull. Soc. Bot. France 65: 24-69. 1885. A revision of some Boragineous genera. 1919. Sertum plantarum novarum. Bull. Soc. Bot. Proc. Amer. Acad. 20: 257-286. France 66: 216-233. 1886. Contributions to American botany. Proc. 1920. Sertum plantannn novarum. Bull. Soc. Bot. Amer. Acad. 13- .36.3-413. France 66: 286-,307. 1886. Synoptical flora of North America. Vol. I. Garrett, A. O. 1917. Spring flora of the Wasatch re- Ed. 2. Washington: Smithsonian Institution. 480 gion. 3rd Ed., Lancaster Press: Salt City. Lake pp. 144 pp. 1889. John Torrey. p. 359-369 in C. S. Sargent, Gates, R. R. 1915. The mutation factor in evolution, Scientific papers of Asa Gray. Vol. 2. Essays; Bio- with particular reference to Oenothera. London, graphical sketches. 1841-1886. Macmillan and MacMillan and Co. 353 pp. Co.; London. .503 pp. in species— XVII. Gentry, J. L. 1974. Studies the genus Hackelia (Bo- Greene, E. L. 1896. New or noteworthy raginaceae) in the western United States and Pittonia3; 91-116. Mexico. Southwestern Naturalist 19: 139-146. 1896. Studies in the Cruciferae-I. Pittonia 3; GooDDiNG, L. N. 1904. Southwestern plants. Bot. Gaz. 117-138. 37: 53-59. 1897. Some Rockv Mountain asters. Pittonia 3: 1932. sea- 212-224. Goodman, G. J. and C. L. Hitchcock. A new pose Draba from Utah. Ann. Missouri Bot. Card. 1899. New or noteworthy species—XXIV. Pit- 19: 77-78. tonia 4: 68-72. 190 Great Basin Naturalist Vol. 42, No. 2

1899. West American Asperifoliae— IV. Pittonia Heil, K. 1979. Three new species of Cactaceae from 4: 86-97. southeastern Utah. Cact. Succ. (U.S.) & J. 51: 1900. A fascicle of Senecios. Pittonia 4: 108-124. 25-30.

. 1900. A fascicle of new Arnicas. Pittonia 4: Heimerl, a. 1902. Abronia fallax. Page 684 in P. A. 159-174. Rydberg, Studies on the Rocky Mountain Flora— 1901. New or noteworthy species— XXVIII. Pit- IX. Bull. Torrey Bot. Club 29: 680-693. tonia 4: 2.3.3-241. Heiser, C. B. 1961. A new annual sunflower, Helianthus

. 1903. Pkitystemon and its allies. Pittonia 5: deserticolus, from the southwestern United 139-194. States. Proc. Indiana Acad. Sci. 70: 209-212. 1905. A proposed new genus, Annotites. Leafl. Heller, A. A. 1905. Western species, new and old— IV.

Bot. Obs. & Crit. 1: 97-112. Muhlenbergia 1: 111-118. 1905. Segregates of the genus Rhus. Leafl. Bot. Hermann, F. J. 1934. A new Cardamine from the Uinta Obs. & Crit. 1: 114-144. Mountains, Utah. Rhodora 36: 409-412. 1905. New species of Penstemon. Leafl. Bot. Obs. 1937. Extensions of range and a new species in & Crit. 1: 161-176. Carex. Rhodora 39: 492-495.

. 1905. Revision of Eschscholtzia. Pittonia 5: HiGGiNS, L. C. 1968. New species of perennial Cryp- 205-293. tantha from Utah. Great Basin Naturalist 28: 1905. Diagnoses Aragallorum. Proc. Biol. Soc. 195-198.

