Hybrid Speciation with External Barriers: Encelia
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December 2012 Number 1
Calochortiana December 2012 Number 1 December 2012 Number 1 CONTENTS Proceedings of the Fifth South- western Rare and Endangered Plant Conference Calochortiana, a new publication of the Utah Native Plant Society . 3 The Fifth Southwestern Rare and En- dangered Plant Conference, Salt Lake City, Utah, March 2009 . 3 Abstracts of presentations and posters not submitted for the proceedings . 4 Southwestern cienegas: Rare habitats for endangered wetland plants. Robert Sivinski . 17 A new look at ranking plant rarity for conservation purposes, with an em- phasis on the flora of the American Southwest. John R. Spence . 25 The contribution of Cedar Breaks Na- tional Monument to the conservation of vascular plant diversity in Utah. Walter Fertig and Douglas N. Rey- nolds . 35 Studying the seed bank dynamics of rare plants. Susan Meyer . 46 East meets west: Rare desert Alliums in Arizona. John L. Anderson . 56 Calochortus nuttallii (Sego lily), Spatial patterns of endemic plant spe- state flower of Utah. By Kaye cies of the Colorado Plateau. Crystal Thorne. Krause . 63 Continued on page 2 Copyright 2012 Utah Native Plant Society. All Rights Reserved. Utah Native Plant Society Utah Native Plant Society, PO Box 520041, Salt Lake Copyright 2012 Utah Native Plant Society. All Rights City, Utah, 84152-0041. www.unps.org Reserved. Calochortiana is a publication of the Utah Native Plant Society, a 501(c)(3) not-for-profit organi- Editor: Walter Fertig ([email protected]), zation dedicated to conserving and promoting steward- Editorial Committee: Walter Fertig, Mindy Wheeler, ship of our native plants. Leila Shultz, and Susan Meyer CONTENTS, continued Biogeography of rare plants of the Ash Meadows National Wildlife Refuge, Nevada. -
KERN RIVER PARKWAY PLANT LIST (Only Plant Species Permitted for Projects Within the Kern River Parkway Area - Includes Streetscape and Parking Lots)
KERN RIVER PARKWAY PLANT LIST (only plant species permitted for projects within the Kern River Parkway area - includes streetscape and parking lots) Scientific Name Common Name Type Acer macrophyllum Bigleaf maple Large tree Acer negundo ssp, californicum California box elder Large tree Aesculus californica California buckeye Large tree Alnus rhombifolia White alder Large tree Amelanchier pallida Western service berry Shrub or small tree Artemisia californica Coastal sage Shrub or small tree Artostaphlos densiflora Manzanita Shrub or small tree Artostaphlos glauca Manzanita Shrub or small tree Artostaphlos manzanita Manzanita Shrub or small tree Artostaphlos parryi Manzanita Shrub or small tree Atriplex lentiformis Quailbush Shrub or small tree Baccharis glutinosa Mulefat Shrub or small tree Baccharis pilularis "Twin Peaks" Dwarf coyote bush Flowering herb or groundcover Baccharis pilularis ssp. consanquinea Coyote bush Shrub or small tree Calycanthus occidentalis Western spice bush Shrub or small tree Carpenteria californica Tree anemone Shrub or small tree Castanopsis spp. Chiquapin Shrub or small tree Ceanothus cunneatus Ceanothus Shrub or small tree Ceanothus gloriosos Navarro ceanothus Flowering herb or groundcover Ceanothus griseus Carmel creeper Flowering herb or groundcover Ceanothus integerrimus Ceanothus Shrub or small tree Ceanothus leucodermis Ceanothus Shrub or small tree Ceanothus purpureus Ceanothus Shrub or small tree Ceanothus thrysiflorus Blue blossom Shrub or small tree Ceanothus thrysiflorus Ceanothus Shrub or small -
Illinois Bundleflower (Desmanthus Illinoensis) Story by Alan Shadow, Manager USDA-NRCS East Texas Plant Materials Center Nacogdoches, Texas
Helping People Help The Land September/October 2011 Issue No. 11 The Reverchon Naturalist Recognizing the work of French botanist Julien Reverchon, who began collecting throughout the North Central Texas area in 1876, and all the botanists/naturalists who have followed ... Drought, Heat and Native Trees ranging from simple things like more extensive root systems, to more drastic measures like pre- Story by Bruce Kreitler mature defoliation, what they actually have little Abilene, Texas defense against is a very prolonged period of no appreciable water supply. nybody that has traveled in Texas this year A will have noticed that not only most of the By the way, even though they are usually the land browned out, but also if you look at the trees same species, there