Dinosaurs I. Triassic Biome
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APPLICATIONS of QUANTITATIVE METHODS and CHAOS THEORY in ICHNOLOGY for ANALYSIS of INVERTEBRATE BEHAVIOR and EVOLUTION by James
APPLICATIONS OF QUANTITATIVE METHODS AND CHAOS THEORY IN ICHNOLOGY FOR ANALYSIS OF INVERTEBRATE BEHAVIOR AND EVOLUTION by James Richard Woodson Lehane A dissertation submitted to the faculty of The University of Utah in partial fulfillment of the requirements for the degree of Doctor of Philosophy in Geology Department of Geology and Geophysics The University of Utah August 2014 Copyright © James Richard Woodson Lehane 2014 All Rights Reserved The University of Utah Graduate School STATEMENT OF DISSERTATION APPROVAL The dissertation of James Richard Woodson Lehane has been approved by the following supervisory committee members: Allan A. Ekdale , Chair May 5th, 2014 Date Approved Randall B. Irmis , Member June 6th, 2014 Date Approved Marjorie A. Chan , Member May 5th, 2014 Date Approved Elena A. Cherkaev , Member June 12th, 2014 Date Approved Leif Tapanila , Member June 6th, 2014 Date Approved and by John M. Bartley , Chair/Dean of the Department/College/School of Geology and Geophysics and by David B. Kieda, Dean of The Graduate School. ABSTRACT Trace fossils are the result of animal behaviors, such as burrowing and feeding, recorded in the rock record. Previous research has been mainly on the systematic description of trace fossils and their paleoenvironmental implications, not how animal behaviors have evolved. This study analyzes behavioral evolution using the quantification of a group of trace fossils, termed graphoglyptids. Graphoglyptids are deep marine trace fossils, typically found preserved as casts on the bottom of turbidite beds. The analytical techniques performed on the graphoglyptids include calculating fractal dimension, branching angles, and tortuosity, among other analyses, for each individual trace fossil and were performed on over 400 trace fossils, ranging from the Cambrian to the modem. -
The Lower Cretaceous Flora of the Gates Formation from Western Canada
The Lower Cretaceous Flora of the Gates Formation from Western Canada A Shesis Submitted to the College of Graduate Studies and Research in Partial Fulfillment of the Requirements for the Degree of Doctor of Philosophy in the Department of Geological Sciences Univ. of Saska., Saskatoon?SI(, Canada S7N 3E2 b~ Zhihui Wan @ Copyright Zhihui Mian, 1996. Al1 rights reserved. National Library Bibliothèque nationale 1*1 of Canada du Canada Acquisitions and Acquisitions et Bibliographic Services services bibliographiques 395 Wellington Street 395. rue Wellington Ottawa ON KlA ON4 Ottawa ON K1A ON4 Canada Canada The author has granted a non- L'auteur a accordé une licence non exclusive licence allowing the exclusive permettant à la National Libraxy of Canada to Bibliothèque nationale du Canada de reproduce, loan, distribute or sell reproduire, prêter, distribuer ou copies of this thesis in microfom, vendre des copies de cette thèse sous paper or electronic formats. la fome de microfiche/nlm, de reproduction sur papier ou sur foxmat électronique. The author retains ownership of the L'auteur conserve la propriété du copyright in this thesis. Neither the droit d'auteur qui protège cette thèse. thesis nor substantial extracts fiom it Ni la thèse ni des extraits substantiels may be printed or otherwise de celle-ci ne doivent être imprimés reproduced without the author's ou autrement reproduits sans son permission. autorisation. College of Graduate Studies and Research SUMMARY OF DISSERTATION Submitted in partial fulfillment of the requirernents for the DEGREE OF DOCTOR OF PHILOSOPHY ZHIRUI WAN Depart ment of Geological Sciences University of Saskatchewan Examining Commit tee: Dr. -