Washington 18: 11-18. Hitchcock, A. S. 1933. New species and new names of 1906. The genus Ptelea in the western and south- grasses Texas. Sci. from J. Washington Acad. 23: western United States and Mexico. Contr. LI.S. 449-456. Natl. Herb. 10: 49-78. Hitchcock, C. L. 1941. A revision of the Drabas of 1906. A study of Rhus glabra. Proc. Washington western North America. Univ. Washington Publ. Acad. 8: 167-196. Biol. 11: 1-120. 1908. Novitates borealis Americanae— II. Feddes 1950. On the subspecies of Lepidiiim montanum. Repert. 5: 241-244. Madrono 10: 155-158. 1909. New western Asteraceae. Leafl. Bot. Obs. 1952. A revision of the North American species of & Crit. 2: 8-13. Lathyrus. Univ. Washington Publ. Biol. 15: 1910. Some western caulescent violets. Leafl. 1-104. Bot. Obs. & Crit. 2: 32-34. Hitchcock, C. L., and B. Maguire. 1947. A revision of 1910. Miscellaneous specific types— I. Leafl. Bot. the North American species of Silene. Univ. 2: 45-48. Obs. & Crit. Washington Publ. Biol. 13: 1-73. 1910. western species of is. Some Arab Leafl. Bot. Holm, T. H. 1891. A study of some anatomical charac- Obs. & Crit. 2; 69-88. ters of North American Gramineae - III. Bot. 1910. Studies of Thalictraceae-II. Leafl. Bot. Gaz. 16: 275-281. Obs. & Crit. 2: 89-94. Holmgren, A. H., L. M. Shultz, and T. K. Lowrey. 1912. New species of Chaenactis. Leeafl. Bot. 1976. Sphaeromeria, a genus closer to Artemisia Obs. & Crit. 2: 221-225. than to Tanacetum (Asteraceae: Anthemideae). 1912. Novitates borealii Americanae— VI. Species Brittonia 28: 255-262. novae generis Cercidis. Feddes Repert. 11: Holmgren, N. H. 1973. Five new species of Castilleja 108-111. (Scrophulariaceae) from the Intermountain Re- Greenman, M. 1914. Descriptions of North American J. gion. Bull. Torrey Bot. Club 100: 83-93. Senecioneae. Ann. Missouri Bot. Card. 1: 1978. Three new species of Penstemon (Scro- 263-290. phulariaceae) from the Intermountain region. 1916. Monograph of the North and Central Brittonia 30: 416-425. American species of the genus Senecio. Part II. 1979. New Penstemons (Scrophulariaceae) from Ann. Missouri Bot. Card. 3: 85-194. Brittonia 31: 217-242. 1917. Monograph of the North and Central the Intermountain region. Holmgren, N. H., A. H. Holmgren. 1974. Three American species of the genus Senecio. Part II. and Ann. Missouri Bot. Card. 4: 15-36. new species from the Great Basin. Brittonia 26: 309-315. Greenman, J. M., and E. M. F. Roush. 1929. New Agaves from .southwestern United States. Ann. Holmgren N. H., and P. K. Holmgren. 1979. A new Missouri Bot. Card. 16: 389-392. species of Asclepias (Asclepiadaceae) from Utah. Hackel, E. 1896. Festuca dasijclada. in Brittonia 31: 110-114. Page 602 W. J. Beal, Grasses of North America Vol. 2. New York, Hopkins, M. 1937. Arabis in eastern and central North Henry Holt and Company. America. Rhodora 39: 106-148. Hall, H. M., and F. E. Clements. 1923. The phyloge- House, H. D. 1906. New and noteworthy North Ameri- netic method in Taxonomy. The North American can species of Trifolium. Bot. Gaz. 41: 3.34-347. 1940. Plants. Leafl. Bot. species of Artemisia, Chrij.sothaniniis, and Atri- Howell, J. T. New western W. plex. Carnegie Institution of Washington. 355 pp. 2: 2,53. L. T. K. 1943. Studies in Phacelia—a revision of species re- Hanks, and J. Small. 1907. Geraniaceae. N. Amer. Fl. 25: 3-24. lated to P. pulchella and P. rotundifolia. Amer. Hanson, C. A. 1962. New species of perennial Atriplex Midi. Naturalist 29: 1-26. Ives, 1860. the Colorado River of the from the western United States. Stud. Syst. Bot. J. C. Report upon (BRY) 1: 1-4. west. Washington D.C. C. Wendell, Printers. June 1982 Welsh: Utah Plant Types 191