is a difference in landscape in the fields and beside the roads, they aren't trees and native trees, which are untended plants looking so good either. It doesn't take a rocket that have to fend for themselves. While they are scientist to realize that extreme high temperatures indeed the same basic trees, the differences be- combined with, and partially caused by, drought tween the environments that they live in are huge are hard on trees. and thus overall general environmental factors such as drought, temperature, and insect infesta- Since I'm pretty sure that most of the people read- tions act on them differently. For the purposes of ing this article understand very well that drought this article, I'm referring to trees that are on their is a problem for trees, the question isn't is the pre- own, untended for their entire lives in fields, pas- sent drought going to have an effect on trees, but tures, forests, or just wherever nature has placed rather, what are the present effects of the drought them and refer to them as native trees. -
Flor De Rocío (Encelia Farinosa) Nombres Comunes: Hierba Ceniza (Cora) / Incienso, Palo Blanco (Español) / Hierba De Las Ánimas, Rama Blanca (ND) / Cotz (Seri)
Flor de rocío (Encelia farinosa) Nombres comunes: Hierba ceniza (Cora) / Incienso, Palo blanco (Español) / Hierba de las ánimas, Rama blanca (ND) / Cotz (Seri) ¿Tienes alguna duda, sugerencia o corrección acerca de este taxón? Envíanosla y con gusto la atenderemos. Foto: (c) Florian Boyd, algunos derechos reservados (CC BY-SA) Ver todas las fotos etiquetadas con Encelia farinosa en Banco de Imagénes » Descripción de EOL Ver en EOL (inglés) → Taxon biology Encelia farinosa has a bioregional distribution that includes California's eastern South Coast and adjacent Peninsular Ranges, as well as a desert distribution outside California to southwestern Utah, Arizona and northwest Mexico. The occurrences are restricted to elevations less than 1000 meters. Chief habitats are in coastal scrub and on stony desert hillsides. This desert shrub, also known by the common name Brittlebush, reaches a height of 30 to 150 centimeters, manifesting a single or several trunks. The stems are much-branched above, with young stems tomentose; older stems exhibit smooth bark, This plant's sap is fragrant Leaves are clustered near stem tips, with leaf petioles 10 to 20 millimeters in length, and with ovate to lanceolate blades ranging from two to seven cm. These tomentose leaves are silver or gray in color. Inflorescence heads are radiate, and generally yellowish, although the disk flowers can be yellow or brownish-purple. National distribution 1 United States Origin : Unknown/Undetermined Regularity: Regularly occurring Currently: Unknown/Undetermined Confidence: Confident Description 2 Brittle bush is a native, drought-deciduous, perennial shrub [7,8,21,28]. It grows to about 5 feet (1.5 m). -
March 6, 2015
Theodore Payne Foundation, a non-profit plant nursery, seed source, book store, and education center dedicated to the preservation of wild flowers and California native plants. This a report for March 6, 2015. New reports will be posted each Friday through the end of May. Again this year, we are at below normal rainfall and time is running out for significant rain events. Snow in the high elevations and high desert regions may contribute to a nice bloom, but not until later in the spring. The drought makes seeking out wild flower beauties more challenging but a lot more fun for Southern Californians to get out there a cheer every flower sighting! Most of the reports this week are out of the desert regions as they have been experiencing an early spring. This is not a banner year in Death Valley National Park, but it is still lovely. The southern part of Badwater Road, is a good spot to enjoy vast carpets of sand verbena (Abronia villosa). Not only a feast for the eyes, the sand verbena is also one of the most fragrant plants in Death Valley. The extremely hot weather earlier this month did do some damage to early blooming species, but recent small rain episodes may save the wild flower season, especially in the higher elevations. Scotty’s Castle Road is looking good, with some brown eyed evening primrose (Chylismia claaviformis), and really nice patches of notch-leaf phacelia (Phacelia crenulata), golden evening primrose (Chylismia brevipes), and brittlebush (Encelia farinosa). Further south on the Badwater Road, the areas between Mile Marker 40 and 48 there are delightful displays of desert sunflower (Geraea canescens) along with the sand verbena. -
