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ACTA ENTOMOLOGICA MUSEI NATIONALIS PRAGAE Supplementum 11 (2007) AACTACTA EENTOMOLOGICANTOMOLOGICA MUSEI NATIONALIS PRAGAE Petr Bogusch, Annotated checklist Jakub Straka of the Aculeata (Hymenoptera) & Petr Kment of the Czech Republic (editors) and Slovakia Komentovaný seznam žahadlových blanokřídlých SUPPLEMENTUM 11 (Hymenoptera: Aculeata) České republiky 2007 a Slovenska www.nm.cz NATIONAL MUSEUM, P R A G U E ACTA ENTOMOLOGICA MUSEI NATIONALIS PRAGAE Supplementum 11 (2007) Annotated checklist of the Aculeata (Hymenoptera) of the Czech Republic and Slovakia Komentovaný seznam žahadlových blanokřídlých (Hymenoptera: Aculeata) České republiky a Slovenska Edited by: BOGUSCH P., STRAKA J. & KMENT P. ISSN 0231-8571 © Národní muzeum, Praha – 2007 ACTA ENTOMOLOGICA MUSEI NATIONALIS PRAGAE Supplementum 11 (2007) Chairman of the editorial board: Josef Jelínek (Czech Republic) Editor-in-chief: Petr Kment (Czech Republic) Technical editor: Martin Fikáček (Czech Republic) English language editors: Lubomír Masner (Canada) Nicole Černohorská (Czech Republic) Advisory board: Jan Bezděk (Czech Republic) Vítězslav Kubáň (Czech Republic) David S. Boukal (Norway) Jan Macek (Czech Republic) Freddy Bravo (Brazil) Wolfram Mey (Germany) Vladimir Gnezdilov (Russia) Carl W. Schaefer (USA) Jiří Hájek (Czech Republic) Aleš Smetana (Canada) Petr Kočárek (Czech Republic) Pavel Štys (Czech Republic) František Krampl (Czech Republic) Manuscripts should be sent to: AEMNP journal offi ce, Department of Entomology, National Museum, Kunratice 1, CZ-148 00 Praha 4, Czech Republic. E-mail: [email protected]. Published occasionally by the National Museum, Václavské náměstí 68, CZ-115 79 Praha 1, Czech Republic. Distributed by the Department of Entomology, National Museum, Praha. Journal web page : http://www.nm.cz/publikace/acta.php. Indexed in Entomology Abstracts, Zoological Records. Date of issue of this volume: November 16, 2007 ISSN 0231-8571 This issue was published with fi nancial support provided by: Charles University in Prague, Faculty of Science (grant MSM0021620828) Český literární fond Foundation, Prague Cover: Coelioxys alata Förster, 1853 (Apoidea: Megachilidae). Orig. P. Bogusch. Contents / Obsah Introduction / Úvod. ........................................................................................... 1 BOGUSCH P., STRAKA J. & KMENT P. Chrysidoidea: Bethylidae (hbitěnkovití). ......................................................... 21 MACEK J., STREJČEK J. & STRAKA J. Chrysidoidea: Chrysididae (zlatěnkovití). ....................................................... 41 TYRNER P. Chrysidoidea: Dryinidae (lapkovití) and Embolemidae (vejřenkovití). .......... 65 MACEK J. Vespoidea: Tiphiidae (trněnkovití). .................................................................. 85 BOGUSCH P. Vespoidea: Mutillidae (kodulkovití). ............................................................... 93 BOGUSCH P. Vespoidea: Sapygidae (drvenkovití). ............................................................. 105 BOGUSCH P. Vespoidea: Pompilidae (hrabalkovití). ........................................................... 111 STRAKA J. Vespoidea: Formicidae (mravencovití). ......................................................... 133 WERNER P. & WIEZIK M. Vespoidea: Scoliidae (žahalkovití). ................................................................ 165 BOGUSCH P. Vespoidea: Vespidae (vosovití). ..................................................................... 171 DVOŘÁK L. & STRAKA J. Apoidea: Spheciformes (kutilky). .................................................................. 191 VEPŘEK D. & STRAKA J. Apoidea: Apiformes (včely). .......................................................................... 241 STRAKA J., BOGUSCH P. & PŘIDAL A. Back issues / Nabídka předchozích svazků. .................................................. 300 Should be cited as follows / Doporučená citace: Book / Celý sborník: BOGUSCH P., STRAKA J. & KMENT P. (eds.) 2007: Annotated checklist of the Aculeata (Hymenoptera) of the Czech Republic and Slovakia. Komentovaný seznam žahadlových blanokřídlých (Hymenoptera: Aculeata) České republiky a Slovenska. Acta Entomologica Musei Nationalis Pragae, Supplementum 11: 1-300 (in English and Czech). Chapter / Kapitola: WERNER P. & WIEZIK M. 2007: Vespoidea: Formicidae (mravencovití). Pp. 133-164. In: BOGUSCH P., STRAKA J. & KMENT P. (eds.): Annotated checklist of the Aculeata (Hymenoptera) of the Czech Republic and Slovakia. Komentovaný seznam žahadlových blanokřídlých (Hymenoptera: Aculeata) České republiky a Slovenska. Acta Entomologica Musei Nationalis Pragae, Supplementum 11: 1-300 (in English and Czech). ACTA ENTOMOLOGICA MUSEI NATIONALIS PRAGAE Published 16.xi.2007 Supplementum 11, pp. 1-20 ISSN 0231-8571 Introduction / Úvod Petr BOGUSCH1), Jakub