Johnston, I. M. 1923. Diagnoses and notes relating to 1940. Studies in Penstemon— VW. The .subsections the spermatophytes chiefly of North America. Gairdneriani, Deusti, and Arenarii of the Gra- Contr. Gray Herb. II, 68: 80-104. ciles, and miscellaneous new species. Amer. Midi. 1937. Studies in the Boraginaceae— XII. .\rnold J. Naturalist 23: 594-616, .\rb. 18: 1-25. Klein, W. 1962. New taxa and recombinations in Oeno- 1939. Studies in the Boraginaceae— XIII. ]. .Ar- thera (Anogra). Aliso 5: 179-180. nold Arb. 20: 375-402. KoEHNE, B. A. E. 1890. Amelanchier utahensis. Wiss. 1948. Studies in the Boraginaceae— XVI. Species Bei. Progr. Falk-Realgym. Berl. 95: 25. chiefly from Mexico and western United States. J. KuNTZE, C. E. O. 1885. Monographic der Gattung Cle- Arnold Arb. 29: 227-241. matis. Verh. Bot. Ver. Brandenb. 26: 84-202. 1952. Studies in the Boraginaceae—XXII. Note- 1891-95. Revisio generimi plantarum. Vol. I & II. worthy species, chiefly Asian and South Ameri- Leipzig. 1011 pp. can. Arnold Arb. 33:' 62-78. J.' Leveille, H. 1908. Monographic du genre Oenethera. Jones, A. and M. Broaddus. 1952?. V. Marcus Eu- J., J. Bull. Acad. Internat. Geogr. Bot. 17: 257-332. (Mark). gene Jones, 1852-1934 In A. J. Jones, Macbride, J. F. 1916. Mertensias of western North Publius Virgilius and Lavinia (Burton) Jones and America. Contr. Gray Herb. II, 48: 1-20. their family-1800-1950. Published by the family; 1916. Revision of the genus Oreocarya. Contr. without locality designated. Gray Herb. II, 48: 20-38^ Jones, M. E. 1881. Notes from Utah. Bull. Torrev Bot. P., Macbride, J. and E. B. Payson. 1917. A revision of Club 8: 70-71. the Erigerons of the series multifida. Contr. Gray 1891. New plants from Arizona, Utah and Ne- Herb. II, 49:72-79. vada. Zoe 2: 12-17. Mackenzie, K. K. 1917. Carex epapillosa. Page 138 in P. 1891. New species and notes of Utah plants. Zoe A. Rydberg, Flora of the Rocky Mountains. New 236-252. 2: York: by the author. 1893. Contributions to western botanv. No. 3. Maguire, B. 1941. Great Basin plants— III. Caryophvl- Zoe 3: 283-309. laceae. Madrono 6: 22-27. 1893. Contributions to western botanv. No. 4. 1942. Great Basin plants— VII. Cruciferae. Amer. Zoe 4: 22-54. Midi. Naturalist 27: 463-469. 1893. Contributions to western botanv. No. 5. 1944. Great Basin plants— VIII. New species in Zoe 4: 254-282. Carex and Erigeron. Brittonia 5: 199-202. 1894. Contributions to western botanv. No. 6. 1946. Studies in the Caryophyllaceae— I. Bull. Zoe 4: 366-369. Torrey Bot. Club 73: 326. 1895. Contributions to western botany. No. 7. 1947. Great Basin plants— IX. Compositae. Ar- Proc. Calif. Acad. II, 5: 611-7.33. nica, Hymenopapptis, Haplopapptis, and Lijgo- 1898. Contributions to western botanv. No. 8. desmia. Amer. Midi. Naturalist 37: 136-145. Contr. W. Bot. 8: 1-43. 1947. Studies in the Caryophyllaceae— III. A sy- 1902. Contributions to western botanv. No. 10. nopsis of the North American species oi Arenaria. Contr. W. Bot. 10: 1-90. Bull. Torrey Bot. Club 74: 38-56. 1903. Contributions to western botanv. No. 11. 1951. Studies in the Caryophyllaceae— IV. Are- Contr. W. Bot. 11: 1-22. naria in America north of Mexico, a conspectus. 1908. Contributions to western botanv. No. 12. Amer. Midi. Naturalist 46: 493-511. Contr. W. Bot. 12: 1-97. Maguire, B. and A. Cronquist. 1947. Erigeron flag- 1910. New species and notes. Contr. \V. Bot. 13: ellaris var. trilobatus. Page 258 in A. Cronquist, 1-87. Revision of the North American species of Erige- 1912. Contributions to western botany. No. 14. ron. north of Mexico. Brittonia 6: 121-302. Contr. W. Bot. 14: 1-52. Maguire, B. and G. H. Jensen. 1942. Great Basin 1923. Revision of North .American species of As- plants— V. Aquatics. Rhodora 44: 5-10. tragalus. Salt Lake City; by the author. 288 pp. Maguire, B. and A. H. Holmgren. 1946. Botany of the 1929. Contributions to western botany. No. 15. Intermountain Region— I. Leafl. W. Bot. 4: Contr. W. Bot. 15: 1-163. 261-267. 1930. Contributions to western botany. No. 16. Maguire, B. and R. E. Woodson. 1941. Two new Ascle- Contr. W. Bot. 16: 1-53. piads from the southwestern United States. Ann. 1933. Contributions to western botanv. No. 18. Missouri Bot. Card. 28: 245. New species and notes. Contr. \V. Bot. 18: Marshall, W. T. 1954. Pediocactus hermannii sp. nov. 20-130. Saguaroland Bull. 8: 78-81. Kearney, T. H. 1898. Studies on grasses. Revi- American Martin, R. F. 1940. A new Juncus from Utah. Rhodora sion of North American species of Calamogrostis. 40: 69-70. 1-62. U.S.D.A Agrostol. Bull. 11: Mathias, M. E. 1930. Studies in Umbelliferae III. A Keck, D. D. 1934. A new Penstemon from Utali. Leafl. monograph of Cymopterus including a critical W. Bot. l:82-&3. study of related genera. Ann. Missouri Bot. Card.

1937. Studies in Penstemon— W . The section Eri- 17: 213-474.

copsis. Bull. Torrey Bot. Club 64: 357-381. 1932. Studies in the Umbelliferae. IV. .\ new spe-

1937. Studies in Penstemon— \ . The section Pel- cies of Cogswellia. Ann. Missouri Bot. Gard. 19: tanthera. Amer. Midi. Naturalist 18: 790-829. 497-498. 192 Great Basin Naturalist Vol. 42, No. 2