Vegetation Classification for the Mojave Desert Inventory and Monitoring Network of National Parks
Vegetation classification for the Mojave Desert Inventory and Monitoring Network of National Parks Julie Evens (1), Keith A. Schulz (2), Marion S. Reid (2), Jaime Ratchford (1) and Kendra Sikes (1) (1) Vegetation Program, California Native Plant Society, Sacramento, CA; contact: [email protected], (2) Ecology, NatureServe, Boulder, CO Background/Question/Methods: Through a collaboration with the National Park Service, CNPS and NatureServe ecologists compiled over 9,000 new and existing vegetation surveys and analyzed over 4,000 surveys from three parks and other areas in the Mojave Desert and related ecoregions. A vegetation classification was developed, identifying approximately 105 alliances and related types from Lake Mead National Recreation Area (NRA), Mojave National Preserve, and Death Valley National Park within the Mojave Desert Inventory & Monitoring Network Inventory & Monitoring (MOJN I&M). This project’s data analyses of three park areas spanned more than 4 million acres of area, plus enabled cross‐analyses with other parks and preserves such as Joshua Tree National Park in California and Red Rock Canyon National Conservation Area in Nevada. We used classification and ordination methods such as agglomerative cluster analysis, indicator species analysis, and Nonmetric Multidimensional Scaling (NMS) to inform the vegetation classification of the park areas (Evens et al. 2017*). *The project classification report is in peer review; vegetation mapping is in progress through 2019, see Salas et al. (2016) for the mapping report for Lake Mead NRA. This project is supported by the National Park Service’s Vegetation Mapping Inventory Program and MOJN I&M, with technical assistance by Karl Brown, Tammy Cook, and others. -
Vascular Flora of West Clear Creek Wilderness, Coconino and Yavapai
VASCULAR FLORA OF WEST CLEAR CREEK WILDERNESS, COCONINO AND YAVAPAI COUNTIES, ARIZONA By Wendy C. McBride A Thesis Submitted in Partial Fulfillment of the Requirements for the Degree of Master of Science in Biology Northern Arizona University May 2016 Approved: Tina J. Ayers, Ph.D., Chair Randall W. Scott, Ph.D. Liza M. Holeski, Ph.D. ABSTRACT VASCULAR FLORA OF WEST CLEAR CREEK WILDERNESS, COCONINO AND YAVAPAI COUNTIES, ARIZONA WENDY C. MCBRIDE West Clear Creek Wilderness bisects the Mogollon Rim in Arizona, and is nested between the Colorado Plateau and Basin and Range physiographic provinces. Between 2013 and 2016, a floristic inventory vouchered 542 taxa and reviewed 428 previous collections to produce a total plant inventory of 594 taxa from 93 families and 332 genera. The most species rich families Were Asteraceae, Poaceae, Fabaceae, Brassicaceae, Rosaceae, Plantaginaceae, Cyperaceae, and Polygonaceae. Carex, Erigeron, Bromus, Muhlenbergia, and Oenothera Were the most represented genera. Nonnative taxa accounted for seven percent of the total flora. Stachys albens was vouchered as a new state record for Arizona. New county records include Graptopetalum rusbyi (Coconino), Pseudognaphalium pringlei (Coconino), Phaseolus pedicellatus var. grayanus (Coconino), and Quercus rugosa (Coconino and Yavapai). This study quantified and contrasted native species diversity in canyon versus non- canyon floras across the Southwest. Analyses based on eighteen floras indicate that those centered about a major canyon feature shoW greater diversity than non-canyon floras. Regression models revealed that presence of a canyon Was a better predictor of similarity between floras than was the distance betWeen them. This study documents the remarkable diversity found Within canyon systems and the critical, yet varied, habitat they provide in the southwestern U.S. -
1999 Comparative Morphology of Disk Floret Trichomes Of
