J. Paleont., 75(3), 2001, pp. 706±720 Copyright q 2001, The Paleontological Society 0022-3360/01/0075-706$03.00 GRADUAL MIOCENE TO PLEISTOCENE UPLIFT OF THE CENTRAL AMERICAN ISTHMUS: EVIDENCE FROM TROPICAL AMERICAN TONNOIDEAN GASTROPODS ALAN G. BEU Institute of Geological and Nuclear Sciences, P O Box 30368, Lower Hutt, New Zealand, ,
[email protected]. ABSTRACTÐTonnoidean gastropods have planktotrophic larval lives of up to a year and are widely dispersed in ocean currents; the larvae maintain genetic exchange between adult populations. They therefore are expected to respond rapidly to new geographic barriers by either extinction or speciation. Fossil tonnoideans on the opposite coast of the Americas from their present-day range demonstrate that larval transport still was possible through Central America at the time of deposition of the fossils. Early Miocene occurrences of Cypraecassis tenuis (now eastern Paci®c) in the Caribbean probably indicate that constriction of the Central American seaway had commenced by Middle Miocene time. Pliocene larval transport through the seaway is demonstrated by Bursa rugosa (now eastern Paci®c) in Caribbean Miocene-latest Pliocene/Early Pleistocene rocks; Crossata ventricosa (eastern Paci®c) in late Pliocene rocks of Atlantic Panama; Distorsio clathrata (western Atlantic) in middle Pliocene rocks of Ecuador; Cymatium wiegmanni (eastern Paci®c) in middle Pliocene rocks of Atlantic Costa Rica; Sconsia sublaevigata (western Atlantic) in Pliocene rocks of Darien, Paci®c Panama; and Distorsio constricta (eastern Paci®c) in latest Pliocene-Early Pleistocene rocks of Atlantic Costa Rica. Continued Early or middle Pleistocene connections are demonstrated by Cymatium cingulatum (now Atlantic) in the Armuelles Formation of Paci®c Panama.