McKelvey, S. D. 1947. Yuccas of the southwestern B. S. 1899. Norton, J. North American species of Eu- United States. Part 2. Jamaica Plain; Arnold Ar- phorbia, section Tithymalus. Eleventh Annual boretum. 192 pp. Report. Missouri Bot. Gard. 144 pp. McVaugh, R. 1956. Edward Palmer. Plant Explorer of Osterhout, G. E. 1926. Oreocarya breviflora. Page 169 the American West. Norman: University of Okla- in E. B. Payson, Tlilaspi, Oreocarya, and Erige-

homa Press. 430 pp. ron. Univ. Publ. Bot. 1: 145-186. Wyoming . Mez, C. C. 1921. Stipa mormonum. Feddes Repert. 17: Ottley, a. M. 1944. The American Loti with special 209. consideration of a proposed new section, Sim- Morton, C. V. 1937. Plujsaria grahamii. Page 26 in E. peteria. Brittonia 5: 81-123. H. Graham, Botanical studies in the Uinta Basin Ownbey, G. B. 1958. Monograph of the genus Argemone of Utah and Colorado. Ann. Carnegie Mus. Vol. for North America and the West Indies. Mem. 26. Torrey Bot. Club 21(1): 1-159. MuNZ, p. A. 1928. Studies in Onagraceae— I. A revision Parry, C. C. 1875. Botanical observations in southern of the subgenus Chylismia of the genus Oeno- Utah in 1874. Amer. Naturalist 9: 14-21, thera. Amer. Bot. 15: 223-240. J. 139-146, 199-205, 267-273, 346-351. 1931. Studies in Onagraceae— VII. The subgenus 1878. Botanical Collection. Proc. Davenport of the Pachylophus genus Oenothera. Amer. J. Acad. Nat. Sci. 2: 279-282. Bot. 18: 728-738. Payson, E. B. 1915. New and noteworthy plants from 1965. Botanical exploration of Marcus E. Jones. southwestern Colorado. Bot. Gaz. 60: 374-382. Leafl. W. Bot. 10: 193-236. 1921. A monograph of the genus Lesquerella. Murray, E. 1980. Acer notes No. 9. Kalmia 10: 2. Ann. Missouri Bot. Gard. 8: 103-236.

S. 1981. Astragahts section Neese, E. J., AND L. Welsh. 1926. Thlaspi, Oreocarya, and Erigeron. Univ. Desperati and a new species from the Uinta Ba- Wyoming Publ. Bot. 1: 145-186. sin of Utah. Rhodora 83: 455-460. Pennell, F. W. 1920. Scrophulariaceae of the central Nelson, A. 1899. New plants from Wyoming— VII. Bull. Rocky Mountain states. Contr. U.S. Natl. Herb. Torrey Bot. Club 26: 236-250. 20: 313-381. 1900. Some Rocky Mountain Chrysothamni. Bot. 1933. A revision of Synthyris and Besseya. Acad. Gaz. 28: 369-377. Nat. Sci. Philadelphia 85: 77-106. 1900. Contributions from the Rocky Mountain 1935. The Scrophulariaceae of eastern temperate Acad. Nat. Sci. Philadelphia herbarium-I. Bot. Gaz. .30: 189-203. North America. 1902. Contributions from the Rocky Mountain Monogr. 1: 1-650. herbarium— IV. Chenopodiaceae. Bot. Gaz. 34: Petrak, F. 1917. Die nordamerikanischen Arten der 355-371. Gattung Cirsium. Beih. Bot. Centralbl. 35(2): 1904. Contributions from the Rocky Mountain 22.3-567 herbarium-V. Bot. Gaz. 37: 260-279. Pilger, R. K. F. 1922. Uber die Formen von Plantago 1904. New plants from Nevada. Proc. Biol. Soc. mapr L. Feddes Repert. 18: 277. Northwestern Washington 17: 91-98. Piper, C. V. 1899. New and noteworthy 1906. Contributions from the Rocky Mountain plants-II. Erythea 7: 159-163. herbarium VII. Bot. Gaz. 42: 48-54. Porter, C. L. 1952. Novelties in Hermidium (Nyctagi- 1907. Some new western plants and their collec- naceae) and Astragalus (Leguminosae) from eastern 54: 158-161. tors. Proc. Biol. Soc. Washington 20: 33-40. Utah. Rhodora 1912. Contributions from the Rocky Mountain Preston, C. H. 1897. Biographical sketch of Dr. C. C. Sci. 6: 35-45. herbarium— XI. New plants from Idaho. Bot. Parry. Proc. Davenport Acad. Nat. A. 1900. Die Succulenten der La Sal Moun- Gaz. 54: 1,38-151. PuRPUs, J. 95-97. 1913. Contributions from the Rocky Mountain tains in Utah. Monats. Kakteenk. 10: herbarium-XIII. Bot. Gaz. 56: 61-71. 1900. Die Succulenten der La Sal Mountains in Utah. Monats. Kakteenk. 10: 110-112. 1922. Flora of the Navajo Indian Reservation— II. 1909. Opuntia utahensis A. Purpus nov. spec. Amer. Botanist 28: 20-25. J. 1924. Taxonomic studies— miscellaneous new spe- Monats. Kakteenk. 19: 13.3-135. Raven, P. H. 1962. The systematics of Oenothera sub- cies. Univ. Wyoming Publ. Bot. 1: 63-68. Calif. Bot. 34: 1926. Miscellaneous new species. Univ. Wyoming genus Chylismia. Univ. Publ. 1-122. Publ. Sci. Bot. 1: 122-143. 1969. revision of the genus Camissonia (Ona- 1934. Rocky Mountain herbarium studies— II. A 161-396. 21: graceae). Contr. U.S. Natl. Herb. 37: Amer. J. Bot. 573-582. 1936. Rocky Mountain herbarium studies— IV. Rechinger, K. H. 1936. Rumex utahensis. Feddes Re- pert. 298. Bot'. 23: 265-271. 40: Amer. J. Reveal, L. 1965. Notes on three Utah Eriogonums. 1945. Rocky Mountain herbarium studies— VI. J. Proc. Acad. 42: 287-292. Bot'. .32: 284-290. Utah Amer. J. V. revision of the Nelson, A., F. 1916. Western plant 1967. Notes on £n'ogonum— A and J. Macbride. complex. Great Basin studies-Ill. Bot. Gaz. 61: 30-47. Eriogonum coryinbosum Nelson, E. 1899. Revision of the North American Naturalist 27: 183-229. 1968. A new annual Eriogonum from Utah. Ann. Phloxes. Laramie; by the author. 35 pp. 74-75. 1901. A revision of certain species of plants of the Missouri Bot. Gard. 55: species of from Utah. genus Antennaria. Proc. U.S. Natl. Museum 23: 1969. New Eriogonum 697-713. Madrono 19: 289-300. June 1982 Welsh: Utah Plant Types 193