Diversity Variation, Distribution Pattern, and Evolutionary Implicat
+Model PALWOR-305; No. of Pages 20 ARTICLE IN PRESS Available online at www.sciencedirect.com ScienceDirect Palaeoworld xxx (2015) xxx–xxx A systematic overview of fossil osmundalean ferns in China: Diversity variation, distribution pattern, and evolutionary implications a,f,g b,c,∗ d b e Ning Tian , Yong-Dong Wang , Man Dong , Li-Qin Li , Zi-Kun Jiang a College of Palaeontology, Shenyang Normal University, Shenyang 110034, China b Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China c Key Laboratory of Economic Stratigraphy and Palaeogeography, Chinese Academy of Sciences, Nanjing 210008, China d College of Geosciences, Yangtze University, Wuhan 430100, China e Chinese Academy of Geological Sciences, Beijing 100037, China f State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China g Key Laboratory for Evolution of Past Life in Northeast Asia, Ministry of Land and Resources, Shenyang 110034, China Received 7 August 2014; received in revised form 9 December 2014; accepted 12 May 2015 Abstract The order Osmundales is a unique fern taxon with extensive fossil records in geological past. Diverse osmundalean fossils have been reported from China, ranging in age from the Late Palaeozoic to the Cenozoic. Most of them are based on leaf impressions/compressions, but permineralized rhizomes are also well documented. In this study, we provide a systematic overview on fossil osmundalean ferns in China with special references on diversity variations, distribution patterns, and evolutionary implications. Fossil evidence indicates that this fern lineage first appeared in the Late Palaeozoic in China. -
Contributions in BIOLOGY and GEOLOGY
MILWAUKEE PUBLIC MUSEUM Contributions In BIOLOGY and GEOLOGY Number 51 November 29, 1982 A Compendium of Fossil Marine Families J. John Sepkoski, Jr. MILWAUKEE PUBLIC MUSEUM Contributions in BIOLOGY and GEOLOGY Number 51 November 29, 1982 A COMPENDIUM OF FOSSIL MARINE FAMILIES J. JOHN SEPKOSKI, JR. Department of the Geophysical Sciences University of Chicago REVIEWERS FOR THIS PUBLICATION: Robert Gernant, University of Wisconsin-Milwaukee David M. Raup, Field Museum of Natural History Frederick R. Schram, San Diego Natural History Museum Peter M. Sheehan, Milwaukee Public Museum ISBN 0-893260-081-9 Milwaukee Public Museum Press Published by the Order of the Board of Trustees CONTENTS Abstract ---- ---------- -- - ----------------------- 2 Introduction -- --- -- ------ - - - ------- - ----------- - - - 2 Compendium ----------------------------- -- ------ 6 Protozoa ----- - ------- - - - -- -- - -------- - ------ - 6 Porifera------------- --- ---------------------- 9 Archaeocyatha -- - ------ - ------ - - -- ---------- - - - - 14 Coelenterata -- - -- --- -- - - -- - - - - -- - -- - -- - - -- -- - -- 17 Platyhelminthes - - -- - - - -- - - -- - -- - -- - -- -- --- - - - - - - 24 Rhynchocoela - ---- - - - - ---- --- ---- - - ----------- - 24 Priapulida ------ ---- - - - - -- - - -- - ------ - -- ------ 24 Nematoda - -- - --- --- -- - -- --- - -- --- ---- -- - - -- -- 24 Mollusca ------------- --- --------------- ------ 24 Sipunculida ---------- --- ------------ ---- -- --- - 46 Echiurida ------ - --- - - - - - --- --- - -- --- - -- - - --- -
Journal of Anatomy