COMPARATIVE MORPHOLOGY OF DISK FLORET TRICHOMES OF ENCELIA (ASTERACEAE: HELIANTHEAE) A Thesis Presented to the Faculty of California State Polytechnic University, Pomona In partial fulfillment of the Requirements for the Degree Master of Science In Biological Sciences By Kevin Joseph Carpenter 1999 SIGNATURE PAGE THESIS: COMPARATIVE MORPHOLOGY OF DISK FLORET TRICHOMES OF ENCELIA (ASTERACEAE: HELIANTHEAE) AUTHOR: Kevin Joseph Carpenter DATE SUBMITTED: Department of Biological Sciences Dr. Curtis Clark Thesis Committee Chair Biological Sciences Dr. Gary Carlton Biological Sciences Dr. Mark Porter ii ACKNOWLEDGEMENTS I would like to thank the following for their help with my thesis and research, and for the great amount they have contributed to my education: My thesis committee members, Drs. Curtis Clark (committee chair), Gary Carlton, and Mark Porter (Rancho Santa Ana Botanic Garden) The faculty, staff, and graduate students of Rancho Santa Ana Botanic Garden, including Drs. Travis Columbus, Elizabeth Friar, and Roy Taylor, as well as Vanessa Ashworth Mike Kinney, Mike McMillin, Rashmi Pant, Eric Roalson, Victor Steinmann, and others. Others including my wife and parents, as well as Drs. Jonathan Baskin, David Edmonds, Don Fosket, Jim Doyle, Daryl Koutnik, David Moriarty, Brian McNamara, and Camm Swift. iii ABSTRACT Disk florets of 19 Encelia taxa were examined with scanning electron microscopy to characterize disk floret trichome complement, density, and distribution on anthers, abaxial corolla lobes, and corolla tubes, to interpret their evolution in light of the phylogeny and ecology of the species, and to determine the utility of these characters for phylogenetic analysis and species delimitation. Trichomes are all multicellular, and include biseriate glands, biseriate achene hairs (Zwillingshaare), and narrow unicellular- based, straight uniseriates. -
Annotated Checklist of the Vascular Plant Flora of Grand Canyon-Parashant National Monument Phase II Report
Annotated Checklist of the Vascular Plant Flora of Grand Canyon-Parashant National Monument Phase II Report By Dr. Terri Hildebrand Southern Utah University, Cedar City, UT and Dr. Walter Fertig Moenave Botanical Consulting, Kanab, UT Colorado Plateau Cooperative Ecosystems Studies Unit Agreement # H1200-09-0005 1 May 2012 Prepared for Grand Canyon-Parashant National Monument Southern Utah University National Park Service Mojave Network TABLE OF CONTENTS Page # Introduction . 4 Study Area . 6 History and Setting . 6 Geology and Associated Ecoregions . 6 Soils and Climate . 7 Vegetation . 10 Previous Botanical Studies . 11 Methods . 17 Results . 21 Discussion . 28 Conclusions . 32 Acknowledgments . 33 Literature Cited . 34 Figures Figure 1. Location of Grand Canyon-Parashant National Monument in northern Arizona . 5 Figure 2. Ecoregions and 2010-2011 collection sites in Grand Canyon-Parashant National Monument in northern Arizona . 8 Figure 3. Soil types and 2010-2011 collection sites in Grand Canyon-Parashant National Monument in northern Arizona . 9 Figure 4. Increase in the number of plant taxa confirmed as present in Grand Canyon- Parashant National Monument by decade, 1900-2011 . 13 Figure 5. Southern Utah University students enrolled in the 2010 Plant Anatomy and Diversity course that collected during the 30 August 2010 experiential learning event . 18 Figure 6. 2010-2011 collection sites and transportation routes in Grand Canyon-Parashant National Monument in northern Arizona . 22 2 TABLE OF CONTENTS Page # Tables Table 1. Chronology of plant-collecting efforts at Grand Canyon-Parashant National Monument . 14 Table 2. Data fields in the annotated checklist of the flora of Grand Canyon-Parashant National Monument (Appendices A, B, C, and D) . -
Encelia Farinosa A. Gray Ex Torr NRCS CODE: Tribe: Heliantheae ENFA Family: Asteraceae Order: Asterales Subclass: Asteridae Class: Magnoliopsida
SPECIES Encelia farinosa A. Gray ex Torr NRCS CODE: Tribe: Heliantheae ENFA Family: Asteraceae Order: Asterales Subclass: Asteridae Class: Magnoliopsida juvenile plant, 3/3/2010, Riverside Co. A. Montalvo , 2003-2010, Riverside Co. Subspecific taxa None currently accepted (JepsonOnline 2nd Ed. 2010). Synonyms Encelia farinosa Torry & A. Gray corrected to current authorship (JepsonOnline) ENFAF Encelia farinosa A. Gray ex Torr. var. farinosa brittlebush ENFAP Encelia farinosa A. Gray ex Torr. var. phenicodonta (S.F. Blake) I.M. Johnst. brittlebush ENFAR Encelia farinosa A. Gray ex Torr. var. radians Brandegee ex S.F. Blake Common name brittlebush Also: brittle bush, brittle-bush, brittlebush encelia, incienso, incienso brittlebush, common brittlebush, white brittlebush; brown-center brittlebush for what was considered to be variety phenocodonta ; incienso is used for taxa in multiple families and desert encelia is also used for other species of Encelia (Painter 2009). Taxonomic relationships There are seven other species of Encelia in North America plus additional species in South America (JepsonOnline 2010). The genus Encelia is in the large tribe Heliantheae which also contains the native sunflower, Helianthus annuus L. Phylogenetic studies based on DNA show that Enceliopsis and Geraea are closely related genera and that diversification of species of Encelia has been quite recent (Fehlberg & Ranker 2007). Relationships among the various species of Encelia confirmed hypotheses by Clark (1998) based on chemical and morphological traits. Related taxa in region Four species and spontaneous hybrids of Encelia occur in southern California and may overlap with E. farinosa in some part of its range (FNA 2010, JepsonOnline 2010): E. actoni Elmer (in the southern Sierra Nevada, Tehachapi and Western Transverse Ranges, San Gabriel and San Bernardino Mountains, Mojave and Sonoran Deserts, and Desert Mountains; overlaps with E. -
A 30-Year Chronosequence of Burned Areas in Arizona— Effects of Wildfires on Vegetation in Sonoran Desert Tortoise (Gopherus Morafkai) Habitats
Prepared in cooperation with the Bureau of Land Management A 30-Year Chronosequence of Burned Areas in Arizona— Effects of Wildfires on Vegetation in Sonoran Desert Tortoise (Gopherus morafkai) Habitats Open-File Report 2015-1060 U.S. Department of the Interior U.S. Geological Survey Cover: Photograph showing burned habitat for the Sonoran Desert Tortoise (Gopherus morafkai) within the perimeter of the Gost Fire, Maricopa County, Arizona. This site burned in 2005 and is representative of burned areas in the Arizona Upland subdivision of the Sonoran Desert. The burned saguaro cactus (Carnegiea gigantea) is evident by scarring (beige) and charring (blackened) at the base and is surrounded by several species of short-lived perennials, as well as a few long-lived perennials that persisted through the fire. Photograph taken by Felicia Chen, U.S. Geological Survey, October 24, 2013. A 30-Year Chronosequence of Burned Areas in Arizona— Effects of Wildfires on Vegetation in Sonoran Desert Tortoise (Gopherus morafkai) Habitats By Daniel F. Shryock, Todd C. Esque, and Felicia C. Chen Prepared in cooperation with the Bureau of Land Management Open-File Report 2015-1060 U.S. Department of the Interior U.S. Geological Survey U.S. Department of the Interior SALLY JEWELL, Secretary U.S. Geological Survey Suzette M. Kimball, Acting Director U.S. Geological Survey, Reston, Virginia: 2015 For more information on the USGS—the Federal source for science about the Earth, its natural and living resources, natural hazards, and the environment—visit http://www.usgs.gov/ or call 1–888–ASK–USGS (1–888–275–8747). For an overview of USGS information products, including maps, imagery, and publications, visit http://www.usgs.gov/pubprod/. -
Alluvial Scrub Vegetation of Southern California, a Focus on the Santa Ana River Watershed in Orange, Riverside, and San Bernardino Counties, California
Alluvial Scrub Vegetation of Southern California, A Focus on the Santa Ana River Watershed In Orange, Riverside, and San Bernardino Counties, California By Jennifer Buck-Diaz and Julie M. Evens California Native Plant Society, Vegetation Program 2707 K Street, Suite 1 Sacramento, CA 95816 In cooperation with Arlee Montalvo Riverside-Corona Resource Conservation District (RCRCD) 4500 Glenwood Drive, Bldg. A Riverside, CA 92501 September 2011 TABLE OF CONTENTS Introduction ................................................................................................................................... 1 Background and Standards .......................................................................................................... 1 Table 1. Classification of Vegetation: Example Hierarchy .................................................... 2 Methods ........................................................................................................................................ 3 Study Area ................................................................................................................................3 Field Sampling ..........................................................................................................................3 Figure 1. Study area map illustrating new alluvial scrub surveys.......................................... 4 Figure 2. Study area map of both new and compiled alluvial scrub surveys. ....................... 5 Table 2. Environmental Variables ........................................................................................