1972. Descriptions of new species and com- 1903. Some generic segregations. Bull. Torrey binations in Eriogonutn (Polygonaceae). Phyto- Bot. Club 30: 271-281. logia 23: 163-178. 1904. Studies on the Rocky Mountain Flora— XII. 1973. Eriogonum (Polvgonaceae) of Utah. Phyto- Bull. Torrey Bot. Club 31:'555-575. logia 25: 169-217. 1904. Studies on the Rocky Mountain Flora— 1974. Two shnibby novelties in Eriogonum (Po- XIII. Bull. Torrey Bot. Club 31; 631-655. lygonaceae) from the deserts of Utali and Ari- 1905. Studies on the Rocky Mountain Flora— zona. Brittonia 26: 90-94. XIV. Bull. Torrey Bot. Club 32: 12.3-138.

1974. Two new varieties of Eriogonum (Polvg- 1905. Astragalus and its segregates as represented onaceae) from the Intermountain Region. Great in Colorado. Bull. Torrey Bot. Club 32; 657-668. Basin Naturalist 34: 245-246. 1907. Studies on the Rocky Mountain Flora—

1976. Eriogonum (Polygonaceae) of Arizona and XVII. Bull. Torrey Club .34: 35-50. New Mexico. Phytologia 35: 409-483. 1907. Studies on the Rocky Mountain Flora— 1977. Yucca angustissima var. avia. Inter- XVIII. Bull. Torrey Bot. Club 34: 417-437. mountain Flora 6: 534. 1908. Rosaceae-III & IV. N. Anier. Fl. 22; 1981. A new pulvinate Eriogonum (Polygonaceae) 239-387. from Utali. Great Basin Naturalist 41; 229-231. 1909. Studies on the Rocky Mountain Flora-XX. 1871. Potamogeton. Pages 247-340 in S. Bobbins, J. W. Bull. Torrey Bot. Club 36:'675-698. Watson, Botany. Report of U.S. Geological Ex- 1910. Studies on the Rocky Mountain Flora—

ploration of the Fortieth Parallel. 5: xiii-525. XXI. Bull. Torrey Bot. Club .37: 127-148. Robinson, B. L. 1917. A monograph of the genus Bri- 1910. Studies on the Rockv Mountain Flora—