Journal of Anatomy J. Anat. (2017) 230, pp325--336 doi: 10.1111/joa.12557 Reconstruction of body cavity volume in terrestrial tetrapods Marcus Clauss,1 Irina Nurutdinova,2 Carlo Meloro,3 Hanns-Christian Gunga,4 Duofang Jiang,2 Johannes Koller,2 Bernd Herkner,5 P. Martin Sander6 and Olaf Hellwich2 1Clinic for Zoo Animals, Exotic Pets and Wildlife, University of Zurich, Zurich, Switzerland 2Computer Vision and Remote Sensing, Technical University Berlin, Berlin, Germany 3Research Centre in Evolutionary Anthropology and Palaeoecology, Liverpool John Moores University, Liverpool, UK 4ChariteCrossOver - Institute of Physiology, Berlin, Germany 5Senckenberg Research Institute and Natural History Museum, Frankfurt (Main), Germany 6Steinmann Institute of Palaeontology, University of Bonn, Bonn, Germany Abstract Although it is generally assumed that herbivores have more voluminous body cavities due to larger digestive tracts required for the digestion of plant fiber, this concept has not been addressed quantitatively. We estimated the volume of the torso in 126 terrestrial tetrapods (synapsids including basal synapsids and mammals, and diapsids including birds, non-avian dinosaurs and reptiles) classified as either herbivore or carnivore in digital models of mounted skeletons, using the convex hull method. The difference in relative torso volume between diet types was significant in mammals, where relative torso volumes of herbivores were about twice as large as that of carnivores, supporting the general hypothesis. However, this effect was not evident in diapsids. This may either reflect the difficulty to reliably reconstruct mounted skeletons in non-avian dinosaurs, or a fundamental difference in the bauplan of different groups of tetrapods, for example due to differences in respiratory anatomy. -
1 Tommotiids from the Early Cambrian (Series 2, Stage 3) of Morocco and the Evolution Of
1 1 Tommotiids from the early Cambrian (Series 2, stage 3) of Morocco and the evolution of 2 the tannuolinid scleritome and setigerous shell structures in stem group brachiopods 3 4 by CHRISTIAN B. SKOVSTED1, SÉBASTIEN CLAUSEN2, J. JAVIER ÁLVARO3 and 5 DEBORAH PONLEVÉ2 6 7 1 Department of Palaeozoology, Swedish Museum of Natural History, P.O. Box 50007, SE- 8 104 05 Stockholm, Sweden, [email protected]. 9 2 UMR 8217 Géosystèmes, UFR Sciences de la Terre, Université Lille 1, 10 [email protected], [email protected]. 11 3 Centro de Astrobiología (CSIC/INTA), Ctra. de Torrejón a Ajalvir km 4, 28850 Torrejón de 12 Ardoz, Spain, [email protected] 13 14 Abstract: An assemblage of tannuolinid sclerites is described from the Amouslek Formation 15 (Souss Basin) of the Anti-Atlas Mountains in Morocco. The assemblage contains two species, 16 Tannuolina maroccana n. sp. which is represented by a small number of mitral and sellate 17 sclerites and Micrina sp., represented by a single mitral sclerite. Tannuolina maroccana 18 differs from other species of the genus in the presence of both bilaterally symmetrical and 19 strongly asymmetrical sellate sclerites. This observation suggests that the scleritome of 20 Tannuolina was more complex than previously thought and that this tommotiid may have held 21 a more basal position in the brachiopod stem group than previously assumed. The shell 22 structure of both T. maroccana and Micrina sp. is well preserved and exhibits two 23 fundamentally different sets of tubular structures, only one of which was likely to contain 24 shell penetrating setae. -
Lifestyle of the Octoradiate Eoandromeda in the Ediacaran