STRAKA2) & Petr KMENT3,2) 1) Department of Biology, University of Hradec Králové, Rokitanského 62, CZ-500 03 Hradec Králové, Czech Republic; e-mail: [email protected] 2) Charles University in Prague, Faculty of Science, Department of Zoology, Viničná 7, CZ-128 44 Praha 2, Czech Republic; e-mail: [email protected] 3) Department of Entomology, National Museum, Kunratice 1, CZ-148 00 Praha 4, Czech Republic; e-mail: [email protected] Abstract. This chapter is an introduction to the annotated checklist of the Acu- leata (Hymenoptera) of the Czech Republic and Slovakia. It summarizes the basic information about the phylogenetic position of Aculeata and relationships among particular families, delimitation of the studied area, brief account of the research history, and some introductory words about this checklist. The numbers of species of particular families known from the Czech Republic (1343 species in total), its historical parts (Bohemia – 1098 species, Moravia – 1279 species), and Slovakia (1453 species) are summarized in a table, and compared with former lists. Key words. Hymenoptera, Aculeata, checklist, phylogeny, zoogeography, history of research, Czech Republic, Bohemia, Moravia, Slovakia The order Hymenoptera is among the most Řád blanokřídlí (Hymenoptera) patří speciose groups of insect worldwide. This mezi nejpočetnější skupiny hmyzu nejen order contains ca. 13 % of all described insect celosvětově (13 % popsaných druhů hmyzu), species. In the Czech Republic and Slovakia, ale i v rámci České republiky a Slovenska approximately 7,500 species of Hymenoptera (okolo 7 500 druhů), kde o prvenství v počtu have been recorded so far. Similar amount of dosud známých druhů soutěží blanokřídlí species is known from our country only within pouze s brouky (Coleoptera) a dvoukřídlými the orders Coleoptera and Diptera. Although (Diptera). Ačkoliv se jedná o velice diverzi- Hymenoptera are very much diversifi ed, they fi kovanou skupinu, nepatří zástupci tohoto cannot be interpreted as a popular and well řádu mezi nejstudovanější hmyzí skupiny, studied group by the researchers, especially což se týká i Českých zemí a Slovenska. in our country. Only several groups have been To se samozřejmě netýká včely medonos- studied more extensively. The main focus is né (Apis mellifera Linnaeus, 1758), která the honeybee (Apis mellifera Linnaeus, 1758) patří mezi užitkové druhy hmyzu. K vše- BOGUSCH P., STRAKA J. & KMENT P. (eds.) 2007: Annotated checklist of the Aculeata (Hymenoptera) of the Czech Republic and Slovakia. Komentovaný seznam žahadlových blanokřídlých (Hymenoptera: Aculeata) České republiky a Slovenska. 300 pp. 2 BOGUSCH, STRAKA & KMENT: Introduction / Úvod classifi ed as an animal kept by people for obecně známým skupinám můžeme zařadit honey, wax, etc. Next, some social groups are i mravence (Formicidae), čmeláky (Bombus generally popular (e.g. ants (Formicidae) and spp.), a některé zástupce čeledi vosovitých bumblebees (Bombus spp.)) or unpopular (e.g. (Vespidae). Naproti tomu zástupci skupin hornets and wasps (Vespinae)) for people, and s četnými malými druhy, zejména nadčeledí thus better known than the others. The opposi- vejřitek (Proctotrupoidea), chalcidek (Chal- te situation is in numerous groups of small, cidoidea) a početných lumků (Ichneumono- usually parasitic hymenopterans, especially idea), byli u nás studováni jen zřídka. Zatímco superfamilies Proctotrupoidea and Chalci- v jiných zemích, třeba ve Spojených státech doidea, and the most numerous group within nebo v Japonsku, patří chalcidky a samotářské Hymenoptera – ichneumonids (Ichneumono- včely mezi oblíbené a často studované sku- idea). Although both solitary bees and proc- piny hmyzu, v našich podmínkách se těmito totrupoids have been intensively studied for skupinami ještě před zhruba deseti lety téměř many years, e.g. in the USA or Japan, Czech nikdo nezabýval. Dokonce i společenské vosy species were largely unknown only several (Vespinae) byly u nás vždy na okraji zájmu years ago. Even the social wasps (Vespinae) badatelů, a proto biologie i ekologické nároky have been studied only beside other related těchto (i pro člověka) významných zástupců groups and now we are only experiencing hmyzu zůstávají jen málo známé. ecological demands and bionomics both of common and rare representants of this for all people ‘well known’ group. Phylogeny of the Aculeata Fylogeneze žahadlových blanokřídlých (Aculeata) The Aculeata is a group of hymenopterous Žahadloví blanokřídlí (Aculeata) jsou insects, easily defi ned by their characteristic díky specifi ckému utváření kladélka dobře modifi cation of the ovipositor. It serves as a defi novanou skupinou hmyzu. Spíše než jako stinging apparatus connected with venomous aparát na kladení vajíček, slouží kladélko této gland, rather than an apparatus
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