ckelki. Mem. Gray Herb. 1: 1-151. XXII. Bull. Torrey Bot. Club 37; 31.3-3.35. L., 1899. Revision of 1910. Studies on the Rocky Mountain Flora— Robinson, B. and J. M. Greenm.\n. the genus Cymnolomia. Proc. Boston Soc. Nat. XXIII. Bull. Torrey Bot. Club 37; 44.3-471. Hist. 29: 87-100. 1910. Studies on the Rocky Mountain Flora— Rollins, R. C. 19.39. The Cruciferous genus Phij.saria. XXIV. Bull. Torrey Bot. Club 37: 541-557. Rhodora 41: .392-414. 1911. Studies on the Rocky Mountain Flora— 1940. Studies in the genus Hedijsarum in North XXV. Bull. Torrey Bot. Club 38: 11-23. America. Rliodora 42: 217-239. 1912. Studies on the Rocky Mountain Flora— .39; 99-111. 1941. Some new or noteworthy North American XXVI. Bull. Torrey Bot. Club 1912. Studies on the Rocky Mountain Flora— cnicifers. Contr. Dudley Herb. 3: 174-183. Bull. Bot. 39: .301-328. 1941. Monographic studv of Arabis in western XXVII. Torrey Club North America. Rliodora 43: 289-485. 1913. Studies on the Rocky Mountain Flora— 1955. The auriculate-leaved species of Les- XXVIII. Bull. Torrey Bot. Club 40; 4.3-74. Flora— querelki (Cruciferae). Rliodora 57: 241-264. 1913. Studies on the Rockv Mountain Bull. Bot. Club 40; 461-485. 1971. Protogyny in the Cruciferae and notes on XXIX. Torrey .389-479 Arabis and Caulanthus. Contr. Gray Herb. 201: 1913. Rosaceae-V. N. Anier. Fl. 22: 3-10. 1914. (Carduales) Carduaceae. Helenieae. N. .34: 1-80. 1981. Studies on Arabis (Cruciferae) of western Anier. Fl. Gaillardia. N. Fl. .34; 131-140. North America. Systematic Botany 6: 55-64. 1915. Amer. 1917. XI. Roses of California RosENDAL, C. O., F. K. Butters, and O. Lakela. 1936. Notes on Rosaceae— and Nevada. 44; 65-84. Heuchera versicolor f. pumila. Minnesota Studies 1917. Flora of the Rocky Mountains. New York; Plant Sci. 2: 85. author. 1143 Rydberg, p. a. 1897. Notes on Potentilla-VI. Bull. Tor- by the pp. Mountains, ed. 2. rey Bot. Club 24: 1-13. 1922. Flora of the Rocky New author. 1900. Catalogue of the flora of Montana and the York: by the 1143 pp. Bull. Torrey Bot. Yellowstone National Park. Mem. New York Bot. 1925. Notes on Fabaceae—V. Club 52; 229-235. Card. 1: i-492. 1927. Senecioneae. N. Amer. Fl. 34: .309-360. 1900. Studies on the Rocky Mountain Flora— I. (Rosales) Galageae. N. Amer. Fl. Bull. Torrey Bot Club 27: 169-189. 1929. Fabaceae, 24: 315-462. 1900. Studies on the Rocky Mountain Flora— II. Proc. International Bull. Torre\- Bot. Club 27: 614-636. 1929. Scylla or Charybdis. 1539-1551. 1900. Studies on the Rocky Mountain Flora-Ill. Congress Plant Sciences. 2: 1925. a L' etude des Festuca Some smaller genera of composites. Bull. Torrey St. -Yves, A. Contribution 1' Nord Bot. Club 27: 614-636. (Subgenus Eu-Festuca) de Amerique du 229-316. 1901. The oaks of the continental divide north of et du Mexico. Candollea 2; K. 1905. Beitrag zur Kenntnis der .\rten Mexico. Bull. New York Bot. Card. 2: 187-233. Schneider, C. Kunth. 1902. Studies on the Rockv Mountain Flora— VII. und Fornien der Gattung Cercocarpus Mitt. Gesells. 14; 125-129. Bull. Torrey Bot. Club 29:' 145-160. Deutsch. Dendrol. 1902. Studies on the Rockv Mountain Flora— 1905. Illustriertes Handbuch der Laubholzkunde. VIII. Bull. Torrey Bot. Club 29: 232-246. Vol. I. Jena. Part 2: 449-592. 1902. Studies on the Rockv Moimtain Flora— IX. ScHULZ, O. E. 1903. Monographie der gattung Carda- 417-623. Bull. Torrey Bot. Club 29: 680-693. mine. Bot. Jahrb. Syst. 32: 1903. Studies on the Rockv Moimtain Flora—X. 1924. Descurainia. Pflanzenr. 4, Fam. 105 (Heft 86); .304-346. Bull. Torrev Bot. Club .30; 247-262. 194 Great Basin Naturalist Vol. 42, No. 2