Lifestyle of the Octoradiate Eoandromeda in the Ediacaran Authors: Wang, Ye, Wang, Yue, Tang, Feng, Zhao, Mingsheng, and Liu, Pei Source: Paleontological Research, 24(1) : 1-13 Published By: The Palaeontological Society of Japan URL: https://doi.org/10.2517/2019PR001 BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access titles in the biological, ecological, and environmental sciences published by nonprofit societies, associations, museums, institutions, and presses. Your use of this PDF, the BioOne Complete website, and all posted and associated content indicates your acceptance of BioOne’s Terms of Use, available at www.bioone.org/terms-of-use. Usage of BioOne Complete content is strictly limited to personal, educational, and non - commercial use. Commercial inquiries or rights and permissions requests should be directed to the individual publisher as copyright holder. BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research. Downloaded From: https://bioone.org/journals/Paleontological-Research on 15 Feb 2020 Terms of Use: https://bioone.org/terms-of-use Access provided by Bing Search Engine Paleontological Research, vol. 24, no. 1, pp. 1–13, JanuaryEoandromeda 1, 2020 ’s Lifestyle 1 © by the Palaeontological Society of Japan doi:10.2517/2019PR001 Lifestyle of the Octoradiate Eoandromeda in the Ediacaran YE WANG1, YUE WANG1, FENG TANG2, MINGSHENG ZHAO3 and PEI LIU1 1Resource and Environmental Engineering College, Guizhou University, Huanx 550025, Guiyang, Guizhou, China (e-mail: [email protected]) 2Institute of Geology, Chinese Academy of Geological Sciences, 26 Baiwanzhuang Street 100037, Beijing, China 3College of Paleontology, Shenyang Normal Univeristy, 253 Huanhe N Street 110034, Shengyang, Liaoning, China Received May 14, 2018; Revised manuscript accepted January 26, 2019 Abstract. -
The Cambrian Substrate Revolution and the Early Evolution of Attachment in MARK Suspension-Feeding Echinoderms
Earth-Science Reviews 171 (2017) 478–491 Contents lists available at ScienceDirect Earth-Science Reviews journal homepage: www.elsevier.com/locate/earscirev The Cambrian Substrate Revolution and the early evolution of attachment in MARK suspension-feeding echinoderms ⁎ Samuel Zamoraa,b, , Bradley Delinec, J. Javier Álvarod, Imran A. Rahmane a Instituto Geológico y Minero de España, C/Manuel Lasala, 44, 9B, Zaragoza 50006, Spain b Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington DC 20013-7012, USA c University of West Georgia, Carrollton, GA, USA d Instituto de Geociencias (CSIC-UCM), c/José Antonio Novais 12, 28040 Madrid, Spain e Oxford University Museum of Natural History, Parks Road, Oxford OX1 3PW, UK ARTICLE INFO ABSTRACT Keywords: The Cambrian, characterized by the global appearance of diverse biomineralized metazoans in the fossil record Palaeoecology for the first time, represents a pivotal point in the history of life. This period also documents a major change in Evolution the nature of the sea floor: Neoproterozoic-type substrates stabilized by microbial mats were replaced by un- fl Sea oor consolidated soft substrates with a well-developed mixed layer. The effect of this transition on the ecology and Attachment evolution of benthic metazoans is termed the Cambrian Substrate Revolution (CSR), and this is thought to have impacted greatly on early suspension-feeding echinoderms in particular. According to this paradigm, most echinoderms rested directly on non-bioturbated soft substrates as sediment attachers and stickers during the Cambrian Epoch 2. As the substrates became increasingly disturbed by burrowing, forming a progressively thickening mixed layer, echinoderms developed new strategies for attaching to firm and hard substrates. -
Plated Cambrian Bilaterians Reveal the Earliest Stages of Echinoderm Evolution