1927. Draba-Section XII. Phyllodraba. Pflan- 1914. A new Delphinium from Utah. Proc. Biol. zenr. 4, Fam. 105 (Heft 89): 173-198. Soc. Washington 27: 61. ScRiBNER, F. L. 1911. Notes on certain species of Muh- 1923. New or noteworthy species of plants from lenbergia. Bull. Torrey Bot. Club 38: 319-328. Utah and Nevada. Proc. Biol. Soc. Washington ScRiBNER, F. L. AND T. A. WiLLiAMs. 1899. Ncw spccics 36: 181-184. of North American Grasses. U.S.D.A. Agrostol. Torrey, 1852. J. Catalogue of plants collected in the ex- Circ. 9. 7 pp. pedition. Pages 381-397 in H. Stansbury, Explor- Sharp, 1935. A critical study of certain epappose genera ation and survey of the valley of the Great Salt of the Heliantheae-Verbesininae of the natural Lake of Utah. Philadelphia. Lippincott, Grambo family Compositae. Ann. Missouri Bot. Card. 22: and Co. 51-152. 1871. Species of Fraxinus, and Phacelia. Pages Sheldon, E. P. 1894. A preliminary list of the North 251,253,283. in S. Watson. Botany. Report of U.S. American species of Astragalus. Bull. Geol. & Geological Survey of the Fortieth Parallel. 5: Nat. Hist. Surv. Minn. 9: 116-175. xiii-525. Shultz, L. M. and Shultz. 1981. A new species of J. Torrey, J., and J. C. Fremont. 1845. Pages 311-320 hi Hackelia (Boraginaceae) from Utah. Brittonia 33: C. Fremont, Descriptions of some new genera 159-161. J. and species of plants, collected in Capt. J. C. Slosson, M. 1914. Notes on two North American ferns. Fremont's exploring expedition to Oregon and Bull. Torrey Bot. Club 41: 307-309. north California in the years 1843-44. House Small, K. 1896. Oenothera and its segregates. Bidl. J. Rep. Doc. 199. Bot. Torrey Club 23: 167-194. Torrey, A. J., and Gray. 1855. Report on the botanv of 1898. Studies in North American Polygonaceae. the expedition. Pages 115-132 in E. G. Beckwith, Bull. Bot. 40-53. Torrey Club 25: Report of explorations for a route for the Pacific 1905. Hydrangeaceae. N. Amer. Fl. 22: 159-178. Railroad, on the line of the forty-first parallel of Small, K. P. A. J. and Rydberg. 1905. Saxifragaceae. N. north latitude. Reports of explorations and sur- Amer. Fl. 22: 81-158. veys, to ascertain the most practicable and eco- Smiley, F. 1916. Mertensia praecox. 10 J. Page in J. F. nomical route for a railroad from the Mississippi Macbride, Mertensias of western North America. Roiover to the Pacific Ocean. 334d Cong., 2nd Contr. Gray Herb. II, 48: 1-20. Sess., House Rep. Ex. Doc. Vol. II, no 91. Smith, C. P. 1912. Notes from northern Utah-IV. Muh- 1870. A revision of the Eriogonea. Proc. Amer.

lenbergia 7: 136-138. Acad. 8: 145-200. 1948. Lupinus. Sp. Lupinorum 36: 625-640. Trelease, W. O. 1902. The Yucceae. Rep. Missouri Bot. 1949. Lupinus. Sp. Lupinorum 38: 657-672. Card. 13: 27-133. 1951. Lupinus. Sp. Lupinorum 40: 689-704. 1907. Additions to the genus Yucca. Ann. Mis- Smith, souri Bot. Card. 18: 225-230. J. G. 1899. Studies on American grasses: synopsis TuzsoN, 1921. Arahis f. laeiis. of genus Sitanion. U.S.D.A. Agrostol. Bull. 18: J. hirsuta Bericht Freien 1-21. Verein. Pflanzengeogr. Syst. Bot. 1919: 43. 1921. L. 1898. Selaginella its Standley, p. C. 1916. Chenopodiales, Chenopodiaceae. Underwood, M. rupestris and allies. Bull. Torrey Bot. Club 25: 125-133. N. Amer. Fl. 21: 1-93. Vasey, G. 1876. A catalogue of the forest trees of the Stansbury, H. 1852. Exploration and survey of the val- United States which usually attain a height of six- ley of the Great Salt Lake of Utah. Philadelphia. teen feet or more. Washington. Govt. Printing Lippincott, Grambo and Co. Office. St. John, H. 1915. Rumex persicarioides and its allies in 1893. Descriptions of new or noteworthy grasses North America. Rhodora 17: 73-83. from the United States. Contr. U.S. Natl. Herb. Stockwell, p. 1940. A revision of the genus Chaenactis. 1: 267-281. Contr. Dudley Herbarium 3: 89-168. Voss, J. W. 1937. A revision of the Phacelia crenulata Stokes, S. G. 1936. The genus Eriogonum, a preliminary group for North America. Bull. Torrey Bot. Club study based on geographical distribution. San 64: 133-144. Francisco. H. Neblett Press. J. Wagner, W. L. 1981. Oenothera acutissima (Ona- SuKSDORF, W. N. 1931. Untersuchungen in der gattung graceae), a new species from northwestern Colo- Amsinkia. Werdenda 1: 47-113. rado and adjacent Utah. Systematic Botanv 6: Swallen, R. 1931. Crassipes, a J. new grass genus from 153-158. Utah. Amer. Bot. 18: 684-685. J. Ward, L. F. 1876. General notes. I—botanv. Amer. Nat- Thurber, G. 1871. Vilfa depauperata var. filiformis. uralist 10: 552-555. Page 376 in: S. Watson, Botany. Report of U.S. Watson, E. E. 1929. Contributions to a monograph of Geological Survey of the Fortieth Parallel. 5: the genus Helianthus. Papers Michigan Acad. Sci. xiii-525. 9: 305-475.

Tidestrom, I. 1910. Species of Aquilegia growing in Watson, S. 1871. Botany. In C. King. Report of the Utah and in adjacent portions of Colorado, Idaho Geological Exploration of the Fortieth Parallel. 5:

and Arizona. Amer. Midi. Naturalist 1: 165-196. xiii-525. Washington: Government Printing 1913. A new Salicornia. Proc. Biol. Soc. Washing- Office. ton 26: 13. 1871. Descriptions of new species of plants, with 1913. Novitates florae utahensis. Proc. Biol. Soc. revisions of certain genera. Proc. Amer. Acad. 12: Washington 26: 121-122. 246-278. June 1982 Welsh: Utah Plant Types 195

1873. Revision of the extra-tropical North Ameri- 1978. Endangered and threatened plants of Utah: can species of the genus Lupintis. Proc. Ainer. A reevaluation. Great Basin Naturalist 38: 1-18. Acad. 8: 517-548. 1978. Utah flora: Fabaceae (Leguminosae). Great 1873. Revision of the extra-tropical North Ameri- Basin Naturalist 38: 225-.367. can species of the genus PotcntiUa. Proc. .4mer. 1980. Utah flora: Malvaceae. Great Basin Natu- Acad. 8: 549-573. ralist 40: 27-37. 1873. New plants of northern Arizona and the re- 1981. New taxa of western plants— in tribute. gion adjacent. Amer. Naturalist 7; 299-303. Brittonia .33: 294-.303. 1873. Revision of the extra-tropical North Ameri- 1982. Utah Flora: Rosaceae. Great Basin Natural- can species of the genus Oenothera. Proc. Amer. ist (in preparation).

Acad. 8: 573-618. Welsh, S. L., and N. D. Atwood. 1977. An undescribed 1874. A revision of the North American Che- species of Thehjpodiopsis (Brassicaceae) from the nopodiaceae. Proc. Amer. Acad. 9: 82-126. Uinta Basin, Utah. Great Basin Naturalist 37: 1874. Contributions to the botany of North 95-96. America— characters of various new species. Proc. Welsh, S. L., and R. C. Barneby. 1981. Astragalus len- Amer. Acad. 10: 68-78. tiginosus (Fabaceae) revisited—a unique new va- 1875. Revision of the genus oCeanothus, and de- riety. Isleya 2: 1-2. 1981.

scription of new plants. Proc. Amer. Acad. 10: Welsh, S. L., and S. Goodrich. 1980. Miscellaneous 333-350. plant novelties from Alaska, Nevada, and Utah.

1876. Botanical contributions— V^I. Proc. Amer. Great Basin Naturalist 40: 78-88.

Acad. 11: 105-148. Welsh, S. L., and G. Moore. 1968. Plants of Natural 1876. Geological Survev of California. Botanv. Bridges National Monument. Proc. Utah Acad.

Vol. 1. ed. 1. 45: 220-248.

1877. of species of plants, with S. L., L. 1968. species of Descriptions new Welsh, and J. Reveal. A new revisions of certain genera. Proc. Amer. Acad. 21: Townsendia (Compositae) from Utah. Brittonia 246-278. 20: 375-377. 1878. Bibliographical index to North American 1977. Utah Flora: Brassicaceae (Cruciferae). botany. Washington, D.C. Smithsonian In- Great Basin Naturalist 37: 279-365.

S. L., stitution. 476 pp. Welsh, N. D. Atwood, and J. L. Reveal. 1975. 1879. Contributions to American botanv. Proc. Endangered, threatened, extinct, endemic, and Amer. Acad. 14: 213-303. rare or restricted Utah vascular plants. Great Ba-

1888. Contributions to American botanv. Proc. sin Naturali.st .35: 327-374. Amer. Acad. 23: 249-287. Wheeler, G. M. 1878. Report upon United States geo- 1977. of Xijlorhiza (As- graphical surveys west of the one hundredth me- Watson, T. J., Jr. The taxonomy teraceae— Astereae). Brittonia 29: 199-216. ridian. Vol. VI. Washington D.C. Government Weber, W. A. 1946. A taxonomic and cytological study Printing Office. 404 pp. of the genus Wijethia, family Compositae, with Wherry, E. T. 1943. Microsteris, Phlox, and an inter- 60-63. notes on the related genus Balsomorhiza. Amer. mediate. Brittonia 5: 1944. New Phloxes from the Rocky Mountains Midi. Naturalist .35: 400-452. and neighboring regions. Not. Nat. 146: 1-11. Welsh, S. L. 1970. An undescribed species of Astragalus White, T. G. 1894. A preliminary revision of the genus (Legimiinosae) from Utali. Rhodora 72: 189-193. Lathijrus in North and Central America. Bull. 1970. New and unusual plants from Utah. Great Torrey Bot. Club 21: 444-458. Basin Naturalist .30: 16-22. Williams, L. O. 1932. Field and herbarium studies— I. 1971. Description of a new species of Dalea (Le- Bull. Torrey Bot. Club 59: 427-429. Naturalist 31: giuninosae) from Utah. Great Basin 1936. Revision of the western Primulas. Amer. 90-92. Midi. Naturalist 17: 741-748. 1974. Utah plant novelties in Astragalus and Williams, T. A. 1899. Poa fendleriana and its allies. Yucca. Great Basin Naturalist .34: .305-310. U.S.D.A. Agrostol. Circ. 10. 6 pp. 1976. Utali plant novelties in Cymopterus and YuNCKER, T. G. 1960. Two new species of Cuscuta from Penstemon. Great Basin Naturalist 35: 377-378. North America. Brittonia 12: 38-40.