Zamora, S., Rahman, I. A., & Smith, A. B. (2012). Plated Cambrian bilaterians reveal the earliest stages of echinoderm evolution. PLoS ONE, 7(6), [e38296]. https://doi.org/10.1371/journal.pone.0038296 Publisher's PDF, also known as Version of record Link to published version (if available): 10.1371/journal.pone.0038296 Link to publication record in Explore Bristol Research PDF-document University of Bristol - Explore Bristol Research General rights This document is made available in accordance with publisher policies. Please cite only the published version using the reference above. Full terms of use are available: http://www.bristol.ac.uk/red/research-policy/pure/user-guides/ebr-terms/ Plated Cambrian Bilaterians Reveal the Earliest Stages of Echinoderm Evolution Samuel Zamora1, Imran A. Rahman2, Andrew B. Smith1* 1 Department of Palaeontology, The Natural History Museum, London, United Kingdom, 2 School of Geography, Earth & Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, United Kingdom Abstract Echinoderms are unique in being pentaradiate, having diverged from the ancestral bilaterian body plan more radically than any other animal phylum. This transformation arises during ontogeny, as echinoderm larvae are initially bilateral, then pass through an asymmetric phase, before giving rise to the pentaradiate adult. Many fossil echinoderms are radial and a few are asymmetric, but until now none have been described that show the original bilaterian stage in echinoderm evolution. Here we report new fossils from the early middle Cambrian of southern Europe that are the first echinoderms with a fully bilaterian body plan as adults. Morphologically they are intermediate between two of the most basal classes, the Ctenocystoidea and Cincta. -
669-14373-2-LE Maquetación 1
AMEGHINIANA - 2013 - Tomo 50 (5): 493 – 508 ISSN 0002-7014 REVISIÓN DE LA PALEOFLORA DE LA FORMACIÓN NESTARES (JURÁSICO TEMPRANO), PROVINCIAS DEL NEUQUÉN Y RÍO NEGRO, ARGENTINA EDUARDO M. MOREL1, 2, DANIEL G. GANUZA1, ANALÍA E. ARTABE1, 3 Y LUIS A. SPALLETTI3,4 1División Paleobotánica, Facultad de Ciencias Naturales y Museo, Universidad Nacional de la Plata, Pasaje Teruggi s/nº, Paseo del Bosque, B1900FWA La Plata, Provincia de Buenos Aires, Argentina. [email protected]; [email protected]; [email protected] 2Comisión de Investigaciones Científicas de la Provincia de Buenos Aires (CIC), Argentina. 3Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Argentina. 4Centro de Investigaciones Geológicas, Avenida 1 nº 644, B1900TAC La Plata, Provincia de Buenos Aires, Argentina. [email protected] Resumen. En esta contribución se realizó la revisión de la paleoflora de la Formación Nestares, aflorante en ambas márgenes del Río Limay, en las inmediaciones del dique de Alicurá, sector noroccidental del Macizo Nordpatagónico, provincias de Río Negro y Neuquén, Argentina. Se midió un perfil sedimentológico, se identificaron cuatro estratos fosilíferos y su estudio permitió la determinación sistemática de 18 taxo- nes, 12 previamente descriptos para esta localidad= Neocalamites carrerei (Zeiller) Halle, Marattia muensteri (Goeppert) Zeiller, Goeppertella diazii Arrondo y Petriella, Kurtziana brandmayri Frenguelli, K. cacheutensis (Kurtz) Frenguelli, Taeniopteris sp., Otozamites albosaxatilis Herbst, O. ameghinoi Kurtz, O. bechei Brongniart, O. hislopii (Oldham) Feistmantel, Ptilophyllum acutifolium Morris en Grant, Elatocladus confertus (Oldham y Morris) Halle y seis nuevos= Equisetites frenguellii Orlando, Archangelskya protoloxsoma (Kurtz) Herbst, Sagenopteris nilssoniana (Brongniart) Ward, Nilssonia taeniopteroides Halle, O. -
A Stem Group Echinoderm from the Basal Cambrian of China and the Origins of Ambulacraria
ARTICLE https://doi.org/10.1038/s41467-019-09059-3 OPEN A stem group echinoderm from the basal Cambrian of China and the origins of Ambulacraria Timothy P. Topper 1,2,3, Junfeng Guo4, Sébastien Clausen 5, Christian B. Skovsted2 & Zhifei Zhang1 Deuterostomes are a morphologically disparate clade, encompassing the chordates (including vertebrates), the hemichordates (the vermiform enteropneusts and the colonial tube-dwelling pterobranchs) and the echinoderms (including starfish). Although deuter- 1234567890():,; ostomes are considered monophyletic, the inter-relationships between the three clades remain highly contentious. Here we report, Yanjiahella biscarpa, a bilaterally symmetrical, solitary metazoan from the early Cambrian (Fortunian) of China with a characteristic echinoderm-like plated theca, a muscular stalk reminiscent of the hemichordates and a pair of feeding appendages. Our phylogenetic analysis indicates that Y. biscarpa is a stem- echinoderm and not only is this species the oldest and most basal echinoderm, but it also predates all known hemichordates, and is among the earliest deuterostomes. This taxon confirms that echinoderms acquired plating before pentaradial symmetry and that their history is rooted in bilateral forms. Yanjiahella biscarpa shares morphological similarities with both enteropneusts and echinoderms, indicating that the enteropneust body plan is ancestral within hemichordates. 1 Shaanxi Key Laboratory of Early Life and Environments, State Key Laboratory of Continental Dynamics and Department of Geology, Northwest University, 710069 Xi’an, China. 2 Department of Palaeobiology, Swedish Museum of Natural History, Box 50007104 05, Stockholm, Sweden. 3 Department of Earth Sciences, Durham University, Durham DH1 3LE, UK. 4 School of Earth Science and Resources, Key Laboratory for the study of Focused Magmatism and Giant Ore Deposits, MLR, Chang’an University, 710054 Xi’an, China. -
Leppe & Philippe Moisan
CYCADALES Y CYCADEOIDALES DEL TRIÁSICO DELRevista BIOBÍO Chilena de Historia Natural475 76: 475-484, 2003 76: ¿¿-??, 2003 Nuevos registros de Cycadales y Cycadeoidales del Triásico superior del río Biobío, Chile New records of Upper Triassic Cycadales and Cycadeoidales of Biobío river, Chile MARCELO LEPPE & PHILIPPE MOISAN Departamento de Botánica, Facultad de Ciencias Naturales y Oceanográficas, Universidad de Concepción, Casilla 160-C, Concepción, Chile; e-mail: [email protected] RESUMEN Se entrega un aporte al conocimiento de las Cycadales y Cycadeoidales presentes en los sedimentos del Triásico superior de la Formación Santa Juana (Cárnico-Rético) de la Región del Biobío, Chile. Los grupos están representados por las especies Pseudoctenis longipinnata Anderson & Anderson, Pseudoctenis spatulata Du Toit y Pterophyllum azcaratei Herbst & Troncoso, y se propone una nueva especie Pseudoctenis truncata nov. sp. Las especies se encuentran junto a otros elementos típicos de las asociaciones paleoflorísticas del borde suroccidental del Gondwana. Palabras clave: Triásico superior, paleobotánica, Cycadales, Cycadeoidales. ABSTRACT A contribution to the knowledge of Cycadales and Cycadeoidales present in the Upper Triassic of the Santa Juana Formation (Carnian-Raetian) in the Bio-Bío Region of Chile is provided. The groups are represented by the species Pseudoctenis longipinnata Anderson & Anderson, Pseudoctenis spatulata Du Toit and Pterophyllum azcaratei Herbst & Troncoso. A new species Pseudoctenis truncata nov. sp. is described. They appear to be related to other typical elements of the paleofloristic assamblages from the south-occidental border of Gondwanaland. Key words: Upper Triassic, paleobotany, Cycadales, Cycadeoidales. INTRODUCCIÓN vasculares (Stewart & Rothwell 1993, Taylor & Taylor 1993). Sin embargo, se reconoce la exis- Frecuentemente se ha tratado a las Cycadeoida- tencia de una brecha morfológica entre ambos les (Bennettitales) y a las Cycadales como parte grupos, ya que a diferencia de las Medullosa- de las Cycadophyta.